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CMYK

14 MM

je L NGO ,dG Gg U CGJ fCG H ,QdG jY


e N c dEG e J j jG ddG g N eh .jG ddG
j dch ,SG bdGh qdGh dG M e jQhV OGe e L jN
100`dGh .JLMG ad HGJ e j e PJG EG aVEH N c Xh j
q Jh H a ee T V J L e j dG N jJ
T N e j Ggh .dP e AH J CG hO ac JLMEG
IAdG H J g JG Th .IAdG e Gk L dY LQO Y H a qJG
H dG dG Gg dCJ EG aO dG SCG MCG EGh .fedG SGH bQG
jhQGda ,aOG KCH q M Z OGe e IG Af dSG KEG g BG jj
M Z OGe e Cf IG qCG dc Yq jh ,Lh Y dG OLH eDj jOGh
jG H gG JG Gg OLh O q dh .f AJ e bGdG aOG KCH
b q dG Gg j CG h ,Y j QN f OLh Y j fedGh
e qdG dGh .dP Y c dO Mh fedG OLhh ,aOG Lh
JQHh H eDG Sf Jh fS dH EG SJ g dG Gg dCJ
J h d Sh dG Gg j CG Yh ,G Y gedGh IOhqdG
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jdGh jOQhGh jdGh SdGh fcdGh fhfGh ShdGh jddGh HdGh
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dG OLH dG Gbh hCG fjOCG Y hdG J ,dG OLh M dHh
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jBG AV Y dG g c dhJ dG fEG YVG L T
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SCG N e QdG gP DhdG hCG d kfe J jH fBGdG jBH
YLG dG L AGq d c JdG YG dG dGh dG
e QdG q dG FGhdG SCG Gg .Jh e J CG dG jGh
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.H YbG ANEG e Gj h dG g JMG dG FH GhKCJ
e dhj hCG Oe H dDG c e Zh dG Gg GC j CG Qd
Ie bG ha e IOSG Zj jdG dhCG eCG .YL be N
.jNBG EG HQ Y dGh JYfH AOEG e S fEG PEG Gk L
eN j dG Ld c dG dG g VYh IAGb gG CG EG aVEG
hjj jd f dd ,bEG jZ SCH dG g FG VY . jd
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id ,dG jf IOLG dG g e P Y j CG Qd G e
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dG HQM jnsdG sd Ibe icP ''j''h ''hQg'' SG e ,QG hCG dG SG j
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Y J dG YGG c HEGh cdGh jOEG SCG f ag JdG Vh dGh BGdG AV h
aGgCG Y YEG H c H e dG G Ggjh ,INCG G c d d ,jd jOG cG
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dH EG Y Jh ,SdG EG fBGdG dSdG f gh MGh g M dDG c L QhJ
.NBG dGh dEG OLdGh fEG YVH dGh
EG GfEG eh ,jeCG EG dG e AGdG e SGh jd Ic H j hQg c J
fCGh jfEGh fdG EG c H LJ bh ;fSEGh jRGdGh SdGh GdHh fhfCG
.fhfCGh ShdGh fdCGh HdGh jOQCGh dJdGh
dG SCa ,eDG EG jJh ,dH EG EG eDG Z IYO JFa dG g KCG d
CG Y G e .QdG Y dGh jdG KCdG fe kFf j dG g H J dG Gh
MCG H dh .jOG dG GfCG e f C ke j CG L H gG jh dG g GC j Qb
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.j hQg c EG Oj dGh dG MoO b dG OLh GH J dG LdjjEG L EG
g dG g f AGQh e JdG gh ;MVhh BGdG M e Ie FG g CG T
fCG Y IhY ,IdG hCG dG dh ,dG EG Ud dG jdG Y j jdG dhCG eN
.g c f AGQh e Oe g Lj
Hbh YCG J dG ,dG g IAGb Y jNBG j jdG Ea ,FG g AV Yh
.H QJ eN ej ,d jOdG jW EG gTJh
Va EG fEG OJh QdG gP jdG e kYf J dG dG bJ j ,iNCG L e
fCG VGdG a dG g eCG ,Gh bd e ,SdG b e dG MGREG KDJ h ,LdjjEG
dG edG dG e cCG Jd HOCG IdG Y cJ fc d QdG Y dG KCG Gg d J
dG jJh dG g g j hQg d MdG dG CG ij CG dH j eh ,dEG j
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f EG d Gg ej eSEG dG j dG YGdGh dG dG CG EG IQTEG e H


EG e EG IOdH EG j d Ggh ,IadG LdjjEG f LdGh G dG jO Y OHG
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f b ...EGh FGOCG e H LGh g ,QedGh OdG jhg f H J dGh jOG dG dM EG dHh
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dG g S dG Hh ,FdG QhdG Gg JY Y NCG b j hQg Y EG :b GPEG df
.dG BGdG H Yh dG IOdGh dGh dG dN e jdG dG T S dG SdG
,dG Y jhQGdG JL dG QGdG ,fSGh jOdG ,jG SG dG :JdG YCG J
,SdG G dG ,gQEG aj SEG ,TdG :jhQGd jedG LdjjEG ,TeCG Y YdG
fehdG :dG S ,3-2-1 YVG ,BGdG nb ,ScddG OGM AGQh ,gQEG OGM AGQh
QdG jf QfG ,dCG hC ,IFdG CG , ,QdG jPcCG ,QdG YN ,dG EG j dG ,2-1 FM
,GdCG dG RYEG ,gdG dG ,Sj dG ,Se dG ,jQdG Y bO HLEG ,kGDS jY
e FGQ Ph dG H ,dG dG ,dG jW BGdG ,dG Gg IM M ,e c dG
,IQdG Ie ,QdG Mh OG ,gJ ,dG N ,ka CGH b jedG ,GG Y dG
IG ,dG IG ,JdG G Ie ,dG IG ,YG RG Ie ,G IG
,GG e S ,G QdG ,Id G UCG ,dG IG ,dG f IG ,VdG
. DNA QGSCG ,JhdG IG ,FdG cdG AGG IG ,dG oj dH
e ,dG bG ,GG Y MQ ,dG MQ ,dG N Ge :WCd JdG c h
,dG ,QdG FbUCG Y ,G dG IM kYS 24 ,fEG L GG ,2-1 G dG
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bNCG n pdG ,BGdG SSCG gG :fBGb YVe hJ dG iNCG JdG YCG Jh
fMhdG dG ,G :eDd G ihCG ,gG g ,3-2-1 Ed jS a ,BGdG
dOGh dSdG J ,dG ASCG ,aG QGSCG ,BGdG aG T ,dG S IdG ,BGdG Y ,BGdG
,dG :oG fEG hY ,SdG ce ,QdG H ,BGdG e HLEG ,BGdG SSCG gG ,BGdG
BGdG MCG ,kGHCG J ,dG j ,BGdG YdG gCG ,BGdG IdG ,dG J ,gG jO ,KdG
,gGdG dG M ,BGdG e eY Qe ,BGdG dG gCG ,gG fEG T ,G
G ,mGB Y dG ,dG SHc ,dG N ,eDG MQ ,SQ J ,J CG b ,eG EG
,jQdG Y SEG d VJ dG GAGaG ,Sj SQe ,3-2-1 dG dL e Y ,BGdG IG
,e c j dG ,BGdG QGSCG H ,BGdG dG j c ,f J GP ,dG c JG gCG
.BGdG EG j jdG dhCG ,BGdG dGh dG

jG
8................................................................................................. NG
14........................ edG IdGh jjdG G :eLCG FdG cG
58.......................................................................... bQq dG IdG :IG jEG
74................................................................. ddG d SG jG
86............................................................................. eLCG dG e
90.................................................................................... qjdG a OdG
116................................................................................... G fedG
134........................................................................ IMGdG G NGO HdG
158............................................................................. G NGO jdG f
172.............................................................. qdG qG UGGh qdG
188....................................................................QG YdG
q :jdG cCG
206................................................................................................... G
210.......................................................................................... Qq dG jN

dh ,G Y IjY ha jd Y cT Qd
dEG j jQ J V J cdG g qCG fQYG
,feEGh gDG Mf e H a j BG e
d j CG dG eh ,Oq g d j G dh
AfCG Ie hah Th
o IQGOEG h fEG cGeh fe cdG
.ac dG
cG HCG b GfEG cdG a qGC G S Y Vd
d HSGh ,GfEG FHdG W Y jeCG FdG
dG QGUEH (IQGOEG ) jeCG cG b dG
QhHh ,djEG jdGh HCG b EG UG GH eRdG
jfEG dG
q jRJh dhCG PdG H dG Gg b
g M CG Hh ,g G id dG OQ ae Vd
cdG g e EG dG QU G H PdG
hdG EG aVEG ,PdG g fEG hdH dG OLGh
.IjG H jd Y jYO M
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q T H
OLh Y ka ,dG LQO YCG d JG FSh
YEG Xeh dGh SGh jjG H eJh J
Jh eJ OLH EG dG j h ,dEG...jdG Xeh
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dh e d NBG kLP iNCG Ie Qdh


dG Gg CG Vd ,HdG PdG e H cCG
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e
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q
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JdG Y ADg e MGh c e JG AGLE Gk e
j CG Y j eRdG dG FYE G
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dG gMCG e j CG :G S Yh
hCG j CG hCG dG IJh j CG e e j
Gk e q K CG Qf Ggh ,G PdG e q j CG
,dG AfCG L edG OGaCG bGj kjce
Gh ,HGh bGGh dG qdj G Ggh
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OY j Cc ,HdG
q G M e Sdh
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cCG dG OdG Gg q j dG G dG dh
e cc dCG 15 g qCG CG ,IAch IQe
J VdG CG CG ,cc L j cGdG g dG q dh ,cc
ShQe ah j kL ADg CG kjCG Vdh ,fEG jJ 100 d
Y d eG q dG ah dPh hj CG Y j e aj Lh ,eeh
.G JdG

10

j hQg

GE q dH Yha Hh Y cd FdG cG H eq dGh JG j


YE G AGNh SGh hjGh IQGOE G AYC a ,kGL Ic LQO
L IOLG qJG T eGC ,Oeh e JG Gfj GC j
..Y cT cC JG T qGC e GqG j kGJ cGC a fE G

,JM ka OLe Pd dh ,J fEG Y j G Gg qEG


fEG L g Pq dG Gg qEG ,kGHCG qbJ hOh QGSH jh e Gk L jb H
.M N jJ 100 Y j dG
ah eLCG NGO qJ FXdG e jG qEa QdG g hDhJ eY fEG
dch ,eLCG AfCG IOLG jG LMG L CH bO HM
11

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dch ,e HG FXdG RE jG EG eRqdG q dG QGUEH G dG


.IM Y N c e L aJ
dG UN f q Kh ,RcdG S EG dG jN QdG g IAGa
e bH j dGh ,e ie Y ajh dG OLG dG f j
H eJ dGh ,ddG e e f Y dGh ,IMGdG bdG
H J dG jG FXd IjY iNCG eCGh ,IMGdG bdG dG e e f
gh ,dGh dG jZ qJd f N e H a Jh eH G ILCG
,N jJ 100`dG GP G ILCG H eG OJ q Y j
q Fc f
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jG H edGh fG dG RG e hdH edG q dG j
qE a FSdG jEG YH J dG ffG H dG RG T dh ,G
Id j KCJ dh AH fCG CG OdG jdG Y FSdG OJ j edG dG
.WCG
J Q Ic M eeCG d fEG L ILCG Y c J dh
jG L da Y JS ie Y gMCG dCS a ,SdG e c GQOEG Y
FG e cCq dH aGj GG Gg dh ,Y ke e fCH eJ Yb Y HLC
.HdG dG
j :dP Y eh ,kFL EG L da j CG j fEa
GSH e kT j CG hCG fd GSH j CG hCG bS GSH CG fEG
dG FjdGh FdG ddG e BG dG j dh ,jj
JM dG j fCG j e Ea dG Gdh ,JOGQEG f QN L NGO
.kL kja CN Jj G L fh
QX dSG J iNCG M eeCG S dG Gg d JAGb H
AYCG H Jd bO f OLh c Y f CG H kUN ,aOH IG e

12

j hQg

g e ZdG Yh .AYCG g V q G Z OGG H eJ OLh ch ,L


Z OGe e IG N gMh aOG qCH jhQGdGh jOG eDj WdG G
L dG HCG q dh .ATCG N Y Lh Y dG IQH dG eDj h ,M
IgH Ff EG UJ b jdG dG
FGOCG ch G FdG LCH J
n r oG Ao n r SnC G pqd hn }
H OLG aGq dG qEG .jG Jdd
n jp sdG Gh Qo Phn n pH o o Yo Or an
j Mh fedGh G jG
ppF n r SGnC pa nho p rjo
AGOCGh IG QX feEG Y KE
,aOH jG Jdd G FdG
Gof cn en nhr n r o n S
EG S b dG Gg Ea dd
|on r jn
dG dOCG OGjEG :hCG ,SSCG ag
180:GYCG
jdG jOGh jhQGdG YGe d
Y ie KEGh ,aOG IH eDj
jHh JQb Hh ac dH eDG EG N LJ :dG dGh ,LGSh gJ
.Od jJ YhQh N Lh U
dG Y c Y YWGh eOdG d JAGb N e - ke Sh
.ATCG N J dG IQb Y -a h fEG L edG

13

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IQGM LQO fc e 37^5 h 36^5 H HK eLCG IQGM
G IQGM LQO Fa J q GC h ,a Lf dG SdG
fEGh .Jb Qh IN Ye EG ODj kYJQG hCG kVfG
je LQO 0^5 OhM EG L IQGM LQO J dG
,fRGe IQH G ILCG Y H MGdG dG N
jedG YhCG NGO dG V Y G j f dHh
Gg ch ,eY jG Y IJh Yh dG AG f Yh
j dG N e j G ILCG Y RGdG
.RGdG Y d Ie IQH SM
q b G ILCG Y RGq dG Gg qCG Qdh
IQGM jRGe Vh Y j dG g a ,kYU DhGLEG
je Vh dch ,fEG L AfCG L IjY
Vhh ,dG ac Sd jedG YhCG NGO IjY
,dG V Sd jedG YhCG S Y Ic GehQH
bGeh jG Y YS Sd IU je Vh dch
ILCG g e IOQGdG fdG ac L dHh ,FGOCG
QG BG SG RL EG fh BH J dGh jG
HG FdG EG UdGh eRqdG HH d Gk L
. c dG GQGdG PJGh
PJG j H dG GQGdG PJG j h
g J Heh IOQGdG d kah eRqdG HGq dG
GH G jG b e gJ c Hh ,HGdG

:eLCG FdG c
IdGh jjdG G
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q T Hh .eCG
L qCG H ,dG LddG FSdH adGh dG
q jeh IQGG jRGc
g CG CG ,kGL Gk Qeh Gk L kbGQ bGGh Sd ef IOdG eh ,fEG
eRdG eG dG V jeh G IQGG Ge e BG
.{SeKdGz Yj G e AL g SG Ggh ,ed SM EG SJh

(jjdG G) SeKdG :eLC G G


FdG Qhq dG j gh ,edG dG Y dG G {SeKdGz j
FS eh G e IOQGdG G SH j gh ,fEG L RGdG Y G
LQO K Y G ke dP e , c dG dG REGh G AfCG
EG LH QdG Mh G FG RGdGh dG V Y Gh G IQGM
,dG M M QH M jh ,ITe q G j gh ,IjY iNCG FXhh dG
cGe e OJ dG eG gh ,dEG H G eG e c AL jEG jh
LGdG GAGLEG Hh RdG dH jh ,ac G AfCG h G OLG G
qG G AYCG EG eGhCG QGUEGh G gJ RdG GdGh gPJG
G) jjdG a ,kGL a eCdG j q e f g eeCG hJh .dH
IMGdG Gh ,(kGQT G) jQT jN Y e qdC J (SeKdG
G IQGM LQO ae J h ,fEG dEG j dG dG QGe ae J
G AL HCG EG IOQGdG G f QOdG eCG J He J h ,G
qfC c dP c H J (SeKdG) eq dG e AG Gg dDJ dG jG d
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16

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.f IAdH edG q dG J a IAH dG fG


AYCG FXh Y IdG gYj Ac Ye jjdG dh
FS EG jjdG G gq J dG GQGdG ac jEH YG Gg jh G
dH UG GG J dG V VfG Y G S a ,G AfCG
FdGh
q QdG AGLEG PJH a jjdG G EG jQdG eG jEH
fhe EG Gg gQGb jjdG G Jh .dG iG EG dG V ad
.AdG
QGUEG jh ,eCH qG G EG dH RdG eCG QGUEH hG Gg jh
QdG eCG Gg eSG H jG Jh ,QdG Yh jG J dG dH eCG
.G iG EG dG V ad RdG Yh a AL e H
edG dG Y KCJ d dG AG g jjdG d AdG hGh
e Ic LQO Y eG OJh Jd f gh .{edG IdGz`H Yjh
J dG e dG Jg Cch ,edG Iq dH jjdG G Hj IAdG
Y edG IdG UN jjdG dh .ed dOJ AKCG ffEG cS

17

edG I`e

I idG gCG GP fedG e jdG RGaEH IdG g a edG IdG


.jjdG G GTEG fEG
SQEH dG Me GRe d jjdG G J dP Y eh
,{dG eg RGaEGz`H GeCG dG g Jh ,edG IdG EG UG dG
.e G QGH dG eg RGaEH edG IdG J dHh
QGUEG j IG g dh ,jG Y IJH j fOQhCG jb NBG eCGh
IdG gh edG IdG EG geCG QJ jjdG a ,Me Y eGhCG
Y YS IOjd RdG edG RGaEH bQdG IdG Jh ,bQdG IdG EG geCG QJ
AH UH edG IdG EG eCG QGUEH SeKdG j dc ,G jG
fedG RGaEH (d e feg RJ dG gh :jdG) jdG a OdG
H edG IdG Jh ,UG Jfeg RGaEH Ad G OdH j a eRdG
G EG ODj NBG eCG QGUEH eCG
edG e
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dG fedH Fb OGYEG h
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bQdG IdG
bQOQG IdG
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eg RGaEG EG ODj dG edG
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18

jjdG Gs
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IM feg

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bQdG IdG

ACTH

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dG

jG ddG J e IjY e J gh ,ITe eq dG SGC Jh dG q M M jjdG G M j


G g OD J GC Jh ,G fGE Y Jh G Y Jh ,jdG OdG f Y GTE Gh ,G
g fh .jjdG G Y J dG OdG g qj YC G dG q dGh .fed IRG OdG GSH
dN OLh Y bY d kq M kdO j ,IQGh GC dG Z Qj Qe bY fGE f j dG IdG qdG dG
.OLdG Gd

edG I`e

e Yf RGaEG Y jG Y TG NdH jjdG G J fMCGh


jg RGaEG j LG Yh ,edG IdG fedG jg N jh ,fedG
:g fedG Ggh,edG IdG e fedG
dG V aQ G ODj dHh Yhd e eg gh hCG SHhRa `dG eg
.SJ chCG eg
M jh jd GU jjdG G gRJ GdG fedG Ggh
bH Y jjdG G feg J h ,eCG VM 3 M e MGdG
J keY feda ,G NGO J dG jdG H a G fedG
G feg eCG ,G dG ZH M IRQdG IdG e Gk QYG kjW kjW
edG IdH Jh jdG YhCG e Ge IY EG jW W j a jjdG
.kGHCG GQhdG RL Nj h ,INCG c
J EGh ,a edG Id IG fedG RGaEH jjdG G J
IdG J Ggh ,kjCG edG IdG f jEG Y dhDeUN feg RGaEH
.IdG g f Y edG

j hQg

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.CG ke He cJ EG BG M EG GUj dP

39

edG I`e

c J CG QU Hb jG e Ya ,dEG IG eeCG hJ gh
M fEG jG g Jh ,LQG dG jh kjM Ode LMG
dG Gg jh .CG M gh dG FdG GG bQCG LfEG Y Y
Fc jG e gc gh ,Cd G OdG Y J jN b e dG
.CG M fEH UN HM AGLEG Gk HCG J h bY Z
?kYfh kc Y IdG J ch ,?CG M RGaEG hdG j a PEG
dGh dG Ra .G dEG IG d j GDdG Gg Y GGh
ShQe jW Y CG M fE H e H eH j edG
.Weh U ee EG eh eeh
jh hCG cdG eg gh HdG dG SG cP UN eg g
eh ,edG IdG b e edG Gg RGaEG jh ,G RGaEG Y G OdG
e hCG MGG hCG cdG eg RGaEG e jY eGY g NBG fL
EG gGh Qe a bGh IQS eGH fG eGdG g J h .G

