ÉÖçÖíàäÄ, 2007, ÚÓÏ 43, ‹ 11, Ò.

1–6

ÉÖçÖíàäÄ êÄëíÖçàâ
ìÑä 635.21: 631.527.52: 575.13

àáìóÖçàÖ ÉÖçÖíàóÖëäéÉé êÄáçééÅêÄáàü ëéêíéÇ

äÄêíéîÖãü ë àëèéãúáéÇÄçàÖå èñê-ÄçÄãàáÄ Ñçä éêÉÄçÖãã
© 2007 „. í. Ä. ɇ‚ËÎÂÌÍÓ, é. û. ÄÌÚÓÌÓ‚‡, ã. à. äÓÒÚË̇
ÇÒÂÓÒÒËÈÒÍËÈ Ì‡Û˜ÌÓ-ËÒÒΉӂ‡ÚÂθÒÍËÈ ËÌÒÚËÚÛÚ ‡ÒÚÂÌË‚ӉÒÚ‚‡ ËÏ. ç.à. LJ‚ËÎÓ‚‡,
ë‡ÌÍÚ-èÂÚÂ·Û„ 190000
e-mail: t–gavrilenko@yahoo.com
èÓÒÚÛÔË· ‚ ‰‡ÍˆË˛ 16.08.2006 „.

ÉÂÌÂÚ˘ÂÒÍÓ ‡ÁÌÓÓ·‡ÁË 98 ÒÓÚÓ‚ ͇ÚÓÙÂÎfl ÓÚ˜ÂÒÚ‚ÂÌÌÓÈ Ë Á‡Û·ÂÊÌÓÈ ÒÂÎÂ͈ËË ËÒÒΉӂ‡-

ÌÓ ÔË ÔÓÏÓ˘Ë èñê Ò Ó„‡ÌÂÎÎÓÒÔˆËÙ˘Ì˚ÏË Ô‡ÈÏÂ‡ÏË. Ç˚fl‚ÎÂÌ ÔÓÎËÏÓÙËÁÏ Í‡Í Ô·ÒÚˉ-
Ì˚ı (atpE, trnG/trnR), Ú‡Í Ë ÏËÚÓıÓ̉ˇθÌ˚ı ÎÓÍÛÒÓ‚ (rps10, atp6). Ç ËÁÛ˜ÂÌÌÓÈ ‚˚·ÓÍ ÒÓÚÓ‚
‚˚fl‚ÎÂÌÓ ‚ÓÒÂϸ ‡Á΢Ì˚ı „‡ÔÎÓÚËÔÓ‚. èÓ͇Á‡Ì ÌÂÁ̇˜ËÚÂθÌ˚È ÛÓ‚Â̸ ÔÓÎËÏÓÙËÁχ Ó„‡-
ÌÂθÌ˚ı Ñçä Û ÒÓÚÓ‚ ͇ÚÓÙÂÎfl – ÔÓ‰‡‚Îfl˛˘Â ˜ËÒÎÓ ÒÓÚÓ‚ (91 ËÎË 92.9%) ÓÚÌÓÒËÎËÒ¸ Î˯¸ Í
‰‚ÛÏ „‡ÔÎÓÚËÔ‡Ï (I Ë II), ËÁ ÌËı 62 ÒÓÚ‡ ËÏÂÎË Ó‰ËÌ Ë ÚÓÚ Ê “ÍÛθÚÛÌ˚È” ÚËÔ ˆËÚÓÔ·ÁÏ˚ („‡ÔÎÓ-
ÚËÔ I). èÓ ˜‡ÒÚÓÚ ‚ÒÚ˜‡ÂÏÓÒÚË ÓÒÌÓ‚Ì˚ı „‡ÔÎÓÚËÔÓ‚ ÒÂÎÂ͈ËÓÌÌ˚ ÒÓÚ‡ ÓÚ˜ÂÒÚ‚ÂÌÌÓÈ Ë Á‡Û-
·ÂÊÌÓÈ ÒÂÎÂ͈ËË Ì ‡Á΢‡ÎËÒ¸.

