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P.

Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

OLIGOCENE - LOWERMOST MIOCENE FOSSIL FISH-FAUNA (TELEOSTEI) FROM
ROMANIAN EASTERN CARPATHIANS

Paul CONSTANTIN

Geological Institute of Romania
1 Caransebeş Street, 78 344 Bucharest, Romania, E-mail: paulct2004@yahoo.com

Abstract: An inventary of the localities with Oligocene - Lowermost Miocene fossil fish in the Romanian East Carpathians and their geological frame
is presented together with a list of the whole Oligocene - Lowermost Miocene fossil fish. From the 53 mentioned localities, different authors
determined 160 species (teleostean skeletons or skeletons imprints only) belonging to 76 genera and 46 teleostean families. Many of the mentioned
species would be the object of further reviews based on Recent fish osteology mainly.
Key words: Romanian Eastern Carpathians, Oligocene, Lowermost Miocene, fossil fish-fauna, Teleostei, localities.

1. INTRODUCTION 2. OLIGOCENE-LOWERMOST MIOCENE FOSSIL
FISH-FAUNA LOCALITIES OF ROMANIAN EAST
The first remarks on the presence of the fossil fish in CARPATHIANS. GEOLOGICAL FRAME.
Romania belong to D i m i t r i e C a n t e m i r , in the XVIII th
century. His famous paper "Descriptio Moldaviae", in the so- The localities from the Romanian East Carpathians (fig.1)
called chapter "About the mountains and the minerals of where Oligocene - Lowermost Miocene fossil fish were
Moldaviae", pointed on the first information concerning the mentioned, belong, from the structural point of view, to some
fossil fish remains from Tg.Ocna. It was he who mentioned formations involved both in post-tectogenetic covers (Post-
"...sometimes there are whole fishes in these holes1 which do Tectogenetic Cover of the Laramian Units), and in some
not differ by anything from those found in the surrounding nappes (Tarcău Nappe, Marginal Folds Nappe and
rivers" (fide Il i e , 1 9 4 2 ). Subcarpathians Nappe) the so-called Moldavidae. (tab.1).
The first scientific paper on the fossil fish from Romania The bearing fossil fish fauna formations belong as well to
is considered to belong to C o s mo v i c i ( 1 8 8 6 ) , being all the Oligocene - Lowermost Miocene lithofacies known in
published in Paris. A second paper was published in Romania the Romanian East Carpathians: Valea Caselor Facies (Inner
by C o s mo v i c i ( 1 8 8 7 ) , in which 2 new teleostean species Bituminous Facies), Fusaru-Pucioasa Facies, Slon Facies,
(Glyphysoma caprossoides and Syngnathus incompletus) are Mixed Facies, and Bituminous Facies with Kliwa Sandstone
described from Piatra Neamţ. Subsequently, the list of the (Outer Bituminous Facies) (tab.I).
teleostean fish fauna mentioned from the Romanian East
Carpathians and the localities which provided it, has been In these lithofacies, the fossil fish-fauna is generally
continuously enriched. provided by fine pelitic, bituminous shales, laminitic
limestones or bituminous marls, found on different
In the present paper, based both on the published data and stratigraphic levels in the Oligocene - Lowermost Miocene
on the examination of some collections from different column (tab.I), being predominant in its lower part
museums or on our own recent field observations, an (Rupelian).
inventary of the localities with Oligocene - Lowermost
Miocene fossil fish in the Romanian East Carpathians and The material mentioned in the present paper can be found
their geological frame (reviewed from latest data point of in the collections of the following museums: The Geological
view) is presented. Finally, a list of the whole Oligocene - Museum of the Geological Institute of Romania, Bucharest;
Lowermost Miocene fossil fish from Romania East The Museum of Natural History, Piatra Neamţ; The Museum
Carpathians is presented (even if later reviews could be of Natural History, Suceava; The Museum of Natural History,
made). The taxons are sistematically arranged according to Ploieşti; The Museum of Natural History "Grigore Antipa",
G r e e n wo o d e t a l . (1 9 6 6 ) classification. Bucharest; the museums belonging to the Paleontological
Department from the Faculties of Geology in Bucharest, Iassy
and Cluj-Napoca.
1
Salt mines - n. a.

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

Fig 1 - Oligocene-Lowermost Miocene fossil fish-fauna localities from Romanian East Carpathians

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

Table I - Oligocene-Lowermost Miocene fossil fish-fauna localities from Romanian East Carpathians and their geological frame

No2 LOCALITIES3 OUTCROPS STRUCTURAL LITHOFACIES STRATIGRAPHIC REFERENCES6
FRAME4 LEVEL5 - AGE
(1) SUSLĂNEŞTI - Mărlăuz; Post-Tectogenetic Cover Valea Caselor LDM – Rupelian [26], [27], [46], [47],
- Satului Valley; of the Laramian Units [51]7, [52], [57], [59],
- Măţău; [63], [69], [72], [75],
[78];
(2) VULCANA ? Tarcău Nappe Fusaru-Pucioasa (?) - Oligocene Materials hosted in
Geological Institute
of Romania;
(3) FIENI - 10 outcrops between Post-Tectogenetic Cover Valea Caselor LDM - Rupelian [4], [6], [22], [23], [24],
Ialomiţa Valley and of the Laramian Units Ty - Rupelian [25], [26], [27],
Ialomicioara Valley (F1 - [29] [37], [52], [62],
F10)-Constantin (2000)8 [69], [76], [77];
(4) - 2 outcrops on Tarcău Nappe Fusaru-Pucioasa PB – Uppermost [23], [25], [26], [29],
MOŢĂENI Ialomiţa Valley Rupelian -Chattian [52], [56];
UDS – Lowermost
Miocene
(5) BUCIUMENI-VALEA - Buciumeni; Post-Tectogenetic Cover Valea Caselor LDM – Rupelian [25], [26], [29];
LEURZII - Leurzii Valley; of the Laramian Units
(6) ROŞOIULUI VALLEY - Roşoiului Valley Tarcău Nappe Fusaru-Pucioasa PB – Uppermost [26], [29], [46]9, [47]10,
Rupelian -Chattian [56]11;
(7) ULMETU - an ancient oil well Tarcău Nappe Fusaru-Pucioasa PB – Uppermost [23], [25], [26], [27],
Rupelian-Chattian [29], [65];
(8) BEZDEADUL DE SUS - Coporadu Valley Post-Tectogenetic Cover Valea Caselor LDM – Rupelian [25], [26], [29];
of the Laramian Units
(9) URSEIULUI VALLEY - Urseiului Valley Tarcau Nappe Slon KVC - Reworked [25], [26], [29], 46]12,
fauna [47]13, [56]14;
ŞOTRILE- - Roşie Valley; Post-Tectogenetic Cover Valea Caselor LDM - Rupelian [25], [26], [29], [79];
(10) NISTOREŞTI15 - Cucuiatu Peak; of the Laramian Units Ty - Rupelian
- "Plaiul Cornului";
(11) COSMINEI VALLEY - 2 outcrops on Tarcău Nappe Fusaru-Pucioasa PB – Uppermost [23], [25], [26], [29],
Cosminele Valley; Rupelian -Chattian [74];
- Teiului Valley; LDS (?)or UDS (?)
(12) BUGHEA DE SUS - 1 outcrop between Tarcău Nappe Fusaru-Pucioasa LDS - Rupelian [25], [26], [29], [40],
Bughea Valley and [46], [47];
Slănic Valley

