cla Jy Glee xe Trichoderma
owe 201610 suis 1 8474
Trichoderma Gish ce 1794 tm civil cy hill le 4 Person . Uti LSBisset,1991a) Jb ena cis pall
tac ysel sl Gastéromycates .
jon Lancia, EG Ud Gad II, pall 0 3 jaiS Le pee I diudall al) GUIS) oe cisTrichoderma Che }Y! Gs aul
Wa Bae anki 5 98h ygab igh ge yal Lt le a yl ly gp gpl Le ball Jaa aang LS Mal yg al yally
AB AAS (inl Das ys Ga edt a oh ( Kubicek et Harman, 2002).
asd TrichodermaStal gl eet Trichoderma ame pele CISA pills partaall sot Cue op sh Global Jal Qalke diy le eat on
SmI al hy AS GANT ANN Gh patna gS LS Gal ple a Up pal glad nas LS 5 yes
bao pAl Ay yila de glad JSS ga bbe il sip cp sdleall 4 all el ja) le asd),
BS aa a gl gly phe La Sl al pall colada Of op CS Lee Lae Co jn Chien a LS yo pall pga Cad pall gly
a aye Dead ly SIS Lia lb GlalLall Jas (Landreau, 2001) (1082).
hb § pariee 1USAT viride
iy ce PDA ks
7253 310%(
Samuels,1996 )
igual Trichoderma
ett, 2004 ) ANS oS phil gh ainadll ee dt
hement : Amastigomycotaoranchement : Ascomycotina
ordariomycetes
ypocréales
aypocreaceae
ypocrea mitosporique Trichoderma
iy! ow sl! Trichoderma
ichoderrma el sl Laugh 55 desuall 85 85: a yp GLa ay J Calis Qs lilll le 45 iy (Esposito et
3), aban a y AUS AL ley CALA Gs US le La gay AISA yall GIES Jnl le Sy gy pl pall
cee Gila 85 Sy halls 85 Cah ada, ye ( Roquebert , 1996 ).
LoS yas tse Trichoderma 2224 pd i sy Assy 4 hl el IE on Ge es
jillus sPenicilliumesss Sey eu= Trichoderma 4% pail) tin dU Sil soa 6 % 6 ot gel sl dan ld
u, 2001 ).
ui T. viride
T. odie es) 8 Trichoderma yj sil padi! iy ye Ge pip Y Aas jhe atl pall ett (Jaklisch etal. , 2006).
NAB ja al ellipsis Ay jl) Sal bd Hal CALS Soe Lae yay Aelia ly hall Yo 9 LEG IS
408 gAlBisset,1991a) ). 4 8) lee snw Tviride sT.ligronum, 4S! tug Ll parfaite dis hil
vaurantiaca, Hypocrea coprosma, Hypocrea rufer
valbo-medullosa, Hypocrea atrogelatinosaDomsch et al.,1993).
‘ST. viride
ride .T tens) 4550) Lyi AAD aye ARAM «Mada Lay Ge lp) AS Ga Ale fH aT ak
(il pabedal o yl) Jue All cnt} all Domsch etal.,1993)).
aslduxT. viride
Thalle 4 Trichoderma os =!
5 US ayn AS yal yl pg Lynd Lgl ce
SSE See He pia Hy SN Sal gall G5 5 Se 4,5 — 3.6 Gan la phat aes hy es S55 SY obs eal jell 4 USE esl
aes LS AB Riga i Reet gad lath) Lo ssc GS le AS pal UGS Ga oll el yaad Mel pls pues Jal gall 9s jac
AN pannel Gb RLgS shay elude Cy 5S Gye Kids Land's ( Samson et al., 1994) (2 Js).
phil 4a pL yall yatlasl:2US¢ T. viride Kananan et al., 2012).
(A435 0 fl pall gosh) all La pleas Ba Sp pl)
sue T. virideWall pss 187. viride Ginny gd %¢ (32 ~ 6) ce 81a Shas le alk 5 we iD get Qual «9 (28 — 20) ce
Sola Re yale cll lA A, NA 990 ale yall Ge Sy 0,37 Lada le yay SAAT CYL
Cape Phage) yall Aba,
Yoda 0h» PH shail sal), viride 5 9-3 ow pH - 4.5 ow dus!
‘manian, 1983)).
5 9T. viride
Viride ogi pashan nib ofall Cait dls Gy la i JSG ge 5D a pl a9 SD al yall JUG tye Yuin SS
alge ISAS vie isa clin o 4 Si (Rifai, 2004),
ANI Gletid T. viride
Viride \pie YI hati yo 8 pies AS 5) a) le Aish
ase,endo 13émodine ,cellulase,cellobiase ,amylase gliotoxine, polypeptide, protease, endo
“olase,exo-gluconase B-glucosidase, trichothécénes, trichodermine (xiaoyil et al., 2012).
35 T. viridestail te
1) Ga) Antibiose
$e Se Trichoderma ‘yall AAS le Lyte yt Ae hh ge LL US A ppedly gall Ll etal Ge
hs 89 (Johanne, 2002),
vet al (2009 ) ab jc Katia 5 bl LS pel ctl Hae Gal BT, vivide y
cull L-hydroxy-3-
thraginone ys Al A saya plea peal ly ll Ge yall ls Ge ad IGS I SA
cénes strichodermine sla ut sits of Jaimall ya asta a G8 Uatilly Le pS y cially sa pt ym le
32 hun twill Elena, 2004).a La compeétition
5B gb oa iy go Sl nll Gila poh pal
(Sat) ject A aay gall CS Ope Gud Cay
CASI Si ie gale: Jpn! piety Jill y asa si allty Joana Re juss jualinll US UKE! (Bolton et al.,1993).
