- DYNASTIDAE DYNASTINAE

MARTIN HARDY COLLECTION (MAHC) -

Personal scientific publications:
Hardy M. 2003 Description d'une nouvelle espèce congolais du genre Cyphonistes Fabreries 28(2):53-56

Hardy M. 2003 Description of a new species of Dynastes

Besoiro 9:3-7

Dechambre R-P & Hardy M. 2004 Quatre nouvelles espèces de Stenocrates

Coléoptères 10(18):209-214

Ratcliffe B.C. & Hardy M. 2005 Collagenus dasysternus, a new Genus and Species of Dynastinae from Eastern
Venezuela with a key to the New World Genera of Pentodontini
The Coleopterists Bulletin 59(1):143-150

For trade - exchange of specimens please contact:
Martin HARDY
1910 rue de l'Alliance
Quebec, Quebec
CANADA G3E1W6

email: dynastinae@hotmail.com

List updated: November 2014

(P) = PARATYPES

CYCLOCEPHALINI
Ancognatha
atacazo Kirsch, 1885
castanea Erichson, 1847
erythrodera Blanchard, 1841
falsa Arrow, 1911
gracilis Endrödi, 1966
horrida Endrödi, 1967
humeralis Burmeister, 1847
hyltonscottae Martinez, 1965
jamesoni Murray, 1857
manca LeConte, 1866
quadripunctata Bates, 1888
scarabaeoides Erichson, 1847
sellata Arrow, 1911
ustulata Burmeister, 1847
vexans Ratcliffe, 1992 (P)
vulgaris Arrow, 1911
Arriguttia
brevissima Arrow, 1911
Aspidolea
bleuzeni Dechambre, 1992
clypeata Burmeister, 1847
cognata Höhne, 1922
collaris Endrödi, 1966
ecuadoriana

fuliginea Burmeister, 1847
gaudairethorei Endrödi, 1980
kuntzeni Höhne, 1922
helleri Höhne, 1922
laticeps Harold, 1869
notaticollis Höhne, 1922
pelioptera Burmeister, 1847
quadrata Endrödi, 1966
singularis Bates, 1888
suturalis Höhne, 1922
suturella Höhne, 1922
testacea Höhne, 1922
Augoderia
nitidula nitidula Burmeister, 1847
nitidula yungana Martinez, 1966
Chalepides
alliaceus Burmeister, 1847
barbatus barbatus Fabricius, 1781
comes Prell, 1936
dilatatus Mannerheim, 1829
fuliginosus Burmeister, 1847
hydrophiloides Burmeister, 1847
luridus Burmeister, 1847
narcisoi Martinez, 1978
paradytis Ponchel & Dechambre 2003
unicolor Endrödi, 1963
Cyclocephala
aequatoria Endrödi, 1963
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affinis Endrödi, 1966
alexi Ratcliffe & Delgado, 1991
almitana Dechambre, 1992
amazona amazona Linnaeus, 1767
amazona boliviensis Höhne, 1923
amazona signata Fabricius, 1781
amblyopsis Bates, 1888
arnaudi Dechambre, 1980
arrowiana Martinez, 1967
atricapilla Mannerheim, 1829
atripes Bates, 1888
batesi Delgado & Castaneda, 1994
bella Endrödi, 1969
berti Delgado, 1992
bicolor Laporte de Castelnau, 1840
bimaculata Dechambre, 1999
binotata Dechambre, 1999
borealis Arrow, 1911
boucheri Dechambre, 1997
boulardi Dechambre, 1979
brevis Höhne, 1923
brittoni Endrödi, 1964
camachicola Ohaus, 1910
carbonaria Arrow, 1911
cartwighti Endrödi, 1964
castanea Olivier, 1789
castaniella Bates, 1888
caussaneli Dechambre, 1999
cerea Burmeister, 1847
chalumeaui Martinez, 1978
colasi Endrödi, 1964
comata Bates, 1888
complanata Burmeister, 1847
concolor Burmeister, 1847
confusa Endrödi, 1966
conspicua Sharp, 1877
contracta Kirsch, 1873
cribrata Burmeister, 1847
cubana Chapin, 1932
curta Bates, 1888
deceptor Casey, 1911
dichroa Dechambre, 1992
dilatata Prell, 1934
diluta Erichson, 1847
discicollis Arrow, 1902
dicolor discolor Herbst, 1792
dispar Herbst, 1792
distincta Burmeister, 1847
durantonorum Dechambre, 1999
emarginata Endrödi, 1966
epistomalis Bates, 1888
ergastuli Dechambre, 1997
erotylina Arrow, 1914
fankhaeneli Endrödi, 1964
fasciolata Bates, 1888
figurata Burmeister, 1847
flavipennis Arrow, 1914
flavoscutellaris Höhne, 1923
forcipulata Howden & Endrödi, 1966
forsteri forsteri Endrödi, 1963
freudei Endrödi, 1963
freyi freyi Endrödi, 1964
fulgurata Burmeister, 1847