40

j hQg

CG CG ,dP e eGdG CG EG QG jf f dG EG IQda ,SCG


Y hCG cdG eg RGaEG Y EGh .G fE bc J eGdG g
c dh ,e IFa jG dG hb hO G Gg RGaEG C jd eh jM
d e dEG J eeCGh fGhCG b edG Gg RGaEG e j c?G Gg j
edG Ggh hCG cdG RGaEG eg RGaEH J jjdG a ,dG
fEG eH edG Gg jh (hCG cqd fG edG ) `d gh PIH Yj
.FHEG hCG hCG cdG
g ?hCG cdG fEG jEG Qj dG e :f j dG GDdGh
q
jjdG G Y KDj dG g {LhSGz`H Yj G Ia N Rj eg
RGaEG j dHh ,LhSCG fEG j dG IOh Yh , PIHeg RGaEG Y dHh
IQdG cM d eGdG Y kjQJ eGbCG aQ J IG gh , PIHeg
H QhH jh ka kT OGOj hCG cdG fEG qCG CG ,SCG f fH
.G fEG Y G OdG

edG I`e

,G jGH Y e j G RGaEG C bdG Lh VGh dEG GHEa


:gG dG Gg Y c kb eCf fYOh
c ?G OdG J CG hCG cdG RJ dG edG IdG j jCG e
?kM RJ CH OdG d Gk eCG QJ CG d j
fE H j bdG qCG hCG cdG RGaE fG fedG J c
?ILh Id jdGh QfG Lj dG CGh ,G
CGh G fE G g hCG cdG CG J CG fG fedG d c
?G fEG e EG kM ODj RGaEG e
c d j dG Ggh ,CG M fE SG bdG j NBG f Lj
e
q a ,G dEG RYEG Y NBG dc fH dG G ILCG Y
GQTEG SJ CG QU IOLG dG GWCG YVdG AKCG CG K VdG
,hCG cdG RGaEG eCG fGG dGREH gQhH Jh J dGh jjdG G EG
.G fEG Wf OGOJ G OdG CG CG ,RGaEG OGOj dHh
.GdH dG g Y iNCG Ie eCf fYOh
CG L J e T Y g AYCG gh ,CG QU M AYCG LJ
G EG djEG j Gg dG OQh ,VdG H j dG G
q Y a OQ J
J dG GWCG gh ,IQcG IdH J jW Y GWCG SGH jjdG
H e jjdG G EG G AYCG a OQ e dG GQTEG g jEG
dGh dG e dCG dG G gQhe Y bG MG e fjJ
H Yh ,NBG e CG EG hCG AeCG hCG IG EG jW J GQTEG gh .gZh
geGhCG QGUEGh HSG AGHEH gQa e J IQcG IdG jN dG GQTEG
ICGG QU e eOdG GQTEG g CG J h bY Z jG g dh ,G fE
M RGaEG Lh kjCG J h ,eCG K VdG e SG dG OQ JW

42

jjdG G

edG IdG

G OdG

hCG cdG
G OdG

dG Gd fh ,jjdG G GE dG JGQTGE dG jG H SJ eGC K dG AH Y


Gg OD jh {hq C G cdGz chdG eg RGaE edG IdG GE geGC jjdG G QJ dG
.G fGE Y C G QU IOLG G OdG GE NC G edG

edG I`e

hCG cdG RGaEG Lhh Gg dG OQ d kSSCG kLGh Y CG J h CG


jG Lh dG e PEG ,d LfEG G OdH UG G qfC hCG Mc
?KCdGh dG G LdG g bdG Zh QdG e dG
?CG QU G AYCG Vh dG e
?G AYCG GQTEG d IQG dG L dG e
?jjdG H IQG dG g jf HQ dG e
J J M edG IdG H J CG jjdG G jN Y dG e
?eOdG GQTEG
?edG IdG jN NGO G fE G edG cJ OhCG dG e
EG edG IdG e QG edG jEH j dG GQhdG RL HCG dG e
?CG QU
?dEG OdG edG KCd J dGh CG QU IOLG jG N dG e
Y AdG Y j cdG Y kM RJ CG CG QU j MhCG dG e
?d f U
.VQCGh GdG jH dG Q dG qfGE :SCG g Y dGh MdG GG
ch L Y j CG fEG SG LddG FSdGh dG QJ H
Y dG SGh kbaBG fEG eCG QdG Gg a bh ,bO cCH jG FXd FGOCG
GH G eLCG ILCG cJ c U jH Yh fEG N J dG IQb
.bdGh JEGh dG
dG MdG G qEa (j Y Jh fS) OLh hj jdG eCG
j dG dG aJ dG QaCG d GjJ ejh ,aOG :g H j
.GOLG d Gk Qe aOH Jh ,dGh AMCG H
Y kWb kdO J RjEH dh FN H fcP dG CG M Ieh
e Gk MGh Gk Y aOG LJ CG dSG eh ,jOG ADg YGe H

44

j hQg

jL M hCG jjdG G hCG edG IdG hCG CG QU e H e dG UdG


dG Gg fe qC G hCG dG YGb e dG J aOG c qEGh .edG
RL EG gZ hCG Oda ,HG IQdH Xh RE H EG H j bdG
eH MGh BG J qch ,CGj c J ILCG gh ,dEG... Jh GQhO
dG J dG IQH EG jh Lj CG j eG bdG dG Ggh ,Jh
.SJ chCG g ddG Ggh CG M Ie NBG dO gh ,QG QdG
RJh ,eeh bO f b e CG M RGaEG c HdG QdG MT
e j VdG qC G OdG G jc Y J Ic e eeCG
,YdG G b G UG j c dG M SGH OdG e G
c G RGaEG e IFa a EGh dG M EG IRG OdG e G jEG j Gd
M EG j c dh ,kjM OdG VdG EG j d fC HdG QdG MT
?dG
EG Gg ej M VQCG Lh Y fEG IM AH e ac VdG WCG jj
J EG ODj edG qC CG QU e G VQ Y SJ chCG eg
M OdG e G j qdG Gg N eh ,Od G GdH G dG
.VdG CG Gk e dG M EG jW j
e G Uh gCG J CG SJ chCG RJ dG jd c dh
fc d Mh ?eCG K G j CG VdG Y j EGh G EG OdG
dG J EG ODJ dG FdG cdG J CG d c ,kL dP J
?G GdH G
Yj c f Y fEG Mj CG j dG SCG Y e GDdG Ggh
RJ dG G jN e N c .fGh bOh JEH L ILCG Y c
g fCG dG Hc Jh fS cjh ,SGdG Yh Lh Y dG Y eeCG
:VQCGh GdG HGh dG Mh

45

edG I`e

s mS
n rdGnC o Qo G n r pe cm r jn r ndGE p
o o r jn s oKp VQnC G GE Ap s dG n pe n eC G o Hu n jo }
.5 :IdG |nh s o nJ

:dG AG f J dGB
FG RGdG eg
AG c GJ CG J g ?fEG L IOLG AG c aJ g
GJ CG J gh ,HhGh cCG jW Y eLCG EG NJ dG
AG c GJ CG J gh ?kej G QN MW j dG IFGdG dG
NGO dG c GJ CGh ,dG V GJ CGh ,fK c N dG IOLG
Ggh ,FGLEH ke JM GW d SdG g H gMCG c d ?fCG
M fG GPEG dG C kc AG j CG j fEG L C Gk L e dG
.G N EG VdG j Ga G G AG f e %10
L YCG Lj fC ,L AG f j CG EG j fEG CG H
Gd dGh dG UdG eCJ dh ,Y kH G g j bOh UN f
VdG Y hCG AG T Y Y J dG AG a .IgH Sg Ie Ld dG

dGE j dG OGG cGC AG j


%10 G a dh .fE G L
E a G OLG AG e
.d VY j fE G
j GC GHGC j fE Gh
jh dGE j dG AG c
bO f G g AGOGC Y
.L NGO j jd

46

j hQg

j UN f Lj dh ,G G EG
VfG GH Y dG AG f
L f LG G EG Vdh
AG f Sb j a PEG .G
L
?YJQGh VfG J j ch dG
FdG
IOLe Gk L UN M jN g
AG f Sb J jjdG G
dG
,QdG d JAGb N Mh fK c
dG
fEa AG f VfG CG Lh dh
dG AeCG
fCG Vdh .dH jQa GQTEG J
G jG iMEG gMCG Vh
dG jW Y dG e Zh AG d j
Jh ,jjdG G IOLG
.Gh FdGh AeC Gh
YS 24 GW dG AG f Sb Xh
,G M G g AGOCG j CG Yh ,IMGh d dh fG hO UGe
jG e Y ODJ a PEG .e fEG b e G g CG T
e Gk AL J dG jG g CG GGh ?GPh ?IG GW dG AG Sb e
.G jdG dG Y e dEGh MH ee ODJ jjdG G
GPa ,dG fG AG f CG Vd ,iNCG Mf e dCG g dh
fCG c dh ?j CG cdG fBG G jG e H Vh dG fEG Y
CG hO dG AG f aJ c ?dG g GJ CG cY e jG g
?QYG Y AG T GhNCJ
fK AG Gg LQEGh a OLG AG e dG J GhJ CG V M fEG
AGLEG Gg a d Mh ,AMCG Y kShQO GJ GPEG kUN G EG
a jjdG G IOLG G jG eCG ,kY Y S

47

edG I`e

edG IdG
jjdG G

jT

jQh

G edG IdGr
eeCG edG IdG

jdG YhCG

T eO AYh

jT

` kGe ` kjGC MGh , kGL Qe GE M kHK Y d j e PJGh dG AG Sb AG c d


LJ dh fE G b e dG e hJ G gh ,dG AG dM bG bJ hO kGQfh kd dG GE
.dG e IAH FGOGC Y J IU jN

RGaEG Y edG IdG q M ,dG AG f VfG e cCJ M Y AGLEH


jdG Yc J jL cJ j dG FG RGdG eg hCG ADH eg
dSQ jG gh ,dG IOLG TG EG edG IdG e dSQ j
IOLG AG jL dH jH J dG TG g EG edG IdG e eCG

48

j hQg

.iNCG Ie jedG IQhdG EG YLQEGh TG dG


QGUEG jjdG G jN a c :IOe SCG fgPCG EG QOJh
gJ CG J eCG dSQ Hc YSG c ?Y Gk L IH jc jN EG eGhCG
?dEG IQOdG eGhCG jG g J GP ?H AL e Jh jdG jG
IQhdG EG LJh Ic IQH dG e AG jL dG j SGG g H
e H SCH QGe HQ jh TG dG c J dG d ch ,fK jedG
AG f C ,Y dG J RdG QGG e cCG AG e c HT GPEG eCG .AG
,jjdG G IOLG G jG b e IOjdG g Sb jh ,OGOJ dG
dG e TG AG f J dd ch ,ADH eg RGaEG YS J dHh
ed iNCG Ie gh .HK J dG AG f CG CG ,dG c OGOJh ,dG
OGOj SdG Hh ,jeq dG YhCG Y J GMEG Y JQb gh ADH
MCG Y kb fc Y kdOh fEG I eCG U IG g Jh ,dG V
jPCG IOLe M jN a ,IG g d QHh ShQe J gh .J
JJh ,dG V SH jG g Jh ,dEG IOQGdG jedG YhCG h dG
V j eYh ,edG IdG M dSG dG dG jH G jG
edG IdG EG IQcG G jG e dG GQTEG SQEG j kW dG
8. ADHeg RGaEG e EG IdG g EG GQTEG g Uh ODjh
TCH J dG G QGfEG fCH G jG g Y J h
eYh (AGG TCG) FG Z AdG e keM J fCG g C ,AGG
FG Z AdG Gd Gh SG H JG j G g jZ L j
.fdH QGfEG SL GC j FYh
EG jG g e SQEG SG GPEa ,f cdG Y J G
q jGh
f j e PEG ,e jCG LJ h OY dG CG j da edG IdG
?Gg G QGfEG

49

edG I`e

jjdG

jjdG G

eg j
G ADH`dG
.jjdG

eeCG edG IdG

GQTGE J dG AG f J eY
FV Yh jjdG G GE jQGfGE
RGaE qdH G d dG jG GC J
GE qOD j edG Ggh ,ADH `dG eg
GC Hh ,dG TG AG UeG IOYGE
H dG q Jh dG QGH UeG j
.ADH. `dG eg RGaGE bj

G edG IdGr

jT
SMCG
eg j

QGfEG
jjdG G
OY

ac f O
G edG IdG

HG
ADH
jdG HfCG
dG

j hQg

OLh j dHh ,dG e Ic c j jT f EG fEG Vj eY


IM Y IQN j dHh NGO dG V VfG j Ggh ,jedG YhCG
G jG e SG GQTEG J dG V VfG Yh ,dH G
gQa eh ,edG IdG EG e SJ fc dGh dEG IOQGdG YhCGh dG jPCG
dG J EG ODj dG ADH eg RGaEH CGJh QGfEG Me IdG g NJ
UJ gh ,NGO edG dG IOjR Y j dG Ggh ,jedG YhCH G
.Y ch G QGfEG G dG Gg AGLCH J a eCdG EG aJ IjY
jedG YhCH G dG W J CG jjdG G j jCG a
?J dG ADH eH gKCJ ch
YhCH G dG J g dG V IOjd SCG aCG CG V c
?jedG

GE jedG YhC G fH h dG IOLG UG G jG e adH dG GQTE G SG GPGE


fGh ,dG V j eY j adG Gg q dh ,Gj e Y AT c GC j Ga edG IdG
ILC H eC G Gg J h .dG Ge GE dG aQh jdG Nd edG IdG GE OD j
TC G M aJ SG GPGE Gj e Y eC a .AGG TC G GSH J dG G QGfE G
.kGQa dH GC j QGfE G RL qE a (IQdG g c c S C ) adG Gg fG qeGC ,AGG

51

gGdG hGC aOd


GC aOH UG dG
edG h SE G e ec
AfGE OGeh FHdG SC Gh
WH e je GE iNGC
SM RL J GC hGC dG
..kGL Qq e

cGdG Ad S aOG C H Yf GC G e
aOG C H Yf GC e g c ,kGL IG G
.dG WH e je Qd S

j hQg

?YhCG H G dG d SG FdG cdG J CG d jCG e


Hj dG G QGfEG RL V g IQG JG T HCG dG e
H aOe Cj dG Ggh ,kgeh kbQN kJ eeCG EG ?edG IdH dG
ja ANCG YJ a jOG eCG .J HCGh IQU HCG Y N Y eeCG
Y Gg G QGfEG RL CfCG b G jG CG Yj YCG kJ H e
Mh SEG Y Lj VQCG H dG FYOG J h ,aOGh QdG jW
g d aOG KCHh ,UY g ICah iNCG AH Geh FHc SCG e AH
NGO SCG e EG kjCG aOG OCGh fK UY gh S Wf EG GG
RL SCJ EG kjCG aOG OCGh dK UY gh NGO FHc T qe
EG ,jdG AYOG Gg j CG M bY fEG C h ,NGO G QGfEG
fS dG OLd gQfEG hOjh GAYOG g hOOj GfG e jOG CG
.QdG jf Y YCG kYaO aGjh H ,Yj e H aj h ,Jh
dP J dG jBGh dG Vh VGh G dG dG OLh M qEG
:AH
o d oo nj sfE p an Gk reGnC nbn GPn GE p hp VQnC Gh G n s dG jH}
.117 :IdG |a rc

jG ddG H eGd G fedG


G jd fegh
IY dg CG G q dh ,gh hC JAGb Y IQdG g jJ
YRe bd IjY YS dg qCG AdG KCG bh ,eLCG J YS
e YdG gh ,(LddG) {jG YdG z SG Y GWCGh G AfCG T
j H Gk L IU YS gh ,a LJ dG AG H e M a bJ Y
G IOLe YdG g iMEGh ,{YShjGz hCG bdG YdH
q YJ dd
.9 jjdG

53

edG I`e

Nh ,dG K ddH GC J ,JM IjY MGe Nj fEG qGC f c


gh ,gGG Ia kUN LdH GJ EG fEG Vj MGG g
14-8 H AdG Yh S 16-10 H IQG IdG kUN Lq dG Y iJ IdG
.S
hO J dGh dG e GQG L IOLG LddG YdG g e iJ
e fEG j c M b bdG CG J CG YdG d ch ?Va hCG CN CG
?dH gGe EG W
Xh H e FXh IY jd jjdG H fYO dGh G AL g
fEG H Y EG jODj
QGfEG SL GC a ,gGG Me
,jjdG G fdH
edG XdG g CG j Ggh
.H dG GC S IOdG e Gh
eSG b YdG
q c qEG
e M a Y d AdG
ah fEG L j
b e kee jh e bJ
jjdG G C ,jNBG
,YS jCG Y G
S J e fd dh
FXh e Xh ODJ M jW
e aCG kJ AdG j
XdG g Y d YdG
.dG LDG

54

j hQg

G jN J c dh
?G AGOC M b bdG CG jjdG
Y c EG Oq H Uj dGh
ch Se bJ ah jG g
AGOC e eR H jG g H
.10 e Xh
,JGP H Ie G Gg EG
Gg jN dG j CG G eh
Ggh ,G GdG IG G
Y Y NBG kdO S dG
j eY .Od jJ YhQh dG
Z LdH YS Y jjdG G
G dG jGH ke QGfEG SL
fE G Uh e cC J GC SGH J Fe
h QGSH J YdG gh ,dG Me GE
`dG eg RGaEH CGJ jjdG
.fGE c L N hO
IdG j edG Ggh ,GNRH
Y TDc feg RGaEG Y edG
Gg h ,LH h FSH g fedG Ggh ,fedG jg RGaEG Ye M
dG LdG hCG ICGG L cEG Y hDe fC Ih bQN U fedG
SG cdG fedG jg qC Gk L e dCe gh ,IG eGh
j fCH Mj H fedG Gg jh ,Y GKDj dG AGLCG Y KDGh
dh ,ICGG e IOLG jdG fEG g FSH eg Xa ,fj e c
EG ODj f edGh ,LhSCG eg RGaEG Y G gh iNCG Xh
L g Xa LH eg eCG ,dG jJ Y LdG N jG
G EG aVEG ,d IgLh Wh IM G e J VdG jdG
L IQH j f edGh .ICGG L hLhdG eg RGaEG Y

55

edG I`e

edG IdG
RJ::IGC G edG RGdG
`dG eg IGC d edG IdG
RGaGE GE OD j dG LH
GE h hLhdG eg
IM G e jH hN
FSH `dG eg eGC ,W
f cG GE OD a
.G IOLG jdG

FSH
LH

RGaEG Y d jN `H YJ dGh G IOLG jG H j fC LdG


.{hSJ z`H Yj UN eg
LdG h ICGG e c L H FdG cdH fedG jg RGaEG EG
NG fedG j c ,dGh eCdG EG fYj H Y KCHh
hSJ eg RGaEG EG f edG ODj c ?ICGGh LdG H LddG
f edG j c ?ICGG L hLhH eg RGaG EGh LdG L
eG ICGG jh ,IG G WJh dG fc dLdG U LdG
LdG G G KCdG dP GMEG Y fedG jg eH e ?jfCG
CG ,f jd CG ,aOd ?IG G Gd HG dG Gg e Oj ?ICGGh
?fJ dG GQd

56

j hQg

edG IdG

FSH
LH

RJ::LdG edG RGdG


`dG eg Ld edG IdG
UN jN j dG LH
Y {fd jN zYJ G
,hSJ Yj NGB eg RGaGE
GE OD a FSH `dG eg eq GC
.dG fGE Y G jN

fG GQd h jd h aOG EG Oj Hq G dG Gg CG VGdG


Y GC Jh ,dLQ Jh LdG L Y GC J dG LddG GQdG gh ,jd
,ge IQb b e ke Seh efH Y J EG KfCG Jh ICGG L
.VQCGh GdG jH QG QdG dG T H g gedG IQdG g MUh

