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ជីវវិទ្យា សង្ខេបមេរៀនសម្រាប់ត្រៀមប្រឡងថ្នាក់ទី១២
ជីវវិទ្យា សង្ខេបមេរៀនសម្រាប់ត្រៀមប្រឡងថ្នាក់ទី១២
suImNUEs<m nigGg;sUEs<m
emeronTI1
suImNUEs<m Gymnosperm
1-RbePTepSgnsuImNUEs<m
-etIsuImNUEs<mmanlkNdUcemc?
suImNUEs<mKWCarukC
atimanRKab;TImYy EtRKab;KansMbkkarBarBIxageRkAeT vamanRKab;nnl.
-bcb,nenH etIsmuI NUEs<mmanbu:nanRbePT?GVIxH?
suImNUEs<mman 4RbePTKW
-Rbg; CycadsmanlkNdUcedImetatm:agEdlmanskw penAkMBUlxagcug
Etvamanplitekan. eBllUtlas;ekanrbs;Rbg;manragdUc)al;.
-kUnIEP ConifersKwRsl; EdlmanswkragdUcml lkNEbbenHbgarkar
)at;bg;CatiTwk nigbnSaMCIvitkgGakasFatust.
-KIgkU GinkgoesmanlkNFn;ngw kgVk;briyakas)an.
-sueI NtUPIt Cinetophytes
rUbTI1 Rbg;
rUbTI2 kUnIEP
rUbTI3 KIgkU
rUbTI4 sueI NtUPIt
2-karbnBUCrbs;suImNUEs<m
2>! srIraglUtlas;
-etIedImrbs;suImNUEs<m manlkNdUcemc?
edImmansNanCa edImeTalRtg; nigbBab;edaykUnswk bkenSamEmkenAkMBUledIm.
-cUrerobrab;BTI Rmg;swkrbs;smuI NUEs<m?
swkmansMeNIKuyTInRkas;xag
M sma:tticEtkb;eRCA. swkman TMhMFM RTnugsti nigragEbkKW ragpwt
ragml ragRska.
2>@ srIragbnBUC
-etIsrIragbnBUCrbs;suImNUEs<mmanlkNdUcemc?
srIragbnBUCeQal manlkNCaRskamYyEdlpkmIRkUs,:rg
: ; microsporangium mannaTIplitRKab;
lMGg rWmIRkUs,: microspores . RKab;lMGgmanekasikaBIrKW ekasikabnBUC nigekasikalUtlas;.
srIragbnBUCjI manrUbpE RbRbYlKW
-CYnkalrYjxIkayCa GUvulnnl
emeronTI2
Gg;sEs<m Angiosperm
rukC atiGg;sEs<mmanenARKb;TkI EngelIEpndI. eTaHenAtMbn;RtCak; tMbn;RtUBci enAvalxSac;EdlKanCICati.
-etIrukCatiGg;sEs<mCaGVI?
Gg;sEs<mbrukCatimanpa nigRKab;mansMbkkarBar. Gg;sEs<mmanBIrfak;KWmU:NUkUTIeldUnnigDIkUTIeldUn
1-lkNBiessnGg;sEs<m
1>! srIraglUtlas;
-etIsrIraglUtlas;rbs;rku CatimanGVxI H? cUrerobrab;.
srIraglUtlas;rbs;rukCatiGg;sEs<mman bJs edIm swk.
-bJs mansrsnaMTwk nigGMbilxniCeTAedImnigswk. bJscgPab;rukC atieTAnwgdI ehIykmannaTIsk
GaharpgEdr.
-edIm mannaTIRTRTg;EpkEdlenAdIrbs;rukCati nigdwknaMsarFatu
rvagbJsnigskw . edImxHmananTI eFVIrsIsMeyaKbegItsarFatusrIrag
sRmab;rku C ati. CalikasMxan;rbs;rukC atiKW)ac;srsnaM KWsuIELm nigbEGm
suIELmmannaTIdwknaMTwknigGMbilxniCBIbJseTAedIm swk
bE GmmannaTIdwknaMGaharBIswkeTAEpkepSgnrukCati
-swk mannaTIeFVIrssI MeyaK edIm,IbegItGahar. enAkg swk
manRsTab;ekasikaRbePTxusKaKW
RsTab;):alIsat CaRsTab;ekasikaEdlteRmobKay:agENn
enAeRkamRsTab;ekasikaeGBIEDmelI. ekasika):alIsat
mankrU): aspkkrU:PIl.
RsTab;ekasikaes<atnigrnxl; sti enAcenaHekasika):alIsat nigekasikaeGBIEDmeRkam. kg
RsTab;enHmansrssIuELmnigbEGm.
1>@ srIragbnBUC
-etIsrIragbnBUCrbs;rukCatiGg;sEs<mCaGVI?
srIragePTrbs;rukCatiGg;sEs<mKW pa. papeLIgeday
Rtbk RsTab; kBaeM ksrjI kBaMeksreQal.
1># dMeNIrlMGg
-GVICadMeNIrlMGg? etIdMeNIrlMGgmanbun: an?GVIxH?cUrerobrab;.
dMeNIrlMGg CaeprRKab;lMGgBIe)aklMGgeTAelIsicma:t.
dMeNIrlMGgmanBIry:agKW sVydMeNIrlMGg nig dMeNIrlMGgkat;.
sVydMeNIrlMGg bdMeNIrlMGgg CaeprRKab;lMGgBIe)ak
lMGgeTAelIsicma:tnpaEtmYy.
dMeNIrlMGgkat; CadMeNIrRKab;lMGgFak;eTAelIsci m:atrbs;
panrukCatimYyepSgeTot. dMeNIrlMGgkat;RbRBwte LIgedaysar Twk
xl; stV mnusS.
2-karbnBUCrbs;Gg;sEs<m
2>! kMNRKab;lMGg
-etIRKab;lGM gmanlkNdUcemc?
kge)aklMGgmans,:GabGIutEdlekIteLIgtamcMENkemyUs
: . s,:nmI YyEckxn mYydgtammItUs rYclUt
las;CaRKab;lMGg. RKab;lMGgnImYymanNVyU:GabGIutBIrKW NVyU:lUtlas; nigNVy:UbnBCU .
