The dog (Canis familiaris[4][5] or Canis lupus familiaris[5]) is a domesticated descendant of the wolf.

Also
called the domestic dog, it is derived from the extinct Pleistocene wolf,[6][7] and the modern wolf is the
dog's nearest living relative.[8] Dogs were the first species to be domesticated[9][8] by hunter-gatherers
over 15,000 years ago[7] before the development of agriculture.[1] Due to their long association with
humans, dogs have expanded to a large number of domestic individuals[10] and gained the ability to
thrive on a starch-rich diet that would be inadequate for other canids.[11]

The dog has been selectively bred over millennia for various behaviors, sensory capabilities, and physical
attributes.[12] Dog breeds vary widely in shape, size, and color. They perform many roles for humans,
such as hunting, herding, pulling loads, protection, assisting police and the military, companionship,
therapy, and aiding disabled people. Over the millennia, dogs became uniquely adapted to human
behavior, and the human-canine bond has been a topic of frequent study.[13] This influence on human
society has given them the sobriquet of "man's best friend".[14]

Taxonomy

Further information: Canis lupus dingo § Taxonomic debate – the domestic dog, dingo, and New Guinea
singing dog

In 1758, the Swedish botanist and zoologist Carl Linnaeus published in his Systema Naturae, the two-
word naming of species (binomial nomenclature). Canis is the Latin word meaning "dog",[15] and under
this genus, he listed the domestic dog, the wolf, and the golden jackal. He classified the domestic dog as
Canis familiaris and, on the next page, classified the grey wolf as Canis lupus.[2] Linnaeus considered the
dog to be a separate species from the wolf because of its upturning tail (cauda recurvata), which is not
found in any other canid.[16]

In 1999, a study of mitochondrial DNA (mtDNA) indicated that the domestic dog may have originated
from the grey wolf, with the dingo and New Guinea singing dog breeds having developed at a time when
human communities were more isolated from each other.[17] In the third edition of Mammal Species of
the World published in 2005, the mammalogist W. Christopher Wozencraft listed under the wolf Canis
lupus its wild subspecies and proposed two additional subspecies, which formed the domestic dog
clade: familiaris, as named by Linnaeus in 1758 and, dingo named by Meyer in 1793. Wozencraft
included hallstromi (the New Guinea singing dog) as another name (junior synonym) for the dingo.
Wozencraft referred to the mtDNA study as one of the guides informing his decision.[3] Mammalogists
have noted the inclusion of familiaris and dingo together under the "domestic dog" clade[18] with some
debating it.[19]
In 2019, a workshop hosted by the IUCN/Species Survival Commission's Canid Specialist Group
considered the dingo and the New Guinea singing dog to be feral Canis familiaris and therefore did not
assess them for the IUCN Red List of Threatened Species.[4]

Evolution

Main article: Evolution of the wolf

Location of a dog's carnassials; the inside of the 4th upper premolar aligns with the outside of the 1st
lower molar, working like scissor blades

The Cretaceous–Paleogene extinction event occurred 66 million years ago and brought an end to the
non-avian dinosaurs and the appearance of the first carnivorans.[20] The name carnivoran is given to a
member of the order Carnivora. Carnivorans possess a common arrangement of teeth called carnassials,
in which the first lower molar and the last upper premolar possess blade-like enamel crowns that act
similar to a pair of shears for cutting meat. This dental arrangement has been modified by adaptation
over the past 60 million years for diets composed of meat, for crushing vegetation, or for the loss of the
carnassial function altogether as in seals, sea lions, and walruses. Today, not all carnivorans are
carnivores, such as the insect-eating aardwolf.[5]

The carnivoran ancestors of the dog-like caniforms and the cat-like feliforms began their separate
evolutionary paths just after the end of the dinosaurs. The first members of the dog family Canidae
appeared 40 million years ago,[21] of which only its subfamily the Caninae survives today in the form of
the wolf-like and fox-like canines. Within the Caninae, the first members of genus Canis appeared six
million years ago,[15] the ancestors of modern domestic dogs, wolves, coyotes, and golden jackals.

