You are on page 1of 192

ARAP TASARIM 001-013 PDF

18/11/02

15:12

Page 1

ARAP TASARIM 001-013 PDF

18/11/02

15:12

Page 3

dG
dG

j hQg

QdG GE
J jdG g CG g jhQGdG jdG Qf UN a J AGQh dG
,G M jhQGdG fCG CG a .jG SdG c Y j dG IYdG
dG OLH dG Gbh hCG fjOCG Y hdG J ,dG OLh M dHh
,jdG Y j kLGh jdG g MO j dd .INCG S HQCGh G N
,c e c e cCG GC j CG Qd UdG J b .e c Y dhDe Jh
.VG Gg a f ka f CG jCJQG dd
jBG AV Y dG g c dhJ dG fEG YVG L T
J dG YVG c MT .fe e dGh dG c EG SdG YJ gh fBGdG
SCG N e QdG gP DhdG hCG d kfe J jH fBGdG jBH
YLG dG L AGq d c JdG YG dG dGh dG
e QdG q dG FGhdG SCG Gg .Jh e J CG dG jGh
,H hj h fMhdG QeCG aj jdG dhCG M ,IMGh L dG IAGb
.H YbG ANEG e Gj h dG g JMG dG FH GhKCJ
e dhj hCG Oe H dDG c e Zh dG Gg GC j CG Qd
Ie bG ha e IOSG Zj jdG dhCG eCG .YL be N
.jNBG EG HQ Y dGh JYfH AOEG e S fEG PEG Gk L
eN j dG Ld c dG dG g VYh IAGb gG CG EG aVEG
hjj jd f dd ,bEG jZ SCH dG g FG VY . jd
.kGc kfY d J dG g EG :jNBG EG jdG f
id ,dG jf IOLG dG g e P Y j CG Qd G e
.IGh G jdG OGH dG QOG g VJ dG dG
J hCG Jd T f Lh ,dG e Z c dG Gg d
Fj VhY hCG Ve YVe VY eZ SCG hCG ,dG c Y J
.dG GfG EG ODJh dG J

dG
dG
dCJ

j hQg

LJ

Y NQhGC

2003

jGa - fSG

dD G M
J fCG CG H ,1956 Y IfCG j hQg QG SG j dG JdG dh
QGUEH GC H fdG h ,SG eL S Qe eL GOBG SQO ,IfCG fdGh FGHG
ITG Jbd VJh jhQGdG MJ dG JHH Qe Jc j hQg . jdGh SdG c
.IeG jedG LdjjEG e
HQM jnsdG s d Ibe icP ''j''h ''hQg'' SG e ,QG hCG dG SG j
Gg dG g j dG G JQ ec dG Y dG G j H ,OEGh dG
Sh Y dG U kG f CGh ,jhdG dG NBG g dG BGdG CG EG dG G j .G
c HEGh cdGh jOEG SCG f ag JdG Vh dGh BGdG AV h .dG N g
H e dG G Ggjh ,INCG G c d d ,jd jOG cG Y J dG YGG
.g aGgCG Y YEG H c
EG Y Jh ,SdG EG fBGdG dSdG f gh MGh g M dDG c L QhJ
.NBG dGh dEG OLdGh fEG YVH dGh dH
EG GfEG eh ,jeCG EG dG e AGdG e SGh jd Ic H j hQg c J
fCGh jfEGh fdG EG c H LJ bh ;fSEGh jRGdGh SdGh GdHh fhfCG
.fhfCGh ShdGh fdCGh HdGh jOQCGh dJdGh
SCa ,eDG EG jJh ,dH EG EG eDG Z IYO JFa dG g KCG d
e .QdG Y dGh jdG KCdG fe kFf j dG g H J dG Gh dG
GfCG e f C ke j CG L H gG jh dG g GC j Qb CG Y G
dG g C ,H WY e e dP a ,Y adG AGd j MCG H dh .jOG dG
dG MoO b dG OLh GH J dG LdjjEG L EG .SSCG e dG J J
.j hQg c EG Oj dGh
g f AGQh e JdG gh ;MVhh BGdG M e Ie FG g CG T
,IdG hCG dG dh ,dG EG Ud dG jdG Y j jdG dhCG eN g dG
.g c f AGQh e Oe g Lj fCG Y IhY
Hbh YCG J dG ,dG g IAGb Y jNBG j jdG Ea ,FG g AV Yh
.H QJ eN ej ,d jOdG jW EG gTJh
Va EG fEG OJh QdG gP jdG e kYf J dG dG bJ j ,iNCG L e
VGdG a dG g eCG ,Gh bd e ,SdG b e dG MGREG KDJ h ,LdjjEG
edG dG e cCG Jd HOCG IdG Y cJ fc d QdG Y dG KCG Gg d J fCG
dG g g j hQg d MdG dG CG ij CG dH j eh ,dEG j dG
.ffEG dG jJh

EG d Gg ej eSEG dG j dG YGdGh dG dG CG EG IQTEG e H


e EG IOdH EG j d Ggh ,IadG LdjjEG f LdGh G dG jO Y OHG f
d dG dN d VY dG fBGdG FGdGh dG EG LdGh ,G gG J Y dGh EG
FGOCG e H LGh g ,QedGh OdG jhg f H J dGh jOG dG dM EG dHh .kGQSO d
.SG bdG f b ...EGh
g S dG Hh ,FdG QhdG Gg JY Y NCG b j hQg Y EG :b GPEG df
BGdG H Yh dG IOdGh dGh dG dN e jdG dG T S dG SdG dG
.dG
Y jhQGdG JL dG QGdG ,fSGh jOdG ,jG SG dG :JdG YCG J
G dG ,gQEG aj SEG ,TdG :jhQGd jedG LdjjEG ,TeCG Y YdG ,dG
S ,3-2-1 YVG ,BGdG n b ,ScddG OGM AGQh ,gQEG OGM AGQh ,SdG
hC ,IFdG CG , ,QdG jPcCG ,QdG YN ,dG EG j dG ,2-1 FM fehdG :dG
dG ,Sj dG ,Se dG ,jQdG Y bO HLEG ,kGDS jY QdG jf QfG ,dCG
dG ,dG jW BGdG ,dG Gg IM M ,e c dG ,GdCG dG RYEG ,gdG
,dG N , ,ka CGH b jedG ,GG Y dG e FGQ Ph dG H ,dG
Ie ,dG IG ,YG RG Ie ,G IG ,IQdG Ie ,QdG Mh OG ,gJ
UCG ,dG IG ,dG f IG ,VdG IG ,dG IG ,JdG G
cdG AGG IG ,dG oj dH ,GG e S ,G QdG ,Id G
. DNA QGSCG ,JhdG IG ,FdG
FbUCG Y ,GhdG Y ,GG Y ,!jhQGO c WCG jCG :WCd JdG ch
.SOdG :IG G IH ,JEH N j dG ,dG ,QdG
npdG ,BGdG SSCG gG :fBGb YVe hJ dG iNCG JdG YCG Jh
dG ,G :eDd G ihCG ,gG g ,3-2-1 Ed jS a ,BGdG bNCG
,dG ASCG ,aG QGSCG ,BGdG aG T ,dG S IdG ,BGdG Y ,BGdG fMhdG
,SdG ce ,QdG H ,BGdG e HLEG ,BGdG SSCG gG ,BGdG dOGh dSdG J
,BGdG YdG gCG ,BGdG IdG ,dG J ,gG jO ,KdG ,dG :oG fEG hY
,BGdG dG gCG ,gG fEG T ,G BGdG MCG ,kGHCG J ,dG j
N ,eDG MQ ,SQ J ,J CG b ,eG EG ,gGdG dG M ,BGdG e eY Qe
SQe ,3-2-1 dG dL e Y ,BGdG IG G ,mGB Y dG ,dG SHc ,dG
j c ,f J GP ,dG c JG gCG ,jQdG Y SEG d VJ dG GAGaG ,Sj
j jdG dhCG ,BGdG dGh dG ,e c j dG ,BGdG QGSCG H ,BGdG dG
.BGdG EG

SEL OFSET
stanbul / Turkey
Tel: (+90 212) 6290615

: dG QGO
ARATIRMA
YAYINCILIK
Merkez Mahallesi, Cumhuriyet Caddesi,
Alimanolu Merkezi, No: 40 Zemin Kat
Gneli / stanbul - Turkey
Tel: (+90 212) 511 72 30

: dG GY
KELEBEK MATBAACILIK
stanbul / Turkey
Tel: (+90 212) 5776009

:ffG bG
w w w . h a r u n y a h y a . c o m

dSdG SD e
ee

GG GW G dG

14

QdG :G GdG ILGC

40

dG j fGC h JG

74

jOGJQG MdG fGC

102

FdG adG fGC h HC G dG Ge

114

Id gGdG FdG :dG

122

Gh dG

134

ee
e hCG g G e IOLG edG ;bdG e gH jSCG M d
e c EG .YL f dhj jdG dhCG IY edG g U .dH S
f dG cG e Gk AH , J d ,jSCG M WH j dh M ,dM OLe
.H Y dG RH AfGh dG EG H
eh .e g d e H AGLCG Id Ze g dG ,QNH
C dP ,edG GfCG e f IQdG CG j CG fMCG Y j jdG Gg
AGLCG J j dG Gg Mh ,dGh UTCG f g J e dG
.G h ,IQdG Lh GQWEGh G e G
dG e l f jd GdG ejOh FdG j g ?G FdG Y GPe dh
g dG :IQdG e Y jLCG dG dG ,GDdG Gg Y HLEG b d ?kjCG
dG jdG QdG Y e G AaG dG J dG Gd ,GdG g
e aGe H LCG J a .AGdG RGdG d jdG e dG g
e c QGe EG LM e FdG id SG Y fG c ,FGdG ejdG
.UN J hP e g FdG CG VGdG e ,bdG
LJ .f G LGa ,IG e iNCG kTCG dhJh kfL FdG e cJ GPEG
f fCG a ,dG Gg HJ GPEGh .Gh G dG Y c eCG
.dG Gg e AL fCG
8

j hQg

,H j CG BG LQ C H J dG H dG jj sEG
CG dG jJ dBG aC jH jDhdG Y J SCG g BGJ dG Yh
.gJ
S Y IOLG bG c CG j ,edG dG g EG fMCG j eYh
H :PEG ,e e d H J ch ,ja J G g ,f a ,VQCG
.e Mh IH gAH Mh ,Aj e bh N dN e FG d
.Y SdG dG e J dG QdG jf b e aQ G g CG EG
GfCG L CG YJ gh ''GfCG UCG'' jhQGO dQJ c SCG Vh jdG g
.NBG Y e c QJ aOG GMCG e S Y OLdG EG X b bG
dh .FGY GJ Y IG TCG L Ea ,jdG d SSCG eG Mh
LCG SGh gZ Y d ,IG T QJ QhO FGdG GdG d c
.ddG
jhQGO jf Vh CG EG ,eh Gk L Y fCch keY 140 I jQdG Gg UGJh
jdG g CG J MG g .bG Y dG eCG e fQbh ,c
e be ISa IFGO e cCG J CG hJ fEG :NBG ,kGL IQU SJ
.f
Gd eT jJ J e jha j .FGdG GdG Y khCG AdG d
LGdH J jdG jQdG
q a ,bdG dP G H aG f H G
L G Z aGfG hCG fG e Y :g IdGh .''IdH'' Gdbh
IW h jQdG Y j fCG g VG Gg Gh dG AdG CG EG .G FdG
jdG aY dG GdG M c .G Fd G eG jJ EG OCG IMGh
KCJ GP J G aCG h ,Y CGJ dG G FdH J hCG J fc kjJ
UUdG WEG j IdG N e QJ CG bG CH dG Ea dd .OM
dH .kM MCGh U ahCG UTCG Y HUEG J CG eCG Y SdG e M Y
e dM QJ EG ODJ CG e IW CG gMCG YOG EGh M !AGdG e f Gg
. Qf kag dGRe jhQGdG Ea ,dG eG c f Yh ,jdG G
9

dG dG

j .''F dG IG dG'' Y j dG G g dP AGQh dG


OLd f J G FdG AYCGh fCG dZ CH j e Y G Gg
hCG jdG H d gMCG J hCG AdEG jh ,eGe H J e AGLCG
.cCH dG
.IdG AYCG e S Y e GPEG a GUCG GPBG J :G S Y
J dG ,SdG PCG Y iMEG G S Y dh ,e kjCG qY hCG fG
.J H dG SM
bh ASG hO FGLCG c J CG j ,LQG GUCG e PCG YJ d
GdGh bG gh SdG PCG Y ,dG AZ ,LQG dG IdG :eGe
jdG jG e dG G ,PCG bb C dG FdG ,cdG Yh
cG e G YCG T .GUCG Y jG g YJ dG GgCGh
C fC ,e AGLCG H Qj CG dG dG Gd .edG dG
.dG AGLCG bH Y j CG AL
e kb jhQGO c a ,QdG jf MJ F dG IG dG g CG h
:''GfCG UCG'' Hc a ,IgdG FdG g
Ea ,deh V jJ Y J e Y CG OLh Y gdG GPEG ''
1
''SSCG e S jf
dG feEG e G dG Gg e j CG jj j hCG jhQGO YSH j
dGh ,hdG dG H dG bdG dG dG CG EG .dG dG Y SdG
Ggh ,F Ie H g jG fCG dZ CG Y gH ,UdG OCG hJ
.j c c ''SSCG e jhQGO jf f''
g .jhQGO jf MJ dG jG fCG Y IOe eCG dG Gg Vf
N eh ,GWdG L Mh QdG LCG e Gk AHn e CG g CG fCG
AMCG g dN Y S H ,a jhQGO jf MO EG Uf d d SGQO
.ddG Mh
:dG g dG .G dEG G Y G j dG ddG S g
10

j hQg

n go hn p VQr nC Gh pG hn n s dG en o nd o u njo roG Ao n rSnC G o nd Qo u n oG


o Qp n dG o pd nG o dG n go }
.24 :G |
o p nG o j n dG

F dG IG dG Y e :WdG Y
Y fYG fCG EG ,aNG Y G FdG dG dG GfCG e jdG g
Y GedG U dG hCG ,GedG J dGh jQdG H J dG dG e edG
dG NGO SdG N f cjh SdG Y j dG AdG QfG gh ,Fe
.NGdG
g MCG . H J G FdG e iNCG GfCG dG YCG CG EG
.QfG dh SfG GC e Y WdG Y J .WdG g FdG
g VJ .HG e c OY e G S WdG Y FN gCG e
.bdG jZ ddG dG IQdG hj c HG
M ,dG NBG e CG Lj Gk ja kSg kef WdG Y oJ
2
.dG SdG bQh e hJ ,dG He Gk L bO S Y
J H d kjCG dG He jT HfCH J bO H G HG g
TCG e Y e e fCch hCG gd dG Ub j .dG Ub
WdG Y h .SGdG TGd eeCG dG EG d bGdG fCG e ,SGdG
.SGdG TCG HG
J dG IBGG J HG HfCG g e c fGL CG g AG j e CG EG
fGL VJ Mh ,bH dG Y G AdG Gg cj .Y bdG AdG
3
.IMGh f Y c cd J fEa bO jGhH dG NGO IOLG HfCG
HG HfCG g a .G j e eCG dG Gg d jOdG Z dG EG
H VJ jG g e kc Ea ,dP Y IhY .IBGG e J W Y
.IMGh IQDH AdG j H Gk L bO Sg
fCG dP e gCGh .QdG jf Lh Ic Y J WdG Y CG VGdG e
. kLP J
11

dG dG

e c OY e WdG Y dC J
fH JG HG g .HG
bO YHQ HfC jf GE d
.Ad cdG jGG J gh ,kGL
dG Y G AdG j
jGhH dG NGO HfC G fGL VJ H ,ke e H
.IMGh f c AdG cj M de

dG
dG

QdG e

dG e ,dG g ee MCG M d .''F dG IG dG'' Y


g J a ,INDG dG hCG He d cdG fGG hCG dG HG dG
QJ b dG CG Y gdG G e dd .TCG e T CH jDhdG e dG
e .aOe M b fCG Y FGdG dG Gg be kY OQGdG Z e .IN IN
.ee e c p No b WdG Y CG VGdG
M Jh .jQdG YGG MJ WdG Y iNCG S AG j dc
dGh ,cdG dG g .dG r o dG g dG bG EG fMCG j eY Ie
12

j hQg

a LJ ,Y kLP WdG Y
e gh jdG e IMGh Y
GP LQCG jY Y j
FdG g J .jdG LCG
.HdGh SjdGh fWdG
jdG FY e iNCG AYCG
,''Ad ISdG dG'' GP dG
J Y ke GC e Y J dG
fCG EG ,dG UH dG jG e e dG J g .Ad cdG dG dG
IhY .jFGO hCG SGS J CG eEG jG g Ea ,WdG Y jN Y j Hh
EG dd AdG J EGh ,AdG J jG g IdG SdG Ea ,dP Y
.dG IQDH
.Ad ISdG dG GP dG jdG GfCG e CG ,J e AV Y
jf M .Ad cY Y H LQCG jY jdG dP e dG Yh
L Y QJ b J CG j jdG T QJ dG FdG c Ea ,QdG
cCG J dG ,Ad ISdG dG Y IQe cdG dG CG jQdG Yj dd MGh
.kJ HCGh jdG H kYT
UG ef V ee H J dG c C e Z dG Gg CG EG
GPEG dG AUH VJh gH jdG S .''fG'' QW C a e dG
.Ad cdG jGH Ad ISdG SdG dSG e
dG GP r o dG g EG .e H N b dG Jg CG cDG e dd
Ge G g ch .jd Gk e Gk eCG ''aOG'' feEG e FGdG SdG
kdO EG d fEG .OhM H p G G dEG G Y kLP WdG Y J ,G
GO GG g cG j .RCG Yh jHCG Mh dG Y Y
.dG e dG Ic

13

ARAP TASARIM 014-039

22/2/03

11:53

Page 14

hC G dG

G dG
GG GW
g d QdG CG EG .QdG Y EG FdG IQU J GdG cP Y JCf eY
fGW GQe GG GfCG e jdG .GdG Y IQOdG IMdG FdG
ddG jeCG H aG IdG TGdG J CG PEG ,QdG dG J Y J
.edG e Ie AGdG ke dGh HdG j CG c ,jeCG IQdG SGhCGh
aOe j Gg dh ,S e 300 e J CGH GG CG jQdG
q Yj
LddG c .aOd CG j GG LC G da ,j c
:''GG LC fG dG'' :GY Ve Jhh HhQ EG
H U d ...hQCG J d GH c , IG LCG d OCG a c c''
.dG dS J H dG AGLCG jd dBG Uh ,JEH LCG
fCG H h ,Fa fh G AGLCG GSG :jeCG jg H IG LCG
4
''.AGdG Ad eCG GSG LCG e SG idG e hj J
Ggh ;GG Y qG Qd GG e MGh dO Y oj ,iNCG L e
UCG j a dG BG f'' :dH SGZ H H fdG GG Y dEG QTCG e
jQdG S dGh bG d IG FG H SGQH BG d . 5''GG
.eGdG eX

14

ARAP TASARIM 014-039

22/2/03

11:53

Page 15

n r oG Ao B n rSnC G o nd Qo u noG
o Qp n dG o pdnG o dG n go }
o jp n dG n go hn p VQr nC Ghn pGhn n s dG en o nd o u n jo
24 :G |
o p nG

ARAP TASARIM 014-039

22/2/03

11:53

Page 17

dG QG bGj
.SdG JQe L

dG :HdG dGE
J dG jdG CG EG aVEG .L NGO EG LCG dG j CG
Hh .GG e Z IOLG J Y J LCG cM G Y a
.''FGdG GG'' e g IG g CG jQdG j UG g
Y j dG IG g Y GdG f EG ,dP Y g G CG EG
IFGdG cdG CG d .dG Ge e Ie EG d ,''FGdG GG''
c 6.d kLP dG Ie JGFW iMEG ''jQS'' ,HdG GFW YU
Y dG P VH Yhe ''SQS'' YS dG IBM cT CGH
j dG GQhG AV Y JdG Y AGOCG jW dCG J . (IBM 3081) SG
Y dG GFdh HG GFd SQS P AH dHh .AGdG dG H
.dG P QGZ
16

ARAP TASARIM 014-039

22/2/03

11:53

Page 16

,S bSG dG Y SGQO AGLEH ,dG gGd QG JQe L b


.dG jZ GW dBG d bG g CH jh
hjh L e ML dj M ,H G fhG dG dG L j
G Gg j dG g Hh .cGdG MCG EG dG QRCG dG e kLQe fd
d ,ah j dG G hCG YdG Y dG Hh .GQhG Y Y IQH
J CG dP Y ka c .cG G GdH J K ICa bJ CG IG
dc ,ja f fh J CG J VdG Gg dG VH AGdG e
VjdG YS OJ gh ,S / c 40 :I dH jOY Z J dG YS e jJ CG
,jdH YdG g e dG j YS /c 39 YH hCG e 100 S
eheh Fa fh J dG dG SCG Hh ,bdG IjT ICLG eU Jh
e dG jd j e dH Ggh ,JQG eU fG dG Ea IjT
.edG IT e e M hCG Y eo J CG J
17

ARAP TASARIM 014-039

22/2/03

11:54

Page 19

j hQg

18

ARAP TASARIM 014-039

22/2/03

11:54

Page 18

dG dG

GFW J
SQS HcdG
G d c
bQG JQbh d
.IQhG Y

19

ARAP TASARIM 014-039

22/2/03

11:54

Page 20

dG dG

eCG EG bdG :ka SCG cCc gQhO dG bS NCJ GUG H


.j dd jdG dG gQ J CG J dG ehG jdH eEd
YdG H LG JGQh dG Y JQb M g c Gk KDe dG e hj
e c Y dG j .GG d P aCG Ga Y eCG ,ddG
e c M j dGh ` jhc dG dG Jg OhJ . SY dCG KK Y
dG j CG Hh .kGL SGh jDhQ IG ` kjJ SCGdG M f
.X AGQh j WG Y
CG EG ,deh Ija H Y j e c ,fCG e Y dG hj Ggh
c N a dP eh ,NBG dG Y j fCG g e k GC Y GC j b J CG
.jG JQhO J CG IG d eCG Ggh ,T hCG U CG hO fCG g

dG LGC
id GdG dBG T M CH h gCG L bCG cCG dG LCG J
,CGG M a .IN IN QdG jf AV Y ML a j dGh ,dG
cCG LdG Y Y ODJ dG LCG C e J dG IOG EG QdG jf J
.ee IMc QJ GPEG EG
jG jG AGLCG H Y GHh ,IW EG VJ b IG L CG d Vd
iNCG IW M VGaG ke G Z e .G VGh Z J G J dG
L EG J d GdG g C ;G Y UG IdG g Y aOG jH
.cG Y JQb e S H ,IjL Ie CG J d fCG c ,kec kML IG
eCG be S fC ,U d e gh ,FGR m Mp p t EG IG S
ODS dG AUG Ea ,QdG jd FdG GC G Mh ,dP Y IhY .aG
.ILdG IG g VGfG EG
J eYh .hG QOf Gk eCG IdG J VGaG g c eh
20

ARAP TASARIM 014-039

22/2/03

11:54

Page 21

Y Jh ,SY dGC 30 Y Y c .kGJ VQC G Y IOLG GG Y cGC e dG Y J


.G G bGe e J fGE M kGL SGh jhD Q IG Ohj kjJ SGC dG Me f dG
h WH zaOG GC e{ Y J dG QdG jf YGe j ,dG jZ dG LGC H J ,iNGC L e
N d ITe f GE d a IOLG eGh LC G Ad ejOhjE G dG GE .dG e d SSGC
.bdG jZ Sd

21

ARAP TASARIM 014-039

22/2/03

11:54

Page 22

dG dG

GC HC G c .eeC G LC G GE AGG dG Jh ,GdG AKGC dG LGC cM YGC IQdG hJ


J UN Y SGH LC G J .Fa fGW J IG , JGH aJ Gh eeC G LC G
.aGH

G e dG Gd .k kVe VGeEG EG ,G ZCG ODJ h IH QGVEG


dBG EG edG e Qd dG L dG H EG dG GdG ODJ CG
Y r jo GPa ,kU jQdG
q H j dG AYOG CG kL S d dh .fGdG
H NG Y AdG j .G ?''d FGdG'' dG QJ dG IG
`d jH Y j :NBG .dG j Hh dG AS QW j bCG Ge
. r n p dG bG KG g j ''FGH LCH j'' hCG ''j f''
:CG ,Gg ej M j h G FdG c e e ,kee kco dG No d
.''Qj'' h kNn No fEG

GE OJ dG Ke
fL GE S e 250
.jM j

22

ARAP TASARIM 014-039

22/2/03

11:54

Page 23

j hQg

J dG SSCG IOG g dG CG ,Gd dG jdG EG fG GPEG


GdG Y .dG cM d j feh jb IOe gh ,GG cg e
IG G dG Jh ,Gh eCGh SCGh YCG EG LCG J CG
LCG J .NBG Y e gMCG ,LCG e LhR dG .cG g
J .G MG j eeCG MG Jj eY fEG :CG ,eGe Z H
IOLe aGH dG g Jh ,LCG jH cG dG e Y
Y OJ ,LCG hR Y d dG e Y J M h .G NGO
.Jh HdG GFW JJ f GC G Yh ,QfH NBG hd d iNCG
.YH G J hCG LGdGh ,dG h IQhG e dG q e Ggh

j jb IOe IG L J dG dG IOe GE
IG J .d dG dG d
.kGL GL H

ARAP TASARIM 014-039

22/2/03

11:54

Page 25

dG dG

dG (fG) dG QW
dH e j Gg CG EG .NEG H iNCG Ie dG fCG hGJ
HdG SGH fCH cdG Calopteryx Virgo. Za jHdc f e cd
e H cdG j .cdG fP M bS fCG Jh .(1) bQ e jP IOLG
id SdG jdG Nj jP e (2) OGeG SGH QcdG e Z f CG jH
g gh MCG H Gj fEa ,YS IY J dG g CG h .fCG
J eYh .(3) dG dG hCG GdG e VH dG j .G VdG
iJ GdG g Nh .(4) GS HQCG EG K e AG J VdG e bdG
eEd gDJ YH MdG Y Gk QOb QdG Gg L j dG Gd .(5) AG
j dG dG j bdG J eY .jdG jd j jb gsah ,H
Jh AG J ,NCG dG bh j eYh .Ije bhCG HQCG Ma ,H
eDJ dP H .eGbCG J CG EG OdG J .(6) IU hCG kjW kJf

