external ear, which collects sound waves and channels them inward; the middle ear, which convey sound vibration to the oval window; and the internal ear, which houses the receptors for hearing and equilibrium External ( Outer) ear
• The external or outer ear consist of the
auricle, external auditory canal, and eardrum. The auricle (pinna) is a flap of elastic cartilage shape like the flared and of a trumpet and covered by skin. The rim of the auricle is the helix; the inferior portion of the lobule. Ligaments and muscles attach the auricle to the head. • The external auditory canal (audit=hearing) is a curved tube about 2.5 cm long that lies in the temporal bone and leads from the auricle to the eardrum. The eardrum or tympanic membrane (tympan= a drum) is a thin, semitransparent partition between the external auditory canal and middle ear. • The eardrum is covered by epidermis and lined by simple cuboidal epithelium. Between the epithelial layers is connective tissue composed of collagen, elastic fibers, and fibroblasts. Near the exterior, the external auditory canal contains a few hairs and specialized sebaceous glands called ceruminous glands that secret earwax or cerumen. The combination of hairs and cerumen prevent dust and foreign objectives from entering the ear. • Cerumen usually dries up and falls out of the ear canal. Some people, however, produce a large amount of cerumen, which can become impacted and can muffle incoming sounds. Middle ear
• The middle is a small, air-filled cavity in the
temporal bone that is lined by epithelium. It is separated from the external ear by the eardrum and from the internal ear by a thin bony partition that contains 2 small membrane- covered opening: the oval window and the round window. • Extending across • the middle ear and attached to it by ligament are the 3 smallest bone in the body, the auditory ossicles, which are connected by synovial joints. • The bones, named for their shapes, are the malleus, incus, and stapes-commonly called the hammer, anvil and stirrup, respectively. The “handle” of the malleus attaches to the internal surface of the eardrum. • The head of the malleus articulates with the body of the incus. • The incus, the middle bone in the series, articulates with the head of the stapes. The base or footplate of the stapes fits into the oval window. Directly below the oval window is another opening, the round window, which is enclosed by a membrane, called the secondary tympanic membrane. • Besides the ligaments, 2 tiny skeletal muscles also attach to the ossicles. The tensor tympanimuscle, which is innervated by the mandibular branch of the trigeminal nerve or cranial nerve number 5, limits movement and increases tension on the eardrum to prevent damage to the inner ear from loud noises. • The stapedious muscle, which is innervated by the facial nerve (cranial nerve VII), is the smallest of all skeletal muscle. By dampening large vibrations if the stapes due to loud noises, it protect s the oval window but it also decrease the sensitivity of hearing. For this reasons, paralysis of the stapedious muscle is associated with the hyperacusia (abnormally sensitive hearing). • Because it takes a fraction of a second for the tensor tympanic and stapedious muscles to contract, they can protect the inner from prolonged loud noises, but not from brief ones such as a gun shot. The anterior wall of the middle ear contains an opening that leads directly into the auditory (pharyngotympanic) tube, commonly known as the Eustachian tube. • The auditory tubes, which consist of both bone and hyaline cartilage, connect the middle ear with the nasopharynx (upper portion of the throat). It is normally closed at its medial (pharyngeal) end. During swallowing and yawning, it opens, allowing air to enter or leave the middle ear until the pressure in the middle equals the atmospheric pressure. • When the pressures are balanced, the eardrum vibrates freely as sound waves strike it. If the pressure is not equalized, intense pain, hearing impairment, ringing in the ears, and vertigo • could develop. The auditory also is route whereby pathogens may travel from • the nose and throat to the middle ear.
Internal (inner) ear • The internal ear is also called the labyrinth because of its complicate d series of canals. Structurally, it consists of 2 main divisions; an outer bony labyrinth that encloses an inner membranous labyrinth. . The only labyrinth is a series of cavities in the temporal bone divided into 3 areas:
• 1. the semi circular
canals, • 2. the vestibule, both if which contain receptors for equilibrium, and • 3. The choclea, which contains receptors for hearing. • The bony labyrinth is lined with periosteum and contains perilymph. This fluid, which is chemically similar to cerebrospinal fluids, sorrounds the membranous labyrinth, a series of sacs and tubes inside the bony labyrinth and having the same general form. The membranous labyrinth is lined by epithelium and contains • endolymph. • The level of potassium ions in endolymph is unusually high for an extracellular fluid, and potassium ions play a role in the generation of auditory signals. The vestibule is the oval central portion of the bony labyrinth. The membranous labyrinth in the vitibule consists if 2 sacs called the utricle (little bag) and the saccule (little sac), which are connected by a small duct. • Projecting superiorly and posteriorly from the vestibule is the 3 bony semicircular canals, each of which lies at approximately right angles to the other 2. Based on their positions, they are named the anterior, posterior, and lateral semi circular canals. • The anterior and posterior semi circular canals are vertically oriented; the lateral one is horizontally oriented. At one end of each canal is a swollen enlargement called the ampulla (saclike duct). The portions of the membranous labyrinth that lie inside the bony semi circular canals are called the semi circular ducts. • These structures communicate with the utricle of the vestibule. The vestibular branch of the vestibulochoclear nerve (cranial nerve VIII) consists of ampullary, utricular, and saccular nervers. These nerves contain both first order sensory neurons and motor neurons that synapse with receptors for equilibrium. • The first order sensory neurons carry sensory information from the receptors, and the motor neurons carry feedback signals to the receptors, apparently to modify their sensitivity. Cell bodies Of the sensory neurons are located in the vestibular ganglia. Anterior to the vestibule is the choclea (snail shape) a bony spiral canal that resembles a snail’s shell and makes almost 3 turns around a central bony core called the modiolus. • Sections through the choclea reveal that it is divided into 3 channels. Together, the partitions that separate the channels are shaped like the letter Y. the stem of the Y is bony shell that protrudes into the canal; the wing of the Y are composed mainly of membranous labyrinth. The channel above the bony partition is the scala vestibule, which ends at the oval window; the channel below is the scala tympani, which ends at the round window. • The scala vestibule and scala tympani both contain perilymph and are completely separated, except for an opening at eh apex of the cochlea, the helicotrema. The cochlea adjoins the wall of the vestibule, into which the scala vestibule opens. The perilymph in the vestibule is continuous with that of the scala vestibule. The third channel is the cochlear duct or scala media. • The vestibular membrane separate the cochlear duct from the scala vestibule, and the basilar membrane separates the cocholear from the scala tympani. Resting on the basilar membrane is the spiral organ of corti. The spiral corti is a coiled sheet of epithelial cells, including supporting cells and about 16 thousands hair cells, which are the receptors for hearing. There are 2 groups of hair cells: • the inner hair cells are arranged in a single row whereas the outer hair called are arrange in 3 rows. At the apical tip of each hair cell is a hair bundle, consisting of 30-100 streocilia that extends into the endolymph of the cochlear duct. Despite their name, stereocilia are actually long hair like microvili arrange in several rows of graded height. • . At their ends, inner and outer cells synapse both with first-order sensory neurons and with motor neurons from the cochlear branch of the vestibulucocholear. Cells bodies of the sensory neurons are located in the spiral ganglion. Although outer hair cells outnumber them by 3-1 the inner hair cells synapse 90- 95% of the first order sensory neurons in the cochlear nerve that relay auditory information to the brain. • By contrast 90% of the motor neurons in the cochlear nerve synapse in the outer hair cells. Projecting over and in contact with hair cells of the spiral organ is the tectorial membrane, a flexible gelatinous membrane. THE END