Domestication

Main article: Domestication of the dog

The earliest remains generally accepted to be those of a domesticated dog were discovered in Bonn-
Oberkassel, Germany. Contextual, isotopic, genetic, and morphological evidence shows that this dog
was not a local wolf.[22] The dog was dated to 14,223 years ago and was found buried along with a man
and a woman, all three having been sprayed with red hematite powder and buried under large, thick
basalt blocks. The dog had died of canine distemper.[23] Earlier remains dating back to 30,000 years ago
have been described as Paleolithic dogs, but their status as dogs or wolves remains debated[24] because
considerable morphological diversity existed among wolves during the Late Pleistocene.[1]
This timing indicates that the dog was the first species to be domesticated[9][8] in the time of hunter–
gatherers,[7] which predates agriculture.[1] DNA sequences show that all ancient and modern dogs
share a common ancestry and descended from an ancient, extinct wolf population which was distinct
from the modern wolf lineage.[6][7] Most dogs form a sister group to the remains of a Late Pleistocene
wolf found in the Kessleroch cave near Thayngen in the canton of Schaffhausen, Switzerland, which
dates to 14,500 years ago. The most recent common ancestor of both is estimated to be from 32,100
years ago.[25] This indicates that an extinct Late Pleistocene wolf may have been the ancestor of the
dog,[8][1][26] with the modern wolf being the dog's nearest living relative.[8]

The dog is a classic example of a domestic animal that likely travelled a commensal pathway into
domestication.[24][27] The questions of when and where dogs were first domesticated have taxed
geneticists and archaeologists for decades.[9] Genetic studies suggest a domestication process
commencing over 25,000 years ago, in one or several wolf populations in either Europe, the high Arctic,
or eastern Asia.[10] In 2021, a literature review of the current evidence infers that the dog was
domesticated in Siberia 23,000 years ago by ancient North Siberians, then later dispersed eastward into
the Americas and westward across Eurasia.[22]

Breeds

Main article: Dog breed

Further information: Dog type

Dog breeds show a range of phenotypic variation

Dogs are the most variable mammal on earth with around 450 globally recognized dog breeds.[10] In
the Victorian era, directed human selection developed the modern dog breeds, which resulted in a vast
range of phenotypes.[8] Most breeds were derived from small numbers of founders within the last 200
years,[8][10] and since then dogs have undergone rapid phenotypic change and were formed into
today's modern breeds due to artificial selection imposed by humans. The skull, body, and limb
proportions vary significantly between breeds, with dogs displaying more phenotypic diversity than can
be found within the entire order of carnivores. These breeds possess distinct traits related to
morphology, which include body size, skull shape, tail phenotype, fur type and colour.[8] Their
behavioural traits include guarding, herding, and hunting,[8] retrieving, and scent detection. Their
personality traits include hypersocial behavior, boldness, and aggression,[10] which demonstrates the
functional and behavioral diversity of dogs.[8] As a result, present day dogs are the most abundant
carnivore species and are dispersed around the world.[10] The most striking example of this dispersal is
that of the numerous modern breeds of European lineage during the Victorian era.[7]

Bangladeshi Dog

Biology

Anatomy

Main article: Dog anatomy

Skeleton

A lateral view of a dog skeleton

All healthy dogs, regardless of their size and type, have an identical skeletal structure with the exception
of the number of bones in the tail, although there is significant skeletal variation between dogs of
different types.[28][29] The dog's skeleton is well adapted for running; the vertebrae on the neck and
back have extensions for powerful back muscles to connect to, the long ribs provide plenty of room for
the heart and lungs, and the shoulders are unattached to the skeleton allowing great flexibility.[28][29]

Compared to the dog's wolf-like ancestors, selective breeding since domestication has seen the dog's
skeleton greatly enhanced in size for larger types as mastiffs and miniaturised for smaller types such as
terriers; dwarfism has been selectively utilised for some types where short legs are advantageous such
as dachshunds and corgis.[29] Most dogs naturally have 26 vertebrae in their tails, but some with
naturally short tails have as few as three.[28]

The dog's skull has identical components regardless of breed type, but there is significant divergence in
terms of skull shape between types.[29][30] The three basic skull shapes are the elongated
dolichocephalic type as seen in sighthounds, the intermediate mesocephalic or mesaticephalic type, and
the very short and broad brachycephalic type exemplified by mastiff type skulls.[29][30]

Senses

Further information: Dog anatomy § Senses

A dog's senses include vision, hearing, smell, taste, touch, and sensitivity to Earth's magnetic field.
Another study has suggested that dogs can see Earth's magnetic field.[31]

Coat

Main article: Dog coat

Dogs display wide variation in coat type, density, length, color, and composition

