GEOGRAPHIC DISTRIBUTION 93

GEOGRAPHIC DISTRIBUTION
Herpetological Review publishes brief notices of new geographic distribution records in order to make them available to the
herpetological community in published form. Geographic distribution records are important to biologists in that they allow for a
more precise determination of a species’ range, and thereby permit a more significant interpretation of its biology.
These geographic distribution records will be accepted in a standard format only, and all authors must adhere to that format, as
follows: SCIENTIFIC NAME, STANDARD ENGLISH NAME if available (for the United States and Canada as it appears in Crother [ed.]
2017. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding
Confidence in Our Understanding. 8th ed. Herpetol. Circ. 43:1–102 [available for download here: https://ssarherps.org/publications/]),
LOCALITY (use metric for distances and give precise locality data, including lat/long coordinates in decimal degrees and cite the
map datum used), DATE (day-month-year), COLLECTOR, VERIFIED BY (cannot be verified by an author; curator at an institutional
collection is preferred), PLACE OF DEPOSITION (where applicable, use standardized collection designations as they appear in Sabaj
Pérez [ed.]. 2016, Standard Symbolic Codes for Institutional Resource Collections in Herpetology and Ichthyology: An Online Reference,
ver. 6.5, available at http://www.asih.org/resources) and CATALOG NUMBER (required), COMMENTS (brief ), CITATIONS (brief and
must adhere to format used in this section; these should provide a geographic context for the new record). Close with author name(s)
in bold, capital letters (give name and address in full—spell out state or province names—no abbreviations, e-mail address after
each author name/address for those wishing to provide it—e-mail required for corresponding author). Please include distance from
nearest previously known record (provide a citation or refer to existing vouchered material to substantiate your report). If publishing
specific locality information for a rare or endangered species has the potential to jeopardize that population, please consult with the
Section Editor at time of record submission. If field work and/or specimen collection occurred where permits were required, please
include permit number(s) and authorizing agency in the text of the note. Generally, this means that permit information should be
included for any specimens COLLECTED. This is unnecessary for photo vouchers.
Some further comments. The role of the “Standard Names” lists (noted above) is to standardize English names and comment
on the current scientific names. Scientific names are hypotheses (or at least represent them) and as such their usage should not be
dictated by a list, society, or journal.
Additionally, this geographic distribution section does not publish “observation-only” records. Records submitted should be
based on preserved specimens that have been placed in a university or museum collection (private collection depository records are
discouraged; institutional collection records will receive precedence in case of conflict). A good quality photograph (print, slide, or
digital file) may substitute for a preserved specimen. Photographic vouchers must be deposited in a university or museum collection
along with complete locality data, and the photographic catalog number(s) must be included in the same manner as a preserved
record. Before you submit a manuscript to us, check Censky (1988, Index to Geographic Distribution Records in Herpetological Review:
1967–1986; available from the SSAR Publications Secretary), subsequent issues of Herpetological Review, and other sources to make
sure you are not duplicating a previously published record. The responsibility for checking literature for previously documented
range extensions lies with authors. Do not submit range extensions unless a thorough literature review has been completed.
For reports concerning introduced species, it is important to note whether a population has become established or if the report
represents an isolated occurrence, such as a released captive—which in most cases will not justify reporting. Additionally, it will be
helpful to include any information that establishes a timeline for the introduction, such as date of first observation.
Please submit any geographic distribution records in the standard format only to one of the Section Editors: Stuart Nielsen
(Africa and Europe), Indraneil Das (Asia, Australasia, South Pacific), Jerry D. Johnson (Mexico and Central America, including the
Caribbean Basin), Travis Taggart (USA & Canada), or Gustavo J. Scrocchi (South America). Short manuscripts are discouraged and
are only acceptable when data cannot be presented adequately in the standard format. Electronic submission of manuscripts is
required (as Microsoft Word or Rich Text format [rtf] files, as e-mail attachments). Refer to inside front cover for e-mail addresses of
section editors. A template for preparation of geographic distribution notes is available online at: https://ssarherps.org/wp-content/
uploads/2014/07/GeoDistNotes_FormattingGuidelines.pdf.