ÉÂÌÂÚ˘ÂÒÍÓ ‡ÁÌÓÓ·‡ÁË ‚ˉӂ ͇ÚÓÙÂÎfl ÎËÁ ÔÓÎËÏÓÙËÁχ ÔÓÒΉӂ‡ÚÂθÌÓÒÚÂÈ ıÎÓÓ-

(Ó‰ Solanum L., ÒÂ͈Ëfl Petota Dumort.), ÔÓËÁ‡Ò-
Ú‡˛˘Ëı ÓÚ ˛„Ó-Á‡Ô‡‰Ì˚ı ‡ÈÓÌÓ‚ ëòÄ ‰Ó ˛Ê-
Ì˚ı „‡Ìˈ óËÎË, ‚Íβ˜‡ÂÚ 228 ‰ËÍËı Ë 7 ÍÛθÚÛ-
Ì˚ı ‚ˉӂ, ‚ ÚÓÏ ˜ËÒΠ‚ÓÁ‰ÂÎ˚‚‡ÂÏ˚È Í‡ÚÓÙÂθ
Solanum tuberosum L. [1]. ä‡ÚÓÙÂθ ̇ıÓ‰ËÚÒfl ̇
˜ÂÚ‚ÂÚÓÏ ÏÂÒÚ ‚ ÏË ÒÂ‰Ë Ì‡Ë·ÓΠ‚‡ÊÌ˚ı
ÔÓ‰Ó‚ÓθÒÚ‚ÂÌÌ˚ı ÍÛθÚÛ ÔÓÒΠԯÂÌˈ˚, ÍÛ-
ÍÛÛÁ˚ Ë ËÒ‡ [1]. Ç ÚÓ Ê ‚ÂÏfl ͇ÚÓÙÂθ ÓÚÌÓ-
ÒËÚÒfl Í ÚÂÏ ÒÂθÒÍÓıÓÁflÈÒÚ‚ÂÌÌ˚Ï ÍÛθÚÛ‡Ï,
‚ÓÁ‰ÂÎ˚‚‡ÌË ÍÓÚÓ˚ı Ú·ÛÂÚ Ì‡Ë·ÓΠËÌÚÂÌ-
ÒË‚ÌÓ„Ó ËÒÔÓθÁÓ‚‡ÌËfl ıËÏ˘ÂÒÍËı Ò‰ÒÚ‚ Á‡˘Ë-
Ú˚ ÓÚ ·ÓÎÂÁÌÂÈ Ë ‚‰ËÚÂÎÂÈ [2]. í‡ÍÛ˛ ÒËÚÛ‡ˆË˛
ÏÓÊÌÓ Ó·˙flÒÌËÚ¸ ÌËÁÍËÏ ÛÓ‚ÌÂÏ „ÂÌÂÚ˘ÂÒÍÓ„Ó
‡ÁÌÓÓ·‡ÁËfl ‚ÓÁ‰ÂÎ˚‚‡ÂÏÓ„Ó Í‡ÚÓÙÂÎfl [3, 4],
˜ÚÓ ÏÓÊÂÚ ·˚Ú¸ Ò‚flÁ‡ÌÓ Ò fl‰ÓÏ Ô˘ËÌ:
1) Ì·Óθ¯ËÏ ˜ËÒÎÓÏ ËÌÚÓ‰Û͈ËÈ ˛ÊÌÓ‡ÏÂ-
Ë͇ÌÒÍËı ÍÛθÚÛÌ˚ı ‚ˉӂ ‚ Ö‚ÓÔÛ [5] Ë Ó„‡-
Ì˘ÂÌÌ˚Ï Ì‡·ÓÓÏ ËÒıÓ‰Ì˚ı ÙÓÏ, ËÒÔÓθÁÓ‚‡Ì-
Ì˚ı ‚ ÒÂÎÂ͈ËÓÌÌ˚ı ÔÓ„‡Ïχı, – ÚÓθÍÓ Ì‡ ÓÒ-
ÌÓ‚Â ÒÓÚ‡ Early Rose ·˚ÎÓ ÒÓÁ‰‡ÌÓ ·ÓΠ1000
ÒÂÎÂ͈ËÓÌÌ˚ı ÒÓÚÓ‚ [6];
2) ÌËÁÍËÏ ÛÓ‚ÌÂÏ ËÌÚÓ„ÂÒÒËË ‚Ó ‚ÌÓ‚¸ ÒÓ-
Á‰‡‚‡ÂÏ˚ ÒÂÎÂ͈ËÓÌÌ˚ ÒÓÚ‡ „ÂÌÂÚ˘ÂÒÍÓ„Ó
χÚÂˇ· ‰ËÍËı ‚ˉӂ – ËÁ-Á‡ ·‡¸ÂÓ‚ ÌÂÒÍÂ-
˘Ë‚‡ÂÏÓÒÚË ‚ ÒÂÎÂ͈ËÓÌÌ˚ ÔÓ„‡ÏÏ˚ ‚ӂΘÂ-
ÌÓ ÏÂÌ 10% ‰ËÍËı ‚ˉӂ ͇ÚÓÙÂÎfl [7];
3) Ó„‡Ì˘ÂÌËÂÏ ‚ÓÁÏÓÊÌÓÒÚÂÈ „ÂÌÂÚ˘ÂÒÍËı
Ë ÒÂÎÂ͈ËÓÌÌ˚ı ËÒÒΉӂ‡ÌËÈ ËÁ-Á‡ ‚˚ÒÓÍÓÈ ÒÚÂ-
ÔÂÌË „ÂÚÂÓÁË„ÓÚÌÓÒÚË Ë ÚÂÚ‡ÔÎÓˉÌÓÈ ÔËÓ‰˚
„ÂÌÓχ ͇ÚÓÙÂÎfl Solanum tuberosum L., ‡ Ú‡ÍÊÂ
ËÁ-Á‡ ÓÚÒÛÚÒÚ‚Ëfl ÍÓÎÎÂ͈ËÈ ËÌ·‰Ì˚ı „ÂÌÂÚ˘Â-
ÒÍË Ï‡ÍËÓ‚‡ÌÌ˚ı ÎËÌËÈ.
é‰ÌËÏ ËÁ ÔÓ‰ıÓ‰Ó‚ Í ÓˆÂÌÍ ÏÂÊ- Ë ‚ÌÛÚË‚Ë-
‰Ó‚Ó„Ó „ÂÌÂÚ˘ÂÒÍÓ„Ó ‡ÁÌÓÓ·‡ÁËfl fl‚ÎflÂÚÒfl ‡Ì‡-
Ô·ÒÚÌÓÈ (ıÎÑçä) Ë ÏËÚÓıÓ̉ˇθÌÓÈ (ÏÚÑçä)
Ñçä. ì ‰ËÍËı Ë ÍÛθÚÛÌ˚ı ‚ˉӂ ͇ÚÓÙÂÎfl Ó·-
̇ÛÊÂÌÓ ·Óθ¯Ó ‡ÁÌÓÓ·‡ÁË „‡ÔÎÓÚËÔÓ‚ [8–
11]. ì ‰ËÍËı ‚ˉӂ ‚˚‰ÂÎfl˛Ú ÔflÚ¸ ÓÒÌÓ‚Ì˚ı
(T, W, C, S, A) [12] Ë fl‰ ÏËÌÓÌ˚ı ÚËÔÓ‚ Ô·ÒÚˉ-
ÌÓ„Ó „ÂÌÓχ [11]. W, ËÎË “‰ËÍËÈ”, ÚËÔ Ô·ÒÚÓχ
‚˚fl‚ÎÂÌ Û ·Óθ¯ËÌÒÚ‚‡ ËÁÛ˜ÂÌÌ˚ı ‰ËÍËı ‚ˉӂ
[12]. í (ËÎË “tuberosum) ÚËÔ Ô·ÒÚÓχ, Ò‚flÁ‡ÌÌ˚È Ò
‰ÂΈËÂÈ ıÎÑçä ‡ÁÏÂÓÏ 241 ÔÌ, – ÛÌË͇ÎÂÌ Ë
‚ÒÚ˜‡ÂÚÒfl ÚÓθÍÓ Û ‰‚Ûı (S. tuberosum L. Ë S. tar-
ijense Hawkes) ËÁ 235 ‚ˉӂ ͇ÚÓÙÂÎfl [14]. í-ÚËÔ
ıÎÑçä ËÏÂÂÚ ÔÓ‰‡‚Îfl˛˘Â ˜ËÒÎÓ ËÁÛ˜ÂÌÌ˚ı ‚-
ÓÔÂÈÒÍËı ÒÓÚÓ‚ Ë ˜ËÎËÈÒÍËı ‡·ÓË„ÂÌÌ˚ı ÙÓÏ
S. tuberosum subsp. tuberosum [15–18]. ì ·Óθ¯ËÌ-
ÒÚ‚‡ ËÒÒΉӂ‡ÌÌ˚ı ÌÂψÍËı ÒÓÚÓ‚ ‚˚fl‚ÎÂÌ˚
β- ËÎË α-ÚËÔ˚ ıÓ̉ËÓχ, ËÁ ÍÓÚÓ˚ı β-ÚËÔ fl‚ÎflÎ-
Òfl ÔÂӷ·‰‡˛˘ËÏ [9, 13]. èÓ˝ÚÓÏÛ Û ‚ÓÁ‰ÂÎ˚‚‡Â-
ÏÓ„Ó Í‡ÚÓÙÂÎfl í/β-ÚËÔ ˆËÚÓÔ·ÁÏ˚ ̇Á˚‚‡˛Ú
"ÍÛθÚÛÌ˚Ï”, ‡ W/α – “‰ËÍËÏ” [9, 13]. Ç Ì‡ÒÚÓfl-
˘Â ‚ÂÏfl ‡Á‡·ÓÚ‡Ì˚ ÒÔˆËÙ˘Ì˚ Ô‡ÈÏÂ˚,
ÔÓÁ‚ÓÎfl˛˘Ë ‡Á΢‡Ú¸ ̇˷ÓΠ‡ÒÔÓÒÚ‡ÌÂÌ-
Ì˚ ÚËÔ˚ „ÂÌÓÏÓ‚ Ó„‡ÌÂÎÎ ÔÓÒ‰ÒÚ‚ÓÏ ÔÓÎËÏÂ-
‡ÁÌÓÈ ˆÂÔÌÓÈ ‡͈ËË (èñê) [9, 13].
àÁ‚ÂÒÚÌÓ ‰ÓÒÚ‡ÚÓ˜ÌÓ ·Óθ¯Ó ˜ËÒÎÓ ‡·ÓÚ ÔÓ
ËÁÛ˜ÂÌ˲ ËÁÏÂ̘˂ÓÒÚË ˆËÚÓÔ·ÁχÚ˘ÂÒÍËı „Â-
ÌÓÏÓ‚ ÒÂÎÂ͈ËÓÌÌ˚ı ÒÓÚÓ‚ Á‡Û·ÂÊÌÓÈ ÒÂÎÂÍ-
ˆËË [9, 13, 16–18]. ÑÎfl „ÂÌÓÙÓ̉‡ ÒÓÚÓ‚ ÓÚ˜Â-
ÒÚ‚ÂÌÌÓÈ ÒÂÎÂ͈ËË Ú‡ÍË ËÒÒΉӂ‡ÌËfl Ì ÔÓ‚Ó-
‰ËÎËÒ¸. ñÂθ˛ ‰‡ÌÌÓÈ ‡·ÓÚ˚ ·˚ÎÓ ËÁÛ˜ÂÌËÂ
„ÂÌÂÚ˘ÂÒÍÓ„Ó ‡ÁÌÓÓ·‡ÁËfl ÒÓÚÓ‚ ͇ÚÓÙÂÎfl
‡Á΢ÌÓ„Ó ÔÓËÒıÓʉÂÌËfl ÔË ÔÓÏÓ˘Ë èñê Ò
Ô‡ÈÏÂ‡ÏË, ÒÔˆËÙ˘Ì˚ÏË Í fl‰Û ÔÓÒΉӂ‡-
ÚÂθÌÓÒÚÂÈ ıÎ- Ë ÏÚÑçä.