fulvipennis Burmeister, 1847
gabaldoni Martinez, 1980
gravis Bates, 1888
gregaria Heyne & Taschenberg, 1907
guaguarum Dechambre & Endrödi, 1984
guianae Endrödi, 1969
guttata Bates, 1888
halffteriana Martinez, 1968
hardyi Endrödi, 1975
herteli Endrödi, 1964
hirta LeConte, 1861
homlbergi Martinez, 1968
inca Endrödi, 1966
insulicola Arrow, 1937
isabellina Höhne, 1923
jalapensis Casey, 1915
kaszabi Endrödi, 1964
kechua Martinez, 1957
krombeini Endrödi, 1979
kuntzeniana Höhne, 1923
lachaumei Dechambre, 1992
laevis Arrow, 1937
laminata Burmeister, 1847
larssoni Endrödi, 1964
latericia Höhne, 1923
latipennis Arrow, 1911
ligyrina Bates, 1888
lizeri Martinez, 1964
literata Burmeister, 1847
longitarsis Dechambre, 1999
longula LeConte, 1863
lunulata Burmeister, 1847
lurida lurida Bland, 1863
lurida coahuilae Bates, 1888
macrophylla Erichson, 1847
maculiventris Höhne, 1923
maffafa grandis Burmeister, 1847
maffafa maffafa Burmeister, 1847
mannheimsi Endrödi, 1964
marginalis Kirsch, 1870
marginicollis Arrow, 1902
marylizae Ratcliffe, 2003
mechae Martinez, 1978 (P)
mecynotarsis Höhne, 1923
melanocephala Fabricius, 1775
metrica Steinheil, 1874
miamiensis Howden & Endrödi 1966
minuchae Joly, 2003
minuta Burmeister, 1847
modesta Burmeister, 1847
morphoidina Prell, 1937
multiplex Casey, 1915
mutata Harold, 1869
nana Dechambre, 1999
nigerrima Bates, 1888
nigricollis Burmeister, 1847
nigritarsis Ratcliffe, 1992
nigrobasalis Höhne, 1923
nigropicta Dechambre & Endrödi, 1983
notata Illiger, 1806
ocellata Burmeister, 1847
ochracea Prell, 1937
ohausiana Höhne, 1923
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olivieri Arrow, 1911
ovulum Bates, 1888
pan Ratcliffe, 1992
paraguayensis Arrow, 1903
pasadenae pasadenae Casey, 1915
perconfusa Dechambre, 1992
perforata Arrow, 1911
pichincana Dechambre, 1992
picipes Olivier, 1789
picta Burmeister, 1847
pilosicollis Saylor, 1936
poncheli Dechambre, 2005
porioni Dechambre, 1979
prolongata Arrow, 1902
proxima Dechambre, 1997
pseudomelanocepha Dupuis, 1996
puberula LeConte, 1873
pugnax Arrow, 1914
pulchra Dechambre, 1999
puncticollis Endrödi, 1966
putrida Burmeister, 1847
quadripunctata Höhne, 1923
quercina Burmeister, 1847
rondoniana Ratcliffe, 1992
rubescens Bates, 1891
rufa Endrödi, 1967
rufescens Endrödi, 1967
rufovaria Arrow, 1911
rustica rustica Olivier, 1789
sanguinicollis Burmeister, 1847
santaritae Ratcliffe, 1992
sardadebiae Dechambre, 2005 (P)
setosa Burmeister, 1847
sexpunctata Laporte de Castel. 1840
signaticollis Burmeister, 1847
similis Dechambre, 1980
simulatrix Höhne, 1923
sinaloae Howden & Endrödi, 1966
sororia Bates, 1888
sparsa Arrow, 1902
spilopyga Erichson, 1847
striata haitiensis Endrödi, 1963
stictica Burmeister, 1847
subsignata Burmeister, 1847
suturalis Ohaus, 1911
sylviae Dechambre, 1995 (P)
tarsalis Dechambre, 1979
testacea Burmeister, 1847
tetrica Burmeister, 1847
tidula Dechambre, 1999
toulgoeti Dechambre, 1992
tridentata Fabricius, 1801
tronchonii Martinez, 1975
tucumana Brèthes, 1904
tutilina tutilina Burmeister, 1847
tylifera Höhne, 1923
undata Olivier, 1789
unidentata Endrödi, 1980
variabilis Burmeister, 1847
varians Burmeister, 1847
variolosa Burmeister, 1847
verticalis Burmeister, 1847
vestita Höhne, 1923

