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ISBN: 978-960-9407-09-0
SET: 978-960-9407-12-0

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A-D
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1
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2009

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A-D
:
, &
2009
THE RED DATA BOOK

OF RARE AND THREATENED
PLANTS OF GREECE
VOLUME ONE
A-D
Edited by
Dimitrios Phitos, Theophanis Constantinidis & Georgia Kamari
Assisted by Pepy Bareka
HELLENIC CAL SOCIETY
PATRAS 2009
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gpf{lfrzl
gtgs|
toktvo|tqgqm
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kp}spt

q{|kt
% LAIKAUQAL MHLCTNCGQ
fzjilsrs
itopsstokt{ito
t{gqi
% LAIKDINMUARR NSEMESGQ
i{lorfrrs
ggtztsg|gg~kqm
vo|tkqh{{tst
qg
t~{qg |qt~pqto
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% LAIKJUIDAJIR GLAIKCNL
zyplrrl
g{sm
q{|kt
LAIKAKIBEQSAMS GLAIKCNL
frr{syps
v{tnmqg
qg
LAIKKALOQGIAM XAHNNGQ
ripysnzsfzs
tg|t{tig|ggrst|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKOAMDQINO BINKTNAGQ
jophiljiops
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKGENQGIAD TOASQARGQ
pfrssnfphfpf
tg|t{tig|ggrst|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKLAQIAMNT BINKTNAGQ
jophsnnp{s
tgtgs|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKJGENQGHI BINKTNAGQ
pjzplf
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKAQSEKAQI TOASQARGQ
|spif{jphs
qqg|g
i|{gt
jophsnnpfrf
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
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% LAIKNGENQGIN TOASQARGQ
fzzfrsnprl
tgs|ztto{g
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kqj
gjq
% LAIKUAKLAQIKI GLAIKCNL
frrfpsps
tg|t{tig|ggrst|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKAXAMMIS BINKTNAGQ

ffr{l}zl
gtgs|
toktvo|tqgqm
kp}qtspt
q{|kt
% LAIKDALAM MHLCTNCGQ
rsf
tgtgs|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKCSHAMNR BINKTNAGQ
jzlypsnlrjzyl
tgtgs|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKOIMDEK BINKTNAGQ
jsiopysnzsorfrrs
qigqtg|tgs|^kphtgs|
~t{gt{tigvo|tkqh{{tst
qt{ktgskztks|m
kg{ts|m
% LAIKJSHENDNQ FNQATSHGQ
llpzzsjphs
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKDILISQG TOASQARGQ
n{l{s
vqsmgq
gqonqggnsg
% LAIKSHXLAJIR AGQNAQSIRSICNCNL
lysnzsfrfhl
qigqt|t{tig
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gg~kqmkqh{{tst
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gskztpgsssps
k}qm
iqst
% LAIKODILNOTK CCTNIGQ
zjnmjpisnzrl
qigqtg|tgs|^kphtgs|
~t{gt{tigvo|tkqh{{tst
qt{ktgskztks|m
kg{ts|m
% LAIKEEKEHE FNQATSHGQ
h{plyps
ggq~ktsm
smohtg
% LAIKDARKILMI NSEMESGQ
frysnzss{s
gkpqi|s|kg{{kkps
smrtztmmiqt|mps
kpqi|s|iggkps
kpzts|gskztnmss
kqj
nsg
% LAIKQICNRSH ATAGQ
ofrril{zfs
gtgs|
toktvo|tqgqm
kp}qtspt
q{|kt
% LAIKMINAM MHLCTNCGQ
ofrrisnnisqf
tgtgs|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKEINAMMI BINKTNAGQ
fhfy{lrrf
gtgs|
toktvo|tqgqm
kp}qtspt
q{|kt
% LAIKAMMA?J EDTBINKNGXTNCGQ
fzshjpysnzspo{ps
spst
nsg
% LAIKOHAQLEJ NSEMESGQ
fzysnk{lzjnmps
tgvgqg|tispg
mkgvo|sqtsps
gvgqg|ko|
ns|gztjqg|gstnmss
nsg
% LAIKEKJAKO OHAQLTNAGQ

fzyszs
tgtgs|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKAOJAKSRIR BINKTNAGQ
fpljophf
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKJALAQI TOASQARGQ
fpfhfrrf{isnfz{
qigqtomg|tgs|
|gvotikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKUJAQAG BINKATSHGQ
fpfhrrlfz{
vtqgg~kqmns|tqotsto
sqtkqhg{{ts|
smqpmtohto

qit{k}g{ts
% LAIKUARNJAQAG TOASQARGQ
fpfyzrljs|rl
tg|t{tig|ggrst|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKSHJAQALOKIAMIR GLAIKCNL
fpsjphs
ns|jqogiqt|qkosg
sttktikg|s
g|s|tomps
k~st{tigg|sqtsps
qg{|st{g
nsg
% LAIKJAGE FQIAGQ
fpssnjhhrf
qigqtomg|tgs|
|gvotikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKJAQNTRNT BINATSHGQ
fpifrrs
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
gqg
% LAIKJAQXDAR TOASQARGQ
fsrl{l
kqks
qit{
k}g{ts
% LAIKLTRCI NSEMESGQ
s{{rlzzf
qigqtomg|tgs|
|gvotikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKJNJJIMI BINATSHGQ
s{snms
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKJNJLNSNR IMGQ
fzlzrl
ns|skp
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kg{ts|m
% LAIKEJNSAKI BINATSHGQ
snyzsgfrkjsr
tg|t{tig|ggrst|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKKJNTLBKI BINKTNAGQ
snsgszsnfjprf
tgtgs|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKJJNTSRNU BINKTNAGQ

ph{f{zfs
qigqtomg|tgs|
|gvotikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKMJQIGAR BINATSHGQ
'EQLAMX
% LAIKWXRSQAJIRFNSIR VAKDBATTMI FQEIBTQGDE
nypo{lf}fpf
qigqtomg|tgs|
gvo|gqgipi
qm
goqpst
q{|kt
% LAIKJXOQINS RSEGSEIHEQGQ
frffpf
qigqt|t{tig
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gjg~kqmkqh{{tst
|gvo|sqps
gskztpgsssps
k}qm
iqst
% LAIKLOAMISRA CCTNIGQ
npf{ysnzsfpzfys
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKHAJXQI XAHNNGQ
orfrriljs|rl
tg|t{tig|ggrst|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKCNMRSGQ BINKTNAGQ
yfkorrl
tg|t{tig|ggrst|
gt{tig
ns|gztjqg|gstnmss
nsg
% LAIKIBAYNR BINKTNAGQ
yfp{fyl
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKBAQEJA TOASQARGQ
np{lus
!KBEQS ,TDVIGR5MIUEQRISX
)MRSISTSENF3IKUICTKSTQE
4EMMEMBACHEQRSQ
$ &QEIBTQGI"Q
{srsisnf
qot{ps
qpsg
LAIKOAO RTJ NSEMESGQ
fyfjphisnf
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKEUAOAO TOASQARGQ
fpf{jnysnzsnhhjzs
gqg||m
qzt{m
LAIKUAG?OAQAR XAHNNGQ
zzlfpmjryl
ns|jqogiqt|qkosg
sqtkpqi|qkosgqkg{{jg
qzklgksk|t{|t

qmkg{ts|m
% LAIKJGEGGB NSEMESGQQAKKIO AGQNATSHGQ
ffplps
)MRSISTSENF3IKUICTKSTQE
&ACTKSXNF&NQERSAMD%MUIQNMLEMSAK3CIEMCER
!KBEQS ,TDVIGR 5MIUEQRISX
4EMMEMBACHEQRSQ
$ &QEIBTQGI"Q
'EQLAMX
% LAIKDILISQINRRALAQAR VAKDBATTMI FQEIBTQGDE
ffpzs
gsgljto
kg{ts|m
% LAIKRSEKRAL HNSLAIKCNL

fp{ffpf
tgksk|tk~st{tig
gt{tig
gskztnmss
nsg
% LAIKLRAQIJA BINKTNAGQ
nprfz{
tgtgs|
gt{tig
gskztnmss
nsg
% LAIKURLXQMI BINKTNAGQ
fnpysnzs{zfs
olgsto
kg{ts|m
% LAIKMRSAUQ NSEMESGQ
j|fr{lrfff
qigqtomg|tgs|
|gvotikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKDIAMI BINATSHGQ
nzzhr{lorrl
isgg
gqg
kg{ts|m
% LAIKRUBBN IALAGQ
kfrsni{llpl
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKSYAMNXD TOASQARGQ
lrf{sphnp
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKSIMIAJNT TOASQARGQ
ph{ffrfhl
ns|jqogiqt|qkosg
sttktikg|s
g|s|tomps
k~st{tigg|sqtsps
qg{|st
{gnsg
% LAIKSQIGAR FQIAGQSQIGAROAM XAHNNGQ
ph{snfpgpf
{{ms|qsnt{ti|gqkg
g{pqg|{kto
nsg
LAIKQSQIGNT NQMISHNKNGIJIGQ
f{plnisqf
qigqtomg|tgs|
votikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKSRAJIQI BINATSHGQ
gjz{fmfrs
g|kjtst~ps
kg{ts|m
% LAIKSRIUEK AGQNATSHGQ
pyilorrl
qigqtomg|tgs|
votikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKSRIQIOID BINATSHGQ
|ynpior
qigqtomg|tgs|
votikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKRSRIRI BINATSHGQ
usslps
qigqttgs|
tgt{tigvos
gt{tig
gskztgqs
qg
% LAIKDOHISNR TOASQARGQ
usnprfp{lprf
ktikg|iqtst|sttgsps
gs
% LAIKNQA LAICHGQ

frzisn|l
qigqtomg|tgs|
|gvotikpiqg}g
gt{tig
qt{ktgskztks|m
kg{ts|m
% LAIKCHAMKIDT BINATSHGQ
sjiops
tqgtok{topgsssps
psssg
% LAIKYNECHISNT GLAIKCNL
!JMBIDDJ3MK
!RHFTQKNMG#KNRE
$NQE
3HEEKD3..
5+
% LAIK4BKNCJEEK ANKCNL
!GJHLF-HDJQ
2LEQRSEIMRSQ
$ +IQCHHEIL4ECJ
'EQLAMX
% LAIKMBNEHKIMG NQA WDE
!MWBD/DRDP
-AKERIAMA4QNOICAKR
,EUEK
4TM*TGAH4NVEQ.N
*AKAM4TMJT!BDTK2AHLAM
+TCHIMG
3AQAVAJ-AKAXRIA
% LAIKBNSAMIRS LAKERIAMACNL
#MWJD4JPHID
3ACHUEQRSMDIGEMQASFQ5LVEKSFQAGEM325
'ERCHRRSEKKE
,TIREMRSQ
$ "EQKIM
'EQLAMX
% LAIKTKQIJEDNXKE TLVEKSQASDE
&PDSRDP6DPLDP
"NSAMIRCHEQ'AQSEMTMD"NSAMIRCHER-TRETL
"EQKIM $AHKEL
&QEIE5MIUEQRISS"EQKIM
+MIGEM ,TIRE 3SQAE
$ "EQKIM
'EQLAMX
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$IOAQSILEMSNDI3CIEMYE"NSAMICHE
5MIUEQRISDEGKI3STDIDI0AKEQLN
UIA,IMCNKM
0AKEQLN
)SAKX
% LAIKVGQETSEQ BGBLNQG
*JDHLQRDSADP LCPD@Q
7EIDNQMVEG
$ +AQKRQTHE
'EQLAMX
% LAIKJKEIMRSETBEQ BNNJR JABEKBVDE
+@LQCMUL1HBG@PC
!QDENKA%MUIQNMLEMSAK3EQUICER
4HE"QIDKE
3SQNTD
'KNTCERSEQRHIQE',31
5+
% LAIKQKAMRDNVM AQDENKADELNMCNTJ
/@NN!D@R@
"NSAMICAK$EOAQSLEMS
(TMGAQIAM.ASTQAK(IRSNQX-TRETL
0/"NW
"TDAOERS
(TMGAQX
% LAIKOAOOBEA BNSMHLTRHT
/DPQQML*@PHL
"NSAMICAK'AQDEM
#AQK3JNSSRBEQGRGASA!
3% 'SEBNQG
3VEDEM
% LAIKJAQIMOEQRRNM UGQEGINMRE
1@SQ3GMK@Q
"NSAMIRCHEQ'AQSEMTMD"NSAMIRCHER-TRETL
"EQKIM $AHKEL
&QEIE5MIUEQRISS"EQKIM
+MIGEM ,TIRE 3SQAE
$ "EQKIM
'EQLAMX
% LAIKSQATR BGBLNQG
1HQRMU,HBG@DJ
6EGESASINM%CNKNGX.ASTQE#NMREQUASINM
)MRSISTSEFNQ"INCHELIRSQX"INKNGX
5MIUEQRISXNF0NSRDAL
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$ 0NSRDAL
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% LAIKQIRSNV TMI ONSRDALDE

2BGSJDP LCPD@Q
/EMHATREQ3SQ
$ .ET 5KL
'EQLAMX
% LAIKAMDQEARRCHTKEQ GLWMES
2LMFDPSN!PHRR
"NSAMICAK-TRETL
5MIUEQRSIXNF,TMD
6AKKGASAM
3 ,TMD
% LAIK"QISS3MNGEQTO BNSLTRKTRE
2LMFDPSN2TDL
"NSAMICAK-TRETL
5MIUEQRSIXNF,TMD
6AKKGASAM
3 ,TMD
% LAIK3UEM3MNGEQTO BNSLTRKTRE
3SPJ@LC-HBGMJ@Q
-IRRNTQI"NSAMICAK'AQDEM
0/"NW
3S,NTIR-/
53!
% LAIKMICHNKARSTQKAMD LNBNSNQG

vv
vv
Acantholimon aeGaeUm&+-EXEQ
AstraGalUsaGraniotii/QOHEW"NIRR127
Achillea occUlta#NMRSAMSIM+AKONTSY
AstraGalUsdrUpaceUs/QOHEW"NIRR
Adoniscyllenea"NIRR(EKDQ/QOH
AstraGalUslaconicUs)ASQN+IS4AM
AeGilopsspeltoides4ATRCH
UAQliGUstica3AUIGM &INQI
AstraGalUsmaniaticUs+IS4AM3SQID
AsyneUmaGiGanteUm"NIRR "NQML
AethionemacarlsberGii3SQID0AOAM
Athamantaarachnoidea"NIRR/QOH
AethionemaorbicUlatUm"NIRR (AXEJ 51
AUbrietaerUbescens'QIREB
Aethionemaretsina0HISNR3MNGEQTO
BelleValiabreVipedicellata4TQQIKK
AJUGapiskoi$EGEM"AKD 56
BelleValiaedirnensisYHASAX"-ASHEV
AJUGapyramidalis, 58
BelleValiasitiaca+XOQ4YAMNTD150
Alkannasartoriana"NIRR(EKDQ 60
Biebersteiniaorphanidis"NIRR152
AlliUmplatakisii4YAMNTD+XOQ
BolanthUscreUtZbUrGii'QETSEQ155
AlliUmrUnemarkii4QIGAR4YAMNTD 66
BonanniaGraeca, (AKCRX157
AlliUmsamothracicUm4YAMNTD
3SQID+IS4AM 68
UpleUrUmaira3MNGEQTO
BUpleUrUmcapillare"NIRR(EKDQ161
AlnUsincana, -NEMCHRTBROincana 70
BUpleUrUmkakiskalae'QETSEQ
Althenialiformis0ESIS 72
AlyssUmfraGillimUm"AKD 2ECHF
BUxbaUmiaViridis-NTGEW,AL$#
"QIDEW-NTG.ERSK166
AlyssUmidaeUm"NIRR(EKDQ 76
CallitrichepUlchra3CHNSRL
AlyssUmlassiticUm(AKCRX 78
CampanUlaaiZoides:AQAMEW'QETSEQ172
AlyssUmsphacioticUm"NIRR(EKDQ 81
CampanUlaasperUloides
"NIRR/QOH %MGKEQ175
AlyssUmteniUm(AKCRX
CampanUlaGarGanica4EMRTBROacarnanica
$ALBNKDS $ALBNKDS177
AnchUsasamothracica"IGAYYI3EKUI 87
Anthemiscretica,RTBROsibthorpii
'QIREB 'NUAEQSR
CampanUlaGarGanica4EMRTBROcephallenica
&EEQ (AXEJ
AnthemisGlaberrima2ECHF 'QETSEQ
CampanUlahierapetrae2ECHF
Anthemismacedonica"NIRR/QOH
RTBROorbelica0AMI /BEQOQ'QETSEQ
CampanUlamerxmUelleri0HISNR185
Anthemissamariensis4TQKAMD
CampanUlapanGea(AQSUIG187
AQUileGianiGricans"ATLG
CampanUlapapillosa(AKCRX
ArabisprocUrrens7AKDRS+IS101
CampanUlareiseri(AKCRX
ArenarialeUcadia0HISNR3SQID
Armeriajohnsenii0AOAM+NJJIMI105
CampanUlasamothracica$EGEM 'QETSEQ
"TQDESRTBROsamothracica
ArUmcyrenaicUm(QTBX107
CampanUlasaxatilis,RTBROsaxatilis
ArUmpUrpUreospathUm0#"NXCE
AsperUlabaenitZii(EKDQEW"NIRR111
CentaUreaachaia"NIRR(EKDQ
RTBROcorinthiaca"NIRR(EKDQ 0HISNR
4'ENQGIADIR200
AsperUlabrachyphylla4QIGAR)ASQN
CentaUreaaetolica0HISNR4'ENQGIADIR
AsperUlaelonea)ASQN4'ENQGIADIR116
CentaUreaarGentea,RTBROarGentea 205
AsperUlamaleVonensis%HQEMD
3CHMBECJ 4ELERX118
CentaUreaarGentea,RTBROchionantha
4TQKAMD,#HIKSNM 'QETSEQ207
AsperUlanaUfraGa%HQEMD'TSEQL120
CentaUreaathoa$#RTBROathoa
AsperUlasaxicola%HQEMD
CentaUreaattica.XLAMRTBROmeGarensis
(AKCRX(AXEJ $NRSK211
AspleniUmcreticUm,NUIR2EICHRS:AQAM125