57

eGh fG H j dG FdG VGG gCG e qEG


G hCG G GMh e IMh a ,z LfEGz g jG
GMdG g Y YS dh ,e IJhh YH J CG j
jEG V g G Y d SSCG da ,gMh ac Z
YS J a ,H a fG h GMdG g Y YS
,AdG H ke kjf hj dP qE a gZ hO e IMh Y
GdH dG Gd ,c G f Y kY e Z dh
SG a j YdG BGh eG J Je g
edG d G dG e YCG IQGOEG UNGh
.d G LfEG EG kUh G GMdG H dGh
G Jh ,kbY ke iNCG Ie Qf BG fYOh
YS 24 fEG G jh G LG e jG
dG e OY G Gg jh ,fG CG hO UGe
LG CG T ,eY jJ 100 ,dG dH QJ
J d YCG IQGOEG UNGh SG e L EG Se
.dG e dG OdG Gg Y Gh QGhOCG jRJh ShQe
j qfCG qEG eLCG Lj dG G Gg e EG
e IU b g dP e kHh ,UNGh Se a
.dG G Gg IQGOEG J feg eh (e) dG
IQGOEG Y hDGh ,fEG L g QcG G qEG
ed gRGaEG jW Yh ,bQdG IdG g G Gg
L J N jJ 100 Y IQGOEG J chdG

bQdG IdG :IG jGE

edG I`e

g G jG Hb GE Lj dP Sh ,jG ddG AGOGC Xe YH dG L j


G chdG eg jh .FGdG OGG e Ic H bdG Y G Y IG fedG MGG
.jG ddG AGOGC YSh G jG d

,bW EG FGdG OGd jG j YS Y dhDe IdG g EG dG h ,fEG


j G S a ,dhj dG FGdG OGG e G IOSG ie j Ggh
OGa ,dY YH R fC jG ddG YS Mf e Gk e dG QW fEG
e dH j CG ddG fEG j NBG Hh ,bW EG Ic YH J FGdG
fEG q T q J dG J eh ,bTQ Y ka j dG Gd ,bW EG cCj
e Y CG CG ,fGH AG OGOj dd bW EG dG FGdG OGG QGe j H cCd
Gg Y IJh Y iNCH hCG IQH KDj Ggh ,Y IJh hDWj G Y
CG H a G Y YSh dG Y W Oj e Lj GPEGh ,c G
.fEG U ce LGj

60

j hQg

e fE G j
YGC GE j c ` e
J c Y ` IOL
.a edG AGOGC YS

jE HV fG a ,H SCH eLCG j HdG d P gh


jG Y Ya ,jhSCG jqdG e e a fEG L IOLG jG Y
d LQO EG dH fh ,dH YH gO EG dhf dG FGdG OGG Jh J
f EG eCG jf G j K ,MCG H jjCG M j e f
fEG j dG VG VGYCG g ch ,gdG EG ODj eCG Ggh ,bdG
.11chdG eg f e j dGh z Oe z Yjh
hO G jN e N c AGOCG YS j CG j chdG egh
jCG hO edG JM UGj CG fEG j Ggh ,dH HUEG h ,f CG
Gk QhO j H Jda AGOCG YS jG cEH chdG eg j h .ce
Gg e e QGe RGaEHh ,dG G Y jJ eY jG g YS e G Gk Te
AGOCG YS OGR GPEG j GPe iJ .Jdd jG AGOCG RGJ GMEG j edG
IOjd fh ,12bQdG IdG H G VH AG j ?jG Jdd jG
RdG Ga e dG V OGOjh G IQGM LQO JJ jG ddG YS

61

edG I`e

IdG Y G GE
HUE G GE OD j bQdG
Ve e IjY VGeC H
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62

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63

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64

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65

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jG jQGG G e jJ 100
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66

j hQg

e Ie T H dG gh ,JM QGS
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67

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68

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69

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70

e fE H eC G Qj
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72

j hQg

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73

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76

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77

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78

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79

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81

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a f JQG GPGE eGC .eG FGdG OGG e aJ GE aVGE jQdG OLG dG Gg e AL UeG GE h
.d UeG e jJ dG jG Lh dG ddG AHE fddG Nj Fa RdG G

UJ dG f g dEG UJ dG jG C Jh fS dG Ge cCG
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EG ddG UeG Y g bdG AeCG fH IOLGh UG jG
jG Lj dGh D edG C M Y eeCG Jh ,dG D ea OLH
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82

j hQg

Kd MGh e OLh j Gg .a IMGh jL LJ FSdG g dh ,dG ddG f ad FSh K LJ


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jL OjEG j eCG CG CG ,dG d M OjEG Y dG HK f Y d


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83

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eg KCH dG ddG f aQ
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98:W
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?dd UeG AKCG dG D ea EG jG jG CGh ,dG ddG
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84

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eg gh ,eeCG jL NBG eg j gh ,eQKGQdG eg RGaEG e J
ed Gk Oe edG Gg jh ,bQdG IdG b e RGaEG j dG {fdcz
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85

G LM e cCG jS Y jZCG dhJ GPEG


dG f JQG hO jh dH UN f GC j FY
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jQGfEG IdG) Yj Gd QG EG Ve jh feg
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b H dG dW e kdZh ,G gh N a H
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IOjR FdG Gg j f bdG h ,dG bdG e je EG dH kjODe jN
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Y Gj CG dGh d Ggh ,cCG e jG Y G dHh
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be SGH J Ye Ib j FdG Gg Ea gGdG H eYh
a IdG b e Rj dG FdG Gg qGE .Ye YH dH kYe J hCG ibCG
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92

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Y jdG IdG h ,kGL e feg fE H Jh ,Fc Gc jdG OdG J


.(QGfE G) jQOC G H dGh (jQGfE G) djQOC G IdH hC G Yj ,e j
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Gh G edG Gg OLh f e dh ,dEG LG M a kfh jh


dG c GPEG :JBG G jW Y HSG jZ f KCG b HCG qEG ?dG
a djQOCG QGe Ea e 2 Yh e 100 gb jFGO IH C eLCG QG
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Gk L c KCJ d edG e Gk L dG CG CG ,YEGh IG a j Ggh ,Gh
dEG GHEG e J djQOCG eg j ddG UG EG ,fEG L Y

93

edG I`e

.gOHCG H dEG IQdG eeCG J djQOCG eg KCJ c SGQO Yh .G


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V IOjRh jS H JY J EG LM j Ggh ,jdG hdG EG MG
LGj hCG YH hj CG j Ggh ,dG HV Xe IOjR EG LM CG ,eO
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94

j hQg

j .G AfCG FS J jW Yh dH djQOCG jL Jh ,djQOCG


V EG ODj KCdG Ggh ,jedG YhCG NGO e KCJ GMEH djQOCG eg
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dGh d dG e cCG c V dG GHh ,IQcG YhCH G
.24 dG dGh
dGh dGh G IOLG jedG YhCG Sj e djQOCG eg qEG
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j qfGC CG dG dG f OGOJ Ggh ,dG e cCG c aO Y jN j


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96

j hQg

G Ea dP dh ,kGHCG bdG J h G Y CGW dG gCdGh QGfEG


dhDe IdG jG g Jh .dH e AdG dH HUEG j
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97

HCG KCGh ,GdG e AL je IY e AL hCG Ib


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fEG je Id jdG IdG GRGaEG jJ EG LM gh
N dh ,hShdCG eg e a MGh GZ Y d
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hShdCG eg f J CG C Gk L SM
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jCG a Lj dG SdG f G EG AG qC dG
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Y hShdCG j jOdG cJ VfG Yh ,PqdG UN kee
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Y G j SdG Hh ,gP jCG jOdG fjCG AG jL Jh .kfK
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98

kGMGh keGZ Qj e Y a G q d fGE e 100 LGC OLG hShdC G eg c q d


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,a SJdG fjCG MQW dG e jOdG fjCG J fCG CG ,SJdG fjCH


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FdG H RGdG GME Gk L e (fG fjCd ) jCG RGdG Gg qEG
G SdG H RGJ GMEG ,LQN OLG FdGh G NGO OLG
cJ j hShdCG eg qGE .dG dG Y J EG aVEG YdGh
J AG Gg IOLG jGh ,(jQGfEG IdG) jdG IdG e LQG AG

99

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dG UG edG RJ CG d c dh ,dG YCG IOLG jG Y AT CG


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100

j hQg

e NBG FLQh ,QdG G Gg J c dG Y H J e C


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AT CG Jh fEG OLh J aOG Ia EG J QdG jf EG
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101

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102

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103

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105

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cdG qdG eq GC ,ICGG L e jdGh QdG e dG
q IOjdG N e
.G d q dG IOjq dG qj gGG QW N L OG da
e c h K j c N J g ICGG L UG dG
q IOjq dGh
.37VQEGh G AKCG ke G LMG j bdG
j gKCJ q dh ,IG edG ICGG U cEH q G fedG J
g jL hCG J c .38I fN GP JGUCG a Lq dG id q Y

119

edG I`e

U fG CG jG g J c ?ICGG Uh Lq dG U H fedG
LdG U CG f jG d ch ?ICGG d edGh LdG d
?kYf ICGG Uh kN
,H M a e Y q M EG ICGG L G fedG RGaEG Y qEG
Vf L EG LM g a ,jRGe fj fCd dGh G qda
fedG RGaEG Ea dG Gd ,OdG dG d TGQ Y EG LM g G
fEG qCG Y NBG kdO q qT H Ggh ,39 e jY Me H Y j G
.Y f V kjH kN N b

HSGC HQC G GP Id eq dG IQh q dG


OGSG) IdG g qEG,HSCG HQCG c IdG j fCH dG
q ICGG L qj
jL fEG SG LCG e G d ICGG G jN b e J (dG
.I EG ehb Jh
i Jh kjq HJh kU H q Jh JLM c q Jh gdh q C G qJ eh
jG a ,jdG f gG J q C d SdG AYCG jN qEa ,WCG
qEa T qHh .jdG J Y J c jdG iG Yh kq e d SQ e J
.iG jdG G Gg J kSSCG Gk QhO J q G fedG
Gg GC jh ,{LH z`dG eg RGaEH edG Iq dG J hCG HSCG HQCG jGH h
Ye j dHh ,GQhdG RL Y jG EG G e jdG MQ edG
.q M b jG
`dG eg KCJ M Yh ,VdG Z jdG e ?G Y jG
g e a IMGh q dh ,edG Gg KCH dH jdG g e b GC j LH
dG eJ jc G QN gRGaEG jh eJ IQH J CG J jdG
Jh .(CGJ IOh Jh MGh BG jH RGaEG dM ) qd IgL
120

j hQg

FGdG OGG Y H W jdG


Y dG gh jQhdG
, {jGz hG { folliculus jGz
IdG e Ob eH jG KCJ
e CGJh {FSH z`dG eg gh eq dG
YJ UN jL RGaEH gQa
.LhSCG eg hCG {LhSCGz`dH
RJ CG jG W J c
?H f Y e ZdG Y edG Gg
G qEG ?FdG RGaEG Gg e dG e
Y NBG dO EG fOJ SCG g Y HLC
.a G fedG cJ j GdG G
jG a ,G G Y hM
jL CG ,jG b e gRGaEG j dG
N hM Y NBG kdO j e ICGG L IjdG FXdG e d LhSCG
:j c FXdG g RjEG h ,e dEG
g NEG q a ,NEG q g LhSCG jL aJ e H e qEG -1
eg KCd a ,dGh
q fd a gh G
q jd G G
VCG 5 EG 2 QG kS OGOJ fGQa ,NEG OGSG GC j LhSCG
J dG g jdG YhCG gh ,jdG jeq dG YhCH WMEG q Jh ,OdG S
.eRdG q FGdG OGH G jdG jhH
jdG D q dH GC J eG Z jG qC dEG Ie G g qEG
gh ,dd WMG e j e PJGh jQhdG OGH gjhJ Y Jh j dG
.ke G jd Gk e j dGh NEG RdG OGYEH J dG
:dq dG Dhq dG IQKEGh kb bdG j gh

121

edG I`e

Jh JCd G e gRGaEG S jdG CG J CG jG d jCG e


Y dhDG g jdG YhCG CG J CG d ch ?HNEG H NEG
d G j qdG EG J CG YSG c ?AGZ e L G jdG jhJ
?jdG
q YhCG e OY cCG aJ
IOjRh NEG Y e q EG jCG LhSCG eg ODj -2
.40a G
q jdG QGSG H G eS Y aj CG fCT e AGLEG Ggh ,Jbq
ed Te KCH j gGG QW ICGG id QdG
q e qJG qEG -3
gq dG Oq GG a q a G g KCJ GMEG Y j edG Ga ,LhSG
OdG OdG S OGSG g ICGG QU dG Gg qEG .41G OdG KJh
.M e ej
,Iq G jfCG eG ICGG L cEG Y kjCG LhSCG eg j -4
,LdG L eCG g c d dG Y jN Y j ke edG Ga
j dG Mq dG e ICGG Y edG Gg KCJh .IG EG cCdH YJ dG gh
ea ,kGe kjfCG kT HcEG
Gg KCJ f ICGG id dG
q
e Oq Dj c j gh ,edG
Gg L c j qfGC CG ,q L H
.kYf kjfCG dG
q
LhSCG eg FXh e -5
jf a ,NEG Y J kjCG
Vq dG jdG J qdG SCG
eCG Gg IGRGh ,NEd JbhCG fCG
ie dG LhSCG f J
.Ld jKdG G gh dG
e UN
q FS RGaEG EG Oq D j Ggh ,Y

122

j hQg

EG fh NGO eG e Y IQdG jd FdG


q Ggh ,G H NEG
.Vq dG jdG EG dUhh NGO dG cM H j c NEG q
AGLEG j GPEG Ic IQH KGG Qf kFe kfe CG MQ j -6
PEG ,AGS M Y Gh q C G IM Y Gk N q UG
q gh ,dP hO j dG
iNCG Ie LhSCG eg QhO JCj g ?G Gg hM RqdG AGLEG g e
dG EG edG Gg jL Uh a ,G hM e Gk c Gk QhO j fC
Y aH j G SdG Ggh ,M IOq e RGaEG Y jN qj Mq d F qdG
.42IG jG Kd kFe kSh J Y ka HhG QNCG e MdG
QL J GME J Vf Z jM b e gRGaEG j Fc jL qEG
ddG S eRdG GOGSG AGLEG Y f bdG jh fEG L
RGaEG jh ,IO jL e GQd q g LhSCG eg qCG Gh ,jG
g q dh ,jCG bY Z jN Y KDJh bY Z jN b e bdG Z jG g
G e MGh QX qG Gg Y jh ,jd V J c QeCG
.FG e Y eeCG J CG q H a gGdG g L eCJ GPEGh .IG g EG
GQqdG d f d LhSG eg N e J dG G g qEG
cP e dG N dG dEG IQdG jBG e jBG g qfEGh aOG h jG h
:J b ,J MCG Yh fCGh
n o r pe sHn hn n Ln hr Rn n r pe n nNn hn mIn Mp Ghn m r nf pe on nNn p sdG o oHs Qn Go sJG o S s dG n jt GnC jn }
|pbQn r or nYn n cn n dG sGE p n Mn Qr nC Ghn ppH n odAn nnJ p sdG n dG G o sJGh Ak npfhn Gk pcn k Ln Qp
.1 :AdG IQS

jdG S D q dGh OGSG


dG H Y CG ,HSCG HQCG GP q JG IQhq dG e qdG SCG jf Y
,MdG H jdG MQ GC J FYh ,G e VdG jdG GNEG j Y HGq dG
123

edG I`e

Sa SHQc
dhCG jM
(UCG G) Jd SHQc

eO jT

Me jM
dG
Vf Z jfK jH N
ee Ya jM
jH
LQN Wl
G

Ie jM
e jM

.a G fedG cJ j dG GdG G

GPEG eq GC ,jL fEG IOh J dG qEa Mq dG g N NEG M GPEGh


G J eY MQ GC J jda .G QN J jdG qEa NEG j
.edG IdG gh Y Gk L IH IZ jYQ J jKdG G g qdh ,ITe
dd ,UN
q jYQ EG LM jdG qCG J fCch J eq dG Iq dG qEG
Y G EG edG Gg jEG q jh ,{LTHz`dH Yj VdG Gd kUN kfeg RJ
Corpus UCG Gz Yj G e AL KCJ CGjh ,GQhdG RL jW
.hLhdG eg RGaEH GC j Gd fh { Luteum
,LfEG e jZ IY gh ,jd bO J f g hLhdG eg qEG
Gg qEG .kGHCG G QN J dG {UCG Gz jN b e LfEG q jh
LJ j KCdG Ggh ,RJ dG jG Y GL IH jN a j edG
L c Y dO Ggh ,MGG Seh MGh V cG f qe jN
124

6
G e IOQGdG G
RGaEG e jjdG G
5
1

eg f OGOJ IQhdG g Hm
dG iNCG Iqe hLhdGh LhSCG

RGaEH J jjdG G
.RGaEG Y gd OdG EG feg

MdG

W IOe

RGaEd fG G

2
eg RJ eeCG edG IdG r 3
eg jG RJ
.LH. `dG egh FSH `dG
egh LhSCG
hLhdG

4
LhSCG feg
hLh`dGh
S IOjR EG jODj
MdG WG AdG

LhSG feg j
Me AH LhdGh
. dG

G
(UCG G) Jd SHQc

h FSH ed jjdG G RGaGE jh ,IGC G L edG RG J T


LhSC G eg RGaGE j eC G jf h ,qKD G Wd RG Gg SQ fG f LH
.IGC G Y dG M fG IQTGE Gj jdG hLhdGh

edG I`e

.aOG dh dh kb fEG
d He H CGj NEG EG j eY hLhdG eg qEG
eg Y Yj fCc GHh jdG S FG VQCG j fCG CG LhSCG
.LhSCG
e j KCdG Ggh ,G Y KCJ edG Gg j iNCG Mf e
OLh e iNCG jH NEG gh QhG j M G e iNCG jH hN
.ke Gh CG Y kL Gk N j T H Ggh qCG MQ NGO L
H NBG NEG hM e Lh hLhdG eg j c dh
e ?G jEG jW Y j dP qCG JjL J ch ,hCG NEG hM
gdGh IG EG T H YJ QeCG g qGE ?dG IG g jG e dG
.MGh BG YEGh
dG SJ chG eg KCJ e dG gh edG Gd iNCG Ie dg
jCGQ ch ,FGdG dG Y gTh iNCG bQN S gh,edG IdG b e Rj
Y dG Y MdG Y Y hDG g SJ chCG eg qEa kHS
KCJ SG GPEGh .eCG H e G hN J EG ODj fCG CG IOdG Oe GbG
NEG hM e hCG jCG N MdG Y Y SJ chCG eg
f kLQN d dHh MdG GQH dG e G jdG Y j Ga
MG g h .dG g GHCG j G CG CG dG d G dG
jdG d eh SJ chCG eg KCJ ed hLhdG eg Nj GdH
.G QN G
SJ chCG ea dN e j kjL AH hLhdG eg EG
ea ,iNCG Ie ke dh .eCG G fG g Ggh ,Xh AGOCG Y
b e Ra SJ chCG eg eCG G IOLe jN b e Rj hLhdG
.G NGO G SCG HdG IdG gh G Y Gk L IH IZ

126

j hQg

MqdG
da Ib

G
6

(UG G) Jd SHQc

IdG g NGO cG J GC Hh ,(1 dG) da Ib GE G e VdG jdG W j


(6-5 TC G ) fH NGO `f aJh MdG GE jdG J (4-3 TC G)

c ?SJ chCG eg OLH hLhd IRQdG jG J c


G jdG d MG dHh MdG Y J EG ODj edG Gg CG Y
?hLhdG ed gRGaEG SG SSCG CG Y?QG EG
AGLCH ec ae j dG Gg e Vhh ,kYQH kJ g CG T
fedG jg HCG dG dG QdG dG g G Gg VGhh ,ac fEG L
.e H
dG Y eCG J H j fEG iNCG Ie hLhdG eg
.Lh Y
QGL EG dUh e Gk QYG CG L EG dH kjZ kFc J G jdG EG
g H CG L IOLG YadG jG g Ea dd ,a efG AHh MdG

