Zitteliana A 52 (2012) 119

Ammonites and stratigraphy of a Lower Bajocian
(Middle Jurassic) section in Sierra Chacaico
(Neuquén Basin, Argentina)

Volker Dietze1*, Axel von Hillebrandt 2, Alberto Riccardi 3 & Günter Schweigert 4

 Zitteliana A 52, 119 – 139 Meraner Straße 41, 86720 Nördlingen, Germany.
 München, 21.07.2012 Institut für Angewandte Geowissenschaften EB 10, Technische Universität, Ernst-Reuter-Platz 1,
10587 Berlin, Germany
 Manuscript received Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Paseo del Bosque s/n,
20.06.2012; revision 1900 La Plata, Argentina
accepted 16.07.2012 Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany

 ISSN 1612 - 412X *Author for correspondence and reprint requests; E-mail: dietze.v@t-online.de


A Lower Bajocian section from southern Neuquén Basin (Argentina), with 3 faunal horizons (sphaeroceroides, submicrostoma and
giebeli horizons) in the Singularis and Giebeli zones, is described. The Giebeli Zone is relatively thick in contrast with the section at Paso
del Espinacito. The ammonite species Chondromileia submicrostoma (Gottsche) is quite variable and consists of large specimens. For
the first time well preserved and complete macroconch specimens of Emileia multiformis (Gottsche) are described. It is shown that the
lithostratigraphic and biostratigraphic boundaries are not coincident in the studied section.
Key words: Ammonites, Lower Bajocian, biostratigraphy, Argentina


Aus dem Unter-Bajocium des südlichen Neuquén-Beckens (Argentinien) wird ein Profil mit drei Ammoniten-Faunenhorizonten (sphae-
roceroides-, submicrostoma- und giebeli-Horizont) der Singularis- und Giebeli-Zonen beschrieben. Die Schichten der Giebeli-Zone sind
recht mächtig im Vergleich zum Profil am Paso del Espinacito. Die Art Chondromileia submicrostoma (Gottsche) ist bemerkenswert
variabel und beinhaltet großwüchsige Exemplare. Zum ersten Mal werden gut erhaltene, vollständige, makroconche Exemplare der Art
Emileia multiformis (Gottsche) vorgestellt. Es wird gezeigt, dass die litho- und biostratigraphischen Grenzen im untersuchten Profil nicht
miteinander übereinstimmen.
Schlüsselwörter: Ammoniten, Unter-Bajocium, Biostratigraphie, Argentinien

1. Introduction with preliminary comments in Hillebrandt (1973). We
illustrate and describe the ammonites collected from
Abundant fossiliferous Lower Bajocian outcrops levels included in the “Cura Niyeu Formation” sensu
exist in the ”back-arc” Neuquén Basin in west-cen- Volkheimer (1973; name no longer in use, see Sec-
tral Argentina (e.g. Westermann & Riccardi 1972, tion 3) of this section, where the strata of the Gie-
1979; Gulisano 1992; Zavala 1996a, 1996b). The beli Zone are extremely thick. The presence of the
area south of Zapala (Neuquén Province), especially ammonite genera Chondromileia and Emileia is of
in Sierra Chacaico, is famous for its well exposed special interest, as they contribute to a better under-
and thick outcrops containing rich Lower Bajocian standing of the evolution and infraspecific variation
ammonite faunas (for details see Westermann & Ric- of these groups in the Neuquén Basin.
cardi 1972, 1979). One of these outcrops – east of
the road from Melipil to Charahuilla (Textfig. 1) and
near the section Charahuilla-1 of Westermann & Ric- 2. Material
cardi (1979), as well as close to the section visited
in 1994 by the fieldtrips of the 4th International Con- In this paper, we describe and figure the material
gress on Jurassic Stratigraphy and Geology (Guli- collected by Volkheimer (1973), supplemented by the
sano & Gutiérrez Pleimling 1994: fig. 21, section 2) bed-by-bed ammonite collection made by von Hille-
– was described by Volkheimer (1973; ammonite de- brandt in 1971 and 1983. Overall, we studied about
termination by AvH) and studied by one of us (AvH) 80 specimens from about 10 succeeding ammonite

Zitteliana A 52 (2012) 120

Textfigure 1: Geographical location of the section near Arroyo Charahuilla in Sierra Chacaico (Neuquén Basin, Argentina).

levels. The illustrated ammonites are kept in the La caico (Textfigs 1, 2). The outcrop was originally
Plata Museum, Argentina, except of one specimen described as the type locality of the 126 m thick
from another locality illustrated for comparison. “Cura Niyeu Formation” (Volkheimer 1973; Quat-
trocchio & Volkheimer 1993; for a description of
Abbreviations used in the text and figure cap- the complete site, we refer to Volkheimer 1973 and
tions: Hillebrandt 1973).
HT = Holotype Gulisano et al. (1984), Gulisano & Gutiérrez
LT = Lectotype Pleimling (1994), Zavala (1996a, 1996b), and Quat-
var. = variety (used for varieties of infrasubspe- trocchio et al. (1996) studied in detail the lithology, se-
cific rank) quence stratigraphy and palaeogeographic changes
MLP = Museo de la Plata, Argentina of the Cuyo Group in southern Neuquén Basin. The
SMNS = Staatliches Museum für Naturkunde results presented by Zavala (1996a, 1996b) are of
Stuttgart, Germany especial interest. This author measured twelve sec-
AvH = Axel von Hillebrandt, co-author tions and identified depositional sequences of third
and fourth order. In the east there are mainly fluvial
deposits, while in the west coeval shallow marine
3. Section and litho-/sequence stratigraphy sediments occur. The section studied here (Textfig.
3) is in the middle and is equivalent to section 3 of
3.1 Introductory remarks Zavala (1996b), which is found 7.5 km north of sec-
tion 2, described in detail by Zavala (1996a). Both
The section described here is situated at the sections belong to the same series of outcrops, in
eastern slope of Charahuilla valley in Sierra Cha- which the same fourth-order sequences (JC4.1 and

. The images show the section in the year 1971. 1). Zitteliana A 52 (2012) 121 Textfigure 2: Section (indicated by arrows. bars show the layers described here) east of road from Charahuilla to Chacaico (see Textfig.