2
Number corresponding to the one from the sketch with Oligocene-Lowermost Miocene fossil fish-fauna localities from Romanian East
Carpathians (fig. 1);
3
According to our oppinion, localities mean the place bearing fossil fish-fauna, a place where can exist between 1 (most frequently) and
10 outcrops (as Fieni locality);
4
Reviewed acoording to the latest geological data;
5
The formation bearing the fossil fish-fauna; LEGEND: LDM – "The Lower Horizon of the Dysodile Shales with Menilites"; PB - Pucioasa
Beds; UDS - Upper Dysodilic Shales; LDS - Lower Dysodilic Shales; BM - Bituminous Marls; UM - Upper Menilites; KS - Kliwa
Sandstone (dysodilic shales type interbeds); FS - Fusaru Sandstone (dysodilic shales type interbeds); SBS - Slaty Bituminous Shales;
SH – "Shally Horizon"; Ty - Tylawa Laminitic Limestone; J – Jaslo Laminitic Limestone; KDy - Klippen bilt up of dysodilic shales
(Rupelian) in Vineţişu Formation (Uppermost Chattian - Lowermost Miocene); KVC – Klippen bilt up of "The Lower Horizon of the
Dysodile Shales with Menilites" (Valea Caselor Facies)(Rupelian) in Slon Facies (Uppermost Chattian - Lowermost Miocene);
6
The number from the brackets corresponds to the number of the paper mentioned in the references found in the end of the paper;
7
The underlined papers represent the first mention of the locality as bearing fossil fish fauna; when 2 papers are underlined, the first
mention refers to different outcrops from the same locality;
8
From the 10 outcrops (F1 - F10 - C o n s ta n t i n , 2 0 0 0 ) the F10 outcrop corresponds to Valea Caselor locality (P r ie m , 1 8 9 9 ); the F6
outcrop corresponds to Ţâţa locality (Paucă, 1934); the F4 and F7 outcrops were discovered by prof. D a n G r i go re s c u (pers. comm.);
the other 6 outcrops were discovered by Co n s ta n t i n ( 1 9 7 5 ; 2 0 0 0 ) ;
9
Mentioned by Ko t l a r c zy k J . , J e r z man s ka A . (1980) as Bezdead locality;
10
Mentioned by Ko t la r c z yk J . , J e r zma n s ka A . (1988) as Bezdead locality;
11
Mentioned by Pa u c ă ( 1 9 3 2 ) as Bezdead (Roşoiului Valley) locality;
12
Mentioned by Ko t la r c z yk J . , J e r zma n s ka A . (1980) as Bezdead locality;
13
Mentioned by Ko t la r c z yk J . , J e r zma n s ka A . (1988) as Bezdead locality;
14
Mentioned by Pa u c ă ( 1 9 3 2 ) as Bezdead (Urseiului Valley) locality;
15
Name introduced by C o n s ta n t i n ( 2 0 0 0 ) in order of grouping the outcrops: Roişe Valley (P ro te s c u , 1 9 3 8 ), Cucuiatu Peak (col.
Po povi c i Ha ţ eg) and Plaiul Cornului (D a n G r i go re s c u - pers. comm.);