vidanoi Dechambre, 1992
vidua Endrödi, 1966
vincentiae Arrow, 1900
vinosa Arrow, 1937
weidneri Endrödi, 1964
williami Ratcliffe, 1992
zodion Ratcliffe, 1992
zurstrasseni Endrödi, 1964
Dyscinetus
dubius Olivier, 1789
gagates Burmeister, 1847
laevicollis Arrow, 1937
laevipunctatus laevipunctatus Bates, 1888
laevipunctatus minor Chapin, 1935
mendax Joly & Escalona, 2010
morator Fabricius, 1798
olivacea Höhne, 1923
plicatus Burmeister, 1847
picipes Burmeister, 1847
questeli Chalumeau, 1982
rugifrons Burmeister, 1847
subsericeus Burmeister, 1847
Erioscelis
emarginata Mannerheim, 1829
peruana Saylor,
proba Sharp, 1877
Harposcelis
paradoxus Burmeister, 1847
Mimeoma
acuta Arrow, 1902
maculata Burmeister, 1847
signatoides Höhne, 1923
Peltonotus
morio Burmeister, 1847
podocrassus Jameson & Wada, 2004
similis Arrow, 1931
Ruteloryctes
morio Fabricius, 1798
Stenocrates
agricola Dechambre & Hardy, 2004 (P)
ariasi Ratcliffe, 1978
batesi Dechambre, 1979
bicarinatus Robinson, 1948
bolivianus Dechambre, 1979
bollei Dechambre,
canuli Delgado & Castillo, 1991 (P)
carbo Prell, 1937
carinatus Endrödi, 1966
celatus Prell, 1937
clipeatus Endrödi, 1966
cultor cultor Burmeister, 1847
cultor inelegans Arrow, 1913
dubius Endrödi, 1966
duplicatus Endrödi, 1967
eniocanoi Ratcliffe & Cave, 2013
haackae Ratcliffe, 1978
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hardyi Dechambre, 1985
holomelanus Germar, 1824
howdeni Dechambre & Hardy, 2004 (P)
impeditus Dechambre & Hardy, 2004 (P)
inpai Ratcliffe, 1978
laborator Fabricius, 1775
lachaumei Dechambre, 1985
laevicollis Kirsch, 1870
lichyi Dechambre, 1979 (P)
ligneus Arrow, 1911
mahunkai Endrödi, 1973
minutus Endrödi, 1966
mollis Endrödi, 1966
nasutus Dechambre,
omissus Endrödi, 1966
pereirai Endrödi, 1964
popei Endrödi, 1971
porioni Dechambre, 1985
pseudoligneus Dechambre & Hardy, 2004
rabbanii Ratcliffe, 1978
rionegroensis Ratcliffe, 1978
rufipennis Fabricius, 1775
rugulosus Endrödi, 1966
varzeaensis Ratcliffe, 1978
Surutu
dytiscoides Martinez, 1955
seabrai Andretta & Martinez, 1956

ORYCTODERINI
Chalcocrates
borchmanni Endrödi, 1957
bretoni Dechambre, 2001
felschei Heller, 1903
Coenoryctoderus
candezei Lansberge, 1880
Hatamus
gressitti Endrödi, 1971
tarsalis Sharp, 1877
Neohyphus
kleinschmidti Fairmaire, 1881
philippinus Arrow, 1931
semivelutinus Fairmaire, 1883
Onychionyx
minor Dechambre, 2003
Oryctoderus
albertisi
godeffroyi anguliceps Prell, 1934
godeffroyi godeffroyi Fairm. 1877
latitarsis Boisduval 1835

AGAOCEPHALINI
Aegopsis
bolboceridus Thomson, 1860
curvicornis f.atra Sternberg, 1904

curvicornis trinidadensis Ster. 1904
westwoodi Thomson, 1860
westwood ab. rubricollis Sternberg, 1904
Agacephala
bicuspis Erichson, 1858
cornigera Serville, 1825
duponti Laporte de Castelnau, 1832
inermicollis Arrow, 1914
mannerheimi Laporte de Castel. 1832
urus Thomson, 1858
Brachysiderus
elyanae Dechambre, 2009
matogrossensis Ohaus, 1930
mielkeorum Grossi & Grossi, 2005
minicola Ohaus, 1930
quadri. quadrimaculatus Waterhouse, 1881
Democrates
burmeisteri Reiche, 1852
Gnathogolofa
bicolor Ohaus, 1910
Lycomedes
buckleyi Waterhouse, 1880
hirtipes Arrow, 1902
ohausi Arrow, 1908
reichei Brême, 1844
velutipes Arrow, 1902
Mitracephala
humboldti Thomson, 1859
Spodistes
armstrongi Dechambre, 1994
batesi Arrow, 1902
grandis Sternberg, 1903
mniszechi Thomson, 1860
monzoni Warner, 1992