1
2 ɇ‚ËÎÂÌÍÓ Ë ‰.
åÄíÖêàÄãõ à åÖíéÑõ
Ç Ì‡ÒÚÓfl˘ÂÈ ‡·ÓÚ ·˚ÎË ËÁÛ˜ÂÌ˚ 98 Ó·‡Á-
ˆÓ‚ ÍÛθÚÛÌÓ„Ó Í‡ÚÓÙÂÎfl Solanum tuberosum L.
ËÁ ÍÓÎÎÂ͈ËË Çàê, Ô‰ÒÚ‡‚Îfl˛˘Ëı ‰‚ „ÛÔÔ˚
cÓÚÓ‚ ‡Á΢ÌÓ„Ó ÔÓËÒıÓʉÂÌËfl: 1) 63 ÒÓÚ‡
ÓÒÒËÈÒÍÓÈ ÒÂÎÂ͈ËË Ë ÒÂÎÂ͈ËË ÒÚ‡Ì ëçÉ; 2) 35
ÒÓÚÓ‚ Á‡Û·ÂÊÌÓÈ ÒÂÎÂ͈ËË (ÒÚ‡Ì˚ Ö‚ÓÔ˚ Ë
ëòÄ), ‚ ÚÓÏ ˜ËÒΠÒÚ‡Ó‰‡‚ÌË ÒÓÚ‡, fl‚Îfl˛˘Ë-
ÂÒfl Ó‰Ó̇˜‡Î¸ÌË͇ÏË ÏÌÓ„Ëı ÒÓÚÂÌ ÒÓ‚ÂÏÂÌ-
Ì˚ı ÒÓÚÓ‚.
Ç˚‰ÂÎÂÌË ÚÓڇθÌÓÈ Ñçä ÔÓ‚Ó‰ËÎË ËÁ ÏÓ-
ÎÓ‰˚ı ÎËÒڸ‚ ‡ÒÚÂÌËÈ Ò ËÒÔÓθÁÓ‚‡ÌËÂÏ ÏÂÚÓ‰‡
ÇË̇̉‡ Ë î‡ÈÍÒ‡ [19] Ò ÌÂÍÓÚÓ˚ÏË ÏÓ‰ËÙË͇ˆË-
flÏË. Ä̇ÎËÁ ÔÓÎËÏÓÙËÁχ Ô·ÒÚˉÌ˚ı „ÂÌÓÏÓ‚
ÔÓ‚Ó‰ËÎË ÔË ÔÓÏÓ˘Ë èñê Ò ËÒÔÓθÁÓ‚‡ÌËÂÏ
Ô‡ÈÏÂÓ‚ ALC_1/ALC_3 Ë NTCP9, ÒÔˆËÙ˘Ì˚ı Í
ÎÓÍÛÒ‡Ï atpE Ë trnG/trnR ıÎÑçä ÒÓÓÚ‚ÂÚÒÚ‚ÂÌÌÓ [9,
20], ‡ ËÁÛ˜ÂÌË ÔÓÎËÏÓÙËÁχ ÏÚÑçä – ÔË ÔÓÏÓ-
˘Ë Ô‡ÈÏÂÓ‚ AL_Mt2/ALM_3 Ë ALM_4/ALM_5,
ÒÔˆËÙ˘Ì˚ı Í ÎÓÍÛÒ‡Ï atp6 Ë rps10 ÒÓÓÚ‚ÂÚ-
ÒÚ‚ÂÌÌÓ [9, 13]. àÒÔÓθÁÓ‚‡ÌÌ˚ ‚ ‡·ÓÚ ÒÔˆË-
Ù˘Ì˚ Ô‡ÈÏÂ˚ ÔÓÁ‚ÓÎfl˛Ú ˉÂÌÚËÙˈËÓ‚‡Ú¸
ÓÒÌÓ‚Ì˚ ÚËÔ˚ ˆËÚÓÔ·ÁÏ Í‡ÚÓÙÂÎfl. í‡Í, ÔË
‡ÏÔÎËÙË͇ˆËË Ñçä ÍÛθÚÛÌÓ„Ó Í‡ÚÓÙÂÎfl Ò
Ô‡ÓÈ Ô‡ÈÏÂÓ‚ ALC_1/ALC_3 Ë ‡Á‰ÂÎÂÌËË ÔÓ-
ÎÛ˜ÂÌÌ˚ı ÔÓ‰ÛÍÚÓ‚ ‚ ‡„‡ÓÁÌ˚ı „ÂÎflı ‚˚fl‚Îfl-
˛ÚÒfl ‰‚‡ ÚËÔ‡ Ù‡„ÏÂÌÚÓ‚ ‡ÁÏÂ‡ÏË 380 Ë 620
ÔÌ, fl‚Îfl˛˘ËıÒfl χÍÂ‡ÏË “ÍÛθÚÛÌÓ„Ó” (í) Ë
“‰ËÍÓ„Ó” (W) ÚËÔÓ‚ Ô·ÒÚˉÌ˚ı „ÂÌÓÏÓ‚ ÒÓÓÚ‚ÂÚ-
ÒÚ‚ÂÌÌÓ [9, 13]. èË ‡Á‰ÂÎÂÌËË ‚ ‡„‡ÓÁÌ˚ı „ÂÎflı
ÔÓ‰ÛÍÚÓ‚ ‡ÏÔÎËÙË͇ˆËË Ñçä Ò Ô‡ÓÈ Ô‡ÈÏÂ-
Ó‚ ALM_4/ALM_5 Ó·‡ÁÛÂÚÒfl ‰‚‡ ÚËÔ‡ Ù‡„ÏÂÌ-
ÚÓ‚, Ëϲ˘Ëı ‡ÁÏÂ˚ 2.4 Ë 1.6 ÚÔÂ Ë fl‚Îfl˛˘Ëı-
Òfl χÍÂ‡ÏË α- Ë β-ÚËÔÓ‚ ÏÚ-„ÂÌÓÏÓ‚ ͇ÚÓÙÂÎfl
ÒÓÓÚ‚ÂÚÒÚ‚ÂÌÌÓ [9, 13]. é·ÓÁ̇˜ÂÌËfl ‡ÎÎÂÎÂÈ Û͇-
Á‡ÌÌ˚ı ‚˚¯Â ÎÓÍÛÒÓ‚ Ó„‡ÌÂθÌ˚ı „ÂÌÓÏÓ‚ ͇-
ÚÓÙÂÎfl Ô˂‰ÂÌ˚ ‚ ÒÓÓÚ‚ÂÚÒÚ‚ËË Ò Í·ÒÒËÙË͇-
ˆËÂÈ ãÓÒÒÎfl [9, 13] Ë Ò ‰ÓÔÓÎÌÂÌËflÏË Òӄ·ÒÌÓ
ÄÌÚÓÌÓ‚ÓÈ Ë É‡‚ËÎÂÌÍÓ [8].
ìÒÎÓ‚Ëfl Ôӂ‰ÂÌËfl èñê ÒÓÓÚ‚ÂÚÒÚ‚Ó‚‡ÎË ÔÓ-
ÚÓÍÓÎ‡Ï ‡‚ÚÓÓ‚ – ‡Á‡·ÓÚ˜ËÍÓ‚ Ô‡ÈÏÂÓ‚ [9,
13, 20]. ê‡Á‰ÂÎÂÌË ÔÓÎÛ˜ÂÌÌ˚ı Ù‡„ÏÂÌÚÓ‚ ÔÓ-
‚Ó‰ËÎË ˝ÎÂÍÚÓÙÓÂÁÓÏ ‚ ‡„‡ÓÁÌ˚ı „ÂÎflı ‚ ·ÛÙÂ-
 íÇÖ, ‡ Ú‡ÍÊ ‚ 6% ËÎË ‚ 10% èÄÄÉ Í‡Í ‚ ‰Â̇-
ÚÛËÛ˛˘Ëı, Ú‡Í Ë ‚ ̉Â̇ÚÛËÛ˛˘Ëı ÛÒÎÓ‚Ëflı.
éˆÂÌÍÛ ‰ÓÒÚÓ‚ÂÌÓÒÚË ‡Á΢ËÈ ÏÂÊ‰Û „ÛÔ-
Ô‡ÏË ÒÓÚÓ‚ ÔÓ‚Ó‰ËÎË, Ò‡‚ÌË‚‡fl Ëı ÔÓÔ‡ÌÓ
ÔË ÔÓÏÓ˘Ë ÍËÚÂËfl tϕ [21].
êÖáìãúíÄíõ
èñê ÒÓ ÒÔˆËÙ˘Ì˚ÏË Ô‡ÈÏÂ‡ÏË ÔÓÁ‚ÓÎflÂÚ
ÓˆÂÌËÚ¸ ‚ÌÛÚ˂ˉӂÓÈ ÔÓÎËÏÓÙËÁÏ ÔÓ ÎÓÍÛÒ‡Ï
ıÎ- Ë ÏÚÑçä. Ç Ì‡ÒÚÓfl˘ÂÈ ‡·ÓÚ ËÁÏÂ̘˂ÓÒÚ¸
Ó„‡ÌÂθÌ˚ı Ñçä ·˚· ÔӇ̇ÎËÁËÓ‚‡Ì‡ Û 98
ÒÓÚÓ‚ ͇ÚÓÙÂÎfl ÔÓ ‰‚ÛÏ Ô·ÒÚˉÌ˚Ï (atpE,
trnG/trnR) Ë ‰‚ÛÏ ÏËÚÓıÓ̉ˇθÌ˚Ï (rps10, atp6)
ÎÓÍÛÒ‡Ï.
Ç ËÁÛ˜ÂÌÌÓÈ ‚˚·ÓÍ ÒÓÚÓ‚ ·˚ÎË Ó·Ì‡ÛÊÂ-
Ì˚ ÚÓθÍÓ ‰‚‡ ‡ÎÎÂÎfl Ô·ÒÚˉÌÓ„Ó ÎÓÍÛÒ‡ atpE
(ËÒÛÌÓÍ, ‡). ÅÓθ¯ËÌÒÚ‚Ó ÒÓÚÓ‚ (63 ËÁ 98) ÒÓ‰Â-
ʇÎË ‡ÎÎÂθ í, Ú.Â. ӷ·‰‡ÎË “tuberosum”-ÚËÔÓÏ
