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Pando (tree) - Wikipedia, the free encyclopedia http://en.wikipedia.


Pando (tree)
From Wikipedia, the free encyclopedia

Pando (or The Trembling Giant[1]) is a clonal colony of a single male Quaking Aspen (Populus tremuloides) tree located in the
U.S. state of Utah, all determined to be part of a single living organism by identical genetic markers,[2] and one massive underground
root system. The plant is estimated to weigh collectively 6,000 tonnes (six million kilograms),[3] placing it as the heaviest known
organism in existence.[4] The root system of Pando is estimated to be among the oldest known living organisms in existence at
80,000 years of age.[5]

1 Name
2 History
3 Size and age
4 Debate
5 Location
6 Trivia
7 See also
8 References

The name "Pando" was chosen because it is Latin for "I spread",[1][6] while the nickname "The Trembling Giant" can be attributed to
the common name of Populus tremuloides, the trembling aspen, whose leaves frequently rattle.

Pando is thought to have grown for much of its lifetime under ideal circumstances: frequent fires
have prevented its main competitor, conifers, from colonizing the area, and climate change,
transitioning from a wet and humid weather pattern to semi-arid, has obstructed widespread seedling
establishment and the accompanying rivalry from younger aspens.

During intense fires, the organism survived by its root system, sending up new stems in the
aftermath of each wildfire. Due to its age, the climate into which Pando was born is markedly
different from that of today, and it may be as many as ten millennia since Pando's last successful
flowering, according to an OECD report: An aspen grove at Fishlake
National Forest (probably not
Clonal groups of P. tremuloides in eastern North America are very common, but
“ generally less than 0.1 ha in size, while in areas of Utah, groups as large as 80 ha
have been observed (Kemperman and Barnes 1976). In the semi-arid western United
States, some argue that widespread seedling establishment has not occurred since
the last glaciation, some 10,000 years ago (Einspahr and Winton 1976,
McDonough 1985). Indeed, some biologists feel that western clones could be as
old as 1 million years (Barnes 1966, 1975). It has been claimed that a single
clone, nicknamed "Pando" (Latin for I spread), covers 43 hectares, contains more
than 47,000 stems and weighs in excess of 6 million kg, making it the largest
known organism (Grant et al. 1992, Mitton and Grant 1996)

—OECD, Consensus Document on the Biology of Populus L. (Poplars)[3]

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Pando (tree) - Wikipedia, the free encyclopedia

Pando was discovered by Burton Barnes of the University of Michigan in the 1970s, and studied in detail in 1992 by Michael Grant of
the University of Colorado at Boulder.

Size and age

The clone encompasses 43 hectares (107 acres) and has around 47,000 trunks, which continually
die and are renewed by its roots. The trunks are connected together by its root system. The average
age of Pando's trunks (or technically, stems) is 130 years, as deciphered by tree rings. Michael
Grant in BioScience said:

...quaking aspen regularly reproduces via a process called suckering. An individual

“ stem can send out lateral roots that, under the right conditions, send up other erect
stems; from all aboveground appearances the new stems look just like individual
trees. The process is repeated until a whole stand, of what appear to be individual The trunk or stem of a Quaking
trees, forms. This collection of multiple stems, called ramets, all form one single,
genetic individual, usually termed a clone.
” Aspen

In comparison to Pando's most widely held total age of 80,000 years, the most accepted view among current anthropologists is that
Homo sapiens first migrated out of Africa to Eurasia and Oceania only 40,000 years ago, and finally to the Americas 10,000 years

Some experts speculate that Pando's reign since 1992 as the heaviest organism in existence may be
short-lived. Less well-studied Quaking Aspen in Utah may be 80 ha in size and one million years in
age, and other large colonies could exist elsewhere. A clonal colony of at least 7 Coastal Redwood
could weigh more,[8] although no such stand is known to exist. Other scientists feel that portions
of Pando's root system may be dead and might have led the plant to split into separate groups, and
thus would not be one organism, though the collective groups would remain the same singular
genetic individual.

Tree experts also note that the organism's age cannot be determined with the level of precision
found in tree rings; some claim Pando's age is closer to 1 million years[4]. Its current 80,000 years
designation is based on a complex set of factors including the history of its local environment
such as, the evidence indicating that there are few if any naturally occurring new aspens in most of
the western United States since a climate shift took place 10,000 years ago and eliminated
favorable soil conditions for seedlings, the rate of growth (including the differences of rates in Quaking Aspen in Utah
distinct climates when accounting for its local climate history, that males grow slower than
females, and that aspen grow slower at higher elevations with Pando at 2697 m or 8,848 ft above
sea level), its size, and its genetic code in comparison to the mutations found among aspens born in the modern era. Michael Grant
summed it thus:

Despite enormous crops of viable seeds, successful seedling establishment appears to be a rare event in the semiarid
“ West, but the establishment of new trees from seeds appears to be common in the moist, humid forests of New
England... aspen establishment from seeds probably has not occurred in the western United States since the last
glaciation, some 10,000 years ago... Part of the rationale behind current age estimates for aspen clones is that
sexual reproduction is effectively frustrated by the rarity of a favorable suite of conditions in semiarid
environments... High levels of genetic variation and excesses of heterozygotes are found in [the aspen of] semiarid
environments... Clonal reproduction is more common in arid environments... Heterozygotes often exhibit superior
longevity in forest trees [across many species]... growth rate of aspen decline with elevation, steepness of slope,
age of the ramet, and exposure to wind... growth rate decreased dramatically with elevation... The researchers
reported that the area of the female clones was 41% greater than males, the number of female ramets 52% greater, and
the basal area of females 56% greater [when compared at the same age and environment]...

Other candidates for oldest living organism include possibly older fungal mats in Oregon, older Creosote Bushes, and strands of the
marine plant posidonia oceanica in the Mediterranean Sea.

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Pando (tree) - Wikipedia, the free encyclopedia

Pando lives in the Wasatch Range of the Rocky Mountains in Utah near Fish Lake and Bryce Canyon National Park.

The United States Postal Service has made a stamp in commemoration of the aspen, calling it
one of the forty "Wonders of America".[9]
Alternate claimants to the title of largest and longest lived include Posidonia oceanica, a
seaweed off the island of Ibiza and Armillaria ostoyae, a fungus in the Blue Mountains of
eastern Oregon.

See also
Largest organism
Vegetative reproduction
Basal shoot

1. ^ a b Michael C. Grant (October 1993). "The Trembling Giant
( ". Discover 14 (10). Retrieved on
2006-10-08. US Postal Stamp depicting
2. ^ Pando ( by the National Park Service Pando
3. ^ a b OECD Environment Directorate. "Consensus Document on the Biology of Populus L. (Poplars)
" (Microsoft Word document). OECD. Retrieved on 2006-10-08.
4. ^ a b Genetic Variation and the Natural History of Quaking Aspen, Jeffry B. Mitton; Michael C. Grant, BioScience, Vol. 46, No. 1. (Jan., 1996),
pp. 25-31.
5. ^ Quaking Aspen ( by the Bryce Canyon National Park Service
6. ^ Research and Development Information Outreach, USDA Forest Service (Fall 1999). "Quaking in Their Roots: The Decline of Quaking Aspen
( ". Natural Inquirer 2 (1): 7–11. Retrieved on
7. ^ Templeton, Alan (2002). "Out of Africa again and again" ( Nature 416: 45 - 51.
8. ^ Bob van Pelt (
9. ^ Wonders of America: Land of Superlatives ( at

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Categories: Individual trees | Salicaceae | Flora of Utah

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