ARAP TASARIM 014-039

22/2/03

11:54

Page 24

j hQg

G J IMGh dR C ,eGbCG jf IOLG HdG IY HK VH f


.MG g
EG G Gg j Lh Y dG CG HdG HQCG QGWCG Y QdG Gg j
.FGQ Y N e FW
jL e j kSGh kNT d dG j K ,(7) khCG bdG X j
.(8) HdG G Y I HQCH kes D e bdG G Gg j .hCG G Y kc
,M Y QJ h f dc J dh ,de feh aT H HQCG g No
.TJh AG bdG S j
dG L j .L W dG YJ UN dBG g ,dP Y IhY

2
3

ARAP TASARIM 014-039

22/2/03

11:54

Page 27

dG dG

H UN FS j dG Gg ''j'' dh .dG dG NGO ke jh


dG e hG H G e G AGLCG J .dG REG J UN eh
dG HQCG H J Fc IOe RGaEG j AKCG g .(9) UG FdG V H
.iPCG CH dG hO dH IjG dG J
,ke kS j CG LQCG MCG c CG e ZdG Y ,JEH dG g J
.H CH dG b CG EG bO jY f dG Jh
Vh J fCG EG J H QJ b LCG J g
J Y G e V a UG FdG eCG .dG
10
gf H d LCG J .(10) LCG jN EG IjT
.(11)
HH j ,ke jh dG L dG QOj CG H
ajh IMGh IMGh fS h a ,LCGh fS
.ge QNG LCG jh
Y dG e .GdG dG EG Gk NCG IG J
G dP f g IFdG IG g CG j CG AG
H dG j .(12) AG j kYhjo c dG
,dG RGJ Y d FGdG FdG
12
11

ARAP TASARIM 014-039

22/2/03

11:54

Page 26

j hQg

.GdG Y IQOb IG UCGh fG b dG MGe J gh


hCG UCH fH dG hf eY QdG jf GAYOG dSG LGf iNCG Ie
GJ N e IG EG JCG bG sEG :jdG J J.p G dG Gg Ce
e QW CG dBV H e ,CN CG Ie Y dG Y CG EG ,FGY
IG g AGjCG S QGWCG e QW CG kH c dh FY CG ha ;QGWCG
.p oG dG Y VGh gH dHh ''F Ie Y'' dG .cg hCG
dG j :QNH
dG FdG MCG dG
9
G dG N Y d M
e WdG JCj dG Gh
a .N e h jj H
,GG M J dG
...dG bG J

ARAP TASARIM 014-039

22/2/03

11:54

Page 28

dG dG

GdG dGB
:G S Y .YCG Y J dG HdG GQTEG H GG LCG J
gQhH ODJ dG dG J Y TGdG Y dG GQTEG g e IQTEG c J
''aGh aGdH'' J dGh ,cG dG e Y a .G j EG
eY SCGh YCG EG cH LCd J
A
LCG
.ce gH aJ
LCG
e
e
Ie 15-12 Qj e MH TGdG aJ
e EG UCG GG CG EG ,fdG
-200 dG J .GdG e d YCG
1000 EG dG g JJh ,fdG Ie 400
j NBG dO 7.iNCG Wh edG HHP
LCG
LCG
Je
1000 ML aaQ e j e 1 H FW
Je
LCG NJ
LCG NJ hCG hCG GMG CG dG hO fdG Ie
EG eCG e
kfW
.mTJ
G
b Y IFdG bG g f eY
.H HYEGh gO OGOJ
LCG
B
e
LCG
SGH J LCG aaQ CG kHS fcP
e
aY e GPEG dh ,YCG J Hc GQTEG
dG
,fdG IQTEG 200 g dG G OhM CG
aGdG
FdG
G dBG
LCd
aJ CG dG gG IG d a
?fdG Ie 1000
Ie 200 aJ dG IFdG GG
dG
aG FdG
dG J Y J Y - Y bY fdG
LCd
EG oJ IMGh Hc IQTEG g .TGdG
YG RGdG L f j
ahG dG CG EG aVEG .aaQ IY c
.aadG e e dG Y GG
28

ARAP TASARIM 014-039

22/2/03

11:54

Page 29

j hQg

Ib ) bGdG dG
(G

G
NGdG

PG dG
e fdG dG
dG
fG dG
Qd

dG AGdG
fG

cG g j kUN kef ceH HdG Gg j .fdG Ie dGC MH HdG GfGC H aJ


. dG ge T Y H Je UN W g H, LH ITe JJ g da .gG

GQTEG e J PEG ,TGdG Y Y J jH J dG dH


J ,JdG e e ie EG J eYh .Gd d dG Y dG
.f AJ e
cM EG LCG aaQ jH j HfdGh dG GM Y f g
H ,G Y e Te H GdG dBH J dG dG J PEG ''JeJhCG''
LCG dG dG J ,QG Y J Ue SGH QdG e J
G QdG j dG g J eY .QdG e dGh dG dH
.SCG EG aO QhH dj dG ,cG
H cG dG e Y J dG e Y fG Y C jh
cM J jdG H ,''kJeJhCG kY kef'' IG EG :CG ,JeJhCG
8
.dG Gg J YCG Y iNCG QGfEG GQTEG QJ CG EG bJ hO dG
e H AGLCG VJ PEG .HdG GC e Y J dG YdG g GdG dBG J
.G Gg G GHEG g G e .G EG IU cM CG ODa ,ke
.G dG N fEG

29

ARAP TASARIM 014-039

22/2/03

11:54

Page 30

dG dG

aGdG Id dG dG
jGhdG J j H ,Og GW Y d SCGh YCd LCG aaQ J
e LQH LCG J .YCG EG aGQh eCG EG aGO Ib d a aJ Ie c
GdG Y gaJ fhG g .IG f Y Jh GQhdG Y gYJ fhG e
.Gd jQhdG bdG kjCG dJ dG FdG
,YCG EG OdG Y G Ue H IOLG dG g J :G S Y
YdG GP aCG J SGH J Y G cM X .G jhGR e jd
SCGh YCG EG IG J :NBG H .GdG AKCG kgEG Gk Qe SJ gh ddG
dG eDJ cG g ,AG fCG d c FGO H J EGh ,a
.SSCG
AGdG ehe G fGW IdG LCG GP GG LGJ e cCG J
AGdG dG ODj PEG .AGdG HfCG LQO H dPh gQG fSG dG
.c H LCG da N EG IdG GG g LCH
CG GG iJ dGh IdG LCG GP GG Y Lj dG Gd
g M CG hj AdG CG EG ,AGdG ehe Y d fdG Ie 1000 aJ
.dd iNCG fCG J fEG :CG ,AGdG IG ad ac Z gMd YdG
. iNCG dSCG ,SQfEG e IdG dG GfCG d :dP Y e
J K H MG Uh J j a.zdGh edG{ jWh SCG e
a LJ M,khG LCd eeCG GG J g MG fEG Yh MG
adG Gg dj .SdG dG dG G EG NdH AGd S Sbh U jGT
9
.LCG d aGdG IdG YJ dGh FGg N FGdG
d Id kN No UN NBG f g
Gk MGh kLhR GG H .AGdG HK VH
Y jFGO IY T Y Yh LCG e a
LCc AGdG fRGG J .fpRGG YJ gX
jQfGB
N jOdG
30

ARAP TASARIM 014-039

22/2/03

11:55

Page 31

j hQg

dG jZC G bdG g LGJ .fdG aQ 1000 d bdG e Ic c GE edG HHP


AGOdGh AGdG G H dGE Hj jdG LG IG g h .QgdG e dGh jdH
.dH TGC dGh L H j dG

LG e J fCG G LCG J c AGdH JQG EG ODJ fCG eh ,GdG


dG Gg j .gG Ga e IG ,GdG G j eYh fRGG J .
10
.Gg ej jdG MG j dG ShG
IFdG LCG Qq J N
FGdG UdG

M cM QWEG
UdG

GdG ie
dG aCG N

Fe IFdG
j QWEG
IFdG cM

QdG EG IFdG

kHK UdG e j
,aC G RGGh ShG e ke J Ie IGOC H I a dP e ,IFdG e Ie H 100UGC HHdG
.IFdG e H bQGC IQhG Y JQbh bQG fGdG Je gh ,Gd kGL jQhV dB G Jgh

31

ARAP TASARIM 014-039

22/2/03

11:55

Page 32

d G G

dG e ddG IYG dG

. Y LGC W e GG e jdG J
IQhG Y dG e j dH J eYh
W .eG IOLG AGLC G GSH
W dGB GC dB G g jeC G jG GdG
e , E6B JG JGFW Y ,jG LC G
GC dG hGC HHdG GC g H NG
a IFdG eGC ,ec H LGC J
NB G a G f iS J GC

dG Vh

UN feh fd JhH IOe e QdG id G e j :SQ


hjh.YUCGh dG G UGN J kUGN IOG g h.(SQ)YJ
jW Yh IdG fe H IOG g Jh.GG kYU g FdG SG
bdG IOYCG J Y IKDdG IdG GhR Yh bdG H fCGh dG cM
aQ AKCG a ddG ICdG g Hh,%96 Jh GL dY IOG g ICc Ea Gd.eH
e bdG g SG j G G cM Yh, bdG e %85 N j a G G
.bdG g N d JYh QdG GQL N c 11.jL

32

ARAP TASARIM 014-039

22/2/03

11:55

Page 33

J dG jdG dG j
Gg NGO IG dG
a jh ,LLdG G
GQ GdG dG
JQG e gGh
.dGh

OLG dG j
dG IQhG QdG GE
GFW K H J
JGe cGC J dGh
g GC GE . kTgOGE
b J GG
feGE e fQe Y
J Y HdGh dG
hO GC QG
G Gg .YdG J
V VH jo
Y J dG dG
e fQH KdG IFdG
.IG GG g

4270 m

F-4E

3660 m

F-16
3050 m

2440 m

1220 m

HiMAT
610 m

610m

1220m

1830m

2440m

ARAP TASARIM 014-039

22/2/03

11:55

Page 34

dG dG

GH UG dG RG
YH G S Y dG j .M e fQe Gk L dY YH GG J
CG .YdG eCG BG YH fEG S YS OJ YdG gh ,S/c 40
H IFdG M fMCG Qj eYh .KdG GFdG EG YdG g EG fEG j
.GFdG YS J YH ka J GG g CG jd kVGh j fEG
je EG IG ,Jc YS ad kUN Gk Obh KdG GFdG J ch
ch ,b cCG e Ic c J bdG g .fGW AKCG bdG e dY
e Zh HdG OLe OY Z J RL SGH e J gQhH cCG
.GG
Y d AGdG f f ,fRL Y ke jH j Gg dG RL EG
EG .SGH G Y Rjh dH ITe cCG j HdG Y dh ,FdG
AfCG YRJ b jedG jG EG dfG H j jQa cCd HdG LM
FGdG HfCG J M ,kGja kJ kef HHdG G g e edh .G
,dG GQhdG RG G g ch .G AfCG L EG AGdG H IG L
jN e N c OhJ (dG dH YJ) jedG HfCG e Ie T kjCG g Lj
.RdG cCH G
g e ITe cCG dG Gg H fGdG d fG jG J
.fdG IQhO 1000 dG dG g jJ Y dG Gg Yj c ,HfCG
e .Gc Gk e kef j CG Qda ,G Y dO dG Gg CG G e
RG j PEa .QdG jf YJ c QGWCG Y CG dG Gd dH kjCG G
dG Y H Id Gk e j d fEa ,Se Me hO ,ee H dG
.WEG Y gAH M hCG JM SG MG g OS
ah J OY Z kef CG d J BG M gVSG dG fCG g c EG
dG N Y gJ Ie g HHP c EG .HdG e e IdG M ,c FdG
Y'' dSG Yh ,Gh G
34

ARAP TASARIM 014-039

22/2/03

11:55

Page 35

j hQg

ILCG e MGh RL QJ J Y Y dGh , jhQGdG H J dG ''QdG


.HHdG dG
:J dH G H dG EG jdG c Lh Y dG Yj
p ndhn kH Hn Po Go or jn nd pdG ph Oo pe n Yo r nJ n j p sdG sGE p o nd G o p n r S an l nen
n p o V o S s dG jt GnC B jn }
.73:G | or nG hn
o pd s dG
n o n V o r pe o h o p n rjn s k n T
o Hn t dG o o r or jn GE p hn o nd G o n n Lr G

jQdG G

eZdG
O2

GC

CO2

eZdG e a

f j ,cC G e Ic c GE LG VH d eLGC HdGh dG GM dG dP e f


Gg Y e YC G .UN HfGC jW Y fC G GE Te H GQhdG dG G g c ke cC G
:dG IM dG
dG dH T e hM WN fC G GQL M hj .hdGE dG I FGdG dG -CG
.dG j dG QdG aT j
.e HdG cGC K fh RdG cC H IG L jN jhH FGdG dG J -

ARAP TASARIM 014-039

22/2/03

11:55

Page 37

...kHHn Po Go or jn nd ...
c dEG J HHdG M
dG dG dSCG
ej M jdG fCG
.''M '' fEG ,Gg
HcdGh GFdG
edG e kMOQ J
HHdG eCG ,CGd J K
dG AT e EG a
.G gKH

36

ARAP TASARIM 014-039

22/2/03

11:55

Page 36

sEGp o nd Go p n Sn
r a l nen
n p Vo S
o s dG jt nCG Bjn }
kHHn Po Goor jn nd pdG phOo pe nYo r nJ n jp sdG
kTn
o Hn t dG o o r or jn EGp hn o nd Go n n Lr G p ndhn
@
o o rnGhn
o pd sdG
n o Vn o r pe o hop n r jn
| jpYn
w p n nd n dG sEGp pQp r bn s Mn n dG GhQo n bn en
74 -73:G

37

ARAP TASARIM 014-039

22/2/03

11:55

Page 38

hCG FG SCG Y dH HHdG J CG


I eGbCa ,YS I dG Y HK J
GPEGh ,dGh GQGh LdG Y bd
IOLG UG FSdG Ea ,ac JHc J
OGOJh ,dG Y kSe kabh eDJ eGbCG Y
.UN FS RGaEH FSdG g Ib
AG e ''dG'' G HHP J
.dhJ CG b dG hd dG
j bG e dG hO HdGh
RGaEH dP jh ,L QN dG
FdG Gg j .dG Y Vg FS
IG J FS EG djh dG
OLG AGdG IG J K ,UeG
dG kjCG dG SGH L EG FdG
.H G WG EG FdG

dG AYCG j HHdG J jGdG .dG e Ic LQO Y IgX G HHP J


,eeCG G RGdG AYCG Vh jH dPh Gd gCdG Vh NCJ K ;GdG eS
SM SGH gO dG AdG YSh jdG G Y Ie bEG jhGR H J kGNCGh
,fdG e 100/1 e cCG J dG g CG EG .J K QSG hb Y IOLe
.S/ c 10 EG J M YS IOjR Y IQOb HHdG a
kGQe fGW AKCG J a , Sc ''GdG GdG IS'' HHdG Y j dG Gd
CG Y H CGh ,a J dG G e kjOY bEG c ,bQN jH AGdG kLe
jdG IG g H J dG iNCG FG eh .Fe Y hCG QGfG Y dG H S
j UN H N fCG EG ,SCG EG J CG j HPG fb a .SCG Y abh
eJ eY kLd kFS GOSdG g RJ ,aT GOSh eGbCG ShDhQ Y Lj .ke G
h jJ M e J EG eh ,e J eY dG H fS IG J .dG
e ej kGL kbQ kAZ jh G e e c f j ML G HHP .H
GdG Y cj a dP e ,NBG Y e H MG jg e c j CG .G
dG J .MhCG G GP GFdG G g c ,MGh Q Y Gh eCG EG
LCG cM Y dhDG
38

ARAP TASARIM 014-039

22/2/03

11:55

Page 39

SOGS Y H 6000 e G HHP Y J


jdG g e c NCJ ,''jdG'' SG Y j
Yh a ,SdG ,G hCG eCG EG :k e
e dM e c HHdG iJ CG .fGG L
AT H J CG :NBG H .G L
YCG fK J .LQO 360 jhGR jDhQ M
GHh ,jdG g e IMGh H Ad e
48000 GM dG SG jG j
.fdG IQU 100 dJ CG N

G dG e GdG FdG JQe HHdG J


IOLG IOQhCGh dG jdG J .LCd
IG Y j kGL SM GT LCG Y
.fG dGh AGdG G j

CG EG ,GdG jGH YCG J fCG e ZdG Yh .dG Y Jh bEG Y


.ILh Ia H JeJhCG J G e dG g cM
aOU GPEa .edG EG GdG ee H SCGdG Nh LCG IOLG SG J
,edG EG G IjG eG H SG g J ,fGW AKCG kGjL kFGg kGQJ IG
Y dG e HHdG J jdG H .jG H LCG LH dG GC J SSCG Yh
HHdG .SG bdG eBG e EG dGh ,VEG FGg QJ dH IjL IM CG OLh
AKCG G bdG e Ie e cCG GdG G bdG Jh ,fdG GG e ML
bhCG idG bW j CG fEG C ,kGL b fCG f CG g e .MGdG
dP EG VCG .a jOdG IG YCG G bW VCG IY EG J dGh ( QGdG) dG
dG HHdG Y Y ,YCG c a FbO d bdG g U j CG fEG CG
12
.f YdHh e ae bdG H aJ CG c ,YS f I IJdG g Y J CG
39

dG dG

dG dG

GdG ILGC
QdG :G
e QJ fCG ,NBG Y QJ b QdG CG Y hj jdG ,jQq dG Yj
,G Gg Y dBG e dBG CG j CG YdG Gd ,dP e ZdG Y .MGhdG
QdG LCG J k jGH .YG SSCG dP Y ,jdG H Y kc j dG
bh .gJ Igc jdG g Lh Gk c kFY a FdG UG J dGh
QJ KEG feEG Y b dG GYH ''QhQc jEG'' cdG QdG OCG
:LCG
e fc GPEG a J CG fCG g LCGh YCG H R dG GG e
.J CG f LCGh ,iJ CG kf IQG dG :NBG H .QdG
13
.VdG j Gj eCG Ga ,dG EG AYCG g AL c eCG
N e LCd G dG a J dG dG Y j dG GDdG jh
iME eeCG dG a d dG dGh ,GL hO dG GdG e S
d LCG OLh CG EG .HCG EG dG Z S QY L EG MGhdG
jdG J .kGFW j M VQCG d MdG G

40

en n rpr jn hn mas B n U o bn r an p r s dG GE Ghr n jn ndhn GnC }


| l pHn Am r n T u opH o sfGE p o Ms dG sGE p s o op r jo
19 :G

dG dG

g G S Y FdG Y .GdG FdG ej dG dBG e OY EG


dG dGh Yh FdG H dc ,e H jdG dG J e NCG
cCG QdG Y GQhdG dGh .jdG IOLG J Y kjCG J dGh
Ggh .''cGJ Y'' jW Y edG Y LJ CG dBG g d .kJ
.d e AaL GAYOG QW EG fGG H J dG YGG J

FdG jQ H
aNG J CG ,MGhdG g QdG OGLCG CG YJ dG QdG jd
a ,MGhdG Y kH kaNG QdG J .QdGh MGhdG :dG Jg H FdG
.QG dG GhP e j Gk ja kJ kefh ,kGL Nh a Y e dDe g H
QdG H Ia gh ,fGG e gZ hO QdG GfCH UN iNCG Ie g jdG
.gRLG MGhdGh
''jdc N'' IQY Jh ,QdG H j dG dG MGdG cCG e TjdG J
.jd IG dG c Y

j hQg

jG e J e IOe JGdGh .JGc YJ JhH IOe e jdG J


dGh G e dG dG IOLG AGdGh cCG QOe e Lg dG dG
J Gk L e J jdG J EG .IjG jG eCG G d
Ie jS H d '' :QdG jQ Y Sja BG dG j .jQdG dG AV Y
14
.''iNCG Sh CG J CG SCH GdH J
e H g jdG CG'' kjCG YG Sja CG EG ,QJ Y fCG e ZdG Yh
ejdG e e T GPh jb ,N fC ''GdG e kjJ e H
15
.dGh aG e Ie H dh ,FGdG
ShhdG jQ L d H ;H dG Y f jhQGO dQJ TjdG g LCG d
cJCG'' :1860 f e Y ddG Z RBG jU EG c d .(db M) ''kje''
''..BG Gg Y J fCG EG ,dG H J CG e Mj IOhdH QdG c M ke
:Hj K
TjQh ShhdG jP e EG .JQG H TCG dG FH aCG eY BGh ...{
16
z!VH j

aGh dG
ch .gdG S G FdG jQ H fMCG j eY
j .TjdG bd dH cG J FQ b g kL f
c dG e G FdG dG g Y
G JhG ` dG g Oh ,gG
c h ,Fd FGdG ejdG ` edGh
YJ dGh G e BG b
,IOG dH jDhQ gh ,dG
jT SGH H e dG g HJh
,aG g IY dJG jW Jh ,aN
''qdG'' G dGdG Sj dG dG j H
43

d jdH L j YS 3-2 H UU
.eRdG adG d

.G a fGSGC H e jdG j

dG fL e fL c Y b 650 Y IMGh aGQ jQ :G S Y


J dG g e b ch , b c Y b 600 Qj e jh ,FdG
Y dGdG SCG HJ c H e aG HJ .kaN 390 SGH gZ e
.bGNH Y G AGd M J H jdG H dG HJ ,fL
VdG j CG Fd a ,SCG e S C H Y aG fG GPEG
.VfH hCG ,Q jH eEG jd dG
j d Gd kFGO IgLh Je f TjH j CG FdG Y
fU jdG SCG IOLG jdG IdG IOY FdG j .IG QGSG
J eY dG e J fCG c ,Jh TjQ QdG J jdG Gg SGH .TjQ
.IWG AGLCG J hCG J hCG
QG G eCG ,OQdG G dG e FdG L IQGM LQO TjdG
17
.JOhH d H TjdG a

TjdG GfGC
G Y IOLG Tjda ,FdG L Y jRJ M TjdG FXh J
ch FdG LJ Y TjH jdG j .jdGh MG Y IOLG J Y J
Ib IOjd GdG AKCG dG G SJ Y G TjQ J H ,YdG
Qhe d H e TjdG J ,SCG f e LCG aJ eY .JQG
H Y IYe TjdG J YCG f G Y LCG J eY dh ,AGdG
18
.dH AGd S
Ggh , GdG Y JQH d dG e e Ga N TjQ QdG J
.kGQa KdG hCG HG TjdG GSG j
45

:QhdG Gd G QGWC G dG cG g QJ
.dG ,dG GdG ,bE G

LC G GHGE
e Gd FQ TGC KK g
T ,dG :SC G GE YC G
.HSh ,'' V ''

GE ,fGdG UH QdG e J
a ,IMGh jH J fGC
Y Gd gD J bQN GQb
.VQC G e kGL jb ae
.FdG NH G T j

AdG jQ
jdG P G
.jdG

dG jQ

OD J
TjdG
,e IY
H U bh
kN G TjQ
a jdG eGC ,Gd
LdG Vd U
Y IYGh
.dG

.SQdG FW L

,QG AGdG e dG jh ,G kWb dG LGC IY SQdG Lj


.LC dG dG H AGdG edG e Ia QdG J GC H

dSC H jL NB H dG jQ GSH QdG J


Y '' :Y j jdG Ggh ,GfC G NH J
.IdG b J dGh ''TjdG W
SGC dG Y OLG jdG j
HWdG e QdG LC Gh Gh
Y gYJ c ,IOhdGh
J .AGdG dG
Y IOLG TjdG
G fG
YJh YdG
J Y
LQO
.IQGG

FW jQ
SdG

HGC L e jQ
jQR

G FW jQ

G AfG H
Y AGdG V j
VGC dG dG
,dG dG Y e
FdG aQ GE OD j
G c GPGE .AGdG
AGdG aJ E a ke
kGjGJ j dG Y
Ib j dG
jdG Hh ,g
IQdG fG )FdG j
.(dG SGC

.o dG FW L
dG L

.G AfG GE J AGdG G

dG dG

IFdG dB G TjQ
Yh FGg ScCH L Ohu Ro d .d N fCG QdG LCG bG j
FdG d FdG dG Jh .dG RdG e dHh G Rh e d a
.M EG dH Gk L N a TjdG eCG ,L j dG FGdG AG W Y
:iNCG J IMGdG dha FdG IG dG g e d

dG dG -1
e ZdG Y JLMGh H e fG jZ FdG L H j dG IdG EG
V S 18 dW j dG jdG dG FW j :G S Y .aG eY cJ
,QdG Y H e QdGh dGh dG Y J .jdG IQH d c 68^5 Oj
dG dP e aCG ''J'' gh
d W e

FdG aj eY Jh LG TjQ J.TjdH e FdG L


Qh.YdG J eY LG ShDhQ Y TjdG aJ. MH
.dG jh Jb FdG j TjdG a AGdG

YGO IYCG

jh
bG ShDhdG Y TjdG JJ
AGdG .YdG J eY LCd
Jb FdG jh TjdG N e
dG d

IFW L
a Y

,ke N FdG Y
fc H jb dh
AGdG Y h a
YGO b h NGO
.HU e jJ

G TjQ

YGO IYCG

jh

TjdG IYb J
jdG G H
48

j hQg

dG iNCG GG e .FdG H H J dG IdG Y gj gh ,jdG


dG dG dG e NCG fCG ` kdS fcP c ` Fd dG dG H j
,LEG fRh e % 4^4 ed dG dG Rh j :G S Y .jdG
.TjQ Rh e bCG CG a keGZ 118 (H FW) bdG FW Y Rh j H

dG dG - 2
hCG dG :SCG Id jdG Y ke H QdG Y dG RG j
cCG c J :G S Y .FdG j dG cCd WG LG EG Oj
Lj dG dG V jY FdG Lj dG
c J CG jdG Fa .fEG
FQ U dd ,QdG L dG cCG
.ke H QdG
j :G FK jdG AGdG OJ j
Y bjh GdG e T Y MQ AGdG
cCG OJ Y NCJ gh ,IU FGg ScCG
G AGdG j .fe HdG cCG Kh
FGdG dG f keh FdG kcQJ dG QG
.MW j M
QdG Y dG Ea ,dP e dG Y
QdG Fd UG dG
jh L e dG AGdG Nj M G OMCG
eYG gOGLGC Y ke k kjJ QdG
jJ aJ dG gh .iNCG L e G AGdG
Y kc je H FdG FQ J .''MGhdG''
J .jdG FQ ah J dG dB G
JLM j ,QdG Y cCH Ie
eGC ,f dG jdG e MJh AGdG jdG
LddG j .J dG IdG bdG d
.He a e jh AGdG Na QdG
jd H hG jGO e GSCG
e Ic c Fd d jdG dG Gg N
.GdG dB jQhV J dGh cC G
:j c FGdG FdG jhQGdG