Ô·ÒÚÓχ, Ë 35 ÒÓÚÓ‚ ËÏÂÎË ‡ÎÎÂθ W, Ú.Â. ı‡‡Í-
ÚÂËÁÓ‚‡ÎËÒ¸ Ô·ÒÚÓÏÓÏ “‰ËÍÓ„Ó” ÚËÔ‡ (Ú‡·Îˈ‡).
Ç Ô·ÒÚˉÌÓÏ ÎÓÍÛÒ trnG/trnR Û ËÁÛ˜ÂÌÌ˚ı ÒÓ-
ÚÓ‚ ·˚ÎË ‚˚fl‚ÎÂÌ˚ ÚË ‡ÎÎÂÎfl (ËÒÛÌÓÍ, ·). ÅÓθ-
¯ËÌÒÚ‚Ó ÒÓÚÓ‚ (65 ËÁ 98) ÒÓ‰ÂʇÎË ‡ÎÎÂθ 6; ‡Î-
ÎÂθ 8 ·˚Π̇ȉÂÌ Û 32 ÒÓÚÓ‚ Ë ÚÓθÍÓ Û Ó‰ÌÓ„Ó ÒÓ-
Ú‡ Flourball ·˚Πӷ̇ÛÊÂÌ ‡ÎÎÂθ 7 (ÒÏ. Ú‡·ÎˈÛ).
Ç ÏËÚÓıÓ̉ˇθÌÓÏ ÎÓÍÛÒ atp6 ·˚ÎË ‚˚fl‚-
ÎÂÌ˚ ‰‚‡ ‡ÎÎÂÎfl – 1 ËÎË 0 (ËÒÛÌÓÍ, ‚). Ç ÒÎÛ˜‡Â
ӷ̇ÛÊÂÌËfl ÌÛÎÂ‚Ó„Ó ‡ÎÎÂÎfl ÓÚÒÛÚÒÚ‚Ë ÔÓ‰ÛÍ-
ÚÓ‚ ‡ÏÔÎËÙË͇ˆËË ·˚ÎÓ ÔÓ‰Ú‚ÂʉÂÌÓ ‚ ÚÂı ÌÂ-
Á‡‚ËÒËÏ˚ı ˝ÍÒÔÂËÏÂÌÚ‡ı. ÄÎÎÂθ 0 ËÏÂÎË 65
ÒÓÚÓ‚ ËÁ 98 ËÁÛ˜ÂÌÌ˚ı, ÓÒڇθÌ˚ 33 ÒÓÚ‡ ÒÓ-
‰ÂʇÎË ‡ÎÎÂθ 1 ‰‡ÌÌÓ„Ó ÎÓÍÛÒ‡.
ç‡ÍÓ̈, ÏËÚÓıÓ̉ˇθÌ˚È ÎÓÍÛÒ rps10 Û ËÒ-
ÒΉӂ‡ÌÌ˚ı ÒÓÚÓ‚ ·˚Î Ô‰ÒÚ‡‚ÎÂÌ ÚÂÏfl ‡ÎÎÂ-
ÎflÏË – α, β Ë 0 (ËÒÛÌÓÍ, „); ̇˷ÓΠ‡ÒÔÓÒÚ‡-
ÌÂÌÌÓÈ Ó͇Á‡ÎÒfl ‡ÎÎÂθ β (66 ÒÓÚÓ‚) (Ú‡·Îˈ‡).
åÂÊ‰Û „ÛÔÔ‡ÏË ÒÓÚÓ‚ ÓÚ˜ÂÒÚ‚ÂÌÌÓÈ Ë Á‡Û-
·ÂÊÌÓÈ ÒÂÎÂ͈ËË Ì ‚˚fl‚ÎÂÌÓ ‡Á΢ËÈ ÔÓ ˜‡ÒÚÓ-
Ú ‚ÒÚ˜‡ÂÏÓÒÚË ‡ÎÎÂÎÂÈ ËÁÛ˜ÂÌÌ˚ı ÎÓÍÛÒÓ‚.
Ä̇ÎËÁ ÔÓÎÛ˜ÂÌÌ˚ı ÂÁÛθڇÚÓ‚ ÔÓ͇Á‡Î, ˜ÚÓ
Ó„‡ÌÂθÌ˚ „ÂÌÓÏ˚ ÒÓÚÓ‚ ı‡‡ÍÚÂËÁÛ˛ÚÒfl
ÛÒÚÓȘ˂˚ÏË ÒÓ˜ÂÚ‡ÌËflÏË ‡ÎÎÂÎÂÈ. í‡Í, Û ÔÓ‰‡‚-
Îfl˛˘Â„Ó ·Óθ¯ËÌÒÚ‚‡ ÒÓÚÓ‚ (93 ËÁ 98 ËÁÛ˜ÂÌ-
Ì˚ı) ‚ Ô·ÒÚˉÌ˚ı ÎÓÍÛÒ‡ı atpE Ë trnG/trnR ‚˚fl‚-
ÎflÎËÒ¸ ‰‚‡ ‚‡ˇÌÚ‡ ÒÓ˜ÂÚ‡ÌËfl ‡ÎÎÂÎÂÈ: ÎË·Ó í Ë
6 (“ÍÛθÚÛÌ˚È” ÚËÔ ıÎÑçä), ÎË·Ó W Ë 8 (“‰ËÍËÈ”
ÚËÔ ıÎÑçä). Ä̇Îӄ˘ÌÓ, Û 93 ÒÓÚÓ‚ ËÁ 98 ËÁÛ˜ÂÌ-
Ì˚ı ‚ ÏÚ-ÎÓÍÛÒ‡ı atp6 Ë rps10 ‚˚fl‚ÎflÎËÒ¸ ÒÓ˜ÂÚ‡-
ÌËfl ‡ÎÎÂÎÂÈ ÎË·Ó 0 Ë β (“ÍÛθÚÛÌ˚È” ÚËÔ ÏÚÑçä),
ÎË·Ó 1 Ë α (“‰ËÍËÈ” ÚËÔ ÏÚÑçä) (Ú‡·Îˈ‡1).
ëÓÔÓÒÚ‡‚ÎÂÌË ÂÁÛθڇÚÓ‚ ‡Ì‡ÎËÁ‡ ıÎ- Ë
ÏÚÑçä ÔÓÁ‚ÓÎËÎÓ ‚˚fl‚ËÚ¸ ÒÂ‰Ë 98 ËÁÛ˜ÂÌÌ˚ı
ÒÓÚÓ‚ ‚ÓÒÂϸ „‡ÔÎÓÚËÔÓ‚, Ô˘ÂÏ ÔÓ‰‡‚Îfl˛˘ÂÂ
·Óθ¯ËÌÒÚ‚Ó ÒÓÚÓ‚ (91 ÒÓÚ, ËÎË 92.9%) ÓÚÌÓÒË-
ÎÓÒ¸ Î˯¸ Í ‰‚ÛÏ „‡ÔÎÓÚËÔ‡Ï – I Ë II (Ú‡·Îˈ‡).
ç‡Ë·ÓΠ‡ÒÔÓÒÚ‡ÌÂÌÌ˚È „‡ÔÎÓÚËÔ I, ‚˚fl‚-
ÎÂÌÌ˚È Û 62 ÒÓÚÓ‚, ı‡‡ÍÚÂËÁÓ‚‡ÎÒfl ÒÓ˜ÂÚ‡ÌË-
ÂÏ ıÎÑçä Ë ÏÚÑçä “ÍÛθÚÛÌ˚ı” ÚËÔÓ‚. Ç ÒÎÛ-
˜‡Â „‡ÔÎÓÚËÔ‡ II ÔËÒÛÚÒÚ‚Ë ıÎÑçä “‰ËÍÓ„Ó” ÚË-
Ô‡ ÒÓ‚Ô‡‰‡ÎÓ Ò Ì‡Î˘ËÂÏ “‰ËÍÓ„Ó” ÚËÔ‡ ÏÚÑçä
(Ú‡·Îˈ‡). í‡ÍËÏ Ó·‡ÁÓÏ, ‰‚‡ ÔÂӷ·‰‡˛˘Ëı
„‡ÔÎÓÚËÔ‡ I Ë II ÒÓÓÚ‚ÂÚÒÚ‚Ó‚‡ÎË “ÍÛθÚÛÌÓÏÛ”
(í/β) Ë “‰ËÍÓÏÛ” (W/α) ÚËÔ‡Ï ˆËÚÓÔ·ÁÏ [9]. ɇÔ-
ÎÓÚËÔ III (W/0) Ë „‡ÔÎÓÚËÔ˚ IV–VIII Ò ÌÂÓ·˚˜Ì˚-
ÏË ÒÓ˜ÂÚ‡ÌËflÏË ‡ÎÎÂÎÂÈ ıÎ- Ë ÏÚÑçä ·˚ÎË ‚˚fl‚-
ÎÂÌ˚ Î˯¸ Û Â‰ËÌ˘Ì˚ı ÒÓÚÓ‚: ÄÍÒ‡ÏËÚ, Hilta,
Paul Wagner, Flourball, Alaska Frostless, å‡ÎËÌӂ͇
Ë ÅflÌÒ͇fl ÌÓ‚ËÌ͇ (Ú‡·Îˈ‡).
èÓ ˜‡ÒÚÓÚ ‚ÒÚ˜‡ÂÏÓÒÚË ÓÒÌÓ‚Ì˚ı „‡ÔÎÓÚË-
ÔÓ‚ (I Ë II) ÒÂÎÂ͈ËÓÌÌ˚ ÒÓÚ‡ ÓÚ˜ÂÒÚ‚ÂÌÌÓÈ Ë
Á‡Û·ÂÊÌÓÈ ÒÂÎÂ͈ËË Ì ÓÚ΢‡ÎËÒ¸ ‰Û„ ÓÚ ‰Û-
„‡, Ӊ̇ÍÓ ÔÓˆÂÌÚ ‰ÍËı „‡ÔÎÓÚËÔÓ‚ Û ÓÚ˜Â-
ÉÖçÖíàäÄ      ÚÓÏ 43      ‹ 11      2007
 àáìóÖçàÖ ÉÖçÖíàóÖëäéÉé êÄáçééÅêÄáàü ëéêíéÇ 3