3 eRbobeFobrukCatim:UNUkUTIeldUn nigDIkUTIeldUn
rukC atimU:NUkUTIeldUn
DIkTU IeldUn
-mankUTIeldugmYy
-mankUTIeldugBIr
-pamanRsTab;bI
-pamanRsTab; 4 b5
-)ac;srsnaMsitenAray)a:y
-)ac;srsnaM sitenACargVg;
-manRTnugswkRsb
-manRTnugswkEbkExg
-edImeTal
-edImEbkEmk
-bsEsJ
-bsEkv
4 plRbeyaCn_rbs;rukCatimanRKab;
rukC atimanplRbeyaCn_Nas;cMeBaHCIvPaBrs;enArbs;mnusSeyIg.
-rukCatisuImNUEs<mCaBiessRsl; GacykeTAeFVICaRkdas eRKOgsgarwm nigcMhuyeFVICaeRbgsRmab;
labeQI pH nigBa)alCMgWpgEdr.
-rukCatiGg;sEs<m CaRbPBGahardsMxan; sfsRmab;Ba)alCMgW plitCasMBt; Rkdas nigeFVICa
eRKOgsMNg; eRKOgsgarwmdRbNIt. BiessrukCatipl;GuksuIEsndl;mnusS stV.
CMBUk3
tRmUvepSgrbs;sarBagkay
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emeronTI2
srIragviBaaN Sense organ
RbBnR bsaTeqIytbeTAnwgRBwtkarN_epSgEdlenACMuvji xneyIg.
-etImnusSmansrIragvBi aaNbun: an?
srIragvijaNrbs;mnusSman 5 KW ckviBaaN esatvijaN XanvijaN CIvavijaN kayvijaN.
1-ckviBaaN
-etIEPkCasrIragGV?I cUrerobrab;BKI MehIjrbs;EPk.
EPkCasrIragKMeXIj vapl;BtmanCag80 EdlTTYl)anBIBiPBxageRkA. EPkmannaTIRbmUlpMkaMrsIBnW
EdlenACuMvijxn eyIg.
xYrk,alcab;ykrUbPaBBIEPkxageqVgnigEPkxagsaM ehIysMeyaKbBalKaeTACarUbPaBEtmYy Edlpl;
nUvKMeXIjnvimaRtbIEbbsRmab;eFVkI arvinicyBIRbEvg TMhM nrUbPaB.
1>! TRmg;RKab;EPk
-etIRKab;EPkrbs;mnusSmanTRmg;dUcemc?
RKab;EPkmanragEsVr manGgt;pit 2>5 s>m EdlsitenAkgRbLg;EPknlladk,al. RKab;EPkrbs;
mnusSmanPasbIRsTab;KW ekr:UTic kUrUG: Iut er:Tin nigeRsabedaymCdanfa.
k> PasRKab;EPk
-cUrerobrab;BKI MehIjrbs;EPk.
KMehIj s exA nigBNekIteLIgedaysarmanCatiBNnigvtmanvItamIn Ga. enAeBlBnWb:Hnwgekasik aekan
bekasikadMbg vabMEbksm<nKImIrvagCatiBNnRsTab;erTInnigRbUetGIun begIt[manGaMgPcRbsaT dwknaMeTA
kan;xYrk,al tamsrsRbsaTGubTic ehIyxYrk,aleFVIkarbkRsayCaKMehIj.
srsRbsaTGubTicekItBIsrsRbsaTecjBIekasikaekan bekasikadMbg rYmKa.
mnusSxVak;man; bNalmkBIxVHvItamInGa EdlminGacemIleXIjkgBnWexSayeT.
1>$ KMehIj
-etIEpkNanRKab;EPkEdltRmUvbrimaNBnWcUleTAkgEPk?
karcab;ykBn)w anGaRsyedayekasikarYsnwgBnWenAelIerTIn EdlGacR)ab;eTAxYrk,al RtwmBnN
W aEdl
cUleTAkgEPkbu:eNaH. dMbUgBnWnImYyR)ab;eTAxYrk,alRtwmEtbMENktUcnrUbPaBEdlvacab;)an. bnab;
mkxYr k,aldak;bMENktUcTaMgenaHbBalCamYyKa[eTACarUbPaBmYyeBjelj. GgsulI IEy nigsrs
cMNgCYyEkvEPk edIm,IbegItrUbPaBc,as;l.
k>naTIrbs;EPk
-mannaTIbrfamBlBnWeTACafamBlGKIsnItUc. karbrfamBlRtUveFVIeLIgenAelIerTIn eRBaHTIenaHman
ekasikaRbsaT120lan.
K>Rtecokkg
-Rtecokkgman bMBg;ragKUTxg nigbMBg;ragBak;kNalrgVg;.
bMBg;ragKUTxg CasrIragl,W eBjedaysarFaturav ehIymanRsTab;ekasikaFlEdlmaneramlit enAelI
PasxagkgnbMBg;. ekasikaFlrYsnwglMjr.
bMBg;Bak;kNalrgVg; eBjedaysarFaturavnigBasedayekasikaFl manBnyqar EdlCYyeGaysarBag
kayrkSalMnwg)an.
2>@ l,W
-etIl,WekIteLIgenAeBlNa?
eBlrlksUreFVd
I MeNIreTAkan;Rtecokkg ekasikaFl brrMjreTACaGaMgPcRbsaT. bnab;mkGaMgPc RbsaT
dwknaMeTAkan;xYrk,al tamsrsRbsaTesatvijaN.
2>3 tMEhrkSalMnwg
tMEhrkSalMnwg CanaTIrbs;RtecokkgnigxYrtUc. enAkg Rtecokkg manbMBg;Bak;kNakrgVg;cMnYnbIEdleBj
edaysarFaturavnigmanRsTab;ekasikaeramjr.
