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New combinations in Chrysanthemum (Compositae-Anthemideae) of Asia

with a list of Japanese species

Article  in  Journal of Japanese Botany · June 2004

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Hiroyoshi Ohashi
Tohoku University
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 植物研究雑誌
J
.J
pn
.Bo
t.
7
9:1
86-
195(
200
4)

NewC
omb
ina
tionsi
nChrysanthemum(Co
mpo
sitae
-An
the
mid
eae
)
ofA
siawi
thaL is
to fJap
ane
seSpe
cie
s
a b
H
iro
yos
hiOHASHl andK
ojiYONEKURA

a
Bot
an
ica
lGard
en，G
rad
uat
eS c
hoo
lofSc
ie
nce，To
hokuUni
ve
rs
it
y，S
en
dai，980-0862JAPAN;
b
Th
eMt
.Ha
kkod
aBota
ni
calLa
bor
ato
ry，G
rad
uat
eSch
oolofSc
ie
nce，To
hok
uUn
ive
rsi
ty，A omor
i，030・0111JAPAN
(
Rec
eiv
edo
nJa
nu紅 y10，2004)

Newc ombina
tionsornewnamesf ortwosect
ions，28spe
ciesandt h
ei n
fra
spe
cif
ic
t
axaareprop
osedundertheconse
rvedge
ner
icnameCh ηsanthemumL.Theg enu
siscir
-
cu
mscribe
dh er
ea lm
ostasi de
nti
calwihD
t en
dr a
nthe maofKitamur
a( 1978
)i nc
ludi
ng
αn
A
j ia，Arctan
themum，De nd
ranthemaandPh αeos
tigmαint h
es ens
eo fBremera nd
Hump
h r
ies(1993
).A lis
to fth
eJ ap
ane
sespec
iesofC hrysan
themumisp ro
vide
d.

Keyw
ord
s:A
si，C
a h
rys
ant
hem
um，D
end
ran
the
ma，newc
omb
ina
tio
ns，newn
ame
s.

Theg ene
ric name ChrysanthemumL .i s of Dendranthema a sA jania( A.p acific
a
conservedw ithac onservedtypeC .indicum (Nakai)K . Bremer & Humphries and A .
L.(Brummitt1998，G reutereta.2
l 000).The shiwogiku(Ki t
am.) K. Bremer & Humph-
genercnameDendranthemaDesMou
i l
.b e- ri
es)
. However，m orphologicald isti
n c
tion
C創 ne a synonym o f Chrysanthemum，a 国l andg en
eticisolati
onbetweenthema rec on
-
thought henamehadbeenr ecentl
ya ccepted sider
edt obei nsuff
icien
tt or ecogni
zethem
inE .Asia( Kitamura1978，1 981，Shihand asdiffe
rentgenera.Manyh ybrids紅 eknown
Fu 1983，Barkalov 1992，Koyama 1995， between A j
an ia and Chrysanthemum i n
Penge tal
.1 998)anda lsoinag enericmono- Japan(Table1).Thechromosomeso fA jani
a
graphbyBremerandHumphries( 1993). andChrysanthemum訂 es imilartoeacho ther
BremerandHumphries( 199
3)d i s
cussed (Nakataeta l
.1 987).Distinc
tn e
sso fA jani
a
rela
tions between Ch ηsanthemum ( as fromChrysanthemumwasa l
r eadysuspected
Dendranthema) and i t
ss atel
lit
eg e
n erai
町n byTzvelevbasedone xtensivee xamination
cluding A jania， Arctanthemum and of Chinese m at
erial ( se
e Bremer and
Phaeostigma， which were i ncluded i n Humphries 1 993
). We r eg紅 d A jan
iaa sa
Dendr，αnthema by K itamura (1978). The sect
iono fChηs anthemum.T hi swasa lread
y
genusi全janiaP o
ljak.(P olj
akov1 955)based proposed by Kitamura ( 1978) under
onA .p alla
sianα(Turcz.exB esser)P ol
jak. Dendranthem α.Ph αe o
stigmai ss ep紅 ated
hasbeena ccep旬 db y many a uthors，espe- fromAjaniαbyMuldashev( 1981，1 983)，but
cial
lyi nRussiaandChina，a sdisti
nctfrom ShihandFu( 1983)includeditinベ ja
nia.We
Chrysanthemumo rDendranthem α(Tzvelev inc
1udePh αeo s
tigmαintoC hrysanthemum.
19 1，S
6 h
ih 1979，1994，Muldashev 1983， Chrysanthemum a rctic
umL .( in
cluding
ShihandFu 1983，Barkalov 1992，Bremer Dendranthema k urile
nseT zvele v
)i so ften
and Humphries 1 9
9 3)
. Bremer and regarded a s c ompr
ising t he genus
Humphries( 1
993)k ep
tb othg eneraseparae，
t ArctanthemumTzvelev( 1
985)b asedoni ts
andt reate
dtwod isciformJ apaneses pe
cies dis
tinct
1yribbedc yp
selaswitho utmyxogenic
-186-
J
une2
004 J
our
nalo
fJa
pan
eseB
ota
n .79N
yVo
l o.3 1
87

T
abe1
l .I n
ter
sec
tio
nalhy
brisbetweenChrysanthemums
d ect
.Ch
rys
ant
her
nun(=D
r endranthema)ands
ect
.Aj
ani
a
(=A
jania
)foundinJap
an

Hybrid(
set
.C
c h
rys
ant
her
nur
nxs
ect
.Aj
ani
a)

C
.makinoixC.paUαsian
ωη =ChrysanthemumxkonoanumMakino トガクシギク
仁 m
orifol
iumxC.pacificum=Chrysanthemumxmarginatum(M
iq)N
. .E.B. ハナイソギク
r
C
.i n
dicumxC.kin
okuniense=Ch ηsanthemumxogawaeK it
ar
n. ヒノミサキギク
C
.makinoixC.rup
estre= Chrysanthemumxto
daie
nseH.Ohashi& Yon
ek.* リュウノウイワインチン
C
.makinoixC.kin
okuniense
仁 m
orifol
iumx仁 kinokunien
se
C
.makinoixC.shiwo
giku
C
.morifol
iumxC.shiwogiku