Recommended citation for new distribution records appearing in this section is: Chacón-Juárez, F., and V. Vásquez-Cruz. 2018.
Geographic distribution: Mexico, Veracruz: Clelia scytalina. Herpetol. Rev. 49:717.

CAUDATA — SALAMANDERS
AMBYSTOMA MACULATUM (Spotted Salamander). USA:
TENNESSEE: CroCKett Co.: Humboldt, 1942 Clarence Norfleet
Road (35.731450°N, 88.965693°W; WGS 84). 1 December 2018.
Chris Brauss. Verified by Chris Gienger. David H. Snyder Museum
of Zoology, Austin Peay State University. (APSU 19900; photo
voucher). One adult found on screened-in porch in area of mixed
woods and fields. New county record (Redmond and Scott 1996.
Atlas of Amphibians in Tennessee. Misc. Publ. No. 12, The Center
of Excellence for Field Biology, Austin Peay State University,
Clarksville, Tennessee. 94 pp. Hard copy and Internet version

Herpetological Review 50(1), 2019

94 GEOGRAPHIC DISTRIBUTION
current through September 2018 [https://www.apsubiology.
org/tnamphibiansatlas/]; 5 Dec 2018). The nearest record (APSU
18895) for this species is ca. 30 km to the northeast at the Milan
Army Ammunition Plant (Johansen and Ream 2008. Herpetol.
Rev. 39:360).
CHRIS BRAUSS, Brauss Farm, 1943 Clarence Norfleet Road, Humboldt,
Tennessee 38343, USA (e-mail: cbrauss@eplus.net); MARK GREENE,
Gibson EMC, 1207 S. College Street, Trenton, Tennessee 38382, USA (e-mail:
greenesnake@yahoo.com); A. FLOYD SCOTT, David H. Snyder Museum of
Zoology, Department of Biology, Austin Peay State University, Clarksville,
Tennessee 37044, USA (e-mail: scotta@apsu.edu).
AMBYSTOMA MACULATUM (Spotted Salamander). USA:
TENNESSEE: Marshall Co.: Chapel Hill, Moss Road (35.668653°N,
86.755015°W; WGS 84). 18 September 2011. Daniel Moss. Verified
by A. Floyd Scott. David H. Snyder Museum of Zoology, Austin
Peay State University (APSU 19159; photo voucher). One adult
observed under a pile of logs and brush. Another individual
(APSU 19897; photo voucher) was found on 21 October 2018
at this same locality while turning soil in an old hog lot that
was last used 20 years ago. New county record (Redmond and
Scott 1996. Atlas of Amphibians in Tennessee. Misc. Publ. No.
12, The Center of Excellence for Field Biology, Austin Peay State
University, Clarksville, Tennessee. 94 pp. Hard copy and Internet
version current through June 2018 [https://www.apsubiology.
org/tnamphibiansatlas/]; 1 Oct 2018).
E. DANIEL MOSS (e-mail: dmoss5@earthlink.net) and CHING LI
WANG, 395 Janet Drive, Pleasant View, Tennessee 37146, USA.
AMBYSTOMA TIGRINUM (Eastern Tiger Salamander). USA:
TENNESSEE: hiCKMan Co.: Centerville, 2135 Jamie Drive
(35.803946°N, 87.426925°W; WGS 84). 8 November 2018.
Rebecca Ruiz. Verified by Chris Gienger. David H. Snyder
Museum of Zoology, Austin Peay State University. (APSU 19899;
photo voucher). One adult found under wooden board next to
driveway adjacent to grassy field. New county record (Redmond
and Scott 1996. Atlas of Amphibians in Tennessee. Misc. Publ.
No. 12, The Center of Excellence for Field Biology, Austin Peay
State University, Clarksville, Tennessee. 94 pp. Hard copy and
Internet version current through September 2018 [https://www.
apsubiology.org/tnamphibiansatlas/]; 13 Dec 2018). The nearest
record (APSU 3161) for this species is ca. 31 km to the north-
northeast at the rear entrance to Montgomery Bell State Park.
REBECCA RUIZ, 2135 Jamie Drive, Centerville, Tennessee 37033, USA
(e-mail: rweikel@gmail.com); A. FLOYD SCOTT, David H. Snyder Museum
of Zoology, Department of Biology, Austin Peay State University, Clarksville,
Tennessee 37044, USA (e-mail: scotta@apsu.edu).