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

No LOCALITIES OUTCROPS STRUCTURAL LITHOFACIES STRATIGRAPHIC REFERENCES
FRAME LEVEL - AGE
(13) HOMORÂCIU - many outcrops on Tarcău Nappe Fusaru-Pucioasa LDS - Rupelian [23], [24], [25], [26],
Teleajen Valley [27], [29], [41], [42],
[43], [44], [45],
[46],[47], [72];
(14) VITIOARA DE SUS ? Tarcău Nappe Bituminous Facies LDS (?) - Rupelian [48];
with Kliwa
Sandstone
(15) BUŞTENARI ? Tarcău Nappe Bituminous Facies (?) - Oligocene [48];
with Kliwa
Sandstone
(16) COADA MALULUI ? Tarcău Nappe Bituminous Facies (?) - Oligocene [24], [65];
with Kliwa
Sandstone
(17) OGRETIN - Păcurita Valley Tarcău Nappe Bituminous Facies LDS - Rupelian [26], [29];
with Kliwa
Sandstone
(18) TÂRLEŞTI - Seacă Valley Tarcău Nappe Bituminous Facies BM - Rupelian [23], [25], [26], [29],
with Kliwa [65];
Sandstone
(19) CĂTINA-CALVINI ? Tarcău Nappe Bituminous Facies (?) - Oligocene [48]
with Kliwa
Sandstone
(20) SIRIULUI VALLEY -1 outcrop at the Tarcău Nappe Slon Kdy - Reworked fauna [14], [23], [24], [25],
confluence of Siriu [26],[29];
Valley with Roşia
Valley
(21) GURA ARŢAGULUI - 2 outcrops on Tarcău Nappe Fusaru-Pucioasa SH - Rupelian [7], [23], [25], [26],
Buzău Valley [29];
(22) TEHEREU BROOK - Nowdays under Tarcău Nappe Fusaru-Pucioasa SH - Rupelian [36];
the Siriu Lake
(23) SIRIU LAKE - 1 outcrop on left Tarcău Nappe Fusaru-Pucioasa J - Chattian [10]16, [23], [24], [25],
border of Siriu Lake [26], [29];
(24) CAŞOCA BROOK - Prunecea Valley Tarcău Nappe Fusaru-Pucioasa SH - Rupelian [25], [26], [29];
(25) JEŢU - Buzău Valley Tarcău Nappe Mixed Facies LDS (?) or UDS (?) [25], [26], [29];
(26) PĂTÂRLAGELE ? Tarcău Nappe Bituminous Facies BM - Rupelian [23], [25], [26], [29],
with Kliwa [62];
Sandstone
(27) SIBICIULUI VALLEY - Sibiciului Valley Tarcău Nappe Bituminous Facies BM - Rupelian [23], [25], [26], [29],
with Kliwa [86];
Sandstone
(28) PATACU BROOK - Patacu Brook, Tarcău Nappe Fusaru-Pucioasa SH - Rupelian [25], [26], [29], [73];
tributary of Bâsca
Mare Valley
(29) BÂSCA MICĂ - Bâsca Mică Valley Tarcău Nappe Bituminous Facies BM - Rupelian [25], [26], [29], [73];
VALLEY with Kliwa
Sandstone
(30) COMANDĂU - Băsca Mare Valley Tarcău Nappe Fusaru-Pucioasa SH - Rupelian [8], [24], [25], [26],
- Kitereş Brook; Ty - Rupelian [29], [46], [47], [50],
[73], [81];
(31) CREMENII VALLEY - Cremenii Valley, tri- Tarcău Nappe Bituminous Facies BM - Rupelian [25], [26], [29], [39];
butary of Zăbala with Kliwa
Valley Sandstone
(32) ZĂBALA VALLEY - Murdanu Springs, Marginal Folds Nappe Bituminous Facies LDS - Rupelian [25], [26], [29], [50];
under Lăcauţi Peak with Kliwa
Sandstone
(33) NĂRUJA - an outcrop on the Subcarpathian Nappe Bituminous Facies UDS (?) - Lowermost [11], [23], [25], [26],
left bank of Zăbala with Kliwa Miocene [29], [68];
Valley Sandstone
(34) HERĂSTRĂU ? Subcarpathian Nappe Bituminous Facies UDS (?) - Lowermost [23], [26], [29], [68];
with Kliwa Miocene
Sandstone
(35) COZA VALLEY - Coza Valley Marginal Folds Nappe Bituminous Facies LDS - Rupelian [24], [25], [26], [29],
("Groapa cu pini" with Kliwa [88];
site) Sandstone
No LOCALITIES OUTCROPS STRUCTURAL LITHOFACIES STRATIGRAPHIC REFERENCES
FRAME LEVEL - AGE
(36) MĂGURA SPINEŞTI ? Marginal Folds Nappe Bituminous Facies (?) - Oligocene [23], [29], [66];