The coats of domestic dogs are of two varieties: "double" being familiar with dogs (as well as wolves)
originating from colder climates, made up of a coarse guard hair and a soft down hair, or "single", with
the topcoat only. Breeds may have an occasional "blaze", stripe, or "star" of white fur on their chest or
underside.[32] Premature graying can occur in dogs from as early as one year of age; this is associated
with impulsive behaviors, anxiety behaviors, fear of noise, and fear of unfamiliar people or animals.[33]

Tail

There are many different shapes for dog tails: straight, straight up, sickle, curled, or corkscrew. As with
many canids, one of the primary functions of a dog's tail is to communicate their emotional state, which
can be crucial in getting along with others. In some hunting dogs the tail is traditionally docked to avoid
injuries.

Health

Main article: Dog health

Some breeds of dogs are prone to specific genetic ailments such as elbow and hip dysplasia, blindness,
deafness, pulmonic stenosis, cleft palate, and trick knees. Two severe medical conditions significantly
affecting dogs are pyometra, affecting unspayed females of all breeds and ages, and Gastric dilatation
volvulus (bloat), which affects larger breeds or deep-chested dogs. Both of these are acute conditions
and can kill rapidly. Dogs are also susceptible to parasites such as fleas, ticks, mites, hookworms,
tapeworms, roundworms, and heartworms, which is a roundworm species that lives in the hearts of
dogs.

Several human foods and household ingestible are toxic to dogs, including chocolate solids, causing
theobromine poisoning, onions and garlic, causing thiosulphate, sulfoxide or disulfide poisoning, grapes
and raisins, macadamia nuts, and xylitol.[34] The nicotine in tobacco can also be dangerous to dogs.
Signs of ingestion can include copious vomiting (e.g., from eating cigar butts) or diarrhea. Some other
symptoms are abdominal pain, loss of coordination, collapse, or death.[35][page needed]

Dogs are also vulnerable to some of the same health conditions as humans, including diabetes, dental
and heart disease, epilepsy, cancer, hypothyroidism, and arthritis.

Lifespan

Further information: Aging in dogs

The typical lifespan of dogs varies widely among breeds, but for most, the median longevity (the age at
which half the dogs in a population have died and half are still alive) ranges from 10 to 13 years.[36][37]
The median longevity of mixed-breed dogs, taken as an average of all sizes, is one or more years longer
than that of purebred dogs when all breeds are averaged.[36][37][38] For dogs in England, increased
body weight has been found to be negatively correlated with longevity (i.e., the heavier the dog, the
shorter its lifespan), and mixed-breed dogs live on average 1.2 years longer than purebred dogs.[39]

Reproduction

Main article: Canine reproduction

A female dog nursing newborn puppies.

In domestic dogs, sexual maturity happens around six months to one year for both males and females,
although this can be delayed until up to two years of age for some large breeds, and is the time at which
female dogs will have their first estrous cycle. They will experience subsequent estrous cycles
semiannually, during which the body prepares for pregnancy. At the peak of the cycle, females will
become estrous, mentally and physically receptive to copulation. Because the ova survive and can be
fertilized for a week after ovulation, more than one male can sire the same litter.[12]

Fertilization typically occurs two to five days after ovulation; 14–16 days after ovulation, the embryo
attaches to the uterus and after seven to eight more days, a heartbeat is detectable.[40][41]

Dogs bear their litters roughly 58 to 68 days after fertilization,[12][42] with an average of 63 days,
although the length of gestation can vary. An average litter consists of about six puppies.[43]
Neutering

Neutering is the sterilization of animals, usually by removing the male's testicles or the female's ovaries
and uterus, to eliminate the ability to procreate and reduce sex drive. Because of dogs' overpopulation
in some countries, many animal control agencies, such as the American Society for the Prevention of
Cruelty to Animals (ASPCA), advise that dogs not intended for further breeding should be neutered, so
that they do not have undesired puppies that may later be euthanized.[44]

According to the Humane Society of the United States, three to four million dogs and cats are
euthanized each year.[45] Many more are confined to cages in shelters because there are many more
animals than there are homes. Spaying or castrating dogs helps keep overpopulation down.[46]

Neutering reduces problems caused by hypersexuality, especially in male dogs.[47] Spayed female dogs
are less likely to develop cancers affecting the mammary glands, ovaries, and other reproductive organs.
[48][page needed] However, neutering increases the risk of urinary incontinence in female dogs[49] and
prostate cancer in males[50] and osteosarcoma, hemangiosarcoma, cruciate ligament rupture, obesity,
and diabetes mellitus in either sex.[51]