PENTODONTINI
Aceratus
davus Burmeister, 1847
Adoryphorus
canei Carne, 1957
couloni Burmeister, 1847
Alissonotum
binodulum Fairmaire, 1891
cribratellum Fairmaire, 1893
elongatum Arrow, 1910
nitens Paulian, 1970
piceum krombeini Endrödi, 1976
piceum piceum Fabricius, 1775
yamayai Drumont, 2013
Anomalomorpha
anthracina Arrow, 1908
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Anoplognatho
dunnianus Rivers, 1889
Aphonus
brevicruris Cartwright, 1944
castaneus Melsheimer, 1846
densicaudus Casey, 1915
texanus Gill & Howden, 1985
tridentatus Say, 1823
variolosus LeConte, 1847
Barutus
hartmanni Ratcliffe, 1981
Bothynus
alvarengai Martinez, 1983 (P)
ascanius Kirby, 1818
complanus Burmeister, 1847
cribrarius Fairmaire, 1878
cunctator Mannerheim, 1829
cylindricus Arrow, 1937
dasypleurus Germar, 1824
deiphobus Burmeister, 1847
entellus Serville, 1825
exaratus Burmeister, 1847
herbivorus Arrow, 1937
herteli Endrödi, 1968
horridus Endrödi, 1968
laevipennis Arrow, 1937
laticifex Burmeister, 1847
medon Germar, 1824
nyx Ratcliffe, 2010
quadridens Taschenberg, 1870
stenelus Burmeister, 1847
striatellus Fairmaire, 1878
villiersi Endrödi, 1968
Calicnemis
atlanticus Mosconi, 1996
latreillei Laporte de Castelnau, 1832
sardiniensis Leo, 1985
Cheiroplatys
bifossus Blackburn, 1896
borealis Carne, 1957
excavatus Lea, 1917
inconspicuus MacLeay,
laevicollis Arrow, 1937
latipes Guérin-Méneville, 1830
nitidus Carne, 1957
solidus Carne, 1957
volsellus Carne, 1976
Corynophyllus
interocularis Lea, 1919
modestus Blackburn, 1888
Coscinocephalus
cribrifrons Schaeffer, 1906
Dalgopus
aethiopicus Dechambre, 2001

Dasygnathus
dejeani MacLeay, 1819
globosus Arrow, 1914
trituberculatus Blackburn, 1889
Diloboderus
abderus Sturm, 1826
Dipelicus
alveolatus Heller, 1897
cantori Hope, 1842
centratus Endrödi, 1969
daedalus Fabricius, 1792
deiphobus deiphobus (Sharp, 1873)
deiphobus robustus (Heller, 1897)
duplex Sharp, 1875
fastigatus Endrödi, 1969
hircus Fabricius, 1775
integriceps Fairmaire, 1877
lacordairei Sharp, 1873
monotuberculatus Miyake&Yamaya, 1997
optatus Sharp, 1875
quadratifer Heller, 1897
sinuosus Silvestre, 2006
tridentifer Silvestre, 2012
tuberosus Silvestre, 2012
Enarotadius
gressitti Endrödi, 1971
Eucopidocaulus
platypygus Prell, 1912
tridentipes Arrow, 1911
Eutheola
bidentata Burmeister, 1847
humilis humilis Burmeister, 1847
humilis rugiceps LeConte, 1856
latipennis Arrow, 1911
subglabra Schaeffer, 1909
Eutyctus
deserti Semenov, 1889
Heteroconus
major Dechambre, 1976
paradoxus Endrödi, 1968
Heteroligus
meles Billberg, 1815
peringueyi Arrow, 1911
trunc. truncaticeps Fair. 1879
Heteronychus
abyssinicus Jack, 1923
arator
annulatus Bates, 1891
arator Fabricius, 1775
arator australis Endrödi, 1961
arator rugifrons Fairmaire, 1871
bituberculatus Kolbe, 1900
citernii Paoli, 1934
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congonis Kolbe, 1900
curtulus Fairmaire, 1887
fossor Reiche, 1849
inoportunus Ferreira,
licas Klug, 1835
lioderes Redtenbacher, 1868
minutus Burmeister, 1847
mozambicus Péringuey, 1908
pauperatus Péringuey, 1901
plebejus Klug, 1832
rusticus niger Klug, 1855
rusticus rusticus Klug, 1832
tenuestriatus Fairmaire, 1893
Hylobothynus
arnaudi Dechambre, 1979
pontis Ratcliffe, 1982
Hyphoryctes
maculatus Blackburn, 1895
Lonchotus
lentus Burmeister, 1847
punctatissimus Arrow, 1908
rugosicollis Sternberg, 1910
Metanastes
picipes MacLeay, 1873
vulgivagus Olliff, 1889
Microryctes
monodon (Fairmaire, 1893)
silvestrei Drumont & al., 2011
Neodon
bidens Blackburn, 1896
laevipennis Blackburn, 1896
laevis Burmeister, 1847
pecuarius Reiche, 1860
simplex Carne, 1957
Novapus
adelaidae Blackburn, 1888
chararus Carne, 1957
laticollis Blackburn, 1890
nitidus Blackburn, 1895
obscurus MacLeay, 1871
rugosicollis Blackburn, 1890
Orizabus
batesi Prell, 1914
clunalis LeConte, 1856
cuernavacensis Delgado & Deloya,
isodonoides Fairmaire, 1878
pyriformis LeConte, 1848
Oryctomorphus
bimaculatus Guérin-Méneville, 1830
laevipennis Germain, 1855
maculicollis Guérin-Méneville, 1838
Orsilochus
kochi Ferreira, 1965