with Kliwa (?) - Lowermost
Sandstone Miocene
(37) BUCIAŞ VALLEY - Buciaş Valley Marginal Folds Nappe Bituminous Facies SBS – Lowermost [23]17, [25], [26], [29];
with Kliwa Rupelian
16
Mentioned by B r u s t u r, A lexand re s c u ( 1 9 9 8 ) as Buzău Valley locality;
17
Name introduced by B r u s t u r (pers. comm.); material hosted in the collections of Paleontological Department from the Faculties of
Geology in Bucharest (c o l . B r u s t u r );

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

Sandstone
(38) OITUZ VALLEY - an outcrop on the Marginal Folds Nappe Bituminous Facies UDS - Lowermost [23], [24], [25], [26],
left bank of Oituz with Kliwa Miocene [29], [30];
Valley Sandstone
(39) REPAT BROOK - Repat Brook Tarcău Nappe Slon Kdy - Reworked fauna [15], [25], [26], [29];
(40) SLĂNIC MOLDOVA - an right tributary of Marginal Folds Nappe Bituminous Facies LDS - Rupelian [23], [25], [29], [49],
Dobru Brook; with Kliwa [69];
- an left tributary of Sandstone
Slănic Brook;
(41) TÂRGU OCNA -2 outcrops on the Marginal Folds Nappe Bituminous Facies BM – Rupelian [23], [25], [26], [29],
right bank of Slanic with Kliwa LDS - Rupelian [38], [62];
Valley Sandstone
(42) SUSĂU BROOK - an outcrop on the Tarcău Nappe Slon Kdy - Reworked fauna [25], [26], [29];
right bank of
Susău Brook
(43) VĂSIEŞTI - Bucioc Brook Tarcău Nappe Bituminous Facies Ty (?) - Rupelian or [48];
with Kliwa J (?) - Chattian
Sandstone
(44) MOINESTI - Paltinisu Brook Tarcau Nappe Bituminous Facies (?) - Oligocene [23], [48];
with Kliwa
Sandstone
(45) LUCĂCEŞTI ? Tarcău Nappe (?) Bituminous Facies Ty (?) - Rupelian or [48];
with Kliwa J (?) - Chattian
Sandstone
(46) SOLONŢ ? Subcarpathian Nappe Bituminous Facies (?) - Oligocene [48];
with Kliwa
Sandstone
(47) STĂUNOIU BROOK -Stăunoiu Brook, left Marginal Folds Nappe Bituminous Facies UDS - Lowermost [13], [24], [26];
tributary of Calu with Kliwa Miocene
Brook Sandstone
(48) PIATRA NEAMŢ -Cernegura (48a); Marginal Folds Nappe Bituminous Facies BM – Rupelian [2], [3], [4], [6], [16],
-Agârcia (48b); with Kliwa LDS – Rupelian [17], [18], [19], [20],
-Pietricica (48c); Sandstone UDS (?) - Lowermost [23], [26], [27], [31],
-Cozla (48d); Miocene [32], [33], [34], [46],
[47], [55], [58], [67],
(69], [72], [82], [83],
[84], [85], [94]
(49) BICAZ -Mărceni Quarry Tarcău Nappe Fusaru-Pucioasa - FS – Chattian - [28];
(49a)
-Bicaz Valley - Ty - Rupelian - [7], [46], [47];
(Capşa) (49b)
(50) GURA HUMORULUI - an outcrop on right Tarcău Nappe Bituminous Facies BM (?) - Rupelian [1], [3], [9], [12], [46],
bank of Moldova with Kliwa LDS (?) - Rupelian [47], [62], [65], [69],
Valley Sandstone KS - Chattian [89], [90], [91], [96];
("Piatra Pinului" site)
(51) HUMORULUI - Dulcea Brook (a); Tarcău Nappe Bituminous Facies - BM+LDS - Rupelian - [12], [21], [23], [28],
VALLEY - Văcăreni Brook (b) with Kliwa - KS (Chattian)+ [80] [87];
Sandstone UDS (Lowermost - [12], [21], [23], [28],
- Rău Brook (c); Miocene) [87], [95];
- Ty - Rupelian - [28];
(52) STRUJINOASA - an left tributary of Tarcău Nappe Bituminous Facies LDS – Rupelian [3], [5], [27], [28];
BROOK Strujinoasa Brook with Kliwa
Sandstone
(53) STRAJA18 - Suceava Valley Marginal Folds Nappe Bituminous Facies Ty (?) – Rupelian [23];
with Kliwa
Sandstone