-eBlbgVilxn sarFatueramjrmanclna. clnaemkanicbMEbgeTACaGaMgPc RbsaT rYceFVIdeM NIreTAkan;xYr
tUc tamsrsRbsaTesatvijaN. xYrtUceFVIkarbkRsayTisedA ehIybBanGaMgPceTAxYrFM rYcePJaceTA
sac;duEM pkk,alnigk[rkSaTItaMgk,al.
3 CIvavijaN
-etIGVIEdleFV[I eyIgdwgBIrsCati?
edaysarGNateFVI[eyIgdwgBIrsCati.
-etIenAelIpGNatmanlkNdUcemc?
enAelIpGNatmanRKab;tc
U lit KWCaBnkCIvavijaN. BnkCIvavijaN
CaFlvijaN EdlrYsnwgrsCati. ekasikaTaMgenHsitenAkgGBaajelI
pGNat. GNatrYsnwgrsCatii manmUldanbYnKW CUr lVgI Rb nigEpm.
4 XanviBaaN
-cUrerobrab;BlI kNrbs;XanvijaN?
RcmuHCasrIragmYymannaTIsMxan;kgdMNkdegm
I nigkin. kgRbehagRcmuH manRsTab;esImEdlekItBI
ekasikabIRsTab; Kw RsTab;ekasikaRTRTg; RsTab;ekasikamUldan nigRsTab;ekasikaXanvijaN.
-PasesImEdlBasrnRcmuHsMbUredaysrsQam manBNkUlab. PasesImenHmanRkeBjrMGilCaeRcIn
EdlCYyrkSaPaBesImCanic. rMGli enHeFVI[xl;ekA esIm nigGacsMlab;emeraKEdleRCotcUltamxl;degIm
-PasesImelOgmannaTIEpkvijaN. PasenHminsUvsMbUrsrsQamnigRkeBjeT EtmanEpkcugbMputn
srsRbsaTXanvijaN.
nRsTab;eGBIEDmplitemLanInEdlpl;BN[Es,k.
EDm CaRsTab;EdlekItBICalikasnansrsyWtnigkULaEsn collagen
sm,rsrsQamnigcugPIbRbsaT. RsTab;enHptp; g ;sarFatuciBawmdl;ekasika nigtRmUvsItuNP aBkg Es,k.
5># srIragbnab;bnSMrbs;Es,k
-etIEs,kmansrIragbnab;bnSMGVIxH?
Melanocyte
emeronTI3
RbRBnGg;dURKInEndocrin system
-dUcemcehAfa efrlMnwg?
efrlMnwg KWtMEhrkSamCdansarBagkay[mantulPaBCamYymCdanxageRkA.
-enAkg sarBagkaymanRbBnCaeRcIn EtmanRbBn2EdlsMxan;CageK etICaRbBnGIVxH ?
KWRbBnR bsaT nigRbBnGg;dRU KIn.
I>RkeBj nigGrm:n
U
1>! RkeBj
-etIGVICaRkeBj? vamanbu:nanRbePT?GVIxH?
RkeBjCasrIragEdlekIteLIgBIekasikaeGBIetlm.
vamankeTskmkgkarbeBajsarFatu sMrab;sarBagkay. RkeBjman 2y:agKW
RkeBjGicsURKIn nigRkeBjGg;dRU KIn.
RkeBjGicsURKInbRkeBjbeBajeRkACaRkeBjmanbMBg;naM mannaTIbeBajrstam
bMBg;naM. RkeBjsURKInman RkeBjejIs RkeBjTwkmat; RkeBjrsrMlayGahar
RkeBjePT lMEBg.
RkeBjGg;dRU KInbRkeBjbeBajkg CaRkeBjEdlKanbMBg;naM mannaTIbeBajGrmU:n
eTAkgcrnQamedaypal;.
1>@ Grm:nU hormone
-dUcemcehAfaGrm:Un? GVCI aekasikaeKaledA?
Grm:UnCasarFatuKImIEdlman\TiBleTAelIemtabIlIsnekasikaeKaledA.
enHGaRsyedaykarERbRbYlnGRtaRbtikmKImICIvNamYyenAkgekasikaeKaledA.
1># clnkarnGMeBIrbs;Grm:Un
-kgRbBnG g;dURKIn etIeKEckGrm:nU Cabun: anRkum?
eKEckGrm:n
U Ca2 RkumKW
-Grm:Unbubi TIt CaRbUetGInu FMEdlekItBIRcv:ak;nGasuItGamIen. vaminrlaykglIBIt
dUcenHvaminGacqgkat;Pasekasika)aneT.
-Grm:UnesrGU: Iut CalIBItEdlRtUv)anpliteLIgBIkUeLesru:l. vamanlTPaBqgkat;
Pasekasika .
k>Grm:UnbuibTIt
-etIGrm:Unbibu TItman\TBi lelIdMeNIrRbRBwtrbs;ekasikaeKaledAdUcemc?
Grm:UnbuibTItrYmmanGrm:Unrbs;RkeBjGuIbP
U: Is ):ar:aTIr:GU Iut GaMgsyu lIn nigKykakug
vaRtUv)ansMeyaKeRkamkardwknaMrbs;EsnkgekasikaRkeBjGg;dURKIn.
Gem:nU buibTItman\TiBlelIdMeNIrRbRBwtrbs;ekasikaeKaledA edayeFVI[Gg;suIm
GaednIlsuIkas EdlPab;elIpxagkgnPasekasikaeKaledAskmeLIg.
x>Grm:nU esrU:GtIu hormone strod
-etIGrm:Unman\TBi lelIdMeNIrRbRBwtrbs;ekasikaeKaledAdUcemc?