49

dG dG

HfCG EG QdG Y eZdG dG GC j


dG YdG g jdG h .kGL IU
kfGQhO kef d iNCG Ie dG dG J
...MGh H FdG N AGdG a
GfCG FGdG ScCG OLh e ZdG Y
QdG Y jFdG dG CG EG ,MGhdG e e
eZdG G Lj .ke ja Y H dG dG Yh
eeCG
dG J J jQdG Y jFQ H CG
eZdG G
H de dG g CG Y IhY ,QdG
G
19
...UJ
dG G
EG'' eRCG jf'' Hc jGO Vjh
OMGC FGdG adG FGdG dG f j
:QdG jW Y G dG Gg J
q dSG
H J K AGdG dG g .G
kGe kaJ jdG g J .FdG GE e dG Gg CG jJ Gk L dG e''
Fd dG Gg N d .FdG GE dG AGd dG dG Y ke G dG
.cC G e IdG dG GE FdG LM
,QdG dG e kjQJ QJ b jQdG
Y G CG H fNCG e GPEG UN
Oj b kH c e N CG CGh ,Fd dH e hCG IM Ve dG dBG
GPEG fGdG dBG c J CG TjdG CG ch .FbO N JH
Y FGdG FdG EG dH eCG dc ,e H aG e dG HJ
j dG dG T RG j e dG RG c J CG ,FdG
jH LhCG b dG HfCd FGdG YdG J dG FGdG ScCGh
20
''.ee H dN e J CG
dG FdG C ,e eCG FGdG dG EG QdG dG e FdG QJ Ea :QNH
M dh FQ hO M Fc CG j CG PEG ,Ya FQ J CG dfG M
GW d dG je j CG G Y Lj WHh dd .FbO d
.JM J FGY
eZdG
dG Y
bhdG H G

50

j hQg

IdG dH OhdG j e J Y J FGdG FdG Fd IjdG dG EG


d H SMEG e cCG EG eCG j .fGW FdG Lj dG cCG e
CG jd kL UCG Ggh .jOGQEG Gd WYG f d Fd KG Z jdG CG
.IQdG H N dG g dG CG Y J dG FdG e NBG dO g FdG FQ

RGdG f -3
J G gh .bG bH CT fCT N CG hO J MCG QdG dG N
N MG CG j UN J QdG LCG N .UdG e J c
AKCG FdG j M N RH d nu Uo ke dG SCGQ .GdG AKCG RGdG
.a L Rh e %1 FdG SCGQ Rh j Y Hh .GdG
,FGdG ejdG e SG TjdG H ,FdG id iNCG RGdG FN e
.FdG RGJ Gk L a H LCGh jdG TjQ UNh ,TjdG J M
Y VfG AKCG ge RGH j dG dG FG g RYEG j
. !YdG c 384 YH gT Y e ja

bdGh IdG e - 4
Y J ,AjdGh AdGh AMCG Y OGG e S ah J Y c EG
H d bdG e e c Y G'' :QNH GC G Gg j .''bdG M'' GC e
.''e
.bdG M GC e Y kVGh ke FdG GW j
j fCG EG ,MQ AKCG J bdG e e c QNOG ILG QdG Y Lj
FGdG RdG W j CG a eCG j eh , Rh NCG bdG H J CG
LQO bCH ObdH MG e
J CG j ,Jje fOCG ObdG Rh j CG j M :NBG .YdG
.QdG eCG kFY J dTEG g c .Je bCG bdG

51

dG dG

jT AH j CG FdG Y c GPEa .Gd idG YdG j g hCG IG


bdG Ea Gk L dY YH j c GPEG eCG ,AGdG AdG Y d bdG SG j
S de YS Y G j fCG j Ggh .AGdG ehe Y dG J
dG d FGdG ejdG dG Y OYH .ObdG e c bCG SG
.QdG GfCG d dH de J G YdG Ea ,LCGh
pluvialis dominica fulva: OdG gdG GbdG FW Y bdG e BG d
Jg jW CG h .g AdG a d GhdG QL EG SCG e FdG Gg Lj
Ggh ,jf M MQ jGH e (c 4000 CG) e2500 b EG j fEa ,QG em N
.YS 88 J MdG g CG EG f CG H ,bJ hO L HV 250000 j
gO ( 70) e fhCG 2^5 , 200 Oj e CG fhCG 7 MQ jGH FdG j
AdG M c ` GW YS d FdG Lj dG bdG CG EG .bd Qc j
ObdG kf g CG CG ,jd fGdG YdG j Obc (82) fhCG 3 hJ `
CG b Obh hO eCG e j CG FdG Y CG j ( 12) fhCG 0^4 Oj RdG
.Ghg EG j
,H Ghg QL EG gdG FdG j ,HG g c e ZdG Yh dh
?Gg AGQh dG a ,S c OYG c e CG LGe hOh
fGW e MdG g e jW J CG N dG QdG g EG J dG MhCG
QhH dG Ggh ,S V dh FGY H QdG g J .kYah kS Gk eCG
bdG e c EG QdG dG Gg Hh "v" . M H AGdG j
(7-6) fhCG 0^2 jd jh ,bdG e %23 f aJ Ggh ,LGJ dG AGdG ehe
Gg . dH ?d e Fa IFGdG bdG g g dh .nnG eY gdG e
21
.ce FGg GQJ dG LGj eY QGdG M GSd FdG
:SCG g eeCG J gh

52

j hQg

?eRdG gdG hCG bdG c QdG J CG c


?GdG b eRdG bdG c QdG g ueDJ CG c
?d RdG ObdG ch MdG ae CG c
?SCG e aCG Ghg jG hdG CG J CG c
H J hCG ,HG g hCG ,eG g EG QdG UJ CG G e
j e c Y dJh LJ Y Ib :dEG MdG fEG .HG g Y AH dG
QdG H J iNCG jW EG gfG dG BGdG j dc .dG Gg jQGSG d
:dG dEG g CG AMG g Y OLG GQOEG Y jBG fJh ,( msaBUn )
m as B n U o r s dGh p V Qr nC Ghn
p Ghn n s dG en o nd o u n jo n dG sGnC n nJ r ndGnC }
o nJnn U n p Yn r bn w co
.41 :QdG| n on r nj n pH l p Yn odG hn on pr nJhn

| l pHn Am r n T u opH o sfGE p o M s dG sGE p s o opr jo en n rpr jn hn m as B n U o bn r an p r s dG GE Ghn jn ndhn GnC }


.19 :G

dG RG

W d .jdG MdG GSGC V dG QdG J


.bW e %23 Qj e aj GC '' V '' dG

53

Gd ,Ic Ib EG GdG j
e f cCG QdG dG
H G dG /dG jG
Sj dc .WEG Y FdG
ddG jG e jd SG
Yj kSh .dG Id
JJ eY FdG Gg Y SG
jfa 50 L IQGM LQO
.(je LQO 10)

J dG JG IQGG g .je LQO 42 M L IQGM Jh ,bdG bO 460 QhdG QdG b j


Gg jW Y FdG ej dG bdG e dY je dJ j M ,FdG Ad kGL eg ,jdG AMC d dH G
.jdG SG

LQO 42) jfa 108 ddGh ` G S Y ` Qht dG FdG IQGM LQO J


HSG dG YS ie Y ,(je LQO 43^5) 109 ddG QhdGh ,(je
dH Gk L jQhV ,iNCG AMCd dH Jb J dGh ddG IQGG g .jd
.FdG Ib e dHh bdG SG e jJ fC ,IG b Y FdG jQGS
jj :G S Y .de jH eW H bd WG LM H QdG J
f J H .dG e eGZ c 3 dhJ Y MGh GZc QG dG U Rh
dH eCG dc .dG e eGZc 10 d MGh GZc jdG IOjdG
dG j H .ddG bdG e e aGd U dG QdG Y GQhdG Rd
Qht dG FW Y bdG N 460 EG dG g JJ ,bdG N 78 dG
ILCG g c Ohj dG FdG be FGdG FdG Jh .QhdG Y 615h
.d RdG cCH AGOCG jdG
54

j hQg

J bdG SG Ic Ya J a ,Ic YH bW QdG Jh


2^5) G Qdcc 4 fdG j :G S Y .d jdG SG
.Qdcc 41 U jK GM j H ,(edG
d Mh .jdGh QdG H dG J CG Id
IW jW Y KM FG g MCG CG kL S
IMGh UN Ea ,dH e VGaG gh ,FGY
dJ dG SG EG .kT J de J
e CG j bdG e dY je Y
Gg CG Y IhY ,e FGg FQ hO
jdG f e Ife Gd j b
dG RG Y IW CGW GPEGh .cCH
FdG j CG j Ga ,iNCG ILCG b
Sh ,LM j kcCG
g c ke IQV IOjdG
c a .dG d dH eCG dc.dG
c e e HSG fh FGg FH Gk FdG
FdG Ib H a .GdG Y Gk LY a ,JLMG
d kf ICGGh dG dG H GdG VQCG
fdG W
j e J EG kjCG LCG J j .dG
.CG
J G e :IMGh f f FG g c J
e BG .QdG jf hCG OG dG N e QdG UCG
Y kc dG N :NBG H .''IMGh bO'' fG FN H No QdG GfCG
.In Mp

55

dG dG

G GdG dGB
e q fGW dBH ,dG Mh SQdG e QdG GfCG c Lh Y dG L
H QdG No a ,bdG e dG j GdG CG h .jdG e IOSG
MhCG a .eLCG Y QdG N Ie J h .N Yh Ic bh jb QU
.d eRdG bdG e J GdG e jW JG QdG e dG EG dG
j CG PEG ,UN IQ j gh ,SBGh jaEGh HQhCG e H FW SdG
SCGQ CG EG AGdG LQCj L CG eh .jdG LGe fGW AKCG HK VH SCGQ
GC G Y .Lf j b dG cG c e ZdG Y bK jDhQ d j kHK j
j dd ,QdG HG dG fRG j dG ShG RL j f
22
.''fRGG UdG SCGQ'' d SdG SCGQ Y AdG

bdG dGB
Y iJ CG SdG Qa .Fa IQ H UG dG hL QdG J
dG bh eGj .YS c OUd Jh VQCG IOY J GdGh ,GdG
J g PEa ,dG M Y cCJh QdG N a ,GdG Y e SdG Y
kahh ,eRdG bdG e J jdG g .kN kfRh d j ZQa I QdG
23
.jdG H bW e %7 f aj FdG Ea ,AdG H

56

j hQg

AGdG dG
Qda .AGdG eGSH G bdG e J e QdG J dc
M AGdG e J CG Jh ,LCG a FGdG QdG IT e jJ eY
.d dH aVEG Ie IYdG AGdG GQJ Jh .jdG GQdG
SdGh ,''dG'' fGW AKCG bdG ad FGdG GQdG FdG GSG Y j
.QdG Y dG FN e J dG feEG EG .IQG H J dG QdG MCG g
Y dG AKCG AGdG Ad eRdG bdG aj fCG hCG :SSCG JFa d
j . GdG ae IOjH Fd j fCG fdGh ,VQCG ja Y VfG hCG dG
24
.dG AKCG bW e %70 aJ e SdG

FGdG GQdG e bdG Y G


,JG b e Qj dG SdG j :jH FGdG GQdG QdG J
.QG dH Gg Yjh ,FGdG GQdG e WdG G Uj dG SdGh
J dP eh ,cdG AGdG e Le J ,JG b a jb jQ eY
GQdG g e jdG QdG e Zh WCG FW j .dG g CG QdG
GOG H gJ dG GQdG e j MCG H h ,QG dG cdG
.SdG ba j dG ,dG e
.Qd aGdG GQdG FGdG G J
jh .adGh MCG G FGdG dG H UdG dG dG g Gh
dGh G g cG .''IQG UdG'' dG SCG g Y QdG Y
e hCG QGOGdG jW Y hCG dG e eOdG FGdG GQdG H WGdG Y J e kdZ
:dG e GNBG Yf g .GSCG T Y a QJ gh jdG QdG bGe N
.ejdG dGh QGG dG

57

dG dG

Lh Y IQG QG We QGG dG IgX MJ


VQCG J ,VQCG EG dG TCG J eY .UG
bCG j AGdG j eYh .d eG AGdG j gQhH
kjCG IgdG g Me .JQH NCjh kfRh
.FGdG GQHdG hCG edG UGdG

QdG Y dG jW
LG e IOSG UN jW QdG
e VQCG e e d dG Y jQGG
Le G jQGM Le e J a . Se Y
Me a Ggh dG GW iNCG jQGM
.MGdG dG Ic
dG Y FGdG LG GC J
J eYh ,VCG GQdG ee dH khCG IdG QdG J .JQH
M dRdG GQdG g eCG H kdZ QdG J .kM cCG QdG J ,GQdG
V FGhO V QdG .FGdG QdG e YS cCG aGdG GQdG VJ
.QdG ce e J dH ZJ eYh .HPG Ibh kdY dG H RGdG eDd
GQdG g ke dG a ,IQG GQdG dG QW e iNCG kYGfCG J
jaEG EG Ljh SdG HQhCG HCG dG j .UN H IdG MQ NdG
kee Oa H Lg GPEGh .( c 7000) e 4350 a j MQ g AdG d
HCG dG CG EG ,bCG Y Ge HQCG MGSd bj CG a ,LCH aadG jW
YS 7-6 I IQG GQdG kee a HSCG KK N MQ j
.bdG c aJ EG Ljo Ggh ,dG

58

j hQg

g Ggh ,QdG a NdG GQdG J dd ,VQCG e bCG YH AG j


QWh dG j .jW MQ J eY QdG a LJ QdG j dG dG
eh dG VGQCG eEG ,jaEG EG Jg jW J CG HQhCG SGhCG J iNCG
.QW L e a jjEG IjG hCG ,QSdG
GQdG iNCG jH QWh SdGh WCGh ,SQdG j ,iNCG L e
IhQP Y IOLG aGdG GQdG e QdG g J .ddG LG J dG FGdG
EG dH Ja jdG LGjh j FGdG GQdG SQdG AKCGh .GeCG
AGdG Gk e 15-10 JQG H Hh ,YCG

dG dG
G SCGdG hP dG

jZ f

EG SG dG j
dG b G
dG

''jdG'' f YGC G G Qj .IQG GdG dGB H dG ae b FGZ GE QdG J GC


dG ddG YdG e ZdG Yh .dGh SGC dG G dG e c fG YSG L Y VfG hj dG
L GE dG j GC .SG bdG G GE Ud J fGC GE ,S/c 40 J dGh dG H R
.S/c 70 YH FbO K N SGC dG G dG dGE j H ,bO 4^25 N c 3^5 J

.jdG gG J e bdG Y QdG .dG jh jL e gG j


GQJ SQdG LGj dG Gdh .AG S eJ eY YS FGdG GQdG J
kHe jL e Qd Oj ,SG YdG j CG Hh ,dG VhdG ibCG
.dG S e
QdG e jdGh ` MG jW H FW gh ` AG L j
.dG a dG f SCG ,iNCG jdG
KK W Y G jdG MH SQdG FW j
Gg j .dG QdG cGC MGC QeGC
bdG e dG GE dG FdG
SGC j a dd ,fGW
jW ae d dG
e dG SG Y aj
.bdG
60

J H GdQ
H e

JQG
Y
GeCG

Je GW
Qe
QG dG
nnJd
e AGdG Jj
rnnJoG JQG

AGdG cM Y j QG dG
.JG b f JG
dd a , dG e ML j dG jdG GE e HGC FdG j
eGC , jWh kGL SM Qe e dG AG .ja Y kH Uj
AG S e kGL kjb GdG e e j jH U a ML
e dG AG H FdG j .H j GC hO jW bdh
.dG j ja GC d Qe

Je NS AGg
VQCG H G FGdG QdG
rodGh
o
jhQ GdF OGN GdQ
GdGF

j QGG d eGhdG M P
.IYCG e c HS IYb fe

FGhO V JQG

jG QdG f QGfG

G dG ac J .ec 8 JQG M RhE G FW j


JQG Gg Yh ,%65`H dG S e bGC c 5 JQG Y
jG j ,cGC YH MH aj GC FdG Y Lj
e dG Y FdG Gg FQ U ddh ,cC G e
Gg e aGd cC G e c cC H d ,jdG
.JQG

a OdG QGG dG P
.IQG WG
e H j FW
IYCG bJ
FGdG

aGdG idG N

L AdG Y a UdG dG
.FGg

dG
dG

GG GE dH G Qf j GC
c ,fH QTC G hL NGO G bdGh
GSH QgdG MQ fdG QdG GC
.bdG cdG fd

AdG dG
AdG dG

FK jDhQ

FK jhD Q
G SGC dG fL Y IOLG dG OhJ
AGdG WG) SGh jhD Q
.(dJdGh

dG QdG J .dGh jhD dG g QdG kGQJ SGG cGC


dG J H ,IQe jhD Q SH QdG eW OJ
AG OJ dG QdG H .jb S SH dG OJ
a ,AG jhD dG e I Y H I J
dG VJ .aGC jhD H j kJ cGC gY dG
jhGR jd J dHh ,SGC dG fL Y QdG e Y
OJ dG QdG SGC Q ee dG q VJ eGC .Ic jhD Q
G jhD dG GE fC G dG e a ,dG
hGC G jhD dG GC e ZdG Yh .edG jhD dG e cGC
eGC h kY cGC J fGC GE ,V jhGR J FdG
.fE G Y jhD dG e ,kGcJ
G JH d J ge iNGC kSGM QdG
Y IhY ,J dG GdG GQe VQC d WG
.GdG AKGC AGdG GRGgG GQOGE Y JQb

QeC G FW j
Qhjh Ic YH
a dd ,YGH G
SGh jhD Q j
e Z e cGC
dG aJ .QdG
Y IOLG IdG
G Gg SGC dG fL
.jhD d SGdG

d dH dG SM J
Oj GC .kGL eg QdG
e G e OSC G dG
dG SM H IH ae
.Y IQG

ee edG Y LJ
k d aj SGC Q
.dG H jhD d kGR
OLh e ZdG Yh
jhD dG YGC M
Y KD j fGC GE ,G
GC j fC ,FdG
LQO 270 SGC Q jj
.G G d

Y-iNG AMG e e- QdG LL


cG UN K a a a bOh ec J
dGh dGh jhD dc SGG

fG K

Oj dG FdG j
jhD dG H J IOM jhD H QdH eW
ndG nPG ofE G ij GC .jdG
H ,cJ hOh HV H
e JGP GG dG ij
.cGC VH f aG

dG j

PG Ga

Y dG IaGQ J
GdH d QhdG FdG
dG Gg j .jW Gd
.QdG Y

Fd G TjdG H IQdG VJ
VJ .jG MG N e dG
GP j H a IdG TjdG
.ejOhjGE FGa

dG dG
ea ,G j kjQhV j Ggh ,kf fe Z QdG Y QdG Y
.dGh GdG AKGC QdG dG M j ML dG j
dG TjQ

G OGb

IdG G TjQ

G TjQ

FdG

FGdG dG

fdG
jG
dG

dG

IG
Se FW

'' IAGdG QdG'' J


jdG fH edG e
Y gYj ,jdG JYh
jW ae G
H .Fa YHh
SG QdG J
OYh kf IU LC H
.jdG cd gD j a
fG ,VJ SQ Lj)
.(dG

AGqY FW

SC G GE LC G J
. G dG H
Jh MG Jj eY
,IdG QdG Y

Y
Y IU jQU
Ic jQU

edG dG
MdG ,Gd
cdGh

,IdG dG NJ
IdG J eYh
jh IdG NJ
.MG
fG ,VJ SQ Lj)
.(dG

Ic jQU Y

dG Y
VG Y
Qe

IbdG Y

dG
fE a Gd N b QdG GC
,H eh a keY
hO RdG G Ie d aj
.AGOC G Ib Y KC dG
.(dG fG ,VJ SQ Lj)

edG
QdG Y
dG HUCG
MdG Y

J Yf Ib fEG RMCG
G .jdG dG GdG
SGQO g dG d FdG
.QdG G ed AdG
dSCG e jdG Ic
ghLh dG ejOhjEG
J Y QdG LCG
J dP Y fh ,GFdG
EG bJ dh dh ,kGL Lf
e dG C ;dG ie
.G dG N Y

ond p sdG pd o r nG }
ondhn p VQr nC G nehn pGhn n dG
s ne
op nG n go hn pIn Np B G o r nG
nehn p VQr nC G op nj ne onr nj @ o pnG
n pa oo r nj nehn Ap B n dG
s npe op `nj nehn rpe oo r nj
2-1 :CS

| Qo ondG oMp s dG n go hn

YS EG .cG e jH P QdG GW
MdG hCG cdG VjH Jge GdG
.bCG bd cSG j dP eh ,SQ dG

S 55 H dG LCH dG edG J
SCGQ ee gY VJ .kYQH kd kUb
dG Jg ,ja OjEG dS d j
fCG EG aVEG ,dG jDhdG Y IQdH GJ
,IFGdG HQCG KK SCGH jJ CG J
d jh jd jDhdG e q Sj dG eCG
. Ha Y gQY e
dG dG e J CG a kGL SM FdG Gg fPCG
LCG GPG QJ dG OdG dG Y J
,U QGUEG hO LCH edG aJ . IOLG
Jh jH eEG Y IOG d Jh
d N G Gg CG if Ggh .QTCH
.kSe kd kSe

j hQg

FdG H dG
gGj H ,IdG JGQe hCG TjQ hCG LCG Y QdG N RYEG j
.VH EG
HW GRZ J e 15000 Y d dH jOY hJ dG LLdG H EG
dG g aJh .G EG IdG Vd SEG dG jDhQ .ddG
.ed ehe e jJh d fhe SEG
GfCG c aJ a ,JGP H Ie IMh g dG
dG QU Nj .NGH G Qd RdG AGdG
j H ,OGh edGh ,gdGh ,JhdG
.VG FdG QhO VdG
cCG j CG EG G jh
EG kjCG j ,HdG cCG K jh
Ghh eY d ddGh ,IQGd Qe
J .FjdG edGh jdG e Fd
e j dG ,d Gg c dG Ib
aJ H ,H j dG FdG dG N
cCG dG Gg j dG jedG YhCG
.HdG cCG K Y NCJh d RdG
gh ,e dh ,ge M EG bQ dG Ib
.G EG VG dGdG IQGM J FG H

69

FGg Ga
bdG dG e LQG dG

QhdG dG

c R

%16 fG Ia N dG J
H .Ae QH T Y FG gG e
bdG d NGdG dG
PDe Gg CG hj jW Id AdG
GHJ
UCG G
.dG Ib IOLG eG Sh
Gg CG XCG IjG HCG CG EG
j M d dH QhV dG
IdG
EG UdG j .dG e hG e
j e d Gh d cCG
TdG UdG
AG Jh ,dG AKCG dG d j SCGQ
dG e
.G jg eDj
f M de hX X %20 EG 15 H e FG dG f J ,dP Y IhY
e Z e cCG edG
q UGdG FW VH FG dG j :G S Y .dG Ib
.Ge H kaL cCG hX J dG QdG
BG

a eJ dGh ''dG S'' j e GdG


Jh ke dG b J dG g .dG e d
.dG H

70

I G j j b dG Z
S a dP eh ,fG e kej 20
.j GC dG jj eY dG

jM

dG

QU
dG

j dG JhH
g RGaEG

RdG
IdG FZ RGaEG j
dGh dG M

SdG dG

jdG IdG
G

rRn ndG

J Yf Ib fE GRMGC
G .jdG dG GdG
SGQO g dG d FdG
Ic .QdG G ed AdG
dG ejOhjE G dSC G e jdG
,GFdG J Y QdG LGC ghLh
GE bJ dh dh , kGL Lf J dP Y fh
.G dG N Y e dG C ;dG ie

fG e j dG J
edG MG ,IQGGh AGdGh AG :a VJ dG hdH J dG fe EG
.VG dGdG Rhh LQG
dG dG dG N bh ,fe cCG d U VdG CG jdG dG j
J H ,fhe bCGh bCG IOY J IdG QdG a . H dG dGh
.fheh IhGW cCG IdG QdG VH
f bdG ,H a J eY J fCG EG ,Uh Sb LLdG VH J
VH Ibh HH IdG QdG VH fc d .dG e dG IdG g
VJ eY dG e fheh Jb CG EG ,cCG dH f f LdG
.ed
71

Je

cCG

Ae

IdG
cCG K
HdG

eG IdG

Je

IdG ZCG

cdG je
gGh

G AdG

cDG dG
chG e G jQdG O

HdG cCG K GQe dG j .G I RdG cCG dG Ib dG N dG eG eDJ


.AGh cCGh
c djc W
jYdG GdG W
SEG dG
IdG e SG

IdG
jQG
jQG
LQG IdG AZ

LQG IdG AZ

NGdG IdG AZ

NGdG IdG AZ

.WQ W H qdG H Ib YCG dG j


dG ''jYdG GdG W'' YJ H Ie IdG
.NG e dGh fG e eG

j .Ijeh hX X QdG VH J
g .G FW H Id e YCG dG
e VJ M ,dG LQG djdG IdG
.edGh ThG Y kGH eBG

.jdG GdC H QdG e dG VH J


gh ULE G T q dG H J
VJ eY .HG Id G dG
GE h ,dH J ed dG g
.FGhdG e kGOY qe QhJ

dG jdG j EG gGJ H ,dG jM Y d fG dG IFa J


jdG K a EG SdGh dG IdG e S dG UdG j .a J
dG jd aH IG EG UdG j M ,ebh SCGQ j CG b d dG
25
.dG T Y J dGh b T Y J
G Id LQG dG