Inbreeding depression

A common breeding practice for pet dogs is mating between close relatives (e.g., between half and full
siblings).[52] Inbreeding depression is considered to be due mainly to the expression of homozygous
deleterious recessive mutations.[53] Outcrossing between unrelated individuals, including dogs of
different breeds, results in the beneficial masking of deleterious recessive mutations in progeny.[54]

In a study of seven dog breeds (the Bernese Mountain Dog, Basset Hound, Cairn Terrier, Brittany,
German Shepherd Dog, Leonberger, and West Highland White Terrier), it was found that inbreeding
decreases litter size and survival.[55] Another analysis of data on 42,855 Dachshund litters found that as
the inbreeding coefficient increased, litter size decreased and the percentage of stillborn puppies
increased, thus indicating inbreeding depression.[56] In a study of Boxer litters, 22% of puppies died
before reaching 7 weeks of age. Stillbirth was the most frequent cause of death, followed by infection.
Mortality due to infection increased significantly with increases in inbreeding.[57]

Behavior
Main article: Dog behavior

See also: Dog behavior § Behavior compared with other canids

Dog swimming over to catch a ball, pay attention to the leg and tail movements

Dog behavior is the internally coordinated responses (actions or inactions) of the domestic dog
(individuals or groups) to internal and external stimuli.[58] As the oldest domesticated species, dogs'
minds inevitably have been shaped by millennia of contact with humans. As a result of this physical and
social evolution, dogs have acquired the ability to understand and communicate with humans more than
any other species and they are uniquely attuned to human behaviors.[13] Behavioral scientists have
uncovered a surprising set of social-cognitive abilities in domestic dogs. These abilities are not possessed
by the dog's closest canine relatives or other highly intelligent mammals, such as great apes, but rather
parallel to children's social-cognitive skills.[59]

Unlike other domestic species selected for production-related traits, dogs were initially selected for their
behaviors.[60][61] In 2016, a study found that only 11 fixed genes showed variation between wolves
and dogs.[62] These gene variations were unlikely to have been the result of natural evolution and
indicate selection on both morphology and behavior during dog domestication. These genes have been
shown to affect the catecholamine synthesis pathway, with the majority of the genes affecting the fight-
or-flight response[61][63] (i.e., selection for tameness) and emotional processing.[61] Dogs generally
show reduced fear and aggression compared with wolves.[61][64] Some of these genes have been
associated with aggression in some dog breeds, indicating their importance in both the initial
domestication and later in breed formation.[61] Traits of high sociability and lack of fear in dogs may
include genetic modifications related to Williams-Beuren syndrome in humans, which cause
hypersociability at the expense of problem-solving ability.[65]

Intelligence

Main article: Dog intelligence

Dog intelligence is the dog's ability to perceive information and retain it as knowledge for applying to
solve problems. Studies of two dogs suggest that dogs can learn by inference and have advanced
memory skills. A study with Rico, a Border Collie, showed that he knew the labels of over 200 different
items. He inferred the names of novel things by exclusion learning and correctly retrieved those new
items immediately and four weeks after the initial exposure. A study of another Border Collie, Chaser,
documented his learning and memory capabilities. He had learned the names and could associate by
verbal command over 1,000 words.[66] Dogs can read and react appropriately to human body language
such as gesturing and pointing and human voice commands.
One study of canine cognitive abilities found that dogs' capabilities are no more exceptional than those
of other animals, such as horses, chimpanzees, or cats.[67] One limited study of 18 household dogs
found that they lacked spatial memory, and were more focussed on the "what" of a task rather than the
"where".[68]

Dogs demonstrate a theory of mind by engaging in deception.[69] An experimental study showed

compelling evidence that Australian dingos can outperform domestic dogs in non-social problem-
solving, indicating that domestic dogs may have lost much of their original problem-solving abilities once
they joined humans.[70] Another study revealed that after undergoing training to solve a simple
manipulation task, dogs faced with an unsolvable version of the same problem look at the human, while
socialized wolves do not.[71]

Communication

Main article: Dog communication

Dog sounds

0:15

A dog making noises and barking

Problems playing this file? See media help.