5] ?Chondromileia/Emileia [m] [Pl. month and day [of collec. 1.5 m: Siltstones with interbedded calca- Cuyo Group (from bottom to top): reous sandstones (CC 12): at base siltstones Los Molles Formation: (CC 11) with many ammonites in calcareous • 1 (27. Figs C. subfrancisci Sturani posterum Gottsche [M] [Pl. Figs 5–8. For an interpretation of our sec- tion. Fig. 2. altecostata Tornquist [M] S. in the labeling of the successive layers we follow beds of marls with numerous large sonniniids) Volkheimer (1973: layers 1 through 21).6 m: Alternation of siltstones. 7. sandstones) mer should be abandoned. Ammonites: Sonninia sp. 2] (bed CC 14 = vH 711126/4 [collection notice AvH: ca. 7] Ammonites: Sonninia altecostata Tornquist var. Ammonites: Chondromileia submicrostoma (Gottsche) [M] • 16 (0. Fig. altecostata Tornquist [M] [Pl. slightly higher within layer 4: Ammonites: Emileia multiformis (Gottsche) [M] [Pl. 2. subdeltafalcata Tornquist [m] [mentioned by Volkheimer 1973] • 13 (up to 1. 3] below 26/3]).6 m: sandy siltstones) [Pl. 1996b) are exposed. from slightly deeper] • 11 (4. to avoid confusi. Fig. 1. (bed CC 10).8 m: banks of sandstones) Bivalves: Pleuromya sp. [M] [mentioned by Volkheimer 4. 4.3). 3..8 m: Shales and siltstones. In its lower third. 5. 4] • 17 (1. Pl. Figs 1–3] ?Chondromileia/Emileia sp. We follow the authors mentioned above and use the name Lajas Formation • 7 (3. Volkheimer (bed CC 9). Figs 1–4] 4. ?Emileia/Chondromileia sp. • 3 (2. 2.8 m: sandstone) Molles and Lajas formations (sensu Zavala 1996a. [m] [Pl. Pl.g. 4. 2. [m] [Pl. Figs 1.2) of the third-order sequence JC4 of the Los • 5 (1. 1. 6] 3. Figs 1. 6] (bed CC 2 = vH 711126/9) S. 4. Ammonites: Chondromileia submicrostoma (Gottsche) [M] [Pl.0 m: siltstones) and ammonite fauna • 10 (3. 8] (bed CC 5). Fig. Ammonites: Emileia multiformis (Gottsche) [M] [Pl. 3–5] ?S. 3. Figs with interbedded sandy siltstones) 7. 3] and shales. Fig. basal 5 m with several layers (bed CC 11 = vH 711126/6) of calcareous nodules containing ammonites) Ammonites: Sonninia espinazitensis Tornquist [M] [Pl. Hillebrandt labeled the beds in which he collected • 8 (1. 5. boden- 8.5 m: alternating sequence of siltstones. labeled each of the fossil-bearing beds with the pre. gracile ?Emileia/Chondromileia sp. (bed CC 7). nodules): marls and shales. stones and sandy. Fig. 7] 1973] S. • 6 (3. 3 m below 26/4] ). muddy sand- for most of our section. Figs Tornquist [M] [Pl. in the lower • 2 (5 m: marly or sandy siltstones.6 m: sandstone) ammonites with the year. 7. Fig. Figs 6. as not all of the beds are menti- oned in his paper. Modiolus sp.0 m: calcareous and marly sandstones). calcareous concretions with ammonites are • 12 (2. Pl. in its lower part with interbedded sandstones. lithostratigraphy • 9 (4. this numeration is not continuous. 6] From between layers 6 and 7: Sonninia espinazitensis Tornquist [M] [Pl. 4–6] • 4 (7. 4.4 m. 5. Figs C.5 m: sandy siltstones) Lajas Formation: (bed CC 13s = vH 711126/5 [notice collection AvH: ca.5 m: Alternation of siltstones. • 14 (2.2 Description of the section.5 m: calcareous sandstones up to 0. [m] [Pl. Fig. However. at the base on. marls and The name “Cura Niyeu Formation” sensu Volkhei. Figs 1.8 m: Alternation of partly sandy siltstones Lytoceras aff. muddy marls. 5. 9] benderi Tornquist [m] [Pl. Very fossiliferous in the uppermost 3 m) • 15 (1. 6. however. see below (Section 3. 2] [loose. alternating half banks of sandstone). submicrostoma (Gottsche) [M] [Pl. submicrostoma (Gottsche) [m] [Pl. 4. with marls and sandstones.3 m: sandy siltstones) . 711126/1].8 m: siltstones with interbedded sand- common) stones) (bed CC 4 = vH 711126/8). (bed CC 6 = vH 711126/7). 8. tion] and “bed” [e. Figs 1–4] 3. 2.Zitteliana A 52 (2012) 122 JC4. (bed CC 13): Ammonites: Sonninia espinazitensis Tornquist [M] [Textfig. 3. submicrostoma (Gottsche) [m] [Pl. 2 m C. Ammonites: Sonninia espinazitensis Tornquist [M] [menti- oned by Volkheimer 1973] fix “CC”.

6 m) Emileia [vH 4] 15 (1.8 m) CC 7 CC 6 4 (7.2 12 (2.3 m) [vH 3] CC 14 16 (0.8 m) Chondromileia giebeli JC4.5 m) [vH 9] Textfigure 3: Chart of bio-. litho.1 Los Ps.6 m) CC 10 7 (3.5 m) CC 5 CC 4 2 (5. singularis CC 2 1 (27. sphaeroc.and sequence stratigraphy of the described section. Molles JC4.0 m) [vH 1] CC 16 19 (2.0 m) CC 13s 14 (2.6 m) 5 (1.0 m) [vH 2] Chondromileia giebeli 18 (4.5 m) CC 9 6 (3.0 m) [vH 8] Formation S.0 m) 8 (1. altecostata Ps.8 m) [vH 7] 3 (2.8 m) CC 13 11 (4. chrono-.3 m) ?CC 15a multiformis CC 15 17 (1.5 m) CC 11 [vH 6] 9 (4.0 m) CC 17 ? 20 (20.8 m) Lower Bajocian CC 12 Chondromileia submicrostoma Chondromileia submicrostoma 10 (3. Zitteliana A 52 (2012) 123 Formation Sequence Volkheimer (CC) Faunal Layer Subzone Zone Stage (Cuyo Group) (Zavala 1996b) Hillebrandt [vH] Horizon 21 (25.5 m) [vH 5] Lajas Formation 13 (1. .