Ib
G

jQG

dG IYCG
IdG

LQG IdG AZ
NGdG IdG AZ

73

H j
dG Ib

ddG dG

fGC h JG
dG j
TG Y idG j f
.GdG JQe g IQKEG JGQe cCGh ,ge TG
e S N e acG idG jW Y ag j Y TG IQe EG
QdG g H Y AdG d .AdG H b QdG
d G dG Y G c S
26
.bG
aZ TG J QdG g hCG
jGhR iMEG HHP Vhh ,ke e
d Gec Vh .d jc adG
HHdG CGH EG eh ,VdG bG
G jH TG M GdH
AdG SG .Y fGh gf
j TG CG HdG g e
dG M Gk L IOM GQOEG SH
.dG
74

j hQg

ij G Gg CG CG ?dG SH bY d g GQOEG SM g :GdAJ dh


adG jGhR e jhGR Yhjo GVa ,iNCG HH Geb dP Y HLEd ?dG
Yho dG dGh dG aQ kbh r p jo TG WCG eYh ,H gZh f
.jDhdG SH d bY cGQOEG TG IQa :PEG
TN nVp ho M ,jW gO IjL HH Gea ,TG Y HQ AdG HJ
GQG e Iee LGM Vh .iNCG jhGdG TGdG e Yh jGhdG iMEG
J dG g dh e TG Qh j c K Y e c j ,fG
CG TG Y Lj fCG CG ,NBG Y j LM c IOLe fc H ,be
.gRd kLe kjW j
TG Uh eY .dG QdG gdG Gg TG GhQM M bGG AdG CGH
.HU TCG dG bH e eCG j h ,dH dG RLG e hCG LG EG
eHh ,kjCG dG e j e H ,a LG j e TG j Ggh
.dG H Je Ce b TG c NCG dG RLG
GQOEG SM ie Gd INCG H e G GhQa ,hgT AdG U
VH G Gg Y G GQOEG OhM ae HdG g e dG c .TG Y
J e 0^6 e c b j jPa SCH e jW f iNCG Ie .cCG
.dG e FGY H
e ZdG Y FGdG g e kjCG j CG hO MQ TG HJ ,gdG e jh
GQOEG Y Ie IQb TG id CG HdG g J dG HCG XCG bh .dBV
N e a Lj dG dG WQN j e TG j ,jd idG j H J
Gg 27.J CG jdG PCd dGh ATCG Y QJ dG JdG SfG
f j .dEG e fG dG GUCG H c HHdH TG SMEG CG j
idG e UCG GUCG Qjh IQOdG GUCG f c TG Y idG
Y kbO kJ OQGdG idG SGh QOdG dG dJ H UdG bdG j .FdG
TG a Lh dG HdG :G S Y .dGh TG H UdG aG
dY HHP QGUEH adG Th Yho dG OLh TG QOCG ,VQCG Y Yho dG

75

dG dG

TG Qb Ggh ,GUCG VQCG Y ;Y G GQTEG Jh dG e


TG EG bCG fc YhodG Ea ,dP e dG Yh .VQCG Hh H UdG aG
H .G GOOdG e j QhH Ggh ,N cM c fCG c ,S 0^5 GH
JU IFGO 20000 GM TG Qj .VQCG Y Yho dG OLH TG Y jdG
gh aj TG Ea ,dP Y IhY ,G GUCG c j CG h ,fdG
.FdG Gd G dG J FG g c Y fCe If .G H j
e gY QGOGdG f CG g TG QW H J dG gG iNCG UG
,kGL V bG d dG GOOdG W EG M ,JU iS J H
Qe e c c GPEG fCG j HhdG KCJh .HhdG KCJ H Ic e d j
76

j hQg

dG M GQG MG dG ae dS H j TG GC QdG KGC


.dN e GdG j fGB h,eGdG

GPEG dh ,QG gQj dG f GOOdG j G Ea ,kf kHK Gh dG


gQj dG J Y J G J dG GOOdG Ea kce gMCG c
dG OOJ aHhO IgX M fC Td dH e e g J Gd.QG
OOJ Ca Y Ie HHP Y JU Le j eY fCa Gd S e L e G
.TG j Gk OOJ j b G dG
LG jH j fCG g H d G g e TG j Gg c eh
:G S Y .HhdG KCJ S c j fCch cG IOG EG Sj dG JdG
77

dG dG

AG G LG J dHh ,IG HHdG EG JdG LG YCG TG Sj


. JdG HHdGh GOOdG W e TG j dG
?JdG GOOdG jH TG j c dh
hCG :QfdG fCH J dG dG jG e Yf TG eO Lj
.idG ce GAGf dd dG EG eGhCG QJ fdGh ,G idG J
Gk QGfEG j G GQTEG kH kaGfG EG M e eGH jG g J
Y idG LQO J Ggh .idG OOJ e Zj dG AGdG Gk OOJ djh iNCd
.AGOCG aCG Y d G e aGdH TG
EG .aOG jW Y dGh QdG jf GC e Y gGdG g J G e
OLda ,FGdG ''IdG'' GC e Y h Gk L e TG id QfdG f
TG e J .FGOC dH kjM j ee H dG Gg f eGG

c ,kGe 3050 Y M TG Gg j .jeGC Jh e 50 GE fS OY j dG Td Ie cGC


28
GQG QGOGQ j GC H kGL c fGC

78

j hQg

IQhGh G GQTEG e EG dP iJ H ,GUCG e dY LQO WEG Y


.MGh bh dP c H UG QfdG GUCG jJh
AT Y EG j Gg EG H ,dG aOG GC e AV Y dP T ,dH
.FdG Gg N dEG RYEG :MGh
dj e cG c Yh ,TG N IjL Ge Y AdG HCG c
EG IjG HCG UJ a .dG dG Gg Y dBG ah c cCG NGO
e Y bGe GhOGQCG jdG AdG e OY b 29.TG Y Gk L Ie acG
QdG g CG Md .YG AYCG H Y SQEG ILCG cH ,dG MCG aG
g Y e H MdG MG .dG b OJh nJor b d dG e dG J
70-50 H e hGJ ae EG MQ j YG g AYCG H CG GhLh .MdG
.kTgOEG MGG cCG fc H dG hH b TG gDhj dG IOdG MQ CG EG.c
gb dG aG fc e e GLN dG dG EG jFY kL TG OGaCG j
dG GDdGh .dEG GUh dG Gh
G TG j c :f j
c Y H Oj ch a j dG
aG Ie
?e N dG
j c f f BG M
ie AdG j .IOdG Md TG
dG
MQ Y TG id dG dG KCJ
ij TG CG fcJ GPEGh .IOdG
e jG bj AdG Ea ,ke AdG
dG Gg d j dG BLG
WG e OY j TG GC o cG dhj AdG GR e ,QNH .G
e ke kjW j Oj eY fGC GE ,G
jdG dGReh .dG GE a c dG G dG Gg Y fj dG cG
!dG
.VGh Z IOdG MQ fGW j dG
79

dG dG

FHdG dG
G S Y YdG bdG
J .hReCG f kjJ je dG e j dG FHdG G j
.FHc U 6000 EG 5000 H e gh dG g L K FHdG AYCG
g J .eCG 2 d da 500 QG FHc T dJ CG FdG g
.FHdG dH ja J a ,dGh adG Vd FHdG UG dG
g dJ CG .je ae e IH d dG g J dG FHdG dG J
.fdG e 1000 /3-2 EG J YH T FHdG dBG
dG jZ dBG .JGP M Ie J dG g H J dG IdG IdG g
CG FHdG dG C ,''IN IG'' QdG CGe N e WH MT h
H LhCG b dG Gg CG j ,Jee eH j GPEG IFa jCG G Gd j
.MGh bh ee

FHdG G SGH ''iJ'' dG dG


FHc T dJ iNCG S g ,FHdG dH G dG Y Gk H
ad FHdG dG J dG g dh .da 3-2 H e hGJ H
? PEG eJ GPa ,dGh
LCG kM kef J dG N .M c dG GQTEG g dG J
30
.GQTEG g jh Qj CG dG
e FHdG HHdG QJh ,jP Lj UN Y FHdG GQGUEG dJ
gh ,dM kFHc kM QdG Y dJ GQTEG T Y gX IOLG eG BG
CG AHd bf g g ,G Gg V j AT CG Yfh M j e dG J
J CG FHc e dG L Y Lj .cM O CG c ?
.ke QGOGdG j c G
80

j hQg

bf Z IOe

FHdG IQdG

FHdG dG
bdG IOG

dG GG jh FHdG GQTEG dEG j Gk QGOGQ dG g ,QNH


UCG eYh .H L dG GQTEG g J dG OGG H G NGO
.dG d G G cG fEG id kahe QGOGdG

e e
J HGG a .G e GfCG SCG g LCG Lj
g .GG bj hCG SCG Y QJ dG G OOdG GP FHdG HHdG
CG c ,VQCd WG G J fCG LQd SG IjT G
. SGh jdG Y eG
,dG gQJ dG dG OOdG GP GQTEG HGG G J CG
Jh JGP dG a J SG g .HfCG H We G a
H QJh H a SCG UGJ CG jdG H .a LGJ dG G SQ Y
.Gh Gh dGh GfCG M eG OJ fCG c ,G QNCG e
81

dG dG

G H aNG J dG GQTE G
OLh Y IhY ,Ija FHc GQTEG SCG g GfCG e f c j
H UdG H J .HK J edG dG CG EG ,MGdG dG OGaCG H aNG
.eG d kah Jh G aJ cdG fH fCG ea ,OGaCG

dG GQTGE
dG jM SCa .dG Y Y ee FHdG GQTEG
JHHP dG SCG Jh ,dG SCG dG J Y J GQTEG
dG d K JHHP J gY ,Id LhN e Y HGdG dG M UG
jW a ,IG IHCGh eeCG bY Gk c Gk QhO J UG g .dG SCG
.kS ihCG EG H Ojh U Y dGdG j
82

j hQg

GQTE G Y J dG fC G
GfCG c .Gh dG ee Y J ee kjCG SCG J CG
S J :G S Y .jj SQ FHdG GQTEG J FHdG SCG
dGh ,fdG HHP 10 CG Jg 10 OOH FHc GQTEG Mormydae GeQG
G jH cdG Mormydae dG g J .Jg 120-100 H e EG d JOjR
MCG e .dG AH b dG MEG J dBG g .Y S dG dG e
WEG Y jdG bJ cG AfG H .jdG Y G AbEH jdG Gg j
dGh , cS cG e J dG dG J .bO 30 I FHdG GQTEG
J fCG PEG ,kfK H G ATCG h ,khCG HU cCG Y QdG L g dP AGQh
.FHdG dG H ke AY

d UN f
FHdG GVG
e S HbG d j GPe BGh
FHdG GQdG J gh iNCG
e QGOGdG GQT NGJ CG ?f
gM dG dh ,W f g ?H
j .GVG Gg W YaO dBH
fG HSG'' SG dG Gg Y AGG
S LGJ eY JAR . Gk QNG ''MORd
,YH gOOJ J f OOdG iNCG
FHdG NGdG hM j Ggh
. e bH
FHdG dG J Y j Gg c
IgdG g J h ,dG Y

iNGC S e FHdG dG O
.FHdG GQTE G jW Y

83

dG dG

g J .He GQTGE dG f OGaGC j .LG g N e FHc Le SJ dG SC G J


dG feGE j ,GVG d dG f e iNC H J eY FHdG GOOdG T e SC G
.IH dh He GQTGE Y

dG g fCG T dSH ''GfCG UCG'' Hc jhQGO YG .''QdG'' AV Y


jhQGO eh .31''jdG LGJ dG HdG'' GY j a jdG g Y AH
.kGJ cCG H Y Ad J dG gh dG Mh
H dG OLh Y dJ bG e gc gh ,Lh Y dG N e SCG g
...Shb S ,JQbh

84

j hQg
YGfGC M FHdG SG j dG GQT G GfGC

Gymnarchus nilotikus

Gnathonemus pertersii

Gnathonemus moori

Mormyrus rume

Gnathonemus moori

Mormyrops deliciosus

NGB f J GC FHdG d
.FHdG GQdG N e SC G e

ddG L NGO QfdG


H Yh eGO X AG OdG e b H ddG CG
CG hO dP fEG , ?aG g e ddG S iJ a ,(kGec 3) e
dG idG j f SGH bdG jdG GH J CG .GJ
Hh YSh , Mh ,T Y e ee H d j dGh ,L
.AdG
j a ,G dG Gg j dG GQG jJ kbh ddG j
.HdG e GS EG dG ddG j ,ATCG e e ddG ddG
,a a j dG G cG idG j f ddG j
kJGUCG QJh dG bh MCG H adG e HQCG hCG KK H
ddG adG QJ.dG S h ja HJ Gk L jbh Je LQH
e e JdG LG g ddG Qj hCG Jg 000.02 fEG j kJGUCG
a Zj dG G G LG g LJ ddG jh fG .SCGQ ee
L CH g QfdG Le eUCG Ca.G J f SCGQ jH dPh
gO IO IA ZGa NGdG P Gh S G dG H Lj. LJh J
eG J K .NGdG P G G LG g f Xhh (d) SG e
gH j dG edG EG dG
f He gO IOe LJ .h
GUCG J PEG , G id QfdG
gO OGe Y JdG a LG
g J CG j . Hh
dH Ve G gdG OGG
f e J M SG dGh

86

fE d Ye Z kJGUGC ddG adG QJ


IOLe e e e GUC G g J .a a Jg
''G fG'' YJ SGC dG ee
ZQ M LG g LJ GC ad
QfdG Le J .SGC Q jH
dG j .FY C H J eY ITe
dG PC G GE GQTE G j xnnonc dG
.gJh H d edG GE gQhH eG LJ

(20.000 Hz.)

dG
fCG

dG G fG
fCG
JU a Le

idG
NGdG PCG

,OdG G gO Y jh JGP H ja gdG e f c .JdG LG


d .G fCG e OY EG Ie Fc Y N e jh
g CG gh H d QdG jf YJ c ,jQdH Qj CG ddG Y Gg QfdG
FdG Vhh T J e ddG Y G gQhO ODJ CG IG gdG
ceh NGdG PCG ,dG e IYG fCG J CG j ,eCG dc .Gg
,''F Ie H '' g idG j f EG .FdG H edG GUCG
.p G dG N e fEG .ke Gk eCG QGWCG J CGe N e gJ hj dHh

87

dG dG

JGC b
dG KOG g CG J a,d jU Hh Y a J dG dG cj c
?Ieh jW jH IA Ib GC
e ZdG Yh ,MGh e GYe jU L eCG iMEG fCG Vd
jU H gMCG dG .H NBG OLH j e kjCG CG EG G bGdG bY
L e G AdG Nj :FdG dG e S GC J gh ,H dG
Gg K ,fdG (FdG jG) Ja jJ IY YH dG SY EG jU
e e f dG .dG EG dG Ic C dG FdGh SdG Y AdG
jQG eCGh ,dG Y AdG a dG eCG ,''dG'' h ''jQG'' Y j N
.GdCG a
.dG Y G FdG LG J ,kLQN IOLG ATCG Y OYH
b H J .jU iCGQ dG g fMCG CG Vfh jdG e EG Od

MdGh fdG
W gh ,d SSC G HQC G fG iMGE g fdG
gY QhH Ad fdG J .dG ee fe aT
j GC aOe d .dH IadG LR j c ke
G G c e d Lj dG dG g Ve
a MdG eGC .dG e eeC G dG GC ,ke SG
c MdG J . ke fdG N IOLe gh fd d
-dG D HD H SJh J jW Y dG GE IOQGdG AdG
,QdG Y G AdG D HD dG j .SdG jFGdG dG
GE NdH AdG e c cC H d Sj G AdG h
GE ,OQGdG AdG jd GedG GC G Gg OYG .dG
.G dG Ge Y IH J fGC

88

j hQg

H dG N e fE G Y J
Zh ,G fG e HQGC
dG j fG g e IMGh
Y :G S Y .dG Y WY
dG edG OdG iMGE Z
G dG Gg .dG Y bJh
''jQdG QdG'' jW Y J
b dG GC j ,jQdG Yj c
e G H OLd X
.knN oN fGE :GC ,hC G dG
.(dG fG ,VJ SQ)

dG
G

G
DHDH IOQhCGh jGT
dG
dG

HdG dG

dG IOQhCG

MdG
D HDdG
SdG
LQG FdG
fdG
e HQCG

G
dG
dG

IOQhCG
dG

89

dG dG

e eb cCG dG J :G S Y .dG Y AdG e ac Lh


ibCG FV ac dG IQhG jG J H ,e aH AdG ,LG
.dG Y adG FV Le LdG b J Ggh .jdG L e
d G GDdG Gg Y GG ?LG g J dG GG ge dh
.d p G dG eCdG Y HLEG EG Ud .kS

SOhQ

AdG
SOhQ

Yj kGL U AL H AdG j H J g jhD dG hC G IG


gh H j c JhH kGJ j dH gh ,dG cis-11
.ShOhdG

90

j hQg

jhD dG Fc
.ehdG GC e Y FdG YdG e S GC J ,dG EG fJdG J eY
.fJd SM AoL gh cis-retinal-11 Y j AoL g ehdG Gg e hCG dG
ShjdG JhH T J EG ODj ,T j AoG Gg EG JdG j eY
d j kT dP H Rhodopsin ShjQ JhH j .MEH H Jj dG rhodopsin
transducin ShfGJ Y j G NBG JhH e dH
NBG AL e Jj ShOhdG rhodopsin e transducin ''ShfGJ'' Yj CG bh
j NBG ALo e Jjh GDP ALo Qj rhodopsin e transducin j eY ,GDP Yj
FLo h transducin rhodopsin),) : JhH e G IdG g J dG Gd GTP. Y
Transducinrhodopsin GTP-. : `H (GTP ) UCG Fc
Yj G NBG JhH e YH GTP-'' ''Transducinrhodopsin NCG Gg Jj ,BGh
Yj G NBG JhH J e phosphodiesterase JhH q Ggh phosphodiesterase.
e jG H fe NCJ dG g C ,Le H G cGMP cJ bj cGMP .
.JhdG
KDj .GG YdG Gg INCG dG J ?jDhdG Y Gg c gj c
JhH e dDe H g g jCG Ib .G fjCG Gb Y cGMP cJ VfG
jOdG fjC IdG g J jOdG G .G jOdG fjCG J
eYh .RGdG Y d IFGdG fjCG H NBG ALo j H ,G EG adH
NG EG Gg ODj ,jOdG fjCG OY kjCG j cGMP jL OY j
Y f e d ,jG g e JG dG jG j ,AdG Y dG RGJ
.jDhdG gh edG EG dG YCG .''FHdG dG''
f G Y dJ YdG e S Yh IMGh N MGdG JdG j QNH
g c CG g g gG G .AdG IT CG ,JdG bW M IQSG J .FHc
IOYEH G NGO iNCG e JhH Jh .fdG e 1000/1 e bCH J dG
dG J .UCG dM EG cis-retinal , transducin, rhodopsin-11 e dG UdG
NGO J dG FdG YdG N eh ,UGG fJdG e T QGSH
32
.fJdG g e Ja H SMEG e dG J SG jG
91

dG dG

Rhk
PDE

IdG

Rh

SdG

LQG AdG

JdG
Axoneme

IGd LQG GG

: GE RedG J ,jhD dG d jG FdG YGC SdG Vj


SS fGZ = GC jQCG = A Rc ShOhQ =Rhk ShOhQ = Rh
jj OSa = PDE ShfGJ = T

LG Gg j dP e ,dG SCG g cP Gk J TCG G jDhdG dBG


Fc YJ ,JGh bdG jZ kJ fEG .d G G Y dd
Oj) Hc L gcP dG ehdG H ` JQbh dG N J ` TCG UGe SH
QJ CG PEG ,(jd a dG Gg
S e ceCG H .hCG d a IMGh H dG BG e S
EG b dG aGdG e YdG e IjL S GC d ILCG cJ j ehdG
.YdG e IjL S iNCG Ie GC d SG G Jh NBG e
92

j hQg

jW Y fe NCG b dG g CG j CG bY fEG C dH
G e .VQCG Ig Y f hCG a tdG e JCG hCG jdG L b J Cc aOG
.ge bOh jT QH feG Vh dG g e b c CG d dH
e kHV fc Y hJ aOe c CG dG FdG YdG S fcJ dc
IO FXh ODd e f ah JGh G FGLCH G dG Gg eCG d d .AGdG
.,G J fRGe SS N e
hU'' Hc N e QdG jfh dG Fc Y H Fe AMCG Y j
:''OSCG jhQGO
fCG.dG jJ J h,jDhdG Y J Y SdG dG hQGO j {
UG Gg QdG jf eb dG edG CG. QdG H Gg J
j fCG LQO G dG dG IjdG IG dG J IUbh V
33
.zHc M

jhD dG H e
Y EG jdG L Y J dG hCG JG fJa Y BG M K
J .Ie Fc Y N e FHc GQTEG dJ dG jN CGh ,LdG
IdG IG edG M ,T d ,L ,jdG Lh e GQTEG g UJ
.edG EG GQTEG g J CG j BGh .Lh IOLG
.NBG f e Fc YH dG jL b e IQG dG jG NJ
cM j ,dG Y IOLG JhdG jL T j dG G QJ eY
kaNG kFHc fG GQd IG cG dJ .HjEG dG GP jOdG GQP
CG H dG G SCGQ EG IQTEG J ,FHc IQTEG QhH dj Ggh ,G da
FHdG IQTEG Yh Y N H IK g J dh ,S 1 e bCG ae J
g dG G H LGJ e Fc .e gh ,IdG g J CG
e fH dSdG Jh ,G 1/5 EG 1/4 GM dSdG J jdG H ,IQTEG
93

dG dG

.edG EG jdG J CG EG iNCG EG Y N


IY e edG jDhdG ce dCj .edG jDhdG ce EG GQTEG g oJ
e IMGh c ,Me He e 13^5 h S 2^5 H ,iNCG a e c VJ We
,khCG eOdG GQTEG HGdG G J .Y N e 17 GM WG g
N C QGWCG e QW CG .iNCG WG EG eG H J hCG Hh
.iNCG N CG e IQTEG J CG Y
fQe J CG j BGh .edG jDhdG ce LdG IQU J jdG H
d M .kbO c e J CG ZEG hO kjCG dH j Ggh ,IcGdG jH IQdG
Lh hj GP'' :khD J j edG Ea ,IOY Y c kHT cCG jdG Lh GH
''?dG Gk L kMT jU

dG
.fK e bCH ''dG ''h ''jDhdG'' Y WCG dG JG jdG H
fQeh e J M ,edG EG jG e jG e J dG G J
.jU LdG d IcGdG jH
jN N e jDhdG J dG dG IOQ fEG j .dG Y dG J
dG eGhCG Y AH ''kMe'' :S fCG Qj :G S Y .fK e bCG IcGdG
Gg .''eHG'' f e SGQ a LdG H J dG ZedG jG gQJ
.IG dG e SS jh dG jG Y He H j eCG
U Qd dG dGh dGh JdG G EG eGhCG QJ f bdG h
H cH JdG jG CGJ dG QhU Hh .dG cM e ''kMe''
S YH J dG g JdG LG GdG j .dG d SQCG dG dG
gh ,SdG PCG EG YH fPCG NGO HG AGdG j K ,fdG e 5/1 QeCG
dG EG GRGgG g J .RGgH e 7^6 b j dG dG AZ GC j
PCG EG gQhH J fe GRGgG EG J M ,SdG PCG dG
.PCG bb NGO Lj UN FS NGO Le gKEG Y da NGdG
94

j hQg

NGO GUCG IY j
G e OY Lj M . bdG
dG J J dG G
U e QJhCG g J .IQdG
e ''kMe'' U GC a jdG
QJhCG J . Jj K fdG LQdG
h .bdG QJhCG K khCG dG
dG J AYCG J jdG
,BG GY gOYh ,IdG
.dG YCG EG GRGgG

cdG Y jFGO T M
GdG bdG Gb
bG
gdG dG
Oval
ZdG dG
window

GdG

dG IdG
LQG
PCG W

TShCG f

IOe RJ jT jN dG Id NGdG dG J .dG IdG gcJh GUC G d GdG U


AZ N .dG AZ j SdG PC G jGH H fPC G IdG jf h .LQG ShC G e PC G T
h ,SdG PC G PC G V fRG TShGC f j .cdGh GdGh bG :gh Y K LJ dG
.UN FH h SG IjT S dB H J dG bdG J jf
95

edG EG PCG e dG MQ
Y J .''aOG'' SSCG Y J dG GdGh QdG jf kje j dG G dG g PCG
.(2) dG AH dG j K GUCG (1)GdG j jGdG ,dG jZ f eJ Y dG
gQhH J fe GRGgG EG JdG LG LJ Ggh .J (3) SdG PCG Y j Gg
Y f EG fG GRGgG J g .(5) bdG OLG FdG a (4) ''gEG IadG''
.(6) jgdG YCG Y edG EG MQ GC J
.FdH G (7) bCG H Y (SdG q G dG) bdG .kGL Ie bdG NGO fG
Gg j .dG SM Y gh (dG bCG qG dG) (9) ''JQc Y'' Y (8) bdG IdG
(11) Yb AZ N e jG g EG bdG FS GRGgG J .(10) ''jdG jG'' edG

Q n Hr nC Ghn n r s dG o ond nC n fGnC p sdG n go hn }


nC Ghn n
.78 :eDG| nh o or nJ es k p bn nIn p ar
jdG jGh ,(CG12) NGdG jdG jG ,jdG jG e Yf g .JQc Y Y Vj
j dG Ggh ,J dG JdG HHdGY kGOYG H jG g J .( 12) LQG
.f dG G GUCG H
.(14) gdG dG EG Jh FHc f EG JdG GRGgG (13)LQG jdG jG
JCj c edG dG Gg H .(15) Ic gEG IY fPCG c e eG dP H J
.34(18) dG IdG jdG h (17) SG cdG G (16)HQ G GQG:
bdG SJ .G GUCG MCG Gh ,JG GUCG jW edG NGO QRCG G Qj
.ZedG IdG f EG GUCG SCGQ JOLG
.UJ UCG M bH J H e dG Y fYj dG dG dCj ,VGh g ch
c dG S e U AL CG Ga C ''IN IG'' jH OLd X b j CG dG Gd
.d G JEG Y VGh NBG ke PCG hJ Ggh .dG Y kGLY

dG dG

IdG
jgdG

IadG
bG ZdG LG gG

bdG
dG

bdG i
dG IdG

JdG LG
LQG dG IdG

LQG PCG

dG Le

PCG W

PCG
SdG

NGdG PG

dG - dG g OD J .NGdG PC Gh dG AZ H j c SdG PC G dG dG dG J
dG ZdG LG J .NGdG PC G GE gQhH J dG JdG LG J e - U NB H e c Jj
FdG Gg b e QJ dG SG GC Jh bdG FS NGO gE G IadG AH cdG Y JG Y f
.GUC G e MQ

YCG N e edG EG YH J ,FHc IQTEG ''kMe'' U UCG ,BGh


f .edG dG cG EG J CG EG YCG NGO MdG g UGJ .dG
H .G FGh dG eG H edG YCG fdJ J dd
e dG Y J .dG OH BG G j .gh jU jdG j jdG
IdG LJ :fdG e U AL N eGe H J dG dG e e N
Y j dGh edG dG ce Sj K ,c EG dG Y Oc edG U dG
.bdG GP dG EG GQTEG ''chH e''
dG .d G IOG g NdG AGdGh ,''NdG AGdG'' FdG dJ jGdG
N e AGdG j .FdG EG dUhh cCH dG AGdG SG g g SSCG
FdG OLG dG b e cCG UeG j .FdG EG dG Y jh fCG
.FdG IQO Gk gL AGdG j gY ,HdG cCG K QhH j dG