CentaUreacharrelii(AKCRX$Q216
ColchicUmsoboliferUm
&IRCH#!-EX 3SEF
CentaUreacithaeronea0HISNR#NMRSAMSIM218
ColchicUmsteVenii+TMSH
CentaUreaheldreichii(AKCRX220
Consolidaarenaria#AQKRSQL
CentaUrealancifolia3IEBEQEW3OQEMG222
ConsolidabreVicornis6IR 3N
CentaUrealeonidia+AKONTSY#NMRSAMSIM
ConsolidatenUissima3L 3N
CentaUrealitochorea4'ENQGIADIR0HISNR227
ConsolidatUntasiana(AKCRX 3N
CentaUreamessenicolasiana4'ENQGIADIR
'$ILISQEKKNR2NTSRI
ConVolVUlUs mairei-AIQE0ESISL
Coriariamyrtifolia,
CentaUreaniederi(EKDQ
Corydalis thasia3SN+ISAM 3SN+ISAM
CentaUreapaxorUm0HISNR4'ENQGIADIR
CrepisarcUata+ALAQI3SQID
CentaUreapeUcedanifolia"NIRR/QOH
Crepisathoa"NIRR
CentaUreapocUlatoris'QETSEQ
CrepisaUricUlifolia3IEBEQEW3OQEMG
CentaUreacarystea4QIGAR#NMRSAMSIM
CentaUreaprinceps"NIRR(EKDQ
Crepisheldreichiana+TMSYE 'QETSEQ
CentaUreapseUdocadmea7AGEMISY
CrepismerxmUelleri+ALAQI(AQSUIG
CentaUreapUmilio,
Crepissibthorpiana"NIRR(EKDQ
CentaUrearechinGeri0HISNR251
CrocUsrobertianUs#"QICJEKK
CentaUreasUbciliaris"NIRR(EKDQ
RTBROacarnanica-ASSHR
CrocUsspeciosUs-"IEBRTBROspeciosUs
CentranthUssieberi(EKDQ256
CrUciatataUrica0AKKAREW7IKKD %HQEMD
RTBROeUboea%HQEMD %HQEMD
CephalantheracUcUllata"NIRR(EKDQ258
CUscUtaatrans&EIMBQTM
CephalariasQUamiora3IEBEQ 'QETSEQ261
CyanUsbaldaccii"AKD (NKTB
CerastiUmdominici&AUAQGEQ
Cynaracyrenaica-AIQE7EIKKEQ
CerastiUmillyricUm!QDRTBROcrinitUm
,NMRIMG 0$3EKK7HISEHEAD265
CynoGlossUmsphacioticUm"NIRR(EKDQ
DactylorhiZaincarnata, 3N
CerastiUmrUnemarkii-RCHK2ECHF268
DactylorhiZamacedonica*(KY+MJEKE
ChaerophyllUm creticUm"NIRR(EKDQ270
DactylorhiZapythaGorae'KY(22EIMHAQD
CicUtaVirosa,272
Datiscacannabina,
Clematiselisabethae-carolae'QETSEQ
DianthUsarpadianUs!DE"NQML
ColchicUmasteranthUm6ARRIKIAD+0EQRR277
ColchicUmbUrttii-EIJKE280
DianthUshaematocalyx"NIRR(EKDQ
RTBROphitosianUs#NMRSAMSIM
ColchicUmchimonanthUm+0EQRR282
DianthUsinGoldbyi4TQQIKK
ColchicUmeUboeUm"NIRR +0EQRR
DianthUsjUniperinUs3L
RTBROkaVUsicUs4TQKAMD
ColchicUmlinGUlatUm"NIRR3OQTMEQ
RTBROlinGUlatUm286
ColchicUmparnassicUm
3AQSNQI/QOH(EKDQ288
ColchicUmpeloponnesiacUm2ECHF
0($AUIR
ColchicUmpsaridis(EKDQEW(AKCRX
ColchicUmpUlchellUm+0EQRR
ColchicUmraUsii+0EQRR
DianthUsxylorrhiZUs"NIRR(EKDQ
Drabalaconica3SEUAMNUI+IS4AM
DrabanUda"K !K 3HEHBAY-+NCH
Drabastrasseri'QETSEQ
DroserarotUndifolia,
Dryasoctopetala,
Drymocallishalacsyana$EGEM
+TQSSN3SQID

Acantholimon aegaeum F. K. Meyer
54
Plumbaginaceae
Acantholimon aegaeum F. K. Meyer in Haussknechtia 3: 38 (1987).
3ys ;{{ E t t= b#HINRc PaUli
{ozt*%
zngsjmtoqinm|gskrggmh|mm|oqp
knkkghgnghkt{n|jg}totoi|k|qsk zqtk htzto zkqtqltos t kjt
k|qstggpsqsgzgzqtgsg}kqnsgqmksk|kgskzkqt~kk|knkstghk{ntozq~tossttqtq|mmtot
qt qt t kjt ostjkkg gz g Silene UrVillei
PetrorhaGia armerioides VincetoxicUm canescens RTBRO
pedUncUlatUm AsyneUma VirGatUm RTBRO cichoriiforme
Stipa holusericea&estUca jeanperti|ggq|kjk|jk
gz |tskqg kjm zto }tsg k ghkt{n| zkqt~
nrrnf Acantholimon UlicinUm 7IKKD EW 3CHTKSER

"NIRRIM$#REMRT"NJHAQI%DLNMDRNMIM&K4TQ
|g$NAM!JAXDIM
A echinUs , "TMGE REMRT 2ECHIMGEQ F IM &KNQA
!EGAEAA
jphpf|t{okozginstkzqt|k}g{tq}t~gg|gsnjkh{gt|gzo|s}{{g
kgq }{{g |g gg zkqgsnto {ko| {ip m
zgqrm |qs }gq|s gztzksps {kzps snt}qt h{gt |to CL gz{t hqg~pjg|{gjlkstgrgsnkv{{g W
LL iqgtkj {ti~tkj k gh{kg hm zto
~kjszkqh{{ktsh{gq~gpk{g}qq~p~kjsi{go|k{kz|ggqgtjspm|g
|{mqg|gsntkjgz{mrmqg}{{gmstrm
z{gkqg|g{kzkqggzgotonqtogrgsng
k hqg~ {k~t g|qkqm p |g CL ps }{{psksgjtj~to~k|to
LL~tsgt|gnsgpgqgg~jgq|g
z{gg|jpzqsgzkq|{ktsgkq|gsnm
{o|g LLoz{ko|tkhqgsjmkg{g|q~pgtsp{sg|gkoqkg~tgstkjgz{mrm
|qgkkqonqskqggtztgjks|gg{itosg
g|gssg tjtsp zkqnqg g{g k{g}q p
ksspqjsghqg~poskspsgmhmtok
g|qso~gtskkz{gosksmhmzqtgqmsmhkpszk{ps{tgqz|g
ztozkq~ksgzqg|gzkq|{kkgts|{o|g
fqrs{}zfg}otoAcantholimon aeGaeUm
zkq{ghstsgs gq~| g kjm A echinUs , "TMGE
|g A UlicinUm 7IKKDEW3CHTKSER "NIRR O -EXEQ
tzkqiqgkpjgkqtkjtkggsgnkqmmps
koqpzg|s SAWA to isto Acantholimon sg gqn gz kstzk |g tq}t{ti| jg|q tq}
gz ms sgt{g ~tos zq}gg gsgispqnk t
kzzkjt to kjto gz to $NAM !JAXDIM
ttzttosgsgkqgzzqtmitkskjmtkktoz{mkqtoiiksktoA aeGaeUm
ksg~pqg}ht{ggtjggzkjmm|g
sgt{g zto gs|tos ms RTBRECS Caryophyllacea
ozkq{ghskkjmzptA phryGiUm"NIRRgz
N -AMIRA $A |ts ms qsm |g g A acerosUm
7IKKD "NIRR|gA diVersifoliUm3CHVAQY-EXEQgz
posof{fpmispt
jpsisfrms|spfztstptststk
msgr|gqzgsgzgqgsk~qts|hqt
johpf|{ jqyzol sg ksjm| kjt ps sps to |g to isp st gz om{ nk
gqmqthg|gqtmst|gn|ggqm
q|m|gzk{tgqhtsmqt}m{g ms
t}gslkgkokqt Lkszngsk|kskg~qggskqgmkgpsom{tqps
|tqo}s
systAcantholimon aeGaeUm}kg~kjsgzt|{k|kgst|zkqjkk|k|gqk|gn|gkgstigg}qoispsgskqktqks
zkqt~ zto h|tsg stsg ks hqnm|k g }tq
kghkt{n||qmskhqkg|nkm{ttisptz{mnotk}gsltsgkjksjqkzkqt~zto
kpiqg}|krz{pmtoksjm|tkjtoAcantholimon aeGaeUm

mskzgq~g-TKAgtgtoA aeGaeUm gzmt

k}gsltos ~gm{kqt t |g |qkqg }{{g k
~mkgomtowgzt{kgtjtgo
tjkoztz{mnosngzqzksgnkpqmntspjgkqkgzmsztmmzqtggto
qtmtoksmst~khqknkkzkqt~
ms|tqo}toqtoq|m|gkjzgqkzkqt~tqtzk{tk|nkz{mnooztz{mno
sgzkq{ghsksgzkqtqstgqngpsrggmzgqtg|ggmo~s|{g|g{{gi
g{{gim~qmmimzkqt~zto}kgt
kjtztqksgktosgiksk|gztngtok
|qtmkt
fflyzlmnrtAcantholimon aeGaeUm }kgkt{iqnktjkg|qhpszqt|k}g{tq}psgpsk{ikzkqt~zto|g{ztoszkqhg{{ts|togzgktjkpsgpsg

jg}tqk|qmksgtog|gzttspskkgr
tog{{ksjk~tspk~gszkqkqkko|gqksg
{ntoskkz|tspsggqg|nksmt |gtzgqk{nsknqgomgr|gqzgztozgqgskt}o
igg|q~qts|jmgzqt}qk|g{ko|gqk
igjgztqoi|k|qskzkqt~ktki{t
okqtkrggpso~ss~oqsgsps
pf ypsff {{gi ms ~qm m im

zkqt~ztogzgstAcantholimon aeGaeUmngzqzksggzt}ko~ntssggskqgggtoqto
q|m |gnkqpnts p zqtgkoksm zkqt~ k jg~kq| ~jt zto ng zkq{ghsk |g to gst|tghkt{n|thtztom|tqo}gong
jg}{lkmos~kgmzgqrmtokjtogms
zqtgg ps oztz{mnos m to t qt qt
zqt}qk|g{kqkjosgmk~tsgzkqt~
kghk{ntghgnjg}t|gz{tgtz|~{pqjgkokqt L
rinrs {f }fpf{lp {flhspf fyjz

t kjt }psg k g |qqg m )5#.
ABII A |g $ zto z{mqtsg ~gqg|mqlkg p
qp65 ~khqknkstsgoztz{mnok|nk
2TDL2LMFDPSN!PHRR2LMFDPSN
Acantholimon aeGaeUmgztqtqtmstotovpgqjg

Achillea occulta Constantin. & Kalpoutz.
54
Compositae
Achillea occultaConstantin. & Kalpoutz. in Bot. J. Linn. Soc. 147(2): 250 (2005).
ys ;{{ 0E k{tzssmt t g|psg=
b-S+NTKNCHEQASHETOOEQOAQSRNFSHELNTMSAIMFQNL
SHECHAOEKNF0QNSIR)KIARSNSHE.%TOOEQOAQSRLNRSKX
AKNMGSHECQERSKIMEAMDSHE.%RKNOERRSEEOCAKCAQENTR
QNCJR RKNOER AMD SHICJESR VISH Acer semperVirens AMD
Amelanchier AKS C L .
%c -AX Constantinidis & +alpoUtZakis
{ozt!4(5ozt"&)50!
to qto zto jgmqtsg ozt{kgg h{mm

k nsto |g jsqg g zkqkqt |ts ngsjm
|g jksjqjm kjm ksg g g|{tong k gkq|t
mkstsg g ksjm| }o m {{jg Acer
semperVirens
Amelanchier parViora RTBRO chelmea
&raxinUs ornUs RTBRO ornUs |g Phillyrea latifolia ks
to |qmst k}gslkg t
RhamnUs sibthorpiana
A occUlta zqt|kqm|kqqmgtztg
mh{mmtkrqkpshq~psmszqtgktos
gz ms stsm m{g| g|stht{g stt tg
gottokjtoztqksghqkntskzmk~g{|jk
k k|knksk zkqjk tztnkk gl k ms
A occUlta }tsg tqsg gosng kjm zp g
*Consolida tUntasiana
CerastiUm illyricUm RTBRO
brachiatUmCrepis fraasii
Cymbalaria microcalyx RTBRO
microcalyx
-inUartia pichleri
-elica rectiora
Stachys
chrysanthaThalictrUm orientale |A occUlta ztqk
sgksg}ntsm|gsggztk{ktkz|qgtskjtk
zqtgkosk qpi hq~ps zto opqkkg
|qztmgitsto~gt
oztz{mnot zto }tsg ms tozqlg |g
ms zgqzgq zqtts zgqk}kq zkqh{{tsg
{t t oztz{mnot k}gsltsg g L
zkqzto
jphpf| t{ok kjt ~mglts ojk snt}qgk{~m~pqjg|{gjk|to CL

kgqgq~pmqg}{{ggqgq~pp~kjs{kggz{zkqt~j|to W LL
klkimgzz{gkgsptkjk|qto{tht|g
zt{ |qkqto {tht tjsk |p gz gotqt{thtghgn|gg|gssgtjtsptp
~stkgg}{{g~ggsptkjpz{gpk{{kz| W LLzgqtgkg}{{g
mhmk {thtk}{tkstkpk
ztj|to|to LLkqh{mg|k}g{toz{to LLhq|gzkqh{gt|to LL
ptkjpk{{kz|zt{gqgq~pp~kjs
{kgkkskhqgsjmk{g}q|ggszkqnqg zg gsntj~m k{{kz| jg}gs {kg g{{
kgqgtgjskkq}kqkg|gsnjgn{kg{ko|
gs |k}{t i{pjg k ghgnk {tht
jgkps W LL snjg j|to kqg}qjg{ko|gjksjmg}jkoqosksgmhm
toigsgzkqh{tosms|tqo}tog~gsto~gsg
ksgsptkj|ggs W LL~pq
zzzt k gg} |tqsg ~ h{kssjm gs gztqqt}tsoiqgg
fqrs{}zfAchillea occUlta zkqiq}m|kzq}gg#NMRSAMSIMIDIR+AKONTSYAJIR kpqnm|kgs|kkg|qksmgkjsztogztk{ts
posof{fpmn #NMRSAMSIMIDIR
+AKONTSYAJIR
jpsis frms|spf oqp t t p ztqk
sgkzk|gnkptstst
johpf|{jqyzoltz|ksjm|kjtm{{jgspstgzgqmto{t~qg|gtozqlg k{tzssmt |gn |g gz g hqg~jm
rgqmisppzgqzgq|tsggsg}kqnsg
qm
sys Achillea occUlta k}gslkg |oqp g
gskqg gg to qto to{t~qg sg qt
kgto otqto om{kqm |tqo} g L
t tztt kzm ztqk sg nkpqmnk p zqt|gm to
qtoqspsgzqtgsttgsgt{|g}ogot
to kjto }tsg osnp k hqg~jk z{gi |g
|qmst g htqktgsgt{| |g gsgt{| gg
kpiqg}|krz{pmtoksjm|tkjto!chillea occUlta

tksjm|kjtAchillea occUltagztqtto{t~qgmk{tztsstovp%+g{ztol|m

ms tjg Anthemoideae m tztg oiikskk ks k

msAchillea RECSINMPtarmica tjgpsAnthemoideaegsghgnnm|kmsA RECSIN Anthemoideae 'TN
AK3ATJEKAK%HQEMDNQFEQ'TN
g{{mgrst|tqtnmmgomRECSIN~qklkg
zkqkqmqkosg
gkjmztoksggrst|kiikqgmsAchillea
occUlta ksggA barrelieri 4EM 3CH"IOA mUcronUlata "EQSNK A oxyloba $# 3CH"IO|gA schUrii3CH"IO
oi|k|qsmtjgkjskrgz{skggzsssg
qm{zk|gggqzngks{mmtjgA barellieri A mUcronUlata A oxyloba }tsgms
|gg{gksozq~tosgsg}tqpszgqgzsp
kjsgzms{{jg|gmA occUlta nkpqkgikpiqg}|gzttspsmgztooiikskm
Achillea occUltajg|qskgk|t{ggz{toto
oiikskmkrggmtq}t{tigps}{{psg
}{{g m hm |g g |gkqg }{{g to h{gt
msA occUlta }qtoslkimgzz{gkgsptkjk
{tht {tht got |gn |g t g|qgt {th to
}{{toztqksgjgqtsgzkqgqpzkqzto~q
t ts to sg kzz{ts |q {th tjsg
ztqksgozq~kt~tto}{{to|pgzto
|qto {tht {g g {{g oiiks m kjm ~tos
}{{g hm |g |gkqg }{{g h{gt k zkqkqto {tht zto ksg iqgtkjk p ptkjk k{{kz|totztqkmos~kgsg~ltsg
zkqkqt zp ms zkqzpm m A mUcronUlata
|gmA schUrii
gskpzlk |ztts kt |sjost krg}sm t

kii{{tstz{mnotmngzqzksgk{i~tsg
igzngsg{{gitokzzjtokz|sjosmgkj| ms zkqzpm zto gqsm| krk{rk kzmqktos
t}o|mzkqh{{tsgzgqjkiggzoq|gi
p|gqk}kmsgqgro{jmh{mmztozqt}qk|ztgzqtggtkjt
(tqksh{mmmto{t~qgoz|kgkstsmh|mmgz|tzjggitzqthpsgqimsstrm
|g t |g{t|gq sg jkokqkps jqt zto tjmik
|kqgk m{kz|tspsg ms |tqo} to qto
jko|t{sk ms tj| kg}tq gsnqzps |g lps
pmkz|qgtgh|mm}gskgjksnkk
ozkqht{| |sjost ms kzhpm ps oztz{mnos
m A occUlta 4g ~tqt}ig lg |ggsg{stos |oqpggsnsggg|gg}{{gtokjtog}stsggsii~gg|gkqgggtoh{gt|g
to t}ng{t gsgspm |ts t ql| mg
t}ng{tgsgspmo~szqtgktsggk
|t{mkqpihq~psztoj|t{gzqtkiiltsggzki{g~tqt}iglgqsgtgmA
occUlta zto}tsg|ts|kqgkm{kz|tspsg
zqtkiiltsg ko|t{kqg |g ksg zkqkqt k|knksg k |sjsto |g zngs |ggqt} {{k
ojkp}tsgkgz|qmsggzqgqm
ztozqt}qtoszqtggtgzmsh|mmt
|ggzgsnqzskjqgmqmk
pf ypsff ztkjztk g{{gi t tztk
ng ztqtgs sg kzjqtos gqsm| sg ozthgntosmsztmgtozkqh{{tstmAchillea occUlta
ngzqzksggzt}ko~nts
g }o m A occUlta |gg}qstos sg kzhstos
zgqmszgqtogzkqtj||gzgh|mmto
htztotoztgjztkksg|tztmmmh|mm p ng ztqtk sg kk ms zgqgipi ps
g~gsps to kjto |gn |g sg gsgk{k ms gsgspmtoz{mnotgzzqggwk|ttog
tgksqikgngzqzksggzt}ko~nk
toi|k|qst}oksgk{|o||gztqkk|t{g sg |g{{kqimnk k tgs|t zto jgkqg
p}ohqg~|mzpsgg~gstokts|g{{m{t
~kqk}gsltosom{ztt}qpm|ggztk{ts|g{qztjgqmm|gjgsttokjto
t qt to{t~qg ~k kz{kik sg zkq{m}nk t
|ot ps t|t{ti| mgs|s zkqt~s v
zqtggtoqtogsgskgsg~knk|
kzjqgmmsk{{ts|kzhpmmA occUlta
ffl yzlmnr Achillea occUlta ~mglk

sg ki{t |g ksgt z{mno t qt to{t~qg
ztotgqnpsgpstokjtoozt{tilkgk
gq|k~{jkg|qhgqnps}osztok}gsltsghqg~jkkrqkksgj|t{tsgozt{tink g }o o~s qltht{ts g |gkqg ggtok{~toto|g~mgltosstoqjg|k
}{{psztoosk~ltostsgikszt{{gz{ggg
zqmk|mmtoz{mnotmto{t~qgt
jkrkgogskq~gszkqztoktgp
gzkqkqtg|qhkzzgkzgsk|mmt
ozt{iktsjtz{mnog tg
jkkqtispoztz{mnomzgqzgqzkq{ghskzkqztotgksmszkqt~mtozqlgk}gsltsgzkqztotg
rinrs{f}fpf{lp{flhspffyjzt
kjt ~gqg|mqlkg p qp 65 k hm ms |gmitqtztmm m )5#. }psg k t |qqt
$|oqpkrggmzt{zkqtqsmzkqt~k}s to s |g m Achillea occUlta jks }gskg sg
js|rlorfrril
$zjnmpsfzysnk{l

Adonis cylleneaBoiss., Heldr. & Orph.
o-3
Ranunculaceae
Adonis cyllenea Boiss., Heldr. & Orph. in Boissier, Diagn. Pl. Orient. ser. 2, 5: 5 (1856).
j{nys 3SQID B ;{{ 0E k{tzssmtttqsng=bIMLNMSE:IQIA#XKKEME OQNOE
&KALBNTQISRALAKSOEDc!OQIK-AOrphanides ' "/)3t{k|ozt,$75
tkjt}kgkzmkghkt{n|z{gikgr
gqgsjsjqpsAbies cephalonicao~szqtgkostgzkjmzpgPteridiUm aQUilinUm Vicia cracca |g
-arrUbiUm cylleneUm kms|gqzt}tqggqit|g{t|gq g ksgqg gg to A cyllenea rmqgstsg
|g sg k{~m k}gsltsg spq ms strm gz g
ozikgqlgg
A cyllenea }kgkgr L
gssgqt g{kjm g zgqgqmgs
t kjt ksg gzs gz gq|k jt{sk ~gm{tqpsotqpstoqtogpkzkjtoi|k|qsk zkqt~ ~qmtzttsgs ig |g{{qikg k
zg{gkqgm
nrrnf ChrysocyathUs cylleneUs "NIRR (EKDQ