18
Mentioned by Co n s ta n t i n ( 1 9 9 6 ) as Gura Putnei locality; material hosted in the Geological Institute of Romania collections; (co l .
M i c u );

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

(11), (12), (13), (15), (19), (21), (23), (24),
(25) , (28), (29), (32), (33), (38), (40), (41),
(47), (48b), (51c);
Genus Opisthonema GILL, 1861
3. SYSTEMATIC PART Opisthonema persicum ARAMBOURG, 1966 -
(48d), (51b);
A synthetic list of the teleostean fish-fauna from the Opisthonema antethrissa CIOBANU, 1977 –
Romanian East Carpathians, and the localities which (48d), (51a, b);
provided it, is presented below: Genus Sardina ANTIPA, 1905
Osteichthyes (Teleostean fish)19 Sardina necteodosciobanensis CIOBANU, 1977 –
(48 a, c, d);
DIVISION I Genus Sardinella CUVIER & VALENCIEN., 1847
SUPERORDER Elopomorpha Sardinella denticulata CIOBANU, 1977 –
Order Anguilliformes (48c);
Suborder Anguilloidei Sardinella rata DANILCENKO, 1959 -
Family Muraenidae (48a, c, d), (50), (51a, b);
Genus Eomyrus AGASSIZ, 1844 Sardinella engrauliformis SMIRNOV - (48d),
Eomyrus aff. ventralis AGASSIZ, 1844 – (51c);
(48d)20, (51b); DIVISION III
Family Nemichthyidae
Genus Oligonemichthys CIOBANU, 1977 SUPERORDER Protacanthopterygii
Oligonemichthys photophorae CIOBANU, 1977 – Order Salmoniformes
(48d)21; Suborder Argentinoidei
SUPERORDER Clupeomorpha Family Argentinidae BONAPARTE, 1846
Order Clupeiformes Genus Glossanodon GUICHENOT, 1867
Suborder Clupeoidei Glossanodon musceli (PAUCĂ, 1929)25 - (1),
Family Clupeidae BONAPARTE, 1831 (3), (5), (8), (9), (12), (21), (13), (48c);
Genus Pelonulla GÜNTHER, 1868 Glossanodon inclinata (DANILCENKO, 1960)
Pelonulla grasionescui CIOBANU, 1977 – (35)26, (48d);
(48c); Glossanodon sp. - (3);
Genus Dussumieria CUVIER & VALENCIEN., 1847 Suborder Esocoidei
Dussumieria cf. elami ARAMBOURG, 1966 – Family Esocidae, GÜNTHER, 1866
(48d); Genus Esox LINNAEUS, 1758
Genus Etrumeus BLEEKER, 1853 Esox moldavicus TRELEA et al., 197727 - (50),
Etrumeus hafizi ARAMBOURG, 1943 - (48d); (51b);
Etrumeus sp. - (43); Suborder Stomiatoidei
Genus Alosa LINK, 1790 Family Gonostomatidae GILL, 1893
Alosa sculptata (WEILER, 1920) - (1)22, (13), Genus Scopeloides WETTSTEIN, 1886
(48a,c,d), (50), (51a, b); Scopeloides glarisianus (AGASSIZ, 1844) –
Alosa aff.sculptata (WEILER, 1920) - (13), (1)28, (20), (23), (48b,c), (50), (51a);
(48c, d); Scopeloides paucai CIOBANU, 1977 - (48c),
Alosa crassa SAUVAGE, 1873 - (13); (51a);
Alosa cf sagorensis (STEINDACHNER, 1863) Scopeloides sp. - (16), (23), (48b);
(38); Genus Idrissia ARAMBOURG, 1954
Alosa sp. - (1), (13), (38), (44), (45), (47); Idrissia carpathica JERZMANSKA, 1960 –
Genus Pomolobus RAFINESQUE, 1820 (50), (51a);
Pomolobus facilis DANILCENKO, 1960 – Idrissia carpiromanica CIOBANU, 1969 –
(48d), (50), (51a,); (48d);
Pomolobus aff. facilis DANILCENKO, 1960 – Family Photichthyidae WEITZMAN, 1974
(51b); Genus Vinciguerria GOOD & BEAN, 1895
Genus Clupea LINNAEUS, 1758 Vinciguerria talgiensis DANILCENKO, 1946 –
Clupea longimana (HECKEL, 1850)23 - (1), (3),
(4), (10), (26), (27), (28), (29), (30), (31), localities;
(35), (41), (42), (48a,c,d), (50), (51a, b); 25
This species was originally described by Pa u c ă ( 1 9 2 9a)
Clupea cf. longimana (HECKEL, 1850 - (13),
(50); from Suslăneşti as Nemachilus musceli n. sp.; mentioned also as
Clupea sardinites (HECKEL, 1850) - (1), (3), Nemachilus musceli PAUCĂ by Pau c ă from Moţăeni and
(4), (13), (21), (30), (47), (48a,c,d), (51b); Suslăneşti ( 1 9 3 2 ;1 9 3 3a ) an d b y J o ne t ( 1 9 5 2 ; 1 9 5 7 )
Clupea voinovi PAUCĂ 1929 -(1), (51b); from Homorâciu;
26
Clupea aff. arcuata KNER, 1863 - (30); Mentioned f r o m Coza Valley by Ş te fa n ( 1 9 8 8 ), and from
Clupea sp.24 - (1), (2), (3), (4), (5), (6), (8), (10), Piatra Neamţ (Coza outcrop) by C io ban u ( 1 9 7 7 ), and Tuc
(1989) as Proargentina inclinata DANILCENKO. Genus
19
The taxons are sistematically arranged according to Proargentina DANILCENKO is synonym with Glossanodon
G ree n wood et al . (1 9 6 6 ) classification. GUICHENOT (J e r zma n s ka , 1 9 6 7 );
27
20
The number in parantheses is identically with the one of the Needs reviews because in the Carpathians areas the species
locality which provided the species (the same number in fig.1 and initially determined as belonging to some fresh water genera (e.g.
tab. 1); Nemachilus, Leuciscus, Propercarina, etc.) proved lately to be
21 wrong determined (J e r z man s ka , 1 9 6 8 );
The underlined number in parantheses means that the holotype 28
was described from the mentioned locality (the underlined locality This species was originally described by Pa u c ă ( 1 9 2 9a ,
represents "locus typicus" for the respective species); 1 9 3 3a , c ) as Mrazecia mrazeci n. sp. and mentioned also by
22 Pa u c ă ( 1 9 3 4 c ) as Scopeloides mrazeci (PAUCĂ); mentioned
This species was originally described by Pa u c ă ( 1 9 2 9 b ) as
from Gura Humorului (Piatra Pinului) by Pa u c ă ( 1 9 3 4 b ) as
Clupea sculptata (WEILER);
23 Mrazecia mrazeci PAUCĂ and by T r e lea et a l . ( 1 9 7 3 ) and
Also mentioned as Meletta crenatta by Po pe s c u Vo i teşt i Vo i c u , I g na t ( 1 9 7 4 ) as Scopeloides mrazeci (PAUCĂ);
( 1 9 0 9 ), Clupea crenatta (HECKEL) by Pa u c ă ( 1 9 2 9 b ) or mentioned from Siriu Valley by B u c u r ( 1 9 6 7 ) as Scopelus
Clupea (Meletta) crenata (HECKEL) by P r o te s c u ( 1 9 3 8 ); mrazeci (PAUCĂ); mentioned from Humorului Valley (Dulce
24
Mentioned as Meletta sp. (isolated scales only) by Krejci-Graf Brook) by Sa r a i man ( 1 9 8 4 ) as Scopeloides mrazeci
and Weiler (1928) from Buştenari, Cătina-Calvini and Solonţ (PAUCĂ);