Oxygrylius
peninsularis Casey, 1915
ruginosus LeConte, 1856
Oxyligyrus
cayennensis Endrödi, 1972
nasutus Burmeister, 1847
peruanus Endrödi, 1966
zoilus Olivier, 1789
Papuana
angusta Arrow, 1914
biroi Endrödi, 1969
fortepunctata Heller, 1913
hardyi Endrödi, 1971
huebneri Fairmaire, 1879
japenensis Arrow, 1941
lansbergei badia Prell, 1912
lansbergei lansbergei Schauf. 1887
miyakei Yamaya, 2006
oryctoides Dechambre, 2005 (P)
philippinica Arrow, 1937
splendens Prell, 1912
szentivanyi Endrödi, 1971
trinodosa Prell, 1912
uniformis (Prell, 1934)
woodlarkiana woodlarkiana Montrouzier, 1855
woodlarkiana tenuistriata Aulmann, 1911
Paranodon
coquereli Fairmaire, 1871
Parapucaya
amazonica Prell, 1934
nodicollis Kirsch, 1873
Parisomorphus
descarpentriesi Lebis, 1961
niger Dechambre, 1976
Pentodon
algerinum algerinum Herbst, 1789
algerinum bispinifrons Reitt. 1894
algerinum dispar Baudi, 1870
algerinum indicum Endrödi, 1967
algerinum tchadense Paulian, 1954
bidens bidens Pallas, 1771
bidens punctatum Villers, 1789
bidens sulcifrons Küster, 1848
caminarum caminarium Fald., 1835
idiota idiota Herbst, 1789
quadridens bidentulum Fairm. 1887
quadridens quadridens Gebler, 1845
variolopunctatum Fairmaire, 1879
Pentodontoschema
caper Prell, 1936
Pericoptus
stupidus Sharp, 1878
Philoscaptus
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bonariensis Burmeister, 1847
Phyllognatus
excavatus Forster, 1771
burmeisteri Arrow, 1911
dionysius Fabricius, 1792
orion Olivier, 1789
Pimelopus
dubius Blackburn, 1885
fischeri Montrouzier, 1860
minor Carne, 1957
nothus Burmeister, 1847
Podalgus
cuniculus cuniculus Burm. 1847
cuniculus infantulus Seme. 1890
cuniculus reichei Paulian, 1954
Prionoryctes
breviusculus Arrow, 1911
camerunus Prell, 1914
capreolus Quedenfeldt, 1884
rufopiceus Arrow, 1911
Pseudohomonyx
borneensis Arrow, 1908
crassus Arrow, 1911
negros Miyake & Yamaya, 1997
samarensis Miyake & Yamaya, 1997
Pucaya
castanea Ohaus, 1910
pulchra Arrow, 1911
Pycnoschema
antinorii (Gestro, 1881)
cuspidatum Thomson, 1858
dechambrei Silvestre, 2000
diversum cavifrons Lesne, 1906
diversum diversum Péringuey, 1901
operculatum Thomson, 1858
scrofa Harold, 1880
simplicicolle Kolbe, 1905
stricticeps Fairmaire, 1893
subulatum Quedenfeldt, 1884
suillum Kolbe, 1905
Semanopterus
angustus Blackburn, 1888
leai Blackburn, 1897
rectangulus Blackburn, 1895
solidus Burmeister, 1847
subcostatus Laporte de Castelnau, 1840
Siralus
insularis Dechambre, 1997
Temnorrhynchus
baal baal Reiche et Sauley, 1856
bourgeoni Paulian, 1946
clypeatus Klug, 1855
coronatus coronatus Fabricius, 1781

coranatus diana Beauvois, 1805
coranatus natalensis Kolbe, 1887
descarpentriesi Paulian, 1946
flavipennis Arrow, 1937
raffrayi Fairmaire, 1887
repandus Burmeister, 1847
retusus Fabricius, 1781 (Afr.+Aust.)
rugatus Kolbe, 1905
stromsi Duvivier, 1891
truncatus Klug, 1832
Tomarus
bituberculatus bituberculatus Beau. 1805
bituberculatus maximus Arrow, 1913
burmeisteri Steinheil, 1872
cicatricosus Prell, 1937
cuniculus Fabricius, 1801
ebenus De Geer, 1774
fossor Latreille, 1833
gibbosus gibbosus De Geer, 1774
gibbosus obsoletus LeConte, 1856
gyas gyas Erichson, 1848
gyas amazonicus Arrow, 1914
laevicollis Bates, 1888
maimon Erichson, 1847
maternus Prell, 1937
nasutus Burmeister, 1847
neglectus LeConte, 1847
relictus Say, 1825
rosettae Endrödi, 1968
sallaei Bates, 1888
selanderi Cartwright, 1959
similis Endrödi, 1968
villosus Burmeister, 1847
Trissodon
curtum Burmeister, 1847
subopacum Arrow, 1914
tibiale Carne, 1957
Thronistes
rouxi Burmeister, 1847

