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TWO NEW BAJOCIAN MICROCONCH OTOITID AMMONITES AND THEIR SIGNIFICANCE by ©. F, PARSONS [Ansnacr. Two new jan (Md Juan), mcoconck species Delnging to hp amma fmiy Otis, Trdbiceray(Trdicerar)cnht nv. a EO) die ova deere, tn re pated withthe [roable mctoconc pura, 7 Emde) marten Backman) and E(B) seas Backes rope {ely The raiaph dbo of theta subs the subtly Otte in southern Engand shows ‘hat the wo ne spe flan Important sap in our know oe sna Imprceaae (7) of nts mporinee, a tis te doubed ancestor (0 both ia and Popdontes ind pos aso Pied tnt Wi he ee rar rr a ie ea ton hs oops a ae Stattarph tne or delaitng the bam ofthe os Suro of the leactde Zone. The most important ‘xosares ofthe oats Subrone are diced andthe abu un ceed ro he south of France by Baur DURING the course of a recent stratigraphic study of the Bajocian rocks (Middle Jurassic) of southern England (Parsons 1974, pp. 164-171), large ammonite collec- tions have been made from some long-neglected horizons. In particular the ovals, (0rs0 called ‘Lower white iron-shot’ bed of Dundry Hill, near Bristol (Buckman and Wilson 1896, pp. 708-709, table iv), has yielded an interesting assemblage, which may be correlated with the ovalis Subzone of the laeviuscula Zone (Parsons 1974, . 169)—see Table | for the zonal scheme used here. The majority of ammonites ‘aE |, Zones and relevant sbzones of the Lover Bajctan subsiage(—~ Middle ‘Bajeian tra Aral 986, ater Parsons 174 Stephonoceras hamphrisamm mila (rots sae Wl Wit rea f Hypertoceras dts collected from this horizon (+3694), belong to the Witchella|Pelekodites dimorphic group. However, intensive coliecting revealed that a small proportion of the total auna consists of two, as yet undescribed, species belonging to the microconch sub- genera Ofoites and Trlobiticeras, A study of the previous literature and pre-existing ‘museum collections showed that these stratigraphically important species had been collected and recognized as distinct for some considerable time; it has merely taken a hundred years for them to be formally described. The small physical size of these two new mictoconch species, as well as their relative rarity in comparison with the abundant IWitchellia, goes some way towards explaining their absence in previous ‘works on the Bajocian Otoitidae (Westermann 1954). Urns Yo 3 Pa 178K A Lowen narocias W.()laciseda Soma ovatis wo PALAEONTOLOGY, VOLUME 20 AAs already noted, these two new species form only a minor part of the total ‘ammonite fauna of this agein England. However, at comparable horizons in southern, Europe the sonninid ammonites are less dominant, whilst the stephanoceratids are relatively more abundant. Its thus not surprising that this area of Europe has pro- duced the majority of previous records of these two new species, particularly the south of France, Portugal, and Sicily, With their wide geographic distribution and relatively restricted stratigraphic range, these ammonites are of great value for corre- lation purposes, particularly for between sonninid-cominated north-west Europe and the Tethyan region. These taxa are also of considerable phylogenetic importance, as they form a critical link between the early stephanoceratids of the concavum/discites Zones and the more abundant forms of the upper laeviuscula/sauzei Zones. As Emileia (Otoites) dowillei nov. and Trilebiticeras (Trilobiticeras) cricki nov. are ‘characteristic of the ovalis Subzone of the faeviuscula Zone, some discussion of the stratigraphy of this horizon in southern England and elsewhere in Europe is given, [Numbers preceded by these abbreviations refer to ammonites in the Following oletions: BMNH, The British Museum (NH), London. Cy Bisel City Museum, TGS. The lstute of Geological Sciences, London. M. The Fsoledes Mines Collection, Universit de Paris