Dog communication is how dogs convey information to other dogs, understand messages from humans
and translate the information that dogs are transmitting.[72]: xii  Communication behaviors of dogs
include eye gaze, facial expression, vocalization, body posture (including movements of bodies and
limbs), and gustatory communication (scents, pheromones, and taste). Humans communicate to dogs by
using vocalization, hand signals, and body posture.

Ecology

Population

The dog is probably the most widely abundant large carnivoran living in the human environment.[73]
[74] In 2013, the estimated global dog population was between 700 million[75] and 987 million.[76]
About 20% of dogs live as pets in developed countries.[77] In the developing world, dogs are typically
feral or communally owned, with pet dogs uncommon. Most of these dogs live their lives as scavengers
and have never been owned by humans, with one study showing their most common response when
approached by strangers is to run away (52%) or respond aggressively (11%).[78] Little is known about
these dogs, or the dogs in developed countries that are feral, strays, or are in shelters because the great
majority of modern research on dog cognition has focused on pet dogs living in human homes.[79]

Competitors and predators

Although dogs are the most abundant and widely distributed terrestrial carnivores, feral and free-
ranging dogs' potential to compete with other large carnivores is limited by their strong association with
humans.[73] For example, a review of the studies in dogs' competitive effects on sympatric carnivores
did not mention any research on competition between dogs and wolves.[80][81] Although wolves are
known to kill dogs, they tend to live in pairs or in small packs in areas where they are highly persecuted,
giving them a disadvantage facing large dog groups.[80][82]

Wolves kill dogs wherever they are found together.[83] In some instances, wolves have displayed an
uncharacteristic fearlessness of humans and buildings when attacking dogs to the extent that they have
to be beaten off or killed.[84] Although the numbers of dogs killed each year are relatively low, it
induces a fear of wolves entering villages and farmyards to take dogs and losses of dogs to wolves have
led to demands for more liberal wolf hunting regulations.[80]

Coyotes and big cats have also been known to attack dogs. In particular, leopards are known to have a
preference for dogs and have been recorded to kill and consume them, no matter what their size.[85]
Siberian tigers in the Amur River region have killed dogs in the middle of villages. This indicates that the
dogs were targeted. Amur tigers will not tolerate wolves as competitors within their territories, and the
tigers could be considering dogs in the same way.[86] Striped hyenas are known to kill dogs in their
range.[87]

Diet

See also: Dog food

A Golden Retriever gnawing on a pig's foot

Dogs have been described as omnivores.[12][88][89] Compared to wolves, dogs from agricultural
societies have extra copies of amylase and other genes involved in starch digestion that contribute to an
increased ability to thrive on a starch-rich diet.[11] Similar to humans, some dog breeds produce
amylase in their saliva and are classified as having a high starch diet.[90] However, more like cats and
less like other omnivores, dogs can only produce bile acid with taurine and they cannot produce vitamin
D, which they obtain from animal flesh. Also more like cats, dogs require arginine to maintain its
nitrogen balance. These nutritional requirements place dogs halfway between carnivores and
omnivores.[91]

Range

As a domesticated or semi-domesticated animal, the dog is nearly universal among human societies.
Notable exceptions once included:

The Aboriginal Tasmanians, who were separated from Australia before the arrival of dingos on that
continent

The Andamanese peoples, who were isolated when rising sea levels covered the land bridge to
Myanmar

The Fuegians, who instead domesticated the Fuegian dog, a different canid species

Individual Pacific islands whose maritime settlers did not bring dogs, or where dogs died out after
original settlement, notably the Mariana Islands,[92] Palau[93] and most of the Caroline Islands with
exceptions such as Fais Island and Nukuoro,[94] the Marshall Islands,[95] the Gilbert Islands,[95] New
Caledonia,[96] Vanuatu,[96][97] Tonga,[97] Marquesas,[97] Mangaia in the Cook Islands, Rapa Iti in
French Polynesia, Easter Island,[97] the Chatham Islands[98] and Pitcairn Island (settled by the Bounty
mutineers, who killed off their dogs to escape discovery by passing ships).[99]

Dogs were introduced to Antarctica as sled dogs, but were later outlawed by international agreement
due to the possible risk of spreading infections.[100]