AMBYSTOMA TIGRINUM (Eastern Tiger Salamander). USA:
TEXAS: burleson Co.: ca. 7 km N of Caldwell (30.5995°N,
96.6881°W; WGS 84), 105 m elev. 9 January 2019. Andrew R.
MacLaren. Verified by Carl J. Franklin. Amphibian and Reptile
Diversity Research Center, University of Texas at Arlington
(UTADC 9238–9253; photo voucher). New county record (Dixon
2013. Amphibians and Reptiles of Texas: with Keys, Taxonomic
Synopses, Bibliography, and Distribution Maps. Texas A&M
University Press, College Station, Texas. 447 pp.). This record
fills a gap within surrounding Brazos, Robertson, Milam, and
Lee counties where it is known to occur (Dixon 2013, op. cit.).
Two adult specimens observed in fine sand soil, within a rodent
(potentially Geomys sp.) burrow; only one captured (female, 102
mm SVL, 208 mm TL, 41 g). The nearest specimen in collection
Herpetological Review 50(1), 2019
comes from Milam County, ca. 28.3 km northwest (Biodiversity
Collections, University of Texas at Austin [TNHC] 6168).
Specimen collected under Scientific Permit SPR-0102-191 issued
to MRJF by Texas Parks and Wildlife Department.
ANDREW R. MACLAREN (e-mail: arm107@txstate.edu) and MICHAEL
R. J. FORSTNER, Department of Biology, Texas State University, 601
University Drive, San Marcos, Texas 78666, USA.
DESMOGNATHUS FUSCUS (Northern Dusky Salamander).
USA: INDIANA: SpenCer Co.: Luce Township (37.85110°N,
87.16342°W; WSG 84). 6 July 2018. Michael J. Lodato, Nathan J.
Engbrecht, and Elizabeth M. Johnson. Verified by Daniel B. Wylie.
Illinois Natural History Survey (INHS 31187). New county record
(Minton 2001. Amphibians and Reptiles of Indiana. Indiana
Academy of Science, Indianapolis, Indiana. 404 pp.). In addition
to the collected voucher specimen, numerous individuals of
various age classes were observed under cover in a forested
seep spring complex proximate to the Ohio River. This locality
represents the westernmost outpost for this species in Indiana
and is disjunct and isolated from the more contiguous range for
the species to the east. The nearest Indiana record, also disjunct,
is ca. 67 km to the northeast in adjoining Perry County, Indiana
(Minton 2001, op. cit.). In addition, this new county record is
isolated and disjunct from any proximate records in the Ohio
River border counties and adjacent region of Kentucky, where
contiguous populations are known both east and west of the
locality reported here (J. MacGregor, pers. comm.). Specimen
collected under Indiana Dept. Nat. Res. Fish & Wildlife License
No. 1804290185150.
MICHAEL J. LODATO, 8438 Pelican Pointe Drive, Evansville, Indiana
47725, USA (e-mail: mikelodato@wowway.com); NATHAN J. ENGBRECHT
(e-mail: nengbrecht@dnr.in.gov) and ELIZABETH M. JOHNSON (e-mail:
ejohnson1@dnr.in.gov), Indiana Department of Natural Resources,
5596 East State Road 46, Bloomington, Indiana 47401, USA; and LEE A.
CASEBERE, 9202 N. Delaware Street, Indianapolis, Indiana 46240, USA.
DESMOGNATHUS FUSCUS FUSCUS (Northern Dusky
Salamander). USA: SOUTH CAROLINA: piCKens Co.: Table Rock
State Park (35.032714°N, 82.701303°W; WGS 84). 5 June 2016.