*Newn
arn
ess
eet
ext

ce
lls( Br
emer and Humphries 1993).T hi
s Chrysanthemum s ect
. Arctanthemum
spe
ciesis，however ，v erys im
ilart oC . (Tzve
lev)H .Ohashi& Yone k
.， comb.nov
.
yezo
enseMaek.whichi softenreg紅 dedasa Dendranthema s ect
. Arctanthemum
sub
specie
s o f C. a rc
ticum a s s ubsp
. Tzvelev，Fl
.URSS2 6:385( 1961)
.
maekawanumK itam.Morep recisea na
lyse
s Arctanthemum ( T
zvelev
) T zvelev in
on achenec h
arac
tersa ren eededf o
rC . Novit
.S yst
. Pl
.V asc.2 2: 274 ( 1
985
);
arct
icum，but we c onsidrA
e r
c tanther
um BarkalovinKha r
kev.，Pl
.V a
sc.O ri
ent
.Ext
r.
shoul
dbet r
eate
dasas ectio
no fChrysanthe
- Soviet
.6:1 09(1992
) ;Bremer& Humphries
mum. inBull
.Na t
.Hist
.Mus.L ond.(Bo t
.
)23(2
):
114( 1993)
.Types pec
ie:A
s .a rc
ticum(L.
)
New c omb
in ations for s ections i n Tzvel
ev .
Chrysanthemum
Chrysanthemum s e.A
ct jania (Pol
jak.
) New c ombin
ati o
ns f or speci
es and
H.O hashi& Yonek.，comb.n ov. in
fraspe
cif
ictaxai nalphab
etica
lo rderof
Ajania POりak.i nNo .
tS yst
.H er
b .I st
n. th
es pec
ifi
cepithe
t
Bo t
.A cad.Sci
.URSS1 7
:419( 1955)，p.p.
; Chrysanthemum a bol
inii( Kova
levsk
.)
Tzvelev，Fl.URSS2 6:398( 196
1);C .S hih H.Ohashi& Yonek.，comb.n o
v.
&G .X .Fu， Fl
.R eipub.P
l op.S i
n.76( 1
):1 02 Ajan
iaabol
ini
iK ovalevs.i
k nNovi t
.Syst
.
(1983);B arkalovi n Khar k
ev.，Pl.V asc
. Pl
.V a
sc.23:247( 1986
).
Orient
.E xt
r.S oviet
.6:1 62( 1992
);Bremer Di
str
.:C.Asia.
& Humphriesi nB ul
l.Na t
.H ist
.Mus.L on
d.
(Bot.
)2 3(2)
:1 1 5(1993)
.Types p
ecies:A. Chrysanthemumalabasium(H.C.F u)
pallasia
na( Fi
scherexB ess
e r
)P olj
ak. H
.O hashi& Yone， comb.n
k
. ov.
Dendranthema s ect
. A jania ( Pol
jak.
) A ja
niaalaba
siaH.C .Fu，Fl
.I n
tramo
ng.
Kitam.i nActaP hyt
otax. Geobo.
t2 9
:1 6
8 6
:3 25，.
t34( 19
82)，e
d.2，4:585(199
3).
(1978). Di
str
.:China(In
nerMon
golia
).
Pha e
ost i
gmαMuldashev i n Bot
. Zhum.
SSSR6 6:586( 1981)
;Bremer& Humphries .，S
ChrysanthemumarcticumL p.Pl.2
:
inB u
ll.Na t
.H ist
.Mus.L ond.(Bo t
.
)2 3(2)
: 889(175
3).
1 1
5( 1993)
. su
bsp.yezoens
e (Maek)H
. .Ohas
hi&
Yone，s
k
. tt
.n
a o
v.
1
88 植 物 研 究 雑 誌 第79巻 第3号 平成 1
6年6月

Chrysanthemum a rct
ic umv a.y
r ez
oens e Chrysanthemum arcti
cumis var
.y ezo
ense
Mae k.i nT ra
ns.S apporoNa .H
1 is.S
1 oc.8: Maek.( 192
1)，b utthenamei sconfusa
ble
14，f ig
s.1 -4( 1921);K i
ta m.inMem.C ol
l. with C. y ezoe
nse Mae . (
k 192
1) (=C .
Sci. Kyoto I mp. U ni
v. s e
r. B， 1 5
(3) arct
icumsubsp.maekawanumK i
tam.)which
[C ompos.J ap
.2 ]: 347 ( 1940)，non C . occu
rss i
mpat
rical
lyw it
ht hef or
meri n
yezoens eMae k.(1921). Nemuro P e
ninsula ( Nis
h i
kawa and
Dendranthema k uril
ense T zvelev， Fl. Kobayashi1989)
.
URSS 2 6: 385 ( 1961)，nom. n ov.f orC .
arc t
ic umv紅 .yezoenseMae k
. Chrysanthemum brachyanthum ( C
.
Arctanthemum k ur
ilense ( Tzvel
ev ) Shih
)H .O hasi& Yone
h ，c
k
. omb.n ov
.
Tzv elevi nNovi .S
1 ys1. Pl
.V a
sc .2 2: 274 Ajaniab rachy
anthaC .S hihi nA c
t a
(1985);Bremer& Humphriesi nBul .Na
l 1. Phy
totax.Sin.17:114(1979)，inC .Y.Wu，
His1.M us.L ond.(Bo 1
.)23(2):1 14( 1993). Fl
.X izang
.4 :736(1985)
;C .S hih& G .X.
Arctanthemum αu cticum ( L.
) T z
vele v Fu，Fl.R ei
publ
.P op
.S i
n.76( 1
):1 10，1
.1 7，
subsp.k uril
ense( Tzv
elev)T zvelev，Fl.Arc 1
. fi
g.2( 1983
);Bremer& Humphriesi nBull
.
URSS1 0:1 1
5( 1987)
;B arkalovinKh a
rkev.， Na.H
1 is1
.M us. Lond. (Bot
.)2 3(2
):1 15
Pl.V a
s c.Orien1.Ext
r.S ovie.6
1 :1 10， i
nn o
t a (19
93).
(1992). Di
str
.:C hina(S
.T ib
et( X
izang)
).
?C hrysanthemuma rcticumvar.k urilen
s e
Kudo， Fl
.P aramush
ir.:1 67( 1922
) . ChrysanthemumdelavayanumH .Ohashi
Pla
ntso f Chrysanthemum a rcticumi n &Y onek.，nom.n ov.
Nemuro P eninsu
lao fe astem Hokkaido， Ajaniaseric
eαC.S hihinBull.Bot.L a
b.
Kur i
leI sland
sandS outhS akhalin紅 ech 佐 N.ーE
.Fo r
.Inst
.6 :1 4(1980
);C .Shih&G .
acterizedbyt hes m
allerh abitw ithu sual
l y X.Fu，Fl .R eipub.P
l o
p.S in
.76( 1
):1 18，t
.
sim ples temandmores triat
edl igu
le softh e 19，fi
gs.1& 2( 198
3);Bremer& Humphries
ray-flor
etst hant hoseo ft het ypi
ca lf o
rm . inBull.Na t
.H ist
.Mus.L ond.(Bo t
.
)2 3(2)
:
Acc ordingly，t he
yhavebeend istinguish
e d 115( 1993)，non Chrysanthemums e
r i
c eu
m
fromt hetypic
alforma sad iffer
entt a
xona t Adam ( 18 05-1
8 0
6)，n ec Hoffm. & Link
the r ank o f v ari
ety (Maekawa 1 921， (182
0).
K itamura1 940)，s ubs
pecies( Tzvel ev1 987， Distr
.
:China(NW.Yunnan).
Ba rkalov1 992)orad is
tinctspecies( Tzvel
e v The e pi
thet o f t h
is s p
ecies u nder
1 961，1 985).K itamura(1967，1 97 8，1 98
1 ) Chrysanthemum i s c hanged t o a void
treatedt hist axona s a synonym o fC . homonymya ndrenameda fte
rJ.M.D elavay，
ar cticum，s in
ce he r egarde dp lants from coll
ectoroft h
et ypespecime
n.
Shi re
t okoPeninsulaofe astemHokkaidoa re
ind i
stinguish
able from t he t y
pi c
al C . Chrysanthemumdichrum( C.Shih)H.
ar cticum，butwec onsidert ha
tt het a
xoni s Ohashi& Yone ，c
k
. omb.n ov
.
distinctbecausei t
sd i
agnosticc h
a r
ac te
rsa re Dendranthemad ichru mC.S hi
hi nBull
.
use f
ulandi t
sd istr
ibuti
oni ss ep
a r
a tedfrom Bot
.L ab.N .-E
.F or.I nst
.6: 8( 19
80);C.
thet ypicalform.Wer ecogni z
et het axona s Shih& G .X .Fu，Fl.R eipub.P
l op.Sin.76
as ubspeci
eso fC .αr ct
icum.U nfort un
ately， (1
):47( 1983
);Bremer& Humphriesi nBull
.
thea vailabl
ee pit
hetk u
rilenseatt hes ubspe- Na.H
t ist
. Mus. Lo n
d. (Bot.)23(2)
:1 1
4
cies l eve
l c an n ot be u s ed u nder (199
3).
Chrysanthemumduet ot hee xistenceo fC . Di
str
.:N .C hi
na( Hebe
i )
.
αr cticumv a.k
r u
rilense Kudo ( 1922). The ThisC h
ineses p
ecieswaspresumeda sa n
earlie
st e pi
thet f or t he t axon u nder in
terse
ction
alh yb
ridi nt h
eo r
igi
naldesc
rip
-
J
une2004 Joumalo
fJa
pan
eseB
ota
n .
179No.3
yVo 1
89