127

edG I`e

bG dG Gg EG ,e e k Gk eCG j jN IY EG Gh G jdG
,CG MQ d M CG j d dHh L EG j CG b fEG IM j
YadG jG g hLhdG eg C ke dP Z bGdG UG CG EG
ka dG j eCG Gg EG .MdG QGL IOLGh (jGdG) G jdG H
g EG ,iNCG IG FXd FGOCG fL EG jGdG jH j edG Gg C
.Jh fS dG b e b fc gh Gk MGh kT J IG
IQhdG e dG dG j hLhdG RGaEa HdG QdG cP c
dG hLhdGh LhSCG e c f Ea NEG j GPEGh jdG
hCG dG g EG.jL fEG hd FG VQCG d fG b LG C J
d j (MdG QGL jdG YhCG XG IOjdG J dG) GOGSG
.AdG id G hCG jdG IOdH Yj e gh kLQN
G iNCG jH GC Jh FSH eg RGaEH edG IdG CGJ HSCG HQCG Hh
.iNCG jT IQhO GC J Ggh ,dH

L q d SdG RG
GC J fedG a ,cdG SdG RG FdG Qhq dG kjCG fedG J
edG KCdG Gg GC jh .dG IOh e GS Y H hCG dG Ia jGH Y KCdH
dG g SCGQ Y jjdG G Jh ,fedG eGhCG e S N e
.fedG
YS HQCG EG YS I {LHRHz `dG eg RGaEH jjdG G GC J
,b J edG Gg e RJ dG dG qCG qGE ,IOdG J dG GdG N kej
Yh edG Gg e cCG q c RGaEH jjdG G GC J GS Y H dh
HcEGh cdG dG L d M b bdG qCG a qfCc WCG eR Ga
G EG {LHRH z`dG eg G edG eCG j .43eRqdG jcdG eG
128

j hQg

z Yj NBG eg RGaEH gQa e GC J dGh edG IdG gh eGhCG S fdG


RGaEG hdG Y (G OdG) G H edG Gg jh ,{LH
.edG
.q e eR H dH GC Jh ,jW GS eN fedG qdG g L J
Gg qEG ?dG hd SG bdG dG g fe J c dh
dG ` edG dG Gg EG ,dG fO dGc HLEG Y j Gj e GDdG
VQCG Y fEG OLh jGH e j Gj e ` H fEG b e S j
.BG Mh
eg RGaEG FY G CGJh ,GQhdG RL Y G EG LH eg j
b GhCG CG J edG Gg RJ dGh G IOLG jG qCch hSJ
gRJ dG hSdG eg ja ,gGG Me EG ddG Me e d M
Jh ,dG Me EG ddG Me e dG QY SSCG QhO H J jG g
G e AGLC UN eGhCG qdh ,ac G AfCG edG Gg jL
e cf Lq dG L edG Gg FXh eh .AGLCG g EG ce e Y AGOCH
:j
hj dd ,dG jd OdG qdG Y hSdG jL YJ -1
LdG L j dG g dG dGh ,ICGG L e ibCGh fe cCG Lq dG L
.G LdG G
d
q jQG jG Y KCdH edG Gg jL J f bdG -2
.G EG G dG
q q M jh ,q dH QGdGh dG GC J dHh
q TCG LdG U j Gdh ,JdG G Y kjCG edG jL qKD J -3
Y IQdG LdG L J jG g qCG EG aVEG ,ICGG U e fN
Z jL H J FXdG g L CG g q Gh ,ICGG id IOLG jdG NEG

129

OJh LdG L Ich IU c J fCH Mj kcS J jG gh ,bY


.dLdG qJ LQ qd G jG fS
j H ,a HYCG edG Gg Y FXdG dEG RYEG j h
Ga ,QN Jh e J M Y VGh dO gh H KDj dG dG
,(jcdG SdG AYCG jN) Y KDj dG dG jN NGO ddH j edG
jL ce j jdG Hh G NBG fCG e q j NGdG EG ddG Yh
ce e q j jG cG Gg qGE H IG g Y eCG j h ,da cCG
a IG jG e YG g h .{bdGz `H Yj VdG Gd NBG Fc
JhH d e eRdG dG J fCch DNA `dG hdG H J H
Y Jh ,fCG ICGGh Gk cP LdG dG g GdH IG gh ,G jL
.IG G dG
q e c
- hSdG e dCG Kda ,ge LQHh bH d Lh b dG Gg qEG
c DNA `dG jL Y IjdG jG e fCG AG d Qj bdG - fCG
dP Y eh ,d Oq G JhdG H j FV Y dGh SG eG j

j hQg

GAL d Qj KqdG Ga ,d jQG G KdG Gg KCH j dG qdG


G EG dH eCG dch ,dG JhH d G H UG DNA`dG kq e
.fCG U Y cdG U dG JdG
q
HdG e qdC j C19H28O2 hSda ,gCG jZ j fcP e qEG
Z edG Gg j jCG eh ,GQdG e OGYCH cCGh LhQdGh
e ZCGh ?DNA`dG IOLGh eG Mf e d fCG AG J CG bdG
YdG H DNA`dG IOLe hM GQe 3 H e eG d g c dP
IAGb M) DNA`dG IOLG eG IAGb qCG Y GPEG UN ?CN hOh gG
q dh ,IaG LddG bQCG GSHh GS Y AdG e e J (a
LhSG jL eq GC ,J IQH ch eG g f j BG q M EG j
g ge j e c a C19H28O2 hSdG jLh C18H24O2
dG je eh CN CG hO Gg Y Iq e gh ,ge H eG
.dG e jdG LCG NGOh
.jdGh G dEG IQdG Y kdO Md QG dG Gg jh
b e RGaEG j dG LH`dG eg IS j hSdG eg RGaEG qEG
OGR GPEa ,Oe fedG jg H IdGh qdG qGC g dGh ,edG IdG
qJ c eq dG Iq dG Y KCdH edG Gg jL J dG hSdG cJ
dHh ,jL e LH`dG RGaEG GC j hSdG cJ qb GPEGh ,LH `dG RGaEG e
.iNCG Ie hSdG RGaEG Y G
edG Iq dG H Kh ee OJ g qCG Wb IQHh J q q jh
gMCG j e Y Id fGh eH J G Jg qEG f ,Gh
dG hSdG eg c Y IdG
q q J dHh ,NBG j e Y Gk OYG
qE a NBG Hh ,edG Gg c dG hCG IOjdG Y JG QGVCG J jh
fG H J Gh edG Iq dG e c IOLe fCGh dBG g
131

f e dG G
jjdG G

feg RGaEH J jjdG G


RGaEG Y gd OdG EG

jfEG eg
f e j
e eeCG G
edG IdG

RGaEH J G
jfEG eg

e LH egh FSH eg RGaEG


eeCG edG IdG b

FG G
G EG LG
jjdG

G fEG

eg G RJ-3
hSdG

LdG id G jdG
QX e gGG Ia AHh
G T aNG

LdG Y edG AdG j J

eg RJ G
hSdG

j hQg

dG b e q U b qfCH bG EG Oj QG q dG Ggh .H a hJh


.Jh fS
fEG Y G H eq dG Iq dG b e Rj dG FSH `dG eg j c
EG ODG FSH `dG RGaEGh ,jdG NEG Y dhDG jKdG jG gh dG
RYEG h dG YhQ Y NBG kdO e bJ h dG
q QW dG fEG
.dEG

KG G OGG GP G q G
hSJ )fc gh ,ce H jfCGh jcdG G fedG R
J dG ja ,IMGh IOe e J ICGG hLhdGh LhSCG ,LdG (
.fEd N IOc hddG jL J fedG g RGaEH
jN qEG ?f G OGG GSH G jdG CG jG J c
dch ,dLdG S AG J dG OGG e J d gOGe J G
a ,hLhdGh LhSCG eg d f G Oq GG G jN J
,G jN KCH EG Lq dG EG dH KDGh FdG T NCJ CG J G IOG
.dG fEH GC jh c jh JU jh LdG J KCdG Gg jW Yh
g e jh ,G KCH ICGG KfCG Y KDGh FdG T NCJ f G IOGh
IgL Iq G cH Jh ,JU efh gQU RhHh ICGG VM SJ
q dG
OGe fE f G OGG J bdG Z jG a PEG .IOdGh d
dG Ggh .KCdG Mf e CG dG Mf e AGS bQN q dh H Y H
.IOG dH jDhQ jN f O H M OLh Y NCG e fcP
:J b
nYn n dG sGnC G o nr n pd s o n r Hn o er nC G o s n n jn s o nrpep V Qr nC G n pehn mGn n S n r n S n nNn p sdG o dG }
.12 :dG | kn rYp Am r n T u opH n Mn GnC r bn n dG sGnC hn l j n bn Am r n T u co

133

c Yh jG H JG OLh c Y ke aJ g EG
dG fedG SGH j KCdG Ggh ,iNCG N Y N KCJ
h .jG g K dG KCdG g Oq J e FXh
dG FdG dSq dG jEG H qj dG dG if S dG Gg
S CG G AZ Rh e Gk QYG G IGf EG edG j
.G IOLG JG T Y q c Y ke

G NGO Jh JG cGe
jCGQ e Gk ch ,Jd G ddG GHCG jCGQ e Gk c
,RdG TT Y QNCG If JG q a M
dY GHCG T Y gKCJ edG gG g cJ Qh
ka GKCdG g qEa bGdG h .Ie fhdEG Ge hCG
G g IOLG LddG GG g Y dG qC
fL EG ,fhdEGh JG Jee N j
g G g qCH SqdG ZCG id SGQ T Yb g dP
qCG q J dh .eG OJh JG ebE JG G e
,dG Y bJ dG JG cGeh GHCGh G L
q j qbdG Gg e hM qCG VGdG ?iJ j S GPe
e qJC G Oq G QdG
q qCG H ,G L kHGVGh Va
.TCG e H aJ Gk c c e VdG g
L N jJ100EG dH j eCG qdh
T bJ hCG jG H a JG fG a fEG
e dG e AL N dG Y G NGO JG

NGO HdG
IMGdG G

edG I`e

LdJ Ge GSH J jG JG FSh qEG .k G d fdG


a G G NGO JG FSh eq GC ,fhdG CG fc fe AGS IQe
NGO qJGh ,fEG Y kaN fGL e c GReh QdG e bGQ LQO Y
.JG Sc JhH ce GSH q j G
,UG TCG hCG fhdEG FGhq dG Y Y J JhdG g qEG
qCG cdH jG eh .LhdGh cCGh LhQdGh HdG GQq P Y qfEGh
a e %2 h dCG KdG Y Hj fEG L IOLG JhdG GfCG OY
ZCG Jh ,44dGh
dG j GRa bdG eCG ,jG FXh Y dG eCG
q

RGaEG dG JhdG

IGdG IOLG DNA `dG


G GA

j hQg

.Gh FXdG Mf e d G JhdG


JG f Hh IUG JM JG f H c M EG HJ g
FSdH KCJ d eRdG AZ Y M cGJ LJ G S a ,G NGO
cGdG g h ,d {SG FGgz H Jh G EG eOdG FdG
H Jh eOdG FdG FSq dG LJ e iNCG cGJ LJ ITe
AdG IOLG a ,G e (fedG) FdG eGhC d eG jG GbG H G JG GC j
GE GC ,gUe GE eGhC G g M Y dhD G G NGO iNGC jL GE Jh FdG eC G J G
.FdG eC G Y LfGE G JhdG U Y M dHhDNA hdG dG IOLG G
Fc dSd YH N
e Nr
FdG dSd bf FL

FdG IQTEG J dG jdG

jN e

edG I`e

GP jG JG FSh J
,fE G IM kGL Ic gGC
Y a fE G L eq GC
kGL dY IAc GP JG ILGC
Lh e QGSH ee OD Jh
.fE G

AGLCG Jh qG e dCG Fe EG 1 dW j IQcG G FGdGh ,qJG


G e dCjh ,{Rjc jGRhjKz Yj G q FGdG Ggh ,ac G AdG
FGdG c M EG j eRdG AdG QN Gk M j dG AGh ,jdGh Gh
dSG NG EG Gk OSG G J eh ,FdG dG S eG

138

j hQg

gKCJ N kW G AZ IOLG G J dc ,G GOOd


.G fedH
LJ dh ,jG G H J jG bH e eOdG feda
CG edG qC Gk L q e dG gh ,e eg e AJ a IMGh e
fGh DhH j c b jG Gh eda ,iNCG N CN qj
MGh e Ljh ,Gh dH AMCG Y c fj AdG L jc
edG j J CG J a IMGh q N qGC CG ,dG dG a j
.iNCG jG hO
SCG bQN jh dH GC j UN
q f g G EG edG j Mh
OLG DNA`dG EG fedG dSdG jEG e j dG g qdG Ggh ,Y
g d :edG IG e ke OQf fYO IG G Gg Sh .G
Gg jh ,hCq G EG fhdEG dSQ SQEG jh ffG Y NBH Je SM
dG gh ,QdG Y d ke fhdEG HdG EG dSdG g SQEH SG
jG G H
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cGJ Y G AdG j
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jGH dG edG JM Na ffEG eCG ,kjCG RG EG NOh fdG
.q Y kjZ G Gg j dG Gd ,q d qdG q dG
LQd jd V H U kHSM g CG Oe dG
q Gk N kej fCGb dh
HUC iNCG SG ILCG e HGJh JG dM qfGC h ,IOG dH jDhQ
JM ka OLe G Gg qCG qGE ,RG Gd gG dG
q e dG h gq dG
. c dTG EG aVEG IOG dH ij h dG e ie YCG Yh
gG YdG H G IGf EG G J dG FdG dSq dG f qEG
U e Ie cCG j dSdG g f Gk L ee LdJ GSG EG aVEG

ffEG

H a J IjY ILGC V SM RL GE ffE G Y e ee SQGE d


L Ljh ,QdG Y H J fhdGE HW GE f eG g E a T
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140

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IdG gOGYCH dDJ dG gh EG d e J G qC dG e SM RL


SGH dG dGh dG Gg IAGb f SGH dG dG Y hG L
ch ,M N jJ 100 Y j fEG Lh ,dSh H J f
.bQN Ie eCG Gg CG Th ,Jd Gk L G dG Gg Y e IMGh
fh ,G AGLCG NGO FSq dG f H J dG jdG Y ke qf Sh
.G e FH 1 e UCG j M J dG dEG IG QGSCG Y dc

G NGO JG OH UG edG j dG jq dG
AZ IOLG H dG g G EG j eY edG j e hCG qEG
J gQhH Gh ,dEG j dG FdG dSdG H j H bdG Yh ,G
G L jh ,{jdGz`H YGh G NGO HdG FL EG dSdG g H
H S M GPEGh ,JGfh G AZ H OLG G j dG eRHdG e Gk AL
Y EG G fG jh Fc YJ j Gh edG
f {IdGz `H J dG gh ,djP TCG J EG aVEG IOe fc CG H
G IOLG fCG b f j jdG T dG Gg qC Sa EG
.jdG EG SdG e c aVEH
dP Y eh ,JhdGh jG e jdG b e YO j dG g qEG
cCG q dG a ,G jL Gk QNG YJ dGh GTP `H IG jG
.SG dG Nq dH GC j Ye eY IN d Ljh ,IG g
NGO eGh FSq dG f Y kq e Gk AL J IQcG dG g qEG
Jhd AH IQTEG fc e J gCGh ,fCG SGH J q H ch ,q G
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141

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HdG GC j CG b
eg
jG G
G AZn
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G NGO bf jL

fd da Z JG Sh

j dG G GE edG j eY
j e jh ,G AdG IOLG H
Gh ,H dG dG G GE Fc eGC e
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H j b GE G J eC G OHh
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G IOLG fC G J GC H fdG
H J GG gh ,a SdG SH
G JG FSh hGC jG cGq d

SH JG Sh

jHdG dG f

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bf jL

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Sar

fd dG JG Sh

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G JG f CGj Ggh jG G hCG Rjc jGRhjK `dG e kJG jh
.dG
VG M k G Jd QG dG Gg c
b e ahe dG ahG dG UdG J h ,jdG
hO ed G JG Gg j c :Cj FS
q Qh
eh eG f Y UM CN Ca ?CN CG JQG
N hM dHh WN JhH J EG ODj kW c
UN FSh OLh INCG HCG KCGh .d dG dG
e MGh f iS d QcG SH2 `dG h ,jG H Jd
.G JG FSd G GfCG e
G ALQCG Ie q G JG FSh qEG
G AGLCG H eG Od T H a Jh ,ac
FdG G IQdG ij
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143

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cGdG g J h ,FXd FGOGC bQG jG cGdG jW Y IGdGh G AdG H JG q


.G JdH JG AGLGE e dG FdG H Y

fCG J jG G g SGHh ,G JdG SGH JG ebEG e


.hCH khCG Y H UG eG J CG G IGf edG

JG FSh
MdGh AdG L b jG G g AH Y jLCG dG HCG Ff qEG
FG P JhH e dCJ FSdG g e jN JG Sh q a ,gq dG e GaCG Za
dG Gdh ,jNCG JT TCG Y G OHCG KK T e dh ,eCG M
N f j eG qCG CG ,a JGf e J CG JhdG g J GdH
d Gb a ,G YGPEG dG e d bdG JhdGh eG f M G
.G YGPEG GOOdG J

144

AdG IOLG QhG Gb g gf


e JhH e GdG dC Jh ,G
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J G NGO JG Gb e dC J
c GE Lj Ggh .H e kH ee
g ch OHC G KK JhdG jL
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q Ea (MODUL) {JG FShz d eGSG gh e f cf CG jh


jG g Y dG eSG dh ,kec JhdG g M J dG g
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GP j G g EG SdG aVEa .AdG j dG g SdG Ggh
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.MGh L b fCc SH2 `dGh G
G Jd jG qG g EG q FLdh J CG dG HCG YSG
Gg eh ,Ie e jG cGdG g J j dG hdEG G GSH
je Jd jG cGdG g e dG qCG hcDj AdG Gj e SG q dG
H a J fCG jG G g dGh .45dG dG EG LG e
C N hCG N hCG b CGh ,GHCG j H eG Od jW S d
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146

j hQg

.SG SH2 JG Sh e

.SG a SH3 JG Sh e

gh ,e cGJ Y ebCG H j G JG f EG
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S Y OLe GdH O AL EG dSdG g jEH J cGdG g qCG CG
GHh ,V YS U e Ae j eG e Fg c H e DNA`dG
e AGOCG FGdG JEG Gg qCG
q T h .CN q GC hO JhdG AH G J dG
.H RYEG g G e H 1 dW j dGh JhdG jL b
J dG dGh q dG ie d Vj FXd e AGOCG e HS cP e qEG
eh ,G FS IQe G JhdGh GdH e J dGh G G H
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G NGO qJG Y IqdG dGB


gj dG G fh W M c dEG LG jG Y fedG KDJ
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e q a ,dG dG f j GdG ZcdGh dfCG feg dP Y
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147

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JhdG Y QdG Y IQdG jG dh


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JG N hM j Ga Me jq CG CN
G qGC H .kGcDe dG j dHh G
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fCG j cGdG d q G bG Ggh .CN
CG jG G J c :IjY DhJ
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j G JG FSh AGOCG JEG CG FdG g e q jh ,HdG
q SCG Y
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JhdG Gg qGE .d G dG Y d GdH JhdG GH GC f fYO dG
dG a dG AG eq GC ,jG G jH j dG g hCq G :jCGL e dCj
.IdG Rc j dGh IG YdG U SH2 `dG j
J jWh gOYh eCG VG H OJ d eOdG IdG gh
f b e EG Yf Y dG j IdG gh ,jG G jH J dGh
f dEH j NBG AGh ,jdH dG j dGh SH2 `dG e a MGh
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,O UN f EG Gk OSG H FGY IQH dG g qEG
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j ,fEG j L hM Y j e e NCd jG H dG SdG

148

j hQg

AG dG G IOLG H f dEH PDGF Gh bdG JhdG


H d NGdG AG GC j dG Gg fh ,G e edG AYd
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.G JhdG H Fc HGQ JhdG Gg QYG h
dGh {SOSz Gh bdG JhdG gH Grb2 j IG g Hh
GC j M IN h ,(G FS) G eRHS OLG gh SG fCG j
H edG JQG f KOG dG
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H FdG eC d bdG edG j


