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An indicator-based assessment
111Impacts of Europe's changing climate — 2008 indicator-based assessment
5.7 Terrestrial ecosystems andbiodiversity
5.7.1 Introduction
Climate (change) is an important driving forcein the distribution and functioning of naturalsystems (Parmesan and Yohe, 2003). Europe's biodiversity (its species, habitats and ecosystems)has been modified repeatedly during past glacialand inter-glacial periods, with some speciesrecolonising the continent from ancient
refugia
.Today, ecosystems have an essential role inproviding services to humankind such as nutrientcycling, pest control, pollination, quality of life, andhydrological, atmospheric and climatic regulation(Díaz
et al.
 , 2006; IPCC, 2007). Impoverishmentof Europe's biodiversity may affect the deliveryof ecosystem services with potentially seriousconsequences (Lovejoy and Hannah, 2005).Maintaining and enhancing healthy ecosystems arean important element in climate change mitigationand adaptation actions.About 60 % of the world's known ecosystemsare currently used unsustainably (Reid
et al.
 ,
2005). In Europe, the richness and abundance of biodiversity is undergoing significant decline. Thisis in large part due to changes in land use andmanagement, which are resulting in degradationof (semi-)natural habitats, declines in traditionalagricultural and forest management on whichmany habitats depend, and now large-scale landabandonment. Urbanisation, industrialisation,modification of rivers and watercourses,fragmentation of habitats by infrastructureand growing pressure from public access tothe countryside for tourism and recreation arealso causing widespread biodiversity losses(Millennium Ecosystem Assessment, 2005).It is likely that these losses of biodiversity will beexacerbated by climate change. Projections suggestthat between one fifth and one third of Europeanspecies could be at increased risk of extinction ifglobal mean temperatures rise more than 2 to 3 °Cabove pre-industrial levels (Lovejoy and Hannah,2005; IPCC, 2007). A combination of climatechange and the drivers of change outlined abovewill reduce the adaptive capacity (and resilience)of many species, possibly resulting in differentecosystems and landscapes across Europe. Localand regional extinctions are likely (McKinney andLockwood, 1999). Species at greatest threat includespecialists, those at the top of the food chain, thosewith latitudinal and altitudinal restrictions, andthose with poor dispersal abilities.The European Commission, through its targetto 'halt the loss of biodiversity by 2010 — and beyond', is addressing observed and projecteddeclines in biodiversity and their consequences forhuman well-being. As part of this process, reducingthe impacts of other drivers of change will enhancethe ability of species to adapt to climate change(IPCC, 2007). But new areas for conservation arealso needed, together with measures to improveconnectivity, thus facilitating species movement infragmented landscapes. As such, the robustness ofthe European ecological network of Natura 2000sites should be strengthened, including throughmore widespread implementation of Article 10of the Habitats Directive (which relates to thenetwork's coherence).This section outlines the impacts of climatechange on biodiversity by showing both observedand projected changes in the distribution andphenology (changes in the timing of seasonalevents) of plants and animals, and the implicationsfor communities.
 
An indicator-based assessment
112Impacts of Europe's changing climate — 2008 indicator-based assessment
5.7.2 Distribution of plant species
Figure 5.31 Increase in species richness on
Swiss Alpine mountain summits
in 20th century
Note:
Endemic, cold-adapted species are declining as
pioneer species drive them out of their characteristic
niches due warming conditions. ×: 1900s; ◊: 1980s;●: 2003; open symbols indicate a (temporary)decrease in species number (Piz Trovat, PizLanguard).
Source:
Walther
et al.
, 2005.
Key messages
 