GEO-ECO-MARINA, 4/1999
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Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

(48d); Order Gadiformes
Vinciguerria merklini DANILCENKO, 1946 – Suborder Gadoidei
(35), (48d); Family Bregmacerotidae THOMPSON, 1840
Vinciguerria obscura DANILCENKO, 1946 – Genus Bregmaceros THOMPSON, 1840
(48d), (51b); Bregmaceros filamentosus (PRIEM, 1908) –
Vinciguerria distincta DANILCENKO, 1962 – (13)33;
(51a); Bregmaceros sp. - (3);
Vinciguerria macarovicii CIOBANU, 1969 – Family Gadidae RAFINESQUE, 1810
(35), (48c), (51a); Genus Palaeogadus RATH, 1858
Vinciguerria praeatenuata CIOBANU, 1977 – Palaeogadus athanasiui (PAUCĂ, 1929)34 - (1),
(35), (48d); (3), (5), (6), (8), (9), (13), (51a);
Family Sternoptychidae DUMERIL, 1806 Palaeogadus simionescui (SIMIONESCU, 1905)
Genus Maurolicus COCCO, 1838 (3), (12);
Maurolicus sp. - (48d)29, (50)30; Palaeogadus aff. leptosomus35 (KRAMBERG., 1879)
Genus Argyropelecus COCCO, 1829 (13);
Argyropelecus cosmovicii COSM. & PAUCĂ, 1943 Palaeogadus intergerinus DANILCENKO, 1947
(2), (4), (48d)31, (50), (51a); (48c);
Genus Polyipnus GÜNTHER, 1887 Palaeogadus atropatanus (BOGATSHOV, 1933)
Polyipnus anteasteroides CIOBANU, 1977 – (48c), (50), (51a);
(3), (48d); Palaeogadus abbreviatus (BOGATSHOV, 1933)
Family Astronesthidae RICHARDSON, 1843 (48d);
Genus Criptostomias GIBSS & WEITZ., 1965 Palaeogadus sp.36 - (3), (8), (13), (41), (50);
Criptostomias antiquus CIOBANU, 1977 – Genus Raniceps OKEN, 1817
(48d); Raniceps parcus DANILCENKO, 1960 - (48c);
Criptostomias aff antiquus CIOBANU, 1977 – Genus Palaeomolva DANILCENKO, 1947
(35); Palaeomolva tarchanica DANILCENKO, 1947
Order Scopeliformes (48d);
Suborder Myctophoidei Family Merlucciidae
Family Paralepididae RAFINESQUE,1810 Genus Merluccius RAFINESQUE, 1810
Genus Holosteus AGASSIZ, 1844 Merluccius ovalis CIOBANU, 1970 - (48d);
Holosteus mariae (MENNER, 1948)32 – Merluccius romanicus PAUCĂ, 1929 - (3), (4);
(1), (13) Merluccius sp. - (45);
Holosteus fieniensis CONSTANTIN, 2000 – Suborder Ophidioidei
(3); Family Ophidiidae JORDAN & EVERMANN, 1898
Family Myctophidae GILL, 1892 Genus Propteridium ARAMBOURG, 1966
Genus Eomyctophum DANILCENKO,1947 Propteridium profondae CIOBANU, 1970
Eomyctophum koraense DANILCENKO, 1947 (48c);
(48d); Genus Ophidion LINNAEUS, 1758
Eomyctophum menneri DANILCENKO, 1947 - Ophidion longipinatus (PAUCĂ, 1931)37 - (48);
(48d); SUPERORDER Atherinomorpha
Eomyctophum limicola DANILCENKO, 1960 – Order Beloniformes
(23), (48d); Suborder Exocoetoidae
Eomyctophum cozlae CIOBANU, 1969 – Family Hemiramphidae CUVIER, 1817
(48d); Genus Hemiramphus CUVIER, 1817
Genus Myctophum RAFINESQUE, 1810 Hemiramphus georgii JERZMANSKA, 1968
Myctophum weileri CIOBANU, 1977 - (48c); (3), (13)38;
Myctophum praepterotum CIOBANU, 1977 – Family Belonidae
(48c); Genus Belone CUVIER, 1817
Myctophum robustus CIOBANU, 1977 - (48c); Belone menilitica PAUCĂ, 1938 - (13), (18);
Myctophum antelaternatum CIOBANU, 1977 – SUPERORDER Acanthopterygii
(48c); Order Beryciformes
Genus Lampanyctus BONAPARTE, 1840 Suborder Berycoidei
Lampanyctus longaevus CIOBANU, 1977 – Family Trachichithyidae BERG, 1940
(48c); Genus Gephyroberyx BOULENGER, 1902
Genus Diaphus EIGENMANN & EIGEN., 1890
Diaphus duosensitivus CIOBANU, 1977 – 33
Mentioned as Bregmaceros prahovanus n. sp. by Jo ne t
(48c);
( 1 9 5 8 ) based on a poor preserved material. Reviewed by
SUPERORDER Paracanthopterygii
C o n s ta n t i n ( 1 9 9 7 ) based on photographic material only. The
Order Lophiiformes
original material needs reviews.
Suborder Antennarioidei 34
Family Onchocephalidae REGAN, 1912 This species was originally described from Suslăneşti as
Genus Onchocephalus HUBBS, 1948 Merluccius athanasiui n. sp. by Pa u c ă ( 1 9 2 9a , b ); was lately
Onchocephalus dudensis CIOBANU, 1977 – mentioned also as Nemopteryx athanasiui (PAUCĂ) from
(48d); Suslăneşti (Pa u c ă , 1 9 3 3 c ) , Roşoiu Valley (Pa u c ă , 1 9 3 2 ),
Urseiului Valley (Pa u c ă , 1 9 3 2 ), Homorâciu (J o ne t , 1 9 5 8 )
29
This species was originally described by C io ban u ( 1 9 7 7 ) and Humorului Valley (Dulce Brook) (Ş te f a n , H o ra i c u , 1 9 8 4 );
both as Polyipnus oligocenicus n. sp. and Polyipnus subnoviensis Mentioned also as Gobius elongatus n. sp. by S i m io ne s c u
JERZMANSKA; re-examined and replaced among the genus ( 1 9 0 4 ;1 9 0 5 ) ;
35
Maurolicus by Ba c i u , C o n s ta n t i n ( 1 9 9 8 ) and Ba c i u , Because of the succint description of a single specimen,
C o n s ta n t i n (in press); J e r z man s ka ( 1 9 6 8 ) considers it as "nomina nuda",
30
Mentioned by Voicu, Ignat (1974) as Polyipnus subnoviensis mentioning that all the specimens belonging to this species are
JERZMANSKA; poor preserved and described;
36
31
Also from Piatra Neamţ (Cozla outcrop) Pa u c ă ( 1 9 3 1a) Mentioned from Ţâţa (Fieni locality) and Piatra Neamţ by Pa u c ă
mentioned it as Sternoptyx prisca n. sp., reviewed by Ba c i u , ( 1 9 3 4 c ) as Nemopteryx sp.
37
C o n s ta n t i n ( 1 9 9 8 ); Described by Pa u c ă ( 1 9 3 1 ) as Ophidium (?) longipinatus
32
Mentioned from Suslăneşti by Pa u c ă ( 1 9 2 9a , b ; 1 9 3 3 c ) PAUCĂ;
38
and from Homorâciu by J o ne t ( 1 9 5 8 ) as Pronotacanthus Mentioned by Jone t ( 1 9 5 8 ) as Belone tenuis
Sahel Almae (DAVIS); KRAMBERGER and reviewed by J e r z man s ka ( 1 9 6 8 ) ;

