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THE TRILOBITE GENUS PHILLIPSINELLA FROM THE ORDOVICIAN OF SCANDINAVIA AND GREAT BRITAIN by D. L. BRUTON Asta. Pilpsnt formant mona Pprecla s. n.of desre rom the Carudc Lome Chas ‘Rope Sai i athe Upper Chasops Limes i) seco tne Ono Region Nery, pth witigued but tanned seem om cguratet hones tnd Sern Seem Matar fF. procures. os efi tom ie bie Panam of nother agp. The Normepn material provides ne formation {moons pternsand eensitl race ure. Wel dened gute tbe Turow: ne dst faethe ie tee bin tive best -yrsar onc lobein Pformons A nem ages pe othe ‘Sc ach tly Sc ent pa gu hay ‘Svlow knoe cmd flows te recogston two specs wou sn Arent wp euro ts fiom Bake Saocinai enern Iedand, Turley, Kxathnan, and Ustetsan, and an Agi Secs 28 ‘Sted mound praolo son to ows rm Bra, Bobena Polina out Swen, Tuts work represents part of a larger project concerned with the vertical stati {graphical distribution and description of Middle Ordovician fossils from several Sections inthe Oslo Region, Norway. Detailed coleting has produced a wealth of thateral amongst which ae specimens Belonging to two mow species of Philfipsinelta described herein from the Caradoc Lower Chasmops Shale (4bs) and the Upper Chasmops Limestone (dbo, .) respectively, Further specimens fom equivalent forzony or older are desrbed from Sweden, none of which has been. previously figure, although occurrences have ben listed by Jaanusson (1960, 1963) from the Uhaku Stage cone of Gyprograprs teretiaaeulu) of Olan andthe Sijan district. “The importance of the Norwegian and Swedish material is that it all of pre- Ash age and thus haps to shed some light on the origin and distribution of the fges Philipsinela otherwise best known through is type species, parabola {farrande), which i abundantly represented in rocks of Asal age in Bohemia, Britain (for dscusion see Whittington 1980, 1966, 196; Ingham 1966, 1970; Price 1973) eastem Ireland (Dean 1974), Poland (Kielan 1960), Kazakhstan (Appotonoy 10974), and Ustekistan (Abdullaey 1972), In discussions on tnilbite migration, Kielan (1960, p38) suggests « Scandinavian origin for Phifipsinla. The material describ) herein and tat recently described by Den (1973, p. 308) from the Arenig of Turkey, shows tht the early history ofthe genus appears tobe within the Asaphid Province. Family puut.arsineiupat Whittington, 1950 (Genus PHILLIPSINELLA Novik, 1885 ‘Type spcien By original dsination, Phacups parabola Barrande, 1846, p 6, from the Kriluy Dvdr Formation Asha, Sober eng, 1, Pe 41%, TK, ph 110) 70 PALAFONTOLOGY, VOLUME 19 Diagnosis. Cephalon as long (Sag.) as wide, moderately convex; glabella clavate, defined by deep furrows, narrowest between palpebral lobes, widest across frontal lobe. Three glabellar furrows on suitably preserved material, otherwise shallow to lacking. Dorsal furrows end in anterior pit alongside frontal Jobe: second (posterior Pit) in furrow nearly opposite front of palpebral lobe. Facial sutures opisthoparian, anterior branch diverging forward, curving to meet along anterior margin, Con: nective sutures diverge inwards and backwards, rostral platelong (sag ), lying beneath frontal glabellar lobe. Hypostoma elongate (sag.) covering posterior haif of glabella and slightly beyond posterior margin; middle body oval in outline, anterior wings short. Free eheek with genal spines extending the length of thorax. Thorax with six segments, pleural furrows shallow. Pygidium semicircular to quadrate in outline with small postero-median notch. Rachis long, slightly tapered, moderately convex With post-rachial ridge best seen on internal moulds; three o six rachial rings; pleurae smooth or with at least three shallow furrows. Glabella with raised line pattern with ‘or without associated pits, On pygidium, raised lines along postero-lateral bordet and doublure. Stratigraphical and geographical range. Ordovician (Arenig to Ashgill); Scandinavia, Turkey, Kazakhstan, Uzbekistan, Ceechoslovakia, Poland, Bintan, and eastern Ireland. Discusion. The new diagnosis is emended from that of the Treatise Harington et al. n Moore 1959, p. O40S) so asto incorporate new material topether with study of ‘opoiype material of the type species from the Kraluv Dvir Shale, Czahosiovakia, and from the Upper Drummuck Group, Girvan (Begs Collection, Hunter Museum, Universit’ of Glasgow), the Ashgill of Wales and northern England (Sedgwick Museum, Cambridge), and from northern England figured by Ingham a7, The large colletion of Philipsinelapreclara sp.nov. is excellently preserved and provides much new information on morphology, but dificulties aren comparing ‘with material assigned to P. parabola and ales which are mainly described from sillstone and shale, or a external moulds from decaeied limestone REMARKS ON MORPHOLOGY (Gatton. P.precara stows fos theirs ime details of aber ohes nd farow which, llong aanuson (1956, pp 36-37), are numbered and lere fom the posterior forwards, ng the notation L “ahabelar foe) ai S(giatcie furrow): se PL 106, fi 2). The abe and furrows cut om the waren ‘Posterior half ofthe labella adjacent tothe wide da faiom, the lobes Being acentted on ecu ‘ofthe granulation covering