Climate change, in particular milder winters,is responsible for the observed northward and
uphill distribution shifts of many European plant
species. Mountain ecosystems in many parts of 
Europe are changing as pioneer species expanduphill and cold-adapted species are driven out
of their ranges.By the late 21st century, distributions of European plant species are projected to have
Key messages
shifted several hundred kilometres to the north,
forests are likely to have contracted in the
south and expanded in the north, and 60 % of mountain plant species may face extinction.The rate of change will exceed the ability of many species to adapt, especially as landscapefragmentation may restrict movement.
Relevance
The rate of climate change is likely to exceed theadaptive capacity of some wild plant species(IPCC, 2007), whilst others are expected to benefit from changing environmental conditions(Sobrino Vesperinas
et al.
 , 2001). Consequently, thecomposition of many plant communities is changingto the extent that completely new assemblagesare appearing. In addition, there is a parallelchange in plant distribution and the increasedthreat of extinction of species at the edge of theirgeographical and altitudinal ranges — particularlypoorly-dispersing endemics. The ecologicalimplications of these changes and the effects onthe services that these ecosystems provide are notalways clear. Together with the emergence of invasivenon-native species, these factors will have challengingconsequences for long-term biodiversity conservation(Gitay
et al.
 , 2002) and the ability of Europe to meet itstarget to halt biodiversity loss, not least in relation tothe favourable status of Natura 2000 sites.The adaptive capacity of species is linked to geneticdiversity and this too might change under climatechange; sensitive and valuable relic populations will be particularly affected.
Past trends
Warmer temperatures in the past 30 years havesignificantly influenced seasonal patterns acrossEurope. As evidenced during glacial and inter-glacialperiods, the predominant adaptive response oftemperature-sensitive plant species has been to shiftdistributions, resulting in northward and altitudinalmovements. One such climate-limited species is holly(
Ilex aquifolium
), which has expanded in southernScandinavia in a manner consistent and synchronouswith recorded regional climate changes, linkedin particular with increasing winter temperatures(Walther
et al.
 , 2005).Mountain ecosystems are particularly vulnerableto climate change (IPCC, 2007). There has been ageneral increase in mountain summit species inEurope since the Little Ice Age in the 18th century. InSwitzerland, for example, the uphill shift of Alpineplants showed an accelerating trend towards theend of the 20th century that is likely to be linked
0102030405060Number of species
  P  i  z   L  a  g  a  l  b   (   2   9  5  9   m  )  L  a  s   S  o  u  r  s   (   2   9  7  9   m  )  P  i  z   d  a  l  s   L  e  j   s   (   3   0 4  1   m  )  P  i  z    T  s  c  h  ü  f  f  e  r   (   3   1  2  3   m  )  M  u  n  t   P  e  r  s   (   3   2  0  7   m  )  P  i  z   L  a  n  g  u  a  r  d   (   3   2  6  2   m  )  P  i  z    T  r  o  v  a  t   (   3   1 4  6   m  )  P  i  z   M  i  n  o  r   (   3   0 4  9   m  )  P  i  z   C  h  a  t  s  c  h  e  d  e  r  s   (   2   9  8  6   m  )  P  i  z   A  l  v   (   2   9  7  5   m  )
 
An indicator-based assessment
113
Impacts of Europe's changing climate — 2008 indicator-based assessment
with the extraordinarily warm conditions of the1990s (Walther
et al.
 , 2005) (Figure 5.31). Evidencealso emerged of declines in cold-adapted speciesas warming conditions and pioneer species drovethese from their characteristic niches. Similarobservations are expected from current Europeanmonitoring programs (e.g. GLORIA) for whichresults will be available by the end of 2008. In theSwedish Scandes, the tree line of the Scots pine (
Pinussylvestris
) rose by 150–200 metres as warmer winterssignificantly lowered mortality and increased ratesof establishment. Observations from other continentsshow that uphill tree line migration is a globalphenomenon that could become a major threat to biodiversity in high mountains (Kullman, 2006; 2007;Pauli
et al.
 , 2007).
Projections
Projections indicate that, by the late 21st century,the potential range of many European plant speciesmay shift several hundred kilometres in a northerlydirection. This is several times faster than past ratesas estimated from the Quaternary record or fromhistoric data (Huntley, 2007). The distribution of treespecies is also likely change significantly, with forestsexpanding in the north and contracting in the south,and broadleaved species replacing native coniferousspecies in western and central Europe (IPCC, 2007).Modelling of late 21st century distributions of1 350 European plant species under a range ofscenarios led to the conclusion that more than halfwill be at the edge of their geographic and altitudinalranges and could become threatened by 2080, withhigh risks of extinction (Thuiller
et al.
 , 2005). Thegreatest changes are projected for endemic plantspecies in Mediterranean, Euro-Siberian and manymountain regions. Mountain communities may faceup to a 60 % loss of plant species under high emissionscenarios, reversing the 20th century trend outlinedabove (Thuiller
et al.
 , 2005; IPCC, 2007).Bakkenes
et al.
(2006) obtained similar results frommodelling stable areas of plant species distribution forthis century under different climate change scenarios(Map 5.30). This study suggests that 10–50 % of plantspecies in European countries are likely to disappear by 2100 from their current location in the absence ofclimate change mitigation. Again, species in southeastand southwest Europe are likely to be worst affected.This number will be higher if migration is restricteddue to continuing fragmentation or if there iscompetition with invasive species.
Map 5.30 Projected changes in number of plant species in 2050
Note:
Results for stable area per grid cell, using the EuroMove model with HadCM2 A2 climate scenario.
Source:
Based on Bakkenes
et al.
, 2006.
Number disappearing
Projected changes in plant species in 2050, compared to reference year 2000
0
0
Number appearing
1–2526–5051–100101–150151–200> 2001–2526–5051–100101–150151–200> 200
60°50°40°30°30°20°20°10°10°-10°-20°-30° 60°60°50°50°40°40°
0 500 1000 1500 km
60°50°40°30°30°20°20°10°10°-10°-20°-30° 60°60°50°50°40°40°
0 500 1000 1500 km
Number present (year 2000)
0
1–2526–5051–100101–150151–200> 200
60°50°40°30°30°20°20°10°10°-10°-20°-30° 60°60°50°50°40°40°
0 500 1000 1500 km
of 00

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