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

Gephyroberyx aculeatus CIOBANU, 1976 – Genus Scorpaena LINNAEUS, 1758
(48d); Scorpaena pilarii KRAMBERGER, 1882 - (1),
Family Berycidae AGASSIZ, 1844 (50);
Genus Beryx CUVIER, 1829 Scorpaena boulei ARAMBOURG, 1928 - (13);
Beryx altus CIOBANU, 1977 - (48d); Family Triglidae JORDAN & EVERMANN, 1898
Beryx longus CIOBANU, 1977 - (48d); Genus Trigla LINNAEUS, 1758
Beryx prosus CIOBANU, 1976 - (48d); Trigla disodilica CIOBANU, 1976 - (48d);
Beryx ovatus CIOBANU, 1976 - (48d); Order Perciformes
Beryx sp. - (51b); Suborder Percoidei
Order Zeiformes Family Percidae JORDAN & EVERMANN, 1896
Suborder Zeoidei Genus Lates
Family Zeidae BONAPARTE, 183139 Lates sp. – (30), (48d);
Genus Zeus LINNAEUS, 1758 Genus and species undetermined (24)47;
Zeus hoernesi KRAMBERGER,1891 - (7); Family Serranidae RICHARDSON, 1846
Genus Zenopsis GILL, 1863 Genus Serranus CUVIER, 1817
Zenopsis clarus DANILCENKO, 1960 - (48b)40, Serranus budensis (HECKEL, 1856) - (1)48,
(52); (3), (4), (5), (9), (13), (27), (28), (30), (38),
Zenopsis sp. - (13)41; (39), (48b,d), (50);
Family Caproidae LÖWE, 1844 Serranus simionescui PAUCĂ, 1929 - (1), (3),
Genus Capros LACEPEDE, 1802 (5), (9), (13), (26), (50);
Capros radobojanus (KRAMBERGER, 1882)42 Serranus comparabilis DANILCENKO, 1960 –
(1), (3), (6), (13), (33), (48c,d), (50); (48d);
Capros longirostris (KRAMBERGER, 1882) – Serranus sp. - (3), (4), (5), (10), (11), (12),
(1), (3), (13), (48c,d)43; (17), (21), (31), (38), (40), (42), (50);
Capros sp. – (3), (13); Genus Properca SAUVAGE, 1880
Order Gasterosteiformes Properca sabbai PAUCĂ, 1929 - (1), (3), (48d);
Suborder Aulostomoidei Properca paucae JONET, 1958 - (13);
Family Aulostomidae LACEPEDE 1803 Properca sp. - (3), (5);
Genus Aulostomus LACEPEDE, 1803 Genus Morone MITCHILL, 1814
Aulostomus sp. - (3); Morone major (AGASSIZ, 1844) - (48d);
Family Fistulariidae BLAINVILLE, 1818 Morone sp. - (48d)49
Genus Fistularia LINNAEUS, 1758 Family Priacanthidae BERG, 1940
Fistularia aff köenegi AGASSIZ, 1842 - (13); Genus Priacanthus OKEN, 1817
Fistularia sp. (13); Priacanthus sturi (KRAMBERGER, 1880), (13);
Family Centriscidae RAFINESQUE, 1826 Priacanthus pietrensis CIOBANU,1970 ;(48d);
Genus Aeoliscus JORDAN & STARKS, 1902 Genus Pristigenys AGASSIZ, 1844
Aeoliscus heinrichi 44(HECKEL, 1850) – (30), Pristigenys spinosus (BLAINVILLE, 1818) - (48c);
(36), (48d), (51c); Family Dipterichyidae ARAMBOURG, 1966
Aeoliscus teleajensis45 (JONET, 1948) – (13); Genus Dipterichthys ARAMBOURG, 1966
Suborder Syngnathoidei Dipterichthys originis CIOBANU, 1976 - (48d);
Family Syngnathidae RAFINESQUE, 1810 Family Leiognathidae BERG, 1940
Genus Syngnathus LINNAEUS,1758 Genus Leiognathus LACEPEDE, 1803
Syngnathus incompletus COSMOVICI, 1887 - Leiognathus altapinnus (WEILER, 1955) - (48c);
(38), (48d), (50); Family Carangidae RAFINESQUE, 1810
Syngnathus cf. incompletus COSMOVICI, 1887 Genus Caranx LACEPEDE, 1802
(45); Caranx gracilis KRAMBERGER, 1882 - (48d);
Syngnathus altus DANILCENKO, 1960; (48d); Caranx petrodavae SIMIONESCU (48d);
Syngnathus incertus DANILCENKO, 1960; Caranx macoveii PAUCĂ, 1929 - (1);
(48d), (51b); Caranx aff. macoveii PAUCĂ, 1929 - (51a);
Syngnathus anteacum CIOBANU, 1976; (48d); Caranx sp. - (3);
Syngnathus sp.1 - (3)46 Family Pomadasyidae
Syngnathus sp.- (13), (33), (51b); Genus Lednevia DANILCENKO, 1960
Order Scorpaeniformes Lednevia oligocenica (SMIRNOV, 1936); (50),
Suborder Scorpaenoidei (51a);
Family Scorpaenidae RISSO, 1826 Suborder Trachinoidei
Genus Scorpaenoides LINNAEUS, 1758 Family Trachinidae RISSO, 1826
Scorpaenoides popovicii PRIEM, 1899 - (1), Genus Trachinus LINNAEUS, 1758
(3), (9), (13), (27), (29); Trachinus minutus (JONET,1958) - (13)50, (48c);
Suborder Ammodytoidei
39
Presently reviewed by Tyler, Baciu (pers. comm.); Family Ammodytidae BONAPARTE, 1846
40
Material from the Geological Museum (col. Cosmovici) wrong Genus Ammodytes LINNAEUS, 1758
named as Capros sp. (sample P56-9), reviewed by C o n s ta n t i n Ammodytes antipai PAUCĂ, 1929 - (1), (3),
( 2 0 0 0 ) and in the Museum "Grigore Antipa" - Bucharest (one (9), (13), (48);
undetermined specimen); Suborder Gobioidei
41 Family Gobiidae REGAN, 1911
Mentioned by J o ne t ( 1 9 5 8 ) as Zeus aff. Hörnesi Genus Gobius LINNAEUS, 1758
KRAMBERGER and reviewed by S w i d n i c k i ( 1 9 8 6 ) and Gobius pietricicai PAUCĂ, 1931
C o n s ta n t i n ( 2 0 0 0 ) ;
42
Mentioned from Piatra Neamţ also as Glyphiosoma 47
Specimen hosted in the collection of the Geologic Museum of
caprosoides n. sp., by Cosmovici (1887) and as Proantigonia the Geologic Institute of Romania; (c o l . Co mea gă ) and
caprosoides COSMOVICI by Simionescu (1904); considered by C o n s ta n t i n ( 2 0 0 0 ) to belong to the Percidae
43
Mentioned from Piatra Neamţ also as Proantigonia longirostris family;
KRAMBERGER by Simionescu (1904); 48
This species was originally described by Pa u c ă ( 1 9 2 9a ) as
44
Mentioned by Pa u c ă ( 1 9 4 2 ) and B ö h m ( 1 9 4 1 ) as Serranus elongatus and by Pa u c ă ( 1 9 2 9 b ) as Serranus
Amphisile henrichi HECKEL; oligocenicus.
45 49
Mentioned by Jo ne t ( 1 9 5 8 ) as Amphisile teleajensis JONET; Mentioned by S i m io ne s c u ( 1 9 0 4 ) as Labrax sp.;
46 50
Mentioned by C o n s ta n t i n ( 2 0 0 0 ) as a possible new This species was originally described by J o ne t ( 1 9 5 8 ) as
species; Megalolepis minutus n.sp.; reviewed by J e r z man s ka ( 1 9 6 8 ) ;

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
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P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