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M 1 2 3 4 5 6 7 8 9 10 11 M 1 2 3 4 5

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M 1 2 3 4 5 1 2 3 4 5 6 M
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èÓ‰ÛÍÚ˚ ‡ÏÔÎËÙË͇ˆËË Ò Ó„‡ÌÂÎÎÓÒÔˆËÙ˘Ì˚ÏË Ô‡ÈÏÂ‡ÏË Û ÒÓÚÓ‚ ‚ÓÁ‰ÂÎ˚‚‡ÂÏÓ„Ó Í‡ÚÓÙÂÎfl. ëÚÂÎ͇ÏË Ò

ˆËÙ‡ÏË Û͇Á‡Ì˚ ‡Á΢Ì˚ ‚‡ˇÌÚ˚ ‚˚fl‚ÎÂÌÌ˚ı ‡ÎÎÂÎÂÈ. ‡ – Ô‡ÈÏÂ˚ ALC_1/ALC_3 (Ô·ÒÚˉÌ˚È ÎÓÍÛÒ atpE).
ëÓÚ‡: 1 – Early Rose, 2 – Jubel, 3 – Sante, 4 – ë͇·, 5 – ã‡ÁÛËÚ, 6 – ç‚ÒÍËÈ, 7 – ä‡ÏÂ‡Á, 8 – á‡‚Ó, 9 – íÂÏÔ,
10 − ÖÎËÁ‡‚ÂÚ‡, 11 – àÒÚÓÍ. å – χÍÂ EZ100 bp, Bio-Rad. ùÎÂÍÚÓÙÓÂÁ ‚ 1.4% ‡„‡ÓÁÌÓÏ „ÂÎÂ; · – Ô‡ÈÏÂ NTCP 9
(Ô·ÒÚˉÌ˚È ÎÓÍÛÒ trnG/trnR). ëÓÚ‡: 1 – Apta, 2 – Ç˚ÚÓÍ, 3 – Flourball, 4 – Ella, 5 – ÉÓËÁÓÌÚ. å – χÍÂ EZ100 bp, Bio-
Rad. ùÎÂÍÚÓÙÓÂÁ ‚ 10% èÄÄÉ ‚ ‰Â̇ÚÛËÛ˛˘Ëı ÛÒÎÓ‚Ëflı (ÓÍ‡Ò͇ ÒÂ·ÓÏ); ‚ – Ô‡ÈÏÂ˚ AL_Mt2/ALM_3 (ÏË-
ÚÓıÓ̉ˇθÌ˚È ÎÓÍÛÒ atp6). ëÓÚ‡: 1 – Adretta, 2 – ìÚÂÌÓÍ, 3 – Epicure, 4 – ëËÏÙÓÌËfl, 5 – êÓʉÂÒÚ‚ÂÌÒÍËÈ. å – χÍÂ
EZ100 bp, Bio-Rad. ùÎÂÍÚÓÙÓÂÁ ‚ 1.4% ‡„‡ÓÁÌÓÏ „ÂÎÂ; „ – Ô‡ÈÏÂ˚ ALM_4/ALM_5 (ÏËÚÓıÓ̉ˇθÌ˚È ÎÓÍÛÒ
rps10). ëÓÚ‡: 1 – Early Rose, 2 – Provita, 3 – ãÛ„Ó‚ÒÍÓÈ, 4 – Hilta, 5 – Dorisa, 6 – ã‡ÁÛËÚ. å – χÍÂ Jetway 5000, Genomed
GmbH. ùÎÂÍÚÓÙÓÂÁ ‚ 0.8% ‡„‡ÓÁÌÓÏ „ÂÎÂ.

ÒÚ‚ÂÌÌ˚ı ÒÓÚÓ‚ ÔÓ Ò‡‚ÌÂÌ˲ Ò Á‡Û·ÂÊÌ˚ÏË
·˚Î ‚ 2 ‡Á‡ ÌËÊ (4.8 Ë 11.6% ÒÓÓÚ‚ÂÚÒÚ‚ÂÌÌÓ)
(Ú‡·Îˈ‡).
éÅëìÜÑÖçàÖ
èÓÎÛ˜ÂÌÌ˚ ÂÁÛθڇÚ˚ Û͇Á˚‚‡˛Ú ̇ ÌÂÁ̇-
˜ËÚÂθÌ˚È ÛÓ‚Â̸ ÔÓÎËÏÓÙËÁχ Ó„‡ÌÂθÌ˚ı
Ñçä Û ÒÂÎÂ͈ËÓÌÌ˚ı ÒÓÚÓ‚ ͇ÚÓÙÂÎfl. èÓ‰‡‚-
Îfl˛˘Â ˜ËÒÎÓ ÒÓÚÓ‚ (91 ËÁ 98 ËÁÛ˜ÂÌÌ˚ı, ËÎË
92.9%) ÓÚÌÓÒËÎËÒ¸ Î˯¸ Í ‰‚ÛÏ „‡ÔÎÓÚËÔ‡Ï (I Ë
II), ËÁ ÌËı 62 ÒÓÚ‡ (63.3%) ËÏÂÎË Ó‰ËÌ Ë ÚÓÚ ÊÂ
“ÍÛθÚÛÌ˚È” ÚËÔ ˆËÚÓÔ·ÁÏ˚ („‡ÔÎÓÚËÔ I).
ëÚÂÔÂ̸ „ËÒÚËÛÂÏÓ„Ó ÔÓÎËÏÓÙËÁχ Û
ÒÓÚÓ‚ ͇ÚÓÙÂÎfl, ‚˚fl‚ÎÂÌ̇fl ‚ ̇ÒÚÓfl˘ÂÈ ‡·Ó-
ÚÂ, Ó͇Á‡Î‡Ò¸ Á̇˜ËÚÂθÌÓ ÌËÊÂ, ˜ÂÏ ËÁÏÂ̘Ë-
‚ÓÒÚ¸ ÚÂı Ê ˆËÚÓÔ·ÁχÚ˘ÂÒÍËı ÎÓÍÛÒÓ‚, ӷ̇-
ÛÊÂÌ̇fl ‡ÌÂÂ Û ‚ˉӂ Ó‰‡ Solanum L. [8]. í‡Í,
ÂÒÎË Û ÒÓÚÓ‚ ͇ÚÓÙÂÎfl ‚˚fl‚ÎÂÌ˚ ÚÓθÍÓ ÚË
‡ÎÎÂÎfl Ô·ÒÚˉÌÓ„Ó ÎÓÍÛÒ‡ trnG/trnR (6, 7, 8), ÚÓ Û
ËÁÛ˜ÂÌÌ˚ı ‚ˉӂ Ó‰‡ Solanum L. ӷ̇ÛÊÂÌ˚ ‚Ó-
ÒÂϸ ‡ÎÎÂÎÂÈ ˝ÚÓ„Ó ÎÓÍÛÒ‡ (1–8) [8]. Ä̇Îӄ˘ÌÓ,
Û ÒÓÚÓ‚ ͇ÚÓÙÂÎfl ·˚ÎË ‚˚fl‚ÎÂÌ˚ ÚÓθÍÓ ‰‚‡
‡ÎÎÂÎfl Ô·ÒÚˉÌÓ„Ó ÎÓÍÛÒ‡ atpE (í Ë W), ‡ Û ‚ˉӂ
‰‡ÌÌ˚È ÎÓÍÛÒ ·˚Î Ô‰ÒÚ‡‚ÎÂÌ ˜ÂÚ˚¸Ïfl ‡ÎÎÂÎfl-
ÏË [8]. àÁ ‚ÓÒ¸ÏË ‡ÎÎÂÎÂÈ ÏËÚÓıÓ̉ˇθÌÓ„Ó ÎÓ-
ÍÛÒ‡ rps10, ‚˚fl‚ÎÂÌÌ˚ı ̇ÏË Û ‚ˉӂ Ó‰‡
Solanum L. [8], Û ÒÓÚÓ‚ ·˚ÎË Ó·Ì‡ÛÊÂÌ˚ ÚÓθÍÓ
ÚË ‡ÎÎÂÎfl (α, β Ë 0). ç‡ÍÓ̈, Û ÒÓÚÓ‚ ÔËÒÛÚ-
ÒÚ‚Ó‚‡ÎË ‰‚ ‡ÎÎÂÎfl ÎÓÍÛÒ‡ atp6 ÏÚÑçä (0 Ë 1), ‚
ÚÓ ‚ÂÏfl Í‡Í Û ËÁÛ˜ÂÌÌ˚ı ‚ˉӂ – ÚË ‡ÎÎÂÎfl (0, 1
Ë 2) [8]. çÂÓ·ıÓ‰ËÏÓ ÓÚÏÂÚËÚ¸, ˜ÚÓ „ÛÔÔ˚ ÓÚ˜Â-
ÒÚ‚ÂÌÌ˚ı Ë Á‡Û·ÂÊÌ˚ı ÒÓÚÓ‚ ÒÚ‡ÚËÒÚ˘ÂÒÍË ÌÂ
‡Á΢‡ÎËÒ¸ ÔÓ ˜‡ÒÚÓÚ‡Ï ‚ÒÚ˜‡ÂÏÓÒÚË ‡ÎÎÂÎÂÈ
ËÁÛ˜ÂÌÌ˚ı ÎÓÍÛÒÓ‚ ıÎ- Ë ÏÚÑç: ‚ Ó·ÂËı „ÛÔÔ‡ı
ÔÂӷ·‰‡˛˘ËÏ ·˚Î „‡ÔÎÓÚËÔ I (Ú‡·Îˈ‡).
è˘ËÌ˚ ÔÂӷ·‰‡ÌËfl Û ÒÂÎÂ͈ËÓÌÌ˚ı ÒÓ-
ÚÓ‚ “ÍÛθÚÛÌÓ„Ó” ÚËÔ‡ ˆËÚÓÔ·ÁÏ˚ („‡ÔÎÓÚËÔ I)
Ó·˙flÒÌflÂÚ ‡Ì‡ÎËÁ Ëı Ó‰ÓÒÎÓ‚Ì˚ı [6]. ä‡Í ·˚ÎÓ
ÓÚϘÂÌÓ ‚˚¯Â, ·ÓΠ1000 ÒÂÎÂ͈ËÓÌÌ˚ı ÒÓÚÓ‚
ÔÓËÒıÓ‰flÚ ÓÚ ÒÓÚ‡ Early Rose, fl‚Îfl˛˘Â„ÓÒfl ÒÂ-
fl̈ÂÏ ÒÓÚ‡ Garnet Chili, ÔÓËÁ‚Ó‰ÌÓ„Ó ÒÓÚ‡
Rough Purple Chili – Ô‰ÔÓÎÓÊËÚÂθÌÓ ˜ËÎËÈÒÍÓ-
„Ó ‡·ÓË„ÂÌÌÓ„Ó Ó·‡Áˆ‡ (ËÌÚÓ‰Û͈Ëfl ÉÛ‰˘‡
1853 „.), Ëϲ˘Â„Ó “ÍÛθÚÛÌ˚È” ÚËÔ ˆËÚÓÔ·Á-
Ï˚. ùÚÓÚ Ê ÚËÔ ˆËÚÓÔ·ÁÏ˚ ËÏÂÂÚ ÔÓ‰‡‚Îfl˛˘ÂÂ
˜ËÒÎÓ ˜ËÎËÈÒÍËı ‡·ÓË„ÂÌÌ˚ı ÒÓÚÓ‚ S. tuberos-
um subsp. tuberosum [10, 15, 22].
àÁÛ˜ÂÌË ˆËÔÓÍÌ˚ı „Ë·ˉӂ, ÔÓÎÛ˜ÂÌÌ˚ı
‚ ÏÂÊ- Ë ‚ÌÛÚ˂ˉӂ˚ı (c S. tuberosum subsp. an-
digena) ÒÍ¢˂‡ÌËflı ˜ËÎËÈÒÍËı Ó·‡ÁˆÓ‚, ‚˚-
fl‚ËÎÓ ÔÓÎÓÊËÚÂθÌÓ ‚ÎËflÌË “ÍÛθÚÛÌÓ„Ó” ÚË-
Ô‡ ˆËÚÓÔ·ÁÏ˚ ̇ ÛÓʇÈÌÓÒÚ¸ ‡ÒÚÂÌËÈ, Ò Ó‰ÌÓÈ