KykUscUlkg crnQam
ekasikaebtanlMEBgbeBajGaMgsuylIn
GaMgsyu lInePJacekasika[cab;ykKy kUs
daRkam tMNbRtLb;GviCman
bgak;karsMeyaKGaMgsyu lIn
briePaKGaharsm,rKykUs
>RbBnGg;dURKInmnusS
-cMeBaHmnusS etIRbBnGg;dRU KInekIteLIgBIGVI?cUr[]TarhN_bBaak;.
cMeBaHmnusS RbBnG
g;dRU KInekIteLIgBIRkeBjGg;dURKInepSgCaeRcIn dUcCa
III
RkeBjePTjI RkeBjeQal>>>.
3>! GIub:tU aLamus Hypothalamus
-etIGIbu U:taLamusekItBIGVI? vamannaTIGVI?
GIub:UtaLamusCaEpkmYynxYrk,al EdltRmUvmCdankgsarBagkay.
GIub:UtaLamus mannaTI RtYtBinitkarbeBajrbs;RkeBjGIbu :UPsI .
-etIGIub:UtaLamusplitGrm:Unbu:nany:ag?GVxI H?
GIub:UtaLamusplitGrm:Un2y:agKW ADH nig GuksIutUsuIn.
-Gg;TIDIGuyerTic ADH= Antidiuretique Hormone mannaTIePJactRmgenam [man
sRmbTwkeLIgvij.
-GuksIutUsIun Ocytocine mannaTIePJacs,n[kRnak; kgryeBlsRmalkUn.
3>@ RkeBjGIubUP: Is Hypophyse
-etIGIub:UPsI CaGVI? GIub:PU IsEckCabun: anEpk?GVxI H?
GIb
u U:PsI CaRkeBjtUcmYy manGgt;RbEhl 1s>m EdlsitenAxageRkam GIub:Uta
Lamus. GIub:UPsI EckCa 2EpkKW GIubP:U Ismux nig GIbu :PU IseRkay.
k> GIubP:U IseRkay
-etIGIub:UPsI eRkaymannaTIGVI?
GIubP
:U IseRkaymannaTI beBajGrm:Un Gg;TIDGI uyerTic nigGrm:UnGuksuItsU nIu .
x> GIubP:U Ismux
-etIGIub:UPsI muxplitGrm:nU )anbu:nanRbePT?GVIxH? GrmU:nnImYymanmuxgardUcemc?
GIubP
:U Ismux plitGrmU:n 6 RbePTxusKaKW
-GrmU:nlUtlas; GH= Growth Hormone mannaTICMrju karlUtlas;nsarBagkay.
-GrmU:nRbULak;TIn PRL= Prolactin Hormone mannaTICMrju karlUtlas;RkeBjTwkedaH
nigplitTwkedaH.
x>BgsVas Testes
-etIBgsVasmanmuxgardUcemc?
BgsVas beBajGrmU:nGg;RdUEsn. Gg;RdUEsnmannaTI sMxan;KWplitGrm:Un
etsesrU:n Testosterone. etse srU:n rYmCamYy LH nig FSH ePJackarplit
Es<m:atUsUGIut. Grm:UnetsesrU:neFVI[kan;EtRbesIreLIgnUv karlUtlas;lkNePT
bnab;rbs;mnusSRbus dUcCasMeLgRKlr duHBukmat;>>>.
3>* RkeBjTImus Thymus
-GVICaRkeBjTImus?etIRkeBjTImsu manlkNdUcemc?
CaRkeBjedlsitenAEpkxagelInRTUg Ek,rebHdUg. vamanTMhMFMkgvykumar
ehIyrYjtUceRkayeBlRKb;kar. RkeBjTImusplitGrmU:nTImUs: unI . GrmU:nTImU:suIn
ePJac plitLaMpUsuItfIenAkgRkeBjTwkrgnigkgsrIragdTeTot.
3>( RkBH nigeBaHevontUc
-etIRbdab;rlM ayGaharmanplitGrmUn: EdrbeT?
Rbdab;rMlayGaharkmanplitGrm:UnEdrKw
ekasikaPasRkBHbeBajGrmU:nkaRsIn Gastrine EdlePJac[manrMhUrnrsRkBH
ekasikaPaseBaHevontUc plitGrmU:neseRkTIn SecretineEdlePJac[manrMhUrn
rslMEBg.
CMBUkTI4 naTIrbs;RbUetGIunksarBagkay
emeronTI 1
GasuIGamIen Amino Acid
-etIRbUetGIunCaGVI? ehIyvamannaTIbu:nany:agenAkgsarBagkay?
RbUetGIUnCasmasFatuKImInCIvitRKb;Pavrs;. RbUetGIunmannaTIksarBagkay 6
y:agKwCa -RbUetGIuTRmg; vabegItCaTRmgekasika
-Gg;suIm CakatalIkrKImICIvEdlCmrujel,nRbtikmKImIkgekasika.
-Gg;TIkrmannaTIkarBarsarBagkayTb;ngw Pak;garbgeraK.
-GrmU:n r:ab;rgkgbNrrUbFatu.
-eGm:Ukbn manaTIdwknaMGuksuEI snkg degIm.
-GkeFVIclna RbUetGInu cUlrYmRKb;clnarbs;ekasika.
1-TRmg;GasItu GamIeN
-etIskmPaBTaMgGs;rbs;sarBagkayrs;RtUvRbUetGIunEdrbeT?
RKb;skmPaBTaMgGs;rbs;sarBagkayRtUvkarRbUetGIunCacaM)ac; eRBaHvaCasmas
FatuRKwHnCIvit. RbUetGuInCasarFatusrIragsaMjaEM dlpk kabUnC GIuRdUEsnH
GuksuEI snO nigGasUtN CYnkalmanpUsVatP nigs<an;FrS pgEdr.RKb;
mU:elKulRbUetGIunrtUv)anbegIteLIgBIsarFatugayKw GasuItGamIen.
GasuItGamIennImYyekIteLIgBIbNMkabuksulI -COOHbNMGamIn-NH nigr:aDI
kal;R. rUbmnTUeTArbs;GasuItGamIenKW
2
xageRkamCarUbmnrbs;GasIutGamIenTaMg 20RbePT.