Zitteliana A 52 (2012) 124 Plate 1 .

1–2 m Sequence stratigraphy and lithostratigraphy on below layer 19)] the one hand and bio-/chronostratigraphy (see Sec- tion 5 below) on the other hand do not precisely • 20 (20 m: sandy and clayish siltstones. Entolium sp. . 4. 1898 [M] preted as tidal sand-waves.. Ctenostreon sp. Los Molles Formation. Emileia multiformis (Gottsche) [m] [Pl. the section within off-shore marine claystones. fig. pl. Pl. Lower Bajocian. 2. Figs 3–6] with. The sequence ends ♀ – Westermann & Riccardi: p. interpreted by Zavala (1996a. The boundary between the Submicrostoma and Multi- • 21 (25 m: Alternation of sandstones) formis subzones corresponds to a change from an (= CC 17).. 8. (3) MLP 33741. 3–5. Fig. submicrostoma (Gottsche) [M] [Pl. by complete internal casts of endobenthic marine lost] bivalves. * = beginning of the body chamber. This platform area cannot have been the biotope of the ammonites • 19 (2 m: siltstones to silty sandstones) found there and they must have been washed (= bed CC 16 = vH 711126/2).2) of the Lajas Formation. 21. 2 Plate 1: (1.1. at top match (Textfig.1) and the main part belongs to the lower part (JC4. 4. 4. 8] cur in the lower part. with ammonite bed CC 13. Los Molles For- mation. 1 [LT] tepping set of delta-front deposits related to input. 22. pl. 7. and not at the place of burial. Altecostata Subzone. The ammonite fauna Our section starts in the upper part of the Los Molles Formation (JC4. figs 1. Fig. pl. (2) ?S. Dietze et al.3 Interpretation of the lithologic succession 4. 3 meters of related (sub-)genera. Cuyo Group. 1. 3. 3-5) Sonninia altecostata Tornquist var.2 starts with c. Layer 2 [CC 4 = vH 711126/8]. pl. Sequence JC4. (4) MLP 33742. They are sharply overlain by off-shore marine claystones with *v 1898 Sonninia altecostata Tornquist – Tornquist: p. v 1898 Sonninia gracilis Tornquist – Tornquist: p. 20. where ammonite horizons (CC 14 to CC 16) oc- Figs 4. see exhaustive discussion in sandstones. and (3) a delta plain (HST). bodenbenderi Tornquist [m]. 4. Sequence JC4. interpreted as an estuarine channel fill.2 has been interpreted as a plat- 9] form sequence.)]. which are • 18 (4. Sequence JC4. 7. The platform evolution started with sandstones of estuarine channels. 3. C. Layer 1 [CC 2 = vH 711126/9]. [mentioned in field-book of AvH. 8. Submicrostoma Subzone.1 Genus Sonninia Bayle. Figs 1. (1–5) Cuyo Group. This is not surprising. Lower Bajocian. input-dominated deltaic system to stream mouth- bar deposit. Figs 1. and 4. sphaeroceroides horizon. whilst in the upper part (CC17. with sandstones and claystones of a delta plain ori- Ammonites: Chondromileia giebeli (Gottsche) [M] [Pl. (2005). 19.3 m: siltstones. Singularis Zone.. which are followed by a series of coarsening-upward facies with sigmoidal tidal-bundles which are inter. 5. Giebeli Zone. constricta Imlay [M] [Pl. [m] [Pl. submicrostoma horizon. – All specimens: x1. ashore. (6–7) Sonninia espinazitensis Tornquist [M]. Sonninia altecostata Tornquist. fig.. 1879 1996b) as estuarines. Fully marine biotope conditions are proved Ammonites: Sonninia sp. as most of horizon with bivalves the ammonites lived on the continental shelf area (= vH 711126/1). pl. 8. 77. MLP 33740. Ammonite beds CC 6 (Pl. Trigonia sp. 4 dominated deltaic systems (Zavala 1996a.1 Family Sonniniidae Buckman. 1892 is overlain by cross-bedded sandstones (JC5) of the Lajas Formation. 3 posits show gently deepening foreset beds and lack v 1972 Sonninia (Papilliceras) espinazitensis Tornquist. 4). 7. swamp deposits are common. (7) MLP 33745. 4. Figs 7–9 (found loose ca. (6) MLP 33744 [pathogenic bubble probably caused by an injury of the shell followed by an infection of the epithelium [Keupp in litt. gin. partly sandy) followed (1) by a transgressive surface and tidal [most probably ammonites labeled by AvH as “CC 15a” sandwaves (TST). The boundary between Bivalves: Modiolus sp. 7) to CC 11 occur within these sequences. Emileia aff. 1878 CC 4) is formed by off-shore marine claystones and ends with lower delta front sandstones (Zavala Remarks: For an emended usage of Sonninia and 1996b). 1898 any tidal influence (bed CC 13s). 2] without ammonites). (5) MLP 33743. ammonite bed CC 12 and are followed by a fores. fig. Zitteliana A 52 (2012) 125 (bed CC 15 = vH 711126/3 = vH 831209/3). (1) MLP 33739. fig. gracile Tornquist [M]. Stream mouth-bar de. Camptonectes the Singularis and Giebeli zones is lower down in sp. and (2) a maximum flooding sur- are from here): face and an input dominated sandy deltaic system Ammonites: ?Emileia/Chondromileia sp. 1996b) v 1898 Sonninia curviplex Tornquist – Tornquist: p.1 (ammonite horizons CC 2 and Type species: Waagenia propinquans Bayle.

Zitteliana A 52 (2012) 126 Plate 2 .