t
ion，b
utthel
att
era
uth
orst
rea
tei
tasan
atu
- Her
b.I nst
.Bo t
.Acad. Sc.
iURSS 1 7: 424
r
alspe
cie
s. 19
( 55);C .Shih& G .X . Fu，Fl. Reipubl
.
Pop
.S in
.76(1:1
) 9，
0 inn o
ta( 198
3);Bremer
Chrysanthemumf asti
giatum(C.Winkl.
) &H umphriesinBu
ll.Na t
.H st
i .Mus.L ond.
H.O h
a s
hi& Yonek.，comb.n ov
. (Bot
.)23(2
):115(1993
).
Art
emisiaf a
stigi
ataC . Wink .i
l nA cta Di
str
.:China(N.Yunnan).
Hor
t.P e
trop.11:3 73(1891
).
Ajani
af ast
igiat
a( C. Winkl)P
. ol
jak
.i n Chrysanthemum k hartense( Dunn)H .
Not
.S yst
.H erb.Bo t
.Inst
.A c
ad.URSS1 7: Ohashi& Yone ，c
k
. omb.n ov.
428 (1955
);T zv
elev，Fl. URSS 2 6: 405 Tanacetumkharten
seDunni nKewBul l
.
(19
61);C .S hih& G .X.Fu，Fl .Reipubl
. 1922:150(1922)
.
Pop.Sin. 76(1
):1 25，t
. 20，fig. 3(198
3); Ajani
ak ha
rten
sis(Dunn)C .S h
ihi nActa
Bremer & Humphries i nB u
ll. Nat
.H ist
. Phy
totax.Sin
.1 7
:1 15( 1
9 7
9)，inC.Y .Wu，
Mus.L ond
.(Bo t.
)23(2):115(1993). Fl
.X iza
ng.4:737( 1985)
;C .S hi
h& G .X.
Di
str
.: C . A sia， S. S iber
ia， China Fu，F.lReipubl
.P o
p.S i
n.76( 1
):1 3，t
1 .17，
(Xi
njiang
)andM ong
olia. fi
g.3& t .1 ，f i
g.9( 1983
); Bremer &
HumphriesinB ul
l. Nat
.H it
s.Mus.L ond.
Chrysanthemum g r
acile( Ho
o k.f.& (Bot
.)23(2
):1 15(1993
).
Thomson)H.O hash i& Yone，c
k
. omb.nov. Di
str
.: Pamir，t h
eH im a
layas and SW.
Tanacetumgrac
ileHoo .f
k .& Thomsoni n Chin
a.
Hook.f
， Fl
.B rit
.India3:318(1881).
ベjani
ag rac
ilis (Hook.f. & Thomson) Chrysanthemumk inokuniense(Shimo t
.
Pol
jak.e xT zvelev， Fl
. URSS 2 6: 407 &K itam.
)H .O hashi& Yone ， s
k
. tat
.nov.
(196
1);C .S h
ih& G .X .Fu，Fl.Reipubl. Chrysanthemum s hiwogiku Kitam.v a
r.
Pop.Sin
.76( 1
):1 24，i
nn o
ta(1983);Bremer k
inokuniens
eShimo t
.& K itam.inK itam.in
&H umphriesinB u
ll.Na t
.His.
tMus.L ond. ActaPhytotax.Geobo .
t4 :73( 19 35
)& 22
(Bot
.)23(2
):1 15(1993)
. (4
- 6
):1 20(1967).
Distr
.
:C .AsiaandW.H imal
aya . Ap lanto fChrysanthemum s ect
.A jania
growingincoastalregio
no fKiiP e
ninsulain
Chrysanthemum gr ub
ovii( M
uldas
hev) so
u th
western Honshu h a
s been r eg
arded
H.O has
hi& Yonek
.，c omb.n o
v. t
ax o
nomicallyasav ari
etyo fC .s hi
wogiku
Ajani
ag r
ubo
viiMuldashevinBo t
.Zhu
rn. d
istrib
utedi n S hikoku ( Kitamura 1 9
35，
SSSR 6 7
: 1 529 ( 1982)
; Bremer & 1940，1967，1 978，1 9
81).Shimotomaie ta l
.
HumphriesinBull
.Na t
.H ist
. Mus.Lond. 19
( 68)r eve
al e
dita sb e
ingah ybr
idd eri
va-
(Bot
.)23(2
):115(1993
) . t
iveb etweeno ctoploidC .s hiwogiku and
Di
str
.:Mongo
lia. d
ec a
ploidC.pac がcumd istrib
utedinKanto
andT okairegions.InKi iP enins
ula，how-
Chrysanthemumhypoleucum (Y.L in
g e
ver，t h
ep lantso ccu
rringine asto fCape
exC.S h
ih)H.O h
ashi& Yone，c
k
. omb.nov
. Sh
i ono
misaki，t h
et ipofKi iP en
insula，are
Aja
niah yp
oleucaY.LingexC .S hi
hin p
rincip
allyd eca
pl o
id，w her
ea sthoseg row-
ActaPhyto
tax
.S in
.32:366(1994
). i
ngi nwesto fthecape紅 eu s
uallyo c
toploi
d，
Di
str
.:China
. a
lt h
ough many a neuploi
dp lants were ob-
s
er v
ede special
lya t and a roundt h
et ip
Chrysanthemum junnanicum (
Pol
jak
.) (Shimotoma
ie ta .1
l 9
68). Nakata e ta l
.
H
.O hashi& Yone，c
k
. omb.no
v. 19
( 87)r egardedt h
eo ctopl
oidp lant
sa sC .
A j
aniajunnan
icαPoljak. i
n Not
. Syst
. s
hi wo
gikuandd eca
ploidasC .pacificumex-
1
90 植 物 研 究 雑 誌 第79巻 第3号 6年6月
平成 1