/QOH #HQSEJ3KAUIJNUIM0QERKIA
jphpf| t{ok zg to CL k
hqg~ jg|{gjst qlpg v{{g hm k
g|q~tztojgz{gskgmhmtok|t{k
{gg CLqips| ptkjo~szkqt~j
gqg|gg{g|q~pgzt{m||gg{ti~tkj |to W LL tr{m|g snt}qt {k~t sg {ig {{ts kqpg g|qgg k {ik
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var.ligustica (Savign.) Fiori
54
Gramineae
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54
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54
Labiatae
Ajuga piskoiDegen & Bald. in Oesterr. Bot. Z. 46: 414 (1896).
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57
Ajuga pyramidalis L.
54
Labiatae
Ajuga pyramidalis L., Sp. Pl. 561 (1753).
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65
Allium runemarkiiTrigas & Tzanoud.
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Alliaceae
Allium runemarkii Trigas & Tzanoud. in Nord. J. Bot. 20: 89 (2000).
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67
Allium samothracicum Tzanoud.,

Strid & Kit Tan
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Alliaceae
Allium samothracicum Tzanoud., Strid & Kit Tan in Portug. Acta Biol. 19: 356 (2000).
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Alnus incana (L.) Moench subsp.incana
o-3
Betulaceae
Alnus incana (L.) Moench, Methodus: 424 (1794) subsp.incana.
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71
Althenia liAormis Petit

Zannichelliaceae
Althenia liAormisPetit in Saigey & Raspail, Ann. Sci. Obs. 1: 451, t. 12 (1829).
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Alyssum Aragillimum (Bald.) Rech. f.
54
Cruciferae
Alyssum Aragillimum (Bald.) Rech. f. in Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 80(12): 58 (1943).
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75
Alyssum idaeum Boiss. & Heldr.
54
Cruciferae
Alyssum idaeum Boiss. & Heldr. in Boissier, Diagn. Pl. Orient., ser. 1, 8: 35 (1849).
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bADRAWACACTLIMTLLNMSIR)DAEACS;RICAKS=
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t goq t om{kqt mkt m qm g
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nrrnf AlyssUm cUretUm 'AMD "TKK 3NC "NS

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77
Alyssum lassiticum Halcsy
54
Cruciferae
Alyssum lassiticum Halcsy, Consp. Fl. Graec. Suppl. 1: 10 (1908).
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80
Alyssum sphacioticum Boiss. & Heldr.
54
Cruciferae
Alyssum sphacioticum Boiss. & Heldr. in Boissier, Diagn. Pl. Orient., ser. 1, 8: 35 (1849).
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Alyssum tenium Halcsy
54
Cruciferae
Alyssum tenium Halcsy, Consp. Fl. Graec. 1: 93 (1900).
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86
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Anchusa samothracica Bigazzi & Selvi in Nord. J. Bot. 20(2): 141 (2000).
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subsp. sibthorpii (Griseb.) Govaerts
$-
Compositae
Anthemis cretica L.subsp. sibthorpii (Griseb.) Govaerts,World Checklist Seed Pl. 1(1): 7 (1995).
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Anthemis glaberrima (Rech. f.) Greuter
"1
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Anthemis macedonica Boiss. & Orph.

subsp. orbelica+<ID*=@MKM"M@PO@M
54
Compositae
Anthemis macedonica Boiss. & Orph. subsp. orbelica+<ID*=@MKM"M@PO@MDI2DGG?@IJRD<
(2003).
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Anthemis samariensis Turland
54
Compositae
Anthemis samariensis Turland in Willdenowia 38: 63 (2008).
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i|topsko|sqpsqm ispstgz
kpiqg}|krz{pmtoksjm|tkjtoAnthemis samariensis

tksjm|kjtAnthemis samariensisis kzsp |gthtzto|p gzgko|qmmqmvpTQKAMD

jg}tqk|gzmA samariensisttq}t{ti|
t|gt|t{ti|{zkts4TQKAMD igzkqkqk{kztqkk
rmqgg4TQKAMD ozt{ikt|qmstzto
krkqkosnm|gs gszqtpzktos t zt{ t ps
zngss |gg{{{ps htzps ig t kjt | zto
oztjk|skzngssgozq~ksgikntz{mnotmrmps~{jpspqpsgps
glkstgzqjqtggojkjtsgzqtkskgt~gqg|mqqp65 igtkjt}psg
kms)5#. g}tz{mqtgt|qqt$j
tz{mno|gg{ghskgzt{|qzkqt~kintostJL2kjtnkztogz~toszkqzto
JLkgrto
fflyzlmnrsgisptstjtoztz{mnot to kjto oztz{mno gz zto t

kjtzkqiq}m|kzkq{ghskto{~ts{ikk|gtsjk qg tg sg jkkqt oztz{mno
JL stp m hqkg z{koq m |tqo} o{gs|tozkq{ghskto{~tsqgtgks
ozq~tosjkjtsgztosgoztjk|stososk~lksk
kk g|qgk jg|osk to gqnt ps
gpstoz{mnot
pfypsffoztz{mnotngzqzksgzgqg|t{tontsg jgq| ks ng zqzk sg zqgigtztmntszqtk||qkoskpsqiqptqpsk

sg kzmgsnts kzzqnkt oztz{mnot ngs
t zt |g{{m{k zkqt~ ksg go zto hq|tsg
sgm|tqo}o{gs|tomks|tqo}tiqg|gzkq{ghstostqt|stztojg~pqlk
~gqjqkmggq|gto{i

rggto|qmsjtohtzto~kjs{ggtgksggzqgtgh|tsglgt|gts
snqpzt ks }gskg sg ozq~tos |sjost krgg
}o|s|ggqt}sztongztqtgssgg}gstos
sgktosjqg|tsz{mnotokjtoz{mno~kjmkzhkgzikitsgztoosjtsg
k|gq|}gsksgzp~oqtsktzgik|g
-HBGMJ@Q3SPJ@LC
sp|g|pkz}skgmgrgsngtoksjm|tkjtoAnthemis samariensisgzgko|qmmqmvpTQKAMD

54
Aquilegia nigricans Baumg.
Ranunculaceae
Aquilegia nigricans Baumg. in Enum. Stirp. Transsilv. Princip. Vindob. 2: 104 (1816).
ys t kjt ~k zkqiqg}k gz m togsg
/{ozt#,
sysispzkqt~ztomAQUileGia niGricans

k}gslkg ms {{jg hq|tsg k okqt gz
p L k r}pg |g zgqo} jgs k
&aGUs |g PinUs |g k | }o{{th{g jm tqkss
zkqt~ssgzkgkzkqkqt{ikqtoiq
qmkjzgqmpqg|{omgz}o{{th{to
nsto|gjsqg
nrrnf AQUileGia haenkeana +NCH 3XM &K 'EQL

(EKUED nom illeG
A VUlGaris , UAQ niGricans "ATLG 3CHTQ IM 6EQH
-ISSH3IEBEMBTQG6EQEIMR.ASTQVIRR(EQLAMMRSADS

A VUlGaris , RTBRO niGricans "ATLG $NLIM IM
0QERKIA
fqrs{}zfqstoiiqg}k~tosozthhkmsAQUileGia niGricanskoztkjtzt|{gm
A VUlGaris,g{{gzi~qtstohtgs|tzpt
!JEQNXD |g3SQIDB ~kiskgztjk|p
jgkqtkjttistAQUileGia,ngzqzksgk{kmnkjkrtj||gsgzqgigtztmntszkqggjggoqkpskgrpskjsgtztgzkq{ghsk
ksg|gntqnkkht{ti|qztmtqtnmmps
kjs
jphpf|{itkzgksgkqpjg|{gjlkstro{jkqlpgk{~mqng CLkqp

~stjmkmgjksjkq~pgmhm|ggsgkistgjksjk|gmgjksjkggskqggg
v{{g hkp {ig k g|q ~t {gg koqp
qips| W CLq~jgskqggg
gsq}pqips|pkoqpgsptkjosnp
W CLk{g}qphgnpq{thg!skqg
}{{g hgngp |qkqg gsgg ~g k zt{
|q iknt snm {ig ~mgltsg gst| snkthqok{g}q|k|{sggsksgskgqhgnp
|ogs p k{gstjm rpkq| gg zkqgsnto
jksg gzt|{stsg koqp k{{kz|
LLpkq|gg}qtsgsk|qg pLL
kz|mksg|oqz{|qt|to LLztojkoqskgms|tqo}tskgq|ktk}gsg|qkqtpsmpstozkqgsntotqst~tos
kgqgzkkmtsjm+gqzgztk{tkstgz
k{g}q|oqtqg}jk~stjkn{g|kkzt{{ gjksjk q~k kspsto m hm |to
LLkzt{{zqgg
fflyzlmnrk|nkz{mno~toszgqgmqmnkst{igpkq|jk|jk}oqkltsgpzkqkqt{kztkqkzgqgmqkzkjto
t{{ts
kkoqpzg||{g|gtkjtjksgzk{kgg{{p
{tmk{{ms|~{pqjgnkpqkgqp65
jg|k|tsmkrz{ptokto{ito|qt|g
posof{ fpm n UAM ,NNM

6ARRIKEUA
jpsisfrms|spftstpgq~to{to
johpf|{ jqyzol AQUileGia niGricans ksg
kjt m oqzm t tztt k}gslkg k tqks
zkqt~gzmsoqg~qms{{jg|ggzms
togsg~qmsko|tqpgms{{jgksgispstgznk3SQIDB ztogs|tosk
zkqt~ g qm q{g|g t qkhkss qt
tt gng |g {{m }m t pgsssps
|gtsqtttpgsssps|ggtq
gn t jgjk tqks zkqt~ ~gm{tqto
otqtom|g{{jgjks~tosg|mkqkosmnk zt{ |g{ ksg zngss sg ozq~k t ks {ip
kjtkzkqkqkzkqt~g{{zspzgqgsk
zt{zst
kpiqg}|krz{pmmQUileGia niGricansms{{jg

jzgqtoz{mnotmgsksgmgs|gtoztqksg|ggqg}kgzsg|gstgo~
ikitszpsgtz|zqiqgggsgjpmg
jg|zoq|gi
mgs| t k|kgsk ht|tksk |g jg| zkqt~ztogogsgzkgsgzgqgkstoskms

zgqgjtg|totq}tmjmtoqigzo|ssjgst|gtzoq|giztqtssgktosmgs|tok{{ms|tz{mnot
pfypsffks~tos{m}nkkj|qgig
t kjt ~q kqg g ms QUileGia niGricans ksg
tkjtAQUileGia niGricansgztqtqtvp!3SQID
100
Arabis procurrens Waldst. & Kit.
o -3
Cruciferae
Arabis procurrensWaldst. & Kit., Descr. Icon. Pl. Hung. 2: 154 (1804).
ys;togsg=7aldstein no 02
nrrnf A praecox 7AKDRS +IS EW 7IKKD %MTL
(NQS"EQNK
A recta"ATLGMNM6IKKIM(IRS0K$ATOH6IKKAQR

jphpf|t{okzgkg|qzgqg}ojkk{~m|to CLgstqnpsgpqngj~p
jg|{gjkiosk{ikj~jkq~kv{{g
qjg|ggsptkjpgs{ti~tkj W LL
tr{m|gg|qggk Njskk|nkz{koqo~skjmgz~qpmms|pkz}skgiosk|
gzzgqo}|ggskqgm|pkz}skgzto
}qtosj~jkq~kv{{gh{gt ~gptkjpkz|mk|o|{|p|gqjtkjhm
W LLgrgsngzt{ogsnhtqotkj~pqhq|g gz{ k {ik jg|{gjk m hm kzm|osksmms|gqzt}tqgtj|ttqlst LL
zg{g kz|m ptkj LL ios g z{koq|
g|tkjmhmg{g LL{ko|gr|gqzg
gqgqgiqgtkj W LLtqlsthg{hjkkkoj|qtkgtskqt|gqzt}qt
ztj|t ~q LL zqgg kz|m LL
{kg~pqzkqig|ggs
posof{ fpm n *AKAR
3TNLIMEM
skg ps }{{ps ps ptkjs p kzm|ps }{{ps

h{gtk|o|{|p|gqjtkjhm|gps{kzs
tqltsps |kqps |to LL k ztj|to
g|qkqtopsLLoiiks|mkjmzpmA
Vochinensis3OQEMG|gm ferdinandi-cobUrGii*+EKK
3TEMDEQL jks ~tos ~q kqg gsg}kqnk gz ms
{{jg
ffl yzlmnr k{{ms| z{mno to
kjto hq|kg ms |g tjzm |g jg|qskg
to{~ts k oztz{mnot %KEHEQIADNT
AK kig{kqt oztz{mno hqnm|k
ms zkqt~ qo zto k|nm|k ozq~tos
zkqkqggztgoz{tztoztz{mnot
ksg zt{ |qkqt |g k|nm|k gztk{tsg
ost{|gz{ikqgpsgpsost{|
gqn ps pqps gps ms zkqt~ krz{pm
mArabis procUrrensms{{jgksgj|t{tkzto
zgqst sg k|mnk k g|qhkg ng zqzk p sg
zkq{ghsk zkqkqg gz tg zkqt~
k}smtokjtoksg~k||qJL2
tkjt~gqg|mqlkgp~kjszk{tkst.4 k
hm ms |gmitqtztmm m )5#. |oqpkrggmzkqtqsmzkqt~k}sto
ms{{jg
Arabis procUrrens jks }gskg sg gskpzlk
jpsisfrms|spfzq{tpt
johpf|{jqyzolost{|krz{pmtokjtozkq{ghskggqzng|gtqkszkqt~
to gt m g{|gs| kqtsto to{igqg{{jgzqmstoi|t{ghgtogsg sg}tq
gzm{thg|gjkskzhkhgnm|gsmngiksms
k~gzto|g{{kqikgzkqgg|khqg~|mzto
|gztqksgjg}kikms{{jgksgispst
gzmstqtkqmtjzmvqg|sgkzjg
qo %KEHEQIADNTAK4RIQIOIDIR!SHAMARIADIR4AMAK
systkjt}kgkhq~tokhqgkkj}mk{kz}o{{tzmgkjm&aGUszgskqgk
|qgstiggjgpspsk|kpsk1UercUs ROO
PinUs niGra|gP sylVestriskiqgsmghk{ntk
okqt L%KEHEQIADNTAK
4AMAK
fqrs{}zf Arabis procUrrens jg|qskggz
{{gk{{ms|kjmtoistokrggpsg|qszgqg}ojpstoj~jtq~gtms|pkz}-
kpiqg}|krz{pmmArabis procUrrensms{{jg
101
|ztts kt |sjost krg}sm t kii {{ts

ptgsnqzskjqgmqmkmszkqt~ztqk
sg zqt|g{tos kpm to z{mnot gqgmqnm|kmkrz{pmtoz{mnottokjtokostkg
gzmjzgmm|ttimtojtomszgqrm
gstips |g ioss zqgss jg|s jqps ztsptkz{k||o{ttkmjgqmmpsgstips|gmosqmmpsjg|sjqps|gn|g
m jstrm sps jg|s jqps |g qg|kqtjqps
kosttsmjgqmm|gmsgrmmtoz{mnotto
kjto ks m zo|s oi|pm ps jgs m jpm
psgstips|gmmosqmm|gjstrmjg|s
jqpskzmqkltoszngsgqsm|tooztz{mnottokjto
oztz{mnotmszkqt~tovqg|tzngs
sgjgq~toskig{kqt|sjostkzkjjksiskg|g-
g jg~kq| kzhgm ks to jgmqmto smktotogqnstotovqg|t

pfypsff{{giztongztqtgssgkzjqtosgqsm|sgozthgntosmsztmgto
zkqh{{tstmArabis procUrrensngzqzksggzt}ko~nts w |qt qt zqtgg zqtkskg t
m ksmqpm ps gqjps jg~kq|s gq~s t
|gtokoqkqto|tstztojghkjqgmqtztkg
g jm m zkqt~ zm zqzk sg isk z{qm
|ggiqg} to z{mnot k sg |ggk josg m
k{|sgrtomkggz|gmitqkgzk{tkspskjsm)5#.
orfrrsjsiopysnzs
zrlzjnmjpisnorrlpyil
tkjtArabis procUrrensgzmstqtkqmtjzmvp+ktjpqzto{t
102
Arenaria leucadia Phitos & Strid
N$-
Caryophyllaceae
Arenaria leucadia Phitos & Strid in Bot. Chron. 11: 55 (1994).
3ys;{{)N) stttko|=b.RIDE
CJL.7NFSHESNVMRAMDXREARHNQECLN
.N %c-AXConstantinidis & Strid
/{ozt50!)ozt#
jphpf| tstk zg k zt{oqnto {kzt
h{gt|to CLgkqtqtto}ot
zo|s~stjkmgjksjktgskqt|g{ozkst
kgjksjk|gmgjksjkq~kgkqg}{{g
toh{gtzgto{tkj|to LL~gk skqggkgg|ggskqg~gptkj
q~pgrgsng~k|zo|ssnkthqok
snm |g{osm k zo|s kz|k gjksjk
q~k q|g |to LL ptkj {ti~tkj
trgzg{g|to LLptkj {ti~tkj
tr{m|ggjksjmk|qzkjgz{gpkhqgsjm
g{g gsptkj t|m k{g}q g|qkqg ps
kz{ps{ko|gptkjk|{mqt~pgt|mzqtgzg{gzqggzkqztoWLL
sk}qtkjgqgk|qgzkqtiio{psg}og
posof{ fpm 2n 0HISNR 3SQID

jpsis frms|spf !zggzq{to~qg
{mgto
johpf|{ jqyzol tz| ksjm| m sto
ko|jg gs isp ~q zq}gg st gz t
locUs classicUsmsgjmzgqg{gtog|qpmqtooqzkqg 0HISNR 3SQID q}gk ~{pqj|
o{{timstmko|jgqi|to k~gs
p gzt{kg ms kqkm to kjto |gna {t t |tpsgtnssm|gzgqg{g|lsmzto
~mgltsgghqkgm{stn{ggmz{m
m ko|jg gz t g|qpqt oqzkqg ~q to
{to zp kzm |g k g|m |qt oztz{mnotmszgqg{gngmszgqg{gg{|g
tqtg|qt}pmA leUcadia gsg}qnm|k
gzmsk}g{{msgkjg}tqk|zkqh{{ts|gn
|gktqkszkqt~mko|jgTQSNM
skgonktokjto~qkltsgkzhkhgpm
sys Arenaria leUcadia ksg g{}t kjt t
~gqg|mq| htzt to tztto ksg t gjk
zgqg{ktosmtzto}kgk|g{gskzoisk
|tspsk ~gqg|mq|s gt}{ps kjs zp g
Cakile maritima %rynGiUm maritimUm 'laUciUm aVUm
-alcolmia maritima OtanthUs maritimUsSilene colorata
-edicaGo marina %chinophora spinosa CrithmUm
maritimUm Anthemis pereGrina | t qt g|
k}gslkg k |qt oztz{mnot gjk
ki|t{zk zto ~mgltsg to zgqgng{to