ÉÖçÖíàäÄ      ÚÓÏ 43      ‹ 11      2007

4 ɇ‚ËÎÂÌÍÓ Ë ‰.

ê‡ÒÔ‰ÂÎÂÌË ÒÓÚÓ‚ ͇ÚÓÙÂÎfl ‡Á΢ÌÓ„Ó ÔÓËÒıÓʉÂÌËfl ÔÓ „‡ÔÎÓÚËÔ‡Ï

ɇÔÎÓÚËÔ* ÇÒÂ„Ó (%) óËÒÎÓ (%) Á‡Û·ÂÊÌ˚ı ÒÓÚÓ‚ óËÒÎÓ (%) ÓÚ˜ÂÒÚ‚ÂÌÌ˚ı ÒÓÚÓ‚
I. í/6/β/0 62 (63.3%) 22 (62.9%) 40 (63.5%)
Agave, Baltica, Bintje, Cobbler, Deodara, De- Ä„ÓÌÓÏ˘ÂÒÍËÈ, ÄÚ·ÌÚ, ÅÓÌÌˈÍËÈ,
siree, Early Rose, Ella, Epicure, Garnet Chili, ɇژËÌÒÍËÈ, ÉÓËÁÓÌÚ, ÑÂÚÒÍÓÒÂθÒÍËÈ,
Hindenburg, Jubel, Paterson Victoria, Provita, ÖÎËÁ‡‚ÂÚ‡, ÜÛ‡‚ËÌ͇, á‡‡‚¯‡Ì, á‡Ë̇,
Reichskanzler, Quarta, Russet Burbank, Sante, á‡‚Ó, àχ̉‡, àÒÚÓÍ, ä‡ÏÂ‡Á, ä‡ÚÛÌ-
Sonate, Xenia, ëËÏÙÓÌËfl, ëËÌ˛ı‡ ÒÍËÈ, äÓÏÒÓÏÓΈ 20, ãÓı, ãӯˈÍËÈ,
ãÛ„Ó‚ÒÍÓÈ, ã˚·Ë‰¸, å‡Ú¯͇, åË·‚ˈ‡,
åÛχÌÒÍËÈ, ç‡Ó˜¸, é‰ÂÊÒÍËÈ, èÂÚÂ-
·Û„ÒÍËÈ, èӷ‰‡, èÓ‚Ëӂˆ, è‡ÏÂ̸,
èËÂÍÛθÒÍËÈ ‡ÌÌËÈ, èËÓ·ÒÍËÈ, ꇉۄ‡,
ë‚Â̇fl êÓÁ‡, ëËÌ„·Á͇, ëÛÁÓ¸Â,
í‡ÎËÒχÌ, íÂÏÔ, îËÚÓÙÚÓÓÛÒÚÓȘ˂˚È,
ùÌÂ„Ëfl, üÌÚ‡Ì˚È
II. W/8/α/1 29 (29.6%) 9 (25.7%) 20 (31.8%)
Adretta, Apta, Carla, Delikat, Dorisa, Galina, ÄθÔËÌËÒÚ, ÄË̇, ÇÂÒÂÎÓ‚ÒÍËÈ 2–4, ÇÂÒ̇
Karlena, Rasant, Turbella Å·fl, Ç˚ÚÓÍ, ÑÂθÙËÌ, ÑË̇, ܇‚ÓÓÌÓÍ,
ÜÛÍÓ‚ÒÍËÈ ‡ÌÌËÈ, áÓ‚, ã‡ÁÛËÚ, ã‡ÒÛ̇Í,
å‡Ò, ç‚ÒÍËÈ, êÓʉÂÒÚ‚ÂÌÒÍËÈ, êÛÏflÌ͇,
ë‚ËÚ‡ÌÓÍ äË‚ÒÍËÈ, ë͇·, ìÚÂÌÓÍ, ó‡Ó-
‰ÂÈ
III. W/8/0/1 1 (1.0%) 0 1 (1.6%)
ÄÍÒ‡ÏËÚ
IV. W/6/0/1 1 (1.0%) 1 (2.9%) 0
Hilta
V. W/6/β/0 2 (2.0%) 0 2 (3.2%)
ÅflÌÒ͇fl ÌÓ‚ËÌ͇, å‡ÎËÌӂ͇
VI. W/8/α/0 1 (1.0%) 1 (2.9%) 0
Paul Wagner
VII. W/7/β/1 1 (1.0%) 1 (2.9%) 0
Flourball
VIII. í/8/β/1 1 (1.0%) 1 (2.9%) 0
Alaska Frostless
* ì͇Á‡Ì˚ ‡ÎÎÂÎË, ‚˚fl‚ÎÂÌÌ˚ ÒÓÓÚ‚ÂÚÒÚ‚ÂÌÌÓ ‚ ÎÓÍÛÒ‡ı atpE/trnGtrnR/rps10/atp6.