BV sels T, viride Cun gal hugh yn yl Sg Lal Jute CL a yal Cy ily
roderma ted ot pl it tea IS Jey J Sal Ja! Jal ge da ba (Johanne, 2002). gs a oa
aS ce Sis Le 18 cs il cla le Bape pall ce yall iy hall ony gat GS le Lecall jal coat «lil
viride Sis tae joa Shy shill Ge sel) go pda gle Alla) 44 xis: Verticillium, Rhizoctonia, Pythium,
3....@l! Samson et al.), 1994),
bit) Jaki] Mycoparasitisme
Fichoderma ¢ ca jaa) pail Jy pail (in Gti) nay Ln y a pall Jabal pba gay Lon aca yall Cy gill le
richoderma 53} SLaily JJ Jabal a (Johanne, 2002), i rb vg sail ial p jilSe Gus
rderma ah Gibsons oS:
Nita:
PP. Ge LT 5 phall 0 Trichoderma 4imslyal Lhd) fab las le Cas CLS ye 5 8 gil Gin yal pall oe
limiotropisme 4:3 cS) GY) aa} Gi yx ol jini Se Lala lal ga 9 a5 plan die Chan da Gus Gi)
a: jail yeBday:
pall jhill Ge Trichoderma sos suLectines . bs)! yb ce Gaal!) cy Cus agglutinine v2 jaa hill
shall piesa dl Ch Sal) Gwe ee Trichoderma.
br ajuda
Aaya:
Sey Lan sll ell ghby oro yaa) phil Glia Gye Gilly! Ces Trichoderma bel isl sl Libel cli yb ge
3] Se Shy 539! Gili 5 pit Lytique (>%ecellulase « 548 chutinases 5u% glucanase J&s pectinase
sana gliotassine cy sail Liki psi Jal ts 55 Chet,1987)). cw lS Inbar et Chet,1992)) ataty' of
eal Gi jlall p jt ilall Clie Ja lectine 41s) G22 ye Ls 3 sl agglutinine + Sclerotina rolfsii, jx
hoderma jhi dealye ole lei sé Sclérotina rolfsiis se Gxt yND-glucose »D-manose ce tte as
dhall fas oi CL Su Trichoderma
ThcA tay)
Jb sie og phil! Judai Jal x : 3 J&4 Trichoderma
bi on yg shill Jikitl -4US:Trichoderma » Rhizoctonia solani
! Lantagonisme
ok Gx Lice yaall Gil pall bl ja lil Jat A Sic mycostase «Sb plaill pylue las 5 pla Ce el se mycolyse))
xiuall dus J bactériolyse)) Curl et Truelove,1986)).
a Gatliad quia 55 Chet et al,,1997)), o« JS a8 1971 4 SUI Webster et Dennis atid aun sy
DEE JAE le aay g al GaN agg sll Ja 3 Ji Lal Lajas
35 Gals Trichodermats jad! Gl phil Go yyS axe wee saa le 448 (Comporta, 1985) . Sasi cay LS.
988) CY Gay ol Trichoderma Cistall (aay sai joins Llbad soles he 3 psi YS,het al 1991) o26 Trichoderma ot so8! Sem ce ek CL pi juin AS cn peal abel Gt Gi pall pi le
Ae Ale 3 8 Lill (Fusarium: Pythium _ Rhizoctonia.
vy JS) Trichoderma &
ichoderma flee ysis sal ys¥! taj) Lill aan se gl 4gs ay ye CLR! Ge Gy
serra shal! Aascil JSlo dace paall ily all J hs ally Gund «shall Jom Gils Jala ps5 3,851 obs eas Harman,
Trichoderma % sia) Wal 4 sti) aL I
{sla aac Yedidia
3).
ichoderma hiss pil gb CLS yal 038 Lh Ace peal Jal pal ns Chall a da gle Gaya ll CLS pall Ga gol RDG
(Ce YT Jae et) Hace yell alll hy sine as le LG jet Oligosaccharide Sb CLS i yaw Lal 5
BEE Gye jad gill Gaal 5 jal 5 sTrichoderma Harman et al., 2004). Jalal oss ot!
cepa Sym hey) sill endl Lal ge 5 ce pall Trichoderma hy shill J mes Jd Sal! clay WY
By yshall o shaue le GE cy Sip Ad end (Bly SealS) Lee peal) Cy gel ace Sell Lea Gi ail Wks Sed iat ca peal
Sag gshell Lala dae peal! Jal yall Trichoderma 42 jaall LISI) Las Ci Johanne, 2002 )).
8G Trichoderma Chill «jal yline gle
| cual 222 asi Trichoderma les Eli ly 3 58] 5a4,j ke seLy Chal a! peroxidases , chitinases,
anase : Harman et al., 2004).
Trichoderrma s Jeitial «sisi coal) asl) 45RD olin inns y ela paleatal 5p) Ge jiny Chall sie le
paket pale Gent le Si SAM pail Galata Sal 5D oy sll Jone ye (eset nall (Vedidia
1), o9 «58! Sea Windham et al.,1986)) shi 9! ofTrichodermadtl 5 3 si9l) ual jl ge Uli Chall yin ge
vias ats sh ce,