ghkt{n|t hq~to m zkqt~ ks ts g{
~mglk |q oztz{mno ms sp lsm m
gjto zgqg{g gl k {{g kjm }os zp g
%chinophora spinosa -atthiola tricUspidata &Umana
thymifolia Anthyllis hermanniae %rynGiUm maritimUm
CoridothymUs capitatUs|
fqrs{}zfArenaria leUcadiaksgoiiks
k ms A conferta "NIRR |g ms A serpyllifolia , ms
ko|jgosgsgk|tz{mnotkmskoqpkrgz{psm|gosnpkqgz{tkjA serpyllifolia
, m tztg ~k hqg~kqg zg{g g|qkqg ktitsg jggg |g |qkqg zqgg k tr{m|g
}og z ms kzm jz{tkj A conferta jg}qk
ttiknt|gmstq}pszkqps|gto
q~gmt|gtshtztg}tmk{kogg
ksg sg zkq{}t tqks SAWNM t tztt gzgs k
okqt L0HISNR#NMRSAMSIMIDIRA
ffl yzlmnr Arenaria leUcadia ~mglk |qt oztz{mnot ms zkqt~ to g|qpmqtooqzkqgtgst~toztz{mnoosgg
gz zkqzto tg |g ms zkqt~ ps {ps
gz zkqzto tg kgr gos ps jt nkpshqnm|gsk{~g|gjg|tqzsgtggot
tokjtomszgqg{gtogngt~mglksgs
kpiqg}|krz{pmtoksjm|tkjtoArenaria leUcadia

g|m |q oztz{mno gztk{tkst gz zkqztotg|gmsgjmzgqg{g|lsmtog{t

t oztz{mno gqnk zkqzto tg t qt
g|kqnm|gszkqztotgost{|k{m
ms zkqt~ krz{p m |ggkqnm|gs {ikqg
gztgqi|to
zgqgng{thtzttotztto}kgmArenaria
leUcadiaj~tsgki{tsgqnskz|kzs|gms
nkqszkqtjt|ggogztk{kmgs|gzk{igt
kjtzkjpt}ogsnlk|ggzgsnlkkspq ms strm |g ms nkqs zkqtjt koq|kg oz
mstq}zkqpshonspsmstgzt}kikgtkt|sjost|ggqt}to|gms
jq|kgmzgqgnkq|zkqjtog|gzk{ig
t}ogztk{km|ggzmmmzgqg{g|lsmm
tztgt}k{kgkkkqigtg{tztmmpsnssk
|gz| m~gsgg gz ts t ko|jg zkqt~
{stn{gg kkkqigtoq|oztjtzgqg{gmzkqt~ng kkj}tqkkzlkjqgmqmktztkkzjjtsgtkz|zk
pszgqg{stosmtktjtm~gst|spsps
gtnskqmgztqqps|
rlksgmkpnkgs|gmzkqt~m{stn{ggmz{mmko|jg~kksg~nkt|ot
zqtgkotsps zkqt~s v '2
t|gnkzqtggjks~ktog||ggzqg||k}gqti

qtkstok sg |ggg~nk m Arenaria leUcadia ms
|gmitqg ps sjoskosps %. m )5#.
krggmzkqtqsmzkqt~k}smps
gzk{sztogskpzltostoztz{mnotm|gm
ozthnmmztmgpst|tzpsmg|qqg"ABIII |g#gI |g{ztosmszkqzpmmA
leUcadia
pfypsffztkjztkg{{giztongztqtgssgkzjqtosgqsm|sgozthgntosms
ztmg ps htzps m Arenaria leUcadia ng zqzk
sggzt}ko~ntsjgkqgmszkqt~togngt
ztoj~kgki{mtoq|zm|gmnkqszkqtjtzqtkskgmk{ki~ksm|smmpst~mpsk
khgzqtt}o|zkqh{{ts
k zgqtk zkqt~ toq| jqgmqmg
ksj~kg g qg zqtgg ps htzps sg ksg
j|t{ggsgztk{kg|szkzkqzkzqtkstokms|g{{qikggottokjtoktgs|t
zto|gm}{grmpszkqpstokqzklk
ksk|t{|tkks|ztk~qkgnksgkzgsg}kqnktkjtkg}g{kqknkpsmkqss
{{pszkqt~sk}gskto
tksjm|kjtArenaria leUcadia gzmsstko|jgvpgqm

lpsussfpgpfph{sn
54
Armeria johnseniiPapan. & Kokkini
Plumbaginaceae
Armeria johnseniiPapan. & Kokkini in Willdenowia 12: 221 (1982).
ys ;{{ 7!E t htg= bAD OQNLNSNQITL
ATRSQN NCCIDEMSAKELIMRTKAEIMRAWNRIRKISNQEIRc*TM
+ H RechinGer f {ozt7
jphpf|vozt{okzo|szqt|k}g{tq}t
v{{ggz{ksiqgtkjksgskqtg|qgg
{kgi{go|tro|qo}gkkz}okzqgst|qsto
gjsk |g jt kz}skk jtq}| snt}qt
h{gt{kzt|to CL|o{sjq|tp{mstkjk {kt k kz}ok zqst|qsto gjsk |oqp
mhmtok}{t|qjgqto LLq|g zkqh{gt |ggs khqgsjm g krpkq|
ptkj ptkj qips| W LL k
gz{mrmqgsmzqtgp|gz{gkhqgsjm
zkqnqggkpkq|gsptkj }tqt|tpskrpkq|skz{gkqgkhqgsjmzkqnqg|gk|tqo}k{g}qgh{kgg~jtksg
jtsnmzkqhg{{kstgzsgki{tgsn|hq|t
tj|tgsnps|to LL{o|g~tgstkj |to LL ozkq{ghgststo to igsto|gtoz{|qto kq~k|g|tps|qps
skqpsz{|qthqg~|to LLk|o|{|
pptkjto{p{sg|tozkqztoLL~k{t
|{o|gkkhqgsjkzo~kk}sm{ko|
posof{fpmn qi|g
jpsisfrms|spftst itot
zgq|tohtztoztogsgzkgksggkr
CichoriUm spinosUm Silene sedoides RTBRO sedoides
CentaUrea raphaninaRTBROmixtaAnthyllis hermanniae
AlliUm GUttatUmRTBROsardoUmSedUm litoreUmPhleUm
arenariUmDesmaZeria marinaReichardia picroides|
fqrs{}zfArmeria johnseniiksgoiiks
kmsA canescens(NRS "NIRRsgtqkskjtkki{mzt|{ttq}gztokrgz{skgms|gkiktg~kjsjtq}|}{{gkkskhqgsjm
zkqnqg|ggkrpkq|hq|gztoksghqg~kqg
gzggh{gkpkq|ksggmgs|kqg|ts
~gqg|mq|psjtkjswtmA johnseniijg|qskgk|t{ggzmsA canescensgztokz}ok
zqgst|qsto gjsk to gsnt}qto h{gt
|g g }{{g g krpkq| hq|g k ~gqg|mq|
qgsm zqt g p gz{mrm ts |qkqt |{o|gkq~kst|g|tps|qpsskqps|g
tohqg~kqtoztj|topsg~ojpszmmA
johnsenii gsgzkg k zgq|k zkqt~ ks m A
canescensksgtqkskjt|ggsgzkgosnp
k okqg kig{kqg ps L qi|g
|qkqktq}t{ti|ttmkkmsA johnsenii
zgqtolkkzmmA icarica%DLNMDRNMksjm|kjtpstqps{g|gnqgm|gqg
}psgkto0AOAMICNKANT+NJJIMI g
o{{tigzmsstgolkgkmsA johnseniik
{to to tq}t{ti|t ~gqg|qk k| gz ts

spstgzmskoqkqmzkqt~tog|qpmqto
g}mqghtqt t|qtmhtg
sysArmeria johnseniigsgzkgkzgq|g
hq~g |g zkqjk nk t ikp{ti| ozqpg
mszkqt~krz{pmtokjtogztk{kg|oqp
gz |qog{{|t ~t{nto }gslkg k okqt L k zt{ |q gzgm gz ms g|
kk{kzkqzkgtgtokjtosg
j~tsgggitsjgng{gstskqtgzts~oq |og zto kz|qgk m n{gg m zkqt~
qi|g
zkqkqt oztz{mnot m A johnsenii
zgqgmqnm|gs ms zgqg{g| lsm k nk zto
~gqg|mqltsg gz om{ g{gmg m tztg
|gnjgkqgg}{rkskiggzkqkqg}o|
kjm t ztt }ot|{om nk go ksg
~gm{|gosnpjksrkzkqstktqsk
zkqzkzgqgmqnm|kjkjomgpstokjto
k iksltok jgz{k }qoisps k CentaUrea
spinosa RTBRO spinosa |g SarcopoteriUm spinosUm
{{gkjmztoo~sostjktosmsA johnseniito
105
kpiqg}|krz{pmtoksjm|tkjtoArmeria johnsenii
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106
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54
Arum cyrenaicum Hruby
Araceae
Arum cyrenaicum Hruby in Bull. Soc. Bot. Genve 4: 159 (1912).
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108
54
Arum purpureospathum P. C. Boyce
Araceae
Arum purpureospathum P. C. Boyce in Aroideana 10(1): 8 (1987).
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110
Asperula baenitzii Heldr. ex Boiss.

Rubiaceae
Asperula baenitziiHeldr. ex Boiss., Fl. Orient. Suppl.: 280 (1888).
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Asperula brachyphyllaTrigas & Iatro
54
Rubiaceae
Asperula brachyphyllaTrigas & Iatro in Phyton (Horn) 43(1): 30 (2003).
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zt{ok SAWA zp ksg g CerastiUm candidissimUm
Scabiosa arGentea Sideritis eUboea CentaUrea pichleri
RTBRO pichleri Silene mUlticaUlis RTBRO sporadUm
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fqrs{ }zf AsperUla brachyphylla gs|k
mRECSCynanchicae|gksgksoiikskmsA
abbreViata(AKCRX 2ECHFksjm|kjtmrto
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sg}pkszqstq~tgqgq~p}ozz{ts mgs| ksg t tq}t{ti| jg}tq ps
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oiisk m k ms A nitida 3L RK wt jg}tqtztkg gz gos krgg m zkqkqt zo|s
k}s m ps zt q~ps h{gs ps hqg~kqps|g~kjskonokss}{{pskhqg~kqmog{jm|tqo}pshqg|psztojks}qtosq~kg
zkqnqto|gps|qtqps~lt|gqzpstlk tq}t{ti| zkqkqt k ms A nitida RTBRO
jpsisfrms|spftstpgto{to
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zkqt~osgg|oqpgzt~th|tkzjt|t|g~{pq|t~{nto|gzt{stqgqg
tO(tokj}to|ogskgkgr|g
AsperUla brachyphyllaksgsgoz|~g}ot
qsggzgkjmztozgqgmqnm|gssgostjktos
ms A brachyphylla k gos ts htzt ksg g
kr ThymUs lonGicaUlis RTBRO chaUbardii HypericUm
olympicUm AnthoxanthUm odoratUm &estUca jeanpertii
HieraciUm sartorianUm-ycelis mUralisAUbrieta deltoidea
TrifoliUm UniorUm,eontodon GraecUsSolidaGo VirGaUrea
RTBRO VirGaUrea Arenaria licaUlis RTBRO Graeca
Anthemis cretica RTBRO cretica | tg to kjto
gsgztsgk|qtjkkhqg~jkzkqt~
kpiqg}|krz{pmtoksjm|tkjtoAsperUla brachyphylla

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kqk{{jg|gmsjto3CHMBECJ 4ELERX
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hqkk gz|qmsk z{gi m |tqo} tjg
|gk|ggqnk{ikk|gtsjkgpst
jk{tkgrps|tqo}stjg|gqt}m{g ~k kstznk g |q tjg }os ks
ms |tqo} qt}m {g ~tos zgqgmqmnk jt
|qtjk|gjzgqgtgztoost{|gsq~tsgk |gtost{|z{mno
tokjtojksozkqhgskgtg
AsperUla brachyphylla gsgzkg |g |qt {it
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tmgs|kqtgtoz{mnottkjtptoz|kgkh|mmktqsknkto|gk
gos g|qh ms kzjqgm ksg zngs sg t}k{kg
mgszorm|gg|kspsh{gsgtgztogsg-
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|gnsgmsjmsggzgmgs|kqggt{|
zq|gm~qgksk||qkkm|gg|kozt{{sg|m gsktikssmqs kz|gm ps gt{|s zq|ps
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tqt~m~kkz{kiksgzkq{m}nkt|otps
t|t{ti|mgs|szkqt~smv
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pfypsff{ki~tmh|mmtqt
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brachyphyllazp|gkzt{{{{gksjm||gzsg}o|kjmmzkqt~zz{tszqtk||~pqtnmmpsgt{|szq|ps|ggsg{o|k{kps
zkqhg{{ts|skzzkpsksggzgqgmkksqikk
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}o|t z{mnot zt{{s zsps |g ksjm|s
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A brachyphyllatkjtngztqtkkzmsg|g{{kqimnkktgs|tztop}ohqg~|mzps
tksjm|kjtAsperUla brachyphylla gztqt~mmhtgvpqi|g
115
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Asperula elonea Iatro & T. Georgiadis
+"
Rubiaceae
Asperula elonea Iatro & T. Georgiadisin Willdenowia 14: 55 (1984).
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DECKIUIBTR NQIEMSAKIBTR LNMSIR 0AQMNM OQNOE LNMARSEQITL%KNMAIMRAWNRIRCAKCAQEIRCLc*TK
Iatro & +okkinos /{ozt50!
posof{fpm2n gqt
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|tm ~kqqps zto g jg~ltos ~k kstznk g
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kzzkjknk|gkz{gikki{m|{mqsk}tqgztk{khg|~{pqj|t~kt}qoigs|s t|tomps ks gsgzkg |g{ k zo|s
gqg ktikg| g|g h{mm t okqt |ogskggz Ltom{kqtmktt
tztt~kzgqgmqmnktkjtksgmjo|z{gim
|tqo} tq{g to qto tsthts L ks
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tkzzkjtmn{gg
kgq|knkmAsperUla eloneaksgtkz|qgts
}o|kjtzgkjmztoosozq~tostohtzto
mgsg}qtokksjk||gAbies cephalonica*UniperUs
oxycedrUs 1UercUs coccifera CotinUs coGGyGria ArbUtUs
Unedo Phillyrea latifolia Acer semperVirens 'enista
acanthoclada'lobUlaria alypUmCistUs creticUsHypericUm
empetrifoliUmRTBROempetrifoliUmBUpleUrUm frUticosUm
Calicotome Villosa CoridothymUs capitatUs %rica arborea
% manipUlioraThymelaea tartonraira
jpsisfrms|spfztstst~qt{tto{tog{{|oqptzqtjk|gzksnkqttoto{to
fqrs{ }zf ( AsperUla elonea gs|k ms

RECS Thliphthisa 3CHMBECJ 4ELERX %HQEMDNQFEQ
jphpf|t{okmngstro{jmmhmk
zt{oqng qng gstqntksg jg|{gjsg {kg
k{~m|topCLv{{gktjkps
g}ksg p gstqnpsg jgkps W
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{ti~tkj h{k}gqjp snm ~kjs ~g k}sm
~tgstkj |to LL zqgspz p |ggstjm krpkq| jm kpkq| p{sg LL
{tht qips|t {ti~tkjk |to LL k g|qqips| smgtkj|gzkstkrqmg|to
LLsnqkk{{kz|tk{gst|to
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zkqt~ to qspsg m ~gqjqg to kpsjto
t |g sg |q z{mno |ts t ~pq oq
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qkosg m zkqt~ m k{tztssto +AKONTSYAJIR #NMRSAMSIMIDIR g{{ |g skkqk zgqgmqk htnmgs ms |g{kqm ism m krz{pm to kjto elonea krgz{skg hqkg ~q
z{gi m |tqo} tq|t to qto qspsg
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ksttsg~toszgqgmqmnk~qkqgjo| z{gi to qto qspsg jo|kqg {t t
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sysAsperUla elonea k}gslkgkki{mzt|{ghtzpszstkpkghkt{n|kj}m
kpiqg}|krz{pmtoksjm|tkjtoAsperUla elonea
116
sg sg koj|qt ksjm| kjt m k{tztssto zto ttlk k ms A riGida 3L m tztg
ksgksjm|kjtmqm|gnkzmkmsA
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gk{qlt|gm)ASQN'ENQGIADIR zg
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ki{g snm m elonea |to LL k g g|q krgqgg |to LL ps zk{ps ksg zt{ ~gqg|mq| stt k tqsg }o
zpigzgqjkigkgogzmstztnkgmis
tooqtzgqgmqnm|gssnmkgosnghqg~kg
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k}smgsg}jk
ffl yzlmnr ost{| gqn ps
gpsmAsperUla eloneaksgj|t{tsgk|mnk
gsq~kgpkgq|kjk|jk~{jkgsnkgk
zkogs ~q zq}gg kpqtk g|qhk
kqkztqksgjkrtostost{|z{mno
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mkgrps~pqst{nqg|gk{kkgrto
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|tqo} zgn zsp gz ts oq iqp gz t ~pq
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jk|jk~{jpstg|gkkq|nkgztk{ts
t|qt~{pqj|t~kttohtzto
kjt p 4qp 65 p g sg jkjtsg jks osmitqtsozqmjgqmmtojto~gqg|mqt

gsgissmm ps z{mnos ksg |g{ ig ~tos o~s zgqgmqmnk }o jg}qps m{|s gsnqzsm
jqgmqmgksgstsmktqstohtztoto
kjtoozq~tospgq|kt{{tkk{~m~kjs mjks| gsnqzsm kzjqgm oiiqg}k jks
zgqgqmgs t~kg mgs| h|mm to kjto
k |ztt oztz{mno zgq ms stsm |mstqt}|
jqgmqmgk{mmzkqt~krz{ptozm
}gskgtoztz{mnotgsg|ztos|gstztm|
kgzzoq|gi
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thgqt {it zto sg osmitqts ig ms sgrm m
AsperUla elonea k tztgjztk |gmitqg kg gzk{tosps kjs }psg k ms )5#. k ms
zqtznkmjksngzqgigtztmntsjqg|g{{gitohtztomgztogkps|sjspsm
kqkmoisoztz{mnoskzt{oqngtg|gt
ost{|gqnps}osztogiiltosrkzkqsts
g kzqztos ms kg|smm to kjto ms
|gmitqggm{tsjsto,2 +AKONTSYAJIR#NMRSAMSIMIDIR stt m A elonea zgqgsk sg
ksjm|kjtk|q~k|krz{pmigttztt
ngzqzksg{m}nts{ggqgksgosk~nk
gzq|tzgmjgqmtot{{ts
pfypsfftmgs|kqtqtzqtggmAsperUla eloneaksgmjgqmmps}o|shtzpsmmsmkqsto|ggm|gmgzt}oi
ksspsgsnqpzspskzkhkpsk{{ts|

g{gkqg jkjtsg )ASQN A ~gqg|qlgs t
tksjm|kjtAsperUla eloneagzms~gqjqgtokpsjtovpg{ztol|m
117
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Asperula malevonensis Ehrend.

