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.


, 5060. 
, 
1970
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, 
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, 

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35

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, CO
2 . ..#
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. , , 


, 
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.

,
.. ..
1970 . 

, 
.
90 .. 

. 
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: CO + H 2O CO 2 +
+ H 2. Carboxydothermus
hydrogenoformans, 
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36

( ),
CO H2 ( ..
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9 2013

,

. 

,

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( 5060 7075 
), 
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Thermococcus 4 

. 85 , 
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: 
45% 
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Thermofilum carboxydotrophus, 
.

.
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, .. , 
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,

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.
9 2013

Thermococcus, #
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.

Thermococcus , 


Thermococcus 4 ,
CO H2 . #
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37

, CO #
CO# (coos) # (fdh) ( ..).
, .
, 

, , ( H 2 + CO 2) 
, 
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, .
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Thermococcus onnurineus,
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[2].


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1970 .. 
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[3].
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, 
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38

#
.
..
9 2013

Geoglobus acetivorans #
. ...


Geobacter Shewanella. 

Carboxydothermus ferrireducens,

(
). C. ferrire
ducens [Fe(III) + e
Fe(II)]
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( .).
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(FeOOH0.4H 2O) [4]. , 

, 
, 
,
Fe(II).


Fe(III)

. 

, Fe 3+.
, 

Geoglobus acetivorans, 
[5].
, 10 (

), 
. 
G.acetivorans 

. ,
, 
Fe(III) 
9 2013

. G.acetivorans 
,

.
, 
.

Meliori
bacter roseus, ,

.

Ignavibacteria [6].
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. M.roseus 

. 

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,

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1990 . [7]. , 
,

Desulfurella;
Desulfurellales.
Desulfurella 
,
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, , , 

.
,
39

Desulfurella acetivorans ,
: ()
.
( 
), 
.
Desulfurella 
10 1 3. 
,
Desulfurella D.acetivorans, 
,

 
.


: Ammonifex thiophilus Caldimicrobium
rimae [8, 9]. 
. 
, 
A.thiophilus,
. 

, .

40

, C.rimae, 


( 
96) 
. D.acetivorans, 
Brockia lithotrophica [10]

( 
) ().

:

,
,
,
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2H + + SO 42 + 3H 2S [11]. ,

, ,

. 
Thermosulfurimonas dis
mutans 

, 
2 
.

, 
,
. 
T.dismutans 

.
, 


,
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.
9 2013

Thermosulfurimonas dismutans #
, #
: () . #
...





.

***
XXI .

, 

, 


 .

, 
. 


.

1. Sokolova T.G., Henstra A.M., Sipma J. et al. Diversity and ecophysiological features of thermophilic
carboxydotrophic anaerobes // FEMS Microbiol. Ecol. 2009. V.68. P.131141.
2. Kim Y.J., Lee H.S., Kim E.S. et al. Formatedriven growth coupled with H 2 production // Nature. 2010. V.467.
P.352355.
3. .. // . 2005. .6. C.581595.
4. Gavrilov S.N., Lloyd J.R., Kostrikina N.A. et al. Fe(III) oxide reduction by a Grampositive thermophile:
physiological mechanisms for dissimilatory reduction of poorly crystalline Fe(III) oxide by a thermophilic
Grampositive bacterium Carboxydothermus ferrireducens // Geomicrobiol. J. 2012. V.29. P.804819.
5. Slobodkina G.B., Kolganova T. V., Querellou J et al. Geoglobus acetivorans sp. nov., an iron(III)reducing archaeon
from a deepsea hydrothermal vent Int. // J. Syst. Evol. Microbiol. 2009. V.59. P.28802883.
6. Podosokorskaya O.A., Kadnikov V.A., Gavrilov S.N. et al. Characterization of Melioribacter roseus gen. nov., sp. nov.,
a novel facultatively anaerobic thermophilic cellulolytic bacterium from the class Ignavibacteria, and a proposal
of a novel bacterial phylum Ignavibacteriae // Environmental Microbiol. 9 Jan. 2013
(DOI: 10.1111/14622920.12067).
7. BonchOsmolovskaya E.A., Sokolova T.G., Kostrikina N.A., Zavarzin G.A. Desulfurella acetivorans gen. nov., sp. nov.,
a new thermophilic sulfurreducing bacterium // Arch. Microbiol. 1990. V.153. P.151155.
8. Miroshnichenko M.L., Tourova T.P., Kolganova T.V. et al. Ammonifex thiophilus sp. nov., a novel
hyperthermophilic anaerobic bacterium from Kamchatka hot spring // Int. J. Syst. Evol. Microbiol. 2008. V.58.
P.29352938.
9. Miroshnichenko M.L., Lebedinsky A.V., Chernyh N.A. et al. Caldimicrobium rimae gen. nov., sp. nov., a novel
extremely thermophilic facultatively lithoautotrophic anaerobic bacterium from the Uzon Caldera, Kamchatka //
Int. J. Syst. Evol. Microbiol. 2009. V.59. P.10401044.
10. Perevalova A.A., Kublanov I.V., Baslerov R.V., BonchOsmolovskaya E.A. Brockia lithotrophica gen. nov., sp. nov.,
a novel anaerobic thermophilic bacterium from a terrestrial hot spring // Int. J. Syst. Evol. Microbiol. 2013. V.63.
P.479483.
11. Slobodkin A.I., Reysenbach A.L., Slobodkina G.B. et al. Thermosulfurimonas dismutans gen. nov., sp. nov. a novel
extremely thermophilic sulfurdisproportionating bacterium from a deepsea hydrothermal vent // Int. J. Syst.
Evol. Microbiol. 2012. V. 62. P. 25652571.

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41