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Two Formulae

d n n 1 x ) = nx ( dx

df ( x ) dg( x ) d ( f ( x ) + g( x )) = dx + dx dx

Product rule

df ( x ) dg( x ) d g( x ) + f ( x ) [ f ( x ) g( x )] = dx dx dx

Product rule

df ( x ) dg( x ) d g( x ) + f ( x ) [ f ( x ) g( x )] = dx dx dx

E.g. Let: f ( x ) = x

g( x ) = x

dx dx d [x x ] = x + x dx dx dx

= 1.x + x .1 = 2 x

Prof. Ranjith Padinhateeri, Biosciences and Bioengineering, IIT Bombay

5

x x x x + ... e = 1+ x + + + + 2 6 24 120

x 2 3 4 5

d x x e )= e ( dx

Prof. Ranjith Padinhateeri, Biosciences and Bioengineering, IIT Bombay

6

x7 x3 x5 + ... sin( x ) = x + 6 120 5040

d (sin( x )) = cos( x ) dx

d (cos( x )) = sin( x ) dx

Sin(x) Cos(x)

d (sin( x )) = cos( x ) dx

d (cos( x )) = sin( x ) dx

8

Applications in Biology

Example 1: Polymerization of filaments (Growth of actin filaments)

Polymerization of actin

k p Rate of

Rate of depolymerization

polymerization

kd

k p = k 0 [C ]

[C] : unpolymerized G-actin monomer concentration

Prof. Ranjith Padinhateeri, Biosciences and Bioengineering, IIT Bombay

10

Polymerization of actin

kp

k p = k 0 [C ]

Growth speed

v = k p kd

v = k0 [C ] k d

11

kd

Polymerization of actin

v = k0 [C ] k d

12

v = k0 [C ] k d

Slope

dv = k0 dc

13

1.Draw Velocity vs concentration data 2.Find the derivative (slope). This gives k0 3.Find the Y-intercept. This gives kd

14

For actin

k0 = 11M s kd = 1.5 s

1 1 1

15

Applications in Biology

Example 2: Enthalpy and Entropy of a chemical reaction

16

Enthalpy, Entropy

G0 = RT ln K eq

( )

S + R

RT ln K eq = H 0 TS

0 eq

( ) H ln( K )= RT

G0 = H 0 TS

17

ln K eq

( )

H 0 S = + R RT dy dx

d ln( keq )

Derivative (slope)=

H 0 = = d (1/ T ) R

S Y intercept = R

Prof. Ranjith Padinhateeri, Biosciences and Bioengineering, IIT Bombay

18

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