ÒÚÓÓÌ˚ [23, 24], Ë Ë̉ÛÍˆË˛ Ô˚θˆÂ‚ÓÈ ÒÚÂËθ-
ÌÓÒÚË Û ÒÙÓÏËÓ‚‡‚¯ËıÒfl „Ë·ˉӂ, Ò ‰Û„ÓÈ
[25, 26]. ùÚË Í‡˜ÂÒÚ‚‡ ˜ËÎËÈÒÍË ‡·ÓË„ÂÌÌ˚Â
ÒÓÚ‡ S. tuberosum subsp. tuberosum, Ú‡Í ÊÂ Í‡Í Ë
ÒÓÚ Early Rose, ÔÂ‰‡‚‡ÎË ‚ÒÂÏ Ò‚ÓËÏ ÔÓÚÓÏ͇Ï
ÔÓ Ï‡ÚÂËÌÒÍÓÈ ÎËÌËË [25, 26]. èÓ˝ÚÓÏÛ ÔË ÒÓ-
Á‰‡ÌËË ‚˚ÒÓÍÓÛÓʇÈÌ˚ı ÒÓÚÓ‚ Ô‰ÔÓ˜ÚËÚÂθ-
ÌÓ ·˚ÎÓ ËÒÔÓθÁÓ‚‡Ú¸ Early Rose ‚ ͇˜ÂÒڂ χÚÂ-
ËÌÒÍÓÈ ÙÓÏ˚, ‡ ÔÓÎÛ˜ÂÌÌ˚ ‚ ÒÍ¢˂‡ÌËflı Ò
Early Rose ‡ÒÚÂÌËfl ӷ·‰‡ÎË ÒÚÂËθÌÓÈ Ô˚θ-
ˆÓÈ Ë ÔÓ˝ÚÓÏÛ ÏÓ„ÎË ‚ ‰‡Î¸ÌÂȯÂÏ ËÒÔÓθÁÓ‚‡Ú¸-
Òfl ÚÓθÍÓ ‚ ͇˜ÂÒÚ‚Â ÊÂÌÒÍÓ„Ó Ó‰ËÚÂÎfl. í‡ÍËÏ
Ó·‡ÁÓÏ, ˝ÙÙÂÍÚ˚ “ÍÛθÚÛÌÓ„Ó” ÚËÔ‡ ˆËÚÓÔ·Á-
Ï˚ ̇ ÛÓʇÈÌÓÒÚ¸ ‡ÒÚÂÌËÈ Ë ÒÚÂËθÌÓÒÚ¸
Ô˚θˆ˚ „Ë·ˉӂ fl‚ËÎËÒ¸ Ô˘ËÌÓÈ ¯ËÓÍÓ„Ó
‡ÒÔÓÒÚ‡ÌÂÌËfl ‰‡ÌÌÓ„Ó ÚËÔ‡ ˆËÚÓÔ·ÁÏ˚ Ò‰Ë
ÒÂÎÂ͈ËÓÌÌ˚ı ÒÓÚÓ‚ ͇ÚÓÙÂÎfl.
ê‰ÍË Ê „‡ÔÎÓÚËÔ˚ ÒÚ‡Ó‰‡‚ÌËı ÒÓÚÓ‚ Ò‚fl-
Á‡Ì˚ Ò ‰Û„ËÏË ÌÂÁ‡‚ËÒËÏ˚ÏË ËÌÚÓ‰Û͈ËflÏË ÌÂ-
Ó‰ÒÚ‚ÂÌÌ˚ı Ó·‡ÁˆÓ‚ ˛ÊÌÓ‡ÏÂË͇ÌÒÍÓ„Ó Í‡-
ÚÓÙÂÎfl. ç‡ÔËÏÂ, ÒÓÚ Paul Wagner („‡ÔÎÓÚËÔ
VI) ÔÓÎÛ˜ËÎ ˆËÚÓÔ·ÁÏÛ ÔÓ Ï‡ÚÂËÌÒÍÓÈ ÎËÌËË ÓÚ
ÒÓÚ‡ Industrie, ‚ÓÒıÓ‰fl˘Â„Ó ÔÓ Ï‡ÚÂËÌÒÍÓÈ ÎË-
ÌËË Í Erste von Nassengrund – ÒÓÚÛ ÒÚ‡ÓÈ ÍÓÌÚË-
ÌÂÌڇθÌÓ-‚ÓÔÂÈÒÍÓÈ „ÛÔÔ˚, ·ÂÛ˘ÂÈ Ò‚Ó ̇-
˜‡ÎÓ ÓÚ S. tuberosum subsp. andigena ËÁ ÔÂ‚˚ı ËÒ-
Ô‡ÌÒÍËı ËÌÚÓ‰Û͈ËÈ 1580–1600 „„. [6]. ëÚ‡˚È
ÒÓÚ Flourball, Ëϲ˘ËÈ „‡ÔÎÓÚËÔ VII, ·ÎËÁÓÍ Í
„ÛÔÔ Daber – ·ÓΠ‡ÌÌÂÈ ËÌÚÓ‰Û͈ËË 1830-ı
„Ó‰Ó‚. чÌÌ˚È ÒÓÚ ÙÂÚËÎÂÌ Ë ËÒÔÓθÁÓ‚‡ÎÒfl ‚
ÒÍ¢˂‡ÌËflı ÔÂËÏÛ˘ÂÒÚ‚ÂÌÌÓ Ò ÓÚˆÓ‚ÒÍÓÈ ÒÚÓ-
ÓÌ˚ [6], ÔÓ˝ÚÓÏÛ ÒÂ‰Ë ÒÓ‚ÂÏÂÌÌÓ„Ó ÒÓÚËÏÂÌ-
Ú‡ Â„Ó ˆËÚÓÔ·Áχ Ì ÔÓÎۘ˷ ¯ËÓÍÓ„Ó ‡ÒÔÓ-
ÒÚ‡ÌÂÌËfl.

