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2019 Haplogroup L
2019 Haplogroup L
Downloading this spreadsheet as Excel retains format. Links may not copy to new s
The entire work is identified by the Version Number and date given Main Page.
Directions for citing the document are listeMain Page.
Version History Last revision date for this specific page: 1 October 2019
Because of continuing research, structure of the Y-DNA Haplogroup Tree changes and will be updated when
LINKS: Main Page. Listing Criteria Papers/Presentations Cited Y-DNA Tree Trunk
Glossary SNP Index
FONT COLORS: Newly confirmed in 2019 within subclade Confirmed within subclade
SYMBOLS: ^ Indicates a next-generation sequencing entry which does not yet meet quality guidelines for
M20/PF5570,^^L656, Indicates
L855,an entry
L863, which does not
L878/PF5524, meet qualityL1304,
L879/PF5697, guidelines
M11,but may be helpful.
M61/Page43, M185/PF5755, M23
M2362/PF5692, ~ Indicates only an M2368/PF5693,
M2365, M2366, approximate location on the tree.M2370/PF5695, M2379/S10007, M2381, M
M2369/PF5694,
M2382/PF5701,
The criteria for a M2390/PF5705,
representative SNP M2403/PF5528,
printed in boldM2404/PF5715,
for a subcladeM2407/PF5717, M2411/PF5718,
is: traditional usage, testing doneM2413/P
in multi
M2444/PF5729,
and/or being M2453/PF5737,
foundininthat
thethe
area M2454/PF5738,
of the chromosome M2457,
usedhave M2463/PF5743,
in recent research M2464,
studies. M2466/PF5744, M2470,
SNPs
DID listed
YOU
M2482/PF5750, below
NOTICE italics
M2485/PF5754, (colored
items inblack or red) are
gray M2491,
M2489, sometimes quality variants
a long
M2492/PF5756, name from next-generation
preceding
M2495, M2497,their sequencing
listing
M2503, thereport
that is M2518/P
M2504, same
consistently
above? In showing
this as
situation, representing
evidence that
seems subgroup.
to indicate they also seem to belong
M2525/PF5771, M2528, M2530, M2531, M2532/PF5773, M2536/PF5775, M2537, M2541/PF5777, M2548,to the same subgroup, but some
confirmatory
M2554/PF5785, evidence is not yet available.
M2556/PF5787, M2559, M2569/PF5792, M2572/PF5795, M2578/PF5805, M2579/PF5806,
M2594, M2595/PF5813, M2596, M2597/PF5563, M2598/PF5815, M2607/S7544, M2609, M2611/PF5564, M
M2615/PF5823, M2619, M2620, M2621, M2624, M2628, M2629, M2633, M2638, M2640/PF5829, M2642, M
M2655/PF5832, M2657/PF5833, M2660/PF5837, M2661/PF5839, M2670/PF5841, M2675/PF5842, M8987/P
PF5554,CTS2285/M2465,
PF5557, PF5583,M2374, PF5732,M2383,
PF5757, PF5779,
M2384, PF5790,
M2386, PF5791, PF5802,
M2389/PF5703, PF5804,
M2391, PF5830,
M2392, PF5835,
M2394, M2395/PF5Y551
L Y5530/Z20173,
M2418,
FGC11566, M2363, Y5564/Z20190,
M2419, M2420,
M2456, M2474,Z20117
M2421, M2427/PF1963/Z1075,
M2500, M2526/PF5772, M2527, M2430/PF5725,
M2552, M2564, M2435/PF5726,
M2584/PF5560, M2437, M243
M2612, M
L~ M2652,M2490,
M2680,M2494,