c
lusi
velybasedont heirploid
yd if
fere
n c
es. Thi
sv ar
ietyi
sd u
biousinm o
rphol
ogical
T
heirrecogni
tion，however，makesi tdiff
i- di
stin
ctne
ssfromv a
r.ornαtumi nSatsuma
c
ulttod isc
rimina
tet he two sp
ecies from Pe
ninsu
la. Duetoi t
sr em
oteg eo
graph
ical
e
acho t
her，beca
u s
et h
eirmorphol
ogicald i
s- sep
arati
onwet en
tati
vel
yr ec
ognizei
ta sit
t
inc
tion
sa reo fte
nc ontinuo
us among t he wasdescrib
ed(Ho
ttaeta.
11 9
96).
p
lan
tsinK iiPenin
sula
.Wep refe
rr ec
ogniz-
i
ngt h
ep l
antso fKiiP en
insulaasa ni n
de- Chrysanthemum p rz
ewalsk
ii( P
olj
a k
.)
p
endentspeci
esd e
rivedfromah ybr
idor i
gin H.O hasi& Yonek.，comb.n
h ov.
d
ist
inc
tfromt hepare
nts. Ajani
ap rzew
alsk
iiP olj
ak.inNot .S yst
.
Herb.Inst
.Bo t
.A cad.Sci.URSS 1 7: 422
Chrysanthemum kokanicum ( Kr
asc
h.) (195
5);C .S hh& G
i .X.Fu，F1 .Reipub1.
H.O h
ashi& Yonek.，comb.n o
v. Pop.S in
. 76(1:1
) 09( 19
83); Bremer &
Tanacetum kokan
i c
um Kr a
sch
.i n Not. Humphriesi nBul
l. Nat
.H ist
. Mus. Lond.
Syst
.Herb.Hor t
.Pet
rop.4:7(1923)
. (Bot
.)23(2)
:1 15(1993
).
Ajan
iak o
kanicα(Krasc
h.)T zve
lev，Fl. Disr
t.
:China( Sic
huan，Qing
hai，E.Gansu
URSS2 6:408( 19
61);Bremer& Humphries andNingxia
).
inBul
l.Na t
.H ist
.Mus.L on
d.(Bot.)23(
2):
115(1993). Chrysanthemum purpureiflorum H .
Di
str
.:C .Asi
a. Ohashi& Yonek .，nom.n ov
.
Aja
niap u
rpur eaC.S h
i hinActaPhytot
ax.
Chrysanthemumk orov
inii(Kov
alev
sk.) Si
n.1 7(
2):1 15( 1979)，i nC.Y . Wu，F1.
H
.O hashi& Yonek.，comb.nov. Xiza
ng.4:738( 1985
);C .S hi
h& G .X.Fu，
A ja
niak or
oνini
iK ovalevs.i
k n Novt
.
i F1
. Reipub.
1P op.S in. 76(1
):1 15( 1983
);
Sy .P.
st 1Va
sc.23:246( 1986
). Bremer & Humphries i nB ull
. Na t
.H ist
.
Di
str
.:C.Asia
. Mus. Lond.( Bot.
)2 3(
2):1 15( 1993)，non
ChrysanthemumpurpureumP er
s.( 1
807).
Chrysanthemum nitidum (C.S h
ih)H . Di
str
.:C hin
a( T
ibet(Xiza
ng)).
Ohashi& Yonek
.，comb.n ov.
Ajan
ian i
tid
αC.S
， hihi nBull
.Bo t.L a
b. Chrysanthemum ramosum ( C. C .
N.-E
.F or
.Inst
.6:1 5(1980
);C .Shih& G . Chang)H .Ohashi& Yone ， c
k
. omb.n o
v.
X.Fu，F1.Reipub1
.P o
p.S in
.76( 1
):1 23，t
. Chrysanthemumvariif
oliumC.C .Chang
20，fig
. 2(1983
);Bremer& Humphriesi n va.ramosumC
r .C .ChanginS i
nensi
a5:1 63
Bul
l.Na t
.Hist
.Mus.Lond.(Bo t
.
)23(2)
:1 15 (1934
).
(199
3). Ajn
，α iaramosα(C.C .Chang)C .S hihin
Di
str
.:China(SW.S ic
huan)
. ActaP h
ytot
ax.Sin.17:114( 197
9)，inC.Y .
Wu，Fl.X iz
ang.4:735(1985);C.S h
ih& G .
Chrysanthemum ornatum Hems 1
.i n X. Fu，F.1 Reipub.
1P op
.S in. 76(
1):1 08
Cu
rti s
'sBot.Mag.130:t.7965( 19
04)
. (1983
).
var.tokaren
se( M
.H otta& Y .Hir
ai)H. Phaeostigm
av arii
foli
um( C.C . Chang)
Ohas i& Yonek.，comb.n
h o
v. Muldashev va. ramosum (
r C.C . Chang)
Dendranthemao rna
t m (Hems1
u )K
. itam
. Muldashev in Bot
.Z h
Uffi
. SSSR 6 6: 587
V紅 . t
okar
enseM.H ott
a& Y .HiraiinActa (1981
).
Phytota
x.Geobo t
.47
:1 03(1996 “t
)， o
karen
- Dis
tr.
:C hina(E.T i
bet( Xiz
ang)andW.
si
s". Sichua
n).
Dist
r.:Jap
an:S.Kyushu(ls.
1Yaku-shima，
1
s1KuchinoerabuサimaandTokaraI
. sla
nds
). Chrysanthemumrhombifolium(
Y.L
ing
J
une2004 J
our
nalo
fJa
pan
eseBotanyVo
l.79N
o.3 1
91