98

j hQg

fCG jdG

dG
IdG
G

bQdG jdG

FGdG dG

c dP Yj GC j H ,dG e fMGC j d fH FGdG iGh FdGh JdG G J GC j


.JdG G Y AGdG Qhe e dG AKGC J dG GUC G J .H JG dG dG

,bdG FdG J G g .IG JdG G Y FdG e QG AGdG


VH dG b JdG G J .H G jQdG SGH dGSEG dG
Oj Gg .j dG AGdG H RGgH GC Jh dG AKCG H e Jh ,dG
.YCG G U c c IOhe G fc c ,dG U LQO
99

dG dG

.fCGh dG N e dG EG jh G Y CG H dG e AGdG j
kjb dG j .G d Ija iNCG T Ge dGh fCG H J
Y IY J dG g N e .kGO kT dG NCJh G Y Gk Hh
35
.ge YHh
Qj CG fQHh ,JcGP iNCG GUCH j dG dG jdG Qj
NBG Y dG e c j Ggh . CG jd kadCe dG Gg c GPEG e Gk Qa
,NBG Y e c jh jU Hj eY HdG GG c.jh
f f .NGO Qhj GPe Qf CG b M ge bOh J YH ch
fCG Ea dP e ,WdG jZ QeCG Cch Ic dH fh ifh
.dG jZ g da SMCG g c dG dGh
QdG jd dGh IjdG edG e J eCG Y j G dG Gg
Y .QdG aOG GC e jW Y dGh dGh jDhdG UCG T .gJ CG
JJ fe jQZ e JdG G dC J
J eY . LQG dG Y H
Me G J MGSG QW dG
,(SGC ) dG AKGC G J .(QdG GE )
fc c cGC IOhe G fc ch
.YGC dG If

bQdG jdG
JdG G

eZdG

100

j hQg

dG Jh fS dG MCG dG b e d gh c fCG a dP e dG
.G e IQP c e SCG G JQbh dEG M J
:fCG g Y J dGTh GH dG Y BGdG j
nIn p ar nC G hn Qn n rHnC G hn nr s dG oond nn nLhn kn T n o nr nJ n ropJn seGoC p o oH ue onLn Nr GnC odG hn }
.78 :dG | nh o or nJ osn nd

:iNCG jBG h
nC G hn Qn nHr nC G hn n r s dG o ond nC n fGnC p sdG n go hn }
.78 :eDG | nh o or nJ es k p bn nIn p ar

Gg .fdG pr pY N J IQdG IOQGdG MG c .Fa YS GP GeH JdG G Y jJ


.f j JdG d G dG

101

dG dG

HGdG dG

MdG fGC
jOGJQG
dGh .VQCG Y GdGh MdGh G bG SCG jQdG J
j dG dG e Sb IOe e j dG dG dG OLh g dP AGQh FdG
H IG cG Jh ,FJQGh dG d kde kYO dG J .T
.cG UG
Y dG H H ,jQdG YS e bCG YH a jQadG eCG
.Y
dG g J .''S'' YJ fCG e keY H dG jQadG e QG
J S AZ L j . gG dG Gg H IQhG Y jOY Z IQ
Gg aO SGHh gL AG N J G Gg IOLG dG g SGHh
GG Gg SCGQ fL Y Lj .QG Y Gk L Ie dBG .MdG J AGQdG G AG
EG dG g e G NJ .G J a
K .G NGO fGSCG T Y j
,c V SQ Gk Qa G g oJ
G jdG cH d j
e Gk L Fe MdG dBG g .IdG YQG IdG H cG
Todarodes Y j dGh HdG QG j .edGh YdG :Mf
102

e dG AKGC kGc kYO QG S J


g J .L H j dG G
L NGO e dG J dG SG
ja dG OJ eY .QPC G
dG J .Jh SG g oJ
.(fK Y) YQPC G dG Y
WdG GC QG j iNGC L e
,IQ ej dG Qe SGH ATGC GE
GSGh WdG QO K e a
36
fH NGdG OLG dG

dG dG

SG j dG QG
''jQda dd'' dG
g Loligo Vulgaris
j .Yf OGaGC H UC G
OGJQG MdG f d
30 GE J YH dG
37
. S/c

IdG aG h .jdG H S/c2 YH c 2000 dW j MH Pacificus


.37S/c 30 EG YS J CG GfCG H .S /c 11 M YS jj CG
dG dG Y J dGh Gk L jdG JcH AGYCG QG j CG
H c AGOS HS QG j ,eCG d jdG cG J eYh .jdG
hd c eR gh GK d hdG G Gg Lj .L NGO cH j G
.G G HS N IOLG dG QOJ H ,QG
M ,dG UbG QG dG OQ Y IG MdGh adG fCG J dc
dG jdG cG dG ef j .Ic YH Meh ja Le e j
ef J G c. MdG g eRdG dG GAJQGh dG H
jS jG Y CG SG Y Qd eDj dG dG Ga,e J dG
.MdG Y AG d jQhVh gh
Gg j kjCG NCG ,YQG jdH j dG MdG GG d QGh
JbhCG e J a ,MdG Ige d bG a dP e .MdG e dG
.dG b dG GGh QdG H GdG

104

j hQg

e NCd NGdG G j
a VJ dG e W IY
fdG dG
GfCG KK J dG g .H
jFGdGh fdG :dG e
b dG g J .YdGh
jW Y cG e jdH NCG
dG
.d YOh fRGe
YdG dG
jFGdG
J kLQN AG j eY
H L NC G j
h fCG h ,kfW jFGdG dG
gD j fdG dG iMGE
VY e jJ a M Y aG
.AG Md
f bdG .j G j
hM AKCG AJQG dM YdG dG J .dSG JG fdG dG
J dG Y J CG Hh .LQG AdG S IOjR EG ODj ,HdG dG
e AH AeGh AdG bJ j ,NCG hCG QG W e Jh YdG dG
.jL

jFGO Y
YdG dG
dG H Y
dG jS Y
d W

LJ fdG dG Y kVY dh ,jFGOh YT Y QG L Lj ,NC G Y G g c h


dG OhJ fGC GE aVGE iNC G dG J eY ,OdG e L dG g .dC G e W Y
.jb IYH YdG

105

dG dG

AG n r en

YR
adG fCG
LQG dG hCG LQG AdG QGL
NGdG AYCG
dG

G VdG

NG VdG
G VdG

AG
r n r en

AG f

fCG a

fCG

IQhdG GC J.QG e Weh adG IQhdG IQdG J


G e %10 d LQG dG Oj .(1)SdH
AG Nj .%22 FdG jdG M e jj ,dG
T Y fGC N h SGC dG fL Y IOLG dG e
IC a dG dG OGOj .(2) OdG G e %75 GE G b j ,YC G M GE OdG j eY .b
M e %60 Oj e) kjJ AG c j .aG AG i a fC G a Y OLG NGdG dH aj
PD J b IFGR J GC .(3) NGdG AG jW Y dG T G j.IH QG GE fC G e (dG
.UeG Me a MGdG QhdG Ge 10-6 H e QJ GC h fK f dG Y J .dH G
.UC G M e %90 M j AH QG j eY

106

j :NG H e ,NCG Y dG dP QG id dG dG j
H .NCG Y IOLG fdG dG e kH gEG J QJhCG e W QG
QNh NGO J W e QJhCG g dCJ .jFGdG dG LGJ dG g
VJ H ,jFGdG dG J dG Jg H .fdG dG e ke G
. OY H Jh g H YdG dG
IQD H dG J . kGL Ie QG Y H GE
.dG e SdG J eY D HD dG
ZE H d OQGdG AdG M jJ kjGC
fL GE IOLG IdG ZC G ahGC
.dG
AYC G g OLh J dH
fE G :bG e Yf IG
YG d .QdG jf AV Y QGh
38
.Hc Gg dSH jhQGO

107

p VQr nC Ghn pGhn n s dG


o reo pd hn }
nYn o dGhn Ao B njn en o or jn n o n r Hn en hn
17 :IFG | l j pbn Am r n T
u co

adG fGC h jOGJQG MdG fGC GE


bdG jhD dGh dG Y J dG
FG g c ,G d J feGE h
.pG G Y ge FO iS g e

j fG dG G e G c W LJ
hGC QG j dG g GSH .''dG'' Y
Y cdG j .LQG G d J e NC G
dG J Y J hGdG Y kfGdGC G S
.dG hGC dG ej
eYh ,QRC G dH j fC H cdG j eY
Gd LGG dG j dP AKGC NGB cP Nj
MC G dG QG j .MC G dH NdG
.ddG bGGh dG bGe jd

j Y AG dG J .QG Y OGJQG MdG f YO Y Gh QPC H dG bdG dG YJ


.bdG AKGC Hc J c dG AKGC G fRGe Y QPC G J ,iNGC L e .IQdG j dG AdG Gg

GFdG GC e j GC e Y QGh NC G e c id OGJQG MdG f j


jH U b dG dG GC f eC dG e b eh ,KdG
.kaOe Gg j GC dH .ke SJ

| no pbjo m nr ud
l jn BG mHs BGOn pe to jn en hn r op rNn hn }
.4 :KG

f e J
.QG Y KdG
kSGC dG g VH
jhdH dd gD J Ld
.QdG YGC IOLG
GC GE dG OLG AGdG LC G
dG G j .d IgL J
g Jh ,jP ITdG j e SGH
j U J c .39 H dG H UG
.dE G IQdG jGB e Ggh UN f ah

eG dG

dG Ge
adG fGC h HC G
FdG
JJ ge kTYCG Jh ,MOe Ge J IU b HCG dG
j dG edG dG CG g VG gG CG EG .AdG a Ge e hd VQCG Y
.ke YCG gG AdG GH
Ie OG e GMh g .dG jZ kef HCG dG Y f hj
Je OL Lj c .Y aG I OG gh ,Y adG HCG dG
Vd G fM e kAGHG ;dG TYCG NGO Y c J .''jQfG a''h
Jh .aGG OG AGOCG Y ,dG OH AfGh GQGh fCG AH Gk Qhe
G GC J .KdH j jdG Gh G OLh Y HCG dG Ie jQGSG
eCG ,ke fEG dBH TCG hd GS 9 EG dW j bh ,hCG dG H dH
M Jh HC G dG e J
Gdh ,S 9 GE dW j
IG OGaGC e Y J
.SGMh jJh H

114

j hQg

ah G SJ IG S OGOj eYh ,VQC G S Y TYGC AH HC G dG GC j


.QeGC 5-4 He GE j GC YJQG GE M ,OjORG Gg

UN Y LJ VdG Vh iS d Y fCG h . HdG e a cM


Qj e :CG ,dG H dCG 30 GM G J .Jh eWEGh H AYd dG e
.JM Ia H je10
dG OGaCG QYCG hGJh ,YCG fc e ZdG Y dG QeCH edG dG j
J H ,Y Gh HCG dG Y AH e Y J .GS HQCG EG K H
.Gh QdGh VdG bG iNCG Y
,hdGh dG e SGH UGjh ,e ke IG AYCG j
OH J J h J dGh AdG bG g .FdG GQTEG OJ j M
Y Gg c ,adG GQheh QNCG e jdGh ,OGh ,dGh ,AdG e G e
.FdG GQTEG OJ jW

115

dG dG

qdG
AGdG Ib i
V
V
fG IM
Gb

B
G G

eCG
ja
dG

fG IM

IYdG
IcdG

A: 860F (300C) 2.7% CO2


B: 770F (250C) 2.7% CO2
C: 750F (240C) 0.8% CO2

dG Y ee fGC LJ
jdG fGC ,dG e HC G
VoJ dP Y IhY .WdGh
AGLGC IQGM LQO
LQO J .Y G jh G
HdG cGC K fh IQGG
.40 G NGO G M

IG VJ eY .dG cBGh dG g HCG dG Ge AGYCG NCG EG


ge Je HCG jaEG dG .UN jQfG SCG J jhdG jg MCG g EG
.ATCG EG hdG L j CG IdG T d SCH j
dG H e c fCG LQG dGh HCG dG Y H IMdG JG Sh
dG Y j dG SQG .''jJ'' EG j fCG g Y QhG CG EG ,IMGdG
SCGQ ODj CG .FGQ jW Y CG fS e dG EG NGdG c GPEG e dH
g EG VdG M a,ke G ihJ dG ,fCG g e C AdG QhO dG
.a ShDhQ H LQG aG Jh dG dG dG J

HC G dG J
H dG Je Y adG HCG dG j dG iNCG dG e
j jQfG G g J CG EG ,AGYCH FG EGh eS Y G S
FGSG HdG j dG HCG dG N hJ :G S Y .NBG EG f e
M ''VQCG Y J Hb'' fCch GG g J M .UN H Ie jde
j L NGO kUN kc a ,cSh H
116

j hQg

GE M ,Y adG HC G dG N OGaGC fj .dG cGB h dG :FGYGC dGC V e ce HC G dG jj


NJ bh IQdG edG dG hj .AGYC G Y dGh OG IY cG H j YC G edG dG
.IdG ShhD H jG OG Yd cG

eYh AGYCG b e g EG VJ Y Gk LY ghY j Fc ce Y


jhdG fCG jh H Y H HCG dG j b hdG j
Lh jaEG L HCG dG j .S S UCG FH hdG J EG ODj
QGVCH J Gg cH AdG dG C ,kjQfG dG Gg jh .kHe kf jeCG
.ILh IH dP H Je EG ODj NGdG JLC dH
.OG IY dG M Nj ,WdG eM cG fc GPEG
YGG HCG dG id dG GC eh G dG e fhG dG g MJ
Ia J eCG g CG Y IhY .''UG j c'' :J dG jhQGdG
c dh ?kjL J CG ke AdG dG ZJ GPa , ge IjH J b Y
d c d ?gZ hO dH dG e Jh KCG J CG QJ GPa QG d
H J CG QNG fCG kL S d .kJ bCGh G SCG QN QJ CG
117

dG dG

(eSe Z fG IOQhCG ) G IOQhCG

ddG SCGQ
G SCGd S e

dG dG
d
G SCGQ

edG SCGQ

ddG SCGQ

e LCG EG QfG dG Gg QJ CG G e a ,IG Y adG


CG j CG h YCG edG HCG dG . dG G jW Y JOGQEH gH
.JOGQEH M LCG
,IG g fRGeh e IM j CG j e g a HCG dG dN
dEG dG H MhCG dG LGdH j HCG dG EG .IO dhDe GP kba jh
p GB n go sGE p msHGB On pe ne...} :dG BGdG J j .OLGh H
.56 :Og |.. n pnp U npH l N

dG e fGC
jdG dG XJ L NGO IOLe UN fCG HCG dG j
hG NGO eS Fc OGe H GfCG H J :G S Y .KQGG YadGh
dG J M ''ITdG'' dBG iNCG lGfCG J H .J eY gAGYCG H J dG
.ITc dG dG k ee YJh LG L Y
g J .fEG JY Y jaEG HCG dG Y dG Y adG dhDe J
` kM cCG fj IOYh ` G SGG eCG .kf IU OL gh AY fEG
M .G G NO e NO CG h, G hdGh bdH AYH ea
.dG JJh dG L dG QdG OG Yj
118

j hQg

HCG dG j
d M Je Y adG
hj .JM dP c
HCG dG OGaCG MCG IQdG
dG Y kLd kFS Rj gh
.LG

Idc IOM kcah ,ShdG J kShDhQ OGaCG PEG ;d G SG No


S e FdG Gg jh ,fRh e %10 j SQG L FS Lj .ad I
SQG j ScCG FdG Gg j (alkenes and alkanes). Me JGQgHc
d gS dG AGYCG hL NGO FdG Gg H G SG j .L ee
.ca GSH
.GDdG Gg Y HLEG GaY eY AdG U ?hdG Y FdG Gg KCJ g e
,"haemolymph" Y j dG LCG FS Lj .dG d fe iS FdG Gg j
d dG J Y dH IjL Fc IOe GC J ,dG oJ eY .dG Y j
.IG IOG g Y j HCG dG Rj dG FdG .G
RYEG j h .JGP H e eCG IdG bG g NGO J f OLh EG
H J dG AYCG EG Gj H ,d fG IOG g RGaEG Y HCG dG
119

dG dG

cCdH HCG da ,aOG jH j CG Gc SJ cCdH .ddG IQdG


Gd G IOG OGYEH dP Y AH jh ,dG Y dG J dBG j kFc kY d
.MCG MGdG dG N bG g CG dG dEG bj NBG dO Gg .dG

HC G dG SGC
dG e OG j .HCG d G G Y Ic FO fMCG j CG
IU kca Go pgho G g REh .gOLCG H cj dG dH gAGYCG HCG
. jM Ge jJh cH J CG kjCG OG d .ITdc jY kgTh
j Ggh ,fRh e %35 EG J YaO FGS J CG LPdG AdG IG
.dG e BG d
dBG Prohinotermes GjQa HCG dG
Fc dG Gg j .dH YdG
iNCG GfCG e dG .eS IOc "nitroalkane"
NG g VG gG CG EG ,dG j
:G S Y .dG g d FdG cGdG
kS Schedorhinotermes jaEG HCG dG j
dG j H , "vinylketones" FdG IOG e
dG TYGC eGC j OG MGC
f e Sh d FS TH jh
"B- ketoaldehydes". IOe c HCG
.FdG SC G
dG ''j HCG dG e NBG f g
''cdG'' hCG ''SEG'' YJ Fch ,c
o jG
.G EG ODJh jdG jG e QdG Y YJ dG g c .UN SCc
WGG J GAf Nasutitermitinae HCG dG ISCG AYCG L Y Lj
hdG H WGG g AdG dG OJ G dM .NGO UN ScCG Y
41
.FdG chRdG c ke j dG Gg .dG FdG Jh

120

j hQg

gh ,Fc SCG hO Lh b ''FGdG AdG dG'' Ea ,QdG jf M


.H G EG VGaG Gg CG EG .QdG LQO M H a Lh SCG
c J CG j H ,da GP dG dG j d FdG IOG aJh OLh j
g J CG j ,iNCG L e .eG ddG e fL Y d G AYCG
AYCd dH eCG dc ,a dG j d G AGLCG bH Y dhe AYCG
fe f EG adG fCG j .kjCG dheh JEH e J CG j YRG
,edG Y QdG jW Y J CG AYCG g d .e Y YJ e
EG ODj Ggh ,IFdG Y c dG Gg S dG g e IMGh M Ga C
No b g ''FdG SCG fCG'' CG g MdG dG Ea dd .IG VGfG
dG N d .''r nG'' g Gk Oe Gk HJ dP AGQh CG j Ggh ,IMGh tc
g LCG dG fE Gk ce dG N d .IMGh bG e c HCG
:dG JjBG Gg dG Uh .eGS dG aCG dEG MhCGh ,bG
p G hn n s dG en o nd o u njo n roG Ao n rSnC G o nd Qo u noG
n go hn p VQr nC Gh
o Qp n dG o dp nG o dG n go }
p nG o j p n dG
.24 :G IQS | o

121

SOdG dG

FdG :dG
Id gGdG
d edG FXdG
j a GQhdG FdG Gg SG Wh .IG eLCG d dG N FS dG
hDG fEG .edG OGG e G H eRdG jG eDjh AGdGh IOhdGh AdG
GQ eRdG edH j fCG EG aVEG ,G NGO JG Y kjJ MdG
.QGSH dG Oj Ggh jedG YhCG
dH QhJ dG e Gd 5 Qj e c 60 j dG dG fEG L j
f Ea VjdG jQdG hCG cdG SQ dM dh ,IMGh bO N G AfCG
QhL e AGHG G e e c dG Qhj .aVCG N EG JJ g GQhdG
H e JEH U H J MCG YhCG NGO GbCG HUCG Mh dG
IQGG e Ije c G dG Gg j c ,SdG hCG FGdG H J
.AGdGh

cC G eM
jd QhV Y cCa .d dH IG ee gCG g f dG AGdG
jG EG FdG e cCG f IQhV S g Ggh ,bdG fE RdG dG
.VdG Gg j ,HfCG e Ie T j dG ,GQhdG RLh
122

.57 :bGdG| nbo u n oJ nr nan r co n r nNn o r nf }

dG dG
(jT)

Yf K Y
dG
LQG

SdG dG
Yf bQ Y

AH d fCG
fe

(e jN) NGdG fdG


U
(jQh)

jf

G jG e IOe W

WGC
fC ,gGbGC g jGdG
GE cC H dG dG jRJ Y hD e
AYGC e dG f Y dhD a IOQhC G eGC ,G AfGC
Y dhD G jdG YhC G dE G IQdG Jh .dG GE G
.G IdG ceC G GE dG jGE

(T AYh)

N ch ,cCG AGG jedG jG NGO IOLG dG N jL


N) HZg AL e 300 f UdG T J dG AGG dG jN e
Q H a cCG a ,dG e ke kef AGG dG jN J .(dG
AGG dG jN J dG dG jN :dP Y e ,kjQhV dP c c kjCG
EG H IFY dG GMG Y j dG HdG cCG K h ,fCG EG cCG
.fCG EG NCJh cCH jL e WJQG J dP H .g cJh FdG

124

j hQg

kFS dG d kGOLe dG j d
.d e kLd

dG fRGe FS
(NO). LhdG cCG OMCG RZ cCG EG aVEG dG N jL

kjCG dG N j .QGSH Gk e S dG V Ea dG RdG Gg aj GPEG


Gg Qe CG gdG j e .LhdG cCG OMCG SGH fCG EG UGdG dG c
Gh Y h eO h d YCG GQdG e Y :CG jG e Gk OY EG d ''dG''
G dEG G jBG EG g e eLCG GQdG e Y OJ dG dG Gg .j
.JEG GH eLCG N dG dd

Y
Y
.jedG YhC G dG fC G e W J
e jj ,VGC AYdG j dG J eY
jT e ke dG GE dG Qj .dG V
M Lj .kLhe NGdG S hj M e
.(QRGC ) YGC h (MGC ) Y QJhGC jdG AYdG
125

dG dG

e H jN
Y cdG dG O j .jedG jG e dG ddG AGG dG jN J
d J dG g jG g h ,b Ic e C d J c gh ,AGM N H 30
. Gh dG
YhCG NGO J CG FdG fhe Hh , UGbCG e AGG jG hJ
e W S fEa IdG fhG H J GPEG eCG .bdG dGh jdG
) ehe 7^5 AGG G b j H ,ehe 5 f dG AYdG b j .G
.(G e dCG e 1 hj ehG
H dG Ve MdG YCG ?fhG H AGG jG J d j GPe
g .fhe dG Ve Y AGG dG jN J M ,GDdG Gg Y HLEG
b Ggh dG fCG e bdG fCG jedG GdG h UdG J dG
.dG EG ODj

JeJhC G QGdG f
J dG Gg .dG jW Y J gHh kej 120 AGG jedG G IM J
AGM N e 2^5 fEG j jOY hX .IjL j G fEH fRGe
jHhjQEG Y j eg j .dP IQhdG bG PEG jj CG bdG Ggh ,fdG
YQ hCG OM OG dG c jJ j M ,fEG ie ''erythropoietin''
Y :G S a .AGdG cCG e f GPEG fEG e OGOj dc .G
bCH OLG cCG e IOSd kJeJhCG AGLEG Gg G j ,G J
.JQG OGORG c J cCG f C ,OhM

G dG f
FS g eRdG .jedG jG IOLG iNCG OGG e Gk c Gk OY eRdG
126

j hQg

.OdG G Rh e %5 j ,f UCG
LGOdG jQdG
fRh e %90 j dG FdG Gg j Gdj GdJ
jFdG YhCG
HCG SdG
gOh jSh Oeh eCG Y ,Ae
dG
dG jdG
JhH H .G JhdG GfCG e eh LCG jQdG
h gdG HH J bf JhH g dG HCG
jdGh jdG YhCG
dG RG YhCGh
H gdG JhdG GPEG .fG EG
bG jGdGh YhCG
c FGY H J S fEa ,jdG
IOQhCG
.kMe G VGeCG eCG jdG d e
jGdGh
jdG
eG QhO eRdG fedG ODJ
AYCG H JG H J gh .UG
jGdGh YhCG
jdG
.FdG FSdG jW Y jGh
g eRdG keMORG fedG cCG
e edG Gg Jj .bf eg gh ,edCG
jJ G e N c GQhdG dG j
,HhdG ,fedG ,hddG e gdG
IQdG .dG G J dG N
j .dG e jhOCGh ,edG AGdG dG
dG Y dd AGG YhC G J YGC
OGG NCjh dG edG OGG edG Gg
Y dd QRC G dG j H ,cD G
.L dG eCG EG FGdG
.cD G Z dG
e j ,ATCG g c eCf eY
H dG Y MGdG JhdG IQe EG.ge UH N b G CG jd VGdG
IQTEG d J j CG j dG dGh G jh jhOCGh fedGh gdG
UCG e GY EG g e gG eCG g EG .dG Gd G dG Y VGh
fH J jG e fdJ ;G dG FcdG OGG e BG
MQ J IMGh N e AL e jG g J bGdG .eLCG NGO ge
fEG N dG dG Y Y dO dG G dG Gg CG VGdG e .CG
.AG e f e
127

dG dG

UN dGB

IG J GPGE
IOQhC G jedG
SGh cE G
GE a ,SdH
J .(YGC ) b eRGC
H dG fC G
JQG H MC G
e dG aj .dG
e j fC ch dG

jGdG e AGdG j CG VG e
GNG e j M ,jdG QGG Y
IOLG eG CG e .dG fCG
fCG EG ,kGL IU jdG QGL Y
.f AJ e dEG NJ CG IOe C
.NdG Gg j dG g dG V EG
EG J dG FGdG jG J dP e
.(SGC ) WN
kJ RdG e cCG H fCG
Y d UN dBG LJ Gd .fCG
e cCG a ''edCG'' dhDe g .dG EG jL e FdG Sh ,dG V RGJ
j GPEG .SEG e dG e AG Ue j jdG QGL Y IOLG eG
.AG YG aG AdUdG c a G edCG g
EG j CG NO C G Z a .cdG g Y J QeCG CG EG
jd Gk c Gk QV J NO GPEG H ZG Z OGG C ,HbQ hO ZedG fCG
W J M ,QdG H J b dG MG c e edG Ea dd .dG
J e J fCch eG g e jQhdG OGG c J .eG dG G
.G SM cCG dG EG FGdG OGd RGe aJ j ,eCG

G SeJ
IQGG dG j ,iNCG OGGh edGh AGG dG jNh q dG
t Y Gk H
IQGM LQO e aGdH G IQGM fRGeh jRJ EG .jG bd Ide iNCG OGeh
fd ,eLCG IQGG jRd f g j a .gCG e Ic LQO Y eCG G
Gk c Gk QV j ,kGOQH L bH j H ,f eY jjCG IQGM jT JQH
RGh G AfCG L Y he H IQGG jRJ j Gd .SG Y
128