ORYCTINI
Blabephorus
pinguis Fairmaire, 1898
Ceratoryctoderus
armatus Dechambre, 2001
candezei Snellen van Vollenhoven, 1866
Clyster
itys Olivier, 1789
retusus nakanei Mi. & Yamaya, 1999
rosaemariae Dupuis, 2002
trachypigus Prell, 1911
Coelosis
bicornis Leske, 1779
biloba Linnaeus, 1767
denticornis Arrow, 1937
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hippocrates Blanchard, 1841
inermis Sternberg, 1908
sylvanus Fabricius, 1775
Cyphonistes
arrowi Prell, 1912
brachti Prell, 1912
burmeisteri Dohrn, 1876
camurus Karsch, 1881
corniculatus Burmeister, 1847
gasanus Prell, 1912
glabricollis Burmeister, 1847
inermicollis Fairmaire, 1894
latiusculus Fairmaire, 1887
tuberculifrons Quedenfeldt, 1884
vallatus deserti Prell, 1912
vallatus guineensis Paulian, 1954
vallatus septentrionis Prell, 1912
vallatus vallatus Wiedeman, 1823
Dichodontus
coronatus Burmeister, 1847
crassus Silvestre, 2001
croesus Fabricius, 1801
grandis Ritsema, 1882
punctipennis Arrow, 1937
Dinoryctes
truncaticollis Fairmaire, 1887
Enema
endymion Chevrolat, 1843
pan Fabricius, 1775
Gibboryctes
gracilicornis (Prell, 1912)
szelenyii Endrödi, 1974
Heterogomphus
achilles Burmeister, 1847
aidoneus Perty, 1830
bispinosus Burmeister, 1847
bourcieri Guérin-Méneville, 1851
chevrolati Burmeister, 1847
coriaceus Prell, 1912
dilaticollis Burmeister, 1847
flohri Kolbe,
hirtus Prell, 1912
hopei Burmeister, 1847
laticollis Prell, 1912
mniszechi Thomson, 1859
monotuberculatus Dupuis & Dechambre,
2008
pauson Perty, 1830
pilosus Dechambre, 1998
porioni Dechambre, 1998
rugicollis ohausi Prell, 1912
rugicollis rugicollis Prell, 1912
schoenherri schoenherri Bur 1847
schoenherri whymperi Bates, 1891
telamon Prell, 1912
thoas Burmeister, 1847
ulysses ulysses Burmeister, 1847

Hoploryctoderus
caledonicus Prell, 1914
Licnostrategus
endymion Olivier, 1789
Megaceras
brevis Dechambre, 1999
celatus Dechambre, 1998
crassum Prell, 1914
jason Fabricius, 1775
morpheus Burmeister, 1847
philoctetes Olivier, 1789
porioni Dechambre, 1981
quadraticollis Dechambre, 1975
septentrionis angusticollis Ponchel, 1998
septentrionis crassicornis Dechambre, 1975
septentrionis septentrionis Bates, 1888
stuebeli Kirsch, 1885
Oryctes
agamennon arabicus Fairm. 1896
amberiensis pauliani Dech. 1986
anguliceps Fairmaire, 1901
augias Olivier, 1789
boas Fabricius, 1775
centaurus Sternberg, 1910
colonicus Coquerel, 1852
comoriensis intermedius Decham. 1986
congonis Endrödi, 1969
erebus Burmeister, 1847
forceps Dechambre, 1980
gigas gigas Laporte de Castelnau, 1840
gigas insulicola Dechambre, 1986
gnu Mohnike, 1874
gracilis Prell, 1934
latecavatus Fairmaire, 1891
mayottensis Dechambre, 1982
monoceros blucheaui Fairmaire, 1898
monoceros monoceros Olivier, 1789
nasicornis grypus Illiger, 1803
nasicornis hindenbergi Minck, 1915
nasicornis holdhausi Minck, 1914
nasicornis kuntzeni Minck, 1914
nasicornis laevigatus Heer, 1841
nasicornis nasicornis Linnaeus, 1758
nasicornis punctipennis Motschul. 1860
ohausi Minck, 1913
owariensis Beauvois, 1805
pyrrhus Burmeister, 1847
ranavalo Coquerel, 1852
rhinoceros Linnaeus, 1758
simiar dujardini Dechambre, 1987
simiar nesiotes Paulian, 1960
simiar simiar Coquerel, 1852
sjöstedti Kolbe, 1905
Podischnus
agenor Olivier, 1789
oberthuri Sternberg, 1907
sexdentatus Taschenberg, 1870