&C.Shih)H.Ohashi&Yonek
.
， comb.n
ov. [Ic onogr
. Cormoph. S in
.4 : 510，f ig
. 6433
Dendranthema rhombi
foliu
mY . Ling & (197 5)，comb. n ud] ex K
. itam.i n Acta
C.ShihinB u
ll.Bot.Lab.N.-E.For
.I n
s t
.6: Phytot a
x.Geobo t
.2 9
:1 67( 1978);Kitam.i n
2(1980);C.S hih& G .X.Fu，F.1Reipub1. Sata ke& a ，
.
1 WildFlow.J ap.H erb.P1.3 :
Pop.S i
n. 76(1)
:3 1，.t 5，fig. 1(19 8
3); 167( 1981);Bremer& Humphriesi nB u11.
Bremer & Humphries i nBul
l. Na t
. Hist
. Na t
. Hist . Mus. Lond. (Bot .
)2 3
(2): 114
Mus.Lond.(Bo t.
)23(2:114(
) 199
3). (1993) ;H.Koyamai nK .I wa
ts.& a ，
.1 F.1
Di
str
.:China( Chon
gqing)
. Jap.I IIb:94( 1995).
Dendranthemal avanduli
folium( Fi
sch.ex
Chrysanthemum roborowskii ( Mulda- T rautv.)K itam.va r
.s eticus
p e(Maxim.)C .
shev)H.Ohashi& Yone k
.
， comb.n ov
. Shi hi nBu 1
1.Bo .
tL ab.N .
-E.F or.Ins.
t6 :6
A j
aniar ob
orowskii Muldashev i n Bot. ( 198 0)，p .p .;C .S hih& G . X. Fu，F. 1
Zhum. SSSR 6 7: 1528 (1982)
; Bremer & Reipub .
1P op.S i.76(
n 1)
:42( 1983)，p.p.
Humphriesi nBu11. Nat
. His.
t Mus. Lond. Dendranthemas eticu
spe(Maxim.)K itam.
(Bot.)23(2)
:1 15(1993). f
.b o r
e ale(Makino)K itam.inActaP hytotax.
Distr
.:SW.C hina
. Geobo t
.3 8:381 ( 1987);K itam.& a ，
.1 Co1.
var.tsinghaicum( Muldashev)H .Ohashi 1 1 .
1H erb.P. 1J a
p.( 63thprin
ting)1:6 1，t
.20，
& Yonek.，comb.n ov. no.144( 1987).
A jania r o
borowskii v a
r. t sin
ghaicum D istr
.: Japan (Honshu and N. Kyushu)，
Muldashevi nBo .
tZhum. SSSR6 7: 1529 KoreaandNW.C hina.
(1982). Chrysanthemums eticusp
ewasr arelycu
同l
D is
tr.
:China( Qingha
i). tivatedi nt heEdoe ra，butconsideredase x-
tincti nt h
eM eijip eriod( Kitamura 1940，
Chrysanthemum s eticus
pe ( Maxim.) 1 967 )
.R ecent lyitwasr ediscovere
di nt he
Hand.-Mazz.，Symb.S i
n.7 (
4):1 111(19 3
6). I mpe r
i alP alacei nTokyo( Kitamura1 987).
五b oreal
e( Makino)H.Ohashi& Yone k
.
， Kit am u
ra( 1987)c onsideredita smerelya
comb.n o
v. cultivatedformo fw ildC .b orealedistr
ib-
Chrysanthemumi ndicumL .v ar.borea l
e u te dinJ apan， KoreaandChinaandmadet he
Makinoi nBo t
.Mag.Tokyo1 6:89( 1902). com binat ion Dendranthema s e
ticuspe f.
Chrysanthemumb oreαle(Makino)Maki- b oreal e
.Wea gr
eew ithK itamura'sp in
ion.
no i n Bo t
. Mag. Tokyo 2 3: 20 ( 1909)
; T hi ss pe
c iesh asthesamechromosomenum-
Matsum.，I ndxP.
e 1J a
p.2 (2
): 636 ( 1912)
; b erands i
milark aryotype
sw ithC hineseC .
Kit am.inJ .Jpn. Bo t
.1 3:1 67( 1937)，i n lav andulifoliumT ur
cz.exT rautv.Ast hedi
帽s
Mem.C o11
.S c
i.KyotoI mp.Univ.s e.B，15 t
r inctionbetweent hes p
eciesa reoftencrit
i-
(3)[Compos.J ap.2]:365， t
.1， f
ig.2( 1940)， cal， f urther s tudi
es a re needed on C .
in Acta P hytot
ax. Geobot.2 2(4-6: 129 s
) eticu s
peandC .l a
vandulif
olium.
(1967);H.H紅 a，Enum.S permat
o ph.Jap.2:
1 57( 1952
); Ohwi，F. 1J ap
.: 1187 (19 53)
; ChrysanthemumshihchuanumH.Ohashi
Kit am.& a，
.
1 Co1.111
.H erb.P1
.J ap.1:6 . & Yonek.，nom.n
1，t ov.
20，n o.144( 1957;K
) ita.，Neo・L
g ineam.F.1 A janial atifol
iaC .S h
ihi nB u
11.Bo t
.L ab.
Man sh.:633( 1
979);Ohwi& K ita.，
g NewF. 1 N.E 闇 .F o
r .I nst
.6 :1 2(1980);C .S h
ih&G.
Jap.:1500( 1982
). X. Fu，F. 1 Reipub.1P op.S in. 76(1)
: 106
Chrusanthemum s eticu
spe v紅 . b o
r eαle (198 3
) ;Bremer& Humphriesi nB ul
l.Na t
.
(M 北i no) Hand.-Mazz.，Symb. S in
.7 (4)
: H ist.Mus.Lond. (Bot .
)2 3(
2):1 15(1993)，
1 1
1 1(1936). non Chrysanthemum l atゆl ium (DC.)
Dendranthemab oreale(M紘 i no)Y.Ling Baksay( 19 57).
1
92 植 物 研 究 雑 誌 第79巻 第3号 6年 6月
平成 1

