150 PALAEONTOLOGy
declined condition, ¢.g., D. nicolsoni. Deflexed condition showed further
divergence of the stipes from each other e.g., Leptograptus. Maximum
divergence was in D. hirundo in which the two stipes were horizontal. From
this point, convergence of the stipes started and the stipes came towards each,
other and theca were in opposite directions, the first stage of convergence ig
known as reflexed condition, ¢.g., D. patuius. D. gibberulus exhibits the second
Stage of convergence, the reclined condition. Maximum convergence Tesulted
in the union of stipes to give rise to biserial colony and scandent condition in
Which theca face upwards, e.g..Diplograptus (Upper Ordovician). The changes
‘were rapid and resulted in the extinction of the class Graptozoa (fig. 15.4).
Geological Distribution
The graptozoans were exclusively marine and bottom-living. Their fossils
are found in carbonaceous shales and clayey rocks. Their geological history is
‘very short, while their geographical distribution is very wide. They exhibited
rapid evolutionary changes and therefore, they are good index fossils.
Simple dendroid forms were in Middle Cambrian, ‘Their number increased
in Ordovician and Silurian. From Silurian onwards, their number reduced and
by the carly Mississipian they become extinct. Typical graptolites appeared in
Upper Cambrian e.g., Bryograptus, Dicranograptus, Dictyonema. During Lower
Ordovician times, horizontal forms appeared e.g. Dichograptus, along with
pendent forms like D, bifids and reclined forms and scandent forms
(Glyptograptus and Nemagraptus|. Upper Ordovician times contain
Climatograptus,, Mesograptus, Pleurograptus and Dicellograptus. Silurian times
Were marked by Orthograptus, Cephalograptus and Monograptus. "This period
was long marked by hooked, lobate and isolated theca. Middle Silurian beds
Some Important Fossils
@ Monograptus—Straight, unbranched, uniserial rhabdosome ; Theca are
Cbliquely and are of various shapes, closely placed, ilu .
15.1 uniserial form), neve “phos Agen. Cel
G2) Diplograptus—Two rows of close! i
(i I T so! 'Y spaced, oblique curved theca attached
to single stipe. Age—Mid Ordovician to Lower Silurian (fig. 15.1 biserial form).
a? Pidymograptus—Rhabdosome bilaterally Symmetrical with two stipes
verging upto an angle of 180°, giving fork like appearance, small sicula,
Cylindrical theca direct obliquely, Age—Ordovician to Silurian
() Bryograptus—Bilaterally symmetrical, uniserial wi in sti
diverging at an angle from the ae cabana Wich too main’ stipes
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