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The scientific procedure of arranging organisms into groups and subgroups on the basis of their similarities and dissimilarities and placing them in a
MAP
hierarchy of categories is called biological classification. The earliest classification systems recognised only two kingdoms of living things: Animalia and
Plantae followed by three and four kingdom classifications introducing Kingdom Monera and Protista. The most accepted and latest five-kingdom
classification was proposed by R.H. Whittaker in 1969 to develop phylogenetic relationships. In this classification, the organisms are classified on the basis
of following criteria : (i) complexity of cell, (ii) complexity of the body organisation, (iii) mode of nutrition, (iv) mode of reproduction, (v) ecological role and
(vi) phylogenetic relationships.
Classification Criteria
Major
Producer Decomposer Consumer
Ecological Role
Mode
Autotrophy (Photosynthesis) Heterotrophy Heterotrophy (Ingestion)
of Nutrition
(Absorption)
KINGDOM FUNGI
R. K. MALIK'S NEWTON CLASSES, RANCHI
Angiospermae Echinodermata
Phaeophyta Ascomycota (Echinoderms)
Tracheophyta
Prokaryotes
Cyanobacteria
(Blue green algae)
Eubacteria
(True bacteria) Archaebacteria
Viruses (Ancient bacteria)
Ÿ Viruses are intermediate between living and non-
living entities. These act as obligate parasites, i.e., KINGDOM MONERA
inert outside the host cell. l It includes basically unicelluar, microscopic
Ÿ They do not grow, divide or reproduce like typical living prokaryotes and the most primitive of living forms
organisms, rather reproduce using host machinery. with different modes of nutrition i.e., – saprobic, parasitic,
Ÿ Viruses are thus described as a nucleoprotein entity chemoautotrophic, photoautotrophic and symbiotic.
which is able to utilise the synthetic machinery of a l Cell wall comprises of peptidoglycan and polysaccharides other
living cell of another organism for its multiplication than cellulose. Cells have one envelope type of organisation but
and division. internal compartmentalisation is absent.
Ÿ They resemble living organisms in possessing genetic l If present, flagella are single stranded, genetic material
material and organic macromolecules and showing is not organised into a nucleus, DNA is naked i.e.,
mutations. without histone, all membrane bound cell
Ÿ They are responsible for a number of infectious organelles are absent. Sexual
diseases in plants, animals and humans. reproduction is absent as meiosis
does not occur.
R. K. MALIK'S NEWTON CLASSES, RANCHI
CONCEPT KINGDOM FUNGI
Fungi is a large kingdom comprising of about 5100 genera and more than 50,000 species. l
CLASSIFICATION
Many botanists have classified fungi in different ways.
MAP They are achlorophyllous, heterotrophic, spore forming, eukaryotic organisms with thalloid
body made up of hyphae (together constituting mycelium). They are cosmopolitan in
distribution. Some fungi occur in fresh or marine water, others are terrestrial and still others
are air borne. The study of fungi is known as mycology.
l Martin’s (1961) classification of fungi is most prevalent. He classified fungi into Myxomycotina
(Slime molds) and Eumycotina (True fungi).
Martin further divided Eumycotina into the following classes:
STRUCTURE
l
l
The mycelium is aseptate and coenocytic.
Phycomycetes
The sporangia has innumerable sporangiospores (zoospores or
aplanospores) formed endogenously.
l Fungi range from unicellular, uninucleate forms like yeast l Sexual reproduction is oogamous in Oomycetes, and isogamous in
and Synchytrium to thread-like structure called mycelium Zygomycetes.
NUTRITION REPRODUCTION which is made up of a net like mass of tubular filaments l Biflagellate motile cells (zoospores) are produced by many species.
l Fungi may reproduce by vegetative, asexual and sexual means. called hyphae. The hypha is usually branched, tube like l The zygote is unicellular and simple.
l They may be obligate parasites (obtain food from host plants
structure, having protoplasm with reserve food and l E.g., Albugo, Phytophthora (Oomycetes), Rhizopus, Mucor (Zygomycetes).
and die with the death of host) or facultative saprophytes
(usually parasitic but able to absorb food from decaying host Vegetative bounded by a wall of chitin, a nitrogen containing
plant as well), obligate saprophytes (obtain food from
polysaccharide (C22H54N4O21)n. Ascomycetes
l The protoplasm of the hypha may be continuous without
decaying organic matter) or facultative parasites (usually Budding Fission Fragmentation l The mycelium consists of septate hyphae, possessing central or septal
Small outgrowths from Splitting of Fragments of cross walls, called aseptate hypha or may have
saprophytes but can live parasitically under some conditions). pores. Motile structures do not occur in the life cycle.
vegetative body, cut off vegetative vegetative hyphae transverse partitions or septa, known as septate hypha.
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Zoospores l Sexual reproduction takes place through fusion of sex cells, somatic cells,
central septal pore possesses a barrel-shaped inflation, as
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Uniflagellate or biflagellate, thin walled, uninu- gametangial contact between an antheridium and ascogonium and
in many basidiomycetes, it is known as dolipore septum.
cleate structures formed in zoosporangia, e.g., Sexual l A membranous vesicle called lomasome is found
autogamy.
To MAKE YOUR CONCEPTS CRYSTAL CLEAR
Phytophthora, Albugo. Sexual reproduction takes place by following processes: l Karyogamy is delayed after plasmogamy. Hence, a new transitional phase
attached to plasma membrane.
called dikaryophase appears in the life cycle. The cells of dikaryophase are
Sporangium Spores
Fertilisation Gametangial contact Lipid globule called dikaryotic cells as each cell possesses two nuclei (n + n).
Oogonium Empty
Sporangiospores tube Here gametes are never released Vacuole l Some dikaryotic cells function as ascus mother cells. Ascus is a sporangial
(Female antheridium
Nonflagellate spores that develop inside gameta- (Male from gametangia, instead the male Dictyosome (Unicisternal) sac peculiar to Ascomycetes. 4-8 haploid meiospores named ascospores are
sporangia, e.g., Mucor, Rhizopus. ngium) gameta- and female gametangia come in Glycogen particle or oil (Reserve food) produced internally in each ascus.
ngium) Nucleus
close contact with the help of a l The asci may occur freely or get aggregated with dikaryotic mycelium to
Endoplasmic reticulum
fertilisation tube, through which one Ribosome form fructifications called ascocarps.
Chlamydospores l E.g., Yeast, Aspergillus, Penicillium, Claviceps, morels and truffles.
Ooplasm or more male nuclei migrate to the Mitochondrion
Thick-walled perennating spores which develop at Periplasm Cytoplasmic matrix
female gametangium. E.g., Pythium
places along the hyphae by accumulation of
protoplasm, rounding off and secretion of thick wall.
Planogametic copulation Antherozoid
Plasma membrane Basidiomycetes
Anisogametes
This involves fusion of two l Basidiomycetes are the most advanced fungi and considered among the
Oidia Isogametes Ovum Hyphal wall
naked motile gametes best decomposers of wood.
Oidia (multiply (Usually contains chitin)
(planogametes). Based l Motile structures or cells are absent. Mycelia are of two types, primary and
Usually formed under conditions of excess by
budding) upon the nature and Fig.: Ultrastructure of part of fungal hypha secondary.
water, sugar and certain salts, e.g., Rhizopus.
structure of gametes, it is l Karyogamy is delayed after plasmogamy. A new transitional phase called
of three types: isogamy, Empty l In some fungi, hyphae may structurally modify in response
Flagella antheridium dikaryophase appears in the life cycle. It produces dikaryotic secondary
Conidia anisogamy and oogamy. to functional needs as:
Conidia Isogamy Anisogamy Oogamy mycelium. Secondary mycelium is long lived, profusely branched septate
(a) Prosenchyma : It is formed when the component
Nonmotile, thin-walled, exogenous spores, hyphae possessing dolipores.
Isogametangia Gametangial copulation hyphae lie more or less parallel to one another and
produced in chains upon the tip of hypha called l Hook-shaped outgrowths called clamp connections are found on the
Conidiophore This process involves fusion of the unite to form a rather loosely interwoven structure
conidiophore, e.g., Aspergillus, Penicillium. sides of septa which are meant for proper distribution of dikaryons at the
entire contents of two compatible where their individuality is not lost.