C. Little, J. Newman, and K. Barrett. Verified by Sean Graham.
James Scudday Vertebrate Collections, Sul Ross State University
(SRSU-D 127; photo voucher). One individual was observed
on a rock outcrop. New county record (http://vertnet.org,
http://carolinaherpatlas.org; 5 June 2016). Specimen found
in an intergrade zone that is inhabited by the Spotted Dusky
Salamander (Desmognathus conanti), which the the Northern
Dusky Salamander is known to breed with. This record fills a
distributional gap between Oconee County and Spartanburg
County, as the nearest known record of Desmognathus fuscus
fuscus is in Oconee County (34.7244°N, 83.0764°W; WGS 84),
which is ca. 47 km away from our record (North Carolina State
Museum [NCSM] 64156)
COLE LITTLE (e-mail: colemal@g.clemson.edu), JILLIAN C. NEWMAN
(e-mail: jcnewma@g.clemson.edu), and KYLE BARRETT, 261 Lehotsky
Hall, Box 340310, Clemson, South Carolina 29631, USA (e-mail: rbarre2@
clemson.edu).
ENSATINA ESCHSCHOLTZII (Ensatina). USA: CALIFORNIA:
solano Co.: Vaca Mountains, Mix Canyon Road, ca. 1 km (by
road) E of intersection with Blue Ridge Road (38.412631°N,
122.101594°W; WGS 84), 666 m elev. 22 November 2018. Erica J.
Ely and F. E. Ely. Verified by David B. Wake. California Academy

of Sciences (CAS 263472). Gravid adult (69 mm SVL, 122 mm
total length) was found dead on road during heavy fall rain
(54°F) at 2200 h. First verified county record that confirms
presence in predicted range (Stebbins 2003. A Field Guide to
Western Reptiles and Amphibians. 3rd Edition. Houghton Mifflin
Company, Boston, Masschusetts. 469 pp.). This location is on
the margin of subspecies ranges of E. e. oregonensis and E. e.
xanthoptica (Kuchta et al. 2009. Mol. Phylogenet. Evol. 50:240–
255). Specimen exhibits E. e. xanthoptica ventral coloration,
but lacks the diagnostic yellow eyes (D. Wake, pers. comm.),
thus no subspecies has been designated. Closest records are
an E. e. xanthoptica, ca. 10 km southwest, near Hwy 121, Napa
County, California (Museum of Vertebrate Zoology, University of
California, Berkeley [MVZ] 100398), as well as E. e. oregonensis
(MVZ 158111, 158112) and E. e. xanthoptica (MVZ 158110)
ca. 12.5 km west-northwest, near Atlas Peak, Napa County,
California. Specimen collected under California Department of
Fish and Wildlife, Scientific Collecting Permit SC-12323, issued
to L. A. Scheinberg.
ERICA J. ELY (e-mail: eely@calacademy.org) and F. E. ELY, Department
of Herpetology, California Academy of Sciences, 55 Music Concourse Drive,
San Francisco, California 94118, USA.
ensATinA esCHsCHOLTZii KLAUBeri (Large-blotched
Ensatina). USA: ARIZONA: gila Co.: Tonto Creek State Fish
Hatchery (34.37796°N, 111.09923°W, WGS 84), ca. 1890 m elev.
Autumn 2003. Andrew Baldwin. Verified by A. T. Holycross. Arizona
State University (ASU HP00371; photo voucher). Two individual
E. e. klauberi were discovered within several hundred meters
of the facilities of Tonto Creek State Fish Hatchery. Subsequent
investigation found that 22 individuals of this species had been
collected from a population located in the Palomar Mountains,
San Diego County, California and were intentionally and illegally
released at this site in April 1980 by an unnamed individual (T.
Jones, pers. comm.). Since 2003, E. e. klauberi has been observed
at this site on several occasions. Sightings of two individuals
(34.37792°N, 111.09889°W) on 22 November 2018 (ASU HP00370;
Dave Weber, pers. comm.) demonstrate the population is
extant. Fragments of mitochondrial DNA (cytochrome b) were
sequenced from the two individuals captured in 2003, compared
to one another and to published gene sequences for E. e. klauberi
from Palomar Mountains, San Diego County, California. The
ca. 400-bp sequences of the two specimens were identical to
each other and 0.01% divergent from the reference population
in California. The original release site is 3.2 km from the only
current site from where they are now collected. This introduced
population of salamanders offers a relatively unique opportunity
to investigate many aspects of the theoretical biology of founder
populations (especially genetic effects), given that the source
population, the date of introduction, and the number of founders
are known parameters.