PF5790,M2496, M2517, M2543,
Y5542/Z20116, M2544, M2545,
Y5553/Z20165, Z20055,M2546,
Z20116,M2560, M2568,
Z20164, M2576, M2586, M2593
Z20192
M2635, M2639, M2665, M2666, M2668/PF5840, M2673, M2676, SK1410/Y5528/Z20169, Y5517/Z201
L1 M22 , L1304, M295,
Y5523/Z20157, Page121
Y5524/Z20161, Y5534/Z20201,
M27, F11470/Z20390, L1319, L1320,Y5537/Z20202,
L1321, M76, P329, Y5545/Z20129, Y5546/Z20158,
SK1423, SK1419/Z20362, Y5555
SK1420/Z
L1~ Z20138, Z20139, Z20140, Z20141, Z20153, Z20154, Z20155
Z8050, Z20336, Z20354, Z20355, Z20356, Z20357, Z20358^, Z20359, Z20360, Z20363, Z20464
L1a M2481Z20373^, Z20376,M2664,
, M2565, M2605, Z20378,Y5525/Z20163,
Z20379, Z20380, Z20382, Z20383, Z20384, Z20385, Z20387, Z2038
Y5560/Z20168
Z20399, Z20400, Z20401, Z20403, Z20405, Z20406, Z20407, Z20410, Z20411, Z20412, Z20413
L1a~ M2357,Z20426,M2375,Z20427,
M2401, Z20428,
M2480, Z20433,
M2498, M2555,
Z20434, M2623,
Z20435,SK1417/Y5544,
Z20436, Z20437,SK1418/Y5565,
Z20438, Z20439,Z20132
Z20440
L1a1 Z20450, Z20452, Z24638.2
L1a1~ Z8048, Z20402, Z20429
L1a1a Z20458, Z20462, Z20463, Z20464, Z20465, Z20466, Z20468, Z20469, Z20470, Z2047
L1a1a~ Z20457
L1a1a1~Z34426, Z34427, Z34428, Z34429, Z34430
L1a1a2 Z20475, Z20476, Z20477, Z20483, Z20484, Z20479, Z20480, Z20481, Z204
L1a1a2~Z5925, Z20478
L1a1a2a~ Z42477, Z42478, Z42479, Z42481
L1a1a2b~ Z42482, Z42483, Z42484, Z42485, Z42486
L1a1b Z5926, SK1426/Z20512, Z20492, Z20493, Z20496, Z20498, Z20499, Z20502, Z20503
L1a1b~ Z20497, Z20505
L1a1b1 Z20519^
L1a1b1~M3904
L1a1b1a~ Z34406, Z34408, Z34409, Z34410
L1a1b1b~ Z34411, Z34412, Z34413, Z34414, Z34415
L1a1b2 Z20520, Z20521, Z20522
L1a1b2a Z5929
L1a1b2a1~ Z34379, Z34381, Z34382
L1a1b2a2~ Z34383, Z34384, Z34385, Z34387
L1a1b2b~ Z34467, Z34468, Z34470, Z34471
L1a1b3 Z5930
L1a1b3a Z5931
L1a1b3a1 Z5933
L1a1b3a1a Z5934
L1a1b3a1a1
L1a1b3a1a2
NOTES:
M22 and M295 were considered equivalent to M11, M20, M61 and M185 in previous versions of the tree. In May 20
small group of members of the Y-Haplogroup L Project were found to be negative for all known subgroup SN
in addition M295- despite being M20+ and/or M61+. Further testing has confirmed that M22 is also negative
been revised to reflect this discovery. A new SNP L595 has been discovered in a member of the M22- M295-
confirmed in a second member. This SNP therefore designates a new subclade: L2. Several new SNPs have b
Identical SNPs that were discovered separately are listed in alphabetical order, not necessarily in the order of discove
and separated by "/". Example: M61/Page43.
Items recurrent or volatile: L147.4, L193.2/S176.2.
Mutations previously on tree but found so far only in one man or in closely related men:
PK3 is downstream of M357. Listed 6 August 2012.
L1305, L1306 are found at approximately M357. Listed 6 January 2016.
L864, L865, L866, L867, L868, L869, L870 are found at approximately L595. Listed 6 January 2016.