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GDP
a Z {SGQz fCG

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dG zSGQz fCG

Sos SH3
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AG GC j edG JG j GC H
G JhdG H d NGdG
H JG Y hD G Grb2
j IG g H ,G JhdG
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NGB JhH H j SOS JhH GE


jf Ggh ,d Ras {SGQz Yj
GE FdG eC G jGE j GG g
Y OLGh dG eC G jGE G G
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GG e S Ggh ,{RASz Yj NBG JhH hCG H {SOSz


GC J dG g AV Yh ,G IGf IOLG G EG G jEH J
.H a G dG d fH G e bGdG jG
N CG UE NJ e dBG dgz :j KHCG Ff Y AdG jh
dG f g dGh G dBG g EG .46 zG NGO G JG j
J Ggh ,jG BG J e ie YCG J ah J d QG
jG G bHh JhdGh FdG UdGh Gh fedG
.VQCG Lh Y fEG Lh e ec fGh JG
h ,aOG eY EG QGh G dG Gg OLh Lf CG G eh
h hdG IOe Y cT Y f e kbQh Gk QJ cCG G dG Gg QYG
cdG fEG cGe H JG e keJ cCG g G JGh ,ac dG AfCG
dG Gh jG L qCG qcf CG jh .hdG H Mh jdG cGeh
Y ka J e J h bY Z OGe g G JG T V ee ODJ
dGh jG H jdG SdG Gg qGE .IOG dH iJ j jL fc
ceh dG AfGC IOe ha d Y cd JG RL e IAc cGC G NGO JG RL j
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dG Gg C fCG dG qGE T Hh ,f AJ e C j CG Y f Y j
.dG Q dG g a qGh

jG NGO QNC d bdG cGd UG GfC G


EG ?z jG fY Gk QJ cCG JG Sh g e z :JBG GDdG Lh d
:NBG GDH HLEG g Y qdh ,ffEG g :cCG GG d af e c
jCGL HCG H Gq L Iah ee jEG feEG g :GGh ?GdH ffEG GP
IQK KMCG ffEG CG SdG H jh ,kGL Ib eR Ia cdG Gg Y
LdJ CG qT .gG dG Gg e IH j NBG dGh ,JG Y
eG fQb d dh ,jdG jQJ dG GREG gCG e gQYG ffEG

e ffE G LdJ J
jQJ GRE G YGC
J dh ,jdG
YS e fQH VGe
jG H e JG
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151

edG I`e

hCG dBV ie cQOC YSh q c jG H dG eG e ffEG Y dG


YG Ja ,dG IOLG J hCG G dG jG kUNh fdH fQe
.fEG Y J dG JG FSh SCG e GJ
J jG g Y Y J YG Jg jN IOLG fCG EG
NBG KCGh ,HG G REG jQGSG e eG f JEGh FdG YdG
dg CG dG jG H JG H j dG jdG Y jLCG dG dG HCG
.47 jJ JG VGe e Fg Gk OGYCG JhdG H
e bdG JhdG Y CG SdG H J JhdG g qEG
dG Gg EG LJ q dG jG H JG FdG YdGh ,QGSHh H
JhdG jh jG H e JG e QGhOCG d UN JhH LJh ,QG
Y dhDG dG jG Gk c Gk QhO JhdG Gd CG AdG jh ,gMCG PSD-95
.dG
hCG ,PDZYJ e KK ,IjY JG VGe Y PSD-95 `dG jL
Ad fjG IdG dH j dGh ,jG d eRHdG AH j

JhH FdG g eh ,dJG jW Y kGOYG IjY Ff d C H PSD - 95 G JhdG j


MC H G cda dG GE IQdG eGC ,PDZ`dG cJ IQdG ijh ,PDZ j
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152

j hQg

,H dGh G eRHS SG JhdG Y dG e ddGh ,G


dJG S Y jh Y j PSD-95 Jhd UG dG qEG NBG Hh
.G NGO IjY
EGh dG jG Y j Jd bQG fCG g e OLh EG
Gg d jDhQ CH cdG jh .kjCG dG jN d T OLh Me
dh ,jG Jd QG dG Gg OLH qGE J gjEG JAGbh dG
M H U e CGb e J
q S qfGC CG Y dM UC dG Gg
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AV
ddG

ddG Ib

dG JhH

IdG d G fCG
PDZ fCG Me
cG

IdG d G fCG
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FdG GQTEG f Xh ODJ dG iNCG JhdG

h ,IOe H Y e ce Y d IG g C H cdG HHP Y jLGC dG HC G KGC


G cGdG g MC ke kT YC G GE IQdGh ,Jd e cGJ Y dG g
. q GE IG Y e dG GQTE G f Y hD G INAD`dG JhdG gh

153

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G Jd jdG dG Gg OLh (qaShQO) cdG HHP Y Y HCG


ODj {INADz Yj UN JhH OLh EG dG Gg jh ,Id cG dG
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q q dG dGh ,G EG dG e eG jEG e
c :HdG
q dG FG Y WG CG H ddG SCG fCG Cd J fh
CG JhdG YSG c ?H UG fCG J CG JhdG g YSG
100 GM gOY j dGh ac jG e d Jd dG dG g J

cGdG J H a eeh cGe b T Y ghD fGE dG dG AdG J


.jG IOLG G

154

j hQg

FdG UdG YSG c fCG Cf CG j dch ?N jJ


?...bQG ddG dG g J CG JhdG
FSh Y e bCG gQYG AfEG b GJ e dGh dG AdG EG
T Y J gh jdG jQJ Sg REG cCG gQYG c ,jG JG
EG X b G g CH fZ hCG f Yf CG h ,H a H cGe b
gKCH hCG dG bdGh YGdG M hCG jdG hCG jG hCG GQq dG KCH OLdG
dG HdGh IG cGJ f OLdG EG X b G g CG Gh .kL
hG Gg RE bO HM AGLEG Y ka dG GH e AdG
G Jd bQG fCG g U dG e ,dG g Y eCG GOeh,dG
?H j dG dG JL dGh
Yh dG IQH N Fc g Y G fCGh bdG JhdG EG
:J j ,d q M jdG SGdG
n go hn Am r n T s co n nNn hn l n Mp n U o nd r onJ r ndhn l ndhn o nd o ojn sfGnC p V Qr nC rdGh pGn s dG o jpHn }
,101:fCG |l p Yn Am r n T u opH
mn n S
n rdGnC o Qo G n r pe n cn m r jn pr ndGE p
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dG fOh G G NGO JG
dG acG e dG G JG T
GN dG Nh jdG dG e INCG GdG
UGh aG QeCG e jdG Y dG IQL
dP Y eh.eLCG YCG JG jH
Z G JG dG acG
f e dG Hf FGL fc VG GdG

155

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edG I`e

fL EG G JG W HCG q S
Gg Y K dG fCGh ,iNCG HCG
n nNn sdG n go h}:J
b
n
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nr njhn unrdHp VQnC rdGh pG s dG
Hc jQJ) 2001 S jG dG dEG UJ
o ndhn t n rdG o odr bn o on an co o o j
dG Y dG G aG g e ?(QdG
q g
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opd Yn Qp t dG on roj nr nj oro rdG
e Ic LQO Y SCG g Y HLEG qEG
o p n rdG n go hn pIOn s dGh pr nrdG
IQH dh Yf HLCG g qGE h ,gCG
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n rdG
WG N e jG d gG q dG FL
dGR eh ,H a hCG a JG f Y
GdG e jdG Ie G Gg HCG
SDe qJ bh .GQhdG e IjY GQe EG fGe J SDe b eh
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dG Hf IFL Y FG c jdCG OG Gg SCGjhAFCS SDe QNH hCG
Gg jh .G JG KHCG f IFG g Y RM bh 1994 S
.AdG e G fL EG eL hY OG
aH dG b dG
q EG q G MG GPEGz :OdG
q Gg jdCG QahdG jh
c H dG b q dG e jG EG dG LMG GPEGh ,dH dH e gh
J G jG H e RJ FdG FSq dG e G gh .e cCG
dG e HGd q e VJ G jGh .NBG dG KDJ c eLCG
... J ch J j a ,eRdG dG dG FSdG d FdG
.48 fCG g H dGh JG ebEG c g idG Gh
c kqe G Gg HCG AGLEH AFCS QcG OG b GdH dG
q Gd
kae aSG KHCG AGLEG OqJG CGS :( dDG J Sb H e) j

156

j hQg

z NBGh {jSje jOQcz hCG jG e Yf JQq dH OGh) ,JQb


.({ B-jSd
IY Y IQY dG Ggh ,JQdG JH UG MGdG
q CH dG EG f h
bdG jGh jG G dH OG) e jdG q G SdGh
q ,fGe
SG Y jh ,(deh G JG T W Y G T IQdGh
eh) ,kGL af dG AG e GC f CG j hCG Ich ,c Yh khCG MdG
ShJz G e) . dgEG Y fL EG (G IG JhdG dP Y
MQ AGLEG jW Y IQdG jN e Of CG Y K eh ,({jcSzh {Sjc
{hRjSz UG UGGh dG GSH MdG e jdG VGQCd aSG
JG T V kjM Gh IOG dG JG OG) ,QfCGh dG JGh
dG YCG f dG g e kbfG ke acG SSh (G
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jGH Gj e dG qCG gOe QdG EG dSQ VG IdG qCG Gh
e dG eOdG GdG
q dEG Sh ,jG FdG Y S jW
.qbG dG acG
hO dG dG jdG SCG j AdG e dG g NBG fL eh
Hf IFL Y FG HH fc QahdG g ADg MCGh ,edG RedG GSG
QahdG Gg b bh .G JG M IgdG KHCG H 1999 d dG
:j e e U Ad AKCG ec Ve
hCG fjdH gOY Qj dG jG ddG e jdG G IGf z
.50{kGL jW jW eeCG GRe....Y kT q e MCG Qj h iNCG a ebQCG
kUNh dG acd Gk Y S jdGh OG dG CG d q jh
d IcJh AdG SdG EG HN j Y c ch .G JG fCG jN
.VQCGh GdG
q Wa dG g AT c dN CH

157

e YEGh IG J FNh S q G q G
f V FXh SQ GYh L Y FGMG Mf
G Ge M UdG e jdh) AGLCG Seh ee
QGO e QOU j hQg dD {G Iez c LGe Lj
dG dGz
q ch 2000 S R fSG d GQhh
fc fSG ,d GQhh QGO e QOU f dDd {qd
IQGOEG jdG dhQc e AY q jh .(2001 dG
.51Qjf e Ic j dGh IQGOEH G dGh
LGJ G dG OY J dG Jhd J Yh
B JhH jL Qe e cCG Y N qa ,WS FM
ke f fYO bdG Gg eV ie QJ LCG eh ,52GfCG
SCG c dGh IMGh N JhdG q H b d :JBG G
Gk Qfh kd fdG MGh JhH YHh QG Y gOY Hj dG
M M d eCG ,q dG f c S 32 EG M f CG hO
kac c dG c dP HT eh dGh cCG FdG bdG
Gg Y f GQe 7 GM dg c d GPeh ,dcE
,N jJ 100 Y L j e MGh ch cdG
,cdG Gg Y IOLG JhdG OY J FY
Ie GC J a ,e O dM JhdG g c EG aVEH
G OGG EG J c e fG eCG VG EG T c
Mh iNCG Ie e JhdG H J dGh G NGO
L j dG G GG Sh ,53 LG
cJz H YJ IMGh jL eCG VG
NH FXh AGOCG JhdG g Jh ,{JhdG
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jdG f
G NGO

cG Gm
G eRHhfG dG
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jQfcjG
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jdG
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IGdG

G AH
NBG dGh IG fedG EG j NBG Hh kjJ jG FdG YdG
J Y a G j IYG EG aVEG ,fCG EG cCG jEG j
cJ Y UdG e jdh).iNCG JM FXhh q dG dG
q J e H
GQhh QGO e QOU j hQg dDd {JhdG Iez c LGe Lj JhdG
.( 2001 S f ,fSG d
G NGO UG G cG g d kb gY bdG j dG dCGh
JhdG g qC G kjM cG JhdG H bGd G OdG f
UN R EG f j Hh ,a S dG bGG EG J CG j kjM YG
QN LGNEG j LQG Sd kjM Ye JhH gh .dEG LG M
J QG e eOb iNCG JhH g AKCG J h ,G AdG b e G
cM e Gk AL d kjCG h AdG ISh HbQ V G EG Nq d jW
.jG JhdG
Mq dG Mh ,IOG dH iJ dG G NGO V cM g RjEHh
160

Gg e ZdG Yh , keMORGh kWf cGC fC H QGdG S q dG cM Y G NGO cG J


.QN f ah q e G qC ,YGC IQdG ggf dc Va jGC Ige a MdG

NGO dG cG eCG G kV j IhQdG YS e j QhG


eh ,G e FG 1`H M Qj N NGO J cG g c Y ka ,G
hM c IGh ,G e H 1 e UCG M j dG JhdG b
J dG G GQe b eh IOG dH ij M V cG g e
heh O JhH c GSG beh ,QU f ah e Xh AGOC kTh k
ehRjdG YJ dG fG kYe CG G QN e eOb JhdG Gg c AGS
JhdG Y j {jQfcjGz dEG dG Jhda ,G eRHS IOLG
dG CG dG S Ic j eCG Gg T dh .GdG bH dEG dG
.G OGG e JLMG NGh G LfEG eG J eH
dGh Qd HdG Z H j dG G qM G JhH Qe cM qEG
JhdG g J jCG e :IjY DhJ a j G e FH Gk MGh j
ceCG Y JhdG g J hCG J c ?a J dG bGG kjN YG
JhdG g J c ?dEG LG jG Mh LfEG cGe e Gk QYG IOG
c ?G fe H j dGh gdG cdG P G AZ N e QhG

161

bdG (Vesicle) jG
hRjdGm
hRhdGl
kjM e JhH m

dG
eRHhfCG

hN
e JhdG
Y G
G

JhdG jJm
I RG jG
G AZ

AL GE M q j hGC G QN GE UN bGf SGH f j ka ,NGO dG j G JhdG fGE H


g ch ,dGE LG q M G c L NGO fN q j hGC ,a dSG OGG G e e
.G NGO Jhd FGO cM OLh GE OD J GG

?CN CG JQG hO Gh G cG g e J
kffEG g CG Vf fYO IG g dh G NGO bG f CG hdh
d CH jG OdG Gg Yfh ,dG jM JhH OLh e kH kjM Gk Ode
gh FGM bH Abh dGh cCG OjEG c j kjjh kjT Y dO
jG Y j IOdG jM dG qGC Th ,f Qe GP je j
OjEG Y j dH Ic HU bS dh ka kT dM e Sh
Gd jG g AdG Y dd ,dH G jG g HG ceCG
e fEGh ,Y j dG G dH j dHh YQGTh bRCG j c jW
dG Y JhdG eq GC ,jW Ih Gk c jh Gk L C j CG Y HdG G dG
.q eCG dH Ggh G NGO OLh e e e g e c j a
ec aOJ dG fGG c IR aGgCG f JhdG c dGh
q
QGSCG c EG G Y HCG UJ bh ,d QGh gG dG
.HdG DhdG Y HLEGh G
162

j hQg

?G NGO JhdG cM J j c
jdG edG f CG j qc
aGgCG EG FSdG jE j
dG Ggh ,AHEG hCG CN hCG S hO
YdG d ke Gk eCG j
CG YEd Gh ,jjdG eG
j e OLh KCG dG HCG
hGh ,54G jdG edG f
OG SGH cJ j JhdG CG
T V eCG VG e G GdG j GC j dGE SG GdG GE e dSQ J d
jL kUN Gk AL CGh ,e Sg a G eC G dc ,dG Y He jdG edGh
J dG jjdG RedG e S kjM j JhH
KK EG IY H e jdHh JhdG
.IO qG Lh GE d d dc
e J dG dG keCG kM
kebQCG j FSd jdG edGh .JhdG dH UG jdG eq dG j jG
,eCG VG e OY Y IQY jdG ea JhdG eCG dG Y J e
j SSCG Gg Yh ,NGO hCG JhdG jL W j jdG edG Ggh
g CG EGh c H UG eG j CG G kjM cG JhdG
.kGL Qe SGH G NGO JhdG S ke dh ,j
RGaEG
c L
G AZ

eRHhfCG dG
(bdG H ) jQfcG
GQTEG S
JhdGi

GQTEG S
JhdG
IGdG
GQTEG S

fG ISdG

SGH JhdG LJ c Vj dG q dG Gg
G H kjGC dG Vjh ,jdG edG f
(a JdG jG Lj L SHhQdG ) ,G
kq UN kjq jH kGeQ j AL Y kjM e JhH c j
VG e S e AG Gg qdC jh ,JhdG dH
JhdG jL W MGC Y dG g J eYh .eC G
.G ag GE ZH Jhd kFS Jh

163

edG I`e

dG EG dfG c ke ,kjM e JhH cM f eY


L b e JhdG jdG eq dG Y q dG j :JBG QeCG if eRHhfCG
OjEG JhdG Yj dG ddG S J L gh SRP YJ jL
Y dH G Gg jh ,eRHhfCG dG Gb V d SG NG
EG j CG Hh ,dG LdG EG jG Gk Qq L H jh JhdG jdG edG
IdG Y UN e Y GdHh eRHhfG dG AZ Y SG G
J a jh G g qJ FYh ,G H j Jhd FGh UG
GG g c ,IdG Y SRP L j f dG h ,IdG
a Gk J cG e JhdG LGJ gh .YEGh gd e e eR J
,KK OHH g G , jeh je eG VMG SS e J Jhda e g
,G NGO Jhd FGdG cG g IQq dG q h ,FGO cM dM fH q G G JhdG R
.jG Jhd ddG e J dG SRP`dG jL gJ IQdG e idG dG jhGqdG h
AdG JhH
Re JhHk

G AZ

U c hG jM

bdG RNA

hRjdG

OLh e j dG JhdG
Ga

c L

H IQTEG

H IQTEG
Ga
jQfcG
(bdG dJ ce)

JhH
SRP
eRHhfCG dG

hdG AdG
hRjdG e
JhH
SRPe

H IQTEG
Ga

j hQg

jG g Y j dG g h
dG AZ N e QhG
IOLG IdG b qCG kY ,eRHhfCG
j AdG Gg Y
10-9 * 2 hCG ( 0^000000002)
gh NBG kbQN kJ Of gh .e
,JhdG fEG AKCG G g M
j JhdG fEG Y hDG hRjda
Ggh ,e S T Y LfEH
UG Jb e JhdG Qhe EG ODj
kM j fE G ej dG jdG eqdG f qGE
EG IdG J JhdG Qhe Hh ,dH
.kjM G JhdG LJ d kde
Y j FYh ,NBG Qhe hM M
Y Hh ,e fCG IY JhdG jL e a jh JhdG jdG edG
AfG J dG g .OHCG KdG T NCd JhdG jL J ITe fG
.ag EG dUh H dSdG Y hG dG gCG
jdG edG Y fCG J c J dG g iNCG IGh
g eb GPEGh ,JhdG e j dG eCG VG BG hCG e e JhdG
Gk KCJh kT JhdG J j Ga iNCG eCG VM H (kVa ) fCG
dh ,dG g GN e IN c Yh e S d j Ggh .gCGh
,GMCG e M CH jG g EG Oj dG hCG G dG Gg CG G
h JhdG edGh SRP L CG CG ,je UJ H J dG CG bGdGh
jLh G GY AZh fCGh JhdG Qhe Ibh Gh hRjdG
JH Y hODj jG ADg L ,G jG dG gQhO ODJ iNCG
,G dEG Ie Md QYG Jhd jdG edGh ,bQN fGh
JhdG jdG edG eq CG kjJ S 40 e jdG edG f j fEGh

