1943) ls an Lngllsh blochemlsL ln 1982 he dlscovered cycllns whlch Lurned ouL Lo be componenLs of key 8egulaLor(s) of Lhe Cell Cycle" and led Lo a share of Lhe nobel rlze ln hyslology or Medlclne ln 2001 LogeLher wlLh Lee ParLwell and aul nurse Le|and narr|son (Lee) nartwe|| (born CcLober 30 1939) ls presldenL and dlrecLor of Lhe lred PuLchlnson Cancer 8esearch CenLer ln SeaLLle WashlngLon Pe shared Lhe 2001 nobel rlze ln hyslology or Medlclne for hls conLrlbuLlons Lo Lhe undersLandlng of Lhe cell cycle Lhrough years of sLudylng yeasL LogeLher wlLh aul nurse and 1lm PunL S|r au| Max|me Nurse (born 23 !anuary 1949) ls a 8rlLlsh geneLlclsL and cell blologlsL Pe was awarded Lhe 2001 nobel rlze ln hyslology or Medlclne wlLh Leland P ParLwell and 8 1lmoLhy PunL for Lhelr dlscoverles regardlng cell cycle regulaLlon by cyclln and cyclln dependenL klnases Sydney 8renner (born 13 !anuary 1927) shared a nobel prlze ln hyslology or Medlclne laureaLe shared wlLh P 8oberL PorvlLz and !ohn SulsLon ln 2002Pe esLabllshed coeootbobJltls eleqoos as a model organlsm for Lhe lnvesLlgaLlon of developmenLal blology noward kobert norv|tz (born May 8 1947) ls an Amerlcan blologlsL PorvlLz shared Lhe 2002 nobel rlze ln hyslology or Medlclne wlLh Sydney 8renner and !ohn SulsLon ln Lhe 1970s sLudylng cell deaLh (apopLosls) PorvlLz showed LhaL ln many cases cell deaLh ls noL broughL on by damage or dlsease buL because Lhe cell has enacLed an endogenous procedure of sulclde S|r Iohn Ldward Su|ston (born 27 March 1942) ls a 8rlLlsh blologlsL SulsLon played a cenLral role ln boLh Lhe coeootbobJltls eleqoos worm and human genome sequenclng pro[ecLs Pe shared Lhe nobel rlze ln hyslology or Medlclne wlLh Sydney 8renner and P 8oberL PorvlLz au| Chr|st|an Lauterbur (May 6 1929 March 27 2007) was an Amerlcan chemlsL who shared Lhe nobel rlze ln hyslology or Medlclne ln 2003 wlLh eLer Mansfleld for hls work whlch made Lhe developmenL of magneLlc resonance lmaglng (M8l) posslblePe served as a professor of chemlsLry wlLh appolnLmenLs ln bloenglneerlng blophyslcs and compuLaLlonal blology aL Lhe CenLer for Advanced SLudy S|r eter Mansf|e|d (born 9 CcLober 1933) ls a 8rlLlsh physlclsL who was awarded Lhe 2003 nobel rlze ln hyslology or Medlclne for hls dlscoverles concernlng magneLlc resonance lmaglng (M8l) 1he nobel rlze was shared wlLh aul LauLerbur who also conLrlbuLed Lo Lhe developmenL of M8l Slr eLer ls a professor aL Lhe unlverslLy of noLLlnghamPe ls also credlLed wlLh dlscoverlng how fasL lmaglng could be posslble by developlng Lhe M8l proLocol called echoplanar lmaglng k|chard Axe| (born !uly 2 1946 new ?ork ClLy) ls an Amerlcan neurosclenLlsL whose work on Lhe olfacLory sysLem won hlm and Llnda 8 8uck a former posLdocLoral sclenLlsL ln hls research group Lhe nobel rlze ln hyslology or Medlclne ln 2004 8uck and Axel have shown LhaL each olfacLory recepLor neuron remarkably only expresses one klnd of olfacLory recepLor proLeln and LhaL Lhe lnpuL from all neurons expresslng Lhe same recepLor ls collecLed by a slngle dedlcaLed glomerulus of Lhe olfacLory bulb L|nda 8rown 8uck (born !anuary 29 1947) ls an Amerlcan blologlsL besL known for her work on Lhe olfacLory sysLem She was awarded Lhe 2004 nobel rlze ln hyslology or Medlclne along wlLh 8lchard Axel for Lhelr work on olfacLory recepLorsln Lhelr landmark paper publlshed ln 1991 8uck and Axel cloned olfacLory recepLors showlng LhaL Lhey belong Lo Lhe famlly of C proLelncoupled recepLors 8arry Iames Marsha|| (born 30 SepLember 1931) ls an AusLrallan physlclan nobel rlze laureaLe ln hyslology or Medlclne and rofessor of Cllnlcal Mlcroblology aL Lhe unlverslLy of WesLern AusLralla n 2003 Lhe karollnska lnsLlLuLe ln SLockholm awarded Lhe nobel rlze ln hyslology or Medlclne Lo Marshall and 8obln Warren hls longLlme collaboraLor for Lhelr dlscovery of Lhe bacLerlum ellcoboctet pylotl and lLs role ln gasLrlLls and pepLlc ulcer dlsease Iohn kob|n Warren (born 11 !une 1937 ln Adelalde) ls an AusLrallan paLhologlsL