3-bibu TIt
-ehtuG)VI anCaeKfaGasuItGamIenCasarFatucMNUlTwk?
emeronTI 2
RbUetGIn Protein
-etIGaharRbePTNaEdleFVI[sarBagkaylUtlas;?etIGaharNaxHsm,rRbUetGIun?
>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
-etIRbUetGIunbgeLIgBIGVI?etIGasutI GamIennImYyxusKaedaysarGVI?
>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
1-rUbprbs;RbUetGInu
RbUetGIunCam:aRkUmU:elKulsaMja M. vaCabU:lEI mnGasIutGamIenyfaRbePTxusKa.
RbUetGIunEdlmanGasIutGamIenticCag 50 ehAfabuibTIt. mannyfa RbUetGIunman
Rcv:ak;bubi TItmYybeRcIn ehIyekItBIGasuItGamIencab;BI50eLIgeTA.
-etIeKEckRbUetGInu Cabun: anTRmg;? etITRmg;niImYymanlkNdUcemc?
edayEpkelITRmg; nigsmasFaturbs;RbUetGIn
u eKEckRbUetGIunCa 4 TRmg; KW
1>! TRmg;TI1 ekItBItMNlMdab;GasIutGamIensuT.
1>@ TRmg;TI2 Rcv:ak;bU:lIbbui TItbt;
RbUetGIunmannaTI 6y:agKW
!> katalIkrbegnI el,nRbtikmKImI. ]TarhN_rIbuyLbp UsVat CasmasFatukg
dMeNIrrsIsMeyaK. nIRtUEsNas; CaRbUetGIunmannaTIPab;GasUt.
@>RbUetGInu TRmg; mannaTICaGkkarBarnigRTRTg;. ]TarhN_ kULaEsnsrs
RbUetGIunkgCalikasnanCasmasFatucMbgkgPab;Calika[rwgmaM.
eGLasnI manenAkg srsywtkgCalikansarBagkay dUcCa srsQam
Es,k eFV[I CalikakgsarBagkaymanPaByWt.
#>GkeFVcI lnacUlrYmRKb;clnarbs;ekasika. ]TarhN_Gak;TInmIRkUPILam:g;
TuybuylInkUnbMBg;tUc>>>. sMNuMRbUetGIunTaMgenHenAkg eRKagqgw ekasika man
skmPaBkgkareFIVcMEnkekasika.
$>GkkarBarRbqaMgnwgemeraKkuM[eRCotcUlkg sarBagkay)an.
]TarhN_Es,k karBarkareRCotcUlnemeraKeBlrbYs. PIRbNUEsnnigRtubM n Ca
RbUetGIuneFV[I Qamkk eBldac;srsQam. Gg;TIkr RbqaMgnwgkareRCotcUlrbs;
emeraK.
%>GktRmUvGrmUn: plitecjBIRkeBjGg;dURKIn.
]TarhN_GaMsuylIn nigKy kakug tRmUvkRmitKykUskgQam. rIGrm:UnlUtlas;
ePJacekasika[eFVcI MENk.
^>GkdwknaM Na , K dwknaMmeU: lKulGIuyu:gsarFatuepSgqgkat;ekasika.
eGmU:kb n dwknaMGuksuEI snBIsYteTAkan;ekasika.
-etIeKEbgEckRbUetGIunCabu:nanRkumbEnmeTot?GVxI H?
tamTRmg;rbs;RbUetGIuneKEbgEckvaCa2RkumKW
RbUetGIunsrsCamUe: lKulEvgminrlaykgTwk ehIysVit. vamanenAkgEs,k sk;
Rkck mannaTICaGkkarBar.
RbUetGIunRKab;CamU:lKulragmUltUcrlaykgTwk. vamannaTICaclkr Gg;TIkr
eGmU:kbn Gal;buymIn CaGkdwknaMGasIutxaj;kg Qam.
-etIeKEcksmasFatuRbUetGIunCabu:nany:ag?GVIxH?
eKEckRbUetGIunCa2y:agKW
RbUetGIungayekItBIGasIutGamIensuT.]TarhN_esrUm: Gal;buymIn ekr:aTIn
RbUetGIUnsaMjaMekItBIGasIutGamIen nigsarFatuminEmnCaRbUetGIunRkumRbUesric.
-etIRbUetGIunmankar)at;bg;KuNPaBeRkamlkxNGVIxH?
RbUetGIunmankar)at;bg;KuNPaBeRkamlkxNdUcteTA
GasuItb)asxaMgbNal[sm<nGIuRdUEsnRtUv)anbgak;
Pak;garepSgbgak;sm<nGIuRdUEsnnigkareRCotcUlnTwk
GgFaturMlaybgak;karRCabcUlrbs;Tkw
sabUbgak;PaBbt;Ebnrbs;RbUetGIun nig[rlaRcv:ak;buibTIt
kMhab;GMbilbegnI smasFaturlaykg Twk
elahFatuFn; )artHgman\TiBlelITRmg;RbUetGIun nignaTIrbs;va.
emeronTI 3
Gg;suIm Enzyme
1-GVIeTAGg;smuI ?
Gg;sum
I Ca sarFatuRbUetGIunEdlmannaTICakatalIkrCIv CYyCMrujel,nRbtikm
KImIepSg.
-RbsinebIKanGg;suImeT etIsarBagkayGacRbRBwteTA)anbeT?
RbsinebIKanGg;sm
uI eT skmPaBepSgkgsarBagkayminGacRbRBwte TA)aneT
-etIGg;suImmanlkNsm,tidUcemc?
KWCakatalIkrKImICIvEdlmanRbtikm
yfaRbePTkRmitx<s; ehIyskmPaBn
RbtikmKImCIvKWeTogTat; nigCak;lak;.
]TahrN_
kMedAxagM
bU:tasmkr:at m:g;kaENsGuksutI
KClO2
KCl + O2
0C
0C
0C
0C
CBUkTI5
BtesenTicnigkarsMEdgnEsn
emeronTI1
ADNCaBtmanesenTic
ehtuGVI)anCaeKfa ADNCaTMrBt_manesenTic?