Di
str
.:China(SW.Sic
huan)
. s
ubs
p.αr
cti
cum(
dip
loi
d)アキノコハマ
The sp
ecif
icepit
hetu n
derChrys
anthe
- ギク
mumi srenamedaft
erShihChu，autho
ro f sub
sp.yezoe
nse(Maek.)H. Ohashi.&
tebasionym，t
h oavo
idhomonymy. Yone
k.(di
plo
id)チシマコハマギク

Chrysanthemumx todaien
se H. Ohashi I
I
.S ec
t.Chrysanthemum
& Yone ， nom.n
k
. ov. 2
. Chrysanthemum c rass
um (Kit
副n.
)
Chrysanthemum m akinoi Matsum. & K
itam.( d
eca
ploid
)オオシマノジギク
NakaixC . rupest
re Matsum. & K oidz.
: 3
.ChrysanthemumindicumL.
Kitam.inActaP h
ytotax
. Geobo t
.22(4-6)
: v
ar.indic
um( t
etr
apl
oidandhe
xap
loid
)
113(1967)，cumd escr
.lati
n. シマカンギク
Type: Japan: Honshu， Nagano P ref
. V訂 a
l
.be
sce
nsMakino シロパナハマカ
(Pro
v.S hi
nano)， Kamiinagun， Hasemura， ンギク
Todai，1400m，i nloc
iscalc
arei
s.T.Shimizu v
ar.i
yoe
nse(
te住a
plo
id)イヨアブラギ
5501，21S ep.1959(KYO-holotype). ク
Thisis.aninterse
ctio
nalh yb
ridbetween v
ar. maruyamanumK
ita
m.(
tet
rap
loi
d)
sect
.Chrysanthemumands ect
.Ajan
ia. オッタチカンギク
v
ar.t
sur
ugi
san
eneK
s i
tam
.(t
etr
apl
oid
)
Chrysanthemumt rilobat
um( P
oljak.
)H. ツルギカンギク
Ohashi& Yone ， comb.n
k
. ov. 4
.Chrysanthemumjapone
nseNakai
Ajaniat
r i
loba
taPolj
ak.，Fl
.URSS2 6
:880 V征 .
jαr
pone
nse(he
xaplo
id)ノジギク
(196
1);C .S hih & G. X. Fu，Fl. Reipubl
. va
r.ashi
zuri
eneK
s it
am.(he
xaplo
id)ア
Pop.Sin. 76(1
): 116，t
.1 8，fig
. 3( 198
3); シズリノジギク
Bremer & Humphries i nB u
ll. Na t
. Hist
. 5. Chrysanthemum makinoi Matsum. &
Mus.Lond.(Bo t.
)23(2)
:1 15(1993). Nakai( d
iplo
id) リユウノウギク
Disr
t.
:NW.China( X
inji
ang). 6. Chrysanthemum o kie
ns e K itam.
(te
trap
loid
)オキノアブラギク
Chrysanthemum t ripinnatisec
tum( Y
. Na ka
taeta l
.(1987
)d oub
teditstaxonomic
Ling& C.S h
ih)H.Ohashi& Yone ， comb.
k
. sta
tusb eca
useith a
st hesamechromosome
nov. numbera sC.i ndi
cum.
Ajan
iat ri
pin
nati
sect
aY.Ling& C .S hih 7.Ch ηlsanthemumornαtumHems l
.
inC.ShihinBull
.Bo t
.Lab.N .
-E.F o
r.I st
n. V紅 .omatum( octo
ploid
)サツマノギク
6:1 4( 19
80);C .S hih & G. X. Fu，Fl . va.t
r okar
ense(M.Hotta& Y.H irai
)H.
Reipubl
.Pop.S i
n.7 (1
6 ): 118，t
.1 9，fig
.5 Ohashi& Yone k.(octo
ploi
d) トカラノギク
(198
3);Bremer& Humphriesi nB ul
l.Na t
. 8. Chrysanthemum s et
icus
pe (Maxim.)
Hist
.Mus.Lond.(Bot.)23(2):1 1
5( 199
3). Hand.-Mazz.
Di
str
.:China(NW.S ic
huan). .s
f etic
uspe(onlyknowni nculti
vati
on)
カモメギク
A L i
st of the Japanese spe
cies of f. boreal
e (Makino) H. Ohashi &
Chη'santhemum Yone
k.( di
ploi
d)キクタニギク
Al
lt eJ
h ap
anesetax
a紅 el i
ste
dherewith 9.Chrysanthemumwak αs
，aens
eShimot.ex
th
eco 町ectname，pl
oidyinparen
the
sis，and Ki
tam.( t
etr
aplo
id;der
ivedfromC.m ak
inoi
J
apanesename. xC.s e
ticus
pef.b o
realeorC.i n
dicum)ワ
カサハマギク
1
.Se
ct.Arctantherum 10
. Chrysanthemum we
yric
hii (Maxim.)
1
.Chrysanthemumarc
tic
umL
. Miyabe& T.Miyake( o
ctop
loi
d;seeNakata
J
une2004 J
our
nalo
fJa
pan
eseB
ota
n l
yVo.79No.3 1
93