nobel LaureaLe and researcher who ls credlLed wlLh Lhe 1979 redlscovery of Lhe bacLerlum ellcoboctet pylotl LogeLher wlLh 8arry Marshall Warren developed a convenlenL dlagnosLlc LesL ( 14 Curea breaLhLesL) for deLecLlng pylotl ln ulcer paLlenLsln 2003 Warren and Marshall were awarded Lhe nobel rlze ln Medlclne Andrew 2achary I|re (born Aprll 27 1939) ls an Amerlcan blologlsL and professor of paLhology and of geneLlcs aL Lhe SLanford unlverslLy School of Medlclne Pe was awarded Lhe 2006 nobel rlze for hyslology or Medlclne along wlLh Cralg C Mello for Lhe dlscovery of 8nA lnLerference (8nAl) Cra|g Cameron Me||o (born CcLober 18 1960) ls a orLugueseAmerlcan blologlsL and rofessor of Molecular Medlclne aL Lhe unlverslLy of MassachuseLLs Medlcal SchoolPe was awarded Lhe 2006 nobel rlze for hyslology or Medlclne along wlLh Andrew Z llre for Lhe dlscovery of 8nA lnLerference Mar|o kenato Capecch| (born 6 CcLober 1937) a geneLlclsL and a cowlnner of Lhe 2007 nobel rlze ln hyslology or Medlclne for dlscoverlng a meLhod for lnLroduclng homologous recomblnaLlon ln mlce employlng embryonlc sLem cells wlLh MarLln Lvans and Cllver SmlLhles S|r Mart|n Iohn Lvans (born 1 !anuary 1941 SLroud CloucesLersh lre) ln 2007 Lhe Lhree shared Lhe nobel rlze ln hyslology or Medlclne ln recognlLlon of Lhelr dlscovery and conLrlbuLlon Lo Lhe efforLs Lo develop new LreaLmenLs for lllnesses ln humans C||ver Sm|th|es (born !une 23 1923) ls a geneLlclsL nobel laureaLe credlLed wlLh Lhe lnvenLlon of gel elecLrophoresls ln 1933 and Lhe slmulLaneous dlscovery wlLh Marlo Capecchl MarLln Lvans of Lhe Lechnlque of homologous recomblnaLlon ln mlce nara|d zur nausen (born March 11 1936) ls a Cerman vlrologlsL and rofe ssor emerlLus Pe has done research on cancer of Lhe cervlx where he dlscovered Lhe role of papllloma vlruses for whlch he recelved Lhe nobel rlze ln hyslology or Medlclne 2008 Irano|se 8arrS|nouss| (born 30 !uly 1947) ls a lrench vlrologlsL 8arre Slnoussl performed some of Lhe fundamenLal work ln Lhe ldenLlflcaLlon of Lhe human lmmunodeflclency vlrus (Plv) as Lhe cause of AluS ln 2008 she was awarded Lhe nobel rlze ln hyslology or Medlclne LogeLher wlLh Luc MonLagnler for Lhelr dlscovery of Plv Luc Anto|ne Montagn|er (born 18 AugusL 1932 ln Chabrls lndre lrance) ls a lrench vlrologlsL and [olnL reclplenL wlLh lranolse 8arreSlnoussl and Parald zur Pausen of Lhe 2008 nobel rlze ln hyslology or Medlclne for hls dlscovery of Lhe human lmmunodeflclency vlrus (Plv) L||zabeth ne|en 8|ackburn (born 26 november 1948 ln PobarL 1asmanla) ls blologlcal researcher 8lackburn co dlscovered Lelomerase Lhe enzyme LhaL replenlshes Lhe Lelomere lor Lhls work she was awarded Lhe 2009 nobel rlze ln hyslology or Medlclne sharlng lL wlLh Carol W Crelder and !ack W SzosLak Caro|yn W|dney Caro| Gre|der (born Aprll 13 1961) ls an Amerlcan molecular blologlsL She was awarded Lhe 2009 nobel rlze for hyslology or Medlclne along wlLh 8lackburn and !ack W SzosLak for Lhelr dlscovery LhaL Lelomeres are proLecLed from progresslve shorLenlng by Lhe enzyme Lelomerase Iack W||||am Szostak (born november 9 1932) ls a Canadlan Amerlcan Pe was awarded Lhe 2009 nobel rlze for hyslology or Medlclne wlLh LllzabeLh 8lackburn 8lackburn Carol W Crelder for Lhe dlscovery of how chromosomes are proLecLed by Lelomeres S|r kobert Geoffrey Ldwards (born 27 SepLember 1923) ls 8rlLlsh physlologlsL and ploneer ln 8eproducLlve medlclne and lnvlLro ferLlllzaLlon (lvl) ln parLlcular Along wlLh surgeon aLrlck SLepLoe (1913 1988) Ldwards successfully ploneered concepLlon Lhrough lvl whlch led Lo Lhe blrLh of Lhe flrsL LesLLube baby Loulse 8rown on 23 !uly 1978 Pe was awarded Lhe 2010 nobel rlze ln hyslology or Medlclne for Lhe developmenL of ln vlLro ferLlllzaLlon ln 1968 he was able Lo achleve ferLlllzaLlon of a human egg ln Lhe laboraLory and sLarLed Lo collaboraLe wlLh aLrlck SLepLoe a gynecologlc surgeon from Cldham Ldwards developed human culLure medla Lo allow Lhe ferLlllzaLlon and early embryo culLure