Ascopores (b) Pseudo-parenchyma : It is formed when the hyphae time of cell division.
gametangia, resulting in karyo-
Ascospores Suspensor become closely interwined, forming a tissue which l Karyogamy and meiosis occur in club-shaped structures known as basidia.
gamy. E.g., Mucor.
Nonmotile meiospores which are produced consists of hollow tubes spread in all directions. These A basidium commonly produces four meiospores or basidiospores
Ascus Somatogamy
inside special sacs called asci and are lose their individuality. exogenously at the tip of fine outgrowths called sterigmata.
characteristic of Class Ascomycetes. Here sex organs are not at all formed, but two vegetative hyphae l The fungi may or may not produce fructifications called basidiocarps
or cells take over the sexual function and fuse together. E.g., (c) Rhizomorph : It is a thick strand or root-like
aggregation of somatic hyphae which lose their that vary in size from microscopic to macroscopic forms.
Morchella, Peziza. l E.g., Puccinia, Ustilago, Agaricus, bracket fungi, etc.
Basidiospores individuality. The entire mass behaves as an organised
R. K. MALIK’S
Sterigma
Nonmotile meiospores formed exogenously Spermatisation unit and have higher infection capacity.
Deuteromycetes
NEWTON CLASSES
on short outgrowths of club-shaped structure In some advanced genera, the sexual process is accomplished by (d) Sclerotium : It is a compact globose structure, formed
Basidium
called basidium and are characteristic of Class minute spore-like spermatia (male gametes) and specialised by the aggregation and adhesion of hyphae. l Deuteromycetes is an artificial class of fungi which has been created to
Basidiomycetes. receptive hyphae (female gametes). The spermatia are carried by (e) Appressorium : It is a terminal, simple or lobed,
include all those fungi in which sexual stage is either absent or not known.
air, water or insects to the receptive hyphae. The contents of the swollen structure of germ tubes or infecting hyphae, l Some of the deuteromycetes are unicellular like yeast.
Binucleate spores spermatium enter the receptive hyphae through a pore. found in many parasitic fungi. l The mycelium is usually septate. Coenocytic forms are not known.
Dikaryotic spores meant for multiplying Receptive hyphae (f ) Haustorium : These are intracellular, absorbing l Asexual reproduction often occurs by conidia along with some other
the dikaryotic mycelium, e.g., aecidio- (Oogonium) structures of obligate parasites meant for absorbing types of spores.
spores, uredospores in Puccinia. Another Trichogyne food material from the host. They may be variously l It is believed that most members of deuteromycetes are actually ascomycetes
type of dikaryotic spore is teleutospore
Uredo- Teleut- shaped and secrete specific hydrolysing enzymes. in which sexual reproduction is either absent or yet to be discovered.
or teliospore . Aecidio- spore ospore Spermatium l E.g., Colletotrichum, Helminthosporium, Trichoderma.
spore
R. K. MALIK'S NEWTON CLASSES, RANCHI
R. K. MALIK'S NEWTON CLASSES, RANCHI
Reproduction
CONCEPT LICHENS Lichens reproduce both by asexual and sexual means.
Asexual reproduction Sexual reproduction
MAP Lichens are composite or dual organisms representing an intimate symbiotic relationship
between algae and fungi (De Bary, 1879). The algal component of lichen is known as
phycobiont whereas the fungal component is known as mycobiont. The fungal component
predominates algal component, such an association is known as helotism (Crombic, 1885).
There are about 400 genera and 1600 species of lichens. They usually grow on bark of trees,
CONCEPT MORPHOLOGY
perform photosynthesis. It is a green lateral flattened outgrowth borne
on the node of a stem or stem branch and bears a bud in its axil.
MAP OF LEAF
Parts of
a leaf
A typical leaf consists of three parts - leaf base,
petiole and lamina. Leaf base is the basal part of the
leaf by which it is attached to the node of the stem.
Different plants have different types of leaf bases viz.
pulvinus, e.g., pea; sheathing, e.g., Zea mays; decurrent, e.g.,
Crotolaria and amplexicaul, e.g., Polygonum. Leaves of some
plants have lateral appendages on each side of leaf base, known
as stipules which may be caducous, deciduous or persistent.
Petiole is the leaf stalk that joins the lamina to the stem or its
Opposite
superposed branch. Sometimes the petiole is absent and then the leaf is
Whorled Phyllotaxy said to be sessile.
Phyllotaxy is the arrangement of Lamina is the expanded, green and conspicuous part of
R. K. MALIK'S NEWTON CLASSES, RANCHI
Stem branch
Leaf
tendril Fig.: A typical
dicotyledonous leaf
es
leav Leaf modifications
ple
Sim
Fig.: (i) Leaf tendrils : Leaves are modified into
Simple and slender, wiry often closely, coiled structures, known as
tendrils, which help in climbing. These may be whole leaf
Compound leaves tendrils (e.g., Lathyrus aphaca), leaflet tendrils (e.g., Pisum
A simple leaf is the one in which lamina is undivided or sativum), petiolar tendrils (e.g., Nepenthes), leaf tip tendrils (e.g.,
incised to any depth but not upto the midrib. In a compound leaf, Gloriosa), Stipular tendrils (e.g., Smilax), etc.
lamina is completely broken up into distinct segments called leaflets (ii) Leaf spines : These protect the plants from grazing animals and
which are separately articulated at the base. Compound leaves are of 2 excessive transpiration e.g., Solanum surattense.
types - (iii) Leaflet hooks : The terminal leaflets of compound leaves become
(a) Pinnate compound leaves : In these type of leaves, incision of lamina is transformed into stiff claw-like and curved hooks. These help the plant in
directed towards the midrib which is known as rachis. Leaflets are arranged on climbing, e.g., Doxantha unguis-cati.
both sides on the rachis or on its branches. These are of following types: (iv) Phyllodes : These are the flattened petioles or parts of the rachis which
(i) Unipinnate : Leaflets are directly attached on the rachis, e.g., Cassia fistula perform the function of photosynthesis, e.g., Acacia species. These help to
(paripinnate), rose (imparipinnate). reduce transpiration in xerophytic plants.
(ii) Bipinnate : Rachis divides and gives rise to secondary axis on both sides on which (v) Insect catching leaves : Leaves are modified to form pitchers (e.g.,
Nepenthes), bladders (e.g., Utricularia) etc. to trap and digest insects.
leaflets are arranged, e.g., Acacia.
(vi) Succulent leaves : These are fleshy leaves that store food material,
(iii) Tripinnate : Secondary axis too, divides and gives rise to tertiary axis on which
e.g., Aloe, Agave etc.
leaflets are attached, e.g., Moringa.
(vii) Scale leaves (or cataphylls) : These are dry, membranous
(iv) Decompound : Rachis divides more than three times and gives rise to small leaves which do not take part in photosynthesis, e.g.,
axis on which leaflets are arranged, e.g., carrot. Casuarina.
(b) Palmate compound leaves : In these type of leaves, incision of leaf is (viii) Floral leaves : These are specialised
directed towards the petiole due to which all leaflets seem to be leaves i.e., sepals, petals, stamens and
articulated on the upper end of petiole. It does not have any rachis. carpels.
Depending on the number of leaflets present, a palmate
compound leaf is called unifoliate (e.g., Citrus), bifoliate
( e.g., Balanites ), trifoliate ( e.g., Trifolium ),
quadrifoliate ( e.g., Paris quadrifoliata ), Leaf
multifoliate (e.g., Bombax). spine
Leaf
pitcher
Venation
Venation is the arrangement of veins
and veinlets on the lamina of a leaf.
Venation is of 3 main types - reticulate (veins
form a network), parallel (veins run parallel) and
furcate (veins branch dichotomously, e.g., Circeaster).