e
ta .1
l 987)ピレオギク Hybridsinsect
.Chrysanthemum
Thiss pe
c i
es was s yn o
n ymise
dw i
thC . 15
.Chrysanthemumxa phrod
iteK it
am.(=
zaw
a d
skii( Kita
mura1 967，1981)，butisdis
- C.indicumxC .morifol
ium)サンインギク
ti
nctfromt hecyto
geneticalandmo 中hologi
- Kitamura(1967)regardedthi
sp lan
ta sa
ca
lp oin
tso fview. di
sti
ncth exa
ploidspeci
esc lo
selyrelat
edto
11. Chrysanthemum y e
zoense Mae k
. C.i ndicum，butNa kat
ae ta l
.( 1987
) and
(d
ecaploid
)コハマギク Nakataa ndTakeuchi(1998)consider
edt h
e
Thisplanthasbeenf requentl
yr eg
ardedas typ
es pecimenofthi
ss pe
ciesisah ybr
idd e
-
a s ub
species o f C. a rctic
um i n s ct
e. ri
vati
veb etweenhexaploidC.i ndicumand
Arctantherum(Kitamu ra1 935，1940，1 967， cu
ltiva
tedC .m o
rifoli
um. Theh ybri
dityo f
1981，Koyama1 995)，buti squit
ed i
stin
cta s th
istax o
ni ssuppo
rtedfromm or
phologica
l
rev
ealed by N ishik aw
a and K obaya
sh i po
into fview.
19
( 89).Itisrath
erne 征 e
rt oC.z aw
adskiiand 16
.Chrysanthemumxc uneifol
iumKi ta
m.
C.w ey
richiiina chen ec h
aracte
rs( Tzvel
ev (=C.i ndicumxC.y oshina
ganthum) ワジキ
1961，Hind1 988). ギク
12. Chrysanthemum y oshina
g a
nthum 17
.Chrysanthemumxl e
ucanthum( Mak
i-
MakinoexKi tam.(tetra
ploid
)ナカガワノギ no)Makino(=C .makinoixC .s et
icusp
ef.
ク bor
eale)シロバナアブラギク
13. Chrysanthemum z awads
kiiH e
rbich 18
.Chrysanthemumm i
yatojimeneKi
s ta
m.
(hex
ap l
oidando c
toploid
)イワギク (=仁 m orifo
liumx仁 y ezoens
e) ミヤトジマ
Hexaploidp lan
tss poradi
callydistr
ibut
ed ギク
inHonshu，S hikokua ndKyushu紅 'ea pp紅 - 1. Chrysanthemumxs
9 h
imotomaiiM紘 i-
ent
lyr el
ict
ali nJ ap
an.J apanesehexaplo
ids no(=仁 i ndicmxC
u .j ap
onense) ニジガハ
dominantlyh aved i
ssecte
dl eaveswithn訂 - マギク
rowerl obes，buti n some l o
cal
itie
sp lan
ts
withw iderl o
beso fl eavesa reg row
ing. 1
11.Sect
.Ajania
Thesep la
ntswereo ftenreferre
dt othevar
i- 20. Chrysanthemum kino
kun
iense
etyo fc onti
nentalA sia， va r
. latilob
um (Shimot
.& K ita
m.)H .O hashi & Yonek.
Kitam.，b u
ta reh erec onsi
dereda sl yi
ng (oct
oploidandd ecapl
oid;d e
rivedfrom C.
with
int herangeo fC.z aw
a d
skiiass t
atedby paci
ficumxC .s hiwo
giku) キノタニシオギ
Kitamura(1967，1 981). ク
The o ct
o p
loidp lant
sr e
centl
yf ou
ndi n 21. Chrysanthemum p a
cificum Nakai
KitakamiM ountainsi nN.Honshu(Na ka
ta (de
caploid
)イソギク
andKumagai 1 999) seems p
ecif
ical
lyd is
- 22. Chrysanthemump alla
sianum'(
Fis
ch.
ti
nctn oto nlyfrom C .z awads
kiib uta l
so exB ess
er)Kom.( hexa
ploid
) オオイワイン
fromC .w eyric
hii. Thesep la
ntsweres u
s- チン
pect
ed a s h ybrid d er
ivativ
es b etween 23.Chrysanthemumr u
pestr
eMatsum.&
hexap
l o
idC .z awadskii and decap
loidC . Koidz.(dip
loid
)イワインチン
yezo
en se(N a
kataandKumagai1 999
). 24. Chrysanthemum s hiwogi
kuK itam.
(oc
toploi
d)シオギク
C
ulti
vate
ds pe
cies
1
4. Chrysanthemum mor
ifo
liu
m Rama.
t IV
.I n
ters
ecti
onalhybridsbetweense
ct.
(
hexap
loidprobablyderi
vedfromahyb
rid Chrysanthemumands ec
t.Ajania
C
.i ndi
cumxC .zawads
kii)キク 25
.Chrysanthemum xkonoanum M紘 ino
(C
.m ak
inoixC.p a
lla
siαnum) トガクシギク
26
.Chrysanthemumxmarginatum (M
iq.)
1
94 植 物 研 究 雑 誌 第79巻 第3号 平成 1
6年 6月

