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Alekseev Etal ProcGeoAssc105!1!1994 12

Alekseev Etal ProcGeoAssc105!1!1994 12

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Onthestratigraphicposition
of
theFamennianandTournaisianfossilvertebratebedsinAndreyevka,TulaRegion,CentralRussia
328
IU
S
UNES
0
I
PAlAEOZOIC"
ICROUEIlTEBIlATEs
AlexandrA.Alekseev,
*
OlegA.
Lebedev,
t
IgorS.
Barskov,
*
Maria
I.
Barskova,
t
Lyudmila
I.
Kononova*andVeraA.Chizhova:j:
ALEKSEEV
,A.S.,
LEBEDEV
,O.A.,
BARSKOV
,
I.
S.,
BARSKOVA
,M.
I.,
KONONOVA
,
L.
I.
&
CHIZHOV
A,V.A.1994.OnthestratigraphicpositionoftheFamennianandTournaisianfossilvertebratebedsin
Andre
yevka,TulaRegion,CentralRussia.
ProceedingsoftheGeologist'sAssociation,
105,
41-
52.Thefindoftheearliesttetrapod
Tulerpetoncurtum
LebedevintheuniqueAndreyevka-2vertebratelocality(closetoAndreyevkavillage,TulaRegion,CentralRussia)madetheestablishmentofthepreciseageofthefossiliferousdepositsofthisandtheneighbouringsitesmostimportant.Thedatingwascarried
out
withthehelpoftheostracodandconodontstudies.Inadditiontothistetrapod,acanthodians,antiarchplacoderms,porolepiforms,osteolepiforms,dipnoans,struniiformsandactinopterygiansarefoundintheAndreyevka-Zlocality.
The
ostracodcomplexhereisofKhovanshchinaageandpointstoacorrelationwiththeStrunian(Fa2d(?)
ofthe
Franco-Belgiumbasin).ThegreaterpartoftheMalyovkaFormationandthelowerthirdoftheUpaFormationbelongtothe
Patrognathusvariabilis
zoneandtheconodontsfromtheANE-3and
ANE-6
sitesclosetoAndreyevka-2arecharacteristicofit.Thefishlistincludeschondrichthyans,acanthodians,osteolepiforms,rhizodontiformsandactinopterygians.TheageofthefishassemblageintheANE-4(Andreyevka-l)siteisbasedupontheostracodsofasinglespecies,
Glyptolichwinella
cf.
limbata
Posner,thatischaracteristicofthezone
R.latiorPs.venulosa-Sh,microphtalma
ofboththeMalyovkaandUpaformations.Thefishassemblageconsistsofdiploselachiids,dipnoans,osteolepidids,rhizodontidsandactinopterygians.
PalaeontologyDepartment,GeologyFaculty,MoscowStateLomonosovUniversity,119899,Moscow,Russia.
t
PalaeontologicalInstitute
of
theRussianAcademyofSciences,123
Prof
soyuznaya
St,
117647,Moscow,Russia.
:j:
All-RussianOilandGasScientificResearchInstitute,10DmitrovskyProyezd,125422,Moscow,Russia.
1.INTRODUCTION
ConsiderablerecentattentionhasbeenfocusedonthefaunisticchangesattheDevonian-Carboniferousboundary,whichmarkoneofthePalaeozoicmassextinctions(Barskov,Kononova
&
Alekseev,1985).Thisinterestisreflectedintheworksofthe
lUGS
InternationalWorking
Group
onthe
Devonian
Carboniferousboundaryanditschoiceofpossiblestratotypes(Paproth,Feist
&
Flajs,1991).Marineinvertebrateextinctionisfairlywellknown,butthechangeofmarineandbrackish-watervertebratefaunasis
rather
poorlyrecognized.
The
LateDevonian-EarlyCarboniferouspalaeontologicalvertebraterecordisstillsparse;thispaperaimstointroduceanimportantregion,theRussianPlatform,forthestudyoftheproblem.TheDevonian-CarboniferousboundarysectionswithinthecentralregionsoftheRussianPlatformareimportantastheyreflecttheenvironmentsofthevastshallow-waterepicontinentalseawithextremelyunstablesalinity;thisseawasonlyimpermanentlylinked
Proceedings
of
theGeologists'Association,
105,41-52.
withtheocean.Theenvironment,beingshallowwateringeneral,butwithsea-levelandsalinityoscillations,resultsinawidespectrumoffaciesatthe
Devonian
Carboniferousboundaryin
the
central
part
oftheplatform,knownastheMoscowsyneclise.
The
depositsofthisagearerarelycharacterizedbywellpreservedandreadilyidentifiablevertebrateremains.FewsuccessivelocalitieswithinFamenniantoTournaisiansectionsareknowntocontainrepresentativevertebratefaunas.
The
problemofthev
ertebrate
faunalsuccessionduringthistimeintervalisrenderedmorecomplicatedbecauseofvaryinghabitatandburial(facial)conditions.Nevertheless,somehorizonswithintheintervalunderdiscussioninCentralRussiacontainrichfossilvertebrateassemblages,whichmayhelpthestudyofthevertebratefaunalsuccessionaroundtheDevonian-Carboniferousboundary.AsdemonstratedbyLebedev,Janvier
&
Esin(1992),theEuramericancontinentalnon-marinevertebratecommunitiesofthelatestDevonianaredominatedby
0016-7878/94$07.00
©
1994Geologists'Association
 