AGG dG jN

eRdG

jedG dG

dG RG j .eCG Gg q j dG g GQhdG
eY da cCG H
J c ,Y H G IQGM LQO JJ
IFGdG IQGG aH d G jedG YhCG
cCG Lh j dG Gd .LQG G EG
.Y Y Wf SQ hCG cf eY Gk QGMG
G jedG YhCG a OQdG G eCG
TCG G EG aG dG c UfEG gj
Jje fOCG G IOhH J Ggh kIQGM
OQdG G HCG LdG d AGQh dG EG .G
.kJeJhCG G gj dG WMG g
e ,GQhdG RG fe NCj M c
AT c .dG jZ Y bGdG g ,bO H
j dG RGdG eCG ,UdG UCG M JEH N
,a N CG ODj CG PEG bdG e a ah dG
N d 42.IN Ye EG ,dBV H e
g dG Gg ,IMGh H Jee H dG dG
n pS hn n go sGE p n ndGE p B p sdG o dG o oo ndGE p n sfGE p } .Lh Y dG
.98 :W | k rYp Am r n T
s co

129

AH O jN jeO YhCG

eOCG
G

eOCG

d M

fe W

dH G GC j eY :dG J dGB
Q dG SHehdG fGC Jj
ddG e jPC G fC G jN
ch .dG OLG ehJhdGh
G T J FdG Yd
dG .bGdG dG d jedG
.IG d Jh jG e dG
dH IG IjG jG GC J
J jPC G jG c OJ eYh
.IG

:dG J RL
N C a e RL
pf hCG ,L CG hM Y jdG bj dG CG j c
HU e jJ jeO IN J f j eY .b L CG
IgdG g J b .SG bdG FSG e q Jh G
d Gc jG AdG MdG CG EG ,d dH jOYh H
kH c e N C a e dG jZ Y Y dG
.MGe e Me CG
a edG dG eCG ,Gh edG gGr Ho d Y dG J
FdG j M Gk L b bH dG hM H IG J
Y IG g J a G dG eCGh .a dG e dG G
G FdG O j M dG Y gQbGh G S ec
HKh e JJ ah UJ OCH dG g J.kjCG a c
.j
,dG bdG fG a jedG jG L J
g .gCG e iG H IG jG hCG ,''jedG FdG''
JhH j .dG J Y AGQh J dG FdG UdG g jG
g J CG Y dG UGG dG N e Von Willbrand
dG g eJ eY .HUEG M EG jW dG
d M Gk OY dN e YJ IOe RJ ,HUEG e jedG

.G S Y jdG jG g J .G EG iNCG dG e
AL EG ge g Jh ,G G e Y EG H dG
.dG Y e
JhdG Gg dj M ,dG H ehdG Yj NBG JhH gj
dG Y j h RGaEG jj CG QhdG eh ,a G e
kL 20 e cCG Lj c .OG bdG b bj hCG GC j h ,jQhdG
J CG eEG fCG g .ehdG dJ gJ dG fCG gh kFc
Y EG dP j PEG JhdG Gg dJ Y eCG g .J hCG RGaEG
SG iG EG d fdG fCG J Mh .M L hM
e Y J b bh Nh ,JhdG ejW dG J G
dG J M ,RdG dG e VJ dG dG H dCG
dG dG g IH Yj e EG .a cGdGh G EG hdG dG
.cGdG g Y C j
.IG J ke G j eYh
dfG eRh YJGh gGe Qjh ,IG H j dG dG EG
43
.F Ie H j CG H
.UJ UCG M bH dG Gg j
S Y ?bdG dG Gg NGO IU e M d j GPe
Y IG dG e c GPEG hCG ,L hO dG J M d :G
cC Id gGdG dG CG g :DhdG Gg Y MdG GG ?G
ODS ,GH ke S FdGh edGh dG e SM AYCG
.G jf G EG

YGd e PEG ,G G Y kdO dG Gg j iNCG Ie


Gg CG T .''jQdG QdH'' J dGh QdG jf VJ dG
dG jBG e jBG bdG jZ je Y FdG ZG SdG dG
dG Gg d N dG g cdG Gg Y N dG dG .VQCG
.JM LGf dG WG e j dG gG
bdG EG gGJ H , IgdG hG Y dG J gCG J
f g .G NGO QGSH dG jdG YhCG J dG
hCG dG aOH YGQP j eY .Gf fCG e G NGO e
Gg bj .J jdG YhCG e G Ea ,be jH
Vh EG jdG YhCG OJh dG f H G dG
,cCG j NGdG jdG Ea IjT edG fc GPEG ..dG
eY Ga e j dG fEG Y .''MCG'' dG edG j
j Gg .kH c e HUEG GfCG e f C Vj CG dG
dhCG eCG .jd dG f Veh e e fj jdG UTCG
EG M ;kjW hj a ee QGWCH gY VG Gg j jdG
Yh .kNGO jdG c dh ,JH Oj CG fG hCG dR fOCG
CG Yh s Ln dG jh N dG e c j CG j eG g AV
N f CG h dG jEG gh G Gg .J MCG N
:d b fEG dG WN eY .jN e IMGh
.57 :bGdG| n bo u n oJ nr nan
r co n r nNn o r nf }

dG J dGB
f IG .dG J dGB 44fOGC VG SdG Vj
h ,e JH OGG e jdG H J Fc Yd
.He Ie H gG iNGC YJ NC J IG

G S

Prekallikrein

Kallikrein

Hageman

Hageman*
HMK
PTA

PTA*
Convertin
Konvertin

Christmas
Chrismas

Antihemophilic
Antihemofilli

Preconvertin
Prekonvertin

Christmas*
Chrismas*

dGDoku
eY

Antihemophilic*
Antiherrofilli*

faktr
Stuart

Stuart
Proaccelerin
Proakselerin

Accelerin
Prothrombin

Thrombin
Trombin

Fibrinjen
Fibrinogen

Thrombomodulin
FSF

Antithrombin
Antitrombin

(Fibrin)
(Yumuak
dGPht)
IG

FSF*

ProteinC*
C*
Protein

(Fibrin)
Sert
dG
IG
Pht

Protein C

Plasmin
a2-antiplasmin
a2
antiplazm

t-PA
Plasminogen

dG dH J dG JhdG

Jh dG J dG JhdG
dG dGRCGh jdG be

dG dG

HdG dG

dG
Gh
M G BGQe c jh .QdG Y SQh PdG H G j
j CG H e C .j dG Yh SSCG IdG ,dH YhT
g a e a ,dM dG e Yhe GAjEG j a ,JM aj hCG j kT
JGH H Fb J
IOe khCG G Qj :jL P J G j dG jdG EG b Y d
dHh G Gg LMh dG Gd G G Oj K ,e jdGh dG
.dG je
,dG AKCG OGd keGSG bCG dG G c H e YdG G j b
, jfdG UdG hCG cdG QaCG GNG J dG g N hCG LG C dP
J AKCGh ,bh AH bQh e cC jGdG G j .OG dG fL EG
.Pd HdG eCG LG j d
g LGe J K ,kGQT a dG j G GQG e e G j
Gd SG fEG UJ SQoJ K ,LfEG d kLh IFa gcCG AfG jh QaCG
.dG
134

o sd onJ r ndhn l ndhn o nd oojn sfnCGp VQr nCGhn pGhn n dG


s o jpHn }
o dG o opdPn @ l p Y Am r Tn u opH n go hn Am r Tn s co n nNn hn l n Mp Un
u co nYn n go hn o ho o Yr na Am r Tn u co o pdNn n go sEGp n ndEGp B r oHt Qn
.102-101 :fCG | l pchn Am r Tn

QaCG EG jdG GP QaCG Lj dG gh ,d Qe U P khCG Vj


b .d G H P cJ dG e jG AGLEG Hh .OHCG KK
KE Ge IY PdG H J IG g N .dG e GS dG g J
.d dH JOL
a dG EG jL J Nj eY
:G b e khCG e J j CG jdG
.DhGOCGh da kfKh ...fdh ,T
Mh hCG JGN e dG J dd
d MdG dGh ,e Me H hG
.QeCG c Y JS j dG g edG
c N e JEH bG c dG N d
:dG jBG d J .)rco ) IMGh
Gk er GnC nbn GPn GE p hn p V Qr nC Ghn pG hn n s dG o j p Hn }
o ond o o nj n sfE p an
.117 :IdG | o onan c

136

dG dG

J Lj
N gj YU
dGB j Lj .dG
FGOC H fE G j c GC
.G

eCG .Mh dH UN eCG d HS P OLh hOh dG e rnG Y EG


dG N .Y JGP H g fCG Y IhY ,PdG g H a a fEG
.fEGh dG GQe fEG ghh dG e dGh bG
J EG g e dG dG Y c AL fCG f dG ATCG e dG EG
e GS H QdG EG J dG IQG JH dG Gh ,dG IOLe eC
.dG je e VQCG kS IOLe ,dG
dG N J G H aQdG MdGh Gh AdG hj Ggh
.IjG Je J
137

FdG IdGh

Velcro

,dG YU JdG g
He He f H a
jf jT e j gh WG
Hh ,HG NBGh hGdG j
a HGh hGdG fhe
dG Gdh ,dH Uhh
AdG OGhQ j

dBG GQdGh GQdG


cG ODJ CG dG
.bJ hOh Qe H f
dGh dG G qb
e dG Gg fE jdG
.dBG QPCG

HY
QL jdG SG NG
Y WCG kGjL kYGNG GehO
Gg e c .''ha HY''
IgL h a .''cdG IdG''
e d kjW kbh e bSG
g e HK Y dG AGLCG
GC e GSG Ge a JdG

jQG jdGh dG H
gG e jdG j
Y kVY dBG dG YdG
dBG g cCGh ,jdG IdG
dBG GQdG g keGSG
jdG GQdG GC e Y G

N dG dG
jQG b d .eh
e dG dG g
.dYCG

IOLG eG eDJ
ehe e kGje dG H
UG Y UNh dG
g .dG J M

fE G gb dG edG Y eGC
e EG .j dEG Qe dG edG
edG J jG dG jdG G
.dG IOLG

UGdGh ddG
,ddG L cJ
YS IOjR Md bh
dG GH Z dG UGdG
.%250 H

SGbC kLP ddG N UCG


aJ dG g e ,jG dG
HQCG H .ObdG e %25 dG
Se Qb .dG e GS
j kaZ Gj CG CG UGZ

fYdGh TdG
GC e Y dG
.f G YR

dG TdG jP j
.aCG e b
jOMCG YdG GC e j

dG jMCGh fGQCG
k edEG Gg c bh
jMCG e J
.dG

VjdG jMCGh G GY
JG d
VjdG G
J IdG aM GC e
.dG jMCG e dG

de G GY J
JG J
M jdG
.dGh G dG

jeCG fQCG
AGdH Ie jY keb
,dH dG e

B G LdGh GG J
Y G SGQO IFa J
GhQW jdG SG b d ,jQG
AS GG SGQH BG LdG
DhH b dG BG LdG XCG .gQaCG
eY .aCG kfRGJ GG LQCG CGe Y
LdG GbCG Y aT FSh cJ
.ke HHdG e GQG Y dG feEH UCG BG
Y dG fHj cT U dG BG LdG
noLQn cdG g J .IG e dG
SM SGH QG d BG
45
.L EG dUe
140

GG Y dG eGC
gh Lyon Stalos J e 1987 Y fdG SdG chCG .jG QdG EG IG e
HGL jH G YS dP AGQh e dG c Santiago Calatrava. QG EG ,jdG Qd TGV
SEG IYCG J d ke GG iMEG QG JG .kGe kjQM ke
.(dG EG dG) QUjdG g j dG dG dG Gd YGdG
AdG Y LJ dG c AbQRh AGN GdCH dG Gg AVCG
QG RGdG Gg aG H .GG d LQG
.Ija a c G H 1994 Y

GbCG M .jW eR e j BG GNH j jeCG G CG he


CG MGh e e UCG kc EJG Johannes Smith S fgL QahdG
DhYCG U c eGS J j BG LdG Gg .dG H kdBG kLQ j
hO AGYCG U j ,dG H
GQGOGQ H jh ,d Hn Dr jo CG
ee Jh KdG cGh
cdG QOH d .AGYd SGG
He bdG fHdG
Gg d cGT H TJeh
LQ Y cGdG g KCGh .VG
jh GZ 0^42 GM j Gk L U BG
.bdG QeCG 4 YH
141

dG dG

bGdG AdG IOe :dG


AGQh FdG dG j bh ,VQCG S Y Gk OLh bG cCG GG J
IOe g H J dG ehG eGY MCG .G hdG L ehe g dP
.dG g e dCj dG dG
a ,kGL Nh bQ IOe dG
e ZdG Yh .H MG HU GG
jb fCG EG ,ec H L J fCG
Ib eh .dG dG Y d j
J CG h fhH IOG g J dG
NGO e H J dG dG jW Y
Gd jdG cG j Ggh .G
,LQG edG IM e j H ,a
Y IG AG d fG IOG g YJh
KCJ GG h 46.L FGH MG
e fGdCG CG c ,TEG hCG IQGH
QJ MCG H h .ke G FJ MCG
.gJ dG gGdG GdCG N e GQGfEG
dc IOe eSG d j GPe
e jZ Gg G ?dGh GFdG
.AdG H j
hGdG dG H
M GG Y dG jdG j
J :G S Y .cMh L H
kGL SM AYGC hGdG dG H
dG a j '' qeG'' Y j
.VH S jGC Qd HdG Ib
142

IOG g dG IOe
J dG dG
jd LQG dG
jb fGE .GG e
UH Jh feh
.dRY

dG dG

AGG dG j G dG
.dG cCG f Y dhDG g AGG dG jN
dG jN fJ dG dG N IOe SGH J dG g
c fc c kSGh G g S c ch ,AGG
AGM O N
CG jG g Y j fCG h .cCG dG cCG
Gk c S j CG CG ,fOCG M j CG j M Ea ,jdG YhCG Y
,e jG g H :jG g ah U AGG jG Ea dHh .kGU Mh
.fG c e WG jjdG G M Jh ,fG c e Weh ,jFGOh
CG dG Gg eh .M UCG e S cCG j dG dG g Gg
CG AGG jG Ea ,dP EG aVEG .HZdG e AL 300 AGM N c
47
.H fhe H eG VCGh jdG YhCG UCG NGO

ddG d fdG dG
G aGdG jdG G ddG S j
e IFGR c J eY .SB bdG
Y dG g Y J ,Y AdG
,''kFc G dG GQdG''
j dG dG g Y
He FN S 2^5 dW
J dG SdG F
.AdG Id kJ a fdG adG
eY :dc dG g J
jG CGJ Gk FGR Gk AV dG LGJ
YJ dG fdG

j hQg

j .OSCG U jH (fdG) adG dG M LJ dGh ,''chromatophore''


jDhd dG gDj ,AdG ac e dG TG Y jh dG dG Gg
48
.AdG e c cCG dG Jh G G dG Gg j .cCG VH
dG Gg RGaEH J jG a .e J f g dG Gg CG VGdG e
dG dG j dG G dG N fEG .aOG dh QYG se H
.d G

.dG QdG f H
H VJ .VQGdGh JdG cBG e YaO FN JdG H LJ
eCG g aCGh ,a J dG dG IOLG J J FN JdG g
Ic LQO EG kGe dG g S j .jaEG dG QdG f IOLe
Y g kGL dG e j edH YdG g eYh ,Gh G H
g J .ehe hO dG cBGh GPd VY dG g UC UG g d .QhG dG
dG G j ,bdG Gg FZ J bG e CG h ,G a jf gJ dGh gGdG GdCG GP gQgRCH dG
.jdG e J dG QgdG g Y CG

YhG dG

Md eG IgdG

Md eG Z IgdG

EG GG MdGh dG dG GP joG IgR J


.SG dG e kMQ dG cQhCG IgR fL
K ,MdG h khCG jG f IQjH SdG dG j
fCG EG dG jG d d dG cQhCG IgR EG gj
.cQhCG IgR J e ODj dh ,kMQ g j

J
UN
GQhCG :JdH
Y dG AYCG g GQhCG
Jh cCG a ,JdG
,bQdG d bO eCHh .HdG cCG K
,ge H IOheh bQh kGL N bQdG J
J .QeCGh AGd IR eh Jh e dP e dh
AYh cCG e AGHG M Aj YhCG e Y bQdG
fG e Jge AYh UCH AfGh bQdG L Y
f Xh EG aVEG fG dG g eDJ .bQd dG
.AGdG

dG dG

dG J
bd fG
d kj cCG cG fCG J j e kdZ
J dG cG J :G S Y .HdG jdG fCG e
,jHEG Y J CG dG J e H cCG hdG G
GC e Y dG j H ,fe SSCG Y J hCG C dP
CG .kGJ cCG J C dG edGh ,FjdG dG
e x GC qJ hCG Z ODj PEG .ke kag dG UY e Y c ODj
.IFdG Y dG j MGh Y dG Y dG Y EG UdG g
dG fCG .dG g ANCG a J dG edG g jf
CG edG g e C ,jJ H cCG ANCG fEG j
dG FGdG QdG YCG H CG e ZdG Y ,CGh HdG jW Y
.ANCG L J G e j fCG EG ,GSCG EG G NOEG j
fG fCG a ,dG fG fCG Y J cG g dh
qf CG . JEH bG L Lh Y dG N .dG LQO EG bJ dG
:ddG eCG N e G G P H Y

G Qf L
Ljh QTCG hL J dG bdGh GG Y G Qf ij
YCG IQe gJ IQ dG QTCG TYCG H QdG g J .dG jW Y
.G a
OGOjh ,IMGdG fdG Gf Y EG J H e j CG bG G Qf j
.NCG G Qf eh kM UCG GfCG Y 20-15 H e EG d OdG Gg
YdG g .S/c 100 EG J Y YS Ea ,Y H NCG G Qf j eY
146

(ed Ue) dG dG
QG YO f
dG

dG QG
eCG H QG SCGQ ajo
Gh

EG dG QG j ,eCG EG dG N dG aJ
.bdG Y dG edG d YCG

.edG g Ue dGB jd LJ dP eh ,Ic eU GE dG Q Id f AKGC G Qf Vj


Qhj .FdG d g :fdGh ,edG IM e J dGh QGh G H UGdG dG fC G g :hC G
gj gh G dG Y ah J dG fG J .FdG SGC Q ee e d G NGO dG
.jdG TJ GC GE (SE G jG dG) edG bJ Ggh .If c Y dG edG J

a iNCGh IdG H UdG edG eCG .RdG M M M j dG ZeO Y KDJ


,ke e N QGh SCGdG j dG FdG GC j eY .fdG e 1000/1 e bCG
.edG EG ODS H GfG Ca
SCGdG V j dG J Y J dG bdG g Y J dG edG EG
.HUEG e f C VdG j G Qf ed p G dG CG EG ,SEG FM
Qe CG EG .dG dG cM e QG jh H QdG e Y G Y J
Y dG edG SEG fCH H Y e G Qf FW Lh
EG .GQdG edG Ue e aCG H Y fG IOG g ODJh .dG Y
JOSGh Gk L Ib UGH dG edG UeG Y JQb e JCJ IOG g IOL
.GTCH jG LddG JRaCG dG OGG JOH J gh ,QdG Y dG d
Y QG a EG .fdG bW Y FdG AGOCG M M dG g J
QG Y Gk H cH FdG eO dG Id J bQG jdG H G
49
.edG UeCG fK dBG Jh J Ggh ,dG Y AKCG dG
147

dG dG

dG d G dG :ZdG
hj e CG ,VQCG Y YJQG V 100 EG J Ib j CG ZdG j
hO dG j CG dP Y IhY .fEG H j dG dG dG e Gk e 200
eEG j H ,eG IdG H bS Y ZdG j Y H .YS 78 I bJ
J H iPCG CG H j h QGhdH j a dP eh ,X Y hCG SCGQ Y
.jdG L
e U W e dG Gg dCj .L NGO IG d dG dG Vj
Gg j .dH jh G ec j dG ''JhdG'' SG Y j ce
Jh edG dh ,cG IOhh Gk L Ic e FGT e LQG dG
.dG Y dG edG
U eO FH NGdG L j H ,jeO YhCG CG ZdG L Lj
dG N e dG J J .LG GdG V e jh NGdG AYCG j
,G AfCG IG FGdG

d ZdG No
dH gT Gb
ij dG
.GG

148

j hQg

b U .UGe H cCG H J V e EG LG Y j Ggh


Y jdG GdG Y KDJ H AH Jf e ,fCG T Y IG g
.WEG
J dG GdG J dG IdH d ZdG S Y CG AdG cG
dG HdG dG e f IY ZdG H j gG AGOCG Gg CG EG ,FGOCH
j PEG ''resilin'' Y j dG JhdG H j dG Ggh .fS EG VoCG
bdG e % 97 j Y JQb g IOG d IRQdG UG .fG bdG ZdG
GSCG IOLG fG OGG J dG dG iJ H ,OdG Y NGH fG
S IOLe Gk L bO FSh fG IOG g VJ .bCG c % 85 EG J dG
fd W AKCG IOG g d fdG QYCG H ZdG j .G IG
.dG NJ CG EG je dH H dG dG .Id jdG MG
dG jjdG IOe fG bdG N e Id HG IdG jH HdG dG J
.Y Gb EG LJ

J gG jG IG g . ZdG Y J dG bdG GG e f gh ZdG e ge NGB


.ZdG L J dG dG
149

e No dG dG SS KJ
.ge H ,ja ''K fGC ''

j hQg

dG dGB h dG SS
IT IK Y dG SS J
kjW keWN IG g .dG
Gg jf h , L e WCG SCGQ
j OM U Qe Lj WG
.SCG
Gg IG iNCG MCG
,L e e aCG H WG
IG J eY .gS AKCG J M
gH eWN LJ dG IK Y
fSCG J K ,dG dBH TCG d
WG YCG IOLG QH dG
e SCGQ H IG GC J .IdG Y
QG YEG j ,NBG EG fL
.SCGQ e G eWN j dG
e ke aGj IG g SCGQ U
e ge i j a dBG g
.fhG
dH IdG K AKCG IG J
.dG SS bj

151

dG SS J
dG Y AKCG SCGQ
Ve g c ke
.IQdG

H SdG J dG Y e AfG H .jQd RdG JhdG J Ggh ,gG


GC Jh IdG NGO bj dG J .d U dG IG J IdG Y IMGh
.edG OGORG c c ch ,c c cCG IdG e bdG dG ch .edH
J dG edG g gh ,dG e jdG WdG J CG EG dG g J
eCG d U dG dG SH bdG g J .IdG IQOe feEH UCG fCG bdG e
f M ag UCG BGh .Ic IH IdG dG bdG QOJ K ,jdG fSCG jW Y
GS N I Jh IG Qpr oJ g .S 30-25 H e hGj H VQCG

j hQg

e gQe Jh dG IT EG J a SS J eYh ,VQCG


p Me dG bdG j .jL
Y jG dG Y Qr G
g IM IQhO N e G N dG Ie d J iNCG Ie . 50IdG
IG H J RL hCG f c EG .ddG QdG jf GAYOG Hh ,IG
SCGdGh ,WdG SCG ,bdG WG : d f g
hCG aOG AV Y J Gg c ,dG Y Yj dG G
K AY e cCG j a ,dG Y WG j d .''dG AUG''
.''Me KEG Me'' QdG VGaG eY cDj ,e IFa Yh
bdG CG j .dG g '' Ie c'' bdG AYCG hJ ,iNCG L e
jG ge Jh VQCG kY UJ CG Y ,dG IK jW d jb kfSCG
.dH YQe
Gg c .jdG VSh kjW J a JG IQhdG g e SdG GPEG
.N M dN U e g H aOG jW Y JCj CG
.AT d ) o pdnG ) g RYEG GH IG g dN dG

GC H GG e d kY WdG J
e c OY j .dG SS eJ
.IQOGh bdG QW dG IK GG

153

fG aC G
Genlisea

dG Qha .bdG fGG AeCG G a j


adG j .e a HfCG Y IQY g VQCG
Yj ,HfCG IOLG dG N e dG NGO EG
GG J M ,bdG NGdG GdG dP Y
GT J .G aJ H NGdG bdG dGh
,FG QdG a dG bCG L SCG f J N
J dG ,VdG OdG e Y jdG LGJ jdG h
,dG NGO EG GH aJ fK Ib Jh c
51.kGjd AGZ jdG d
k
Bladderwort

f a

G Y j FT jH f g Bladderwort
g a OdG e GfCG KK g Utricularia. SG dG
h ,dG QN VJ dG ''jhdG OdG'' hCG dG :dG
IdG'' h ''HG YHdG IdG'' g ddGh dG YdG
OdG g dG J .NGdG VJ dGh ''HG FdG

Genliea GQhC gG dG

dG H Vj (CG) GSCG L
NBG GSCG L j () dG
.T a jf ()

eJ -1 :dG JJh Bladderwort d e


NJh G dG jo -2 dG GT jdG
.jdG AGQh HGdG J -3 .jdG

jEd MGc
.dG
OdG J jGdG
G GOGeG H
.e AG H CGJh ,H
Gk L g L j
Lj NG h .NGdG
e AG dG dG H
Jh .NGdG f adG
NGdG IOLG GdG
eJ ea ,kGL SM
dG hCG GG iMEG
AGQh dG jh ,dG NGO EG IH AG aj .QdG Y dG j GdG g
GC J ,fdG e 1000/1 e cCG J dG dG g H .dG H jdG
52.QdG Y VdG GRG WEH VdG GQdG

dG dG

dG dG
cG Y gYd ,''dG'' Y jh ,dG j e jdG GfCG H J
dG G M cH jd jh G AH dG Gg j .FdG G
.IO YH H ZJ
fc ,kjJ Y b dG Y j dG H CG EG ,eR e dG AdG Y
,dG jZ ''Y '' N e j dG Gg CG GcG d .d Ic ICLe
.kFT c c H jRGgG dBG jW Y dh
h ''HdG'' :d FQ BGL g .FHdG G CGe Y aGdG G H J
.''QGhdG''
J .fe cM dj fCH jdG fCG bH Y dG dG j
eJ dG bda :bd UN Qe jd EGh , ATP jL fG bdG G
Ie d NGdG dG .LQG G ZCG Y aG fjCG Y J jdG
SG fe e c Vj .dG AH JhH 240 e Qj e j .kGL
,bJh G J GQTEG dG g JhdG g CG AdG j .jY H
Uh e iNCG JhH u J c ,jQP j cG J JG f Jh
53.d IG dG Y J dG Gg Y H R dG FG g .G AH dG
M a .QdG jf H Y kac kdO IG cH dG dG j
.jd IFa fOCG hO UCGh dG Y d dG Gg e U AL oa hCG Jh
YGG kjCG J G g .a N dG dG e ke dG Y j CG j
.''IN IN QdG'' GC e J dG jQdG

kjGC dG N J
.cM dG

156

j hQg

eG JGh T
WdG G
G
FdG

LQG AdG
eG
27 nm.