.Br
N.E .(仁 m
ori
fol
iumxC
.pa
cif
icu
m)ハ OhwiJ.，K it amuraS .，W atariS.a ndT ornina
riT .
ナイソギク (e
ds.
)WildF l owerso fJa p
an，Herb.Pl.3 :156-235.
HeibonshaL td.，Tokyo( inJapanese
).
2
7. ChrysanthemumxogαwaeKitam.(C.
一一一 1 9
87. On t hrees pecie
so fS ympetalae.A cta
i
ndicumxC .kinoku
niens
e)ヒノミサキギク Phy
totax
.Geobo .3
t 8:3 8 0-382(inJa p
a n
ese).
2
8.Chrysanthemumx to
daie
nseH.Ohashi KoyamaH .1 995.T ribe5 .A nthem
id e
ae.I n:I wa
tsuki
& Yonek. (
C.m akinoixC.r u
pes
tre
) リユ ， Y
K
. am aza kiT .，B ouf for
dD .E .a ndOhbaH .
ウノウイワインチン (e
ds.
)F l or
ao fJ apan3 b:7 7-97.K o danshaL td
.，
2
9. Chrysanthemum m
akinoixC.k i
no- Tokyo.
MaekawaT .1 921. Ont hew ildChrysanthemumo f
k
unien
se nor
thJ apan.T rans.S app oroNa .H
t ist
.S oc.8:9-
3
0.Chrysanthemummori
foliumxC.kin
o- 18
.
k
unien
se MuldashevA .A .1 981
. A newg enusPhaeostigma
3
1. Chrysanthemummaki
noixC.s hi
wo・ (Ast
eraceae)f romt heE astAsia.Bo .Zhum.SSSR
t
g
iku 66(
4):5 84- 588( i
nR ussian).
一一 1 98
3.A c rit
ica
lr e viewo ftheg en
u sA jan
ia( As
3
2. Chrysanthemum m o
rifo
liumx仁
terac
eae- A
n themideae).Bo .Zhum.SSSR6
t 8(2
):
s
hiwogiku 207-214( inR ussi
an).
NakataM a n dT akeuc hi M. 1 99
8.V ariati
onsi n
References Dendranthemai ndicumv ar.aphroditep opula
tion
BarkalovV .Y .1 992.A steraceae( Co
mposit
ae).I n
: inO oza
k ai，HimiC ity，ToyamaP ref
.B ull.Bo t
.
Charkevicz (Kha rke
vicz)， S.S .( ed
.)P lant
ae Gard
.Toyaman o
.3 :1 -16(inJapanese).
Vasculare
sO rient
isE x
tre m
iS ovie
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idc ytot
ypeo f
Nauka ，S.-
t Peters
burg( i
nR uss
ian). Dendranthemaz awads kii( A
sterace
ae )a ndit
si m-
BremerK .a ndH umphriesC.J .1 99
3.G eneri
cmono匂
pl
icat
ionsi ne v
olutionalh is
tory
.B ul .Bo
l .G
t ard.
grapho ftheA steraceae-Anthemid
eae.B u
ll.Na t
. Toyaman o
.4 :1 -1 5( i
nJ apane
se).
Hist
.M us.L ond.( Bot)2
. 3(2):71-177. 一一一， T anakaR ， T
. an i
g uchiK .a ndS h imotomaiN .
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98.R e
por toft hec om
mitteef or 1987.Sp ecieso fw ildc hrysanthemumsi nJ a
pan:
Spermatophy
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-444. cyt
ologi
c ala n dcy t
og enet
icalv iewo nitse n
tit
y.
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ut e
rW.，M cN e
i l
lJ .，BarrieF.R， B
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t a
x.Geobo .3
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:2 41-259( inJapanes
e).
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asT .S .，N i
colsonD .H .， NishikawaT .a n dK obayas hiH .1 98
9.Chromosome
SilvaP.C.，SkogJ .E .，T reha
neP，T ur
landN .J.， number a n
d d ist
ri b
ution o f Chrysanthemum
andHawksworthD .L .2 000.Intem
ationa
lCodeo f arctic
umL .J .J p
n.Bo .6
t 4:7 7-84(inJ apan
ese).
Botani
c a
l N o
m e
nc lature ( S
aint Louis C ode
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ien
t i
ficBooks，K onigst
ein. Composi
t ae.I n:HuangT .C.eta .(
l edsよFlo rao f
HindD .J .N .1 988. Dendranthemay ezo
ense. Kew Taiwan2 nde d
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rialC ommittee
Mag.5 :5 5-
6 0. oft h
eF lorao fT aiwan，D epartmento fB otany，
HottaM.， YamakawaN .，HiraiY.a ndShiuc
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nalT aiwanU niversity，Taipe
i.
Taxonomicaln oteso nplant
so fsouthemJ ap
an111. Poljak
ovP.1 955.Duog e n町 enovaef am
.C om
p o
sitae.
Distr
ibutio
na ndtaxonomyo ft heDendranthema No .S
t yst
.H e rb.I n
st.Bo t
.Ak ad.Sc i
.URSS1 7:
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t a 418-431.
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a x
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t 7(1
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ti o
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il i
aeC ompos it
訂 umT ib
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mums )
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.P arss ecund
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. 76(1
):1 - 1
4 9
.S ci
e n
c eP res
s，Beijing( i
nC hine
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， Ponanthemum.A c
t a S
himotomaiN .，A dac hiS .a ndM atsum oriS .1 968.
Phytot
ax.Geobo .2
t 9
:1 65-170. Cytolo
gi cal，mo 中h ologicala n
dg eographicalstud-
一一一 1 981.C ompositae( Aster
aceae
).I n
:S a
takeY .， ieso n Chηsanthemums hiwogikuv a
r.k inokuni-
J
une2
004 J
our
nalo
fJa
pan
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ota
nyVo
.
l79N
o.3 1
95

en
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. 一一一 1
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大橋広好a，米倉浩司 b .アジア産キク科キク属の
新組み合わせと日本産種一覧
キクの仲間はかつては広義の ChrysanthemumL . ことが知られている.このため，北村四郎博士は
の下に分類されてきたが 近年はこれを複数の属 これらの属をキク属に含め それぞれ節として取
に細分することが多く，細分した場合の属名とし り扱った.われわれは北村博士のキク属の概念を
てはこれまで Dendranthema( DC.)DesMo
uli
nsが 採用する. BremerandHumphries(199
3)は，A j
ania
用いられてきた.しかし， 1995年に，これまで、シュ とキク属との差異が小さいことを認めながらも，
ンギク Chrysanthemumc oro
nariumL.をタイプと 両者を合一することで必要となる多数の新組合せ
してきた Chrysanthemum L.をシマカンギク仁 を避けて両属を維持し，かえって日本産のイソギ
in
dicumL.をタイプとして保存することが提案さ クとシオギクを A j
aniaに組み替えた.しかし，こ
れ，これがセントルイスの第 1 6回国際植物学会議 こでは両属を同一属として扱ったので，中国から
で認可された.この結果 ， Chrysanthemumは現行 中央アジアにかけて最近記載された多数の A j
ania
の国際植物命名規約付則 1 1
1の保存属名中に掲載 の種を Chrysanthemumに組み替えることが必要と
・
されている ( G
reut
ereta l
.2000. このため，キク
) なった. こ の 群 に 属 す る キ ノ ク ニ シ オ ギ ク 仁
属の学名は Chrysanthemumに戻さねばならない. shi
wogik
uva
r.kinoku
niens .& K
eShimot i
ta.は
m ，
日本産のキク属植物は大部分 Chrysanthemumの下 イソギクとシオギクの交雑由来の植物であること
で記載されてきたので，新組み替えの必要がある が下斗米直昌博士らの研究により既に明らかになっ
ものは少ないが，近年に Dendranthem αの下で記 ているので種として扱った.
載されたトカラノギクや古い栽培植物のカモメギ この他，チシマコハマギクの学名の変更と，リユ
クと品種関係にあることが分かつたキクタニギク ウノウイワインチン(リユウノウギク×イワイン
については組み替えが必要となった. チン)に対する新名の提唱を行った.チシマコハ
中国やロシアでは，頭花に舌状花を欠くイワイ マギクに対して新たに定めた学名は，類似するコ
ンチンの仲間を Ajan 臼や P haeost
igma，痩果の性 ハマギクの学名と紛らわしいが，命名法上やむを
質が少し異なるチシマコハマギクを Arctanthemum えない.
として別属で扱うことが多く，これらの属は世界 最後に，日本産のキク属植物の学名を Chη' sa
n-
のキク亜科植物の属レベルのモノグラフをまとめ themumの下で整理した一覧を示した.
た BremerandHumphries(199)にも採用されてい
3 (
a東北大学大学院理学研究科
る. しかし，これらの属の識別形質は小さい差異 附属植物園津田記念館，
であり，また，イワインチンの仲間は日本ではキ b東北大学大学院理学研究科

クの仲間とふつうに交雑して遺伝的な隔離はない 附属植物園八甲田山分園)

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