Pinnate Palmate
Reticulate venation is found in most dicots. Pinnate
(or unicostate) reticulate venation occurs in Ficus
Fig.: Compound leaves religiosa. Palmate (or multicostate) reticulate venation
occurs in Zizyphus (convergent), and Luffa (divergent). Parallel
Parallel venation occurs in most monocots. Pinnate
Functions of (or unicostate) parallel venation occurs in banana.
the leaves Palmate (or multicostate) parallel venation
Primary functions: Photosynthesis, occurs in bamboo (convergent) and
gaseous exchange, transpiration, protection Livistonia (divergent).
of buds and conduction through veins.
Secondary functions : Storage e.g., succulent
leaves of Aloe, Agave etc.; protection e.g., spiny Reticulate
leaves of Barberry, Opuntia etc.; support e.g.,
leaflet hooks in Doxantha; nitrogen nutrition
e.g., leaf pitchers of Nepenthes; reproduction
e.g., leaves of Bryophyllum help in
vegetative propagation; floral
leaves help in sexual Fig.: Types of venation
reproduction.
R. K. MALIK’S
NEWTON CLASSES
Inflorescence is the arrangement and distribution of flowers on the
CONCEPT INFLORESCENCE
shoot system of a plant. The axis of the inflorescence is called
peduncle, whereas the stalk of individual flower is called pedicel. A
flattened peduncle is known as receptacle. Inflorescence is of five
Compound
racemose
Compound racemose inflorescence is an
indefinite or indeterminate inflorescence in
which the peduncle is branched repeatedly once
Solitary or twice in a racemose fashion. It is of following
terminal types–
Single flower occurs on Solitary (a) Compound raceme or panicle, e.g., goldmohur,
the terminal part of a Flowers occur singly or are Cassia fistula, Yucca.
branch, e.g., poppy. separated from other flowers (b) Compound spike or spike of spikelets, e.g., wheat.
of the same plant by (c) Compound spadix, e.g., coconut, date, banana.
vegetative regions. (d) Compound corymb, e.g., Pyrus, cauliflower.
(e) Compound umbel, e.g., Daucus carota,
fennel, Coriandrum sativum.
Solitary (f ) Compound capitulum, e.g.,
R. K. MALIK'S NEWTON CLASSES, RANCHI
Racemose
It is an indeterminate inflorescence
which shows indefinite growth. The Catkin
arrangement of flowers is either
Dichasial
acropetal (vertical orientation of
cyme Polychasial cyme axis) or centripetal
(horizontal orientation
Cymose of axis).
A determinate inflore - Spike
scence in which the tip of main Raceme
axis terminates in a flower and further Umbel
Helicoid growth continues by one or more lateral
cyme Cymose branches. The arrange-ment of flowers is
head
either basipetal (vertical orientation Simple
of axis) or centrifugal Raceme
(horizontal orientation of
racemose Peduncle is elongated having
Simple racemose inflore- pedicellate flowers in an acropetal
axis).
Cymose scence is an indefinite fashion, e.g., Lupinus, Raphanus,
inflorescence in which Linaria.
head the peduncle is
Sessile or subsessile flowers unbranched.
are borne centrifugally Spike
around a receptacle, e.g., An elongated peduncle bears sessile
Albizzia, Anthocephalus Scapigerous flowers in an acropetal fashion, e.g.,
Umbel
cadamba, Acacia. Head All the pedicellate flowers arise from a single Achyranthes, Callistemon, Adhatoda
The leafless flowering axis point in a centripetal fashion. The peduncle is vasica.
known as scape bears clusters very much reduced, e.g., Hydrocotyle, Prunus.
of flowers that form a head
which is covered by spaths, Spikelet
Biparous or e.g., Allium cepa. Corymb Spikelets are small and few flowered
Dichasial cyme The main axis is comparatively short, and the spikes which are surrounded at the
A terminal flower is subtended lower flowers have much longer pedicels than base by two scales or glumes, e.g.,
by two lateral branches which the upper ones so that all the flowers are brought rice, bamboo, oat, etc.
also end in flowers. The process is more or less to the same level, e.g., Iberis amara .
repeated. Inflorescence axis is Catkin
multipodial, e.g., Spergula, Multiparous Corymbose raceme Pendulous spike which bears naked
Stellaria media, The young flowers appear to be arranged like a
Clerodendrum.
or Polychasial cyme pistillate or staminate flowers, (but
More than two lateral branches corymb but in mature state the longer pedicels not both) e.g., mulberry, poplar, Salix,
continue the growth of of the lower flowers do not bring them to the Quercus.
inflorescence when the parent level of upper ones, e.g., mustard.
axis ends in a flower, e.g.,
Spadix
With flattened
Asclepias. The flattened receptacle bears numerous sessile Spike with fleshy peduncle and
Monochasial cyme and small florets (ray florets and disc florets) in a having both male and female flowers.
A single lateral branch arises from the centripetal manner, e.g., Zinnia, Sunflower, Cosmos. It is surrounded by a large green or
peduncle of old flower which terminates coloured bract called spathe, e.g.,
in a flower. The lateral branch also palm, Colocasia, Musa.
terminates in a flower. It is of two types:
(a) Helicoid cyme – All the flowers are borne Cyathium
on the same side forming a sort of helix, e.g., The inflorescence looks like a
Drosera, Begonia, Myosotis. flower. The bracts or the involucre
(b) Scorpioid cyme - Flowers are become fused to form a cup shaped
Hypanthodium
alternately borne on both the sides, structure. The inflorescence contains It has a flask-shaped fleshy
e.g., Tecoma, Ranunculus, pedicellate, achlamydeous, unisexual receptacle which possesses a
Heliotropium. flowers of both the types, male and narrow apical opening guarded by
female. The cup encloses a single hairy structure. The receptacle bears
female flower surrounded by a male flowers towards the pore and
large number of male flowers. female flowers towards the
E.g., Euphorbia Hypanthodium base. E.g., Ficus religiosa,
Mixed pulcherrima. Ficus carica.
Two or more t ypes of
inflorescences get mixed up to
form a mixed inflorescences. It is of
following types:
(a) Panicle of spikelets, e.g., oat, rice. Cyathium Verticillaster
(b) Corymb of capitula, e.g., Ageratum Verticillaster
conyzoides. Two dichasial cyme inflore-
(c) U m b e l o f c a p i t u l a , scences develop from axil of
raceme of capitula. opposite leaves. They together
(d) Thyrsus, e.g., Special form a false whorl around the
grapevine. node, e.g., Ocimum, Leucus.
A true fruit (or eucarp) is a ripened ovary. It consists of a thin or thick pericarp
MORPHOLOGY formed from the wall of ovary and seeds formed from the ovules. A fruit in which
other floral parts (e.g., thalamus, base of sepals, petals, etc.), participate in its
CONCEPT OF FRUITS
formation is called false fruit (or pseudocarp) e.g., apple and pear. The seeds
within the fruits have reserve food for nourishing the young seedlings till they
MAP
become nutritionally independent.
· Some fruits are formed without fertilisation i.e., seedless fruits. They are called as
AND SEEDS parthenocarps (e.g., banana). Fruits are classified into three main categories –
simple fruits, aggregate fruits and composite fruits.