4–5.: p. The ribbing becomes more or less synonymy and extensive discussion] v 2001 Sonninia altecostata Tornquist – Hillebrandt: p. With the exception of this special feature these Comparisons and conclusions: The specimens of finds are very close to topotypes of S. but helpful to mens of the morphospecies S. papillae on flanks and less dense ribbing on phrag- pical high whorls of the medium stage of S. 1. fig. 53–54. 4. 1 (1869): “Mutationen” with “Varietäten” within the v 1972 Sonninia (Papilliceras) espinazitensis Tornquist. figs 2 [HT]. 3 apical angle. 4. in the sense of Bather (1927) – or to quote Waagen pl. by layer 10 (CC 11 = vH 6). Pl. – All specimens: x1. espinazitensis are part of a chronocline with higher 4. “Formenreihe/Collectivart” (see Dietze et al. 4) holotype in Westermann & Riccardi 1972. (7–8) MLP 33749. Fig. fig. The specimens of S. fig. 7. pl. 5. with ribs that about the mid-height of the flanks continues on the curve forward slightly above the row of papillae and evolute body chamber. 3. pl. with papillae 58 occurring about mid-height of flanks. Figs 6. altecostata var. 22. altecostata (Pl. fig. we refrain separating (Textfig. Hence. the Lower Bajocian of Scotland. 20. Textfig. (4–5) MLP 33748. Material: 8 specimens from Charahuilla. Pl. 1. Fig. fig. although closer to the venter at the 91–95. 2 and 10) differ from each Sonninia espinazitensis Tornquist. here consists of black calcite and has fine folds and in contrast to the specimens described by Morton Remarks: The first representatives of the S. 56–58 aperture and the harmonic margin facing the ventral v 2010 Sonninia espinazitensis Tornquist – Dietze et al. 4. 10. 140. 3. dual transition from S. Fig. figs 1 [LT refigured]. now firmed by Dietze et al. For a description of similar material we refer to Morton (1973). 7. Lajas Formation. 4. 1. although the papillae are now higher on shell plasticity in younger representatives. ♀ – Westermann & Riccardi: p. 7 and Pl. exhibit more evolute coiling and a cies. The shell described 4. Pl. Pl. Fig. There is a gra- v 2010 Sonninia altecostata Tornquist – Dietze et al. Giebeli Zone. 4. 1898 [M] other in a characteristic way. (1–2) MLP 33746. the apex is towards the v 2001 Sonninia espinazitensis Tornquist – Hillebrandt: p. Although flanks and the inner whorls are less densely ribbed. intermediate morphs exist. (1973) shows a – slightly – shiny. pl. layers 2. pl. . 1. 3–4 show the sculpture of the nucleus. widely spaced on the body chamber. gracile. 5) yer 2 with exception of the slightly higher position of from the base of layer 1 (CC 2 = vH 9) show the ty. espinazitensis (Pl. 4. 1. 56. 21. figs 1–5. altecostata and S. (3. 7) reappears. Figs. who described apychi from Material: 20 specimens from Charahuilla. black surface. 3. 4. 6) from Espinacito section and to the lectotype of the spe. As already corded. : p.) discernible papillae. 7). pl. Fig. 3. 20. 6) them at specific level. 1). One stata var. 77. pls 23–26 [with further Aptychi: The large example of the species illus- synonymy and extensive discussion] trated on Textfig. fig. ♀ – at the end of the body chamber. pl. This nazitensis morphospecies occur in layer 2 (CC 4 = could be an argument against its classification as a vH 8. altecostata were re. 247. (2010) from the Paso del trast to those of S. 1 ly ribbed body chamber. Fig. pre- 91–95. gracile Tornquist (see re-illustration of the specimen found between layers 6 and 7 (Pl. 2 [with further body chamber. As in the aptychi Westermann: p. submicrostoma horizon. 3 [HT refigured]. p. describe in short individual characters. espinazitensis are almost identical to those from la- Remarks: Two of our specimens (Pl. Only in layer 10 (CC 11 = vH 6) the morphos. 6). * = beginning of the body chamber. alteco. 2005). 1898. resembles specimens from layer 10. in con- described by Dietze et al. 3. 4. (6) MLP 33750 (1–8) Cuyo Group. 6) Chondromileia submicrostoma (Gottsche) [m]. the morphospecific distinction between the Between layers 2 (CC 4 = vH 8) and 10 no speci. Fig. The innermost and true Cornaptychus – the presumed aptychus of Son- median whorls are densely ribbed without clearly ninia according to Arkell (1957: L439). However. The small specimens figured reach the keel. fig.. in about the same number as S. 7). pl. 29. 20. Pl. 31. (3) MLP 33747. the typical dense ribbing with small papillae at denser ribbing on the body chamber. In contrast to Plate 2: (1–2. altecostata S. Layer 4 [CC 6 = vH 711126/7]. espi. 3. The papillae set on at the end altecostata Tornquist ♀ – Westermann & Riccardi : of the phragmocone and continue to the end of the p. Fig. and 10. 7–8) Chondromileia submicrostoma (Gottsche) [M]. 77. part of the whorl. figs. S. 4 shows one aptychus valve still v 1992 Sonninia (Papilliceras) espinazitensis Tornq. Lower Bajocian. Figs 6. 6). (2010). the older specimens showing higher v 2012 Sonninia altecostata Tornquist – Dietze & Hille- whorls sections at middle growth and a more dense- brandt. mocone (cf. indicated by Westermann & Riccardi (1972) and con- pecies S. 54. Figs 6. pl. the same level. it can be observed that the assemblages from the different levels (layers 1. altecostata from layer 10 (Pl. 4. pl. Figs 1. 4. on Pl. 2 described by Morton (1973). layers 1 The next rich level with sonniniids is represented and 10. 1–3 sent in layer 1. They are considered as mor- phologically variable transients within a chronocline *v 1898 Sonninia espinazitensis Tornquist – Tornquist: p. pl. figs. Submicrostoma Subzone. Zitteliana A 52 (2012) 127 v 1972 Sonninia (Papilliceras) espinazitensis (?subsp. espinazitensis (Pl. 1. 1. pl. taxa used herein is purely artificial.