165

edG I`e

.dG e jdG LCG Gg ej Mh dG Y OBG N e a


,dG Rg OQGg e gh G JG M KHCH Qe Y e Lj
UG RedG f Y dG CG MdG Gg cCG bh ,{HL .h.z G Gg SCGjh
edG CG Y cCG bh ,LddG acG YCG e j G NGO JH
OJh JG Y J M jL Rc Y ODj G JhdG jdG
.55 G f Y eG
,edG JM efh Gf e kdZh ,kjZ j Gg YdG dG fh
,dG Gd G dG dG Gg Y ke CG QdG jY Jh
Gk L ke dG SCG Gg jh .G hO hCG LdG N jDhQ h
H ZGa OLh e UGe jRGe jOY kWN jh ,ZCG hCG YdG d
f f jh .VdG Gd e UN RH qdJ dG QdG q TCG SGH JAGb J
dG G dP J e ee EG dj dG SG RL EG jQdG IAGdG
.dS H

d UN eQ OLh AdG j
.jL RL H JhH

166

j hQg

T .edG IG Y d Sh YdG dG QYG jdG h


AGLEG H Uh QW BGh dSCG SGh jQdG SGh dG N CG
H a eH J dG Jfeh dG f jh ,UN
q eGH Vhh J
G g j CG Gh bdG Ad h ,Sg YCGh HC W f
AGQh aOG CG Yj jdG dhCG be Ea dc eCG GOeh ,Y Yj hCG
.G Y dGh G De Y be g G qJd edG dG OLh
o go r GnC Am r n T p r nZ r pe G o p No r GnC }:dJ b ,j c adG dhCG M BGdG d Qjh
q
e dSG QJ jBG IOQGdG QfEG SG Uh ,35 :QdG |n o pd n rdG
JhH fEG M H a G JhH Qe fEG d g dSGh ,Yj
JEG Gg H JhdG g Y g dSG cCGh ,aOe QG Gg e MGh
.dG d AT e fEG IM QGSG Y ad H a edGh fGh
dG eCH AT ch ,jGh JhdGh jGh GQqdG CG T
qdG Y Lh Gg H ,ah JQbh H fEG e q eh jdG dG
.AdGh dGh G e j Y AdGh Mh dG e

G NGO AqdOC G SRP: L


V eR Ua jdh hCG Iq d QhJh d LCG H f QJ
q
fhj ddG Ggh ,MS Te hCG dO PqJG e H dG g ,IQjd
.dG
q dG EG OTQEG EG aVEG dG dP gCG e gdG
NCJ d dG jG JhdG qdj AqOCG Y G Ea dch
dCJ G gh SRP. L g AqdOCG ADgh ,G NGO dG de
dG e q M EG J G g h ,RNA hdG G e e JhH e
.e 10-9 *24 hCG (0^000000024) dW jh ddG d eG
G eh L e JhdG e gj CG j fH SRP`dG L R
h ,iNCG L e eRHhfCG d UG AdG SG IdG a Y IOLG

167

edG I`e

H f QJ
qdG Jh gGC d
,Ib Ia N FLQGC
kM dG g a
jh , qMS Te GE
TG S SRP`dG
LJ f MdG
Lh f JhdG
.G

Gg M AdG j h ,G Gd G AdGh cdG EG BG q M EG UdG j


SRP`dG jL e Gk AL qj dGh RNA hdG d G G j dG
Ib Y IOLG Gh SRP`dG H bdG OHCH eJ ae ae Y EG aVEG
.56eRHhfCG dG
QG fOhO.CG.L QahdG j JhdGh SRP`dG H bdH j ah
h ,57jd Ie bdG g CH jG iG Y jG AdH UG KHCH
qC OhG HCG EG gd Gk e G Gg j kah .58 jL aGJ fCH dG
JM Xh AGOC Gq jh H e eAj c b JhdGh RNA hdG G
d jGh GQdG H OLG gdG gQhO d aOG CH dGh ,IO
OdG gdH OLdG EG X b G JdG CH AYOG j bQG OGG g e
,2000 Y qGE JhdG Gd QdG cqdG EG UdG j h ,jGh GQdG H
Y J dG IQb Y M dO gh ,QN jJ hP JhdG qCG j cdG Ggh
.G

168

j hQg

dG hG GhGdG j (SdG IQdG) kjM G Jhd kdO j dG SRP`dG T qGE


.ddG d SdG

G IGf UGGh q dG
Y fC KGQdG JU d f J fCG G IGf Y kY hG
GOSG G NGO jG ddG ZCG AGLEG jh DNA `dG hdG G jL
IGf H FGO JhH JG g dG Gdh ,DNA `dG jL Y IOLG eG EG
Y j JhdG qJG Ggh .eRHdG GY iMCH hCG eRHdGh G
.ac G LMG d Sh b
AdG Gg Y LJh ,dG q FK AZ d CH G j bH Y IGdG J
IG cGdH gYf dEGh IGdG e LhNh JhdG NO qJ Ie cGJ
cGdG g OLh dh .kGL Ie cGJ fC GdH J Y kVY
CG hO hdG AdG R CG RNA)h (DNA jhdG Vc bY jL YSG
Y H J IG cGdG g J eYh .QV CH jG g J
dG iNCG Gh .59 ,IMGdG fdG hd iNCG Y Yh NO
EG dNO AKCG JhdG H j dO OLh g dG HCG jW Y dEG UdG
ajQdG e IjY GfCG gh ,{ajQcz Yj ddG Ggh ,e LhNh IGdG

169

edG I`e

dd IGdG f c AKCG JhdG IY J


fCGh JhH g ajQdG fL EGh .dEG
e fG Y ke Gk QhO J GfCG hdG AdG cJ
.dEGh IGdG
IGdG AZ
JG OLG HdGhCG dG Gg qEG
eCG AGOCc e T b (G IGf EG f) hdG
hdG AdG IG HG
fc QahdG gh gMCG jh ,QdG jf IYO
e q dG AG ijh ,d
dN e dG HG IQdG
fG Y J dG IG cGdG qGE z:HH
`dG h DNA `dG M jL
.60{H ge EG UdG j JGh
.RNA.
ddG XH j dG ajQdG ke hdh
QG AGOCG Ggh ,G eeh FXh Y dG G BG c a ,Jhd
c QYG H fNCG dh ,G Y hM Y kWb kdO j MGh
EG dP fOd ddG XH J dGh G IOLG IG cGdGh G
.jdG dG dG gh AT c Y QOb MGh dN OLh gh Z IMGh f

QGSC G aGh bQG fC G


e Y AGVCG J dG IgdG FdG EG j H kej UJ dG HCG qEG
Gg e OLH J dG AGQBG fc Ib eR Ia bh .G jdG d
HCG KCGh ,IOG LCG fEG j SdG Hh ,YG RL jdG dG
G fCG EG e GQhdG RL e J CG J jedG jG CG jCG
.UG jG e Y hCG jL SGH
G fCG OLhh c Y af dG eG c M eeCG Jh
HH fc QahdG f bh .H j g h fQH jd V G
g f bh ,{G jdG eq dG fz ac 1999 d dG Hf IFL

170

j hQg

IFG g fc H dG J Md IFG
dG Gg e LCGh ,AY IY H IOdG RJ
BG fz :gdG j e a AL U Ad
M c f CG dN e f Me
bCG Y hCG G NGO JhdG cM
a b fCG j Gg dh ,d edG G
KHCG BG f dP Y eh ,UJ
H jGh G cM M c Y
Y jH dReh ,eRHdGh G AdG
J ch cG g UJh e
.61{Yh
hCG AdG EG gP jCa ,kGL VGh eCGh
AL YG EG hCG HdG hCG QdG YCG
HH fc
dEG IQdG EG j AT a eLCG
ie Y dG acGh ,gedG
dG Sd Yh Y IYO J dh IHdG eRCG dG Y aN fc G
kSEG j G Y jL Y cG hCG QJ ch ,dG e q dG EG
bdG h ,ja Wg hCG S fc hJ dG Qq dG jf IYO QaCG j
fc Mh {a cz ATCd dEG eCG M d QG dG j f
q Y dG Y Y dG EG acG g fOJh .Yh Mh aJ
:J b ,Y AdGh H dG EG aJ h q Lh
u co
o onen ppn pH p sdG n o an @ oon an co o nd o o jn rGnC kn r n T On G Qn GnC GPn GE p o o er GnC `s fGE p }
.83-82:j | o L
n r oJ p ndGE p hn Am nT

171

AKCG Qe LR Y aG LQCH SO fCG Vd


QG Ld eb VJ H edG Qda ,G d
q
qGE ,fdG dCG e AL EG j Gk L V CH cQT Hh
IdG g Nh ,kGHCG H f CG h jd Ib IdG g
JG Gg EG ,G EG dG e Y IQTEG SQEG j IdG
Y j hCG dG jG SGH j ZGh jdG
.{hQfz `d `H n LdH
,e jZ d e AT c CG fLd dM a ff d
bCGh IOLG QGQRCGh SdG G e G Jda
dG jG dc ,kJU SdG H JG LM J fhdEG
g f eY g OGh) ,LCG e N dG jdG J
S jG g J Yh ,(QG hdEG G jG
e IRQdG dG SGh QGh G L e J qfCG
.iNCG jG c GdG Y FGMG EG aVEG G L
G cG Gg CG EG ,kGL Qe c dG G J h
(0^004) H dW hGjh Gk L U fC G Y d e
N dG SG dch QG jh ,e (0^10) h
.jG bH e JG
dG G VY Se qGE :H HM Y dh
(10-3 *1) hCG (e1000 e1 = hG) kfhe j
ddG G dG jG Uj CG AG OGQCG dh ,e
dH ij e e N d Y N Qe100
,ec1000 j dW dh ,he10 VY C IOG
IG eeCG QJ gh GZc 1400 j fEG q eh
IOLG IGh jdG dG H gOHCG H dEG

UGGh q dG
q dG q G

edG I`e

.fEG e
CG LQO EG Gk L IU dG a ,HG FdG g Y kb bdh
Gdh .62 G g H J dG dG C OJ SG G GP e N N
,ITe Z IQH Y G b dG jG Y dG eG q L Ea dG
q
dG jG Qh Y HCG AGLEG jW Y Y G dG ea
.dG dH dG Q j dG Qq d
dG SG OJ :j e gh hC S dG Jd J Yh
jG bH e eOdG dG GQTEG jEG Xhh Y N c id Y H
e IQOdG dG IQTEG M kdZ dG QG Xh qGE .G L EG dG
.H j dG JGAfh dW OGeG Y G L
YJ UN H e Qa ,dG H UG dG EG f CG g jh
GAf Y AdG Gg jh ,dM Y kFHc QG H j {RdG AdGz`H
SGH J FHdG IQTEG CG dG HCG KCGh ,zfGQ Y z YJ e
HG YdG J Ggh iNCG EG IMGh e z GdG z jW Y dG g
.63 IQTEG f
YdG j fGQ IY e dG Qd RdG AdG EG dG f RjEHh

G L

IGf
GT N
(HS) Y e

QG jGHn

QG
jQfce
jGQfO

jOGQGE Y N e

jdG
f Lt

j hQg

edG
cdG
q Gn
GQdG YCG T
jQdG YCG
jQdGYCG T
bdG YCG T
dG YCG
VG YCG

cdG YCG
dG YCG
idG

YH dG GQTE G jGE q j
G GE G AfGC e Fa
Y J dG dG jG SGH
.ac G AfGC J T T
f J FdG YdG g qGE
QN J GP dG RG c
.jd

ac G AfCG GQTEG f Y dG jG Jh ,dG IQTEG d G


,f bdG Fch FHc GN J jdG gh ,H UN jH
a ,jdG g SGH j G AYCG FSh G H hCG G AGLCG H Ja
jj JAGb hCG jEG J hCG dG GH ceEc jOY cM hCG YCH eb
e dY LQO Y dG fjN GdHh eLCG NGO jM Y fH
b e ed OJh JG Y g dG gh ,dGh HdG
GG e iNCG M Ie qfGC fLd c Y gJ d dG dG jG
.dEG

RdG AdG

175

fGQ IY

edG I`e

{HSz`d G q dG
He Jh ,{HSz YJ Jd IO f Y Y N H JG qj
HG g OLHh ,dG Qd IOG YdG jf hCG ShDhQ Y {HSz
ce e ke MGh BG Y N e cCG e JG q CG dG G J
BG h JdG GSH H e Kj Sq dG e Gc GOY j dG JdG
AGLE J dG JdG ILCG e je dG Lj :H fQe dh .MGh
j Ggh ,e jJH {HSz He OY Qa q G eCG ,MGh BG G
qC H hj AdG L b dG G g qEG .64 MGh BG JG jJc
.65{dG G`g dG GdG MCGz `g fEG G
`e BG IY Y N q a ,NBG fL e dG dG g NCdh
N BG IH JG CG J Y N c CG j Ggh ,66{HSz
RL H JG dM Ln GoJ dG HdG QdG jY QJh ,iNCG Y
IH JG J dG dG G QG RYEG d j g eh .Jg
.G Y hM Y dO EG RYEG Gg eh MGh BG iNCG Y N BG

JG e jG e AGLGE
G eGC dG AfGC h MGh GB
Y jJc a a fE G
1000000000000000.JG

176

j hQg

hQj jb bh M AdG c
qCG qEG ,{HSz e e e OLh oY
{HSz e T EG UJ dG HCG
FdG UdG d kJ J EG dch
.YEGh gdG bQZ AdG L
Gg EG UJ dG HCG g jLCG bh
jQCG QahdG b e dG cG
Hf IFL Y RM FV Yh fc
Gg J h .2000 d dG
c jQGC QahdG
Y HS e j :j dG dG
M j CG G S Yh ,dG IQTEG IT Y Gk OYG T d e dBG
dG jG bH EG IQTEG g jEG j jdG Hh ,jb Y IQTEG QY dM
Y HCG AGLEG Y f dG EG Uq dG NBG fL eh ,kT j CG hO
fEG dG RG CH j fc QahdG Mh ,jdG jdG GG
AGLEG Fa HU kfMCG J HdG e dY LQH j keY jdGh kUN
.67 dG HCG
dG jG FdG JG
U Z Ggh ,a FHc W hP dG jG H JG CG ZCG q j
J N e j Ggh ,e Gk c Gk AL J dG Jd FdG dG qC
J Fc jH e c AL j dGh Y N H QG JG
.JG Gg EG J cGJ IY
,dG FdG q dG OY J {q dG bq dGz YJ UN jL LJ
fN d dG QG OGeG Y M jh G L NGO jG g fEG q jh
bf BG N Y Ya c h ,dG QG R IU Ya NGO
f EG Gk OSG H a J dG jG CG dG HCG KCGh .68kjJ Y
177

edG I`e

bq d kUN ke Y N c J iNCG IQHh ,69J dG dG bdG


.d FG dG
dG G eCGh {SGz`H dG IQTEG e QJ dG dG G J
SGH Gh SG H JG jh ,{Gz`H Ya IQTEG J dG
.70c e (0^00003) G H dG GdG jh ,{HSz He
,IQcG YdG IOLG dG bdG d J Y IQTEG qjCG qEG
J M IN h ,G AZ Y dH IG jH G YdG Jh
f (jG) bdG g qJh ,{HSz e EG hCG dG GdG EG jG H
Y IOLe UN e EG dG IQTEG J jdHh G G AZ
.{ q dG bdGz`H YJ UN jL g dG g UY q gGC h ,Y N H JG Y
G jM

dG RjEG bf
e

HS Gar
G L
Y dSG

dG QGt
jCG Ib

jQfcG
ScCG

hCG AdG
fdG AdG

HG GM n

hCG AdG
fdG AdG
HS Gar

178

j hQg

.a Y hGC H RG j qGE h bO Sg HM Y SG RL GE e GC aVGE J


Gg qGE .aOe a OLG fG j GC h ,SG e kGJ cC a G AdG eGC
..Jh fS dG b e G Y hM Y dO iS d G

dG FdG G Yf H H a bdG Jh ,G G AZ
G G b e dG IQTEG SG j Ggh ,dG IQTEG dEG J
gfcP dG GG e IN c CG EG f CG jh ,dG bdG SGH
AdG gOLH j RdCG gh ,j e EG LH Gk RdCG J jdG WH
.71fCG
G J edG ch ,dG G AZ e YdG eJ c hfh
e aVEH efG J h ,H UN HW P efG hCG OG M Y
CG aVEG qGE :j e QdG gP EG QOj dG g Yh .SG RL EG VEG
Sg FM EG J e GN JG f j SG EG e
a Y M hCG eDhJ Y EG ODJ CG H WYG aVEG jq CG qEa qGE h ,fhdEG
cCG cdG P G AdG e YdG efG j G Hh ,SG
IG MGe Hh c dG gh ,WYG Y SG e Gk J
.qLh q Y dG M jYQ
179

CH QdG

bf Y N
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G Qj e YS qfE a L e H AG j eY
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j hQg

QNC d bdG jG id eGdGh dG


IdG hCG ,Y q N H JG Y ITe IQH GKDj eY dg
Gg bdG g ac dGh ,HS e Ga dG bdG J dG edG
g J dG bdG a ,e dBG ah Y bf c f Ojh ,G
H SGH FGOCG H J j NBG dGh ,dG IQTEG f H gh
fCG SGH J dG {dc SCGz IG bdG dP Y eh ,IYG fCG
.{SCGzh {dcz JOe EG UN
LQEG dBG gh ,iNCG bQN dBG Y dG jG dP Y IhY
Ie eGS fN jh ,e FGOCG H iNCG Ie SG G EG dG bdG
eh ,G jG H SGH IG g h .iNCG f Y iNCG
ie J IG gh .{eJGSzh {eHhO z IG dG bdG dP Y
J dG dq dG OGd SG MU IOYEG Y e SdH dG jG IAc
.jc Gk ah Gk L e Jh VG bdG fEG IM ke kfL
q ch ,IOe MG dG jG H FdG JG CG Y cCdG jh
bf EG JG Y qa ,jd bO jRGe Y Gk OYG e Me
.JG Gg MGe e Me H UN fCGh JhH EG h e Y
,dG bdG jJ Gq e OY J iNCG jQhVh bO jRGe dc gh
dG GdG dch ,G d g bdG g bJ dG IdGh
SG jRGG e aGdG dG dP eh .fN hCG bdG g J Y dN
.dG Jd d MGe g
Y f H HUG h {cQH Vez Yj fEG j Ve K
dGh
q ,dG QGh cG HU VGYCG eh ,dG cM H SJ OLh
SCG IG bdGh eHhO Iq G bdG H RGdG NG VG Gd FdG
EG ODJ G jN H J dG {eHhOz`dG jL e IFGdG q da .dc
b e kjM acG q dG G gh ,dG cM Y IdG Ga
181

edG I`e

dG dSGs
eeCG IdG
G G

Y Nt
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IQTEG

IQTEG
fCG G
dG eHhdG
dG eHhdG
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dG G e

N e dG IQTE G J
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QG jf e FHdG
G GE SG d dG
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g MGC zeHhOz`dG
.fedG dG bdG

EG .2000 d dG Hf IFL Y RM dP Hh dQc aQCG QahdG


jRGG a ,Y M iNCG Ie d j dG jG H QG JG Gg
X H dGG aOG f J HG dBG EG aVEG bdGh SG
a G gh ,fEG e gq S dG gh ,Jh fS dG b e N f
.Aj h Aj a
182

j hQg

fG IQdG ij
e je
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Gg jh ,W
e kGMGh VG
J dG VGeC G
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d FGLGE
GE Ud dG
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dG jG FHdG JG
f g dG Ggh ,a c e hCG J hH dG G J
IQTEG f c dh .FchdG dG SCH bdG SGH dG IQTEG
J dBG gh ,dG G H J dG RGdG dBG Y WEG j dG
G NGO ddG fjCG gh ,G NGO FdG fjCG H UG RGdG
,IMGh Le T Y jj GqdG SJdGh jOdG fjCG :j c g dG
ddG dG hP QdG jCGh LhOe Le Tq Y j dG ddG jCGh
,ddG dG GP JhdG jL H LJ fjCG g EG aVEGh ,IMGdG
,{Y RjEG OLh Y hCG MGdGz Ia Y dS T fe dG G Jh
fh ,dG G NGO dS T GP JhHh fjCG LJ dG g h
kSb bCG QdGh jOdG fjCG fh LQG SdH kSb YCG SJdG fjCG
ke kJJ qJe IQcG fjCG c gh e f EG f CG jh .f SdH
.72G HdG f Y a c UN Ge ah
d G AdG IOLG UG G EG dG IQTEG f Y
J h ,ehdG d b j Y J dG e dGe S dG