-eRBaHpktMNlMdab;nuyekGUTIt KWnuyekGUTItenHehIyCaGkpkBt_manesenTic Ca
bg;ksaglkNRbUTIt.
kgkarbnBCU edayePTktfIekItBIsIukUt)anmkBIkarrlaycUlKarvag kaEm:tjInig
kaEm:teQal. suIkUtmanBt_manesenTicsMrablkarksaglkN rbs;Pars;.
1-smasFatuKImInsMParesenTic
1>1 kaeBiesaFrbs;elakRKIPIt Griffith 1928
-etIPmI UP: UkCa)ak;etrIbgCMgWGVI?- Ca)ak;etrIEdlbgCMgW[manCMgWrlaksYt. vaman
rUbrag2 Ebb KW rag S nig rag R .
-etIrbU rag SmanlkNdUcemc?- KWrbU ragmaneRsamEdleFVIGMBIKysIut. vaman
lkNrhskg karbg[manCMgW.
-etIrbU rag RmanlkNdUcemc?- KWragKRKatKaneRsam nigminbg[manCMgWeT.
tamkarRsavRCavrbs;elakRKIPtI )ansnidanfa )ak;etrI R KaneRsam EbgCa
)ak;etrI S maneRsamEdlGacbgCMgW nigbBaneTAsnaneRkay.
1>2 karBiesaFrbs;elakGavIrI
-karRsavRCavrbs;elak GavIrI nigGkviTasaRsepSgeTot bgajfa mU:elKul
ADN CaTMrBt_manesenTic.
H3PO4
H3PO4
H3PO4
H3PO4
sredGuksuIrIbUs A
sredGuksuIrIbUs T
sredGuksuIrIbUs C
sredGuksuIrIbUs G
2>2 lkN)asGasUt
tamratagbrimaN)asGasUtn ADN kg ekasikaepSgbgajfa
RbePTPavrs;
)asBYvic
A
sVa
RdUsPU lI
pSitNWrsU: ,:r:a
)asBIrmI IDIn
T
dUcenH
brimaNTImIn nig GaednInesIKa
brimaNsuItsU unI nig kanInesIKa
)ak;etrIGWklU I
mnusS efmI
KnHw edaHRsaylMhat;
-rkRbEvgrbs;Esn ADN bRbEvgrbs;mU:elKul ADN
M
l
0.34nm eday l CaRbEvgEsn ADN bRbEvgmU:elKul ADN
2
M
M
C
2
M
C G
A
2
L 2 A 3C 2 A 3G 2T 3G 2T 3C
AT
M= 2A+2C = 2T+2G=2A+2G=2T+2C
cMnYnnuyekGUTItTaMgGs;enAelIRcv:ak;nuyekGUTItTaMg2
-rkcMnYnCMhann mU:elKul ADN bEsn
eday 3.4 nmRtUvnwg!CMhan
cMnYnCMhan= 3.l4
M
eday!0KUnuyekGUTIt RtUvnwg!CMhan cMnYnCMhan= 20
-rkcMnYnnuyekGUTItTaMgGs;rbs;mU:elKul ADN bEsnM
tameKalkarN_bMeBj)asnIRTic A-T , C-G
-edIm,IrkcMnYnnuyekGUTItesrI
ebImU:elKul ADN eFVIsVytMelIgu eTV 1dg M = M
ebImU:elKul ADN eFVIsVytMelIgu eTV ndg M = M 2n-1
ecjBIrbU mnxagelIenH eKGacrk
>TesrI = AesrI = A 2n-1 rW TesrI = AesrI = T 2n-1
>CesrI = GesrI = C 2n-1 rW CesrI = GesrI = G 2n-1
-edIm,IrkcMnYnnuyekGITItesrIsrubtageday m
ebIeKR)ab;cMnYnA nig A m A A 100
ADN
ADN
ADN
ADN
ARNm
ebI m nigA
ebIeKR)ab;cMnYnU
ADNm
ARNm
ARNm
nig U
ARNm
ARNm
m 0 0 A ARNm
100
100
U
m ARNm
0 U
0
ARNm
cMnYnnuyekGUTIt A ARNm
ARNm
ebI m nigU
ebIeKR)ab;cMnYnC
ebI m nigC
ebIeKR)ab;cMnYnG
m 0 0 U ARNm
100
100
C
CARNm m ARNm
0 C
0
ARNm
m 0 0 C ARNm
C ARNm
100
100
G
GARNm m ARNm
0 G
0
ARNm
m 0 0 G ARNm
G ARNm
100
cMnYnnuyekGUTIt U ARNm
ADNm
nig
ARNm
cMnYnnuyekGUTIt
ADNm
nig
ARNm
ARNm
ARNm
ARNm
ARNm
emeronTI2
karsMEdgecjnEsn
-etIGVICa ADN? CasmaresenTicEdlpkBt_manesenTic.
-etIlkNrbs;Pavrs;epNUTIbRtUv)ankMNt;edayGV?I ehIysitenANa?
RtUv)ankMNt;edayBtmanesenTic EdlsitenAkgNVyU:nekasika.
-etIBtm anesenTicRtUv)anbBan BIekasikaemeTAekasikakUntamryGVI?
tamrysVydMeLIgeTV ADN nigr)ayesIn ADN kgvKGaNapasnmItUs.
!> skmPaBnEsn
!>1>BiesaFn_rbs;bd l Beadle nigtaTum Tatum
- etIelakbdlnigTumeFVIkarBiesaFn_dUcemc?
BYkKat;bBaagM kaMrsI X eTAelIs,:rbs;pSitprRkhm NWrU:s,:r:aRkassa ehIy
yks,:enaHeTAciBawmenAkgemtabUlItBIrxusKa KW
mCdansm,rsarFatuciBawm
mCdanxSt;sarFatuciBawm edaymCdanmYylayemtabUlIt C nig
mCdanmYyeTotlayemtabulIt D . tamkarBiesaFn_bgajfa manGg;suIm2
RbePTRtUv)ansMeyaKeday
mIeslm edIm,IRTRTg;karlUt
las;enAkg mCdan C nig D.