– Scale bar: 10 cm. but the latter genus organic content. Giebeli Zone.Zitteliana A 52 (2012) 128 Textfigure 4: Sonninia espinazitensis Tornquist [M] with its aptychus [Cornaptychus s. Los Molles Formation. Lower Bajocian. the specimens described by Morton (1973) our find Several similar forms have been classified as Laevi- shows a black shiny surface. Typical cornaptychi exhibit coarse must be regarded as a nomen nudum because it is folds in contrast to the material figured here and by lacking a type species (Moore & Sylvester-Bradley Morton (1973) and Westermann & Riccardi (1972). possibly due to a high cornaptychus by Trauth (1936). MLP 33738. submicrostoma horizon.] in situ. Later. Layer 2 [CC 4 = vH 711126/8]. Cuyo Group.l. Gąsiorowski (1962) emended . Submicrostoma Subzone. * = beginning of the body chamber. in Arkell 1957).

Argentina). Fig. pl. loc 10 in Hillebrandt 2001). 2. (5–6) MLP 33754. 2) 3a–c). no vH collection number]. (1–8) Lower Bajocian. 6. 2) v 1992 Emileia (E. fig. figs 5–6. 1. espinazitensis Tornquist ♂ [S. 6. fig. 8. other morphological features show some variability. Multiformis Subzone. b. giebeli horizon. Diagnosis: Moderately-sized sphaeroconic ma. ty “Stephanoceras” multiforme var. 90. re- Emileia multiformis Gottsche. (6–9) Layer 10 [CC 11 = vH 711126/6]. figs 7a. 13. (6) MLP 33764. layer 14 [CC 13s = vH 711126/5. is difficult. which con- sists only of incomplete phragmocones. micromphalum (Gottsche 1878. pl. Thus. 3 benderi Tornquist. selected by decide if these two taxa are conspecific. (3) Lytoceras aff. layers 14 v 1992 Sonninia (Papilliceras) cf. (2–3) MLP 33752. Lajas Formation. 2. 27 Cornaptychus s. figs 4. Giebeli Zone. Microconchs [=Otoites] lappeted. 22. Giebeli Zone.. pl. b. figs 3[HT refigured]–5 Material: 4 specimens from Charahuilla. (1–2) MLP 33762. One specimen (MLP 33766. Figs 7.l. figs 2a. 1 specimen from Aguada Profeta (Northern – Westermann: pl. pl. Chile. (1–2) MLP 33756. Submicrostoma Subzone. 7. fig.1 Genus Emileia Buckman. Submicrostoma Subzone. 1a.2 Family Otoitidae Mascke. The ribbing of the specimens figured by Gott- sche is much denser compared to the variety E. 2) the Type species: Ammonites brocchii J. parison with Gottsche’s type material. macromphalum It should be noted that in ?S. multiformis var. MLP 33755. ♂ and 16. Lower Bajocian. Figs 1. b. MLP 33765. 1. bo. no vH collection number]. Figs 7. . – Tornquist: species. Westermann & Riccardi (1979. 22. phragmocone is not preserved and therefore com- 1818. (4) Sonninia espinazitensis Tornquist [M]. Zitteliana A 52 (2012) 129 the latter genus but still failed designating a type 1898 Sphaeroceras multiforme Gottsche sp. However. macromphalum. 21. (1) MLP 33751. 1898] – Westermann & Riccardi: p. 2. 3. 4 The specimen MLP 33768 (Pl. (1–9) Cuyo Group. pl. Plate 4 (p. 24. pl. 54. 130): (1–3. 94. 8. distances between primary ribs of that specimen are close to 4. 1878) ♀ – Westermann & Riccardi: p. (1–8) Cuyo Group. mul- *v 1878 Stephanoceras multiforme sp. Lajas Formation. In specimen MLP 33767 (Pl. pl. pl. * = beginning of the body chamber. pl. (7–8) Emileia multiformis (Gottsche) [M]. 1898 [m] extensive discussion] (Pl. 55. (3–4) MLP 33757. 5. 1878 [M] figured by Westermann & Riccardi 1979. figs. Plate 5 (p. – All specimens: x1. 8. 2 [NT] 3–5.2. b.) multiformis (Gott. nova – Gottsche: p. figs 7a. specimen lost) or with the variety E. 3. (4) between layers 6 and 7. multiformis (Gottsche 1878. 92. pl. pl. The flank. 122. both lost) from Paso del Espinacito (San Juan 3) about each second or third rib does not originate Province. MLP 33760. However. 6. 2. (1–3) Layer 4 [CC 6 = vH 711126/7]. 131): (1–4. (1–8) Cuyo Group. 1898 type (Gottsche 1878. (4–5) MLP 33763. espinazitensis Tornq. (1–9) Lower Bajocian. 6. *v 1979 Emileia (Emileia) multiformis (Gottsche. Pl. figs 6a. espinazitensis/altecostata group. subdeltafalcata (see figured by Gottsche (1878. 5 refi- gured] Description and remarks: The final diameters of the complete specimens illustrated by us are simi- Material: 1 specimen from Charahuilla. 8–9) ?Chondromileia/Emileia [m]. 3. Lajas Formation. pl. pl. tiformis var. figs 5a. 4) reminds Plate 3 (p. (7–8) MLP 33753. close to the incomplete phragmocones of the varie- croconch of the S. submicrostoma horizon. Figs 1–4. b) is too small for a proper interpretation. boden. submicrostoma horizon. 3. we keep this aptychus tentatively in p. 6. Sowerby. pl. 7 [HT]. Pl. layer 1. 1907 those shown in our specimen. (2010) assumed that the group of ?S. v 2001 Emileia multiformis (Gottsche) – Hillebrandt: p.: p. (5–8) Layer 8 [CC 10. figs 4–5 *v 1898 Sonninia bodenbenderi Tornquist – Tornquist: p. figs Westermann & Riccardi 1972. The specimen figured by us at the umbilical seam and begins below the middle seem to be a complete example of this variety. but Westermann & Riccardi 1972. fig 9 [HT] v 2010 Emileia multiformis (Gottsche) – Dietze et al. 2.) – Westermann: pl. 96–95. we do not have enough material to neotype of the species E. 4–6) ?Chondromileia/Emileia [m]. 132): (1–2. figs. mary ribs: ze et al. MLP 33761. 22. (8–9) MLP 33758. (5) Chondromileia sub- microstoma (Gottsche). The dense ribbing of the body chamber suggests a croconchiate ammonite with strong and prominent similarity with Gottsche’s “Typus” [= HT] of the spe- primaries and gradual coiling in all ontogenetic cies E. 54 pl. Pl. pl. lar. (1–5) Layer 8 [CC 10. 4. – All specimens: x1. pl. 4. 5. (7) Sonninia altecostata Tornquist [M]. v 1972 Sonninia cf. 5. In contrast. 4 [non HT as mentioned. (6) Sonninia espinazitensis Tornquist [M]. b) of this species. 1–2 [with further synonomy and ?Sonninia bodenbenderi Tornquist. Whorl section and umbilicus uncoiling are closer to those for the holo- 4. – All specimens: x1. MLP 33759. 3. fig. 5–8) Chondromileia submicrostoma (Gottsche) [M]. stages. multiformis. subfrancisci Sturani posterum Gottsche [M]. excepting the prominent short pri- Remarks: Westermann & Riccardi (1972) and Diet. Giebeli Zone. fig. MLP 33766. pl. figs (Pl. Figs 3. figs 1. 8) is very denbenderi/subdeltafalcata is the corresponding mi. Figs 1.