42
A.A.
ALEKSE
EV
ETAL.
porolepiforms
(Holoptychius),
osteolepiforms
and
antiarch
s
(Bothriolepis
and
Remigolepis).
The
chondrichth
yan-acanthodian-actinopterygian
f
auna
of
thesame
ageisdifferent
and
isrepresentedin
the
oc
eanic
andcontinental
marine
faciesby
Phoeb
odus
'Cladodus'
-
Thrinacodus-Protacrodu
s-
'Acanthodes'<'Moythomasia'
associations.
The
lattersurvives
into
the
Carboniferou
s,whereas
the
former
becomesextinctby
theend
oftheDevonian.Only
some
osteolepiforms,e.g.
Megalichthys,
stayasa
subject
elem
ent
oftheconti
nental
non-marinevertebratecommunities
(Andrews
&
Westoll,1970;Janvier,
Termier
&
Termier
,1979).
The
ageofthelocalities
cont
ainingtheearliest
tetrap
odremainswas
subject
todebateinthemiddle
of
the
century
.
The
longest
known
is
the
argument
between
Jarvik
(1948)
and
Westoll
(1940,1943,1951)
on
the
ageof
the
beds
that
contain
the
remainsof
lchthyostega
and
Acanthostega
in
EastGreenland
(then
called
the
Remigolepis
Series).
It
was
demonstrated
recently(Bendix-Almgreen,1976;
Obruchev
,1977;
Friend,
Alexander-Marrack,Allen,
Nicholson
&
Yeats,1983)
that
theserocksareofindisputable
Upp
er
Devonian
age.
Some
oftherecentfinds
not
onlyconfirmt
etrapod
presence
inthe
Devonian
,
but
demonstrate
their
systematicvariability
and
widegeographicdistributionat
that
time
(Warren
&
Wak
efield,1972;
Ahlberg,
1991).Asaresult,
the
ageofeachnew
Devonian
tetrapod
localityhas
been
particularlycarefullyinvesti
gated.The
levelofthe
De
vonian-Carbonif
erou
s
boundary
ontheRussian
platform
waschangedrecently,
the
Ozerki-KhovanshchinaregionalstagebeingincludedintheDevonianas
thetopmost
unit.Accordingly
the
ageofthet
etrapod-bearing
localityisnowreg
arded
as
Devon
ian,r
ather
than
basal
Carboniferous,
asin
the
past.
The
data
obtained
fromthefishesunequivocallyindicateitsDev
onian
age.
Two
uniquefossil
vertebrate
localitiesofF
amennian
and
Tourn
aisianages
are
nowknowninthewest
ern
part
of
the
TufaRegion(CherepetRiverbasin,closetoSuvorovTown).
These
siteswerecalled'
Andrey
evka
-l'
and
'Andreyevka-2'by
Lebedev
(Lebedev,1986;
Vorob
yeva
&
Lebedev,
1986).
The
f
ormer
wasfoundbythefieldcrewoftheTula-KalugaGeologicalProspectingD
epartment
in1948.Theirmaterialinitiatedincreasedfieldresearch,carriedoutsuccessivelyin1984--6jointly
between
thePala
eon
otologicalInstituteofthe
Academy
ofSciences(Moscow)
and
thePala
eont
ology
Department,
GeologicalFaculty,MoscowStateUniversity,resultingindiscoveryof
the
Andre
yevka-2fossiliferoussite(Fig.1).
The
findoftheearliest
tetrapod
Tulerpetoncurtum
Lebedev(Lebede
v,1984)inthe
Andr
eyevka
-2localit
y
madethe
establi
shment
ofthepreciseageofthesedepositsmost
important.Dating
wascarriedoutwith
the
helpofthe
ostracod
and
conodont
studies,thatbecamepossibleon
the
basisofthelocalz
one
correlation
chart
sf
or
thesouthernpart
oftheMoscow
1
«m
c
~ ~
MOSCOW
o
~
Tuld
B
Fig.
I.
Locationoftheinvestigatedarea:(A)the
Europ
ean
part
ofRussiaandTulaRegionwithinit;(B)TulaRegionandthepositionoftheareainvestigatedwithinit;
(C)
thelocalarea.Blackcirclesrepresentthepositionoftheoutcropsstudied.
 