Testa Tegmen
Perisperm Cotyledon
Endosperm
Testa
Caruncle
Inferior Berry Embryo
Pome of Apple Berry of Tomato Drupe of Mango Radicle Cotyledon Axis
of Banana
Castor seed (L.S.) Gram seed (split open)
Achenial fruits (Indehiscent fruit) Capsular fruits (Dehiscent fruit) Schizocarpic fruits (Splitting fruits)
Achenial fruits are of five types : Capsular fruits are of five types: Schizocarpic fruits are of five types :
(i) Achene : It develops from monocarpellary, superior, (i) Legume (or pod) : The fruit develops from superior, (i) Cremocarp : It develops from a bicarpellary,
unilocular and uniovuled ovary, pericarp is free from seed monocarpellary, unilocular ovary with marginal placentation. It syncarpous, bilocular, inferior ovary. On maturity, the
except at one point, e.g., Mirabilis jalapa. (ii) Caryopsis (or dehisces by both dorsal and ventral sutures, e.g., members of family fruit splits into two mericarps, each with one seed,
Grain) : It develops from monocarpellary, superior, unilocular Leguminosae. (ii) Follicle : The fruit dehisces by only one suture, e.g., members of family Apiaceae. (ii) Lomentum : The
ovary. Pericarp is completely fused with the testa, e.g., members e.g., Delphinium. (iii) Siliqua : It develops from a bicarpellary, fruit is a modification of legume, which is constricted
of family Poaceae. (iii) Cypsela : It develops from bicarpellary, superior ovary with parietal placentation and a false septum called in between the seeds, e.g., Mimosa, Acacia, etc.
syncarpous, inferior and unilocular ovary. Pappus may be replum. It dehisces by two valves, e.g., members of family (iii) Carcerulus : At maturity, the fruit breaks up into
present for dispersal, e.g., Taraxacum. (iv) Nut: Pericarp becomes Brassicaceae. (iv) Silicula : It is a shortened and flattened siliqua, single seeded indehiscent mericarps, e.g., Althaea.
h a r d , w o o d y o r l e a t h e r y. Fr u i t m a y d e v e l o p f r o m e.g., Capsella bursa-pastoris. (v) Capsule: According to the mode of (iv) Compound samara : At maturity, the fruit splits
monocarpellary, superior ovary (e.g., cashew nut); tricarpellary, dehiscence, capsule may be porocidal capsule (e.g., Papaver), up into single seeded winged mericarps, e.g., Acer.
syncarpous, trilocular ovary (e.g., litchi), etc. (v) Samara : denticidal capsule (e.g., Pink), pyxidium (e.g., Portulaca), (v) Regma : It develops from multicarpellary pistil and
Pericarp becomes flat like wings and thus help in wind dispersal loculicidal capsule (e.g., Gossypium), septicidal capsule on maturity, splits into as many cocci as the number of
e.g., Holoptelea. (e.g., Viola), septifragal capsule (e.g., Datura), etc. carpels, e.g., Geranium.
Cremocarp of Coriandrum
Follicle of
Capsule of Delphinium Carcerulus of Lomentaceous pod Regma of
Datura Althaea of Touch-me-not Castor
Caryopsis of
Maize
Nut of Litchi
Siliqua of Silicula of
Achene of Mirabilis Mustard Capsella Double samara of Acer
jalapa
CONCEPT
A flower is a highly condensed and modified shoot. It contains reproductive organs of the flowering plants, which develop fruits and seeds. There are four types
of floral organs viz. sepals, petals, stamens and carpel or pistil. A flower having all the four types of floral organs is known as complete flower e.g., cotton. If
one or more of the floral organs are absent it is called incomplete flower e.g., cucurbits. A flower having both the essential organs i.e., stamens and carpels is
FLORAL called hermaphrodite or bisexual flower, e.g., China rose, whereas a flower having only one of the two essential organs is known as unisexual flower e.g., mulberry.
Flowers having only stamens are called staminate flowers and those having only carpels are called pistillate flowers. On the basis of symmetry flower can be
MAP MORPHOLOGY
actinomorphic (two equal halves in any plane), zygomorphic (two equal halves in one plane), or asymmetrical.
ANDROECIUM
l Third and male whorl of the flower made up of
COROLLA
l Second whorl inner to calyx made up of petals
which protects the inner whorls and attracts insects
stamens (microsporophylls) which consist of for pollination. Corolla may be polypetalous (petals free)
filament (lower stalk-like part which may be absent in or gamopetalous (petals fused).
some), anther (upper swollen part usually having two lobes or l Polypetalous corolla may be cruciform (four clawed or
theca i.e., bithecous or in some monothecous, e.g., Althaea) and unguiculate petals arranged cross wise e.g., mustard), caryophy-
GYNOECIUM connective (sterile band which connects two anther lobes). In Salvia,
l Central, female reproductive part l The stamens may be equal llaceous (five unguiculate or clawed petals with limbs placed at right
or unequal in length. When there is connective forms a long curved structure, its one end has a fertile anther angles, e.g., Dianthus), rosaceous (five or more sessile or shortly clawed
which develops from thalamus and lobe and other has a sterile anther lobe. Stamens may be shorter than other
consists of carpels (megasporophylls). two long and two short stamens, the petals with limbs spread regularly outwards, e.g., rose), campanulate or bell-
condition is called didynamous, e.g., whorls (inserted) or may protrude out of the flower (exserted). shaped (e.g., Physalis), papilionaceous (five unequal or irregular petals viz.
l Each carpel consist of– stigma (the tip which
Ocimum. When out of six stamens, four are longer l On the basis of attachment to the filament, anthers can be adnate (filament runs 1 standard or vexillum – posterior largest–, 2 wings or alae –lateral, smaller– and
receives pollen), style (elongated structure along the back of the anther or becomes continuous with the connective, e.g., 2 anterior petals fused together to form keel or carina).
connecting stigma and ovary), ovary (lower swollen in inner whorls and two in outer whorl are shorter,
the condition is called tetradynamous, e.g., mustard. Ranunculus), basifixed (filament fixed at anther its base, e.g., Datura), dorsifixed l Gamopetalous corolla may be campanulate (bell-shaped or inverted cup
part containing ovules). (filament attached to back and anther immobile, e.g., Passiflora), versatile (filament
l Dehiscence of anthers to expose the pollen grains shaped e.g., Campanula), urceolate (urn-shaped e.g., Bryophyllum), tubular
l On the basis of number of carpels present, it can be attached to back and the anther can swing freely, e.g., grasses).
can be longitudinal (long slits appear lengthwise e.g., (tube-like or cyclindrical e.g., disc floret of sunflower), infundibuliform (funnel
moncarpellary (one carpel only) or multicarpellary l The cohesion between the stamens may be monadelphous (all the filaments
mustard), transverse (breadthwise slits, e.g., Malva), shaped e.g., Petunia), rotate (corolla with short tube having limbs placed
(many carpels) which can be apocarpous (carpels free e.g., united into a single bundle, anthers free, e.g., China rose), diadelphous (filaments
porous (pores appear at the tip e.g., Solanum or base transversely like a saucer or the spokes of a wheel, e.g., Solanum nigrum),
Ranunculus) or syncarpous (carpels fused e.g., Petunia). united in two bundles, anthers free, e.g., pea), polyadelphous (filaments united
e.g., Cassia), valvular (split at several places by lifting of salver shaped or hypocrateriform (tubular corolla with spreading lobes,
l On the basis of number of locules (chamber) present in into more than two bundles, anthers free, e.g., castor), syngenesious (anthers
surface layers, e.g., Barberry), irregular (e.g., Najas). e.g., Clerodendrum), bilabiate (bilipped corolla with gaping, wide open
the ovary, it can be unilocular (pea), bilocular united into a bundle, filaments free, e.g., sunflower), synandrous (anthers as mouth, e.g., Ocimum), personate (bilipped corolla with closed lips,
(mustard), trilocular (Asparagus), tetralocular l Longitudinal dehiscence may be laterorse (slits
well as filaments united throughout the length, e.g., Colocasia). The free e.g., Antirrhinum), ligulate or strap-shaped (short and narrow
(Ocimum), pentalocular (China rose) or on sides), introrse (slits towards the inner side
stamens are called polyandrous. tube-like corolla with upper part flattened like a strap, e.g.,
multilocular (Althaea). or centre of the flower), extrorse (slits lie
towards the outer side of flower). l On the basis of adhesion of stamens to flower, it can be ray floret of sunflower), spurred (one or more petals
epiphyllous (attached to perianth, e.g., Asphodelus), drawn out like a beak or spur, e.g., Larkspur).
epipetalous (attached to petals, e.g., Datura),
gynandrous (attached to gynoecium, e.g.,
Calotropis).
PLACENTATION
l Arrangement of placenta (which bear
R. K. MALIK’S
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Forebrain
covered by two meninges; duramater (outer) and pia-arachnoid (inner).