Roles with humans

Domestic dogs inherited complex behaviors, such as bite inhibition, from their wolf ancestors, which
would have been pack hunters with a complex body language. These sophisticated forms of social
cognition and communication may account for their trainability, playfulness and ability to fit into human
households and social situations. These attributes have given dogs a relationship with humans that has
enabled them to become one of the most successful animals today.[101]
The dogs' value to early human hunter-gatherers led to them quickly becoming ubiquitous across world
cultures. Dogs perform many roles for people, such as hunting, herding, pulling loads, protection,
assisting police and the military, companionship and aiding disabled individuals. This influence on
human society has given them the nickname "man's best friend" in the Western world. In some cultures,
however, dogs are also a source of meat.[102][103]

Pets

Siberian Huskies are pack animals that still enjoy some human companionship

It is estimated that three-quarters of the world's dog population lives in the developing world as feral,
village, or community dogs, with pet dogs uncommon.[104][page needed]

"The most widespread form of interspecies bonding occurs between humans and dogs"[105] and the
keeping of dogs as companions, particularly by elites, has a long history.[14] Pet dog populations grew
significantly after World War II as suburbanization increased.[14] In the 1950s and 1960s, dogs were
kept outside more often than they tend to be today[106] (the expression "in the doghouse" - recorded
since 1932[107] - to describe exclusion from the group implies a distance between the doghouse and
the home) and were still primarily functional, acting as a guard, children's playmate, or walking
companion. From the 1980s, there have been changes in the pet dog's role, such as the increased role of
dogs in the emotional support of their human guardians.[108][page needed] People and their dogs have
become increasingly integrated and implicated in each other's lives[109][page needed] to the point
where pet dogs actively shape how a family and home are experienced.[110]

There have been two significant trends occurring within the second half of the 20th century in pet dogs'
changing status. The first has been "commodification", shaping it to conform to social expectations of
personality and behavior.[110] The second has been the broadening of the family's concept and the
home to include dogs-as-dogs within everyday routines and practices.[110]

A vast range of commodity forms aims to transform a pet dog into an ideal companion.[111] The list of
goods, services, and places available is enormous: from dog perfumes, couture, furniture and housing to
dog groomers, therapists, trainers and caretakers, dog cafes, spas, parks and beaches and dog hotels,
airlines and cemeteries.[111] Dog training books, classes, and television programs proliferated as the
process of commodifying the pet dog continued.[112]
The majority of contemporary dog owners describe their pet as part of the family, although some
ambivalence about the relationship is evident in the popular reconceptualization of the dog-human
family as a pack.[110] Some dog trainers, such as on the television program Dog Whisperer, have
promoted a dominance model of dog-human relationships. However, it has been disputed that "trying
to achieve status" is characteristic of dog-human interactions.[113] The idea of the "alpha dog" trying to
be dominant is based on a disproved theory about wolf packs.[114][115] Pet dogs play an active role in
family life; for example, a study of conversations in dog-human families showed how family members
use the dog as a resource, talking to the dog, or talking through the dog; to mediate their interactions
with each other.[116]

Increasingly, human family-members engage in activities centered on the dog's perceived needs and
interests, or in which the dog is an integral partner, such as dog dancing and dog yoga.[111]

According to statistics published by the American Pet Products Manufacturers Association in the
National Pet Owner Survey in 2009–2010, an estimated 77.5 million people in the United States have
pet dogs.[117] The same source shows that nearly 40% of American households own at least one dog, of
which 67% own just one dog, 25% two dogs and nearly 9% more than two dogs. There does not seem to
be any gender preference among dogs as pets, as the statistical data reveal an equal number of male
and female pet dogs. Although several programs promote pet adoption, less than one-fifth of the owned
dogs come from shelters.[117]

A study using magnetic resonance imaging (MRI) to compare humans and dogs showed that dogs have
the same response to voices and use the same parts of the brain as humans do. This gives dogs the
ability to recognize human emotional sounds, making them friendly social pets to humans.[118]

Workers

Dogs have lived and worked with humans in many roles. In addition to dogs' role as companion animals,
dogs have been bred for herding livestock (collies, sheepdogs),[119][page needed][12] hunting (hounds,
pointers)[120][page needed] and rodent control (terriers).[12] Other types of working dogs include
search and rescue dogs,[121] detection dogs trained to detect illicit drugs[122] or chemical weapons;
[123] guard dogs; dogs who assist fishermen with the use of nets; and dogs that pull loads.[12] In 1957,
the dog Laika became the first animal to be launched into Earth orbit, aboard the Soviets' Sputnik 2; she
died during the flight.[124][125]
Various kinds of service dogs and assistance dogs, including guide dogs, hearing dogs, mobility
assistance dogs and psychiatric service dogs, assist individuals with disabilities.[126][127] Some dogs
owned by people with epilepsy have been shown to alert their handler when the handler shows signs of
an impending seizure, sometimes well in advance of onset, allowing the guardian to seek safety,
medication, or medical care.[128]