& Schnbeck-Temesy
54
Rubiaceae
Asperula malevonensis Ehrend. & Schnbeck-Temesy in Strid & Tan (eds), Mount. Fl. Greece 2: 294 (1991).
3ys ;{{ 0E k{tzssmt tt q|gjg
g|psg=bIMCACTLIMELNMSIR-AKEUN,ACNMIAE
OEDc *TK Orphanidis {ozt 75
)ozt!4(575
nrrnf AsperUla boissieri (EKDQ EW "NIRR UAQ
transiens(AKCRXIMRSEQQ"NS:
jphpf|gm{zt{okzgkro{jmhm|g
{kztoziktoh{gtozqiktg~mglts
zo|s zmg k{~m to CL k kqgips|
jgt g|qgg k {ik jg|{gjk gsnt}qt
h{gt gskq~kst k zo|s q~pm q~k |to
LL zsg ~kjs {kt v{{g iqgtkj
{ti~tkjkzo|stztsj{topskinto
W LLkonoksp|gzo{p{kgq~p
gkzzkjgk{g}qoskqgsg|qzkjgq~k
|topLL kog{jkgzt{m||krqmg
|to p LL |g zqtk|h{{tog kgg skqpm
ms|pkz}skgzto|gg{ghskzkqztot
toz{toto}{{togrgsnghqg~kgk t{igsnk kojtztsj{to q|g gs{tig ps }{{ps k{g}q oskspsg m hm to h{k}gqjp
kg|jpgz{mrmk}sm~tgstkjqjsm|to
LLkp{sgzkqztojz{tps{ths{tht
kz|kgh{kkgqg|{mqq~k|gkrgqggknm{tkjkzqtk|ht{wtn|mnm{tkjqg
|gqzjg ~lt|qzg ptkj sk}qtkj LL
i{go|tzqsgp|ggstzqsgkzo|snm{jgkqgmq~mkstkk|ggs|m{jk
gozqikgggto}ot{ghstosmstq}|qtzmgzsgzgqgmqnm|gs}osgzqtk|h{{tos gz ts g|gsnp AstraGalUs anGUstifoliUs RTBRO

anGUstifoliUsktstzttsosozq~tos
t kjt }kg k okqt L |g t
jt oztz{mnot gz~tos kgr to zkqzto
L oztz{mnot m |tqo} tg qg ~tos
hqkt zqtgsgt{ ks t oztz{mnot m ki{mtq{g~tosjg}qtozqtgsgt{tg
zkqkqt osnm kjm zto k}gsltsg gl k ms
A maleVonensis ksg g Anthemis laconica AstraGalUs
anGUstifoliUsRTBROanGUstifoliUsCentaUrea athoa RTBRO
parnonia'aliUm thymifoliUm Sideridis clandestina RTBRO
clandestina Viola parnonia|
fqrs{}zfAsperUla maleVonensis oiikskk
k ms A nitida 3L kjt m |q g jg}qk
pgzgot~oq|gz{gkgtskqtps
kskqps}{{psmszgqtogq~pmmk}sm|gnm{tkjkzqtk|ht{pskrgqmps%HQEMDNQFEQ 3CHMBECJ 4ELERX z ms kzm
oiiks||gksjm|m{{jg pUlVinaris (EKDQ
EW "NIRR jg}qk t g|qkqt og{jk gzt{m||
krqmg ps }{{ps ms {ikqt q~p k}sm
|gtshqg~kqtp{sgtosntoks}gskgsg
ozq~kmgs|jg}tqt~qgmk}smkgrpsjtk{kogpskjsg}tm pUlVinaris ksg
zt|{tq}mpzqttsoi|k|qst~gqg|qg
posof{fpmispt
jpsis frms|spf z g g tosto ~q g

gto{togqg|gqzjgk}gsltsggqits
t{t|gtsitot
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|tqo}s to qto qspsg ms om{kqm ki{m
tq{g|gmikts|tgqg
sys AsperUla maleVonensis }kg k gst|
zkqt~ m ozg{z| lsm k ghkt{n| ~g{|jk ozqpg zto p oi|qgk gq|k ist
~g zz{ts ~k hqknk k ksk{ zkqjk ozqpg zto }kg k qpi jghqpsps |qs
hq~ps|gkn{g|kkj}tokgrgnkqtztmspsikts|s{npsg}o~mgltosojk
nk zto k}gsltsg t oztz{mnot ~tos |q
p mjks| |{m ks jks ~tos zgqgmqmnk tg
k|gg|qo}toghkt{n|t|qmsthq~to
kpiqg}|krz{pmtoksjm|tkjtoAsperUla maleVonensis
118
ffl yzlmnr }psg k zgqgmqk

t zkjt g tg m AsperUla maleVonensis ms |tqo}tgqgjks}gskgsgrkzkqstsg
jkzgqsgmhqkgz{koqm|tqo}|g|oqp
ms|tqo}tiqgzqtmki{mtq{gmki{m
tq{gozq~tostjk}oskj}tqgmkgm
zkqt~m|tqo}zsgpk|qgqngpsost{|gqnps}oskjozt{tilkg
zkqzto k g zkqkqg }o }gskg
sgksggq|kki{mm{|gksgskgqtgksgjgkqgzkqtqsgsggrtmkptkzmt
ikits|tqo}toqspsgzto~toskqkosmnk
jkrtj|zpmggsqqg~m|gmxgqzgqki{k ttmk to k okqt tq}t{tig ozqpggsnqzskjqgmqmk|{zkmki{m
tq{gjks}{trksts|zttoztz{mnotokjto
AsperUla maleVonensis ksgsgzst|gzt{tz|kjtztoztqksgksg~nkms|gmitqgps4qps
65 m )5#. k hm t |qqt $ ms zkqt~ zto }kg gsgzkg stsm |mstqt}|
jqgmqmgpig|q~k|~qts|jmg
kgrpsmssto{to|gkzkhqto|gstgz
|tzjgzqthpsglgjksnitostkjtpztjtzgtstooztz{mnotto|gkgotsqzt
zgqkztjltos jgkqg ms gszorm skgqs gqh{psgtgmA maleVonensiszto}tsgk~kjs
kzzkjkkz}skk}gskgsgzqtgktsg{ikqt
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gtztg}tsggskgk{ntokqpi|qs
hq~psgzngsg{{gitoqztoh|mmkms
kgipi gis ms zkqt~ ng kzmqkk gqsm|
zqttz| kzhpm to kjto kzkj t gik ~tos

k|t{mzqhgmkhqg~jknkjgkqg|q
gqnskgqsgpsmA maleVonensis ngzqzksg
jkqkosmnkzkqgqptin sitUt|gkzkqgg
}qpmzkqps{tksgg|misptzpng
kzmqktostooztz{mnottokjtotjg}gskskg{{gito|{gtzto~tospgzt{kgm
|qkqm~tszpm|gm|qkqmjq|kg~tst|{ompsom{s|tqo}stoqspsg
pf ypsff / mgs| gqn ps zgsps}o|skjszto}tsgggskqgggto
qtoqspsgngzqzksgzqtgkonkgzksjk~ksk joksk ksqikk zp t g{{gi ts qzt
h|mm m zkqt~ t jgstrk kzz{ts zkqzgm|s tstzgs giqt|s jqps m ozkqht{|
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k{{tqzttkjt
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ex sitUzqtggmp|ttzqtztnkm
k{mmht{tigtokjto|gn|gpsosnm|s
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sgkzk|gnkkzt{{zsgkjmmk{{ms|~{pqjgkzo~msm|g{{qikgtooi|k|qstokjto
|g{{psozg{z|s}osk~gm{okqgzqtztnkmsgsgzgqgipipsosnm|sztokz|qgts
gtqkst|toggk}gmmsgskzm
mgormsmnkqt|qggtonqto
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Asperula nauAraga Ehrend. & Guterm.
$-
Rubiaceae
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54
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Astragalus drupaceus Orph. ex Boiss., Diagn. Pl. Orient. ser. 2, 2:32 (1856).
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54
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Astragalus laconicus Iatro & Kit Tan in Willdenowia 29:56 (1999).
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osnp sg g|q ~qts| jmg ig sg ksm{|pnk|goz|kgk|sjstoo~gk|ggqt} zto ztqk sg gztqtos ms gsnt}tqg to g
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m{|gjgkqgtoz{mnotzto}tsgk~gm{kqgokqg
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pf ypsff sg mgs| sg k{g~tztmntstgsnqzskkzkhkzkqt~zto}kgmAthamanta arachnoidea{{pkgqmto{t~qg|gtsthtsmzkqt~mtsm{psg|gm
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tog| |g zqg|| grg zt{{ zqtiqgg zkqhg{{ts|k|zgjkompsgnmsztomos~kg

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zqtk|||g{{qikg}ostokjtoktgs|t
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Aubrieta erubescens Griseb.
54
Cruciferae
Aubrieta erubescens Griseb., Spicil. Fl. Rumel. 1: 268 (1843).
3ys ;{{ .!E its qt= bIM RAWIR LAQLNQEIR
L!SHTRCNOINREOQ0AMAIAAKSCNMRNCIASACTL
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lkittjspsg|qggsnmkgqghqozg{g|to LLg{g|to LL{ko|
qjsg {t |to LL gqz |qg W
LLk|tzkqzto }tqtz{tto
ozkst k kz}skg zto |g{zkg osnp
stgz|qg|stq}kq~ktqsk}tq
kzmkk{g}qj~g{pq~k{g|qkqkgz
g|stq}k
posof{fpm2n = 3SQIDA
jpsisfrms|spf{mzq{to~q{mtostoztoztqksgzgqggnk~qgq~oitto
johpf|{ jqyzol AUbrieta erUbescens ksg
ksjm| kjt to npsg ms ~kqsmt to ito
qto
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npsgzspgzmskqtskimgqgsg
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oztz{mno zto gztk{kg gz zkqzto tg
hq|kgkjg}tqk|mkgskz{gito
npsgzspgzmstztnkgkqgkokqt
Lkzkqjmgstiggg|gh{mm
qkoztz{mnotmAUbrieta erUbescens ksgjg|tqzstmshqg~jm|tqo}tonpsg|gk
zkqjmj|ksgjgsg|qgzgsnqzskjqgmqmkh|mmjkskzqzkgm~kqsmt|g
jksgztk{kgzk{igtkjtgosttm
}p ng ztqtk sg gztk{k josm| gzk{ ig
tojtoztz{mnotg~gm{kqgokqg
tkjtzkq{ghskgtogg{itom)5#.
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g{titpsgi|tpzk{tospsps|gvos ts g{tit to ohto{to m oqzm t
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L gl k g PrUnUs prostrata PterocephalUs perennis
RTBROperennis Daphne oleoides Berberis cretica Crepis
athoaAlyssoides UtricUlata Isatis tinctoria RTBRO athoa
Thlaspi perfoliatUm SedUm ochroleUcUm SemperViVUm
marmoreUm |zmgsgzkgkzkqjmj|ksg |t jto |g g|g h{mm k ~gm{kqg
okqg L glkg*UniperUs oxycedrUs
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0HISNR
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togztk{tsksgjg|tqzsgk|qtjk
osnpps }oskig{kqtoztz{mno
kpiqg}|krz{pmtoksjm|tkjtoAUbrieta erUbescens

gtkjtmjgztqps{ikqpsgzoztz{mnos |gn |g to |qt ozt{tilksto ost{|tgqntpsksm{|psgpsztojksozkqhgstos

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pf ypsff ~qkqgjksozq~toszqg|| qg zqtgg ig ms AUbrieta erUbescens
qg| g{{gi ts htzt to kjto jgkqg
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Bellevalia brevipedicellata Turrill
$-
Hyacinthaceae
Bellevalia brevipedicellataTurrill in Kew Bull. 1940: 264 (1940).
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+QISI ABNUE 0AKAINCHNQA AS +EOHAKA RSNMX KILERSNME
HIKKR VISH BiarUm daVisiic -AQ DaVis K
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LL zkqnqg {kg |ogtkj {gt
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|to LLqnthqg~kqtgztzkqsnt
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koqp ptkjk k k{g}q qjst ~qg |g k g
|ksq| zqsm iqpm snqk jk grgsng
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k{{ktkjp}gq|
posof{fpmn = 4YAMNTDAJIRAK
"AQEJAAK
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|g t ~g ps |gqzs m ptkj p k{{kz|
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skoqk|gksgg|qkqg|gztksgzgom
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m}gq|
fflyzlmnrstkqoztz{mnot
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t|gnsg
zkqtqsm zkqt~ krz{pm m BelleValia
breVipedicellatagzk{kggz|ggqt}tohtzto
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qit The Red Data Book of the Rare and Threatened
Plants of 'reecepqp4TQKAMDA pk
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tartonraira RTBROarGentea4TQKAMD
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msRECSINPatenssgoiiksmB sitiaca +XOQ
4YAMNTDksg|mksjm|tkjtomqmkstzstosttstgntoqm ttztt
zg{gkqgnkpqtsgsgkqgz{tkjtq}mB
breVipedicellatakn W~qptgg(tq}t{tig m grgsng |g ps gsnps t ms B
sitiaca t |g ms " breVipedicellata ksg zgqtg
stt t jgkqk zkqt~ krz{p to t j-
kpiqg}|krz{pmmksjm|BelleValia breVipedicellata

rkzkqsg
pf ypsff ks ~tos {m}nk qg zqtgg ig t kjt ~q kqg gs |g m st gjt
~kzkq{m}nkzkqt~to|tov
htztngzqzksgzqtgkontsgzmsgskr{ki|mtoq|gszorm|gmso{{tiht{hsqt-
|kstosgkrg}g{nkmkzhpmtokjtotg
hq|tsgjmk|g{{qikgtszkqgg|tgs|
zttoqigmqtotgs|togskzmtogqsgq{{m{gosgmosqmmzkqpsk
qzklkksk|t{|t
ylyfp{f
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}p*"IEMUEMTjkr}pgqm

Bellevalia edirnensis zhatay

& B. Mathew
$-
Hyacinthaceae
Bellevalia edirnensiszhatay & B. Mathew in Bot. J. Linn. Soc. 107(1): 90 (1991).
ys;oqpzg|toq|g=b%DIQMESN5YTMJOQJL
FQNL)BQIJ4EOEQNADTMCSINMAKSLc!OQIK
'ler DalGi {ozt)34%
jphpf| t{ok ht{hjk kjt t{h ptkj }gqtkjjgqto LL~sg~qgtk}qt gqtov{{g ksgs{ti~tkj
g|qkqg gz ts h{g |to CL z{to
LL |ogtkj zkqnqg khqgsjm k{g}qq~p{gt to CLgrgsng
k snmzo|sk{{ktkjp|o{sjq|tj|t|to LLqntmsgq~|gmos~kg trtkjk q|g |q {ko| }qtsg jt {thtsqgsnmkstkjmkqsnt|to
LL p{mstkj p |pjpstkj {ko| k{g}q
zqst m hm to k {tht k{g}q |qkqto
|to gz ts gsn| p{sg k{{kz|t ptkjk
snqk|qstgrgsng|gms|gqzt}tqg|o{sjq| g{hjk |g koqp ptkjk |to
LLzqggkoqpptkj k{{ktkj
posof{fpmn = W= YHASAXAK
"AQEJAAK
jpsis frms|spf snlk |oqp |g ms jq|kgtogtog{{zkqgg||ztgtghqnm|gskgsnt}tqgg{mzq{to
johpf|{jqyzol(BelleValia edirnensisk}gslkgmsoqpzg|toq|ggzazto~kzkqiqg}k

to{{~nm|gstgtoistogzgzkqnqg
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mB kUrdistanica&EIMBQTMmtztgkrgz{skggzt
q||gmstqjgsgpmssgt{ggjtkjm
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~qg to rtsg m grgsng t tztt ms B edirnensisksgzqst|g~jmzpgomB
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B kUrdistanica ksgjkksmsB edirnensis|qstzgkjmztogzgstsms{{jgmB edirnensisksgoiiksmB romana , 3VEES|qk
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edirnensisksmB romana~gqg|mqlkggzg{kg
zkqnqgps}{{ps{thttokig{kqto|togztsgsn|p{sg|ggzjkgsnqk
fflyzlmnr~qkqg~khqknkstssgz{mnotokjtogottsthqto
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tg kstznm|gs |g ks ps |g{{kqiks g
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. }psgkms|ggrmm)5#. |gn
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krgz{skg ksg |g{{kqitksm |g t kjt j~kg
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Bellevalia sitiaca Kypr. & Tzanoud.
54
Hyacinthaceae
Bellevalia sitiaca Kypr. & Tzanoud. in Bot. Helv. 109(1): 85 (1999).
ys ;{{ t qm t gnto=
bOQNOE UICTL 3ISIA IM RAWNRIR CAKCAQEIR OQNLNMSNQII
0HAMEQNLEMIc!OQTZanoUdakis{ozt
50!
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LLkk}gskzgq{{m{kskoqkzkqnqg{kg
{gt |to CLgrgsngk
snm|o{sjq|zo|stj|thqg~kqtgzt
zkqsnt |to zkqzto LL qnt kgq snm
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|o{sjq| p p{mstkj |pjpstkj {ko| k
{tht|to }tqt|ttogsn|tp{sgkoqpptkjkkk{g}qqjst~qg|gg
|ksq|zqsmiqgsnqkjkgrgsng|g
ms |gqzt}tqg |o{sjq| k ztj|to hqg~kqto
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posof{fpmn W4YAMNTDAJIR
AKzgq|g"AQEJAAK
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wt m gzttspsm ikpiqg}| krz{pm k

osjog k tqsk tq}t{ti| jg}tq ps
jt kjs tjimgs ms zkqiqg} m B sitiaca p
rk~pqt kjto mgs|kqk tq}t{ti|
jg}tq ps jt kjs kstzltsg g }{{g g
tztg ms B sitiaca ~gqg|mqltsg gz k}gsk
zgq{{m{k skoqk |g ksg kig{kqg k |t
|g kskqg gz go m B breVipedicellata |gn
|g t ~g ps |gqzs m zqm t tztt ksg
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k{{kz|qt}ptqg||ogqtiksk|k{m
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Bupleurum aira Snogerup in Bot. Notiser 115: 366 (l962).
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160
54
Bupleurum capillare Boiss. & Heldr.
Umbelliferae
Bupleurum capillare Boiss. & Heldr. in Diagn. Pl. Orient. ser. 2, 2: 82 (1856).
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162
Bupleurum kakiskalae Greuter
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Umbelliferae
Bupleurum kakiskalae Greuter in Bauhinia 3: 250 (1967).
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Buxbaumiaceae
Buxbaumia viridis (Moug. ex Lam. & DC.) Brid. ex Moug. & Nestl., Stirp. Crypt., Vogeso-Rhen., fasc. 8: n. 724
(1823).
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vp4TQKAMD

Campanula asperuloides
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54
Campanulaceae
Campanula asperuloides JDNN*MKC IBG@MDI IBG@M+M<IOG
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nrrnf TracheliUm asperUloides "NIRR /QOH IM
"NIRRIEQ$IAGM0K/QIEMS3EQ
T tayGeteUm 1TYEK #NMSAMDQ IM 4AWNM

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$ALBNKDSpTracheliUm tayGeteUm
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latifolia |g Amelanchier parViora RTBRO chelmea k
zqtk|ht{ ps hq~ps z{ gl k {{g zsg
tz| ksjm| zp g -elica rectiora -inUartia
pichleriRhamnUs sibthorpiana ThalictrUm orientale|{z
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grst| oiiks k ms CampanUla t TracheliUm
gztjk~nm|kzpjksksggq|tspjg|qksg
j|gtg t kzzkjt to jgkqto isto |g p k|
ttokspgnm|kmsCampanUla tT tayGeteUm
ttzttzkqiq}m|kgztqtgiktmk{tztssto1TYEK#NMSAMSQINONTKNR nkpqkgjg}qkgzmsC asperUloides t~k
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kkzzkjtoztkjto'QETSEQ g{{mos~kg
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sys t kjt ksg ~gt}o| |g }kg k
gz|qmstoghkt{n|thq~tok|qt|g{tzgi
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jtoAbies cephalonica PinUs niGrag|mknk
~pq jg| h{mm |k ztqk sg }kg gl k
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ts qspsg ~k hqknk k hq~to |g |qmst
hqkto m htqktjo|t zqtgsgt{t gl k
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methanea -inUartia pichleri Stachys chrysantha |{z
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175
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176
Campanula garganica Ten.

subsp. acarnanica (Damboldt) Damboldt
54
Campanulaceae
Campanula garganica Ten. subsp. acarnanica (Damboldt) Damboldt in Bot. Jahrb. Syst. 84: 358 (1965).
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177
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178
Campanula garganica Ten.

subsp. cephallenica (Feer) Hayek
54
Campanulaceae
Campanula garganica Ten. subsp. cephallenica(Feer) Hayek in Repert. Spec. Nov. Regni Veg. Beih. 31(2):
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182
54
Campanula hierapetrae Rech. f.
Campanulaceae
Campanula hierapetrae Rech. f. in Oesterr. Bot. Z. 84: 170 (1935).
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54
Campanula merxmuelleri Phitos
Campanulaceae
Campanula merxmuelleri Phitos in Mitt. Bot. Mnchen 5: 121 (1963).
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186
54
Campanula pangea Hartvig
Campanulaceae
Campanula pangea Hartvig in Willdenowia 28: 65 (1998).
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Campanula papillosa Halcsy
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Campanulaceae
Campanula papillosa Halcsy in Bull. Sances Soc. Sci. Nancy ser. 3,9: 387 (1908).
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54
Campanulaceae
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Campanula samothracica (Degen) Greuter