the. and which absent fom the furrows. There ao trace of farrows It front ofS, onthe args, fornardy expanded ronal abla lobe and which, onthe veal sie, nde. si by the ge esl plate P15, 6: ew ao Whitington 193), pl 7S, 46). The tal label Jobe (Ls the most prominent and forma Interal expansion af the hinnan prt of te labellar {meaty n front ofthe octal row. This expansion Logtber with Sion spon of Porch ‘ts. 17), onthe cranium of Philpsnle sp nde B (1S, Tyand on P- parahol himica {see Dea 1974 i, 29 fig. 6) In these, however, unk in P preckara atonal abe and furows e {aint or lacking’ One specimen of P. parabols strated by Kila (196 PL 3B I, lowermost ‘what may bea basil elabela lobe, while othe pine do not, and lobes ae licking on mate by Whitington (966 1 25, p51, 6 and Ingham (197, pl hes. 13.1 ‘ater mate, absence of lobes maybe a result of preservation, SRUTON: PHILLIPSINELLA FROM NORTH-WEST EUROPE vn Median occipital tuberce. alike specimens of P. parabola snow lo. P.pecara (PL 104, 3 5,6 PIGS fg 8) and the eranigism of Pilnta sp. inet 8 (105 fi 7) poses median oct tubercle In P,precara siubly presered spent show the tubercle o contin fur tiny pis aranged {outline comer of agua. A sitar rangement of pss ao seen onthe octal ing of. forme Ie (P10, ig), Buti thi species there no ociptal taker. Such a feature has bee fermed {median social organ by Whitington (1963, p. 297) connection wih Scuteliuds suc as Bronte Sind Reymond and it also known from several ther totes, among them eran odontopsutis {Whitington 1956, p 177; Burt 1959, p. 220). Mascle sir. Canis of rela and P-frachuonss show smal sighly depresed smooth areas alg the matin ine ofthe onal gabe lobe, These are hee ered frontal glabllar sar). 19 1 precr P10, ig 5: PL. 10, Bp 2) single car occurs onthe Fontmos part of the gabe abe, srterens iP ornebeni (P17, i, 2) tent tree beaeste oval cas cock along the median ine Inboth species thee sas are accented by adaption the ane ine pattern surrounding them and Plate 10m ig shows hows in P prevlara the ano ns Joo eros the ‘On the posi of P.precar (PL 106, bg. 1) hee are smooth ares atthe ner ends of ach ih ring furrow and adjcent fo he dora furrow: thee ae nerpreted ws areas of muscle atacient. Tis ‘Ssupported by apypdiumot Phifipietia pind, (108, 12) tom te Lower Joastop Formation, ‘Vistergitland Soden, bere ilar muscle eas corespond to raised areas on the internal mould. This ‘sda ale sa driv outpost acl ridge of the racks, which sms tobe» goer charctrisic Ia is soon evn on a doves ecobeletn, tua ight rosie area (FPL 108, Sig 4: 108, Ags. 8.0), ‘similar mses pattern was figured by Whalington (196, pl 7, fg. [)on the pygiium af Raymond ‘ips recat Anterior and posterior pit The new species of Pilipinla show two prominent pits in th dorsal furow ‘terantia, termed anterior and posterior pits by Kiclan (1960, p. 7) hse have been dented on Aha ‘terial by Whitington (196, p. and Tashan (1970, p36) and eens by Dean (1973, p. 31) onthe Roloype fF. bores deeibed by Kurmmerow (1927, pl, ig. 7) from an PAfenig esti Boulder in ast erm, “The anterior pit whichis known on certain cwtellids suchas Signa (ee Skeet 1955) and Raymond api (eee Whitington 195, p35, 9: pL 9, 7), overs along he frontal label lobe and Inve where the anterior branch ofthe facial stare erones the bowder. 1 is especially prominent on DP fornehueni (PL 107, I), ks $00 P.preclara (Pi 104, fig. 1) bat in oth its aoe ith {diet weling onthe fontl pata foe wen sewed fom atbove-On the ventral surface the dorsal ‘rontelcton the anterior pit orresponds fo «sigh bos which prouaesYeaally and is covered by the ‘Toubare ofthe fe chek. There is no corresponding Feature on the external surace ofthe double 105.60. 6. "The posterior pt occurs inthe dorel arrow kel with the anterior eof the palpebral lobe wher the sibel becomes consisted. Ualitethe anterior pt which oval, the poster pits narow and slit Fie’ On excrnal oul ths sist to se, but fom sts positon might ie bncath the ote ip of the anterior wing ofthe hypostom, and he some fem of atachnent pot. The reconstruction given for the ental def the cphelon by Kila (1960, p74 fi. 19) sis mleading. Furthermore te tales (. 75 that p | antror pi i stuated at the eel be mos ane portion a the eye (whereas on her {Ext 8s shown a Frnt ofthe ee), ap (posterior pi) ite infront ap TG fact tis behind p's frat canbe judged, however, he poston of the pits shown on her gure (oe. ct, fg 8) corresponds to that observed for P.preslara, formbucns and ths Exiemal exeskletal surface. P.prclara and P.forebwons have een collected fom fesh,darksrey limestone and have note itl prepartion excep oe removal of matrix fom furrows and adjacent wo Stare ines. The dastratd tte shows details of excell sculeue of ps, granulation, and rs lines, of combination of ll te ype, which are now described for P prclara is, dens regular network of piting without associated aise ines oscurs onthe anterior order of {ranidia andr concentrated around the base of te ontal label Jbe (P15, 2), Lately, the ‘conseneatton dss and ajacent fo the anterior taal suture tbe border smooth (PL 104 fi. 3. Gabe outer prt ofthe palpebral lobe a larger, more regular pth oecas in sre of supra es,