Suborder Scombroidei Suborder Pleuronectoidei58
Family Gempylidae GOOD & BEAN, 1895 Family Bothidae NORMAN, 1934
Genus Thyrsitoides FAWLER, 1929 Genus Scophthalmus RAFINESQUE, 1810
Thyrsitoides aff. zarathoustrae ARAMB., 1966 - Scophthalmus pietricicensis CIOBANU, 1978
(48d), (50), (51a, b); (48c);
Genus Hemithyrsites SAUVAGE, 1873 Scophthalmus stamatini (PAUCĂ, 1931)59
Hemithyrsites maicopicus DANILCENKO, 1960 -(50); (1), (13), (48), (50);
Genus Gempylus CUVIER & VALENCIEN., 1834 Suborder Soleoidei
Gempylus rumanus JONET, 1958 - (13); Family Soleidae NORMAN, 1934
Family Euzaphlegidae DANILCENKO, 1960 Genus Solea KLEIN. 1775
Genus Palimphyes AGASSIZ, 1844 Solea prisca CIOBANU, 1977 - (48d);
Palimphyes cf. elongatus (BLAINVILLE, 1818) - Order Tetraodontiformes
(13); Suborder Balistoidei
Palimphyes lanceolata (SIMIONESCU, 1904) Family Incertae Sedis60
(48d)51; Genus Ostracion
Palimphyes rebeli (PAUCĂ, 1929) - (1)52; Ostracion sp.61- (44);
Palimphyes pietschmanni (PAUCĂ, 1929) -
(1)53; Many of the species listed above have not been
Family Trichiuridae RAFINESQUE, 1810 reevaluated for a long time, needing revision of the original
Genus Lepidopus GOUAN, 1770 material to exclude any errors in identification.
Lepidopus caudatus (EUPHRASEN, 1788) -
(3), (40)54; Barbus sp. (P a uc ă , 1 9 3 3 ; S t o i c a , 1 9 4 4 ; J o n e t ,
Lepidopus glarisianus (BLAINVILLE, 1918); 1 9 5 8 ) and Scardinus sp. (P a uc ă , 1 9 3 3 ) described only as
(1), (13), (48a, c, d), (50), (51a, b); isolated scales, weren't listed.
Lepidopus cf. glarisianus (BLAINVILLE, 1918)
(30), (37); Dapolis macrurus (AGASSIZ) (mentioned as Smerdis
Lepidopus hungaricus BÖHM, 1941 - (30); macrurus AGASSIZ by K r e j c i - G r a f and We i l e r , 1928
Lepidopus sp. - (3), (6), (9), (10), (13), (14),
(31), (37), (38), (50); from Moineşti) and Leuciscus carpathicus BÖHM (originally
Family Palaeorhynchidae described by B öh m , 1941 from Comandău), two fresh water
Genus Palaeorhynchus BLAINVILLE, 1818 species, weren't listed as well because in many areas from the
Palaeorhynchus longirostris AGASSIZ, 1844 – Carpathians all the specimens described as fresh water ones
(48d); (e.g. Nemachilus, Leuciscus, Propercarina etc) were wrong
Palaeorhynchus glarisianus BLAINVILLE 1818 - (48d); determinated.
Palaeorhynchus cf. zitelli (KRAMBERGER, 1879) - (1);
Palaeorhynchushumorensis BRUSTUR&GRIGOR.,1973 Cephalacanthus trispinosus CIOBANU (originally
(50); described by C i o b a n u , 1977 from Piatra Neamţ) proved that
Family Scombridae sens GREENWOOD et al., 1966
Genus Cybium CUVIER, 1829 it does not belong to the Cephalacanthidae, needing revision
Cybium lingulatum (v. MEYER, 1846) - (30); and a better systematic placement (Tyl e r , pers. comm.). It is
Genus Scomber LINNAEUS, 1758 now reviewed by Baciu and Tyler (B a c i u , pers. comm).
Scomber voitesti PAUCĂ, 1929 - (1)55, (4), (11),
(13), (48); 4. Conclusions
Scomber saadii ARAMBOURG, 1966 - (48d);
Scomber sp. - (3), (9), (38) (48d); In the Romanian East Carpathians, 53 localities which
Genus Sarda CUVIER, 1829 provided Oligocene - Lowermost Miocene fossil fish-fauna,
Sarda sp. (30), (49a); or from other points where such fauna was mentioned
Genus Gymnosarda GILL, 1862 (teleostean skeletons or skeletons imprints only) could be
Gymnosarda disodilica CIOBANU, 1977 – listed, based on field or bibliographic investigations, or on
(48d), (51b);
Gymnosarda sp. - (30), (48d);
the study of the collections from different museums.
Genus Thunnus SOUTH, 1895 From the 53 mentioned localities, different authors
Thunnus abchasicus DANILCENKO, 1960 –
(48d);
determined 160 species belonging to 76 genera and 46
Thunnus albui (SIMIONESCU, 1905) - (48d)56; teleostean families. That' s why the Romanian Eastern
Thunnus sp. - (13)57; Carpathians represent one of the most important areas in
Genus Pinulothunnus CIOBANU, 1977 Europe.
Pinulothunnus cernegurae CIOBANU, 1977 –
(48a), (51c); Even if the present paper presents the Oligocene -
Order Pleuronectiformes Lowermost Miocene teleostean fossil fish-fauna from the
Romanian East Carpathians according to latest data, we are
51
This species was originally described by S i m io ne s c u aware that many of the mentioned species would be the
( 1 9 0 4 ) as Krambergeria lanceolata n. sp.
52
This species was originally described by Pa u c ă ( 1 9 2 9a ,b ;
1 9 3 3 c ) as Propercarina rebeli n. sp. The type specimen need 58
The specimens from Romania are presently reviewed by
new revision;
53 Ba c i u , C ha ne t (Baciu, pers. comm.)
This species was originally described by Pa u c ă ( 1 9 2 9a ,b ; 59
This species was originally described as Rhombus stamatini n.
1 9 3 3 c ) as Propercarina pietschmanni n. sp.; the type specimen
sp. by Pa u c ă ( 1 9 3 1 ). It was recently reviewed by ( Ba c i u ,
need new revision;
54 C hane t - i n p re s s : "Les poissons fossiles plats (Teleosteens,
Mentioned by Le i de n f r o s t 1 9 1 8 (fide Pa u c ă , 1 9 3 0 ) as Pleuronectiformes) dans les formations oligocenes des Piatra
Lepidopus dubius HECKEL and L. brevispondylus HECKEL Neamţ - Roumanie)".
(reviewed by Pa u c ă , 1 9 3 0 ); 60
Because the structure of the scale plates is similar in the
55
This species was originally described by Pa u c ă ( 1 9 2 9 b ) as Aracanidae and Ostraciidae, the isolated fossil plates cannot be
Scomber oligocenicus n. sp.; assigned with certainty to either family (Ty le r, 1 9 7 3 ; Tyl e r,
56
This species was originally described by S i m i one s c u ( 1 9 0 5 ) G r e go rova, 1 9 9 1 );
as Thynnus albui SIMIONESCU; 61
Isolated plates only (two types) was described by K re j c i - G r a f
57
Mentioned by J o ne t ( 1 9 5 8 ) as Thynnus sp. and W e i l e r ( 1 9 2 8 );

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

object of further reviews based on Recent fish osteology
mainly.
Acknowledgments:
We would like to thank prof. dr. D a n G R IG O R E S C U ,
University of Bucharest and dr. Ti t u s B R U S T U R ,
GEOECOMAR Bucharest, for the information concerning
some new localities or outcrops bearing fossil fish fauna and
for loaning us some specimens for study.
We are very grateful to dr. J a m e s T Y L E R , Museum of
Natural History, Smithsonian Institution (Washington, D.C. -
USA) and geol. So r i n BA C IU , Museum of Natural
History Piatra Neamţ, for the supplied literature and
information concerning the latest news in their work
involving the review of some specimens from the Romanian
Eastern Carpathians.

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GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

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GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998
P. Constantin – Oligocene-Lowermost Miocene Fossil Fish-Fauna (Teleostei) from Romanian Eastern Carpathians

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Citation: Constantin P. (1999), Oligocene - Lowermost Miocene Fossil Fish-Fauna (Teleostei)
from Romanian Eastern Carpathians, GEO-ECO-MARINA, 4:119–134, Bucharest.

GEO-ECO-MARINA, 4/1999
National Institute of Marine Geology and Geo-ecology of Romania
Proc. Intern. Workshop on «Modern and Ancient Sedimentary Environments and Processes» in Moeciu, Romania, Oct. 8-15, 1998