L1318 is found at approximately M27. Listed 6 January 2016.
Geographic distribution so far of ancestry of modern-day haplogroup L subgroup men (not complete):
L1a1 M2481 Pakistan, India, Sri Lanka, Bangladesh, Iraq, Bahrain, Kuwait, Saudi Arabia,
Qatar,United Arab Em.
L1a2 M357 India, Sri Lanka, Bangladesh, Iraq, Kuwait, Saudi Arabia, Qatar, Armenia, Russia, Italy
L1b FGC36877/M317 Israel (Druze), Lebanon, Kuwait, Kazakhstan (Tatars), Armenia, Azerbaijan, Iran,
Turkey, Belarus, Greece, Bulgaria, Germany, Austria, Switzerland, Portugal,
Ukraine, Russia, Mongolia
L2 L595 Germany, Sardinian Italy
References:
Balanovsky et al, Parallel Evolution of Genes and Languages in the Caucasus Region. Molecular Biology and Evolu
Behar et al, Contrasting Patterns of Y Chromosome Variation in Ashkenazi Jewish and Host Non-Jewish European P
Behar et al, Genome-Wide Structure of the Jewish People. Nature, 446:238-42, 2010.
Cinnioglu et al, Excavating Y-chromosome Haplotype Strata in Anatolia. (pdf) Human Genetics. 114:127-148, 2004
Cruciani et al, A Back Migration from Asia to Sub-Saharan Africa Is Supported by High-Resolution Analysis of Hum
Deng et al, Evolution and Migration History of the Chinese Population Inferred from the Chinese Y-chromosome Ev
Francalacci et al, Low-Pass DNA Sequencing of 1200 Sardinians Reconstructs European Y-Chromosome Phylogeny
Hallast et al, The Y-chromosome Tree Bursts into Leaf: 13,000 High-confidence SNPs Covering the Majority of Kn
Karafet et al, New Binary Polymorphisms Reshape and Increase Resolution of the Human Y-Chromosomal Haplogr
Karafet et al, Paternal Population History of East Asia: Sources, Patterns, and Microevolutionary Processes. (pdf) Am
Karmin et al, A Recent Bottleneck of Y chromosome Diversity Coincides with a Global Change in Culture. Genome
Kivisild et al, The Genetic Heritage of the Earliest Settlers Persists in Both Indian Tribal and Caste Populations. (pdf
Mohyuddin et al, Detection of novel Y SNPs Provides Further Insights into Y chromosomal Variation in Pakistan. Jo
Regueiro et al, Iran: Tricontinental Nexus for Y-Chromosome Driven Migration. (abstract) Human Heredity, Vol. 61
Sengupta et al, Polarity and Temporality of High Resolution Y-chromosome Distributions in India Identify Both Ind
Shen et al, Reconstruction of Patrilineages and Matrilineages of Samaritans and other Israeli Populations from Y-Ch
Thangaraj et al, Genetic Affinities of the Andaman Islanders, a Vanishing Human Population. (pdf) Current Biology
Zhao et al, Presence of Three Different Paternal Lineages among North Indians: A Study of 560 Y Chromosomes. (a
A limited number of qualifying investigative items here appear only in the public Y trees
Additional Resources:
ISOGG Wiki What you need to know about Genetic Genealogy.