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Scapanes
affinis Dechambre, 1995
australis australis Boisduval, 1832
australis brevicornis Sternberg, 1908
australis grossepunctatus Stern. 1908
australis salomonensis Sternberg, 1908
Strategus
aenobarbus Fabricius, 1775
ajax Olivier, 1789
aloeus aloeus Linnaeus, 1758
aloeus julianus Burmeister, 1847
anachoreta Burmeister, 1847
antaeus Drury, 1773
centaurus Kolbe, 1906
cessus LeConte, 1866
jugurtha Burmeister, 1847
longichomperus Ratcliffe, 1976
mandibularis Sternberg, 1910
mormon Burmeister, 1847
oblongus Beauvois, 1805
sarpedon Burmeister, 1847
simson Linnaeus, 1758
splendens Beauvois, 1805
surinamensis hirtus Sternberg, 1910
surinamensis surinamensis B 1847
syphax Fabricius, 1775
talpa Fabricius, 1792
validus Fabricius, 1775
Trichogomphus
bronchus Jablonsky, 1785
dechambrei Dupuis, 2003
excavatus excavatus Mohnike, 1874
excavatus fairmairei Arrow, 1911
galeatus Dechambre, 1996
lunicollis Burmeister, 1847
lunicollis alcides Snellen van Voll. 1864
martabani Guérin-Méneville, 1834
mongol Arrow, 1908
simson selangorensis Endrödi, 1974
simson simson Snellen van Voll. 1864
vicinus Dechambre, 1995
zangi Sternberg, 1907
Xenoderus
janus Fabricius, 1801
Xyloryctes
corniger Bates, 1888
ensifer Bates, 1888
furcatus Burmeister, 1847
howdenorum Delgado & Najera, 1992
jamaicensis Drury, 1773
lobicollis Bates, 1888
splendidus Prell,
telephus Burmeister, 1847
teuthras Bates, 1888
thestalus intermedius Sternberg 1908
thestalus thestalus Bates, 1888

HEXODONTINI
Hexodon
latissinum Arrow, 1912
minutum Sternberg 1910
montandoni Buquet, 1840
patella Arrow, 1912
unicolor unicolor Olivier, 1789
unicostatum Arrow, 1912
Hyboschema
contractum Boheman, 1857

DYNASTINI
Allomyrina
davidis Deyrolle, 1878
dichotomus dichotomus Fab. 1775
dichotomus politus Prell, 1934
dichotomus septentrionis K. 1931
dichotomus takarai Kusui, 1976
dichoto. tsunobosonus Kôno, 1931
pfeifferi celebensis Silvestre, 1998
pfeifferi pfeifferi Redtenbacher 1867
Augosoma
centaurus Fabricius, 1775
hippocrates Milani, 1996
Beckius
beccari Gestro, 1876
Chalcosoma
atlas atlas Linnaeus, 1758
atlas keyboh Nagai, 2004
atlas mantetsu Nagai, 2004
atlas sintae Nagai, 2004
caucassus Fabricius, 1801
caucassus ab. belangeri Gué. 1834
caucassus ab. bidentata End. 1976
caucassus ab. chiron Olivier, 1789
caucassus ab. edentata Endr. 1976
caucassus ab. jansseni Beck, 1937
moellenkampi Kolbe, 1900
Dynastes
granti Horn, 1870
hercules baudrii Pinchon, 1976
hercules bleuzeni Sil. & Dech. 1995
hercules ecuatorianus Ohaus, 1913
hercules hercules Linnaeus, 1758
hercules lichyi Lachaume, 1985
hercules morishimai Nagai,
hercules occidentalis Lachau. 1985
hercules septentrionalis La. 1985
hyllus hyllus Chevrolat, 1843
hyllus moroni Nagai, 2005
maya Hardy, 2003 (P)
neptunus Quensel, 1905
satanas Moser, 1909
tityus Linnaeus, 1763

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Endebius
florensis florensis Lansberge, 1879
florensis wetarensis Fujii & Silvestre, 2012
Eupatorus
birmanicus Arrow, 1908
gracilicornis gracilicornis Arrow, 1908
gracilicornis kimioi Hirasawa, 1992
hardwickei Hope, 1831
siamensis Laporte de Castelnau, 1867
sukkiti Miyashita & Arnaud, 1997
Golofa
aegeon Drury, 773
argentinus Arrow, 1911
claviger claviger Linnaeus, 1771
cochlearis Ohaus, 1910
costaricensis Bates, 1888
eacus Burmeister, 1847
gaujoni Lachaume, 1985
globulicornis Dechambre, 1975 (P)
henrypitieri Arnaud & Joly, 2006
hirsuta Ratcliffe, 2003
imperialis Thomson, 1858
incas Hope, 1837
minutus Sternberg, 1910
obliquicornis Dechambre, 1975
pelagon Burmeister, 1847
pizarro Hope, 1837
porteri Hope, 1837
pusillus Arrow, 1911
solisi Ratcliffe, 2003
spatha Dechambre, 1989
tersander Burmeister, 1847
unicolor Bates, 1891
xiximeca Moron, 1995
Haploscapanes
barbarossa Fabricius, 1775
Megasoma
acteon Linnaeus, 1758
anubis Chevrolat, 1836
cedrosa Hardy, 1972
elephas elephas Fabricius, 1775
elephas iijimai Nagai, 2003
gyas gyas Herbst, 1785
gyas rumbucheri Fischer, 1968
janus janus Felsche, 1906
janus fujitai Nagai, 2003
joergenseni Bruch, 1910
mars Reiche, 1852
nogueirai Moron, 2005
occidentalis Bolivar et al. 1963
pachecoi Cartwright, 1963
punctulatus Cartwright, 1952
ramirezorum Silvestre & Arnaud, 2002
sleeperi Hardy, 1972
thersites LeConte, 1861
vogti Cartwright, 1963
Pachyoryctes
solidus Arrow, 1908