Table1.
Thedistributionofconodontsintherocksamples.Figuresrepres
ent
thenumberofspecimensNumberofspecimensAN
E-l
ANE-3ANE-6ANE-4ANE-5
--
---
ANE
ANEANEANEANE
ANE
ANE
ANE
ANE
Cono
donts1/193/13/23/364/14/25/15/2
Tota
l
"!1
p
z
Polygnathuscommuniscommunis
BransonetMehl325
m
--
--
-
-
Z
Bispat
hodu
saculeatusaculeatus
(BransonetMehl)2
-
I
-
-
1
-
-
4
3
B.stabilis
BransonetMehl
I
1215
P
-
-
-
-
-
Z
Patrognathuscrassus
KononovaetMigdisova1691
-
1
-
-
171
P
P.variabilis
Rhodes,AustinetDruce19
-
24
10
-
-
53
z
"
ispathodusaculeatusplumulus
(Rhodes,Austin30
-
22
-
135
-I
etDruce)
0
c:
B.aculeatusanteposicornis
(Scott)51
-
-
-
6
'"
.a.plumulus-Clydagnathusgilwernensis
4
--
1
-
-
-
5
z
p
Clydagnathus
aff,
ormistoni
Beinert,Klap
per
1
--
-
-
-
1
~
etSandberg
p
Z
Polygnathusparapetus
Druce32
-
42
-
-
11
"!1
Pandorinellinanota
KononovaetMigdisova
6
12721
-
-
37
0
til
aff.
~
P.
(?)
ambigua
KononovaetMigdisova1
-
-
-
23
r
:;:I
Spathognathoduspenescitulus
Rexroadet11
6
1
-
9
rn
Collinson
"
n
Polygnathus
aff.
inornatus
BransonetMehl
-
-
I
1
'"
.crassus-P.variabilis
66
c
til
Pseudopolygnathusdentilineatus
E.Branson11
~
Spathognat
hod
us
sp.nov.1818
P
Clydagnathusgilwernensis
Rhodes,Austinet
Druc
e77Ramiformelements96235518
-
8235341
Tota
lquantity159459
10
7421344
9
714Sampleweight(kg)
1.0
0.70.60.50.40.81.92.1
1.3
-4
W

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