Athletes and models

See also: Conformation show

People often enter their dogs in competitions, such as breed-conformation shows or sports, including
racing, sledding and agility competitions. In conformation shows, also referred to as breed shows, a
judge familiar with the specific dog breed evaluates individual purebred dogs for conformity with their
established breed type as described in the breed standard. As the breed standard only deals with the
dog's externally observable qualities (such as appearance, movement and temperament), separately
tested qualities (such as ability or health) are not part of the judging in conformation shows.

Food

Main article: Dog meat

Dog meat is consumed in some East Asian countries, including Korea,[129][page needed] China,[102]
Vietnam[103] and the Philippines,[130] which dates back to antiquity.[131] Based on limited data, it is
estimated that 13–16 million dogs are killed and consumed in Asia every year.[132] In China, debates
have ensued over banning the consumption of dog meat.[133] Following the Sui and Tang dynasties of
the first millennium, however, people living on northern China's plains began to eschew eating dogs,
which is likely due to Buddhism and Islam's spread, two religions that forbade the consumption of
certain animals, including the dog. As members of the upper classes shunned dog meat, it gradually
became a social taboo to eat it, even though the general population continued to consume it for
centuries afterward.[citation needed] Dog meat is also consumed in some parts of Switzerland.[134]
Other cultures, such as Polynesia and pre-Columbian Mexico, also consumed dog meat in their history.
Dog fat is also reportedly believed to be beneficial for the lungs in some parts of Poland[135][136] and
Central Asia.[137][138] Proponents of eating dog meat have argued that placing a distinction between
livestock and dogs is Western hypocrisy and that there is no difference in eating different animals' meat.
[139][140][141][142]

In Korea, the primary dog breed raised for meat, the Nureongi, differs from those breeds raised for pets
that Koreans may keep in their homes.[143]
The most popular Korean dog dish is called bosintang, a spicy stew meant to balance the body's heat
during the summer months. Followers of the custom claim this is done to ensure good health by
balancing one's gi, or the body's vital energy. A 19th-century version of bosintang explains that the dish
is prepared by boiling dog meat with scallions and chili powder. Variations of the dish contain chicken
and bamboo shoots. While the dishes are still prevalent in Korea with a segment of the population, dog
is not as widely consumed as beef, pork and chicken.[143]

Health risks

Further information: Dog bite, Canine vector-borne disease, and Dog bite prevention

In 2018, the WHO reported that 59,000 people died globally from rabies, with 59.6% in Asia and 36.4%
in Africa. Rabies is a disease for which dogs are the most important vector.[144] Significant dog bites
affect tens of millions of people globally each year. Children in mid-to-late childhood are the largest
percentage bitten by dogs, with a greater risk of injury to the head and neck. They are more likely to
need medical treatment and have the highest death rate.[145] Sharp claws with powerful muscles
behind them can lacerate flesh in a scratch that can lead to serious infections.[146]

In the U.S., cats and dogs are a factor in more than 86,000 falls each year.[147] It has been estimated
that around 2% of dog-related injuries treated in U.K. hospitals are domestic accidents. The same study
found that while dog involvement in road traffic accidents was difficult to quantify, dog-associated road
accidents involving injury more commonly involved two-wheeled vehicles.[148]

Toxocara canis (dog roundworm) eggs in dog feces can cause toxocariasis. In the United States, about
10,000 cases of Toxocara infection are reported in humans each year, and almost 14% of the U.S.
population is infected.[149] Untreated toxocariasis can cause retinal damage and decreased vision.[150]
Dog feces can also contain hookworms that cause cutaneous larva migrans in humans.[151][152]

Health benefits

Walking a dog

Dogs suffer from the same common disorders as humans; these include cancer, diabetes, heart disease
and neurologic disorders. Their pathology is similar to humans, as is their response to treatment and
their outcomes. Researchers are identifying the genes associated with dog diseases similar to human
disorders, but lack mouse models to find cures for both dogs and humans. The genes involved in canine
obsessive-compulsive disorders led to the detection of four genes in humans' related pathways.[10]