183

edG I`e

,e H dG g W BG M
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J IdG g jW Yh ,G AdG
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q ddG G
NBG EG Sh e fjCG fG Y
IQTEG dJ EG dG G EG qdH
dG gh FHc QJ hCG FHc jce
IOLG jG G J dG dSq dG qGE
gh jT RjEH HS gMT
e GKC J dG G AZ
dCH MGH QJ jd Ib eR gH
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EG j M Ic YHh dG QG OGeG Y FHdG QdG Gg jh
dGh Fc GJ hM EG ODj dHh ,{HSz e EG eh YG jdG
Gk e 120`H QdG Gg YS QJh ,jG bH EG dG IQTEG jEH J gQhH
432 OJ YdG g CG f CG eC H HM Y jLCG dh ,73fdG
.YdG Gk ec
Ia EG LJ dG IQTEG jEG Mf e dG G e cG Yh
GdG fG EG Gk OSG j MGdGh qdG H hdG Ggh ,iNCG Ie {MGdGz
e dCH Gk MGh OJ jd Ib eR Ia bfGh SJdGh jOd fjCG
SdG Gg e j CG fEG j a ,jd ge Y IgX gh ,fdG
a YdG g eG fCG Vdh .dY J GP bJ YS GSG hH dG
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184

j hQg

jO

jOdG Ib

H G FdG

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H G FdG

eRHdG

eRHdG
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T Y jj GdG jOdGh SJdG jGC gh eh da FXh GP fjGC Y dG G


SJdG jGC fh ,ddG dG hP jQdG jGC h LhOG LG dG hP ddG jGC h ,IMGh Le
GC QYG H NC f GC jh ,a jOdGh jQdG jGC f eGC ,LQG SdG e YGC G NGO
.fRGeh HK dG a dd ,bdG RGdG Y ad IOLe a WYG IQH LJ dG g

Lh EG djEG Gh IMGdG fdG N dG GQTEG je fa dh


dG GQTEG g j fdG e dCH MGh N Jj C N Ca HG
.ec Va

eGq dG G
bdG ja ,jG GfCG bH Y Ie iNCG H dG jG J
kb gOY j dG Hh .g c HK a dG jG eCG e O dM J
d MT g EG .dG M kjhe j NdG Me LEG OdG qCG EG
IM GW FXd FGOCG ch dG jG Y eY If jT RjEHh QdG jY
Oq DJ fedGh jG qCG qEG ,HSGh MT e a HU CG Hh ,fEG
h Gg ej Mh VQCG Lh Y fEG hCG Lh e J hOh IAc H Y
.dG e jdG LCG
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185

edG I`e

.NB G dG H Hj dG UdG GB h dG jG

JG Gg e j c ?fEG L dG jG GQe H dG
q GC hO dG jRGG g J c ?kW c dh CN CG hM hO QdG
?N
jdGh ,kGQGeh Gk QGJ ? {cz SG IGOCG fEG j CG dG e
jOe ie Y d {aOGz VH YCG J AdG H Y Mj CG

186

j hQg

jhCd Sc dG G jhQGOh
SCG gh .bQG dG gGdG g
pG n n s dG
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H EG Y HLEG Y jOG j
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@ np ndn rdG uQn p VQr nC G uQn hn
ie Y d G c J
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fG J e kWf keYh
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Y :e QdG jH G G
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d QfEG gh ic HhcCG fc Y ka Y dO CG EG j AYOG Gg
.gGh Y jQP
gh :YG jGh SG dBGh jRGG g e Qd Mh J Ljh
LhCGh dG jG g N dG a ,dG Q dG g dG Ggh ,d dN OLh
e fEG Nh ,UG jeh jRGe d Lh ,JG dG dG d
qfEG ,fEG e Ie JhdGh jGh GQdG Lh ,a c d bh dG
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187

IFLh AGdG J H G SdG g e :gMCG dCS d


H bdG Gf kjZ GDdG Gg hj gh hC ?edGh Hf
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.LhdG
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gh ,NOFdG eQh ,S KCJ hPh dG Y RZ gh ,LhdG
IQH IMGh cCG IQP OG e fe H jL Y J
e LhdG dch cCG Yh .LhdG e IMGh
SG SGQdG jq CG qGC cj ch ,jd ahG FdG UdG
hRBG hCG LhdG RZ Y j AGdG e %78 CG qY
.cCG RZ Y j e %21h
{dGz c qCG gh Gk L q e f EG f CG Oq fh
,LhdG cCG cJ WH Y a J HdG QdG
q
KCG bh .fEG IM NOQhO EG dH U dGh
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Gg qCG HCG g H bh ,jG H JG Y kSSCG
ddG J kSSCG Gk QhO jh edG S j cG
CG dG RG AGS ge Fc bc jh ,jd
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dAd VG Gg jGH OQGdG GDdG EG LQ dh
ke cG Gg GOe CG ?AGdG H NO bY e :Fb
dG gh ?AGdG H bY a fEG Gk L
Gk L S RdG Gg qC c q M EG gd Ie GdH
Y ka ,G QeCG S jh AGdG jh

:`jdG cGC
QG YqdG

edG I`e

.dGh hRhCG W Y jdG QKBG


GQdG J a IH RdG Gg Lj
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EG RdG Gg Y j c jdG
Ya jCG GfGh G edG
q JG
Gg CH dG FdG Mh ,d iNCG
MqdG J AdG Y YCG dG RdG
Sh AGdG J j RZ jdG cGC qGE
a OQ ch ,dG ShCG dG
W jJ Ff GE OD j dch ,G QeC G
Y M Qhj keY ShCG g
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dG q dh ,dG FdG g jJ
ShCG c iU MU QdG Ggh ,Ib Ia Z FdG g U KCG
{dG jLz LhdG cCG e IQG {Science dGz L M dG
GH dG HCG UGJ ch .74 1992 S hCG fc QOdG gOY
RdGz :e IQH Y jG CGHh ,JTh LhdG cCG gCG OGORG CdG
.{qdG bqdGzh {bQG jGzh {dG
q
cCG QhO JKEG f 1998 d dG Hf IFL Y AdG e KK RMh
{ha HhQz :g AdG ADgh ,jG H a JG LhdG
e IQOdGh IFG YEG F OQh bh .{OGe jazh {hQjEG .L.jdzh
cCG QhO ac IFG GSG AdG ADg CG UN U De AKCG Hf L
dG HCd jb aO hM EG iOCG cG Gg CGh jL bc LhdG
Z iNCG FGL Y GhRM KdG AdG ADgh .75 eeCG MQ baBG ah GG
.dG dG fd Hf
,LhdG cCG M dG AGLEG Mf e kjQfG Gk Y VG dG ch
Hh .H UN
q q QUh ,LhdG cCH UN L SCG M
190

j hQg

GhRM jdG cQa HhQh hQjGE .L jd h OGeja ( dG GE QdG e) KdG AdG


.G JG LhdG cGC QhO ac f 1998 d q dG Hf IFL Y

dCG 32 j Y c dG G U OY qEa LhdG cCG L S


j CG NO Y KHCH ahG AdG MCG VG FdG g Hh .76 jj hCG U
H b b FdG g qCH YGh jG H JG RdG Gg QhO Y f Lh
QcdG eq CG .77{GOfe hOSz QcdG g dG Ggh ,Y Y kSCGQ dG gG
kbaBG a bh Y cG Gg qCH OaCG a OQfS eL MdG {cc Lz
.78dG GH dG HCG eCG SGh
cG Ga ,QdG IYd c GMEG Qe c cG Gg qCG g Gh
dW j dG jG g qC QdG IYd kYe kY kSHc j NBG cG Cch
,fEG L IN e Y dhDe dh Gh dG Y G e QH Gk MGh
e MGh g LhdG cCG CG
q T a ,kGHCG aOe j CG eCG Ggh
.Lh Y dG OLh Y gM hCG gY dG dOCG
dG OLh Y dG dOCG c kWb kaQ aJ GJ a jhGQdG ShCG eCG
Ggh .YCG aQ dH gh ,GH CG GQdH ke kdO cCG AGS Jh fS
J c dP Y c hfh ,e bGe EG iOCG e Gk c G adG

191

edG I`e

Ja ,LhdG cCG FXh CH dG acG Y ADg YEG FSh


AGWEG Gg d YGO dh ,Rj kjQSCG kH RdG Gg e h dG Sd J
g GJ e RGeh UN f V IN kee ODJ jL ka dg C jGh
dG g q Lh ,jGh cG e d hCG RdG Gd G QeCG e dG
dN OLd gQfEG Y Y Gj c fdDj GhOc M RdG Gg FXH jdG
.f AJ eh JOGQEG e J jG g qCch ,LhdG cCG j
Gg fjOh ,{a bh NC qL M ez FdG G Y j g dMh
fC dGh cdG Qe fC IdGh ,dG Qe fC q dG dCJ Y j
.d jG ALRGh jM a Md Y G eCG j Gg geCGh .AdG Qe
dG QdG MGdG dH EG eq EG ,KG e MGh jW EG fEG eCG Lj h
uco nYn n go hn oho oYr an Am r n T uco opd Nn n go sGE p nndGE p n rotHQn odG oopdPn }:dJ b ,d jT
JH j e dCH CGj CH NBG jdG FM j CG hCG ,102 :fCG|l pchn Am r nT
.M Z CG kq M cCG AGS NBG AT CG hCG jG hCG jG hCG GQdc bQG JGeh
dG dEG q dG e IMGh EG d LhdG cCG jL qCG T
N G iG Y dG acG gCG iMEG dc gh ,fEd IG
q
c Y e qf Sh .G Y IgdG dOCG L EG VCG dGh jdG dG
dGh fEG G jN ZCG fG Y Hf IG FXH jG g b
.N jJ 100 gOY j

jedG YhC G dG
AdG YS e hCG fc fC dEG dH jGdG f jedG YhCG d
dGh jedG YhCG qEa he g ch ,LhdG cC dG FG c
dGh ,fEG L jedG YhCG T J h ,GQhdG RL J dGh
,G AfCG L OGeG d jdG dG e H ec dCG Fe e cCG dW j

192

j hQg

YhCG g CG EG Uf H HM Hh
IdH WMC H fL EG H Vh d jedG
.79

fh Je VQCG

G jedG YhCG CG g cPCG CG OhCGh


jdG e dhd jdG dG T e Gk J cCG
VYh e je ah q J jdG dGh ,IG

dhd jdG dG T e eLGC IOLG jeq dG YhC G T qfQb d


.kGJ cGC eLGC T fd jeGC e ee

edG I`e

jedG YhC G SJ qGE


d VjdG SQ AKGC
q d FQdG LMG
g qGC Y dO g
.jd kbQN kJ

NG e J IO G Y h HK e
jedG YhCd NGdG VdG eq CG ,jQhG adG hCG bdG
ddG H kV hCG kYJG J fCG CG ,HK a
kFQ Gk QhO jdG Gg jh .H J dG G
RGdG Gg EG Gk OSGh ,dG V RGJ Y G
,a Lj dG SdG H kjeO bCj CG G j
AKCG V hCG VjdG SQ AKCG jeq dG YhCG SJ
q eh
f V jedG YhCG d f EG e H HUEG
.RGe UN
q
gQbCG J bJ jeq dG YhCG O c dh
GDdG Gg Y GG qEG ?SSCG CG Yh ,SJ e
H jCG h kW c e CN CG CG VGda ,kGL e
ODj ec dCG 100`dG GP jedG dG g Y
VG M AdG ch .gY j IN Ff EG
da e jedG YhCG NGO j e j jdG
Se GL OjEG Y ke jLY Gfc q dh Ie
OLh KCG HCG dh ,kfBG MW dG GDdG Y
J jG a ,LhdG cCG jL gh ,Fc bf
.jedG YhCG EG {JGz `H eCG
YCG jG g dJ M dH ke GC dh
G jedG YhCG q J a .jedG YhCG
hCG M WH Y Y cGJ gfLd hdEG
EG Vh d jdG jedG YhCG e IY qC U
SCGQ IT b QH gb eM d H fL

194

j hQg

IQdG ij c
jedG YhC a
hGC J GC J
g qKC J f J
.IG GQTE H

H Ie NGdG fH c e jedG Id gG dG
q Ggh ,fEG
Gg qCG qGE ,edG AYdG J hCG SJ Y hDG g dG Ggh ,eCG Y
jN Jh ,FW f e FZ W J H q dG e S j dG
S e M qfCc H fL EG H UGH
q edG AYd FdG dG
jdG dG fK M q j ch .edG AYd FdG AdG jq dG d
,dP iS AT h edG AYdG NGO dG HfG e j FdG dG Gg CH
LhdG cCG jL fEG e IOe FXh dG Gd CG H a JG dh
.kFc kbf j dG
.G Gg fEG LhdG cCG Ea ke FdG dG N fYG d
dH g OG dh Ic eh BG a e gP EG QOj ke f eYh
G Gg Lfh ,G e H MGh M j edG AYdG NGO Gk L U M
g dh .G e QH Gk MGh dW j dG LhdG cCG jL EG g e G
FY dG q M H UCG jdG cCG jL CG ke Qd Gk L IdG OHCG
.80VQCG Jc H dG Ic J

195

edG I`e

J Ae H je q dG YhC G GQL
W J hGC q SJ M GPGE h ,fGh dG
NGO j dG jL E a jeqdG dG YhGC NGO
J dHh ,OGf dG edG AYdG j IC ah
e C H QdG GC jh ,dG GE IOQGdG dG c
dG LhdG cGC f JJ dG g .QdG
.dG d jG jedG YhC G fG G OD j

eC G Y qdG :LhdG cGC IS Y I If


j dG GdG g Nh ,kjJ GK Y LhdG cCG jL Y j
,Lh cCG Yh HG G GMdG EG eCG jEG LhdG cCG
L h Ic YHh FdG dG N e LfEG Qa QfH CGJ Jja
.eCG dG dG j jG ZCG GNG j e dGh ,gG
Qh j FfG jq PdG GP AG q dG G AZ qGC dG HCG e jh
fCG OjEH gQa e GC J G NGO Ldh Hh ,dH Y kae jG g
,dEG dH ic gCG j dGh dG eCG dEG d {GCz Yj UN
196

j hQg

NGO HG YdG e S GC J K eh
.G
NO `dG jL g bdH Yf dGh
,e (10-9 *1) dW jh jQqdG GP
j e d jdG Yc j bdG Ggh
jY qcPCGh ,{GCz `dG fCG CG ,dEG dSq dG
G fCG BG Y q G qCG QdG
Mq dG e Zq dG Yh ,kjCG G FXdG GP
fCG OjEG Gk HCG LhdG cCG Aj
cCG h ,JM b Y ka ,d SG
dh jdG Y Qd UH jCG LhdG
.kGHCG jW Y j
gQhd FGOGC LhdG cGC jL YS qGE
e LhdG cCG REG YS fcJ
Y JG jH f j G JG
T Y hdEG jdG SQEG YH
E G jdG
.hd
jdG e e Oq Dj fEG kah ,ffEG
dG dSdG GC fCG SG a .kGL Ib Ia IjY FSQ qj fC hdEG
GTP`dG jL j Y j Yh ,kGQa dH GC j LhdG cCG dEG M
Y jG q dG Gg hM W GJ eh ,GMP`dG jL EG bd eG
.ac MG j e jh ,G
AYd dG jG ddG f J EG fCG g ODJ jT RjEHh
YhCG SJ Ggh ,dG jG JJ dHh YJ dG dCG qGC CG ,edG
eCd bc LhdG cCG j dG G QhdG f j Gg c .jedG
qGE d jG dG g qGC g f CG jh ,dG AJQG EG ODG FdG
.eLCG NGO c dG jG dG jH e IMGh

197

q eg

FW N
qG dG jN
LhdG cCG

IGf

qe jT
GC

GC

jJ
GC
Ca++
cGMP

GTP
cCG
SG

Se jT

Jh ,jdG cGC W j dG Gg Hh .jdG Y IOLG H FdG dSd eG edG j


G jG jH j edG fC G Ggh ,GC `H Yj a kUN kfGC d AG dG f cG Gg jL
fh .G d G RG f ddG fjGC LJ GE OD j QhH Ggh cGMP `dG jL GE GTP bdH
.jedG YhC G OqJh AG dG jG J GG d

j hQg

Z jL YSG CG M c :ke ,dG g Y DhJ IY QJ Hq Qh


hQG AdG Y dGh bQG G fCG Y J CG jdG cCG e bY
G bdG Y IdG IQdG g jdG cC jCG e ?c Y fCG
M dH gQa e GC J CG jG g J c ?Y bdG hCG dG hd
S hCG CN CG hO ShDhG EG eCG Gg jEG J ch ,jG b e LfEG
?bQN YHh
qfGE ,JGP AJ e bQG YCG g H j LhdG cCG qCG T
dG dG IQb Y gTh ,G Y hM Y dO dG jG bc
.dG

LhdG cGC fGE ce :(FdG G) JhfGC q N


-G eCG G : e ZCG hCG d e hJ b dG H g
FK Sa ,LhdG cCG fEG Y hDG fCG ,{L-arjininz LQCG `d
K ,adG ,Jcf hCG ,chCG ,dhOc ,eCG Jf jOG Jcf
G CG qGE ...(jHH hQg KK) LhQdG jHH ,jOCG adG Jcf
.LhdG cCG fEG eJh aG jG g J FdG
FdG jG e cCG FdG OGG fEH J dG eG qCG jd hG e
e Gk c bQCG FdG G eG dG CG qGE ,GG e fjdG fjH
cCG fEG Y dhDG FdG d J Hh .kJ ee f dG dG eG
g L dG bGdG FG g e dh ,SdG dG Gg d S LhdG
?dG Lh Y fEG Lh e bJ hO J G
dGh LhdG cCG fE eJ dG OGG Gk L J G g EG
N O RhdG cCG fEG j ke ,CN C MG Lj h ,e c e eRdG
2
gh ,kGHCG aOG Y j FdG G fEG qEG ,CN (G RdG)

199

edG I`e

fc d FdG G CG QdG jY qcPCGh .fEG Y J jd bO jRGe


dP iOCGh jedG YhCG c J G e bCH LhdG cCG J
Sd G e cCG LfEG c dh ,dG Hq dG EG dHh dG V IOjR EG
dh ,edG
q EG dHh dG V VfG EG dP iOCGh e H jedG YhCG
.(dG G ) Gk HCG G ANCG g e JJ FdG jG
,JM e c d OGSG gCG Y J IQcG jG EG
fEH jG g Jh .LhdG cC MG g j M fEH Gk Qa CGJh
J h bCG h cCG dH LG QH j LhdG cCG e J a G
.G LM J kM S fc iNCG LMG C J dHh ,fH
gh ,Ie fL hCG jfK e jCG Y fCG jG d iNCG IG
hCG AGdG FdG eG K dG dH fQb d gCG Qf CG f IG
dHh ,IG e dN ODJ GK Y H J jdG cCG ja ,dG
FdG jG g qCG J dG gh .kGHCG G edG jG g cGJ
.Je d G jdG J (jG eG)
eJ dG dGh .dGh Sd cG OG UN fG J
IdG IG hCG fd dG dG M dG QdG J fG g
e IQb J FdG G dch ,a MCG j M gh ,fG g U
.Lh Y dG IQb Y gT L N jJ 100 fL EG gh ,Yh N

dG OLG YqdG
GC fh ,jeq dG YhCG LhdG cC JGSG QhdG Y ke WG b c
fEG Y Hf H j dG QhO ,fEG IM hCG dG e QhO hH BG
`H OG gh e eCG jEG j jH CGf CG bh .FdG IQH q j dGh
e LhN hCG JOh d g OG qEG .{fEG IM hCG dG z
200

j hQg

dG J e J dGh N hCG gY J dG dG g OG H ,Mq dG


.jd
FdG YdG e S hM EG ODj jdH dG OG qCG hG
jO j h ,d fc dG MGG e jdG g qdh ,G H J dGh
q g c G fb jGH ez :MGG g iMEG Y AMCG Y UNG jCG
.81{dG b e HNEG H G
q jdG AH c j Y Gk qce AdG
cCG CG GDdG Gg Y GG OjE jLCG dG dG HCG KCG bh
V da ,CG MQ G q dH AdG IQTEG AYEG Y hDG g LhdG
cCG qde CG LhdG cCG qce hCG NOS Yj UN fCG Y Jfe
GL e bHh b GH NEG hM b Nj fCG Ggh ,LhdG
LhdG cCG jL J ITe NEG hM Hh .LhdG cCG fE
OLG ddG j fK KH dG g Hh ,jdG EG dG e bdG
.G q G fG MGe CGJ SGHh G jdG
e jG EG h G G f QG dG Gg e d MGe gh
H JG hM Y IQdG GfG OJ VG G dGh .dG dG
UJ q dG G gh ,LhdG cCG jL OLh Y Y jdGh dG
G g AGOC eh LhdG cCG qCG Y q J 2000 S dEG fEG
.dG MCG dG Qa ,J dG IQH G