Anterior Cerebral
Brain is divisible into three parts: Forebrain, midbrain and hindbrain. choroid hemisphere
Circulatory System
l Body of a frog is pointed anteriorly and rounded Spinal cord is located in the vertebral column and joins the medulla plexus
posteriorly. It is slightly flattened dorsoventrally, oblongata via foramen magnum of the cranium (brain case). Pineal body Diencephalon
l Circulatory system of frog is closed and includes heart, arterial – Peripheral nervous system (PNS) includes 10 pairs of cranial nerves Pineal stalk
streamlined to swim through water and divisible into
Midbrain
Optic lobe
system, venous system, blood and lymphatic system. and 9 pairs of spinal nerves . Rarely 10th (paired or unpaired) spinal
head and trunk without neck and tail.
l Heart is three chambered made up of two anterior atria or auricles nerve is found.
l Skin of frog is thin, moist, smooth, slimy and green Posterior
and a single posterior ventricle. Two additional chambers are sinus
Hindbrain
Cerebellum
coloured with black or brown spots dorsally and – Autonomic nervous system is made up of sympathetic and choroid
venosus and truncus arteriosus. plexus Medulla
lighter pale yellow ventrally. There are no scales or parasympathetic nerves which controls and coordinates the
l The two auricles, right (larger) and left, are completely separated oblongata
any other hard exoskeleton parts. involuntary activities of the visceral organs.
from each other by inter-auricular septum. Both auricles open into Spinal cord
l Skin of back has dorsolateral folds or thickenings
single ventricle by a common large auriculo-ventricular aperture l Five types of sense organs are skin (tangoreceptor), taste buds
called dermal plicae. (gustatoreceptor), nasal chambers (olfactoreceptor), eyes (photoreceptor) Central canal
guarded by two pairs of auriculo-ventricular valves.
l Head is roughly triangular with a short blunt anterior l The inner surface of ventricle has irregular ridges called columnae and ears (stato-acoustic organs). Fig.: Dorsal view of brain of frog
snout terminating in a large transverse mouth. It carneae or trabeculae, with depressions called fissures.
bears external nares or nostrils, eyes, brow spot and
Urinogenital System
ear drums on the upper side. Aortic trunks Carotid arch
l Frogs have two large and protruding eyes, having an Systemic arch l In frogs, the excretory and reproductive systems are closely associated, hence they are together called urinogenital system.
Synangium Pulmocutaneous arch l Excretory system comprises of kidneys, ureters in females, urinogenital ducts in males, cloaca and urinary bladder. Kidneys
almost immovable upper eyelid and a thin semi-
Right auricle Opening leading to
transparent and freely movable lower eyelid. From pulmocutaneous arch are the chief excretory organs which are made up of large number of uriniferous tubules or nephrons.
Wall of truncus l From the kidneys, arise ureter in females and urinogenital duct in males.
lower eyelid arises nictitating membrane that arteriosus Left auricle
protects eyes during swimming. Anterior row of Opening of l Cloaca receives faecal matter, genital products and urine (from kidney). Ventrally it is attached to urinary bladder.
semilunar valves pulmonary veins
l Vocal sacs act as resonators to intensify sound of Sinu-auricular
l In males, near each kidney there is a cylindrical testis from which several thin vasa efferentia, connecting the testes to
Cavum aorticum
Pylangium
croaking during breeding season. aperture kidneys on each side. The vasa efferentia run transversely through mesorchium and open into the Bidder’s canal which in
Spiral valve Inter-auricular
l Trunk consists of thorax, abdomen and a pair of turn opens into the ureter. Histologically, each testis is a compact mass of seminiferous tubules, the epithelial lining of
Cavum septum
forelimb and hindlimb. pulmocutaneum Auriculo which produces sperms. The sperms when mature are dropped into the lumen to pass into the ureter through vasa
l Frog shows sexual dimorphism as male frog Posterior row of ventricular valve efferentia and Bidder’s canal.
possesses developed vocal sacs and nuptial pad semilunar valves Chordae tendineae
l Females have two ovaries where ova are produced by ovarian follicles. On each side of an ovary is an oviduct which starts
during breeding season and their body is somewhat posteriorly and forms uterus, which opens into the cloaca. During breeding season ova are released into the coelom and
slender and darker in colour than female frog. Fissures then they reach the ovarian funnels from where they pass to the ovisacs, cloaca and then outside.
l Egg of frog is telolecithal.
External nares Nictitating membrane Columnae carneae Part of oesophagus
(for respiration) (protects the eye during Cystic duct Oesophagus Ostium
Eye swimming and keeps Wall of ventricle
Right lobe Ovarian funnel
it moist in air) Left lobe of liver Fat body
Head Tympanum (receives Fig.: Internal structure of heart of frog (ventral view) of liver
Bile duct
sound waves) Gall bladder Testis
Digits Pancreatic ducts Mesorchium
Forelimb
Manus
Antebrachium
Digestive System Pancreas Fat body
Stomach Adrenal gland
Brachium l The digestive system mainly consists of alimentary canal and its Mesentery Ovary
Trunk Hepatopancreatic Left kidney
Dorsolateral associated glands. Ovarian tube
dermal plicae
duct
l Mouth leads into a buccopharyngeal cavity which opens into Urinogenital duct
Pyloric constriction Kidney
Thigh oesophagus through gullet. Duodenum (carries urine
and sperm) Adrenal gland
Anus l Stomach is situated behind the oesophagus and divisible into Ileum
cardiac stomach and pyloric stomach. Spleen Part of
Hindlimb rectum Seminal vesicle
l The small intestine is divisible into an anterior duodenum and a Ureter
Crus (shank ) Rectum (carries urine)
posterior ileum. Digestion of food and absorption of digested food Urinary bladder
Urinary Ovisac
Pes (foot) Toes occur in the small intestine. Cloaca bladder Cloacal chamber
l Ileum leads to rectum or large intestine. The rectum opens into the Cloacal chamber
Cloacal aperture Urinary
cloaca through the anus. bladder
l Digestive glands of frog include liver, pancreas, gastric glands and Cloacal aperture Cloacal aperture
Web Fig.: Alimentary canal (except
Fig.: Dorsal view of frog intestinal glands.
buccopharyngeal cavity) of frog Fig.: Urinogenital system of male frog Fig.: Urinogenital system of female frog
R. K. MALIK'S NEWTON CLASSES, RANCHI
R. K. MALIK'S NEWTON CLASSES, RANCHI
R. K. MALIK'S NEWTON CLASSES, RANCHI
PHYTOHORMONES
Plant growth regulators or hormones are broadly classified into two categories.
Auxin Ethylene
Nature: Weakly acidic growth hormone having an unsaturated ring structure. Auxins refer to natural (IAA, PAA, IAN) and Nature: It is the only gaseous phytohormone which
synthetic (Indole 3-butyric acid, NAA, 2, 4-D, 2, 4, 5-T) compounds having similar structure and properties. stimulates transverse or isodiametric growth but
Discovery: Darwin (1880) was first to find sensation of unilateral illumination in the coleoptile tip of canary grass. Later retards the longitudinal one.
Kogl and Smith (1931) isolated three chemicals from human urine which they named as auxin a , auxin b and hetero Discovery: R. Gane (1934) found that H H
auxin. IAA is the universal natural auxin. substance causing ripening was C C
Location: It is found in shoot apices, leaf primordia and developing seeds and is synthesised from amino acid tryptophan, ethylene. But it was recognised as a H H
a precursor of IAA or auxins. plant hormone by Crocker et al (1935).
Bioassay: Avena curvature test and root growth inhibition test are done for examining auxin effect. Location: It is found in almost all parts of plants in
minimal amount but maximum production occurs
Physiological functions:
during ripening of fruits and in tissues undergoing
l Auxins like 2, 4-D and 2, 4, 5-T acts as weedicides by being selectively harmful to broad leaved dicot weeds.
the sensitivity to gravity.
l Helps in root initiation, growth of lateral roots and
l Induces flowering in litchi and pineapple.
root hairs.
l NAA increases the number of dwarf shoots and fruits on them.
l Stimulates flowering in pineapple and other related
l Prevents pre-harvest fruit drop of orange and apple (by low concentration of 2, 4-D) and tomato (by NAA)
plants and helps in synchronising fruit set.
l Induces ripening of fleshy climacteric fruits and
of fungus, Gibberella fujikuroi (causing bakane disease in rice plants) and e.g., banana, mango, apple, tomato.
named it gibberellin. GA3 was first gibberellin to be isolated in its pure form CO l Ethylene is used to induce feminising effect e.g.,
and remains the most extensively studied. number of female flowers and thus fruits in
OH
cucumber.