The scientific evidence is mixed as to whether a dog's companionship can enhance human physical
health and psychological well-being.[153] Studies suggesting that there are benefits to physical health
and psychological well-being[154] have been criticized for being poorly controlled.[155] It found that
"the health of elderly people is related to their health habits and social supports but not to their
ownership of, or attachment to, a companion animal." Earlier studies have shown that people who keep
pet dogs or cats exhibit better mental and physical health than those who do not, making fewer visits to
the doctor and being less likely to be on medication than non-guardians.[156]

A 2005 paper states "recent research has failed to support earlier findings that pet ownership is
associated with a reduced risk of cardiovascular disease, a reduced use of general practitioner services,
or any psychological or physical benefits on health for community dwelling older people. Research has,
however, pointed to significantly less absenteeism from school through sickness among children who
live with pets."[153] In one study, new guardians reported a highly significant reduction in minor health
problems during the first month following pet acquisition. This effect was sustained in those with dogs
through to the end of the study.[157]

People with pet dogs took considerably more physical exercise than those with cats and those without
pets. The results provide evidence that keeping pets may have positive effects on human health and
behavior and that for guardians of dogs, these effects are relatively long-term.[157] Pet guardianship
has also been associated with increased coronary artery disease survival. Human guardians are
significantly less likely to die within one year of an acute myocardial infarction than those who did not
own dogs.[158] The association between dog ownership and adult physical activity levels has been
reviewed by several authors.[159]

The health benefits of dogs can result from contact with dogs in general, not solely from having dogs as
pets. For example, when in a pet dog's presence, people show reductions in cardiovascular, behavioral
and psychological indicators of anxiety.[160] Other health benefits are gained from exposure to
immune-stimulating microorganisms, which can protect against allergies and autoimmune diseases
according to the hygiene hypothesis. The benefits of contact with a dog also include social support, as
dogs cannot only provide companionship and social support themselves but also act as facilitators of
social interactions between humans.[161] One study indicated that wheelchair users experience more
positive social interactions with strangers when accompanied by a dog than when they are not.[162] In
2015, a study found that pet owners were significantly more likely to get to know people in their
neighborhood than non-pet owners.[163]

Using dogs and other animals as a part of therapy dates back to the late 18th century, when animals
were introduced into mental institutions to help socialize patients with mental disorders.[164] Animal-
assisted intervention research has shown that animal-assisted therapy with a dog can increase social
behaviors, such as smiling and laughing, among people with Alzheimer's disease.[165] One study
demonstrated that children with ADHD and conduct disorders who participated in an education
program with dogs and other animals showed increased attendance, increased knowledge and skill
objectives and decreased antisocial and violent behavior compared with those not in an animal-assisted
program.[166]

Cultural importance

Main articles: Cultural depictions of dogs and Dogs in religion

Further information: List of fictional dogs

Cerberus, with the gluttons in Dante's Third Circle of Hell. William Blake.

Dogs were depicted to symbolize guidance, protection, loyalty, fidelity, faithfulness, alertness, and love.
[167] In ancient Mesopotamia, from the Old Babylonian period until the Neo-Babylonian, dogs were the
symbol of Ninisina, the goddess of healing and medicine,[168] and her worshippers frequently dedicated
small models of seated dogs to her.[168] In the Neo-Assyrian and Neo-Babylonian periods, dogs were
used as emblems of magical protection.[168] In China, Korea and Japan, dogs are viewed as kind
protectors.[169]

In mythology, dogs often serve as pets or as watchdogs.[169] Stories of dogs guarding the gates of the
underworld recur throughout Indo-European mythologies[170][171] and may originate from Proto-Indo-
European religion.[170][171] In Greek mythology, Cerberus is a three-headed, dragon-tailed watchdog
who guards the gates of Hades.[169] Dogs are also associated with the Greek goddess Hecate.[172] In
Norse mythology, a bloody, four-eyed dog called Garmr guards Helheim.[169] In Persian mythology, two
four-eyed dogs guard the Chinvat Bridge.[169] In Welsh mythology, Annwn is guarded by Cŵn Annwn.
[169] In Hindu mythology, Yama, the god of death, owns two watchdogs who have four eyes. They are
said to watch over the gates of Naraka.[173] A black dog is also considered to be the vahana (vehicle) of
Bhairava (an incarnation of Shiva).[174]