& Burdet subsp. samothracica
54
Campanulaceae
Campanula samothracica (Degen) Greuter & Burdet subsp. samothracica in Willdenowia 11: 40 (1981).
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Campanula saxatilis L. subsp. saxatilis
54
Campanulaceae
Campanula saxatilis L., Sp. Pl. 1: 167 (1753) subsp.saxatilis
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54
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208
54
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Compositae
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210
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(Halcsy & Hayek) Dostl
54
Compositae
Centaurea attica Nyman subsp. megarensis (Halcsy & Hayek) Dostl in Bot. J. Linn. Soc. 71: 198 (1976).
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Centaurea carystea Trigas & Constantin.
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Centaurea carysteaTrigas & Constantin. in Bot. J. Linn. Soc. 158(4): 764 (2008).
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Compositae
Centaurea cithaeronea Phitos & Constantin. in Fl. Medit. 3: 273 (1993).
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Centaurea heldreichii Halcsy in Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 61: 315 (1894).
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Centaurea leonidia Kalpoutz. & Constantin. in Bot. J. Linn. Soc. 146: 375-383 (2004).
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T. Georgiadis, G. Dimitrellos & Routsi
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Centaurea paxorum Phitos & T. Georgiadis
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Centaurea peucedaniAolia Boiss. & Orph.
54
Compositae
Centaurea peucedaniAolia Boiss. & Orph. in Boissier, Fl. Orient. 3: 647 (1875).
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b0EMIMRTKA !SHNA OQNOE $AOHME /QOH c kqiq}m|k
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54
Centaurea poculatoris Greuter
Compositae
Centaurea poculatoris Greuter in Bauhinia 3: 252 (1967).
FATCIBTR IMFQA OAGTL !RFMDNR L IM QTOITL CAKCAQEAQTL OQAEQTOSAQTL RRTQIR RNKE IKKTRSQIBTR
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gqg q~pg ms sp kz}skg |g ms pqgsm
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54
Centaurea princeps Boiss. & Heldr.
Compositae
Centaurea princeps Boiss. & Heldr. in Boissier, Diagn. Pl. Orient., ser. 2, 6: 111 (1859).
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Centaurea pseudocadmea Wagenitz
54
Compositae
Centaurea pseudocadmea 2<B@IDOUDIII)<OPMCDNO(PN2D@I
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54
Centaurea pumilio L.
Compositae
Centaurea pumilio L., Cent. Pl. 1: 30 (1755).
ys qtgnk p jgmqmt gz ts 4TQKAMD IM
4AWNM ;iozt {krsjqkg= bIM AQEMNRIRLAQISILIRAD3IDI 'ABEQc!OQBornmller
"
nrrnf AeGialophila pUmilio , "NIRR $IAGM 0K
/QIEMS
CrocodyliUm pUmilio , . 'AQCIA 3TRAMMA IM 0K
3XRS%UNK
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qjg|k k q~pg gqg~stkj kqjk {ko|}gt
osnp {oqtkj zkqtkj k ki{to g|qgto
{thtzgspgjgqkgtzkptkj |gqj~mg
k{{kz|k}{gosnp zgspzkqkqg
p khqg~ztj|tkqh{mgptkjkjkqt p LL kgg hq|g zkqh{gt ptkj
k og{jk krqmg |gg{itsg k ~oq |gsng
|to LLrpkq|gsnjgjm{itg|qkqg
gzgkpkq|kq|gzg|ksq|o~sztq}oqzgsp{ko|~gsgqtzg{tkj|to~q
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maritima %Uphorbia paralias Silene colorata | t

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ammophila
fqrs{}zftstgt{tigmCentaUrea pUmilioozqrkigki{tjmgoi|k~osmktkjt
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"NIRRg|m|gzqt}pz~-EKEAK g{{
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tztmCentaUrea pUmilio gvg{gqsgmqmvpgqm

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250
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tgmCentaUrea pUmiliogzgvg{gqsgmqm
%zsp}pgqm|p}p4TQKAMD
54
Centaurea rechingeri Phitos
Compositae
Centaurea rechingeri Phitos in Ann. Naturhist. Mus. Wien 67: 165 (1964).
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b)MR+XQA0AMAGIAIMQTOERSQIBTRCAKCOQNOELNMARSEQITL0AMAGIAc-APhitos {ozt
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|ga ksg{{gis osnp k{{ktkjk g|qgto
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m hm g|qgg {ti~tkj k}{g ktspsg
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251
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252
Centaurea subciliaris Boiss. & Heldr.

subsp. acarnanicaMatths
54
Compositae
Centaurea subciliaris Boiss. & Heldr. subsp. acarnanicaMatths in Bot. Jahrb. Syst., 95(4): 430 (1976).
3ys;{{3S% kqk{{tp{tg|gqsgsg=b-S"NTLIRSNR-ACCHIE1UercUs ilex 7AKDAKS
Lc *TK Damboldt {ozt "
HEQB$ALBNKDSozt"50!
nrrnf CentaUrea acarnanica -ASSHR 'QETSEQ IM
7IKKDEMNVIA
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p CL {g osnp jg|{gjst jg|{gjk |to CL +gkqg }{{g k tq}
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255
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54
Valerianaceae
Centranthus sieberi Heldr. in Verh. Zool.-Bot. Ges. Wien 40: 295 (1890).
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257
Cephalanthera cucullata Boiss. & Heldr.

Orchidaceae
Cephalanthera cucullata Boiss. & Heldr. in Boissier, Diagn. Pl. Orient. 1, 13: 12 (1854).
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258
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260
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Cephalaria squamiora
(Sieber) Greuter
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Cephalaria squamiora (Sieber) Greuter in Candollea 22: 235 (1967).
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261
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262
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54
Cerastium dominici Favarger
Caryophyllaceae
Cerastium dominici Favarger in Bull. Soc. Neuchteloise Sci. Nat., 99: 82 (1976).
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Cerastium illyricum Ard.

subsp. crinitum (Lonsing) P. D. Sell & Whitehead
54
Caryophyllaceae
Cerastium illyricum Ard. subsp. crinitum (Lonsing) P. D. Sell & Whitehead in Feddes Repert. Spec. Nov. Regni
Veg. 69: 16 (1964).
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fqrs{ }zf t CerastiUm illyricUm ksg zt|{tq}tkjtttzttjg|qskgkqgoztkjmt
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265
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266
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267
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Cerastium runemarkii Mschl & Rech. f.
54
Caryophyllaceae
Cerastium runemarkiiMschl & Rech. f. in sterr. Akad. Wiss., Math.-Naturwiss. Kl. Anz. 1962: 231 (1962).
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Chaerophyllum creticum Boiss. & Heldr.
54
Umbelliferae
Chaerophyllum creticum Boiss. & Heldr. in Boissier, Diagn. Pl. Orient., ser. 1, 10: 51 (1849).
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54
Colchicaceae
Colchicum asteranthumVassiliad. & K. Perss. in Preslia (Praha) 74: 57 (2002).
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+0EQRRNM

Colchicum burttiiMeikle
o
Colchicaceae
Colchicum burttiiMeikle in Bot. Mag. 181: 134, t. 735 (1977).
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281
*@PHL/DPQQML
Colchicum chimonanthum K. Perss.
54
Colchicaceae
Colchicum chimonanthumK. Perss. in Pl. Syst. Evol. 217: 56 (1999).
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Colchicum euboeum (Boiss.) K. Perss.
o
Colchicaceae
Colchicum euboeum(Boiss.) K. Perss. in Candollea 53: 400 (1998).
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285
Colchicum lingulatum Boiss. & Spruner

subsp. lingulatum
o
Colchicaceae
Colchicum lingulatum Boiss. & Spruner in Boissier, Diagn. Pl. Orient. ser. 1, 5: 66 (1844) subsp. lingulatum.
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286
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287
Colchicum parnassicum Sartori,

Orph. & Heldr.
o
Colchicaceae
Colchicum parnassicumSartori, Orph. & Heldr. in Boissier, Diagn. Pl. Orient. ser. 2, 4: 122 (1859).
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288
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jt kjm oi~tsgs kgr to jt t htgs|t
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{{ts gztg|qosg t sg gz t {{t tqg|
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parnassicUm gztqtgqsgvp+
0EQRRNM

Colchicum peloponnesiacum
Rech. f. & P. H. Davis
o-3
Colchicaceae
Colchicum peloponnesiacumRech. f. & P. H. Davis in Rechinger f., sterr. Bot. Z. 95: 427 (1949).
3ys ;{{ 0E k{tzssmt t ~gg= bIM
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*@PHL/DPQQML
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Colchicum pulchellum K. Perss.
o-3
Colchicaceae
Colchicum pulchellumK. Perss. in Willdenowia 18: 30 (1988).
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*@PHL/DPQQML
54
Colchicum rausii K. Perss.
Colchicaceae
Colchicum rausiiK. Perss. in Pl. Syst. Evol. 217: 58 (1999).
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*@PHL/DPQQML
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Colchicum soboliAerum
(Fisch. & C. A. Mey.) Stef.
54
Colchicaceae
Colchicum soboliAerum!DN>C(@T.O@ADI.=JMIGBF<?)<PF
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*@PHL/DPQQML
Colchicum stevenii Kunth
o-3
Colchicaceae
Colchicum steveniiKunth, Enum. Plant. 4: 144 (1843).
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*@PHL/DPQQML
Consolida arenaria Carlstrm

Ranunculaceae
Consolida arenaria Carlstrm in Willdenowia 14 (1): 16 (1984).
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54
Consolida brevicornis (Vis.) So
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54
Consolida tenuissima (Sm.) So
Ranunculaceae
Consolida tenuissima (Sm.) So in sterr. Bot. Z. 71: 241 (1922).
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54
Convolvulus mairei Maire & Petitm.
Convolvulaceae
Convolvulus mairei Maire & Petitm. in Bull. Soc. Sci. Nancy ser. 3(8): 176 (1907).
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sg |ggiqg}ts g skkqg jkjtsg zg{gs z{mnostozgh|mmzkqt~zto}kgt
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|g m ttm to k t C arVensis t |gn ksjk~tsp gskznmt k |g{{qikk |g tgs|t
ztosttmEWRISTjgqmtokkrkj|kosgstg|gztogztk{ksgksjk|sokst
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54
Coriaria myrtiAolia L.
Coriariaceae
Coriaria myrtiAolia L., Sp. Pl. 2: 1037 (1753).
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;g{{g= b(ABISAS -NMROEKIIc ;(EQB ,IMM no
,).. =
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qqksg n{kg |g kqg}qjg k gq|k k}gs gq~|t~kgmspsgn{kg|g|gqzt}{{psg
qqksgzg{gk{knkqgptkjtrgzgqgstsg
g{g k{knkqg zqsg hqg~kqg ps kz{ps
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|gqzt}tqg ~oq qtzjp ms kpkq| z{koq zkq|{ktsg ts |gqz ~q ms pqgsm k{|
gzt~pqlksg ig ms gzt|{o to tsk
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st tsk kr~tsk |ggsktsk itst tski|{ktqnt{tk{knkqtsmgtkjk
t ist kr~tsk |g zt{ gqz oi|qzt gz
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m ktikto }kg k rmq jm k ngssk
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ms{ikqgk}gslkgkjmkngsskkgr
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Corydalis thasia (Stoj. & Kitan.)

Stoj. & Kitan.
54
Fumariaceae
Corydalis thasia.OJE&DO<I.OJE&DO<IDI"J?.JENF0IDQ!DU(<O!<F NO@NOQ$NOJMDE<
(1945).
ys;{{.!E tt=~kzkqiqg}kgz
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gsgztsgkj}tqktztnkkptgzo~gssgjmtoqitosstooztz{mnotk~qts|
zkqtjtzgqg|t{tnmmks
t{{ig}oksnmkqgg~gsgzgqgmqnm|gstozkqkqtogztoisptz{mnot
tokjtokrggmozkqh|mmmtztgzngsssg
ksgozknosmigmjo|t{ggottokjtosgkzk|gnkg|m|gsgkzhkqtkskgsgksg~nk
tkjtzqttzgqsms|gmitqgps~kjszk{tosps }psgkg|qqgm)5#.
krggps|qsjzgqpsz{mnostokms
k{zjgtkii{{tssgmsgzt|{skzqtom{kqthgngzk{tqitThe Red Data Book of Rare and
Threatened Plants of 'reece0HISNRAK tkjt
gsg}qkgpzst2AQE +ALAQIB
,pf ypsff sg j|t{t sg zqtgkonts
{t t |qt |g jg|tqzst oztz{mnot
m Crepis aUricUlifolia wt g qg to ns|t
q|to to }gqgiit m ggq ng ztqtgs sg
kzk|gnts zqt|ksto sg ozkq{htos ~ st
|zttsoztz{mnomC aUricUlifoliag{{|gzt{{s
{{psksjm|smks{ipkoqkqmzkqt~h{zk
BUpleUrUm kakiskalae kgzqtznkgsgjg}g{nk
mkzhpmmC aUricUlifolia tkjt|g{{kqikgex
sitU tszkqgg||zttoqigmqtotgs|
to gskzmto gqs p k gz qsg
~qsggsnt}tqgjkszgq~nmgsqgzqgg|g
g}ogsgzgqitsgsth{gm|
jophffpl
tksjm|kjtCrepis aUricUlifolia|g{{kqimstgztqtxm{tqkmmqmvp

+gqm

54
Crepis heldreichiana (Kuntze) Greuter
Compositae
Crepis heldreichiana (Kuntze) Greuter in Candollea 30: 326 (1975).
ys;{{0E k{tzssmttg|psg=bAD
QTOERQEGINMIRAKOIMAE-S4AXGESI,ACNMIAEKNCNTMICN
+AJNCHINMIDICSNcHeldreich ' "/)3
nrrnf Hieracioides heldreichianUm +TMSYE 2EUIR
'EM0K
Crepis diVaricata "NIRR(EKDQIM"NIRRIEQ$IAGM0K
/QIEMS ;MNM&73CHTKSY=
C tayGetica "ABC IM 5MIU #AKIF 0TBK "NS
nom illeG
jphpf| t{ok kjt k ro{jm hm |g k
zt{{ sk|q }{{g zg{gtqps ks {gt to
osnp ~q CL k zt{oqnk kz|k
j|gzkjg|{gjkv{{ghmksgs{ti~tkj k hgn zkqtkjk ki|t{zk k z{koq|
ggks}{{gh{gtosnp|qiqgtkj~pqki|t{zkkqh{mg|k}g{tojgkps W LL|o{sjq|| pjpstkjk{ko|pzzo|sq~pgrpkq|hq|g|totps
kpkq|sgtztgksgh{k}gqjpms|tqo}|g
kkq|hqg~kkgqkq~kmskrpkq|to
kz}skg {kg ms kpkq| sntj~m |ok{p
k}sm|to~qLL|qsmmskzspkz}skgztq}oq kqonqms|pkz}skg|oqpps
krpkq|sgsnps~gsg|to LL|ggsk
qghjk
fqrs{}zfCrepis heldreichiana|gnksg

sg zt{oz{tkj gztksjm| kjt ~k jgkqt ksjg}qts gz }otikpiqg}| |g |ogqtgrst|
ztmoi|q||gqotomg|k{mgotto
SAWNM|gpsoiiksstokjsC sibthorpiana"NIRR
(EKDQ|gC incana 3L+ALAQI jkrkksg
sgjk|gz{tkjn W kjtttzttzngs
zqtq~kggpkpgztzgqtksjm|jz{tkjkjtC sibthorpiana kn~qptgg
C heldreichiana ~k zt{{ tq}t{ti| "ABCNCJ
AB+ALAQI |g|gqot{ti|+ALAQI
ttmkkmsC sibthorpiana|g{ikqkkmsC
incana m tztg ~k kig{kqk grgsnk k qjsg
jmsnm+ALAQI
fflyzlmnrCrepis heldreichiana ~mglk{itog{{{{tski{toz{mnottt
qtgiktt|gtqtqspstotztto
pkrggmksg|h|mmgzgitzqhgg
g }o ksg zt{ j|t{t sg gsgzo~nts |gsts|
zgsp osgssg gsnsg }o to kig{oqtopsCLksmzgqgipig~gspsksgzt{
zkqtqsmkrggm|ggs{pmpsgrgsns
gz g lg t ikits go ksk ms kiiks gsgzgqgipi ms ztmg ps g~gsps zto zgqitsg
|nkt|gkztspmsjmtoqigspsgps
C heldreichiana gsgzgqikgpk|tto|oqp h{gm| ~mgltsg ~gm{t zqt|k}g{tq}to
posof{ fpm 2n W "ABCNCJ

gqt+ALAQI
jpsisfrms|spfztststp|gtskzhqtzgspptzqtoto|phqto
johpf|{jqyzolsjm|m{{jg}gslkgzkqtqstkkq|tjzgqtooztz{mnot t qt gikt ki{g psqg qt}m
{g kqgjthtsg kqthtsg g|t~s |g t
qtqspsgki{mtq{ggjgqtq~m|gzsp
gztsitqt mk{tztssto
sys vkg |oqp k ~gqjqk |g k |gq| |t{gg k okqt L qt
gst|hqg~jk|gm{{toknkzkggz
ksg|h|mmgzgitzqhggg}ojstosms
kszpm rmqs j|gzps zqt|k}g{tq}ps
~mgs }gslkg gl k {{g ksjm| to
gito|gtoqspsgzpgPhitosia crocifolia
Onosma leptantha Crepis hellenica ScUtellaria rUpestris
RTBRO rUpestris-inUartia jUniperina RTBRO GlandUlifera
Scabiosa tayGeteaRTBROtayGeteaSilene tayGetea|
kpiqg}|krz{pmtoksjm|tkjtoCrepis heldreichiana