Location: The major sites of gibberellin production in plants are embryos, HO
CH COOH l Ethylene also permits thinning of excess flowers
roots and young leaves near the shoot tip. Mevalonic acid (derived from 3 CH2
and young fruits so as to allow better growth of
acetyl Co-A) acts as precursor for synthesis of gibberellins. It is transported remaining fruits.
through simple diffusion as well as via conducting channels.
Bioassay: Barley endosperm test and germination of dwarf pea seeds are used as bioassays.
Abscisic Acid
Physiological functions:
Nature: It is a mildly acidic growth hormone which acts
l Stimulates stem elongation and leaf expansion.
as a general growth inhibitor. It is also called as stress
l Overcome natural dormancy of buds, tubers, seeds etc. hormone since its production is stimulated under
l Induces elongation of reduced stem or bolting in rosette plants e.g., henbane, cabbage. conditions of drought, water logging and adverse
environmental conditions.
l Promotes seed germination by inducing production of hydrolytic enzymes for solubilising reserve food.
l Promotes flowering in long day plants during non-inductive period.
H trans CH3 cis
l Controls fruit growth and development as well as induces parthenocarpy.
l Promotes formation of male flowers on female plants e.g., Cannabis. They can also replace female flowers with male H3C CH3 C
C C
ones on monoecious plants of cucurbits. 6 5 C 3 H
4 2
5 1 OH
Commercial uses: H 1COOH
l Exogenous application of GA4 and GA7 mixture to increase the number and size of fruits e.g., apple, grapes, tomato. 4 2
O 3 CH3
l Production of seedless pomaceous fruits by parthenocarpy.
l GA7 delays senescence of fruits and delays its ripening thus, extending its shelf life and storage period.
l Induces offseason flowering in many long day plants as well as those requiring vernalisation.
Discovery: The hormone was first isolated by Addicott
et al (1963) from cotton bolls.
l Application of gibberellins increases length of stem and yield of sugarcane.
Location: It is found in many parts of the plant but is
l Promotes early maturity resulting in seed production in juvenile conifers. more abundant in chloroplast of green cells. It is
synthesised from mevalonic acid or xanthophyll.
Bioassay: Rice seedling growth inhibition test and
Cytokinin inhibition of a-amylase synthesis in barley endosperm
are used as bioassay.
Nature: These are basic hormones, being derivatives of either aminopurine or phenyl urea that promote cytokinesis.
Physiological functions:
Discovery: The first cytokinin was discovered from autoclaved herring sperm DNA which stimulated cell division in l Induces dormancy of buds, seeds and underground
tobacco pith cells. It is called kinetin and does not occur naturally in plants. stems, hence also called as dormin.
CH2OH
The first natural cytokinin was obtained from unripe maize grains, called zeatin CH2 C l Promotes abscission of flowers and fruits.
(6-hydroxy 3-methyl trans 2-butenyl amino purine). NH CH CH3 l Induces senescence of leaves by promoting
It is found in coconut milk. degradation of chlorophyll and proteins.
Location: It is mainly found in roots, however it is also synthesised in endosperm regions N N l Stops cambium activity (in vascular cambium)
of seeds, growing embryos, young fruits and developing shoot buds. towards the approach of winter.
N
Bioassay: Tobacco pith culture, retardation of leaf senescence and excised radish N H l Inhibits seed germination by inhibiting gibberellin
cotyledon expansion are used as bioassays for cytokinins. Zeatin mediated amylase formation.
Physiological functions: l It is antagonist to gibberellin and counteracts the
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MAP
Human skeleton is made up of 206 bones (300 bones in
newborns) which are distributed into axial and appendicular
skeleton.
Sacrum (5 fused
centrum or body and a posterior Femur
pelvis backbones) portion, the neural arch.. The latter
Coccyx encloses the spinal cord. The hole formed Ilium PELVIC GIRDLE
by the neural arch is the vertebral
foramen. The vertebral foramina of all Lumbar · The pelvic girdle
girdle, also
twenty four vertebrae form the vertebral canal or neural canal. vertebra Patella called the hip girdle, is
Sacrum
· Vertebrae are categorised into five groups: cervical (7), thoracic (12), composed of two coxal
lumbar (5), sacral (5) and coccygeal (4). (hip) bones. Ilium
· Vertebral column displays four curves to enhance balancing powers and firmness for · The coxal bones are also
called the ossa coxae or Coccyx
upright posture of the body. These curvatures are cervical, thoracic, lumbar and pelvic Fibula Pubis Acetabulum
(=sacral). innominate bones. Ischium
· Each coxal bone consists Pubic
· Between the centre of adjacent vertebrae there are elastic pads of fibrocartilage, the Tibia Pubic arch symphysis
intervertebral discs which provide mobility to the vertebrae, check undue frictions and of three separate parts :
take up shocks. the ilium (short and
Tarsals
· Vertebral column carries the weight of the body in motion and when the organism is standing. straight bone), the ischium (lower elongated bone, running parallel to
Metatarsals vertebral column) and the pubis (inner, smaller bone).
STERNUM Phalanges · On its outer surface it has a deep depression called the acetabulum which,
Clavicular
with almost spherical head of the femur, forms the hip joint.
notch
· This is a flat bone which is present just under the
· It supports the weight of the body from the vertebral column. It also protects
Manubrium skin in the middle of the front of the chest. It is about
and supports the lower organs, including the urinary bladder, the
Sternal 15 cm long.
angle reproductive organs, and the developing foetus in case of a pregnant
Facets for · Its shape is like a dagger and consists of three parts—the woman.
attachment
Body of ribs 1-7 manubrium is the uppermost part, the body is the middle portion
and the xiphoid process is the tip of the bone.
· The true ribs (7 pairs) are attached to the sternum. HINDLIMBS
Xiphoid
process · It protects the internal organs in the thoracic region and helps in the · Each leg has 30 bones which constitute 1 femur, 1 patella, 1 tibia, 1 fibula, 7 tarsals, 5
respiratory mechanism. metatarsals and 14 phalanges.
· Femur, tibia and fibula bones together support the shank of the leg. The tarsals form the
RIBS ankle, metatarsals form the sole and phalanges form the digits of the foot.
· The ribs are thin, flat, curved bones that form a protective cage around the organs in the upper body. · The femur is the longest, largest, and strongest bone in the body whose head fits into the
· Ribs comprise of 24 bones arranged in 12 pairs. Each rib remains attached to the respective thoracic acetabulum of hip girdle.
vertebra. · The tibia connects to the femur to form the knee joint and with the talus, a foot bone, to
· The first seven pairs of ribs are attached directly with the sternum and are called true ribs. The 8th, 9th and allow the ankle to flex and extend.
10th pairs of ribs do not articulate directly with sternum, but join the seventh rib by hyaline cartilage. These · The tibia is larger than the fibula because it bears most of the weight, while the fibula
are called vertebrochondral ribs or false ribs. The last two (11th and 12th) pairs of ribs remain free serves as an area for muscle attachment.
anteriorly and are not attached either to sternum or cartilage of another rib, and are called floating ribs. · Fibula is shorter, thinner and slender.
· A typical rib consists of 2 parts : vertebral and sternal. The vertebral part is long and bony. It articulates with · Each ankle is composed of seven tarsals which are calcaneum, talus, cuboid, navicular
the thoracic vertebrae. and first, second, third cuneiforms.
· The sternal part is short and cartilaginous. It articulates with the sternum or sternal part of its upper rib. · The leg bones carry the weight of the body and are involved in propulsion and support.
JEE (MAIN & ADV.), MEDICAL
R. K. MALIK’S + BOARD, NDA, FOUNDATION
Synapse is an anatomically specialised junction between two neurons, where the axon (or some
MAP
other portion) of one cell (neuron) terminates on the dendrites or some other portion of another
cell. The term 'synapse' was first introduced by Charles Sherrington (1924). Transmission of nerve
impulse takes place across a synapse between neurons or neurons and an effector. The neuron
which sends messages is called presynaptic cell whereas the neuron which receives messages is
postsynaptic neuron.