Zitteliana A 52 (2012) 130 Plate 3 .

Zitteliana A 52 (2012) 131 Plate 4 .

Zitteliana A 52 (2012) 132 Plate 5 .

Microconchs lappeted. 7. mis and E. pl. 11.) – Westermann (ed. Figs 8–9 about 1/4 to 1/3 of the final whorl is mis- sing. often laterally crushed and damaged prominent and become weaker higher on the flanks. Figs 1. 3. features of C. The final diameter of E. 5. 3. 2 v 2001 Emileia (Chondromileia) giebeli submicrostoma Material: 2 specimens from Charahuilla.2 Genus Chondromileia Westermann & Riccardi. figs 1–8 discussion] aff. figs 1. pl. However. pls 8. mulitformis var. 8. arkelli Maubeuge exhibits the lower third of the flanks. secondaries still visible on we could find in the literature is Emileia constricta body chamber and a broad-oval whorl section – are Imlay. v 1992 Emileia (Chondromileia) giebeli submicrostoma 28. 2. fig. polyschides are coarse and relatively well E. E.135. multiformis. 5–8. 1878) ♀ – Westermann & Riccardi: p. Figs 1–3. 6. Pl. Multiformis Subzone. pl. Plate 6: (1–4) Emileia multiformis (Gottsche) [M]. pl. 4. Figs 1.2. 2. 8. fig. and the broadly oval whorl section. in the specimen figured on Pl. Lower Bajocian. 3 [LT] of Manflas (Chile). figs 1–3. 56. figs 2. . multiformis (15–16 cm vs. whilst primaries in E. b (Gott. pl. 15. a closer examination showed some significant dif. brocchii (Sowerby) differs by a clearly more de- spaced. layer 16 [CC 14 = vH 711126/4. appearance. – Torn- quist: p. figs 1. aff. There are relatively compressed shells (Pl. 5. The phragmocone is preserved. n. but with prominent primaries restricted macromphalum. morphic ammonite with macroconchs of Emileia-like pl. 8. figs 5a. especially on the body chamber (cf. show less morphological resemblance. submicrostoma – prominent. multiformis assemblage of our section: (1) final Material: 16 specimens from Charahuilla. 2) to 100 mm (Pl. in E.: p. to inner part of flank and sudden egression of body Despite these morphological differences. Giebeli Zone. 2001: 54) shows a very broad whorl section. 2. All other nominal species in the literature upper flank. 3. giebeli horzion. Characteristic broadly rounded oval. Fig. Figs 7–9) (Gottsche. 2. 7. a more rounded whorl section and more prominent polyschides are coarse and prominent also on the primaries. We. figs 4a. known main. fig. 3 [LT refigured]. 2. and 8. 55. sphaeroconic. 8. (1–4) Cuyo Group. although the last 4. Figs cies. figs 6a. constricta shows 4–5). 3. MLP 33777. micromphalum but with a more Diagnosis: Moderately-sized. Pl. Pl. – Imlay: p. 5. An additionally specimen (SMNS 70022. in few specimens. (2) primaries are less chambers.): pl. 9 [with further synonomy and extensive 11. pl. 8. (Gottsche) – Hillebrandt: p. Size of complete specimens varies are pathogenic and thus not diagnostic for the spe. layers 4 diameter is larger. from 69 mm (Pl. Pl. 1979 Figs 1. 98. figs. ♀ – Westermann (ed. b) E. Figs 1. Pl. Zitteliana A 52 (2012) 133 at first sight Emileia polyschides Oppel. Comparison and remarks: Two specimens (MLP 33776. 2) from the Giebeli Zone (Multiformis Subzo- ne) of Aguada Profeta in Northern Chile (Hillebrandt Type species: Stephanoceras giebeli Gottsche. 1. Lajas Formation. pl. 1878) [M] primaries restricted to the lower part of the flanks (Pl. relatively For these features the best fitting nominal species wide spaced primaries. layer 19. fig. – All specimens: x1. lustrated specimens belong to E. 2) feature differs slightly in an undescribed fauna of E. b) as E. 27 Emileia aff. Cha- racteristic and common features are the relatively small final diameter –as compared to the “European” Chondromileia submicrostoma species of this genus –. 1964 Emileia constricta Imlay. Although the primaries in both E. multiformis they are prominent only in pressed whorl section. 7. The name-bearing constrictions of this taxon well developed. chamber in the adult stage. the coarse and prominent (Gottsche. as in the 1878. (3) body chamber whorl section is high-oval and not totally or in part. near the aperture. (1–2) MLP 33767. di- evolute umbilicus – as figured by Gottsche’s (1878. 3. multiformis (Hillebrandt 2001) from the Giebeli Zone * v 1878 Stephanoceras submicrostoma sp. 8.): pl. B40. 3 v 2010 Chondromileia submicrostoma (Gottsche) – Dietze et al. multiformis var. Figs 7–9) differ clearly from the E. 2). 7. so that the original diameter of the complete Description: Most specimens show only the body ammonite was about 15 cm. pl. specimen figured by Gottsche (1878. all il. polyschides is larger than in E. 1992 Emileia constricta Imlay. Although the holotype of E. 6. 12–13 cm). 4. and whorl sections are also quite comparable. 4. stermann 1964. (3–4) MLP 33768. the illustrated paratypes (Imlay 1964. pl. pl. constricta Imlay [M] v 1979 Emileia (Chondromileia) giebeli submicrostoma (Pl. 8. 4. ly from the northern margin of the Tethys. 8) show a similar ribbing style and width of umbilicus ferences. 91–94. v 1898 Sphaeroceras submicrostoma Gottsche sp. 4. b and pl. nova – Gottsche: p. a denser ribbing when compared with our material. figs 1a. sp. multifor.