~mgt sg zngss t kii {{ts t kjt

sg|ggg~nkkom{kqm|gmitqggzk{
t{{ig}oksnmkqgg~gsgzgqgmqnm|gstoki{tooztz{mnottokjtotqt
giktikitsttzttkosjogknkqs
zoq|gittztkztqksgohtsmszkqt~zngssgksgozknostigmjo|t{gtokjtosg
kzk|gnkg|g|gsgkzhkg|qtzqnkg
qtkskgmCrepis heldreichiana sg|gggikzqt
t zgqs ms |gmitqg ps qps 65 zp k~k
zqtgnk +ALAQI C |g t qit The Red Data
Book of Rare and Threatened Plants of 'reece0HISNR
AK |gnz{mqtg|qqg"ABU ABU m
)5#. krggmzt{zkqtqsmost{|
zkqt~k}sm|gmjg|k|tsmkrz{pm ps {itqps gz oztz{mnos m k ms
k{zjgmk{{ts|ztqkgmjksnggzgkms
|ggrmkom{kqm|gmitqggzk{
qtogitozqtkstoksggsg|mqo~nkmzkqt~
t g~kqt josgs p ns| qo t kjt ng
ztqtksgkg}kqnkig|g{{qikgktgs|t
zto |g sg kzgsgtztnkmnk k g zqtgkosm
zkqt~ms}mgsgo~qkgnkzz{tsg~gs
tongztqtgssgjgmqmntsigmsex sitUzqtggtokqzklkksk|t{|t
pf ypsff qt t zgqs jks ~tos {m}nk

qgzqtgggtztgngjg}g{ltosmskzhpmmCrepis heldreichiana t{{ts wtz{tsgzt}g|hgigmszqtgg~sttoks{ip
kjto g{{ |g ks z{nto {{ps krgqk| kzmts| mgg zgsps |g ksjm|s kjs to
tksjm|kjtCrepis heldreichianagzNqtgiktmk{tztsstovpoqg|zto{t

jophffpl
Crepis merxmuelleri Kamari & Hartvig
54
Compositae
Crepis merxmuelleri Kamari & Hartvig in Willdenowia 18: 63 (1988).
ys ;{{0I g|kjtsgtqkhkss=b-S
3LNKIJAR GQEAS 3% QAUIME JL 37 NF 3ALAQIMA BX
SHEFNQERSQNADAKNMGSHEQIUEQAKS LQNCJX
AMD LAQRHX RKNOER QNADRIDER AMD RSNMX QIUEQ BEDR IM
LIWEDFNQERSPinUs niGra &aGUsAcer obtUsatUmSorbUs
aUcUparia &raxinUs excelsior ESC REQOEMSIMERTBRSQc
3EOSHartViG & SeberG {ozt#ozt'50!
gztotzttoiskigzqm}tqisp+ALAQI
(AQSUIG zgqtoltos tqsk tq}t{ti| jg}tq |oqp ms zt|{ttq}g ps }{{ps
nk k}sm m C merxmUelleri ksg osnp
gzqgsjqpstzkqjk~nkztgsk|
jm PinUs niGra &aGUs sylVaticaAcer obtUsatUm SorbUs aUcUparia &raxinUs excelsior | |g t hqg~jk
z{gi|g~gqjqkjg|sk|kpsk PinUs niGra
jphpf|t{okzgk~k|{kzro{jm

zgg{jm qlg {g {kz to
CL k {ik k{g}q qnk jg|{gjk osnp
tgskqtgtoh{gt|g{ostk|ggs~qpk p kz t z{kts gjksjk q~k v{{g
hm zt{oqng zgqgstsg kstk p ms zkqtjt m |gqzt}tqg W CL gs{ti~tkj
tjtsp osnp k ki|t{zk |g k ki{t qips| ptkjk{|g|gg{itsghgngpk
~t zo|s |g{osg k kz|k |ggs gjksjkq~kkqh{mg|k}g{to W LL|o{sjq|| pjpstkjkrt{t|{qto|g{ostgzkz|kk{g}q|ggs|g|qspzgjksjkq~k
k{g}qzg~oskskmhmpkq|hq|gzkqh{gt W LL{ko|pz~stjmkzqgspzpog{jmzkqnqgzkqkqt{ikqt
{kg |ggs ms |tqo} rpkq| hq|g |to
LLiqgtkjkintotptto|topskpkq|ssntj~mk|ok{jk|qtp jzgqg h{k}gqjp k}sm |to ~q
LLki{pjgz{to~qLL|qsg~gsg
|ggs|to LLk qghjkzzt
|to LL{ko|
fqrs{}zfCrepis merxmUelleri ~kzkqiqg}k~k|zqt}p+ALAQI(AQSUIG ~k

jgkqt}otikpiqg}|ksjg}qts|gntsksgsg
gst~t SAWNM k t ks oiiks to C tUrcica
$EGEM "AKD +ALAQI |g |oqp k t RTBRO
mUrGanica +ALAQI |g g jt gzgsts k ghkt{n|zkqggzsgk}{~mgqmm"|g
.sjto gzttzttjg}qkktqsgmgs|tq}t{ti|ispqggh{zk+ALAQI
+ALAQI(AQSUIG
fflyzlmnrkhmgskkqgjkjtsg
m~{pqj|qkosgtotqksti|toto|{g
~tos|ggiqg}ktg|{tontoztz{mnottokjto
jtoztz{mnotgzgstsgggtoqto
JL m ggqsg +ALAQI (AQSUIG
sg z{gi k ~gm{kqg okqg
L |g |t to jqto kgr ps ~pqs jk
|gqggkzk|kstsgms~qkqgisp
posof{fpmn W3SQIDTMOTBKIM
+ALAQI|g#NMRSAMSIMIDIRAK
jpsis frms|spf tst |g t{t zgsp
pggoitto
johpf|{jqyzolsjm|kjtm{{jg zkqtqst k ~k| |qt oztz{mnot
ggsgt{|htqktgsgt{||gsttgsgt{|ggtoqto|{gvoztoo{{~nm|gsgzg
gskqg gsgt{| gg to qto zto~ k
gs{titzqpgkgoto|{gttltostq}t{ti|kmsCrepis merxmUellerigzgkgpzkqgqpk{migmsjko|qsmmgrst|to
~m
sysCrepis merxmUelleri gsgzkgkkqzkss|jg}tkokqt Lhtzt
psjt|qsoztz{mnosm|tsmggqsg

kpiqg}|krz{pmtoksjm|tkjtoCrepis merxmUelleri
krz{pm to kjto #NMRSAMSIMIDIR AK sg

~gm{kqk zkqjk z{gi m hgnk ~gqjqg"{gqsgts|{g|gto{~tssg
g|m g om{kqg gg m jg ~gqjqg |g
jg|{gjkmttzttksgmgs|kgqn gps |g ksj~kg ks qk sg ozzk k to
oztz{mnot ps +ALAQI (AQSUIG zm
k}gsltsg jzgqg tg zkqjk z{gi
to qgt to|togsq jmtoqisg sgs g|m
oztz{mno
C merxmUelleri ~mglk sgs gq|k ki{t
g{{|gg|kqgstz{mnokoztz{mnot
k|qspmgs|sgqnsgpsgzkqkqg}o~toshqknkhqg~jkz{gi|g
|q|t{jkztooststz{kigm{g
qsg|ggsq~tsgto{~tsk{ik~{jkpqpsgps
grgsnk |g o~s t{{|mqg g }o ps kjs to
isto Crepis ksg zqt}{ qt} ps }ot}ips
lpszkqt~zto}kgmCrepis merxmUelleriozq~kt|sjosth|mmps}osgzgik{jk|ggitzqhggstttqstoztz{mnot
tokjtogsgztsgkgq|kgzttspskzkqt~kgz|qmsknkztozqt}o{tsggz
mh|mmzpz~toztz{mnotm{gqsg
snptoztz{mnoztogsgzkg|g|t
tojqtogzt~pqjkzqtgqggkoq|kgozkmgzk{gzgsnqzskjqgmqmkt
tztkngztqtgssg~tosgqsm|kzzkz~
jgz{osm|gg}g{qpmtojqtogsgsgrm
k|hqg~pszkqpjsz{gis|{z
qtkskgsg~gqg|mqnktkjtpqp65
zp k~k zqtgnk |g t qit The Red Data Book of
Rare and Threatened Plants of 'reece +ALAQI D
|gnts z{mqt g |qqg "ABIIIIU ABIIIIU |g
|oqp krgg to |qmqto $ 4N |qqt $ jks
~k g}t to{~ts tg gzgqltos ts
ost{|z{mnomC merxmUelleriq|kgg
to|{gjks~toskzgq|kqkosmnkhtgs||g
ksj~kgtoztz{mnottokjtosgksgzt{ogqnkqt |g sg |gg{ghstos kig{kqm |gm gz
goztoksgkqgisp
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merxmUelleri qtkskgksttmzkqt~m{g
qsgmtztgossgksozpg|tztjgkqm
ttq}kmgs|gqnzgsps}osgs|gm
htgs|k{mmjksksg~qkqgt{t|{mqpsm sgknkoz|gnkzqtgg|gsggztqgzk k|k tztgjztk g{{gi m jt m h{mm
|g|nkzgqstmgsnqzsmksqikg
zz{tstkjtzqtkskgsgkg}kqnk|gsg
jgmqmnkex sitU ktgs|tztoksgztqksgk{kmnkmht{tigto|gtqztgsgzgqgipi to qg g~gsg kzm zqtkskg sg jgmqmntskqzklkksk|t{|t
jophffpllpsuss
js|rlorfrril
tksjm|kjtCrepis merxmUellerigzt

qt |{g m sjto }p psgssjm |g gz |g{{kqimst o{|
}pgqm

54
Crepis sibthorpiana Boiss. & Heldr.
Compositae
Crepis sibthorpiana Boiss. & Heldr. in Boissier, Diagn. Pl. Orient. 11: 56 (1849).
3ys ;{{ t qm t gsps= bIM
RAWNRIRLNMSITL3OHACINSICNQTL#QESAEIMLNMSE3SQAUNONDIAcHeldreich !4(5' "/)3
nrrnf Crepis diVaricata "NIRR (EKDQ RTBRO
sibthorpiana"NIRR(EKDQ (AXEJIM2EOEQS3OEC.NU
2EGMI6EG"EIH
jphpf| t{okzgkro{jmzgg{jmqlg
{gtto CLms|g{{qikg~qCL
gz{t kq|t ~oq jg|{gjst v{{g hm
W CL gs{ti~tkj zkqmg k {thttjtspt|g{osgk{ko|pzq~pg~kjs{kg}{{gh{gt{ig|qhqg|tkjkqh{mg|k}g{to W LL|o{sjq|| pjpstkj
|g{ostkhqg~zo|s{ko|pzq~pgosnp
~pq gjksjk q~k rpkq| hq|g hqg~kqg
gztopskpkq|s{ti~tkjtr{m|gkpkq|hq|ggh{gkstkk|ksq|qg~ggiqg
gz hqg~kk k{gs ~oq q~k krpkq| {kg
~stjm kpkq| sntj~m k |q |t{mk
gqg h{k}gqjp k {kz q~k |to ~q
LLk}sm|to~q LL|qsmmskzsp kz}skg ztq}oq kqonq ms |p ~gsg |to LL|ggskzqtkr~tokqghjk
osnpzqtsptgskqto
posof{fpm2n W;0HISNR+ALAQI k }o gz ts xm{tqkm |g +ALAQI
k}ogzgko|qm=
+TMSYE 'QETSEQ |g C incana 3L jkrk ksg

sg gnkq jz{tkj k n W |g zngs
sg zqitst zgqtksjm | m zt{oz{tkjt C
heldreichianagztksjm |kn W~qptgg
+ALAQI C sibthorpiana~kzt{{tq}t{ti|
"ABCNCJ B +ALAQI |g |ogqt{ti|
+ALAQI ttmk k ms C heldreichiana |g
{ikqt k ms C incana m tztg m grgsng ksg
kig{kqm|ggztk{kggzzt{{k{g}qqjsg
jmsnm~mgopsjtkjsC sibthorpiana
-C heldreichiana oztjm{skms}otikpiqg}|~m
kgrqm|gk{tztssto|gksgko}psg
k zg{gkqg ozkqgg g tztg ~tos zqt|k
gz ms k{m m tstkt Crepis cytherea +ALAQI
+gqm'QETSEQA
fflyzlmnrgnt{{|mqtt}o~qmtztkgpqt}gzglgtz{mnotgjg|
|ggq}tsg to{~ts k{gstsg |nk t
ra gg m stsm h|mm st }o zt gsgztsgzqt}o{gisggkg|gsnptnsto
|oqpBerberis ztqtssgpqtos|gsg~mgtosg~gsgkqgmCrepis sibthorpianazkqtqlkg
st k sg |q z{mno t qt xm{tqkm |g
k {ito jg|tqzsto oztz{mnot g ko|
qmzzqtnpjg|tqzsgtgksgisp
gsgztsgkto{~tsjtnktoqto
qotskqmgtztg~mgltost{iqnktjkk
jpsisfrms|spfztstst~q|gtskzhqt
johpf|{ jqyzol sjm| m +qm kqtqlkg g gskqg okqg to qto xm{tqkm
sg |q z{mno |g ps ko|s qps {it
jg|tqzst oztz{mnot ig tg ozq~tos
|gtqt+qotskqmgtztgmkptsksgjgqpzt|{tq}g
sys ( Crepis sibthorpiana gsgzkg k zkqjk |g hqg~jk nk k jt{sk k zkqjk
|ggqi{jkjg}to~s|pgzg|gsnptnsto}qigsgkghkt{n|jg}tkokqt L
3fqrs{}zfCrepis sibthorpiana~kjgkqt
kzmts| ksjg}qts gz }otikpiqg}| |g
|ogqtgrst|ztm|gnksgsgzgqtksjm|
jz{tkj kjt |ogqtgrst| k{m got to
SAWNM |g ps oiiks|s to kjs C heldreichiana

kpiqg}|krz{pmtoksjm|tkjtoCrepis sibthorpiana
tgtoksjm|tkjtoCrepis sibthorpianagzgko|qmmqmvp*"IEMUEMT

nkzqt}o{giskgzmsh|mm
iggsnsg}o}okqgg~gsgzgqgmqnm|gstskig{kqtispz{mnotokjtog
ko| qm zsp gz ms szt{m ikits t tztt
zngsk|}qlkmsjo|t{ggottoSAWNMsgkrgz{pnkg|msgjgmqkgtgtoz{mnot
topmgsgzgqgipigzzqggqtkskg
sg ozg~nk t kjt ms |gmitqg to qpt 65
zp k~k zqtgnk |g t qit The Red Data Book of
Rare and Threatened Plants of 'reece +ALAQI E
j|gg{ghsk|qost{||gmmtztgksg
jg|k|tsmkoztz{mnotztohq|tsg|p
gzzkm|oqp{ipmozkqht{|h|mmwk|
ttoz{mqtg|qqg"ABIUU |g#Bm)5#.

pf ypsff g ztqtk sg zkq}qg~nk g

|qzkqt~krz{pmtokjtokgzoskssmm k tz| gq~ ng gs josg m osk~zgqg|t{tnmmps}osmzngskz|gmto
oztz{mnot |g m zqtk|| o{{ti g~gsps g
tztgngztqtgssgjgmqmntskqzklkksk|t {|t sg ~qmtztmnts ig |g{{qikg |g
zngs k{{ts| ks~om }o|s oztz{mnos t
tztt hgstos zqt mgs| k{pm to gqnt
ps gps to ztgjztk zngs kzgskgipi
}ostokjtoksgsoztz{mnongzqzksgiskg|gzqtmmgz}ozto~toszqt|kgz
|g{{qikg g~gsps to jto oztz{mnot zm
mC sibthorpianangztqtksgkg}kqnkzqt|g{{qikg ex sitU k tgs|t zto |gnts ksg
|g{{m{mighqg~|mzto
jophffpl
tksjm|kjtCrepis sibthorpianagzgko|qmmqmvp.4TQKAMD kztqkkmgrgsngvp*"IEMUEMT

Crocus robertianus C. Brickell
o
Iridaceae
Crocus robertianus C. Brickell in Jour. Roy. Hort. Soc. 98: 362 (1973).
ys ;{{ 30I.0I zkqt t pgsssps=
bLNTMSAIMRBESVEEM)NAMMIMAAMD4QIJAKAcAKSL*
R -arr {oztjkig|ggknkstmshtgs|o{{tim2NXAK(NQSICTKSTQAK3NCIESX7IRKEX
jphpf| t{ok ht{hqqlt }o to
CL|gmszkqtjtmgsnt}tqgt{h|tsjo{tq}t~kjs}gq|jgqto LL~sk kssp j|opt sjk g}o{{g {ko|
khqgsjm v{{g gzsg ms zkqtjt gsnt}tqg hgnozqsg k k}gs z{g {ko| iqg
msspkz}skgz{to LLk{g}qqg~g
g |qzkjg q|t |g hqg|jt g ~kjs g
khqgsjm{ko|kstk|zqsgsnm k
{g ozt|qst kq| }tq {ko| {kt q~pp{sgzkqitsto|to CLqjst jm
p{ko|zg{ggs{ti~tkjpk{{kz|o~s
qtiio{ksg gh{g {ko|pz p qjsg snqk
|to CL |qst sgg |to LL
{ko|t p ozt|qsto ~qgt {kg k |qt|tz|nm{{ttosmnkqgg|qkqtps
msps |qst p kssp ztqt|g{~qpt jg|{gjlkgtgskqtgtokqgggjgz{gosksg t |qt to }qtsg kstk ghgnk
~g|to CLkz|m k{{ktkj
zqgg|ggsk{{ktkjzkqzto LLkjkqtkgst~~qpmqg}
*UniperUs oxycedrUs |g Ostrya carpinifolia kqkm

to C robertianUs t ~gm{ okqt ps L
ksggrtmkpm|goztjk|sktkjtjks~k
gzt|{k| tqks mtqks |ggst k ~gm{
okqttz{mnot}tsgkoiqkj}m|tsk
~kgqqto|gqo|g
fqrs{}zf ksspj|optsjk~sktoht{ht|tsj{totoCrocUs robertianUsozksnoltostsC cancellatUs (EQBEQS|gjgkqgtsC cancellatUs RTBROmaZZiaricUs (EQBEQS "-ASHEVttztt
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subsp. speciosus
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Crocus speciosus M. Bieb. in Beschr. Lnder Terek Casp.: 129 (1800) subsp. speciosus
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Cruciata taurica (Pallas ex Willd.) Ehrend.

subsp. euboea (Ehrend.) Ehrend.
54
Rubiaceae
Cruciata taurica (Pallas ex Willd.) Ehrend. subsp. euboea (Ehrend.) Ehrend. in Pl. Syst. Evol. 124: 178 (1975).
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Cuscuta atrans Feinbrun in Israel J. Bot. 19: 22 (1970).
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54
Cyanus baldaccii (Bald.) Holub
Compositae
Cyanus baldaccii (Bald.) Holub in Preslia 46: 228 (1974).
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Cynoglossum sphacioticum Boiss. & Heldr.
54
Boraginaceae
Cynoglossum sphacioticum Boiss. & Heldr. in Boissier, Diagn. Pl. Orient., ser. 1, 11: 125 (1849).
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54
Dactylorhiza incarnata (L.) So
Orchidaceae
Dactylorhiza incarnata (L.) So, Nom. Nova Gen. Dactylorhiza: 3 (1962).
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Dactylorhiza macedonica J. Hlz. & Knkele
54
Orchidaceae
Dactylorhiza macedonica J. Hlz. & Knkele in Mitt. Arbeitskr. Heim. Orch. Baden-Wrttemberg 20(1):
191 (1988).
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Dactylorhiza pythagorae
Glz & H. R. Reinhard
"1
Orchidaceae
Dactylorhiza pythagorae Glz & H. R. Reinhard in Mitt. Arbeitskr. Heim. Orch. Baden-Wrttemberg 24(1):
58 (1992).
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Datisca cannabina L.
54
Datiscaceae
Datisca cannabina L., Sp. Pl. 2: 1037 (1753).
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gsgispqnkpgzk{ngztqtgssgzkq}qg~nts
gg im zto zkq{ghstos to oztz{mnot
k sg gzt|{ktos ms zqhgm ps gitzqthps
|gsgjtosmsjosgmgg}osggsgzo~nts
sgjmtoqitoszqgg|gsg}oqtosgqh{ggtkzkq}qgiskzkqt~ngzqzksgzgqg|t{tontsg zqtk|| k sg k|mnk m kzo~g
gomjg~kq|zqg||
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qqkskgqkq |gn{kjkr grgsnktokjtoDatisca cannabina gzmszkqt~gtgg

psko|sqpsvp.4TQKAMD

Dianthus arpadianus Ade & Bornm.

