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Feb 8, 2012 Alejandro, G.D. & S. Liede. (2003).

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(Rubiaceae-Ixoreae) with indications of geographical monophyletic unit and hybrids. Am. J. Bot., 96:686-706.

Mouly, A., Razafimandimbison, S.G., Florence, J., Jeremie, J., Bremer, B. (2009b) Paraphyly of Ixora and new tribal delimitation of Ixoreae (Rubiaceae): Inference from combined chloroplast (rps16, rbcL and trnT-F) sequence data. Ann. Missouri Bot. Gard., 96:146-160.