Xylotrupes
australicus Thomson, 1859
beckeri beckeri Schaufuss, 1885
beckeri intermedius Silvestre, 2004
faber Silvestre, 2002
gideon borneensis Minck, 1920
gideon gideon Linnaeus, 1767
gideon philippinensis Endrödi, 1957
gideon sondaicus Silvestre, 2002
gideon sumatrensis Minck, 1920
inarmatus Sternberg, 1906
lorquini lorquini Schaufuss, 1885
lumawigi Silvestre, 2002
macleayi lamachus Minck, 1920
macleayi macleayi Montrouzier, 1855
meridionalis Prell, 1914
mniszechi Thomson, 1859
pauliani Silvestre, 1997
pubescens pubescens Whaterhouse, 1841
pubescens beaudeti Silvestre, 2006
socrates socrates Schaufuss, 1864
socrates tonkinensis Minck, 1920
striatopunctatus Silvestre, 2003
ulysses baumeisteri Schaufuss, 1885
ulysses ulysses Guérin-Méneville 1830

PHILEURINI
Actinobolus
barbatus Endrödi, 1978 (P)
radians radians Westwood, 1841
Amblyoproctus
arianorum Dechambre, 2008
boondocksius Ratcliffe, 1988
chalumeaui Endrödi, 1977
piliger Perty, 1830
rugosus (Erichson, 1847
torulosus Kolbe, 1910
Archophanes
cratericollis Fairmaire, 1894
Archophileurus
chaconus Kolbe, 1916
cribrosus LeConte, 1854
vervex Burmeister, 1847
zischkai Endrödi, 1981
Ceratophileurus
lemoulti Ohaus, 1911
Cryptodus
caviceps Westwood, 1856
concentricus Lea, 1920
foveatus Lea, 1919
fraternus Fairmaire, 1877
obscurus MacLeay, 1871
passaloides Germar, 1848
rotundicollis Fairmaire, 1877
tasmannianus Westwood, 1841
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variolosus White, 1841
Eophileurus
borneensis Endrödi, 1976
chinen. chinensis Faldermann, 1835
chinensis irregularis Prell, 1913
chinensis okinawanus Nomu. 1964
dentatus Blackburn, 1895
felschei Prell, 1913
heyrovskyi Kral & Strnad, 1992
howdeni Yamaya & Muramoto, 2008
javanus Prell, 1913
malyi Endrödi, 1977
multidentatus Mi. & Yamaya, 1993
platypterus (Wiedeman, 1823)
sondaicus Prell, 1913
Goniophileurus
femoratus Burmeister, 1847
Hemiphileurus
dejeani Bates, 1888
depressus Fabricius, 1801
illatus LeConte, 1854
variolosus Burmeister, 1847
Homophileurus
amazonicus Kolbe, 1910
integer Burmeister, 1847
luedeckei Kolbe, 1910
planus Dupuis, 2013
quadrituberculatus Beauvois, 1805
tricuspis Prell, 1914

villaemagnae Dechambre, 1998
youngi Ratcliffe, 1988
Platyphileurus
felscheanus Ohaus, 1910
Prosphileurus
liberianus Dohrn, 1875
Pseudosyrichthus
clathratus Gerstäcker, 1866
senegalensis senegalensis Lap. 1840
Rhizoplatodes
castaneipennis foveiceps Fairmaire, 1893
Rhizoplatys
cribrarius Westwood, 1841
mainty Fairmaire, 1893
mucronatus cedrici Dechambre, 1983
mucronatus mucronatus Beau. 1805
mucronatus trituberculatus (Burmeister, 1847)
Syrichthodontus
cribratus Boheman,1857
Syrictes
syrichthus Fabricius, 1775
Trioplus
cylindricus Mannerheim, 1829

Macrocyphonistes
kolbeanus Ohaus, 1910
Moraguesia
champenoisi Dechambre, 2008
Palaeophileurus
brasiliensis Ratcliffe, 1988
fallax Dechambre, 1997
Paraphileurus
ventricosus Endrödi, 1978
Phileurus
angustatus Kolbe, 1910
carinatus carinatus Prell, 1914
carinatus declivis Prell, 1914
didymus Linnaeus, 1758
excavatus Prell, 1911
hospes Burmeister, 1847
hospitus Prell, 1934
lecourti Dechambre, 1998
limicauda Prell, 1912
toulgoeti Dechambre, 1996
truncatus Beauvois, 1805
valgus antillarum Prell, 1912
valgus meridionalis Kolbe, 1910
valgus valgus Olivier, 1789
verus Endrödi, 1978
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