Caryophyllaceae
Dianthus arpadianusAde & Bornm. in Repert. Spec. Nov. Regni Veg. 36: 385 (1934).
ys ;{{ .!E t gtnq|m= bIM CACTLIME
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arpadianUs ms|gmitqgpssjoskosps
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m gzt}oi |ggqkz|s kskqiks to htzto
ztogsgztsgtz{mnotto
kg|ggiqg}|gzgqg|t{tnmmtokjto
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Dianthus haematocalyx Boiss. & Heldr.

subsp. phitosianus Constantin.
54
Caryophyllaceae
Dianthus haematocalyx Boiss. & Heldr. subsp. phitosianus Constantin. in Phyton (Horn, Austria) 39(2):
279 (1999).
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b%OAQCHIA4HIUNMCJL3NF8IQNMNLIUIKKAGEAKNMG
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VISHOHQXGAMAAMDKNVROAQRERHQTBRRSNMXOKACERAKS
C Lc.%-AXConstantinidis {ozt50!ozt#50!
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kjt zto }kg k go |k ztqk sg ostjkkg

gz |q tstk kjm zp g BiscUtella didyma
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ms zkqt~ ng zqzk sg gztg|qosnk g kqskg
qm ng zqzk sg gsglmmnts zkqkqt oztz{mnot|gsg|ggkqmntsg}oto|gnksks
ksgrk|ngqtgsto{~ts|zttgztooztz{mnotozitsgkzqtgkosmzkqt~gz{ggto|tov}tqkztoksgozknost ig ms jkrgipi qips g qm z~ g tz|
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|g m jgqmm ps zkqps to k qzklk ksk|t{|t|gn|gm|g{{qikgtooztkjtok
tgs|tztoigmsex sitUzqtggto

js|rlorfrril
Dianthus ingoldbyi Turrill
$-
Caryophyllaceae
Dianthus ingoldbyi Turrill in Bull. Misc. Inform. 1924: 314 (1924).
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jphpf|t{oknogstq}t}okzt{oqntomgsnt}qtoh{gt~oqgztro{psto
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}{{psh{gtk{g}qg|qkqtgzmjkqt
to h{gt snt}qg k{~m k kqgsk gsp}kqk |gp}kqk {kz q~k }gg t |p
g to grgsng ~g{gq k snm gg kz|g{o|to g krpkq| zqttjko|
|qkqg |g kskqg tj|t |to p CL
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LL p~q{ko|t ms sp kz}skg ozt|qst
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dactylon ,aGUrUs oVatUs -icromeria jUliana Phillyrea
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cristata SarcopoteriUm spinosUm Silene aVescens
RTBRO thessalonica SolanUm elaeaGnifoliUm TeUcriUm
capitatUm TraGUs racemosUs|g Vitex aGnUs-castUs
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zqtkskg m gzt}oi kght{s ms h{mm m
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vpqi|g

Dianthus juniperinus Sm.

subsp. kavusicus Turland
"1
Caryophyllaceae
Dianthus juniperinus Sm. subsp. kavusicus Turland in Bull. Brit. Mus. (Nat. Hist.), Bot. 22: 168 (1992).
ys;{{++ t+qmtgnto=b3JKIQNR%NF-S+AUTRIc3EOBarneby & DaVis
sn{ozt+
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CLkplkim}{{psgrgsngk
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Drymocallis halacsyana (Degen)

Kurtto & Strid
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Drymocallis halacsyana (Degen) Kurtto & Strid in Ann. Bot. Fennici 40(2): 137 (2003).
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0HISNR$"EISQGEYTQ+EMMSMIRDEQRDGIRCHEMCampanUla !QSEM^"EQ$ETSRCH"NS'ER
0HISNR$$IEPTIMPTEKNJTKQEMCampanUla !QSEM^RSEQQ"NS:
0HISNR$$IE'ASSTMGSymphyandraIMDEQGIR^"EQ$ETSRCH"NS'ER
0HISNR$$IE'ASSTMGAUbrietaIM'QIECHEMKAMD^#AMDNKKEA
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0HISNR$BCampanUlareiseri(AKCRX^0O )M0HISNR$3SQID!3MNGEQTO3'QETSEQ7
0HISNR$CCentaUreaniederi(EKDQ^0O )M0HISNR$3SQID!3MNGEQTO3'QETSEQ7EDR
4HE2AQE$ASA"NNJNF2AQEAMD4HQEASEMED0KAMSRNF'QEECE^!SHEMR77&FNQ.ASTQE
0HISNR$#NMRSAMSIMIDIR4H!MEVROECIERNFCentaUreaRECSPhalolepisFQNL'QEECE^&K-EDIS

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0HISNR$+XOQINSAJIR:AnthemisGlaberrima2ECHF 'QETSEQ^0O )M0HISNR$3SQID!
77&FNQ.ASTQE
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0HISNR$3SQID!ArenarialeUcadiaAMEVROECIERFQNLSHEIRKAMDNF,EUJAR7'QEECE ^"NS#HQNM

0HISNR$3SQID!3MNGEQTO3'QETSEQ7EDR 4HE2ED$ASA"NNJNF2AQEAMD4HQEASEMED0KAMSR
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SHEMNQSHEQM0EKNONMMERE'QEECE^7IKKDEMNVIA
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IM'QEECE^#AMDNKKEA
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Anthemideae IM"TKGAQIA^0HXSNK"AKCAM
3CHMEKK$%#AQMIUNQNTR0KAMSRNFSHE5MISED3SASERAMD#AMADA^0NQSKAMD/24ILBEQ0QERR

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3CHNSRLAM($#NMSQIBTSINMKaSTDEDERCallitrichesDTBARRIMLDISEQQAMEMESDT-AQNC,EMNLBQE
CHQNLNRNLIPTEDECallitrichepUlchra3CHNSRL^"TKKSTD2ECH3C"IAQQISY
3CHQEIBEQ ( 7AKDGQEMYMAHE "TCHEMVKDEQ TMD 'QARKMDEQ DER &AKAJQNM TMD 0AMGNM IM
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3MNGEQTO3AAethionemaretsina0HISNR3MNGEQTO^0O )M0HISNR$3SQID!3MNGEQTO3
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3MNGEQTO3BAlyssUmteniUm(AKCRXÔ )M0HISNR$3SQID!3MNGEQTO3'QETSEQ7
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3MNGEQTO33MNGEQTO"0HISNR$+ALAQI'4HENQANF#HINRIRKAMD'QEECE ^"NS#HQNM

3MNGEQTO33MNGEQTO"3SALASIADNT%"NSHLEQ2UNM'TRSAFRRNM-&KNQAAMDUEGESASINMNF
!MDQNR+IJKADHER'QEECE^!MM-TR'NTKAMDQIR
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3SEFAMNU"-NMNGQAXAMAQNDAColchicUm,^3BNQM"KG!JAD.ATJ
3SEUEMR0&|gkzkg !MGINROEQL0HXKNGEMX7EBRISE6EQRINM*TME;zkqtj|gsgsktkst=HSSOVVVLNBNSNQG-/"/4QEREAQCH!0VEB
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3SQID!B-NTMSAIM&KNQANF'QEECE#ALBQIDGE#ALBQIDGE5MIU0QERR
3SQID ! C Adonis cyllenea 2AMTMCTKACEAE AMD HelichrysUm taenari !RSEQACEAE QEDIRCNUEQED IM
0EKNONMMIRNR^!MM-TR'NTKAMDQIR
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jtn lth kth|n

THE RED DATA BOOK

Assisted by Pepy Bareka

HELLENIC CAL SOCIETY

zgitqkkgmgsgjmtkom mgsgzgqgipi t{| kq|zkq{mz| |gn|gmgzjtmtozkqk~tstotohh{toktzttsjztkqzt m~gs| m{k|qts| }ptoz|{{t ~pqzqtmitksm

zqkto {tzs sg|g{ptqpsnkqgtzgqshh{t|gsgkzgsptooiiqg}k|gmskzqtz|jttozpiqggzg{gkqgbzqtggmktiktot

FNQAMXCNTMSQX AMDCEQSAIMKXFNQ'QEECE7HEM)VARAXNTMGLAM ASASILEVHEMADUNCASER

mskgipimhgqogsmgzt}kpmks|osk{kkpto|phqto  ig ts bgi|t |ggg| qm ig ms vmc gsg}qkg kgr {{ps

zkqhg{{ts|osn|kmkzt~g tosjkjkskkmskzhpmpslspstqigss |gkszqt|kspto}o|t|tom{{jt g{{tososk~kqztqgijgt

g|k}{ggmgipizqtq~tsg kstkk{g}qqtztztmsg gzthh{tho|hh lqiuht{psvttogqm 

kj|kqg gq~lkjmgzto|{g|t~qstoktsk}qgttsqt 

toRelation dbUn VoyaGe aU ,eVant ttzttzkqiq}ksgski{tsgqn}oszto

{hk htgs| 0IEQQE %DLNMS "NIRRIEQ   ztqk sg ms krkqksmk ts

ts jts g|ggs|mm itmkg^ g{{ |g ms qm zt{kg ps kqkoss to ozqrgs

zpgsk}qnm zt{{gzgllsgksggsnqpztikstzqtk{kkp|g{ggq~g}og gtztgo~szkqtqltsgkgzttspsk oi|k|qskzkqt~ ztogskz~nm

i|qm to zttt ps ksjm|s kjs ps ki{ps ikpiqg}|s zkqt~s m

|tksjm|kjmmqm t%benUs cretica|gmCentaUreaarGenteaRTBROchionantha

||tqo}ki{gpsqgtqtgikt zto}tsgzt{{ksjm|kjm |gn|g

sttoigto.!E  k{tzssmt0  kqk{{3S%  stst)N)  g

|{pqj|zkqt~m{{jt qm^qzgnt  o|{jkst+IJ  sgt{|sttoigto%!E  o|sttoigto7!E  qkksttoigto

|  snt}qt |g |gqzt}qt |{jt m ArbUtUs Unedo |s |togq gz ms k}g{{msg

j|gp isposn|kpskqpsg~qg|g |giago iks| jksksggztjk|ztk{kggzzt{{ gskzo~kzkqzkmskzmts|tqt{tig

togigm|t|{zto msqm zkqkqkstotoigto|gkkq|st

|%Uphorbia dendroideskghkt{n|thq~tomsmjgkg{tottp{tg|gqsgsg vpgqm 

|ksjqjmtq}m1UercUs ilex gztqttjmk}g{{msgvpgtsm 

nsmt|ksq|ggomsmjt|gk|gmqkps k}gsltsgjksjqjkojk tztk|oqgq~tsgkjmArbUtUsUnedo Phillyrealatifolia|gPistacia

tks{ipozt{kg|jt~k~gqg|mqnk j|gp phnrnu tn|lu n nstszto

|qttoozt{kg|tjtokjksjqjktq}ArbUtUs Unedo Phillyrea latifolia

|oz|h{mmgk}{{ps |{mqt}{{psnspsg|g mstztg|oqgq~kmArbUtUs

~g{ztk|m |gP brUtiaqg~kgk|m q|kgigjtkjm oiikskgrto|g

|Abies cephalonica gztjttoqtost k}g{{msgzto|gtlocUsclassicUsto

kqt Lzto|gtlocUs classicUs tokjto hmsoztlsmm&AGINMRXKUASICAE

kzm|gag{}ghm|kq gkjm|ggoztkjmg|nkSAWNMgsgiq}kg kzm mt|tiskgmstztgsttgs|khh{tiqg}|gsg}tq ttztkgst~tsk|nk

po5SJLDP@AJD]54nkpqkgtSAWNM ttztt khmz{tsg|qhkjgnkz{mqt}tqk z{mqttzttjztkgzg|qqg m|gmitqgbqpc|gkztsp

% LAIKQIRSNV TMI ONSRDALDE

Corydalis thasia3SN+ISAM 3SN+ISAM

Acantholimon aegaeum F. K. Meyer

nrrnf Acantholimon UlicinUm 7IKKD EW 3CHTKSER

mskzgq~g-TKAgtgtoA aeGaeUm gzmt

fflyzlmnrtAcantholimon aeGaeUm }kgkt{iqnktjkg|qhpszqt|k}g{tq}psgps k{ikzkqt~zto|g{ztoszkqhg{{ts|togzgktjkpsgpsg

pf ypsff {{gi ms ~qm m im

rinrs {f }fpf{lp {flhspf fyjz

Acantholimon aeGaeUm gztqtqtmstotovpgqjg 

Achillea occulta Constantin. & Kalpoutz.

to qto zto jgmqtsg ozt{kgg h{mm

jphpf| t{ok kjt ~mglts ojk snt}qgk{~m~pqjg|{gjk |to CL

fqrs{}zfAchillea occUlta zkqiq}m|kzq}gg#NMRSAMSIMIDIR+AKONTSYAJIR kpqnm|kgs|kkg|qksmgkjsztogztk{ts

ms tjg Anthemoideae m tztg oiikskk ks k

gskpzlk |ztts kt |sjost krg}sm t

ffl yzlmnr Achillea occUlta ~mglk

Adonis cylleneaBoiss., Heldr. & Orph.

nrrnf ChrysocyathUs cylleneUs "NIRR (EKDQ 

fqrs{ }zf Adonis cyllenea o{{~nm|k ig

C cylleneUs p ~qklkg zkqkqm zkqgg|

k ps tr|s og|s sg ksg ~gm{kqk g

fflyzlmnr/kig{kqtisptz{mnottoAdonis cyllenea }tsgtqtg

pfypsffgtohtztotoAdonis cyllenea ng zqzk sg krg}g{nk st| zqtgg k

rinrs {f }fpf{lp {flhspf fyjz

nrrnfAeGilops liGUstica 3AUIGM #NRR !OOTU

kpiqg}| krz{pm to SAWNM AeGilops speltoides UAQ liGUstica

zto jgmqtsg ms qzklg ksk|t {|t |g

sys t kjt }kg |oqp k {hgjtzts|

ffl yzlmnr ~q kqg t AeGilops

jtn lth kth|n

zgitqkkgmgsgjmtkommgsgzgqgipit{|kq|zkq{mz||gn|gmgzjtmtozkqk~tstotohh{toktzttsjztkqztm~gs|m{k|qts|}ptoz|{{t~pqzqtmitksm

zqkto{tzssg|g{ptqpsnkqgtzgqshh{t|gsgkzgsptooiiqg}k|gmskzqtz|jttozpiqggzg{gkqgbzqtggmktiktot

FNQAMXCNTMSQXAMDCEQSAIMKXFNQ'QEECE7HEM)VARAXNTMGLAMASASILEVHEMADUNCASER

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zkqhg{{ts|osn|kmkzt~gtosjkjkskkmskzhpmpslspstqigss|gkszqt|kspto}o|t|tom{{jtg{{tososk~kqztqgijgt

g|k}{ggmgipizqtq~tsgkstkk{g}qqtztztmsggzthh{tho|hh lqiuht{psvttogqm

kj|kqggq~lkjmgzto|{g|t~qstoktsk}qgttsqt

toRelation dbUn VoyaGe aU ,eVant ttzttzkqiq}ksgski{tsgqn}oszto

{hk htgs| 0IEQQE %DLNMS "NIRRIEQ ztqk sg ms krkqksmk ts

ts jts g|ggs|mm itmkg^ g{{ |g ms qm zt{kg ps kqkoss to ozqrgs

zpgsk}qnmzt{{gzgllsgksggsnqpztikstzqtk{kkp|g{ggq~g}oggtztgo~szkqtqltsgkgzttspskoi|k|qskzkqt~ztogskz~nm

|tksjm|kjmmqmt%benUs cretica|gmCentaUreaarGenteaRTBROchionantha

||tqo}ki{gpsqgtqtgiktzto}tsgzt{{ksjm|kjm|gn|g

sttoigto.!E k{tzssmt0 kqk{{3S% stst)N) g

|{pqj|zkqt~m{{jtqm^qzgnt o|{jkst+IJ sgt{|sttoigto%!E o|sttoigto7!E qkksttoigto

| snt}qt |g |gqzt}qt |{jt m ArbUtUs Unedo |s |togq gz ms k}g{{msg

j|gpisposn|kpskqpsg~qg|g |giagoiks|jksksggztjk|ztk{kggzzt{{gskzo~kzkqzkmskzmts|tqt{tig

togigm|t|{ztomsqmzkqkqkstotoigto|gkkq|st

|%Uphorbia dendroideskghkt{n|thq~tomsmjgkg{tottp{tg|gqsgsg vpgqm

|ksjqjmtq}m1UercUs ilexgztqttjmk}g{{msgvpgtsm

nsmt|ksq|ggomsmjt|gk|gmqkpsk}gsltsgjksjqjkojktztk|oqgq~tsgkjmArbUtUsUnedoPhillyrealatifolia|gPistacia

tks{ipozt{kg|jt~k~gqg|mqnkj|gpphnrnu tn|lu n nstszto

|qttoozt{kg|tjtokjksjqjktq}ArbUtUs UnedoPhillyrea latifolia

|oz|h{mmgk}{{ps |{mqt}{{psnspsg|g mstztg|oqgq~kmArbUtUs

~g{ztk|m |gP brUtiaqg~kgk|m q|kgigjtkjmoiikskgrto|g

|Abies cephalonicagztjttoqtostk}g{{msgzto|gtlocUsclassicUsto

kqt Lzto|gtlocUs classicUs tokjto hmsoztlsmm&AGINMRXKUASICAE

kzm|gag{}ghm|kqgkjm|ggoztkjmg|nkSAWNMgsgiq}kgkzmmt|tiskgmstztgsttgs|khh{tiqg}|gsg}tqttztkgst~tsk|nk

po5SJLDP@AJD]54nkpqkgtSAWNMttzttkhmz{tsg|qhkjgnkz{mqt}tqkz{mqttzttjztkgzg|qqg m|gmitqgbqpc|gkztsp

% LAIKQIRSNV TMI ONSRDALDE

Corydalis thasia3SN+ISAM 3SN+ISAM

nrrnf Acantholimon UlicinUm 7IKKD EW 3CHTKSER

mskzgq~g-TKAgtgtoA aeGaeUm gzmt

fflyzlmnrtAcantholimon aeGaeUm }kgkt{iqnktjkg|qhpszqt|k}g{tq}psgpsk{ikzkqt~zto|g{ztoszkqhg{{ts|togzgktjkpsgpsg

pf ypsff {{gi ms ~qm m im

rinrs {f }fpf{lp {flhspf fyjz

Acantholimon aeGaeUmgztqtqtmstotovpgqjg

jphpf| t{ok kjt ~mglts ojk snt}qgk{~m~pqjg|{gjk|to CL

fqrs{}zfAchillea occUlta zkqiq}m|kzq}gg#NMRSAMSIMIDIR+AKONTSYAJIR kpqnm|kgs|kkg|qksmgkjsztogztk{ts

ffl yzlmnr Achillea occUlta ~mglk

nrrnf ChrysocyathUs cylleneUs "NIRR (EKDQ

fqrs{ }zf Adonis cyllenea o{{~nm|k ig

C cylleneUs p ~qklkg zkqkqm zkqgg|

fflyzlmnr/kig{kqtisptz{mnottoAdonis cyllenea }tsgtqtg

pfypsffgtohtztotoAdonis cyllenea ng zqzk sg krg}g{nk st| zqtgg k

rinrs {f }fpf{lp {flhspf fyjz

nrrnfAeGilops liGUstica 3AUIGM #NRR!OOTU

kpiqg}| krz{pm to SAWNM AeGilops speltoides UAQ liGUstica

sys t kjt }kg |oqp k {hgjtzts|

ffl yzlmnr ~q kqg t AeGilops

fqrs{ }zf oz| tq} to AeGilops

AeGilops speltoides UAQliGUsticag gzmszkqt~mmtotthqtoh ~o|gi g~jtvp{{mhk{|g

jphpf|t{okzg{kgkg|q{kzmro{jk qlpg snt}qt h{gt |oqt ~q ~kjs

ffl yzlmnr t Aethionema carlsberGii

gps|gk|gtz{mnoost{|jks

rinrs {f }fpf{lp {flhspf fyjz

pf ypsff ~q i jks ~tos {m}nk

tkstztksjm|kjtAethionema carlsberGiigzms|tqo}kqthtsggiktvpTQKAMD

jphpf|t{okzt{h{gt{kt}okro{jmhmz~to~qCLsnt}qth{gt|to

posof{ fpm 2n 0AOAMICNKANT

tksjm|kjtAethionema orbicUlatUmgzms|tqo}toqtonpsgitsqt vpgqjg

zkqt~zpgScorZonera scyriakstksjm|to

jphpf| t{ok }o k hm ~oq ro{jm

fqrs{ }zf t z{ts oiiks SAWNM to

tgtoksjm|tkjtoAethionema retsina gztqt~o{gmsto|qtoztqjkvpg{

ffl yzlmnr ~q kqg ksg ispt

rinrs {f }fpf{lp {flhspf fyjz

pf ypsff t |q iknt ps jt oztz{mnostoAethionema retsinatqt~o{gt

jphpf|t{okzt{{|gjmzg{gkqgips| to CL q~p |oqp ipskv{{ggsnkggz{~pqzgq}o{{ggkqg

rinrs {f }fpf{lp {flhspf fyjz

pfypsffzkqkqt|sjostztogzk{ts ms AjUGa piskoi ksg |oqp o~gg ikitsg

fflyzlmnrk{kzkqt~tztk~k|ggiqg}ktkjtms{{jghqnm|gsk{~gtg{ikqgps gzkqkqggzgo

jphpf| lpgjk zt{ok kjt ~pq zgqg}ojk {g to CL kqgips| ~kjs

rinrs {f }fpf{lp {flhspf fyjz

jphpf| t{ok kjt k gq|k qztsg p

fqrs{}zf(Alkanna sartoriana ksgsgzstkjtttztto{{~nm|kmsqit{jggq~|gz

sysg{gkqkz{mqt}tqkgs}kqgsmAlkanna sartoriana }kgkzkqt~kg|gjkozqpg|tskg|6ARRIKIADER4AM

kgqtttoksjm|tkjtoAlkanna sartoriana|tstt{mqit{jgvpg{ztol|m

tksjm|kjtAlkanna sartoriana|tstt{mqit{jgvpg{ztol|m