Kat;ehAsmtikmenHfa EsnmYy
dwknaMsMeyaK Gg;suImmYy.
emtabUlIt CasarFatuEdlcUlrYmemtabUlsI .
-etIpSitprRkhmeQaHNWr:Us,:r:aRkassa manlkNBiessdUcemc?
KWmanlkNGaclUtlas;kugmCdanxSt;sarFatuciBawm ehIybnBCU tamrys,:
GabGuIt.
-GVICaEsn? EsnCaGgt;tUcmYyrbs; ADN EdlmanpkBtm anesenTic sRmab;
sMeyaK RbUetGIunyfaRbePTmYy ehIyRbetGInu CaGkkMNt;lkNrbs;kt.
-etIGksnidandUcemcrvagEsn nigRbUetGIun?
EsnCaGgt;mYyrbs; ADN Edlpk BtmanesenTic sRmab;kMNt;karsMeyaK
RbUet GIunmYy. RbUetGIunenHeFVI[elcecjnUglkNmYyrbepNUTIbmYy.
!>2>PaBRtUvKan ADN nigRbUetGInu
-etIRTwsIxagelIEdlfa : EsnmYydwknaMsMeyaKGg;suImmYy : RtUvbr CaGVI?
{ EsnmYydwknaMsMeyaKRbUetGIunmYy :
-cUreRbobeFob ADN nigRbUetGIun
cMncu dUcKa -Cam:aRkUmUe: lKul
-mU:elKulnImYyEdlekItBIktatUcPab;KaCaRcv:ak;mU:elKul.
-mU:elKulnImYymantMNlMdab;nuyekGUTIt bGasIGu amIenCak;lak;.
cMnucxusKa
ADN
RbUetGIun
>ekItBIRcv:ak;nuyeklGUTItBIrbMeBjKa >ekIteLIgBIRcv:ak;GasutI GamIenmYyExS
>nuyekGUTIt#kMNt;GasuItGamIen!.
>GasutI GamIen!kMNt;edaynuyekGUTIt#
>manRbEvgEvgCagRbuetGIunrab;sibeTArab;ry >manRbEvgxCI ag AND rab;sib
> ADN nImYymannuyekGUTtI 4RbePT Edl eTArab;rydg.
teRmobCatMNlMdab;nuyekGUTItCak;lak;. >manGasuItGamIen 20RbePTEdl
teRmobKatamtMNlMdab;Cak;lak;
)anCaRcv:ak;bUlbI uibTIt.
@> karcmgBtmanesenTic
-etIBtm anesenTicEdlCabg;sagsg;RbUetGInu manTItaMgenANa?
KWenAelIGgt; AND kgNVy:U eRkamTRmg;CatMNlMdab;nuyekGUTIt.
-etIeragCagsMNg;RbUetGIunCaGV?I ehIymanTItaMgenANa?
KWCarIbUsUm ehIysitenAkgsuIt)U as.
@>1 rUbp ARN naMsar ARN
-enAkgsuItU)as etImU:elKulGVIEdldwknaM
sMeyaKRbUetGIunedaypal; ? KW ARN
-GVICa ARN ?
Cam:eU lKulEdlcmgBtmanesenTic
BI ADN ehIybBaneTArIbUsUm edIm,I
sMeyaKRbUetGIun.
m
10
10
kgRbetGInu .
-etInuyekGUTIt4RbePTrbs;ADN kMNt;GasuItGamIenTaMg20RbePTrbs;RbetGInu
)an y:agdUcemc?
KWmansmtikmbI
nuyekGUTIt1kMNt;GasuItGamIen1$!=$ enaHsl;GasuItGamIen16 vaminRKb;.
nuyekGUTIt2kMNt;GasuItGamIen1$@=!^ enaHsl;GasuItGamIen4 vaminRKb;.
nuyekGUTIt3kMNt;GasuItGamIen1 $#=^$RKb;RKan;sRmab;kMNt;GasuItGamIen.
naM[eKGacsnidan)anfa nuyekGUTIt # mYyRtIFatukMNt;GasIutGamIenmYy .
GasuIt GamIen mYyTTYlRkmBIRtIFatumYyrWeRcInBI ADN.
#>2 taragRkmesenTic
nuyekGUTIt3=RtIFatu1=1kUdug=1Gg;TIkUdug=1GasuItGamIen
edayARN CaGksMeyaKRbUetGIunedaypal; dUcenHeKGac[niymny Rkmesen
Tic CaRbBnRtUvKantMNlMdab;rIbUnuyekGUTItrbs; ARN nigtMNlMdab;GasuItGa
mIen.
naM[ $#=^$ kUdug-# kUdugsb =^! kUdug RtUvnwgGasuItGamIen @0RbePT.
-tamtaragRkmesenTic etIkUdugNaminkMNt;GasuItGamIen?
mankUdug UAA UAG UGA minkMNt;GasuItGamIen eRBaHvaCakUdugsRmab;
bBab;karsMeyaKRbUetGIun EdlehAfakUdugsb.
-etIRkmesenTicmanlkNdUcemc?
manlkNCaskl eRBaHPavrs;TaMgGs;mannuyekGUTIt 4RbePT sRmab;kMNt;
GasuItGamIen 20 RbePT ehIyRtIFatumYykMNt;GasuItGamIenmYy.
$> karbkERbRkm
$>1 rIbUsUm
-etIFatusMxan;GVxI HEdlcUlrYmkgkarsMeyaKbU:lIbiubTIt?
FatusMxan;bIEdlcUlrYmkgkarsMeyaKbU:lIbiubTItKW ARN rIbUsUm ARN .
-etIrbI UsUmCaGVI? ehIymannaTIdUcemc?
CaFatuekasikamYy vasti enAesrIkgsuIt)U as EdlcUlrYmsMeyaKRbUetGIun ehIy
mannaTICaeragCagsagsg;RbUetGIun.
rIbUsUmman2EpkKW ktargFM nigktargtUc
m
$>3 clnkarbkERbRkm
-etIkarsMeyaKRcv:ak;bUl: IbuibTItRbRBwteTAenANanekasika?