Zitteliana A 52 (2012) 134 Plate 6 .

Zitteliana A 52 (2012) 135 Plate 7 .

croconchiate otoitids from the studied section and 1878) ♀ – Westermann & Riccardi: p. 4. 633. Hillebrandt (2001. whilst Dietze at al. layer 17. The primari. Lajas Formation. submicrostoma costata/espinazitensis group. The similarity of both 1. 7). 1 [HT] v 1898 Sphaeroceras giebeli Gottsche sp. 27 Due to incomplete preservation of most of mi- v 1979 Emileia (Chondromileia) giebeli giebeli (Gottsche. Cuyo Group. Wester- ge-sized C. giebeli represent transients morphology of both associations we dismiss the or succeeding chronospecies within the same chro- possibility that they belong to different (bio-)species. 330. pl. This and the – see the somewhat intermediate morphs figured on larger adult size of E. nova – Gottsche: p. – Riccar. – Giebel: p. Giebeli Zone. 5. 8) *v 1851 A. submicrostma specimens of 7. (7) MLP 33776. Due to their in.5. characteristic for the Chondromileia giebeli (Gottsche.) s. tures to distinguish both taxa. figs taking into account the great similarity between 1 [HT refigured]. – all specimens: x1. cone. Lower Bajocian. which fades out on the body type material comes (cf. Multiformis ship has been pointed out in Westermann & Riccardi subzones of Northern Chile. Fig. Multiformis Subzone. pl. 2. Cha- the Charahuilla/Chacaico area are remarkably larger racteristic are the compressed body chamber and the than those from Paso del Espinacito. multiformis are the best fea. 56. from where the dense and fine ribbing. we are not able nomy and extensive discussion] to say to which of the genera belong most of our v 1992 Emileia (Chondromileia) giebeli giebeli (Gott. 2. pl. 7 specimens. They are very close to the specimens figured by Westermann & Riccardi (1979. nocline. 135): (1–2) Emileia multiformis (Gottsche) [M]. (3. (3–4) MLP 33774. giebeli stratigraphically later. to re. submicrostoma is mixed with that of C. (Pl. giebeli horizon. 8. 2. 137): (1–2) Chondromileia submicrostoma (Gottsche) [M]. (4–5. (1–9) Giebeli Zone. 5 and Pl. submicrostoma and C. figs. fig. The characteristic differences – extre- v 2001 Emileia (Chondromileia) giebeli giebeli (Gottsche) – mely involute inner whorls and sudden egression of Hillebrandt: p. C. layer 18 [CC 15a]. 1A–C) from the Submicrostoma resp. Cuyo Group. submicrostoma occurring stratigraphically Of great interest is the striking similarity of this lar. However. (5–6) MLP 33775. latively broad shells (Pl. Fig.s. but con. figs 6. pl. 1 [with further syno. Common and typical as chrono-subspecies. 53. 7. 2. constricta Imlay [M]. giebeli egression of the adult body chamber. 5. compare Pl. 8. Description: The two figured specimens have com- complete preservation it is impossible to determine plete body chambers with small relicts of phragmo- the number of primaries on the last whorl. 7.6 the body chamber we identify two specimens (Pl. pls 7–9). 2–3. (3–9) Cuyo Group. Lower Bajocian. MLP 33773. (3–6) ?Emileia/Chondromileia [m]. altecostata and S. 6) ?Emileia/Chondromileia [m]. 6. pl. This probable relation. (2010) re- features for both groups are the prominent. – Riccar- di: p. v 2010 Chondromileia giebeli (Gottsche). 7. Comparison: The C. 7–9) layer 17 [CC 15 = vH 711126/3 = vH 831209/3]. 3) figured specimens of macroconchiate C. mann & Riccardi (1979) therefore regarded both taxa mens of Emileia multiformis. figs 1. figs 3A. Figs 4. 91–94. and C. alte- stermann & Riccardi (1979: 126). 1. 5) resembling C. – All specimens: x1. layer 17 [CC 15 = vH 711126/3]. 7 sistently with a more depressed whorl section and a less curved venter than in C. submicrostoma fauna and some speci. layer 19 [CC 16 = vH 711126/2]. figs 1a–b) from a nearby Charahuilla section. 4. Multiformis Subzone. 3 Fig. from both areas in Westermann & Riccardi 1979. 3.g. 1. Lajas Formation. earlier. Figs 4. in Chondromileia is lower: only rarely the morphology ture cannot be seen in our material –. MLP 33769. pl. Northern Chile (Hille- brandt 2001: 54). pl. Figs 1. fig. 1. – Dietze et al. 246 4. 98. Pl. (4–5. 7–8) Chondromileia giebeli (Gottsche) [M]. SMNS 70022 (3. Cuyo Group. The reason for this size differences may Remarks: As in S. be ecological and in view of the otherwise identical C. 7. morphological plasticity exhibits extremely involute nuclei – although this fea. fig. in contrast to the S. submicrostoma (pl. figs 2a–c. figs 1. MLP 33772. Chondromileia [m] and Emileia [m].3 Remarks on the microconchiate Otoitidae *v 1878 Stephanoceras giebeli sp. . giebeli horizon. also the figured specimens chamber with exception of the low primaries. 15. 6. – Tornquist: p. (9) Emileia multiformis (Gottsche) [m]. (7–8) MLP 33771. giebeli. (4–5) MLP 33770. Plate 8 (p. Due to the egression of v 2008b Emileia (Chondromileia) giebeli (Gottsche).5. These specimens show (1985). 7. short garded them as chronospecies. (8–9) MLP 33777. 55. Fig. confirmed here.2. B) and C.6 most of our microconchs. stermann & Riccardi 1979) – cannot be observed in di: p. 1878) [M] genus Chondromileia. fig. nec d’Orb. Multiformis Subzone.Zitteliana A 52 (2012) 136 Figs 5. 2 with Pl.: p. Lajas Formation. taxa suggests a possible common ancestor. These results are primaries. Giebeli Zone. Aguada Profeta (upper course of Quebrada del Profeta. probab. as already pointed out by We. espinazitensis. C. 6. giebeli horizon. giebeli. However. and a sudden of C. es are relatively well recognizable but variable (e. 8). pl. excellently the involute phragmocone and the sudden egression of the body chamber.134. Multiformis Subzone. Plate 7 (p. (7–9) Emileia aff. Pl. 4. ly in the genus Docidoceras. – Westermann: pl. bullatus Gieb. Giebeli Zone. giebeli (pl. Material: 3 specimens from Charahuilla. 6. 6) layer 16 [CC 14 = vH 711126/4]. 4. 1 the body chamber in the genus Chondromileia (We- v 2008a Emileia (Chondromileia) giebeli (Gottsche). Lajas Formation. figs 7.