The Y-Haplogroup L Project Gareth Henson, Peter Hrechdakian
Back to
Main Page
Y-DNA Tree Trunk
SNP Index
Papers/Presentations Cited
Glossary
Listing Criteria
Investigational items
4414, Z34415
Z34385, Z34387
Z20500
L1a1b3a1a1a~ Z34416, Z34419, Z42487, Z42488
L1a1b3a1a1b~ Z34425, Z42489, Z42490, Z42491
Z8033, Z20527, Z20528
L1a1b3a1a2a Z5937, Z20535, Z20537, Z20539, Z20540, Z20542, Z20544, Z20545, Z20547, Z20548, Z20550
L1a1b3a1a2a~ Z20541, Z20546, Z20549, Z20552
L1a1b3a1a2a1 Z5938, A6402.2/JFS00004.2/Z20559.2, Z8034, Z20558, Z20561, Z20562, Z20563^, Z205
L1a1b3a1a2a1a~ Z39168.2, Z38449
L1a1b3a1a2a1b~ Z38450, Z38452, Z38453, Z34113
L1a1b3a1a2a2~ Z34122, Z34125, Z42492, Z42493
L1a1b3a1a2b~ Z34479, Z34481, Z34482, Z42494, Z42495
4368, Z34369, Z42496, Z42497, Z42498
0771, SK1454, SK1455, Y17950, Y17952, Y17953, Y17954, Y17955, Y17957, Y17958, Y17959, Y17960, Y17961,
0,, Y17965,
Z34483,Y17966,
Z34484,Y17967,
Z34485, Y17968,
Z34486 Y18179, Y18181, Y18182, Y18183, Y18185, Y18186, Y18187, Y18188,
, Y18191, Y18193, Y18194, Y18195, Y18196, Y18197, Y18198, Y20047
7956, Y18180, Y18178, Y18184, Y18192, Y20048
GC32255/Y6252, FGC32257/Y6270, FGC32262/Y6256, FGC32264/Y6272, FGC32265/Y6273,
GC32270/Y6277, FGC32272/Y6279, FGC32276/Y6259, FGC32279/Y6282, FGC32280/Y6283,
GC32284/Y6286, FGC32287/Y6289, FGC32292/Y6291, FGC32294/Y6293, FGC32296/Y6295,
285, Y13388,
2254, Y19628,
FGC32256, Y20208
FGC32260, FGC32263, FGC32266, FGC32267, FGC32271, FGC32273, FGC32274, FGC32278,
2288, FGC32291, FGC32293, FGC32295,
247, Y6253, Y6254, Y6255, Y6261, Y20686,
Y6265, Y6266,Z6953
Y6267, Y6268, Y6269, Y6271, Y6275, Y6278, Y6281, Y6292,
n (not complete):
T->A
T->A
A->G
A->G
G->A
G->A
G->A
G->A
C->T
C->T
C->T
C->T
C->T
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A->C
A->T
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C->A
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C->A
C->A
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C->A
A->G
T->A
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A->G
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G->T
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C->T
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C->A
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G->T
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A->C
C->A
T->C
A->G
A->T
G->A
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C->T
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A->C
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C->A
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T->C
G->A
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G->A
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C->A
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C->A
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C->T
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C->A
C->T
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T->A
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G->A
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GA->del
GA->del
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G->A
G->A
A->C
A->C
G->A
C->A
C->A
C->T
G->A
T->C
C->T
A->G
T->A
C->T
A->G
C->T
G->A
G->A
G->A
A->T
C->A
A->T
A->G
C->G
G->A
G->A
G->C
A->G
A->T
C->T
G->A
A->C
A->C
C->T
G->A
C->G
A->T
C->A
A->G
T->C
A->T
A->C
A->T
C->G
A->C
G->A
A->C
C->T
C->T
G->A
G->A
A->G
C->A
C->T
C->T
G->A
T->C
C->T
G->C
T->A
A->G
G->A
G->A
T->A
G->A
G->A
A->G
C->T
G->C
C->T
G->C
G->A
G->A
A->G
G->A
C->T
G->A
T->G
C->T
T->C
A->T
G->A
G->A
A->G
G->A
A->T
A->G
C->T
C->T
T->A
G->A
T->C
G->A
C->T
G->C
G->C
C->T
C->T
A->G
G->C
C->G
G->T
C->T
C->T
C->G
G->A
A->G
T->C
G->T
G->T
G->A
A->G
A->G
G->A
G->T
A->T
A->G
C->A
T->G
C->T
A->G
C->A
A->G
A->C
G->T
A->G
G->T