ÉÖçÖíàäÄ      ÚÓÏ 43      ‹ 11      2007

 àáìóÖçàÖ ÉÖçÖíàóÖëäéÉé êÄáçééÅêÄáàü ëéêíéÇ
Ä̇ÎËÁ Ó‰ÓÒÎÓ‚Ì˚ı ÒÓÚÓ‚ Ò “‰ËÍËÏË” ÚËÔ‡-
ÏË ˆËÚÓÔ·ÁÏ („‡ÔÎÓÚËÔ˚ II Ë III) Ë ‰ÍËı „‡ÔÎÓ-
ÚËÔÓ‚ (IV–VIII) ÔÓ͇Á‡Î, ˜ÚÓ ÔÓ Ï‡ÚÂËÌÒÍÓÈ ÎË-
ÌËË Ú‡ÍË ÒÓÚ‡ Ó·˚˜ÌÓ ‚ÓÒıÓ‰flÚ Í „Ë·ˉ‡Ï 12.
S. tuberosum subsp. tuberosum Ò Ó‰ÌËÏ ËÎË ÌÂÒÍÓθ-
ÍËÏË ÍÛθÚÛÌ˚ÏË Ë/ËÎË ‰ËÍËÏË ‚ˉ‡ÏË: S. tubero-
sum subsp. andigena (Paul Wagner), S. acaule (Alaska 13.
Frostless), S. demissum (ÇÂÒÂÎÓ‚ÒÍËÈ 2–4, ç‚ÒÍËÈ,
ÄË̇, ÅflÌÒ͇fl ÌÓ‚ËÌ͇) Ë fl‰ÓÏ ‰Û„Ëı. ÅÓθ-
¯ËÌÒÚ‚Ó ÒÓÚÓ‚ Ò “‰ËÍËÏ” ÚËÔÓÏ ˆËÚÓÔ·ÁÏ˚ ·˚- 14.
ÎË ÒÓÁ‰‡Ì˚ Á‡ ÔÓÒΉÌË 40 ÎÂÚ, ÍÓ„‰‡ ‚ ÒÂÎÂ͈ËË
͇ÚÓÙÂÎfl Òڇ· ¯ËÓÍÓ ËÒÔÓθÁÓ‚‡Ú¸Òfl ÏÂÊ‚Ë-
‰Ó‚‡fl „Ë·ˉËÁ‡ˆËfl. é‰Ì‡ÍÓ, Í‡Í ÛÊ ÓÚϘ‡ÎÓÒ¸
‚˚¯Â, ˜ËÒÎÓ ‰ËÍËı ‚ˉӂ, ÒÔÓÒÓ·Ì˚ı ÒÍ¢˂‡Ú¸- 15.
Òfl Ò ‚ÓÁ‰ÂÎ˚‚‡ÂÏ˚Ï Í‡ÚÓÙÂÎÂÏ, ˜ÂÁ‚˚˜‡ÈÌÓ
Ó„‡Ì˘ÂÌÓ ËÁ-Á‡ ·‡¸ÂÓ‚ ÌÂÒÓ‚ÏÂÒÚËÏÓÒÚË. ê‡Ò-
¯ËÂÌ˲ „ÂÌÂÚ˘ÂÒÍÓ„Ó ‡ÁÌÓÓ·‡ÁËfl ÒÓÚÓ‚ 16.
͇ÚÓÙÂÎfl, ‚ ÚÓÏ ˜ËÒÎÂ Ë ÔÓ ˆËÚÓÔ·ÁχÚ˘ÂÒÍËÏ
Ù‡ÍÚÓ‡Ï, ÏÓ„ÛÚ ÒÔÓÒÓ·ÒÚ‚Ó‚‡Ú¸ ÒÓ‚ÂÏÂÌÌ˚Â
ÏÂÚÓ‰˚ ·ËÓÚÂıÌÓÎÓ„ËË, ‚ ˜‡ÒÚÌÓÒÚË, ÏÂÚÓ‰˚ ÒÓ- 17.
χÚ˘ÂÒÍÓÈ „Ë·ˉËÁ‡ˆËË ‡ÒÚÂÌËÈ [27].
ꇷÓÚ‡ ‚˚ÔÓÎÌÂ̇ ÔË ÔÓ‰‰ÂÊÍ êîîà
(„‡ÌÚ ‹ 06-04-08352 ).
18.
ëèàëéä ãàíÖêÄíìêõ
1. Hawkes J.G. The Potato, Evolution, Biodiversity and 19.
Genetic Resources. L.: Belhaven, 1990. 260 p.
2. Davidse L.C., Henken J., van Dalen A. et al. Nine years
of practical experience with phenylamide resistance in 20.
the Netherlands // Netherlands J. Plant Pathol. 1989.
V. 95. P. 197–213.
3. Mendoza H.A., Haynes F.L. Genetic relationship among
potato cultivars grown in the United States // Hort- 21.
science. 1974. V. 9. P. 328–330.
4. Ross H. Potato Breeding – problems and perspectives. 22.
Berlin: Paul Parey, 1986. 132 p.
5. Salaman R.N. The history and the social influence of the
potato // Revised impression / Ed. Hawkes J.G. Cam-
bridge (U.K): Cambridge Univ. Press, 1985.
6. äÓÒÚË̇ ã.à. Ç˚‰ÂÎÂÌË ËÒıÓ‰ÌÓ„Ó Ï‡ÚÂˇ·
‰Îfl ÒÂÎÂ͈ËË Í‡ÚÓÙÂÎfl ̇ ÓÒÌÓ‚Â „Â̇ÎÓ„ËË.
ëè·: àÁ‰-‚Ó Çàê, 1992. 106 Ò.
7. ÅÛ‰ËÌ ä. á., ɇ‚ËÎÂÌÍÓ í. Ä. ÉÂÌÂÚ˘ÂÒÍË ÓÒÌÓ- 23.
‚˚ ÓÚ‰‡ÎÂÌÌÓÈ „Ë·ˉËÁ‡ˆËË Í‡ÚÓÙÂÎfl // ÉÂÌÂÚË-
͇. 1994. í. 30. ‹ 10. ë. 1413–1422. (Budin K.,
Gavrilenko T. Genetic basis of remote hybridization in
potato (Review) // Rus. J. Genetics. 1994. V. 30. ‹ 10. 24.
P. 1225–1233.
8. ÄÌÚÓÌÓ‚‡ é.û., ɇ‚ËÎÂÌÍÓ í.Ä. èÓÎËÏÓÙËÁÏ
ÔÓÒΉӂ‡ÚÂθÌÓÒÚÂÈ Ó„‡ÌÂθÌ˚ı Ñçä ‚Ë‰Ó‚
͇ÚÓÙÂÎfl // ùÍÓÎӄ˘ÂÒ͇fl „ÂÌÂÚË͇. 2006. í. IV. 25.
ë. 3–10.
9. Lossl A., Gotz M., Braun A., Wenzel G. Molecular markers
for cytoplasm in potato: male sterility and contribution of 26.
different plastid-mitochondrial configurations to starch
production // Euphytica. 2000. V. 116. P. 221–230.
10. Sukhotu T., Kamijima O., Hosaka K. Nuclear and chlo- 27.
roplast DNA differentiation in Andean potatoes // Ge-
nome. 2004. V. 47. P. 46–56.
11. Sukhotu T., Kamijima O., Hosaka K. Genetic diversity of
the Andean tetraploid cultivated potato (Solanum
ÉÖçÖíàäÄ      ÚÓÏ 43      ‹ 11      2007
5
tuberosum L. subsp. andigena Hawkes) evaluated by
chloroplast and nuclear DNA markers // Genome. 2005.
V. 48. P. 55–64.
Hosaka K. Who is the mother of the potato? – restriction
endonuclease analysis of chloroplast DNA of cultivated
potatoes // Theor. Appl. Genet. 1986. V. 72. P. 606–618.
Lossl A., Adler N., Horn R. et al. Chondriome-type char-
acterization of potato: mt α, β, γ, ε and novel plastid-mi-
tochondrial configurations in somatic hybrids // Theor.
Appl. Genet. 1999. V. 98. P. 1–10.
Hosaka K. T-type chloroplast DNA in Solanum tubero-
sum L. ssp. tuberosum was conferred from some popula-
tions of S. tarijense Hawkes // Amer. J. Potato Res.
2003. V. 80. P. 21–32.
Hosaka K., Hanneman R.E. The origin of the cultivated
potato based on chloroplast DNA // Theor. Appl. Genet.
1988. V. 76. P. 172–176.
Powell W., Baird E., Duncan N., Waugh R. Chloroplast
DNA variability in old and recently introduced potato
cultivars // Ann. Appl. Biol. 1993. V. 123. P. 403–410.
Prowan J., Powell W., Dewar H. et al. An extreme cyto-
plasmic bottleneck in the modern European cultivated
potato (Solanum tuberosum) is not reflected in decreased
levels of nuclear diversity // Proc. R. Soc. Lond. B. 1999.
V. 266. P. 633–639.
Waugh R., Glendinning D.R., Dunkan N., Powell W.
Chloroplast DNA variation in European potato culti-
vars// Potato Res. 1990. V. 33. P. 505–513.
Wienand U., Feix G. Zein specific restriction enzyme
fragments of maize DNA // FEBS Letters. 1980. V. 116.
P. 14–16.
Bryan G.J., McNicoll J., Ramsay G. et al. Polymorphic
simple sequences repeat markers in chloroplast genomes
of Solanaceous plants // Theor. Appl. Genet. 1999.
V. 99. P. 859–867.
ã‡ÍËÌ É.î. ÅËÓÏÂÚËfl. å.: Ç˚Ò¯. ¯ÍÓ·, 1980.
294 Ò.
ò‚‡˜ÍÓ ç.Ä., ÄÌÚÓÌÓ‚‡ é.û., äÓÒÚË̇ ã.à., ɇ‚-
ËÎÂÌÍÓ í.Ä. èÓÎËÏÓÙËÁÏ fl‰ÂÌÓÈ Ë ˆËÚÓÔ·Á-
χÚ˘ÂÒÍËı Ñçä Û ‡·ÓË„ÂÌÌ˚ı ˜ËÎËÈÒÍËı ÒÓÚÓ‚
S. tuberosum ssp. tuberosum // ëÓ‚ÂÏÂÌÌÓÂ ÒÓÒÚÓfl-
ÌËÂ Ë ÔÂÒÔÂÍÚË‚˚ ‡Á‚ËÚËfl ÒÂÎÂ͈ËË Ë ÒÂÏÂÌÓ-
‚Ó‰ÒÚ‚‡ ͇ÚÓÙÂÎfl (Í 75-ÎÂÚ˲ Ççààäï ËÏ. Ä.É.
ãÓı‡). ÇÓÔÓÒ˚ ͇ÚÓÙÂ΂ӉÒÚ‚‡. å.: àÁ‰-‚Ó
Ççààäï, 2006. ë. 205–211.
Maris B. Analysis of an incomplete diallele cross among
three ssp. tuberosum varieties and seven long-day adapt-
ed ssp.andigena clones of the potato // Euphytica. 1989.
V. 41. P.163–182.
Plaisted R.L. Utilization of germplasm in breeding pro-
grammes – use of cultivated tetraploids// Prospect for the
potato in the developing worlds. Lima (Peru): Intern. Po-
tato Centre, 1972. 23 p.
Grun P. Cytoplasmic sterilities that separate the group
Tuberosum cultivated potato from its putative tetraploid
ancestor // Evolution. 1973. V. 27. P. 633–643.
Grun P., Ochoa C., Capage D. Evolution of cytoplasmic
factors in tetraploid cultivated potatoes // Amer. J. Bot.
1977. V. 64 P. 412–420.
ÄÌÚÓÌÓ‚‡ é.û., ɇ‚ËÎÂÌÍÓ í.Ä. ü‰ÂÌÓ-ˆËÚÓ-
Ô·ÁχÚ˘ÂÒÍË ‚Á‡ËÏÓÓÚÌÓ¯ÂÌËfl Û ÒÓχÚ˘ÂÒÍËı
„Ë·ˉӂ Ó‰‡ Solanum L. // ëÚÛÍÚÛ‡ Ë ˝ÍÒÔÂÒÒËfl
ÏËÚÓıÓ̉ˇθÌÓ„Ó „ÂÌÓχ ‡ÒÚÂÌËÈ. àÍÛÚÒÍ,
2006. ë. 5–10.
6 ɇ‚ËÎÂÌÍÓ Ë ‰.

Abstract

ÉÖçÖíàäÄ      ÚÓÏ 43      ‹ 11      2007