Zitteliana A 52 (2012) 137 Plate 8 .

Figs 1. tensis dominated fauna from layer 2 upwards. espinazitensis group marks the beginning ments. Bio. macroconchiate Emileia multiformis (Westermann & Riccardi 1979. The sonniniid fauna has changed In contrast to the section of Paso del Espinacito from a nearly exclusively S. Germany) helped to digitize some slides. This because these groups normally had a pelagic life. H. vH 711126/4) because of its predominant fauna of mon 1985). submicrostoma. Figs 1–4. giebeli even lower nammatoceras sp.. France) for the de- yo Los Molles and Arroyo Picún Leufú an abundant termination of a lytoceratid and H. Dietze et al. H. of the C. forming also the morphospecies S. espinazitensis – now with a large plasticity As in Hillebrandt (2001) and Dietze et al.and chronostratigraphy of layer 10 (CC 11 = vH 711126/6) another fauna with S. altecostata – oc- the beds are subdivided into the standard chro. 3) appears exotic among From the middle part of layer 4 (CC 6 = vH 711126/7) the remaining ammonite fauna. index species Ps. In the basal part 5. G. (2) 18) and Gulisano & Gutierrez Pleimling (1994).. Joly (Beaugency. 2. constricta. Zavala (Bahia Blanca. evolutionary replacement of the S. and Puchenquia sp. Multiformis Subzone cardi 1992. 8. (Stuttgart. cisci posterum (Pl. sphaeroceroides itself.2. Parent (Rosario. Pl. 2010. Zitteli Subzone Beds of this subzone have not been recorded in Acknowledgements our section. Fig. giebeli horizon. Horizon of Pseudotoites sphaeroceroides: The lo. the index of this faunal horizon (Hil- lebrandt 2001. 6. multifor- yielded Aalenian ammonites (Tmetoceras sp. (3) These ammonites are most likely allochthonous elements. Germany) is thanked for acting as referee. upwards the ammonite fauna consists mostly of C. . and with C. We refrain from naming a new horizon for layers 14–16 for several 5. giebeli) dominating. 9) shows the more regular egression of identifying this horizon (Hillebrandt 2001. 2010).3 Remarks on other taxa yers 2 (CC 4 = vH 711126/8) to 13 are assigned to this faunal horizon. nostratigrahic zones and subzones of the Andean Province (Westermann & Riccardi 1972. Dietze et al. Dietze for the lithostratigraphical interpretation and wer parts of Layer 1 (CC 2 = vH 711126/9) belong to subdivision of the section. ammonite fauna from this subzone. Within Horizon of Chondromileia giebeli: The strata from this chronostratigraphical scheme we describe the layers 14 (CC 13s) to 19 yield the C. smallest distinguishable biostratigraphic units that We assign to this horizon layers 14 to 16 (CC 14 = can be recognized as faunal horizons (sensu Callo. 5. mis and C.2 Bajocian Because of (1) and (3) we refrain also from naming a new horizon for layer 19. altecostata ammonite (Dietze et al. Keupp (Berlin. The best determination we can give for all other microconchiate otoitids (Pl. in the section. 1979. Parent translated some this faunal horizon. gracilis) is characteristic enough for (Pl. espinazi- and lytoceratids are very rare. Submicrostoma Subzone Horizon of Chondromileia submicrostoma: The la- 4. Dietl niid fauna with typical specimens of Sonninia alteco. One specimen stata (incl.2 Giebeli Zone 8. 2008b. J. submicrostoma horizon (Dietze et al. the sonni. by the S. 5. Blau (Frankfurt a. aff. 6) as C. 7. submicrostoma [m]. 4–6. of Emileia multiformis [m]. 4. 2010). 5. Dietze.1 Aalenian reasons: (1) we do not have enough material to assu- re that there are two different horizons. 2010). on the base of only two 5. Pla.2. curs within the C. Hillebrandt 2001). 2008a. 5. which were deposited by high tides to their places of burial (see Section 3. lower Westermann & Riccardi (1979: 93) described from a parts of the Los Molles Formation in the area have nearby section a common occurrence of E. The sole specimen of Lytoceras aff. Although we could not prove the Spanish texts for V. 2010). Argentina) provided information to V. This is not surprising. one of them According to Hillebrandt & Westermann (1985: with E. Germany) for his opinion about a pathological speci- Altecostata Subzone men. 7. Westermann & Riccardi (1972.Zitteliana A 52 (2012) 138 Figs 3. Fig. Figs 3–6) is ?Emileia/Chondromileia [m]. altecostata group style and did not inhabit shallow deltaic environ. 1979) described from a section near the junction of Arro. We thank B. submicrostoma horizon. Pl.). Dietze et the body chamber and the less involute inner whorls al. var. A.1 Singularis Zone occurrences of E. subfran. Ric. phylloceratids are totally missing fauna in the lower parts of layer 2 to a S. Main.3). (2010). 9. Pl. C. Figs 3. Argentina) and C. Dietze & Hillebrandt 2012). giebeli. multiformis (resp.

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