ShdGh jdG q V dG
Gd qCG HdG QdG LhdG cCG FN Y jG Ve OQh d
IG gh ,keq S Gk RZ fc fL EG FdG eCd bc q e JM Ge RdG
j LhdG cCG ja ,fEG G ehGh YG R eYO J INCG
iMEG Jh {ahcez YJ dGh Hhd IOq G G b e gRGaEG
201

,LhdG chG ce hGC (NOS ) `H Yj UN fGC Y dG :204 U IQdG


NE G eh .LhdG cGC fE H GC d NE G hM e GK H b fC G Gg Njh
J fK KH QfG Gg Hh .a Jh G jdG NGO LhdG cGC jL J
.OdG KdG jh dH G fG dGB GC J dHh ,ddG fjGC

SCG qJh ,qe 0^01 dW j G g qEG .G Y adG N RHCG


CGj e kHhe OG G Oj ea ,IQdG jGh jdG HG
eg H GC j WMEG Y cG Hh ,ac fGG e WME G JGOGeEG
UGN Y GOYG FdG YdG GC J q K eh ,LhdG cCG jL e HGH
.hG hCG jdG e EG ODJ dG edG
q jdG cCG jL
e M dO d LhdG cCGh OG G H dG Gg e M qEG
cCG j eh NBG QhO Ljh .OLdG Gd dN OLh Y IdG dOCG
da j jdG cCG CG HCG KCG a ,G ehG RL V LhdG
Y hDe fCG Ggh ,(protease) {RJhHz Yj TGhdG H OLe UN fCG
,dG TGhdG LCG AH eGS JhdG d bdG jG J

202

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H TGhdG KJ LhdG cCa


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d jG H bdG g GMC
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jQL HCG CG VCG CG OhCGh
aG dG FG e jG Y d
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GQhCGh WdG V c S
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Y khCG dG j eCG Gg dh
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203

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g qCG kdM hGh ,FH kHYEG hOGOj AdG j j ch Y e


M SCGdG b e CG G AfCG L LfEG j FdG eCd bdG jG
,a a QhO dh G jG ddG ZCG jG g Jh ,dG NCG
QJ :ke YEGh IG a j dGh jG d G QhdG d j JBG Gh
jd J FY , Mh LhdG cCG e QhdG NCG fCG QdG jY
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j FY ,ec dCG Fe j jdG YhCGh IOQhCGh jGdG W CG
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204

j hQg

c aG J CGh , CG OQCG GPEG d HdG JLCGh AYCG g M


c H a eJh H J ILCG g qCG kY ,FXh ILCG g AGOCG
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jCG hCG ejdG Y AGOCG jh jd YCG j fEH HdG G J
f S ,kGMGh CN Jj CG hO S S EG S Ih iNCG Ge
ILCG aCGh GG bQCG IY M eCG GH dG dSG QJ CG EG Gk e
.dG G g eCG LY fCG S kYh Gk N c eh
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dG jd AY gGLCG e HCG N e jdG dG e NCG dG dEG
OH J LhdG cCG FN EG UdG qGC EG ,J dG
aHh G SCGH gH dG SDGh hdG eb H gMh MdGh AdG
gh ,IGh jQG HCd aO YCGh LddG feEGh ILCG
dGh G G Gg H dEG UdG dGh LhdG cC IjdG FG
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205

Fc jL KCJ c dG Gg U Y VSG
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H Y JH N H JG Af c SG dch
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G Y Jh fS dG IQb Y M eCG OLh Y GOT
.eLCG YCG YCG
UdG Y Jh QdG g QdG jY GC J eYh
AKCG M bJ GG g CG J G GH UG
:dP Y eh ,dG Gg IAGd He
ebh ,dG AG f SH L jG H eb
,kGQGeh Gk QGJ dG dG f SH iNCG jG H
f Y G JY Y NCG jG BH Qq jh NBG dGh
dG EG dG e ddG Y f h ,dG fRGe dG
.iNCG kfMCG dHh kfMCG
,d dG jG e dq dG jd IjL jN dJ q h
fjJ eb ,fH jG H KJ J jW Y dPh
IQGM LQO Y d j aGe Y e AGOCH jG
f SH jG H ebh ,e OhM Vh HK G
d e dBG ah da IY KMh ,dG jOdG
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NBG kM fGh kM J a H G dG Ea jedG YhCG eq CG


.J hCG YhCG g O EG ODd
e iNCG kfMCG AGh kfMCG jOdG TH c IOLG jG H eb
.dG EG d dG FS
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dG cE L AGLCG e AL c ISh Jh Sb j QNHh
Y kHc GC J eY fCGh ,G IG UG FdG e dGh IOjdG Vh
.kHS gfcP dG jG ddG H L feg eb fedG
c qC ,dG fEG f,qdG :Nh qe AT JY Y j dG g Yh
g dOCG g jDhd AT aCGh ,Lh Y dG OLh Y dO AdG hCG VQCG AT
.dG
gjJ Yh ,dG e dG hP jJ Y Yj bG ATCG da
JGe h dG jBG jG M Qj
q e BGdG OQhh ,gQb M Lh Y d
:J b :j c
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p VQr nC G hn pG n n s dGp rNn pa nho sn jn hn r p pH o Lo nYn hn GOk o bo hn ek n pb dG nho co r jn n j p dG
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n Gn Yn n p an
n nf r o S kWp Hn Gn gn
n r nNn en n Hs Qn
Y dG jBG e jBG Y hO fEG L j dG edG dG Ga
,ic Z j H LdH f Lh e VG Gg hJ Ea dG Gd ,Lh
bQG ILCG QX ch L GG g hM c fEG a
.Jh fS dG EG bCG G Gg
Jh AG J gh fedG dG Y J e f dgh
dG IQb Y kX AGaG fc hJ dGh QdG jf J dG GAYOG V kja
e U Y Y dO d edG dG AGLCG H fGh edG qEGh ,OLhh

208

j hQg

.QdG jd SSCG G J dG {aOGz


Ge e e qJ f g dG Gg Qb JY Y G NBG LGdGh
j a jdG Hh , Sh aCGh SCG MCH SdG L H IgH dEG
dG ah ,J dG jBG Y YCG ah Sq dG YJ gh G jZ dG Gg
.Jh fS a J M

209

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Qq ``dG `jN
fc M RhJ dh ,AMCG N Ia gd X jf g :jhQGdG hCG Qq dG jf qEG
jdG g YqJh .QfGh CG Y IH jf fc EG aVEG ,U jCH dG EG nrn Sn
N K eCG ihJ e YS AYqOG Gg qdh ,aOG H M OGe e Cf IG CG
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dG N '' aQ IH J W FU J CG d QqdG jfh .Jh fS WdG
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G HCG AGLEH jRGG g CG fh ,jd SM jRGe ah IOLe a GQqdG
SSCG OGe J a ,kGJ cCG NBG JJ ah G dG IOLe f GQqdG g qCG qGE ,Y
.ge LQO EG SM jeh dBG d Sh Jh ,jGh fCGh JhdG cd
. jdG dG jf H jdG g YGe d kFQ kS c QG dG Gg qEG

jhQGdG eg dG YG
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g dG fO b We d Lh jdG g QH dG G qdh ,jZEG IQG EG gQhL
G FdG N Y Hc dDG VQjh .jhQGO dQJ dD '' GfCG UCG '' c QhU
ac G FdG Af Y G OYG EG Yj dP e kHh ,Jh fS dG b e G
.dG GdG hM f AMCG H NG X edG Qhh ,MGh L e
aGYH qGE d ''ke kL'' fc Rhj h ,Y dO CG G j jhQGdG AYOG Gg qEG
aGYG de IQH hJ '' jdG Ye'' SH H Y iMG dG EG M ,kT g
dH d ,SG OhOdG d CG jdG J dG SCG e jdG OLH f jhQGO
.jdG H ja GK
jd Ib e dG QdG d SCG d SG OhOdG QH dG j CG j ch

210

j hQg

kGMGh YGe s ah jhQGO eCG N jG dG dh ,Hc cP W dG Ggh .edG Qh


. NBG H
:gh Y jc jhQGdG AfG EG OCG FQ eGY KK cP h
.Ie hC IG QX c Y Y J OjEG ke J jdG qEG (1
.AMCG H d Sc '' Qd UN dBG'' OLh Ia Yj Y dO CG OLh Y (2
jf YJ e Y IMGh aO AMCG OLh d J GG j dG
q qEG (3
. QdG
:KdG eGdG g dH Sh

Gk HGC bG Z IG : IG UGC
.S Qe3^8 b IMh N e Cf cCH G FdGh IG qCG Qq dG jf Yq J
dG M e G G FdG GfCG e jG EG J CG IMGh M c dh
?dP J Ge jCG LJ GPa ,ka M b dG Gg c GPEG h ,cdGh
AYOG Y bdG j UdG g VG bh ,Y HLEG jdG J DhdG Gg qEG
X G g CG jdG YJ ?OLdG EG X c iJ .''CG G'' J Gh hCG
Z hCG LQN KCJ CG dg j CG hO dG hX X h gMh aOG f OLdG EG
H d KM M Z kGOGe qGC jdG YJ :NBG ,kWb kaQ G Ia aJ fEG CG ;W
. ahG dG YGdG ac e ke aj AYOG Ggh ,M N QX EG dH OCG aOG

''IG e C J IG''
dG gG W qj dGh ,QcG Hc IG UCG Y kGHCG jhQGO dQJ j
GfBG dG ch .kGL H OGe e AMCG J Va RhJ dGh ,Y IFS fc dG
sOGe qCG gOeh ,SdG hdG e JS VJ fc dG ''FdG ddG'' jf KCJ Gj e
.M OGe fCG h aOH b M Z
GG KJ e OYG Gg J EG SdG H J fc edG G H gh
H fc jdG GAYqOG g KEh .G eGU GdG KJh qdG a
M QdH GdG GC J kb QfG Yh , m H Tb b Y G e M Vh e QdG
. IdG J SdG OYG

211

edG I`e

jdG OYG Gg EG Sq dG bS a ,dG OhdG KJ gh iNCG IgX g fch


eG HdG SGH L j OhdG CG gh ;H a NBG AT KEG dh ,d kdO JGh
IdG fc ''GfCG UCG'' Hc jhQGO dN dCG dG IdG h .dG Y j dGh Jbd
e a GS N H dG GQdG KCG dh ,M Z OGe e C J fCG jdG Y IFdG
jd fdG AMCG Y gGLCG dG HCG jW Y dPh ,a AL e U Y dG dCJ
CG J G Z OGG CH FdG AYOG UCG d'' :j c KHCG Ff QSH jh ,QSH
82
.''`jdG e IG J
dh ,QSH dEG UJ dG dG dOCG V jW I aj QdG jf Y aGG Xh
Gg e QX dSG dHh ,G cJ H j dG dG KCG edG Y QH dG
. f AJ e G cdG

jdG dG dG hG
IG UCG Ve hJ e hCG jQHhCG QdCG AMCG Y ShdG UNG c d
J G Z OGG CG KE jdG dG qKK IjY kKHCG iLCGh ,jdG dG
kFb IQG j CG G VGh jQdG dH AH KHCG dh ,aOG jW Y M OGe OjEG
83
.'' eH QdG jf J AGOS f tnoj G UCG qEG'' :
KHCG GhLCGh jQHhCG S dG f jdG GhSGh ,QdG IYO e AdG bH SCj h
M 1953 S e fS jeCG FdG b e LCG H TCGh .IG UCG EG Ud
GRdG g Vhh ,dG VG G dG e GRZ GP OGe OLh VaG
cJ NJ dG eCG VMG H Y j CG SGh ,bdH gLh MGh e
. JhdG
OGG C ;a K e YS dh ,eCG EG e IN GdG
q J HdG g s Yo h
K dG Ggh ,dG VG IOLe fc dG OGG M J HdG eG
84
.MdG GdG cCdH
J HdG AGLEG eSG dG OGG CH j CG f e VG jW U Ia Hh
85
.edG dS G dG LJ fc dG OGG M
dhJ G gh ,jdG dG GW jhQGdG gGLCG dG QdG c dH AHh

212

j hQg

S f e V HS jO S dG G LdG AdG UNG GOH L


: j e G ALh , jhQGdG LdG GP ''VQCG'' U Y 1998
CGH c :gh ;HLEG d e LGf jGH c c dRe h jdG dG Of f ''
86
? IG
Id IG dG
IG UC dG VG Gg CG gG g QJ QdG jf L idG FdG dG
.Y bj H dG G Fd M ,jd e
h VG bh fEG U dG LddG e L e H YCG G FdG N EG
.dG IQG eG cCG M Z OGe H IMGh N fEG
aOG Y OSH MT fCG LQd ,kGL Ic M N jd eRdG hdG EG
dMG :G S Y) d SSCG M g dG Jhd paOJ jJ MG CG Z ,kbWEG
.bGdG VQCG Y ke J 1 / 10950 g eCG M e N d Se JhH jJ
e QJ ed Fg kH j G IGf G eG j dG DNA `dG sEG
e N JU OY ch , e J Y J e fQJ eG a,a
.U
EG fJ DNA dG jdG CG gh dG g jZ iNCG LGhORG kjCG gh
IOLG eG M j fCG g fEG dh ,UG ( fCG) JhdG H
a ,LGhORG J d f bdG ke gOLh e H a ,H KdG WJQ a DNA
G H jh ,JGP e Lh b IG EG :J dG IdG RCe bdG EG ODj
87
.''LQhCG d '' hG jhQGdG
jJ j dG jZ fe dG RNA DNA h jhdG VGh JhdG EG
CG hO gMCG fEG a ,ke Vae aOe jJ MGh ,f Gh f bdG
IG QX dSG gh f G UdG EG kGe fEG j dch ,kGOLe NBG j
. Fc H
g ,d QN H IG H f CG a ke aOG jH IG QX c EG
.G QfEG SSCG Y jdG Jee c H dG QdG jf J G

213

edG I`e

QdG jd dG dB G
AYOG Ga '' QdG dBG '' M QhJ fc jhQGO jf f kS fc dG fdG dG
AYOG Mh .G jdG Hb Y FGMG dh kY U d dG fO e CG j
Gg qGE M ,AYOG Gd FSG gCG YCGh ''dG fG'' f M QdG qEa jhQGO
. '' dG fG jW Y GfCG UCG'' SCG dG dG SG e j b e gG
hdG eAeh Ib J dG G FdG AH GC e EG j dG fG e qEG
hdG SCG jCG Ea SG fGG b e jG e b Ouog d :G S a ,dG
dBG g dh .hdG jS AjbCG jjCG e kdCe dG j Ggh ,IG b Y AdG j
fG'' J dG
q Gd .ke N EG d Cc ,NBG EG T e jjCG QJ CG J
'' GfCG UCG '' Hc V H cPh dP j c f jhQGO M h ,Qd Sc ''dG
88
.''G VdH j dG fG Ea HjEG GJ J W'' :j

Qe KC J
Y jhQGO LCGh ? HjEG GdG g S fc c :f j dG GDdGh
TY fa AMCG Y Qe h ,Qe e Y LQ e S e QaCG EG kGOSG GDdG Gg
,MdG LCG EG KQJ e GJ J AMCG CG Yj c GH jhQGO b eh
aGQq dG qE a FYOG Mh ,IjL GfCG QX EG Oq GC L H kL GdG g cGJ ch
jhQGO YCGh .jW MCG Y ddG QTCG GQhCG Y d Jh f dG e Cf
Y iJ fc dG HdG e Ob UCG G CG YqOG a ''GfCG UCG'' Hc He eCG
cG dG KGQdG fGb CG EG 89 NBGh G H AG EG hdG EG Ie fch FG FdG
fH FdG IQSCG jf EG iqOGC jdG dG G Y Y GC W dG QdGh e
e KCJ hCG IFa GP Z dBG ''dG fG dBG '' X Ggh ,NBG EG L e G dG
.jG dG f Lh

KGQdG GdGh jG jhQGdG


dG KK kUN gLGh dG jdG G eCG gOL H jhQGqdG b
,'' jG jhQGdG''`H Y e hCG ''jG dG
q jf''`H gSCG IjL jf GbSh jdG

214

j hQg

j edG Ggh ,dG fG fL EG QJ KCJ d NBG eY g jdG g Mh


GdG gh ,HG HjEG GdG J h kS J L hCG KGQh GW M
. d KGQdG SG C N f hCG YTEd VdG H qeEG
jdG g YsJh ,jG jhQGdG SG AMCG id QdG Y aGJ dGRe jdG gh
dG hCG PCGh dc cdG IGh AMCG id IOLG G cGdGh AYCG CG
,G cJ GJ hM hCG KGQh GW hM KCH EG J hCG J dEG ... Gh
AMCG Y QV eY J kFGO KGQdG GdG CG gh ;kM kY kqe LGj AYOG Gg dh
.jCG e j IFa GP J h
e FGY jL J CGh ,jd cdG Ie DNA`dG jL Ea kGL VGh dP Sh
jeCG KffGQ.. Y j dG G gh ,S KCG d j CG e H kW c
h ,QdGh FGdGh dH J KGQdG GdG qEG'' :j c G Y UNG
GdG qCG VJ dG edG FG g qEG .GMCG MCG KCJ GP Z Jh kGQOf EG
qeEG j CG d qH G G FGY J qGC qCG kUN ,QdG GMEG kGQhO J CG
EG ODj CG cCG Ma ,gjJ EG ODj dG YS FGY J qGC ka ,KDe Z hCG kGQV
90
.'' IH qKDe Z j CG hCG H QdG EG
.G Fd G dG EG qOD J KGQh IW OLh dG EG j qfC ;ka M e Ggh
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:Ja SZhO e Qe AMCG Y b b G
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94
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95
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,SOdG SGh ,WGG OJ Y hG e AG Gg Vhh ,dG f Y LQG aG
INCG IOG g Y kJ eNGR jh .fEG QJ kGNCGh ,FdG dG hCG QdGh
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99
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Y Id dfCG TCG OLh J G QKBG jCG OLh GfGh ,jdG g dEG YJ dG QdG
dG YGb e be J dG jdH dG Y dG IQhV EG Uf dG Gd ,jQdG
ce VQCG CG H YOG dGh jQdG Y iNCG jf fG c J CG Hh ,dGh
H gh ,dG GMCG LGh jdG g AH Y kY kGQGUEG g dh .dG
. AdGh dG Y g fCH jd f CG jh eJ iOj
G gh ,dG d Sc eGSGh jdG d G H J j dGh
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. dG OG ga jhd d a Pe
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q H G g J kfMCGh
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jOe GJ Lf j dG g EG Ggh ,kS H sn eo ATCH eDfh ,jOH eDf f''
HCG d OhdG YO S g jOH G fEGh ,FOeh dG YGb dh jfdG gGd
a kbWEG U jOG dh ,Y LJ dG ac gGd jOe GJ OjE jQG
100
.''GMCG LGh EG J CG qdEG Gd f CG kGHCG
ka kHGJ HGe fc O jhQGdG H J dG dG ie J dG g EG
Z OGG CH eDj dG Gdh ,IOG a AT CG jdG g UCG IZ jh ,jOG jdH
QdGh aGQdGh ASCGh Qdc G GfCG e jG EG CG ,G OGG OLh S g G
Y GC W dG NGdG d kLf EG d fEG Mh Gh QgdGh QTCGh GGh
.YGdGh YdGh G c IOG
Y aGj GdGRe qjhQGdG qCG EG ,dGh dG YGb e ke VQj OYG Gg CG bGdGh
.'' GMCG LGh EG J CG dEG Gd f CG kGHCG '' aGgC eN FGQBG
WdG G ij CG d H jOe Z f Lh e AMCG UCG b EG j fEG q c h
dN b e npoN :CG ;d M Yh ge Ib KCH npLho b G FdG ac qEG ,dc
. a c :d bh dG e AT c N dG jdG dG dG g dG Ggh ,d

219

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220

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