dG f

SG

dG NGdG AdG
(eRH) jdG

ce aGhO
hM H

aGfd SM dBG

FGdG G HdG

JhH ObH

fhG jdG

fhG jdG H

.dG e fL Y kJ kjCG FGdG IG TCG CG Y dG Gg dj


dG AMCG AY j c e CG cG c ,dG Gg FbO fEG J ch
.gc ke Ie bGdG g ,H Y hCG An MCG ,jhQGO a ,Y SdG
G dG

UdG) N
(eG
LQG AdG
L M
p M

fe fH

b (Le Z)

NGO (eRH) AZ
S M

FGb

G AG
cdG AG

M M

C M

dG eC G MGC dG dG j .''H''fhj TMh jhQGO c dG IdG bG M e J


dH eU dG Gd NGdG UdG Y dG c ,FH G dG Gg jH AG jdG J ,J
J g k kjL Y N e j dG FHdG G Gg .e FHc Y jdG GC J .Ad
.YC G j c JGP M e

157

dG dG

ddG dG
j Ggh ,jdG bH J c ke FdG jW Y Gh ddG e c j
IQG JQjR AGQh dG g Ggh .SCG e AG J CG Y dG e fCG
eDJ jH dG Gg U .AGdG NOEG Y SCGdG YCG b IOLG dG J .d
eYh ,AG J AH kJeJhCG dG J M ,AG UdG Y keBG kbZEG d
.kjCG kJeJhCG AdG j dG EG ddG Oj

dG VdG hO dG j f
dG g .d a j Ie c AGdH FQ e % 90-80 e ddG C
dG VQCG jdG bc dh OGQEG Y ddG Y gh ,fEG Y %15 iJ
54
.OGQEG H dG g a J
L Lj .gf hG fQf c ke j CG ddG Qj :NBG H
edG f FdG ddG j .AG j eY dG e j UN f ddG
dG EG ke dG dG j hCG dG j a ,bO 15 c he H
j dda ,MdG Y j NBG Y g Qe OLG ddG N EG .d
dG ddG N e jG dG J .dY YH jh AG GNG bCG bW
.ddG j c YS IOjd dhQdG ejdG ah G SdG j
H ddG N
H Fj e

jdG YdG
G fG
ddG rnrpe

dG
G
ddG N

FdG

bdG AeCG
fdG

jQdG fYdG

158

Yf L

dO

FG iG

ddG Y YLG IG
OG dGGh fEG jG e je LCG eh .Ic Y ddG j
GC J . CG EG dG V J H ,IMh J a jG OGaCG eCG .dG ce
.dG EG jG OdG a j dG hCG dG e ddG Y adG HGhdG
d j eCH ke j jdG H ,khCG CG MQ e ddG ddG jP j
dG EG G VdG j ,jdG SCGdG j eYh .VdG AKCG RdG cCG
dG CG aGJ IOY ,IjG IG AGg e c hCG d
e gdh CG YJ .iNCG fCG Vd J
G dG j dG OdG j PEG ,IOdG
T e dN e G j jQe e
dG Gg Y aH ddG j eY .CG H
VdG ddG j .G aj dG e
Gg e %50 gdG Jh ,kej G e GdG GY
J Y G Gg j G h ,(dG M %15 f e fQe) G
IdG adG jNCG fEG J .ddG IQGM LQO d jQhdG jG dG
,QfdG J ch dG J j dc .SCG EG aH dG UdG N
M FH kee ddG j MCG H .GS IY J dG dG dG g
.S KK

Afd fG dG
Gd SdG bdG .fEG dEG j CG YCG EG Uj CG ddG j
aGd Gh ddG U .MGh nn H e 3000 M Uj G e f j dG
159

EG IOdGh UdG Y jdG J j .dG UdG e dG Gg e


.dS cCG dG
ddG j eY :FdG UdG YJ dG iNCG AYCG e
FdG NGO dG OGOj dHh ,ba AG OY V hCG Rh OGOj AG
a dG Gg EG fEG FQ VJ d .QG e RGJ OjE
ddG L UN YaO f LhCG ,G Gg Y dG LCG eh .G
FGdGh eZdG jd eRdG jG e ahZ M eDJ M
.ddG FQ NGO IOLG
.Afd fG dG g FGdG G Gg jG fCG e NBG f
.G e dG Gg LGj ,Ic YH YCG EG SdG j eY
YdG J dHh ,dG EG ITe AGdG NO AGdG S jh
bYEG e J G EG YdG g ODJ CG .jGdG FGdG
160

j hQg

J fCG e ZdG Y G g LGj Gh adG CG EG .edG GQhd


fCG g dGh ,FdG jW Y
d a FQ AGg fCG h ,ZQa FH UJ
.Afd VY
AGdG e ZQa gFQ fc GPEG :g GDS EG Oj Gg CG EG
f H J GPa
?cCG
''HZG'' JhH j GDdG Gg Y GG
Gg j .Ic H dG fCG LGj dG
,cCd Gk L jb HPH ,''HZG'' JhdG
EGh FQ cCG G Gg j dHh
j CG GhCG ddG j .JY
Uj CG c ,jW Gd J hO
dG G j .jj dG dG EG
CG d ,HZG Y kjCG
M H hdG f j
dG jdG dG Gg .dG
jBG e jBG Gh ddG H j
N .p G dEG Gh G
fGG bH e jdG dG
J dG hdG SJ H
.H

161

dG dG

aGQdG Y G
cCG e IMGh (QeCG 5) QdG dH aGQdG J
G Gg CG EG ,dWCGh VQCG Lh Y fGG
d Gk ja kfGQhO kef j QdG dGh dG
j ,jGH .fdG g S IM G Gg j
,j dG a Vj dG edG EG dG j CG
b dG N dd ,d jOY Z H j Ggh
.FR e 350 V dG d j kjb aGQdG
j CG G e dGh - dG dG Gg
T J UN aZ V LGj - OdG fEG
.G HUEG e dG e jdG YhCG e
e j dGh SCGdG H e bGdG G U M j l f Lj ,iNCG L e
,f fRG cG H YhCG aJ dG FGdG J .dRdGh IYdG YhCG
.kNGO kaf J b dG dG V IN JQG e GG j
cdG EG .UN jY ,edGh bdG UNh dG j dG dG j
c ,dG dG d fG VGYCG e G Gg WG g d YG
.dG J jedG YhCG eU LJ
.AG d LM Y H J dG IAfG g ,aGQdG d VJ dG cCG G
e J aGQdG CG EG ,kNGO kaf d j kUCG JG dG Jj dG g
dG cdG G FdG gh UN FS j PEG ,G e dG Gd MG a ,dP
YhCG hM d ce V dH ,QdG OdGh edG kjCG Lj
.SCGQ GG j eY Jh MGh H J eU RGdG Gg Yj c .jdG
e jh .eCH AG e dGh SCGQ N e aGQdG d ,dG aJ e edG J
e dCj dGh dG aH edG g J ,dG JQG QNCG OJ Y UG
.W IY
162

jdG YhCG e T

jdG
JdG
NGdG
fdG
jhdG
G

edG
J dG
dG Qh
LGdG

NGO Ye jT

FGdG dG

LQG dG
SdG dG
NGdG dG
fdG

dG
eOCG
jh IZ
gP IOe
rnjoT
Y
r jQn ho

jCG JdG
CG JdG
YG jdG

dG

dG jdG

jdG
ShCG

UEG jdG
NGdG dG

SdG

LQG dG
p jdG
rdG
(edG)
fdG
dG jdG

edG
J dG
dG IOH
FGdG dG

dG dG

dG Y YadG G J
Y AdG Gk QfR 30 j .HQhCG dG AGYCG dCG g HdG bdG HfdG
.a YS K N G Le Y f 30000
gRGaEG N e dP Y jNBG J fEa IjL f Ie HfdG J eY
Y a dG d JGOGSH GC jh FGdG g kjCG dG j .UN FGQ
LQO IOjd RGgH dG GC j .Jb f 500 J HfdG J eY .G Ne
.HfdG jdG h ,QGM a NGO fCch QfdG j c H h .G IQGM
LQO 48 EG a IQGG LQO J dG AGG WG dG Gd jQGG IQdG J
55
.Hfd dH Jb J fCG e IQGG g j CG d hCG ,je

JHGE J eY dh ,JHGE g dG Y YadG dG


LQO e dG jJ .QGG dG GE dG C J da Z
.jdG H HfdG J gh gAGYGC LGd L IQGM
IQGG LQO J M dG Gg e ce IQdG hJ
.je LQO 48 M AGG WG

164

j hQg

OdG Y KdG Ie
dP e .''ZdG'' QJh dG a jW Y KJ OdG CG j e dG
.gdG e dG j e e ,OdG Kd iNCG dG e jdG Lj
CG J CG dd .dG AdG Y gYJ U OdG
dGh HdGh iQdG ke OdG LGJ .dG IQdG GY e VQCG Y e
g jQGG WG CG EG ,kGe 5000 EG J b dG YJQGh jfBGh jdG Lh
40 CG M OdG MOJ IG HdG a ,OdH keMORG WG cCG
.He je c e kYf
iNCG GfCG LhdG Qj H ,VdH jdH a QcdG j GfCG H
bGJ jSc OdG QcP J :G S Y .gMd CG H J hCG G H
ZGdG J IQL GO Hh ,J CG EG kej 12-10 H e hGJ I VdG
IT CG J dP H ,IMGh fCch GhJ M H Jh eCG QX
gQU CG J ,AH IA Jh AdG f J JHh SCdG J Gk QgRCG HdG
.eCH d cJh IgdG g
.gQU jJ LCG e e Z kVH CG J ,AGZ Y G g CG h
56
.GQgHdGh JhdH G VdG g Y ZGdG iJh
.VdG Y dG G S dG OdG e NBG f g ''dG dG''
.jZ NO CG a Td HUCG kcGTCG dG Gg QcP
afra)dG jaEG cdG j
Y AH G dG QN H (Nectophyrne
J dG dG QfCG hCG GdG WGT
S Y OdG J jdG H .VdG H
IU cM ODJ bh .cCG d AG
EG kjb j e hCG NBG V e
165

.WQ We G VH OdG J
Ic ShDhH VdG g e ZGdG J
dG YGQP j SG bdG h .jPh
J kGNCGh ,dG T NCd bSh
.fdG e IQG jdG

dG S V j
J .(1) jSc
,J M VdH QcdG
H ZGdG GC J K
h JeCG QX
.(2) ge

3
eCG e hd H J CG X EG QdG J eY
QdG Gg jh dH CG GC J K .(3) IMGh c
SCc H hJ dGh IdG g QgR EG J eY
GH ZGT CG J .AH h AdG f j
.(4) eCH Gd SCdG

LLdG OdG

j hQg

cP j .cCG hO d J M dG EG dSQEGh VdG e eG g eJ


c e jd bH AG H j g SGG Ia AKCGh ,VdG SGH dG
.FZ Y dG EG NGdG cCG
dCG a ,VdG SGH j ,LLdG dG gh ,OdG e NBG kYf CG EG
QfCG hCG GdG Jfh QU Y VH OdG g J .dd iNCG jW dG
.AG EG ZGdG J VdG J eYh ,jQGG
aGJ dG G OdG GfC dGh YGdG dG e PdG g MJ
J bG L CG SSCG Y J dG jhQGdG YGG gQU Y FSGH
c QU Y YaOh jYQ MGh d dG dG EG .AdG LCG e fCG GU
OLdG jf j bG d cdG dG CG Y IhY .SSCG e G Gg d
G FdG N b dG CG Y gGT bG g J .jhQGO Y b dG OG
r op rNn h}
n :dG Hc d J dG j .de IM d J dG IjdG a Vhh
. 4 :KG |n o pb jo m r n ud
l jn GB mHs G On pe to jn en hn

IG KJ OV
Y j OdG e f j Y KJ jW G G jBG e iNCG jBG
dh ,NEG H VH OdG g J . Rheobatrachus silus'SS SNGJH jQ''
dh .dG H HSCG S I g Jh IG VdG J .d H cCd d
fCG bJ jGdG .H dG gM dG G dG fEG ?oJ CG hO g J c
,a ZGd IdG g W J IG EG :CG ,HSCG S I dG hJ Y
d ac JOG Jg ,dGh jQchQdG M fEG :gOj NBG N g dh
IOLG FGda .UN je Vh H QeCG C ,j Gg dh ,QdG
gRJ prostaglandin E YJh edG J iNCG IOe jW Y Jj J CG Ie
Y IL H QdG j Ggh .iNCG L e ZGdGh L e VdG c
.M cH J fCG e ZdG
2

169

dG dG

LGe Y fG cdG J e kdZ cG g .OdGh cdG H QhJ WdG eM ce


.dS H dG b dGh edG Od
170

j hQg

dCe kjCG g ?ZQa Ie ZGdG iJ c :g f j dG GDdGh


Z QU Y fC ;e H gZ e cCG GfCG g VH EG .d
.HSCG S I QdG jd j ,JhdH
,jdG Y MdG Oj c ke VdG N G Oj .IOdG bh M BGh
57
.eW hH fCG GC Jh dG dG EG IGh G e c Oj QdG J eYh
f fC ,SSCG e jQdG YGG OdG e dG Gg id KdG f j
jH Je dG g e IN c .HGeh e
CG J CG j .ZdG Gg IM d
S I dG hJ Y bJ CGh khCG VdG
feg IOe RJ CG VdG Y j H ,HSCG
VEG JhdG eCG .Id eG SdG q d
J Rheobatrachus YV
.iNCG IQhV a dG QU ej dG
.a e dWGC
d aOG jW Y fCG e Sj CG
dG gEH c GMCG e dG g Gk U c e N CG EG .gQU
.dG VGfG j G Gg QGSGh
e dG Gg Y dG '' IN IG'' jf J CG dG Gd
e JUh Jee eH ''Rheobatrachus silus'' f e V hCG No d .OdG
e J :JGP G Y gJ BG M gVY dG bG c EG .hCG dG
dG dG e fL Y bG dG N d .fcQCG eH dG j
:gedG dEG IQdGh G dG fEG ,J
n go hn p VQr nC G hn pG hn n s dG en o nd o u njo n roG Ao B n rSnC G o nd Qo u noG
o Qp n dG o pd nG o dG n go }
.24 :G |
o p nG o j p n dG

171

IQdG g /dG dG
IQdG g CG EG ,gQGVCG
YCG UEGh f j
.dG J c

CG Y j FQ MG
e f d Vj b eU
CG hCG Qj CG eEG a :fCG
gh fEG g U .Qj
e dGh edG d IQdG

dG
GedG / dG
,jDhdG M V G d JJ
IQDdG jH J dG CG EG aVEH
adG Y Ie JeJhCG H
dG J Ggh ,QG FdG
.dG Gec c Y be

dG / PC G
G JdG
GUCG jH PCG bdG jdG jG J
.dG Ge J c ke FHc GQT EG
-20 H e a GUCG PCG J CG
dH kde dG Gg jh ,Jg 20000
S e d dP e cCG dG c d .fEd
GUCG Mh dG GN e kGQYG GUCG
dH kje d VdG Gg CG EG .G JG
.G dG e dG OLd fEd

cCG IOe Gg ej M AdG j


GfCG VJ . dG dG e SM
aCG SG jG e IjY

W
f

G /dG
keO JM N fEG b j .dG
M e c MS cH 500 A j
CG e C .e e 300
.Ie fU hO dG GH J

eCG MQ gh Y fEG b GC j
bJ hO bO 200-70 jh
bO c N dG Jj .IG ie
bO 10000 JbO OY Jh bO f

dG f / dG
dG CG c ,bCG JM CG EG
bCG H J dG OGd FdG
.dG IOLG J e H kGJ
e fQH kGL Ie fCG dG EG
.YU Ie CG

QGe e H fEG c J
c dG e (kfdZ 37) kGd 140
dG TG e e Y j
J CG .''fhdG''J
cCG c H YdG TG

SG /edG
IH fQe Y gCG GP Z jG
EG eG J Y N jH
YU e Lj fCG EG aVEG ,edG
hdGh dG SM j G MGh
.edG

GMh Y Y N c
ed .eG f Y a dhDe
J dG dP Oj kY dj CG MGh
QjfGJ IMh e 4^5 H
g J .jG jhG G

jdG / fedG
GQhdG dG AGLCG e AL c
,YCG H Vh G
J dP eh
FShh .JG
FSdG GMh
g JG
C ,fe EG dDG fedG
dG AYCG .Ic jL e
g J e Lj
FSdG FSdG GMh
.UN SM
Y J dG
dG fedG
jH J

FGdG G / dGh

RGe H dG
RGJ Y ad
Y Zh ,G IQGM
AGOCG bOh YS CG dG
dG Gg
jdG fCH fQe
fEG gQj dG
. c

dG cG gJ
dG G aJ
jdG H h ,OQdG
e %90 dG aJ
L e .G IQGM
Y dG j ,iNCG
dG G jJ
Jg J .QG

G /YG RL
db JGSGh ,ca
Y f Lj .UN
bO H VQCG Lh Y
G YG dG dah
.dG

H 200 eLCG
keh .YaO AH O jc
,TG Y G g c
jedG jdG g
SCGh ,kjQSG kef

G / G
ObdGh ,bd dG G g G
IU jL g J dG
da EG ATP . Y j
CG H J ObdG Gg M
G g ODJ .fEG U e
dG iNCG G e jdG
U e dBG CG gJ CG
.dG

fE G :G dB G
p p ndG nuHn pH ns nZ ne o n fE p G n tjGnC B nj }
| nndn n an nG s n an nn nNn p sdG @
.7-6 :QfEG

G dGB /YdG

g YGdG QGh aGdG Y YdG j


Y dG cM J dG ,dG
Y G dBG J .ANSGh dG j
dG IdG c IQG J Hh ,CGG
IdG IH YdG j CG ,
.G

dG dG

dG :YC G dG
.AGS M Y AMCG Zh AMCG dG J ,J dG SSCG fGb g
Y VJ gh ,a J dG AMCG J c ke d G G Y J fGdG g
SG Y j dG fGdG g .FjdG cG c Fja fGb Y Gg ej
c :fG UdG e j) .dch dG N Y kfgH EG d ''AjdG fGb''
.(j hQd dG N
:G c Y eCG H NCd
.''AG Lhd'' :G A IOG FG iMEG eCf :G S Y
SJh de AG Lhd CG EG ,LhdG e Jhe LQH aNG Y FGdG J
,Y g H cCG AG Lhd LQO fc a .dG Lh Y AMCG GfCG c
.jdG HfCG Y AGdG f e dG
dHh ,Y g Y Gk c kaNG QfCG jL a ,Y g bCG fc GPEGh
.HdG d QdG J dh ,dGh jOdG J dh ,G J j
.VG OGGh HhG e eLCG dG jedG jG GQhO AG j dc
dG JQh ,YhCG jG g cM dS,Y g Lhd cCG AG c a
ac dG eCG g M .dG Gd eRdG bdG Y G j bh ,dG V
:J dH AG Uh Y dG jBG JCJ .AMCG LCG e kN N b AG CG Vd
n Qr s dG ppH ond
l Gn n T o r eu osd Ak B en Ap B n s dG n pe n n fGnC p sdG n go }
o p jo @ n o poJ ppa l n n T o r pehn
:dG| nh o sn nnj m n ud k jnB npdPn sGE p pGn n sdG uco pehn n nYr nC Ghn n p sdGhn n orjs dG hn
.11-10

idG RGJ
cdG hCG dG S ?BG Y g cCG fc HPG Ib CG d j GPe
f j Ggh ,cG e Gd H cCG bW GGh fEG S .ke
174

e G ATCG J d ?Y g Ib bCG HPG fc d GPe .VQCG bdG OQGe


S .jW Gd AGdG ke edG GQP S ,VQCG Y Vh Y G
h QfCG cM C WS .VQCG EG J CG b J bh G Gb bJ YS
.dG P UN Y j dP c .JGP dH AHdG dJ
175

dG dG

aGh c Y J ATCG H PdG Ib CG Y HPd Jf fb j


J PdG Ib Ea ,Ge K H aG YJ GPEG dd .H UdG
.VCG HQCG HPG Ib OGOJ ,dG EG aG fG GPEG hCG ,VCG J
HPG fb c d .cGdGh dGh VQCd G VdG a Y Yj fdG Gg
OjORH OGOJ HPG Ib fcd :CG ,Gg Z
Sh gEG dG GQGe J a ,aG
d dP e VCG fc dh .dG Y
d :QNH .dG Y Hh HK e VQCG
VQCG eU bdG H HPG Ib J
.AdG YCG Yd hCG dH

?Gk e fH HK cd GPe
IQGG M .bdG e kTCG LGf f
.Ie jRG N b QdG eCG gf dG
jW Y J bdG CG AjdG Vj
.''fGc'' YJ IU jL EGh ,jQGM Le
Y j HK b J G bdG M h
.dgEG H Gk L U bdG Gg .fH HK
b GPEG 6.626x10-34 kjJ gh ,dG HdGh SSCG dG e MGh fCG EG
.kFGO HdG bdG Gg g dG Ea ,TEG M e dM CG OOdG Y JdG bW
.fH Hd J gZh ,AdG ,IQGG e WehdG bdG GfCG L
g Y Gk c QdG eCG gf dG IQGG fd ,kb k HdG Gg c a
U AL j CG M jQdG bdG J CG eEG :bG MCG j b gh .Y
Y dG H jQf Ic J M a jQGG bdG J CG hCG ,bGMEG e
.gSCH VQCG aJ
176

j hQg

MG Ib
c dh .j c dG JM LGf kYe Gk eCG MG Ib J e kdZ
GQhCG dS ?MG Ib g j d dG hj CG
dhdG dSh ,VQCG Y Jh jjCG e bCGh
J ATCG S :QNH ,adG jhGR H
.f NCG jdG bJ CG EG QhJh
Jh ,dG c J CG MGdG Y
j H ,eCG e ZGdGh eG c
.jfdG EG OOj GU jh dG
bGh dG dG fGdG g EG
.Y dEG Ib f EG g e a J dG
Uh iS d FjdG fGdG g bGdG
fGdG dG N d .dG YHCG dG dEG d

WehdG bdG TGC c fH HK j


HdG Gg M J d ...AdGh IQGG e
gf dG IQGG fd kb dG
gh .Y g Y kGc QdG eGC
J GC eGE :bG MGC j b
j GC M jQdG bdG
GC hGC ,bGMGE e U AL
M a jQGG bdG J
dG H jQf Ic J
.c VQC G aJ Y

dG dG

H j LdJ e c
IQdG a , MG I b
.MG Ib H j

.f Y dG jh dEG dG jh jh id fEd gSh fdG


e c . IG dEG IQd VJ dG eCG VY Y fEG bj CG
e IQH dEG dG ZU e N EG d dG je b N CG e dG Gg
.ge Mh

178

in nJ es kbn Wp mGhn n Sn n r Sn n nNn p sdG }


in nJ r gn n n n dGp Lp Qr na mhno nJ pep Ms dGp rNn

n r ndEGp
r p n jn p r nJs cn n n n dGp Lp Qr G s oK @ Qm ooa pe
.4-3 :G | l p Mn n go hn Sp Nn o n n dG

dG dG

:e

Qq ``dG `jN
fc M RhJ dh ,AMCG N Ia gd X jf g :jhQGdG hCG Qq dG jf qEG
jdG g YqJh .QfGh CG Y IH jf fc EG aVEG ,U jCH dG EG nrn Sn
N K eCG ihJ e YS AYqOG Gg qdh ,aOG H M OGe e Cf IG CG
dG T H g bdG jRGG a Vhh dG N da .Lh Y dG b e AMCG Zh AMCG
dG N '' aQ IH J W FU J CG d Qq dG jfh .Jh fS WdG
. Y kH '' aOG'' e Jh '' Fd
H kGMGh gqJ dG dOCG CG ac gOHCG e jdG g fGL f eY dHh
dP Y eh .G Z FdG e kGJ cCG G FdG OLG QG da ,NBG
G HCG AGLEH jRGG g CG fh ,jd SM jRGe ah IOLe a GQqdG
SSCG OGe J a ,kGJ cCG NBG JJ ah G dG IOLe f GQqdG g qCG qGE ,Y
.ge LQO EG SM jeh dBG d Sh Jh ,jGh fCGh JhdG cd
. jdG dG jf H jdG g YGe d kFQ kS c QG dG Gg qEG

jhQGdG eg dG YG
dGh jdG cGdG EG Ie Y SdG dG G IO IQH jq dG g X
g dG fO b We d Lh jdG g QH dG G qdh ,jZEG IQG EG gQhL
G FdG N Y Hc dDG VQjh .jhQGO dQJ dD '' GfCG UCG '' c QhU
ac G FdG Af Y G OYG EG Yj dP e kHh ,Jh fS dG b e G
.dG GdG hM f AMCG H NG X edG Qhh ,MGh L e
aGYH qGE d ''ke kL'' fc Rhj h ,Y dO CG G j jhQGdG AYOG Gg qEG
aGYG de IQH hJ '' jdG Ye'' SH H Y iMG dG EG M ,kT g
dH d ,SG OhOdG d CG jdG J dG SCG e jdG OLH f jhQGO
.jdG H ja GK
jd Ib e dG QdG d SCG d SG OhOdG QH dG j CG j ch
180

j hQg

kGMGh YGe sah jhQGO eCG N jG dG dh ,Hc cP W dG Ggh .edG Qh


. NBG H
:gh Y jc jhQGdG AfG EG OCG FQ eGY KK cP h
.Ie hC IG QX c Y Y J OjEG ke J jdG qEG (1
.AMCG H d Sc '' Qd UN dBG'' OLh Ia Yj Y dO CG OLh Y (2
jf YJ e Y IMGh aO AMCG OLh d J GG j dG
q qEG (3
. QdG
:KdG eGdG g dH Sh

Gk HGC bG Z IG : IG UGC
dh .S Qe3^8 b IMh N e Cf cCH G FdGh IG qCG QqdG jf YqJ
dG M e G G FdG GfCG e jG EG J CG IMGh M c
?dP J Ge jCG LJ GPa ,ka M b dG Gg c GPEG h ,cdGh
AYOG Y bdG j UdG g VG bh ,Y HLEG jdG J DhdG Gg qEG
X G g CG jdG YJ ?OLdG EG X c iJ .''CG G'' J Gh hCG
Z hCG LQN KCJ CG dg j CG hO dG hX X h gMh aOG f OLdG EG
H d KM M Z kGOGe qGC jdG YJ :NBG ,kWb kaQ G Ia aJ fEG CG ;W
. ahG dG YGdG ac e ke aj AYOG Ggh ,M N QX EG dH OCG aOG