Sending neuron
Receiving
neuron
STRUCTURE OF SYNAPSE
Most of the synapses comprise the following structures:
(i) Synaptic knob – Terminal bulbous ending of presynaptic axon which is devoid of neurofilaments but its
Sending
cytoplasm contains: neuron
(a) Synaptic vesicles – Small vesicles present in presynaptic cytoplasm that contain neurotransmitters
(for excitation or inhibition), like acetylcholine, GABA, etc. Action
(b) Mitochondria, ER and microtubules. potential
(c) Presynaptic membrane – Nerve membrane which is in close approximation with membrane of
Vesicle containing
postsynaptic cell. neurotransmitters
(ii) Sub-synaptic and postsynaptic membrane – The surface of the cell membrane involved in the
synapse is called the sub-synaptic membrane and the remaining of the motor neuron cell membrane is
Synaptic gap Axon terminal
called the postsynaptic membrane. Receptor sites for neurotransmitters are usually located on the sub-
R. K. MALIK'S NEWTON CLASSES, RANCHI
Postsynaptic
TYPES OF SYNAPSE
Receptor sites on
membrane
receiving neuron
Neurotransmitter
On
On th ion wi
the
locat
b asi
so
Chemical Synapse
e b thin
fp
hy
asi
io
of chemical messenger – a neurotransmitter,
fp r
lo
ox
g
r
Myelin
Axon
Postsynaptic Postsynaptic
Gap
neuron neuron
junction
Ions flow through gap Synaptic Neurotransmitter released
Presynaptic junction channels
Dendro-dendritic membrane vesicle fusing Presynaptic membrane
synapse Axo-somatic
synapse
Synaptic
cleft
At Electrical
PROPERTIES OF SYNAPSE Synapse
l Gap junctions in electrical
synapse allow the local currents
l Convergence and Divergence : Many presynaptic resulting from arriving action potentials
neurons converge on any single postsynaptic neuron, At Chemical to flow directly across the junction from
e.g., in spinal motor neurons, some inputs come from Synapse one neuron to the other.
dorsal root, some from long descending spinal tracts l This depolarises the membrane of
Mechanism of chemical transmission across a
and many from interconnecting neurons. The axons of the second neuron to threshold,
synapse is as follows:
most presynaptic neurons divide into many branches continuing the propagation of
that diverge to end on many postsynaptic neurons. Action potential arrives at axon terminal the action potential.
l Fatigue : Repeated stimulation of presynaptic neuron
leads to gradual decrease and finally disappearance of Voltage gated Ca2+ ion channels open and electrochemical
the postsynaptic response. This is due to exhaustion of gradient favours influx of Ca2+ and Ca2+ flows into axon terminal
chemical transmitter, as its synthesis is not as rapid as
the release. Ca2+ ions cause synaptic vesicles to move to the
l Synaptic Delay : When an impulse reaches the surface of the knob and fuse with synaptic membrane terminal
presynaptic terminal, there is a gap of about 0.5 msec.,
before a response is obtained in postsynaptic neuron. Vesicles release neurotransmitters by exocytosis
This is due to the time taken by synaptic mediator to be
released and to act on postsynaptic membrane.
Neurotransmitters diffuse across synaptic cleft and bind to
l Synaptic Plasticity : Plasticity implies the capability of
receptors on postsynaptic membrane
being easily moulded or changed. Synaptic conduction
thus can be increased or decreased on the basis of past
experience. These changes can be presynaptic or This causes depolarisation and generation of
postsynaptic in location and play an important role in action potential in the postsynaptic
learning and memory. membrane.
R. K. MALIK'S NEWTON CLASSES, RANCHI
MAP is maintained. It is of two types: asexual and sexual. Asexual reproduction is the formation of new individual without involving
fusion of gametes. It is uniparental as offspring are produced by a single parent.
Fission
l It is a type of asexual reproduction in which
CHARACTERISTICS OF ASEXUAL REPRODUCTION
the parent organism divides into two or
more daughter cells. l It is more primitive than sexual reproduction as it involves only mitotic divisions.
l In this type of reproduction, whole parent l New organisms are produced from the somatic part of parental organism, so it is also called as
body acts as the reproductive unit. (a) somatogenic reproduction.
l It is of three types: l New individuals produced are genetically similar to the parent as well as to each other and are called
(a) Binary fission : In this, parent organism clones. Hence, it plays no role in evolution.
divides into two halves, each half forming l Unit of reproduction may be either whole parent body, or a bud, or a body fragment, or a single
an independent daughter organism. It can somatic cell.
be simple (occurs through any plane, e.g., (b) l It is usually found in lower organisms like protistan protozoans (Amoeba, Paramecium), sponges
Amoeba), longitudinal (plane of division is
longitudinal axis of body, e.g., Euglena), (Scypha), coelenterates, (Hydra, Tubularia, etc.), certain flatworms (Planaria), some worms and
R. K. MALIK'S NEWTON CLASSES, RANCHI
Budding Sporulation
l Budding refers to the process of formation of daughter l Spores are minute, single celled, thin or thick walled
individuals from a small projection or bud arising on the propagules which are dispersive structures released from
parent body. the parent body and form new individuals. Spore formation
l Each bud enlarges, develops parental characters and TYPES OF is common in members of monera, protista, algae and fungi.
separates to lead an independent life. Some of the commonly produced spores are:
l Budding can be either exogenous (formed on the outer surface) ASEXUAL (a) Zoospores : Motile and flagellated spores produced inside
e.g., Hydra, yeast or endogenous (formed inside
parent body) e.g., Spongilla. In Spongilla, bud is called a gemmule. REPRODUCTION zoosporangia. Flagella help in proper dispersal in aquatic
habitat. E.g., algae and lower fungi like Phycomycetes.
(b) Conidia : Non-motile spores produced singly or in chains by
Tentacles constriction at the tip or lateral side of special hyphal
branches called conidiophores. These are dispersed by wind
and germinate to form new individuals. E.g., Penicillium.
(c) Chlamydospores : Thick walled spores produced directly
from hyphal cells. May be terminal or intercalary in position
and capable of withstanding unfavourable conditions. E.g.,
Rhizopus.
A projection grows on Bud grows Splits away from parent Fragmentation (d) Oidia : Small fragments of hyphae that are thin walled and
outer surface of body. externally. and grows independently. do not store reserve food material. Oidia give rise to new
l In this type of reproduction, parent body breaks into
hyphae. These are formed under conditions of excess water,
two or more pieces called fragments.
(a) Nuclear division Bud sugar and certain salts. E.g., Agaricus.
l Each fragment develops into a new organism.
(e) Sporangiospores : Non-motile spores produced inside
Developing l In fragmentation, rate of reproduction is high.
sporangia. Usually get dispersed by wind and germinate to
bud l It occurs in flatworms, sea anemones, coelenterates,
form new mycelium. E.g., Rhizopus, Mucor.
echinoderms, algae like Spirogyra, etc.
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R. K. MALIK'S NEWTON CLASSES, RANCHI
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R. K. MALIK'S NEWTON CLASSES, RANCHI
Type of Germ Type of
Chromosomes
cells Division
No.
Oogonium Mitosis
Multiplication 46
Chromosomes Type of Germ cells Type of Division (2n)
Phase
No.