Sud, Pais GUM. The Oxford University Museum SM The Sedgwick Meum, Cambridae CP. Theauthors collection, Liverpool University. Under type spaces M = mucroconch and m ~ microconch SYSTEMATIC DESCRIPTIONS. Class crPHALOPODA Subelass AMMONOIDEA ‘Superfamily srerHanoceraracta Neumayr, 1875 Family ororipas Mascke, 1907 The classification of the Otoitidae follows Westermann (1964), who divided this family between two subfamilies, the Otoitinae and Sphaeroceratinae. Subfamily ororriNae Mascke, 1907 The generic classification within this subfamily follows Westermann (1964, pp. 51-34), except for the inclusion of Frogdenites, which on both morphological ‘and stratigraphie eriteria has been transferred from the Sphaeroceratinae. Genus Emileia Buckman, 1898 ‘Typ species. Biel (Ene) Broce 3. Sowerby, 1818), (M.)- Diagnosis. A group of macroconch and microconch, sphaeroconic ammonites, which fon the inner whorls bear ‘club’shaped primary ribs, terminated by blunt nodes, from which divide two or more rounded secondary ribs. The nominate subgenus ‘consists of medium to large-sized macroconch ammonites, with a terminal constric~ tion and flared-mouth border, and which vary from involute sphaerocones via cadicones to evolute platycones. PARSONS: BAJOCIAN OTOITID AMMONITES 10s Subgenus Oroites Mascke, 1907 ‘Type specter, Eniela (Oras) sacl (Orble, 1846), (1). Diagnosis. A group of moderately small sized, lappeted microconch ammonites, which are characteristically involute sphaerocones, with a pronounced contraction ‘of the body chamber. The ribbing style on the inner whorls is identical to its macro- ‘conch, Fmileias. str, that is ‘club’-shaped primary ribs, dividing into three to four finer secondaries. On its outer whorls, Oroties tends to develop sharp tubercles or spines at this point of division, whilst on the body chamber the secondary ribs tend to become inflated and coarsened, often with an alternation of strength on the last half whorl, with every second, third, or fourth rib being more inflated than its neighbour. Emileia (Otoites) dowvilei sp. nov. Pie 7,806 67, and 9 tx Sphacrocera uct & Orbigny; Dow no 'Orbign, pl, pI fig. 9a-6 Oroites suc: Rich and Roman non SOrbigny. p18. pl. 6, fi 8 Sphacrooras(Qtots sanz! &'Orbigny; Rens min Orb. p32, pl. fig 8 and Ba Brule soul SOrbigny; Languine non Orgy p.29% pl. Fi (Ooites sp. (0. sane! Roman now Orbiany): Lali, p17. (Oroites sp Dubat, p82 ly fis: 25-26 and 24. Droits ct sace (@ Orbigns) (Deus, 1885, pl 3 i. 9; Ruge-Perto,p 4 Oroiter sp. Gabilly abd Riou. 3. rile (Dias) po. Parsons, p68 nile (Otates sp 0. Morton, pp. 8-85, pl. 16, His. 5-6 ‘non 1984. Ototes fort p Westerman, pp. 103-106, pl. 3, gs 2-4 tex. 10 a2, Material. T we specimens 030m from the top of bed 2, Barn Batch Spinney (National Grid Reference S1557659, Dundey Hil, Bristol (Buckman snd Wiko® 1896, p. 689), BMNH. C73428-7429: fve nc spsimens frm ob mineum celetions, BMH. C7282, IGS. 25189, Ch 474, Cb. 4957-4958, "nd specinen +050 rom the base of bo & Bruton Railway quarry (ST 682345), Bruton, Somers (Richard 1916, . 495), BMH. C7452, atta of eight specimens Dimensions Holotype, Cb. 474, from Dundey Hil, and by marx fom the orais bed. I is complete, ‘rth the base of lappa just over hrc-quaters ofa whorl of body chambor a a maximum diameter oF 267 em. Umbilical —Numberof Wher! Whew! Diameter diameter primary ibs eight. ——_besdth o (Ws) a) (Hy) Gb) 208) HH) 1565) Die 6830) Lorie) 130465) Fist paratype, BMNH. €79823, with «quarter whor of body chamber. Te osrc9) GR) 12S) 17 04s) ” os) LBD 13s ‘Second paratype, BMH, C7228, 1 D835), 08 a) His) 1 039,30) = G6 (46) 09200) E53 Description. A small (c,23-2-7em diameter), sphaeroconic ammonite, with strongly ‘contracted body chamber, which extends for three-quarters to one whorl. The prorsitadiate primary ribs are short, blunt, and expand into ‘club’-shaped nodes at