''IG e C J IG''
dG gG W qj dGh ,QcG Hc IG UCG Y kGHCG jhQGO dQJ j
GfBG dG ch .kGL H OGe e AMCG J Va RhJ dGh ,Y IFS fc dG
sOGe qGC gOeh ,SdG hdG e JS VJ fc dG ''FdG ddG'' jf KCJ Gj e
.M OGe fCG h aOH b M Z
GG KJ e OYG Gg J EG SdG H J fc edG G H gh
H fc jdG GAYOq G g KEh .G eGU GdG KJh qdG a
M QdH GdG GC J kb QfG Yh , m H Tb b Y G e M Vh e QdG
181

dG dG

. IdG J SdG OYG


jdG OYG Gg EG Sq dG bS a ,dG OhdG KJ gh iNCG IgX g fch
eG HdG SGH L j OhdG CG gh ;H a NBG AT KEG dh ,d kdO JGh
IdG fc ''GfCG UCG'' Hc jhQGO dN dCG dG IdG h .dG Y j dGh Jbd
e a GS N H dG GQdG KCG dh ,M Z OGe e C J fCG jdG Y IFdG
jd fdG AMCG Y gGLCG dG HCG jW Y dPh ,a AL e U Y dG dCJ
CG J G Z OGG CH FdG AYOG UCG d'' :j c KHCG Ff QSH jh ,QSH
58
.''`jdG e IG J
dh ,QSH dEG UJ dG dG dOCG V jW I aj QdG jf Y aGG Xh
Gg e QX dSG dHh ,G cJ H j dG dG KCG edG Y QH dG
. f AJ e G cdG

jdG dG dG hG
IG UCG Ve hJ e hCG jQHhCG QdCG AMCG Y ShdG UNG c d
J G Z OGG CG KE jdG dG qKK IjY kKHCG iLCGh ,jdG dG
kFb IQG j CG G VGh jQdG dH AH KHCG dh ,aOG jW Y M OGe OjEG
59
.'' eH QdG jf J AGOS f t n jo G UCG qEG'' :
KHCG GhLCGh jQHhCG S dG f jdG GhSGh ,QdG IYO e AdG bH SCj h
M 1953 S e fS jeCG FdG b e LCG H TCGh .IG UCG EG Ud
GRdG g Vhh ,dG VG G dG e GRZ GP OGe OLh VaG
cJ NJ dG eCG VMG H Y j CG SGh ,bdH gLh MGh e
. JhdG
OGG C ;a K e YS dh ,eCG EG e IN GdG
q J HdG g soYh
K dG Ggh ,dG VG IOLe fc dG OGG M J HdG eG
60
.MdG GdG cCdH
J HdG AGLEG eSG dG OGG CH j CG f e VG jW U Ia Hh
61
.edG dS G dG LJ fc dG OGG M
182

j hQg

dhJ G gh ,jdG dG GW jhQGdG gGLCG dG QdG c dH AHh


S f e V HS jO S dG G LdG AdG UNG GOH L
: j e G ALh , jhQGdG LdG GP ''VQCG'' U Y 1998
CGH c :gh ;HLEG d e LGf jGH c c dRe h jdG dG Of f ''
62
? IG

Id IG dG
IG UC dG VG Gg CG gG g QJ QdG jf L idG FdG dG
.Y bj H dG G Fd M ,jd e
h VG bh fEG U dG LddG e L e H YCG G FdG N EG
.dG IQG eG cCG M Z OGe H IMGh N fEG
aOG Y OSH MT fCG LQd ,kGL Ic M N jd eRdG hdG EG
dMG :G S Y) d SSCG M g dG Jhd paOJ jJ MG CG Z ,kbWEG
.bGdG VQCG Y ke J 1 / 10950 g eCG M e N d Se JhH jJ
e QJ ed Fg kH j G IGf G eG j dG DNA `dG sEG
e N JU OY ch , e J Y J e fQJ eG a,a
.U
EG fJ DNA dG jdG CG gh dG g jZ iNCG LGhORG kjCG gh
IOLG eG M j fCG g fEG dh ,UG ( fCG) JhdG H
a ,LGhORG J d f bdG ke gOLh e H a ,H KdG WJQ a DNA
G H jh ,JGP e Lh b IG EG :J dG IdG RCe bdG EG ODj
63
.''LQhCG d '' hG jhQGdG
jJ j dG jZ fe dG RNA DNA h jhdG VGh JhdG EG
CG hO gMCG fEG a ,ke Vae aOe jJ MGh ,f Gh f bdG
IG QX dSG gh f G UdG EG kGe fEG j dch ,kGOLe NBG j
. Fc H
g ,d QN H IG H f CG a ke aOG jH IG QX c EG
183

dG dG

.G QfEG SSCG Y jdG Jee c H dG QdG jf J G

QdG jd dG dB G
AYOG Ga '' QdG dBG '' M QhJ fc jhQGO jf f kS fc dG fdG dG
AYOG Mh .G jdG Hb Y FGMG dh kY U d dG fO e CG j
Gg qGE M ,AYOG Gd FSG gCG YCGh ''dG fG'' f M QdG qEa jhQGO
. '' dG fG jW Y GfCG UCG'' SCG dG dG SG e j b e gG
hdG eAeh Ib J dG G FdG AH GC e EG j dG fG e qEG
hdG SCG jCG Ea SG fGG b e jG e b Ou go d :G S a ,dG
dBG g dh .hdG jS AjbCG jjCG e kdCe dG j Ggh ,IG b Y AdG j
fG'' J dG
q Gd .ke N EG d Cc ,NBG EG T e jjCG QJ CG J
'' GfCG UCG '' Hc V H cPh dP j c f jhQGO M h ,Qd Sc ''dG
64
.''G VdH j dG fG Ea HjEG GJ J W'' :j

Qe KC J
Y jhQGO LCGh ? HjEG GdG g S fc c :f j dG GDdGh
TY fa AMCG Y Qe h ,Qe e Y LQ e S e QaCG EG kGOSG GDdG Gg
,MdG LCG EG KQJ e GJ J AMCG CG Yj c GH jhQGO b eh
aGQqdG qE a FYOG Mh ,IjL GfCG QX EG qOGC L H kL GdG g cGJ ch
jhQGO YCGh .jW MCG Y ddG QTCG GQhCG Y d Jh f dG e Cf
Y iJ fc dG HdG e Ob UCG G CG YqOG a ''GfCG UCG'' Hc He eCG
cG dG KGQdG fGb CG EG .65NBGh G H AG EG hdG EG Ie fch FG FdG
fH FdG IQSCG jf EG iOq GC jdG dG G Y Y GC W dG QdGh e
e KCJ hCG IFa GP Z dBG ''dG fG dBG '' X Ggh ,NBG EG L e G dG
.jG dG f Lh

184

j hQg

KGQdG GdGh jG jhQGdG


dG KK kUN gLGh dG jdG G eCG gOL H jhQGqdG b
,'' jG jhQGdG''`H Y e hCG ''jG dG
q jf''`H gSCG IjL jf GbSh jdG
j edG Ggh ,dG fG fL EG QJ KCJ d NBG eY g jdG g Mh
GdG gh ,HG HjEG GdG J h kS J L hCG KGQh GW M
. d KGQdG SG C N f hCG YTEd VdG H eq EG
jdG g YsJh ,jG jhQGdG SG AMCG id QdG Y aGJ dGRe jdG gh
dG hCG PCGh dc cdG IGh AMCG id IOLG G cGdGh AYCG CG
,G cJ GJ hM hCG KGQh GW hM KCH EG J hCG J dEG ... Gh
AMCG Y QV eY J kFGO KGQdG GdG CG gh ;kM kY kqe LGj AYOG Gg dh
.jCG e j IFa GP J h
e FGY jL J CGh ,jd cdG Ie DNA`dG jL Ea kGL VGh dP Sh
jeCG KffGQ.. Y j dG G gh ,S KCG d j CG e H kW c
h ,QdGh FGdGh dH J KGQdG GdG qGE '' :j c G Y UNG
GdG qCG VJ dG edG FG g qEG .GMCG MCG KCJ GP Z Jh kGQOf EG
qeEG j CG d qH G G FGY J qGC qCG kUN ,QdG GMEG kGQhO J CG
EG ODj CG cCG Ma ,gjJ EG ODj dG YS FGY J
q GC ka ,KDe Z hCG kGQV
66
.'' IH qKDe Z j CG hCG H QdG EG
.G Fd G dG EG qOD J KGQh IW OLh dG EG j qfC ;ka M e Ggh
kS L dG GdG g qGC j Ggh ,UG GdG L QV KCG dG gGdGh
ZCG be cJ H ,AMCG Y KCdG jJ Sh jG jhQGdG b e AMCG Qd
h (WdG V HUEG g fEG UG KGQdG Id e aCGh ) MCG
.Qq dG Y d Y Ie dBG QdG KCdG GP KGQdG GdG J CG dc Gh
jhQGO aGYG M a gMd IKDe J CG gQhH a dG fG dBG eq CG
.dG AMCG id QdG Y qGE CG ,''QdG''`H Yj e Lj CG dHh ,f

SdG d KGC : GG S
,QdG jf YqJ dG gjQdG e CG hM Y Y dO aCG GqG S tno
185

dG dG

FdG e e a ,NBG H e H Cf GfCG qG FdG qCG YqJ jdG a


,AMCG e G GfCG X SdG Hh ,edG Qh NBG f EG j CG G e G
j AYOG Gg G kGOSGh .dG e jG e SG YdG dG Gg qE a jdG Mh
. AMCG YdG dG M Ia GW ( dfG) Sh M OLh
jGdG fC ;SCGh MGhdG e ce U Fc OLh j :G S Y
U GP Fc OLh Vj hCG ,MGhR EG jQdH dh AG J Fe b fc
bG g dh ,QW EG d K MGhR fc jGdG fC ;MGhdGh QdG e ce
jh ,e J
q hCG bYEH He hCG N Qb GP J CG Ha Ia TY b VGaG
.'' dfGG TCG'' SG dfG FdG g Y QdG IYO
OGYCH Lh fCG H a ,jQdG G ka TY b '' dG TCG'' g CG VaG dh
jh ,G GG V kGKCG J CG H ch ,GQH H jH QJ Ic GfCGh Ic
b dG G FdG g J CG H a jf U GPEG'' :Hc G g Y jhQGO
V Ge cG g gOLh Y dO MCGh ...VQCG S Y e Ie TY
67
.''jG

jhQGO eGB N
CG Y j dh ,BG Mh Y SdG dG e e kGL Ic Jh jM jLCG
c e Y jG f Y G dG GG KCG bh ,'' dfG TCG '' d KCG
. IQU cCG Yh ICa OLdG EG X b YGfCG AMCG L CG e ;jhQGdG bj
Y jdG UNG jGh jjO gh qjhQGdG IZ MCG G H YG bh
ie Y fcCG AGS ,ec M Fc Y dM g G e qEG'' :kFb GG
KCG qGC Y QdG hO kehO LGh dG gh ,G Gd J Y UCG CG GfCG
OLh CG hO ICa ac AMCG QX d J GG EG CG .68 ''kjQJ IQG bG d
G FdG g c Y J Ggh ,jhQGO dEG gP e Y kW Ggh ,kjf dfG TCd
gh ,kb j CG g e L d j CG hO ICa M Fc Qd MdG dG C ;b
:Ja SZhO e Qe AMCG Y b b G
Lh eq EG AMCa ,AMCG OLh d dG fO GFS GJ hCG ee QdGh G qEG''
186

j hQg

OGLCG hCG GfCG Y gQJ f X fCG CG ,dc J hCG IQU cCG Y dG Lh Y ICa
Yh d M Ib e Ha jdGh dG ec IQHh ICa X b fc EGh ,OLdG S
69
.''G FdG g e OjEG dJ AT H
:CG ,jJh T MCG Yh VQCG Lh Y ICa X b G FdG CG J Ga
. jhQGO j c c QdG dh G g GfCG UCG EG

fE G Qq J IQSGC
OdG Gg h ,fEG UCG TH g QdG jf V Td MhG YVG gCG e qEG
,dG VG TY OdH T M Fc e kGQe Cf G fEG CH jhQGdG YJ
IdG N dfG TCG H OLh jdG YJh ,S je 5-4 b CGH QdG Iah
: gh fEG QJ Y V FQ Y HQCG g G AYOG Gg Mh ,IQcG
Australopithecus SHdGShCG-1
Homo habilis Hg eg-2
Homo erectus SjQEG eg-3
Homo sapiens HS eg-4
g dh ,G Ob hCG '' SHdGShCG '' SG G fEd YCG G Y QdG IYO j
e qc gGLCG dG HCG KCG bh ,G OhdG GfCG e V`e f iS d bG
jdG AY TCG e gch eNGR dS OQdG jdGh OQchCG dQJ QahdG jeCG
70
.fEH ke d bY h OhdG e Ve f iS d G FdG Gg qGC G Ob Y
G MGe ac h ,fEG hCG ''eg'' SG jhQGdG b e Y j G Ob J dG MGh
UG GG VH jhQGdG jh ,G Ob e kGQJ cCG G FdG UCG ''eg''
,G QdG hG Gg e OLh Y kGcCJh jf U Y dc VG GfCG H
dG dG g h .G GfCG g H QJ HGQ CG OLh BG EG j fC ;N :fh
eg EG IG dG qEG'' :kFb je fQBG gh jdG dG QdG jf IYO MCG H YG
71
.''IOe H G e HS
eg (SHdGShCG) G Ob e J U KEG jhGQdG hj S gh
gLh dG acG dh ,j qd kGL j ebCG EG CG HS eg ` SjQEG eg ` Hg
187

dG dG
ceCG GhLh b SjQEG eg h Hg eg h G Ob CG KCG GG AY
Ga M TY b SjQEG eg e GfCG OLh g dP e HCGh .72f edG IdG h
73
. (G fEG) HSeg h dJQff HS eg e L EG kL Lhh kf jM
dG jdG G g eCG h ,NBd kGL gMCG c U Y KCG acG gh
GG Y UNG dc L S gh JYO MCG j QdG jhQGO jf LGh
: j e OQaQg eL
UCG Id M GPe PEG ,f edG G TY b fEH T GfCG KK fc GPEG''
H H fQe AGLEG Y dP e gOCG h ,NBd kGL j H e MCG fCG VGdG ?fEG
74
.''H a jQJ bY jCG EG dN e UdG j Hh
jhdGh kJh keYEG Y cCdGh fEG QJ Y dN b NG EG :IQdG jHh
.Y dO CG EG j Y g fEG NBG fh Ob f G FdG e Ve d
UGe S 15 I G Ob Ge Y KHCG jdG eNGR dS OQdG iLCG bh
H e S jCG OLh Y EG UJ bh ,GG Y UNc dG ce d CG kY
. dG jhQGO fc e ZdG Y dG H YGh fEG Hh OdH dG FdG
QeC bGe Vh H j dG dG hdH UN hL dCH b iNCG L e dh
g jOe dOCG G J dGh dG hdG J eNGR hL Mh ,dG f Y LQN
ha JCJ ` hG aM CG ` kGNCGh YLG da AMCG Y jh AjdGh AdG Y
,SOdG SGh ,WGG OJ Y hG e AG Gg Vhh ,dG f Y LQG aG
INCG IOG g Y kJ eNGR jh .fEG QJ kGNCGh ,FdG dG hCG QdGh
:j c hG
,H Y QSG hCG FdG AMCG Y EG H dG hdG EG jOq G dG e dfG Y ''
AG EG dH kUN h FL AT c qGC ,GG SGH fEG jQJ SG Mh
75
.''MGh BG HQG hCG IOG VdG j CG j fEG M ,QdG jH eDG
UC G Z jhCdH SdG H b e YCG kfEG fEG Qd G dG qEG :PEG
. G GG H

188

j hQg

jOe IY
bJ ieh ,dG gGdGh dOCG e bJ ieh ,QdH UG jdG VSG d
dBG ad QdG KCdG GfG c kjCG VSGh ,dG YGdG e IG UCH G ga
Y Id dfCG TCG OLh J G QKBG jCG OLh GfGh ,jdG g dEG YJ dG QdG
dG YGb e be J dG jdH dG Y dG IQhV EG Uf dG Gd ,jQdG
ce VQCG CG H YOG dGh jQdG Y iNCG jf fG c J CG Hh ,dGh
H gh ,dG GMCG LGh jdG g AH Y kY kGQGUEG g dh .dG
. AdGh dG Y g fCH jd f CG jh eJ iOj
G gh ,dG d Sc eGSGh jdG d G H J j dGh
N jhQGdG Jh YCG kJG OG dH e g H ,jOG SQG e Hf fCH gJ j
. dG OG ga jhd d a Pe
,G Y MdG TCG fd OQjQ j c ,HdG
q H G g J kfMCGh
:Oe Y LQ f jh QdG jf Y SdG aGG e g h ,OQaQg eL j dGh
jOe GJ Lf j dG g EG Ggh ,kS H sn oe ATCH eDfh ,jOH eDf f''
HCG d OhdG YO S g jOH G fEGh ,FOeh dG YGb dh jfdG gGd
a kbWEG U jOG dh ,Y LJ dG ac gGd jOe GJ OjE jQG
76
.''GMCG LGh EG J CG q dEG Gd f CG kGHCG
ka kHGJ HGe fc O jhQGdG H J dG dG ie J dG g EG
Z OGG CH eDj dG Gdh ,IOG a AT CG jdG g UCG IZ jh ,jOG jdH
QdGh aGQdGh ASCGh Qdc G GfCG e jG EG CG ,G OGG OLh S g G
Y GC W dG NGdG d kLf EG d fEG Mh Gh QgdGh QTCGh GGh
.YGdGh YdGh G c IOG
Y aGj GdGRe q jhQGdG qGC EG ,dGh dG YGb e ke VQj OYG Gg CG bGdGh
.'' GMCG LGh EG J CG dEG Gd f CG kGHCG '' aGgC eN FGQBG
WdG G ij CG d H jOe Z f Lh e AMCG UCG b EG j fEG qc h
dN b e npoN :CG ;d M Yh ge Ib KCH npLho b G FdG ac qGE ,dc
. a c :d bh dG e AT c N dG jdG dG dG g dG Ggh ,d

189

e
1. Charles Darwin, The Origin of Species, 6th edition, New York: Macmillan
Publishing Co., 1927, p. 179
2. J.R.P. Angel, Lobster Eyes as X-ray Telescopes, Astrophysical Journal,
1979, 233:364-373, cited in Michael Denton, Natures Destiny, The Free Press,
1998, p. 354
3. Michael F. Land, "Superposition Images Are Formed by Reflection in the
Eyes of Some Oceanic Decapod Crustacea", Nature, 28 October 1976, Volume
263, pages 764-765.
4. Robin J. Wootton, "The Mechanical Design of Insect Wings", Scientific
American, Volume 263, November 1990, p. 120.
5. Pierre Paul Grass, Evolution of Living Organisms, New York, Academic
Press, 1977, p.30
6. "Exploring The Evolution of Vertical Flight at The Speed of Light", Discover,
October 1984, pp. 44-45.
7. Ali Demirsoy, Yasamin Temel Kurallari (Basic Fundamentals of Life), Ankara,
Meteksan AS., Volume II, Section II, 1992, p. 737.
8. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology), Istanbul, Grsel Publications, p. 2676.
9. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology) p. 2679.
10. Smith Atkinson, Insects, London, Research Press, Volume I, 1989, p. 246.
11. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology), p. 2678.
12. Dieter Schweiger, "Die Fliegen", GEO, April 1993, pp. 66-82.
13. Engin Korur, "Gzlerin ve Kanatlarin Sirri" (The Secret of the Eyes and
Wings), Bilim ve Teknik (Journal of Science and Technology), October 1984,
Issue 203, p. 25.
14. Douglas Palmer, "Learning to Fly" (Review of The Origin of and
Evolution of Birds by Alan Feduccia, Yale University Press, 1996), New
Scientist, Vol. 153, March, 1 1997, p. 44
15. A. Feduccia, The Origin and Evolution of Birds, New Haven, CT: Yale
University Press, 1996, p. 130 cited in Jonathan D. Sarfati, Refuting Evolution.
16. Francis Darwin, The Life and Letters of Charles Darwin, Volume II, From
Charles Darwin to Asa Gray, April 3rd, 1860
17. Hakan Durmus, "Bir Tyn Gelismesi" (The Development of a Feather),
Bilim ve Teknik (Journal of Science and Technology), November 1991, p. 34.
18. Hakan Durmus, "Bir Tyn Gelismesi" (The Development of a Feather),
Bilim ve Teknik (Journal of Science and Technology), November 1991, pp. 3435.
19. Michael Denton, Evolution: A Theory in Crisis, London, Burnett Books
Limited, 1985, p. 210-211.
20. Michael Denton, Evolution: A Theory in Crisis, London, Burnett Books
Limited, 1985, p. 211-212.
21. Werner Gitt, The Flight of Migratory Birds, Impact, No. 159
22. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology), page 978.
23. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology), p. 978.
24. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology), p. 978.
25. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology), p. 564-567.
26. J. A. Summer, Maria Torres, Scientific Research about Bats, Boston: National
Academic Press, September 1996, pp. 192-195.
27. Donald Griffin, Animal Engineering, San Francisco, The Rockefeller
University - W.H. Freeman Com., pp. 72-75.
28. Merlin D. Tuttle, "Saving North Americas Beleaguered Bats", National
Geographic, August 1995, p. 40.
29. J. A. Summer, Maria Torres, Scientific Research about Bats, pp. 192-195.
30. For details on this system refer to: W. M. Westby, "Les poissons lectriques se
parlent par dcharges ", Science et Vie, No. 798, March 1984.
31. Charles Darwin, The Origin of Species, The Modern Library, New York, pp.
124-153
32. Michael Behe, Darwin's Black Box, New York: Free Press, 1996, pp. 18-21.
33. Michael Behe, Darwin's Black Box, p. 22.
34. Jean Michael Bader, "Le Gne de LOreille Absolue", Science et Vie, Issue
885, June 1991, pages 50-51.
35. Marshall Cavendish, The Illustrated Encyclopaedia of The Human Body,
London, Marshall Cavendish Books Limited, 1984, pp. 95-97.
36. Fred Bavendam, "Chameleon of The Reef", National Geographic, September
1995, p. 100.

37. Stuart Blackman, "Synchronised Swimming", BBC Wildlife, February 1998,


p.57.
38. Charles Darwin, The Origin of Species, The Modern Library, New York, pp.
124-153
39. Fred Bavendam, "Chameleon of The Reef", National Geographic, page 106.
40. The Guinness Concise, Encyclopaedia, London, Guinness Publishing Ltd.,
1993, p. 125.
41. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology), p. 291.
42. R. Von Bredow, Geo, November 1997.
43. Michael Behe, Darwin's Black Box, pp. 79-97.
44. Michael Behe, Darwin's Black Box, p. 82.
45. T.E. Akiowa & F.C. Schuster, Wars and Technologies, Detroit: Anderson
Bookhouse, 1997, p. 83.
46. Ali Demirsoy, Yasamin Temel Kurallari (Basic Fundamentals of Life), pp. 1822.
47. Marshall Cavendish, The Illustrated Encyclopaedia of The Human Body, pp. 5051.
48. Bilim ve Teknik (Journal of Science and Technology), February 1992.
49. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of Science
and Technology), p. 116.
50. Mark W. Moffett, "Life in a Nutshell", National Geographic, pp. 783-784.
51. Bilim ve Teknik Grsel Bilim ve Teknik Ansiklopedisi (Encyclopedia of
Science and Technology), p. 2995
52. Stanley Taylor, "Life underwater" Botanic, Issue 83, February 1988, p. 24.
53. Michael Behe, Darwin's Black Box, New York: Free Press, 1996, pp. 69-73.
54. Betty Mamane, "Le Surdou du Grand Bleu", Science et Vie Junior, August
1998, pp. 79-84.
55. "If Attacked, Japanese Bees Shake and Bake", National Geographic, April
1996, p. 2.
56. "Poison Dart Frogs Lurid and Lethal", National Geographic, May 1995, pp.
103-110.
57. Reproductive Strategies of Frogs, William E. Duellman, Scientific American,
July 1992, pp. 58-65.
58- Sidney Fox, Klaus Dose, Molecular Evolution and The Origin of Life, New
York: Marcel Dekker, 1977. p. 2
59-- Alexander I. Oparin, Origin of Life, (1936) New York, Dover Publications,
1953 (Reprint), p.196
60- "New Evidence on Evolution of Early Atmosphere and Life", Bulletin of the
American Meteorological Society, vol 63, November 1982, pp.1328-1330.
61- Stanley Miller, Molecular Evolution of Life: Current Status of the Prebiotic
Synthesis of Small Molecules, 1986, p. 7
62- Jeffrey Bada, Earth, ubat 1998, p. 40
63- Leslie E. Orgel, "The Origin of Life on Earth", Scientific American, vol 271,
Oct 1994, s. 78
64- Charles Darwin, The Origin of Species: A Facsimile of the First Edition, Harvard University Press, 1964, p. 189
65- Charles Darwin, The Origin of Species: A Facsimile of the First Edition, Harvard University Press, 1964, p. 184.
66- B. G. Ranganathan, Origins?, Pennsylvania: The Banner Of Truth Trust,
1988.
67- Charles Darwin, The Origin of Species: A Facsimile of the First Edition, Harvard University Press, 1964, p. 179
68- Derek A. Ager, "The Nature of the Fossil Record", Proceedings of the British
Geological Association, vol 87, 1976, p. 133
69- Douglas J. Futuyma, Science on Trial, New York: Pantheon Books, 1983. p.
197
70- Solly Zuckerman, Beyond The Ivory Tower, New York: Toplinger Publications, 1970, pp. 75-94; Charles E. Oxnard, "The Place of Australopithecines in Human Evolution: Grounds for Doubt", Nature, vol 258, p. 389
71- J. Rennie, "Darwin's Current Bulldog: Ernst Mayr", Scientific American,
December 1992
72- Alan Walker, Science, vol. 207, 1980, p. 1103; A. J. Kelso, Physical Antropology, 1st ed., New York: J. B. Lipincott Co., 1970, p. 221; M. D. Leakey, Olduvai
Gorge, vol. 3, Cambridge: Cambridge University Press, 1971, p.272
73- Time, November 1996
74- S. J. Gould, Natural History, vol. 85, 1976, p. 30
75- Solly Zuckerman, Beyond The Ivory Tower, New York: Toplinger Publications, 1970, p.19
76- Richard Lewontin, "The Demon-Haunted World", The New York Review of
Books, Jan 1997, p.28

You might also like