KWRbRBwteTAenAkgsuItU)as elIrbI UsUm. karsMEdgecjnEsnman clnkar 2bn
bnab;KW
clnkarcmgRkm CaclnkarTI1 nig
clnkarbkERbRkmkarsMeyaKRcv:k;bUl: Ibubi TIt CaclnkarTI2
-etIclnkarbkERbRkmRbRBwteTAdUcemc?
clnkarbkERbRkmRbRBwteTA bclnkarsMeyaKRbUetGIun man 3 dMNak;KW
dMNak;epmI dMNak;lUtEvg nigdMNak;bBab;.
k>dMNak;dMbUg
-etIkUdugepImelImU:elKul ARN KWGVI? etIkUdugenHRtUvnwgGasuItGamIeneQaHGVI?
kUdgu epmI AUG RtUvnwgGasuItGamIeneQaH emtnIn.
-etIrbI UsUmmankEngBiessbu:nan?
rIbUsUm mankEngBiess 2 KWft P sRmab;emtnInnigsRmab;bU:lbI uibTIt nigft
A sRmab;TTYlGasuItGamIen.
-etIdMNak;dMbUg cab;epImedaymanRBwtikarN_GVI?
m
% tRmUvkarnkarsMEdgepNUTIb
-etIRKb;ekasikakgsarBagEtmYy manEsndUcKarWeT?
RKb;ekasikakg sarBagEtmYymanEsndUcKa
BIeRBaHekasikaTaMgGs;manedImkMeNIt
ecjBIsuIkUtEtmYyEdlEckxnCabnbnab;tammItsU .
-etIEsnTaMgGs;nsarBagkaymYy manskmPaBRKb;eBlrWeT?
EsnTMagGs;KanskmPaBRKb;eBlRBmKaeT
BieRBaHEsnnImYysMeyaKEtRbUetGIun NaEdlcaM)ac;cMeBaHva nigenAeBlNa Edlva
RtUvkarbu:eNaH. dUcenHeKfa { vamantRmUv sMeyaKRbUetGIun : .
-etItRmUvsMeyaKRbUetGIunTTYl\TiB lBIGVI?nigmanEsnGVIxHcUlrYm?
tRmUvsMeyaKRbUetGIunTTYl\TiB lBIsuIt)U as nigmanEsn3RbePT cUlrYmsMeyaK
RbUetGIun
EsnTRmg; CaEsnman Btm anesenTicsRmab;kN
M t;TRmg;RbUetGIun.
EsnRbtibtikar CaEsn EdlmannaTIbBaaelIEsnTRmg;.
EsntRmUvbEsnRtYtBinit CaEsnEdlTTYl\TiBlBIsuIt)U as ehIymannaTIbBaa
elIEsnRbtibtik ar.
KnHw edaHRsaylMhat;
-tamclnkarbkERbRkm
-bkERbRkmtamsmamaRtCa
( A U ) ARNm
2
(C G) ARNm
CEsn GEsn
2
AEsn TEsn
-cMnYnrIbUnuyekGUTItsrub
m A U C G m
M
2
-#nuyekGUTIt=!RtIFatu=!kUdug=
!Gg;kUdug=!GasuItGamIen
M
2 m
3
3
naM[ cMnYnkUdug=
- ARN dwknaM yfa aa RbePT!
KitTaMgemtnIn eBlkMBgu sMeyaK
RbUetGIun cMnYn ARN =cMnYn aa
-GasuItGamIen !manm:as!!0xat
kabUnm:asRbUetGIun=cMnYn aa !!0
-cMnYnEsn= l l ll
-ebIrIbUsUm!qgkat;edayRtLb;mkvij
cMnYnRbUTIt=cMnYn ndgnrIbUsUmqgkat;
t
ADN
ADN
Esn
ARNm
ARNm
-smamaRtPaKryrbs;ARN KW!00
m
A 0 0 U 0 0 C 0 0 G 100 0 0
-kgclnkarsMeyaKRbUetGIun epImeLIg
edaykUdgu epmI ! nigbBab;edaykUdugsb !
cMnYyGasuItGamIen=cMnYnkUdug-@
bcnM YnGasuItGamIen= M6 2 b m3 2
M (aa 2)6
m (aa 2)3
-GasuItgamIen@Pab;KaedaycMNgbuibTIt
mYy cMNgbuibTIt=cMnYn aa -!
-cMnYnEsn = cMnYnRbePTRbUTtI
-ebIrIbUsUm!qgkat;eday\tRtLb;mkvij
cMnYnRbUTIt= cMnYn ARN
m
-krNIdwgryeBlrIbUsUm
l ARNm
trIbUsUm 1qgkat;
-ryeBlsMeyaKRbUTIt=ryeBlEdlPab;GasuItGamIen! cMnYnGasuItGamIen
-rkcMnYnrIbUnuyekGUTtI TaMgGs;rbs; ARN Edltag eday m
m
AARNm
100
0 AARNm
AARNm
U ARNm
100
0 U
0
U ARNm
C ARNm
100
0 C
0
C ARNm
AARNm
m
100
U ARNm
m
100
0
C ARNm
m
100
U A
T T
TEsn = 2 2
1
ARNm
ARNm
CEsn =
GEsn =
C1 0 0 C 2 0 0 C G ARNm
2
2
0 C G
0 G 0 G
0 1
0 2
ARNm
0
2
2
CEsn= GEsn
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karvivtnPavrs;
emeronTI1
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etIRTwsIvivtnPavrs;CaGVI?
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ryeBldyUrGEgVg Pavrs;dbM UgenHvivtCabnbnab; ehIybegItCa)anPavrs;
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-
.
(Ra )
(Pb )
+
- (C14) (C12)
C14/C12 C14/C12
C14/C12 C14
C14/C12 C14
III.
+ ?
-
-
-