Spermatogonium Mitosis
Multiplication 46 Additional
(2n)
Phase oogonia (2n)
Growth Phase
Primary 46 Primary
Growth Phase 46 Additional oocyte (2n)
spermatocyte (2n) spermatogonia (2n)
Meiosis I Meiosis I
23 Secondary
spermatocytes (n) Secondary
Maturation Phase 23 First polar
Meiosis II body (n) oocyte (n)
Spermatids Sperm (n)
23 (n) Maturation Phase
Meiosis II
Second polar Male pronucleus
Spermatozoa 23 Mature
bodies (n)
Spermiogenesis (sperms) (n) ovum (n)
23 (All degenerate)
Female pronucleus
Fig.: Stages in spermatogenesis (diagrammatic) Fig.: Stages in oogenesis (diagrammatic)
Inhibits Hypothalamus
GnRH Hypothalamus
and LH GnRH
production Anterior pituitary GnRH
LH Testis FSH Positive Anterior pituitary Negative
Leydig’s cells Sertoli cells feedback feedback
LH/FSH
Testosterone Ovary
Stimulates
Spermato- Estrogen and
Positive Testosterone genesis
Progesterone
feedback
Negative Reproductive tract Inhibin Uterus
feedback and other organs
To MAKE YOUR CONCEPTS CRYSTAL CLEAR
R. K. MALIK’S VISIT OUR YOU TUBE CHANNEL : MATHEMATICS MADE INTERESTING JEE (MAIN & ADV.), MEDICAL
Office.: 606 , 6th Floor, Hariom Tower, Circular Road, Ranchi-1,
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NEWTON CLASSES
R. K. MALIK'S NEWTON CLASSES, RANCHI
CONCEPT TRANSLATION
Translation is the process by which a protein or polypeptide is
Ÿ
Translational Machinery
Basic translational machinery is same in eukaryotes and
prokaryotes, however few differences occur. It is composed of
MAP synthesised inside the living cell using mRNA as a template. This
biochemical process is called translation because the information
present in the form of four letter alphabet of nucleic acid is
translated into twenty letter alphabets of proteins.
five components, i.e., mRNA, tRNA, amino acid, enzymes
(aminoacyl tRNA synthetase, peptidyl transferase) and ribosome.
Ÿ In eukaryotes, each mRNA is monocistronic (encodes for only
one polypeptide) whereas in prokaryotes, many mRNAs are
polycistronic (encodes for two or more different polypeptides).
Ÿ Ribosome is the site of protein synthesis. 70S (30S + 50S)
MECHANISM OF TRANSLATION ribosomes are involved in prokaryotic translation while 80S
Ÿ The steps of translation are common in both prokaryotes and eukaryotes. Three main steps involved in (40S + 60S) ribosomes are involved in eukaryotic translation. The
translation are initiation, elongation and termination. Before initiation amino acids are activated and attached two subunits of ribosomes associate at the time of protein
to tRNAs in two steps called activation of amino acids and charging or aminoacylation of tRNA respectively. synthesis and then separate after the completion of process.
Ÿ In eukaryotes, the initiating amino acid is methionine, not N-formylmethionine (fMet) as in prokaryotes. Ribosomes have three sites; the peptidyl (P) site, aminoacyl (A)
Ÿ The main difference between initiation of translation in prokaryotes and eukaryotes is that in bacteria, a Shine- site and exit (E) site. Eukaryotic ribosomes do not have E-site.
Dalgarno sequence (4 to 9 purine residues, 8 to 13 base pairs to the 5’ side of initiation codon) guides correct Ÿ tRNAs pick up particular amino acids (at CCA or 3’ end) and take
initiation codon (5’ AUG) and is the binding site for the 30S ribosomal subunit. the same to mRNA over particular codons corresponding to their
Ÿ In contrast, most eukaryotic mRNAs do not contain Shine-Dalgarno sequences. Instead, a 40S ribosomal anticodons. Each tRNA contact with ribosome at T y C loop and
subunit attaches at the 5’ end of the mRNA and moves downstream (i.e., in a 5’ to 3’ direction) until it finds the the enzyme aminoacyl tRNA synthetase at DHU loop. Eukaryotic
AUG initiation codon. This process is called scanning. mRNAs have 5’ - cap and poly A tail at 3’ end.
1
R. K. MALIK'S NEWTON CLASSES, RANCHI
eIF3, 1, 1A
subunit.
Ÿ The small subunit then binds to the GTP
eIF4B eIF5
mRNA via complementary pyrimidine IF2 eIF4E
5’ UA C 3’ Met
rich sequences close to 3’ end of 16S rRNA mRNA AUGUUUGGC CUUGCUA C C GCUU 4G, 4A eIF2
guided by Shine-Dalgarno sequence and 30S Cap
moves 3’ along the mRNA until it locates AUG
IF1 IF3
the AUG initiation codon. 40S
Ÿ The initiator tRNA charged with N-
formlymethionine and a complex of IF2 eIF4E
4G, 4A
and GTP (fMet-tRNAfMet/IF2/GTP) now binds to mRNA and 30S subunit. eIF4B
Ÿ The complex of mRNA, fMet-tRNAfMet, IF1, IF2 and the 30S ribosomal subunit is called the 30S initiation complex. eIF3, 1, 1A Met
Ÿ Structural changes then lead to the ejection of IF1 and IF3 and IF2 now stimulates the association of 50S eIF2 eIF5
subunit of ribosomes. Simultaneously, the GTP bound to IF2 is hydrolysed to GDP and Pi and leading to
release of IF2. This forms 70S initiation complex. Cap AUG
Ÿ When this complex is formed, the ribosome is ready for the elongation phase.
3 3
Elongation A. Elongation
Charged tRNA
Ÿ Elongation requires three factors, i.e., Ÿ The elongation stage of translation in
EF – Tu, EF – Ts and EF – G and enzyme eukaryotes is quite similar to the prokaryotes.
peptidyl transferase. Ÿ It requires three elongation factors, eEF1a,
Ÿ The fMet-tRNAfMet occupies the P site E eEF1bg and eEF2 as counterparts of prokaryotic
site 3’ EF – Tu, EF – Ts and EF – G respectively.
and another aminoacyl tRNA complex
Ribosome ready for mRNA
(aminoacyl tRNA – EF – Tu – GTP) reach at Ÿ The GTP form of EF1a delivers aminoacyl tRNA to
next aminoacyl tRNA P A
A site depending upon the anticodon site site 2 GTP Peptide the A site of the ribosome and EF1bg catalyses the
5’
present on mRNA. EF – Ts and GTP are bond exchange of GTP for bound GDP. eEF2 mediates
required for the regeneration of EF – Tu – 2GDP formation GTP driven translocation similar to prokaryotic
GTP complex. D. B. EF – G.
Ÿ First peptide linkage is now established Ÿ As eukaryotic ribosome do not have E site,
between –COOH group of amino acids at uncharged tRNAs are expelled directly from the
P site and –NH2 group of amino acid at A P site.
Free E E Ÿ The elongated peptide chain or polypeptide
site catalysed by the ezyme peptidyl
tRNA lies in the groove of the larger subunit of
transferase.
Ÿ This produces a dipeptidyl – tRNA in the A P A P A ribosome.
site and now uncharged tRNA f M et
remains bound to the P site. With the help
GDP 4
of EF – G (translocase) dipeptidyl - tRNA
C. Termination
moves from A site to P site. The ribosome GTP
Ÿ Termination in eukaryotes is similar to that in
moves one codon toward the 3’ end of
mRNA (called translocation). Free tRNA prokaryotes.
slips to E site and from there to outside in Ÿ In eukaryotes, a single factor eRF1 recognises
E all three termination codons and with the
the cytoplasm.
Ÿ New codon exposed at A site attract new help of eRF3, ribosomal subunits are released.
aminoacyl tRNA complex and thus P A Ÿ eRF3 prevents the reassociation of ribosomal
peptide chain elongates. subunits in the absence of an initiation complex.
Free
4 Free tRNA polypeptide 50S
Termination Release
Ÿ Termination occurs when a non sense or stop codon (UAA, UAG, UGA) reaches A site. factor
Ÿ Stop codons are recognised by 2 release factors RF1 and RF2. A third factor RF3
mediates interaction between RF1 or RF2 with the ribosome. 5’
Ÿ RF1 is specific to UAG and UAA. 3’ 3’ mRNA
Ÿ RF2 is specific to UAA and UGA. 3’
Ÿ RFs hydrolyse the terminal peptidyl-tRNA bond, release polypeptide and last tRNA 5’ 5’
Release
from the P site and dissociates two subunits of ribosomes to start new cycle of factor
Stop codon
translation. (UAG, UAA or UGA) 30S
R. K. MALIK'S NEWTON CLASSES, RANCHI + BOARD, NDA, FOUNDATION
JEE (MAIN & ADV.), MEDICAL