ANDREW S. BALDWIN, Mesa Community College, 1833 West
Southern Avenue, Mesa, Arizona 85202, USA; e-mail: andy.baldwin@
mesacc.edu.
HEMIDACTYLIUM SCUTATUM (Four-toed Salamander).
USA: ARKANSAS: Saline Co.: 7.8 km E of Haskell, on north side
of County Road 190 (34.498479°N, 92.551939°W; WGS 84), ca.
83 m elev. 2 February 2017. C. Vick. Verified by L. Neuman-
Lee. Arkansas State University, Museum of Zoology (ASUMZ
33851; photo voucher). The salamander was found during
the day beneath a log lying on a moss bed. This locality is
Herpetological Review 50(1), 2019
GEOGRAPHIC DISTRIBUTION 95
situated within the sandy soils of the western Gulf Coastal Plain
ecoregion, in the second growth hardwood canopy forest of an
unnamed headwater stream valley that drains into the Saline
River. Hemidactylium is widely distributed across the Interior
Highlands and Ouachita Mountains of west-central Arkansas
(Trauth et al. 2004. The Amphibians and Reptiles of Arkansas.
University of Arkansas Press, Fayetteville, Arkansas. 421 pp.), yet
this species is infrequently encountered, as preferred sphagnum
moss wetland microhabitat is localized in occurrence. This
record is 18 km SE of the nearest vouchered specimens in the
Ouachita Mountains of central Saline County (Amphibian
and Reptile Diversity Research Center, University of Texas
Arlington [UTA] 56604, 56605). However, it is significant as it
is the first report of the species from the western Gulf Coastal
Plain physiographic province (Powell et al. 2016. Peterson Field
Guide to Reptiles and Amphibians of Eastern and Central North
America. Houghton Mifflin Harcourt Publishing, New York, New
York. 494 pp.). A subsequent search of this site by C. Vick on 15
March 2018 yielded an adult Hemidactylium attending an egg
clutch found under a sphagnum cap on a root overhanging the
margin of a woodland pool formed by an upended root mat of
a large blown down tree. An additional unattended egg clutch
was found under a nearby moss cap on the margin of the same
pool. Caudate species associates observed in proximity to this
stream valley during the course of the two site visits include:
Ambystoma maculatum (Spotted Salamander), Desmognathus
brimleyorum (Ouachita Dusky Salamander), Eurycea paludicola
(Western Dwarf Salamander), and Plethodon albagula (Western
Slimy Salamander). The presence of E. paludicola lends
support in classifying the locality as coastal plain habitat, as
this species does not ascend into the highlands of the Ouachita
Mountains where it is replaced by Eurycea multiplicata (Many-
ribbed Salamander). It should be noted that the range of D.
brimleyorum (ID confirmed via genetic data by D. Shepard,
pers. comm.), extends south-southeast well into the Gulf Coastal
Plain along the Ouachita River basin. Sphagnum moss capped
margins of predator free ephemeral woodland pools is the
optimal microhabitat structure for successful reproduction in
Hemidactylium. These specialized microhabitat requirements
are likely a limiting factor that has influenced the spotty nature
of the distribution of this species. These microhabitats are in
turn very vulnerable to disturbance and/or destruction from
the heavy machinery used in modern industrial timber harvest
practices, which has dominated land use throughout the
western Gulf Coastal Plain of southern Arkansas over the past
50 plus years. It would not be unreasonable to suggest that such
landscape level impacts may have greatly reduced or eliminated
unknown historic populations of Hemidactylium in the Gulf
Coastal Plain of Arkansas. We thank L. Neuman-Lee, K. Roberts,
and D. Shepard for their assistance in the preparation of this
note.
CALVIN VICK, 111 Village Drive, Benton, Arkansas 72015, USA
(e-mail: calvinhalo@gmail.com); KELLY J. IRWIN, Arkansas Game & Fish
Commission, 915 E. Sevier St., Benton, Arkansas 72015, USA (e-mail: kelly.
irwin@agfc.ar.gov).
SIREN INTERMEDIA (Lesser Siren). USA: TEXAS: Kinney Co.:
Imperialist Creek, TX Hwy 131, ca. 2.6 rd km S jct Co Rd 1908
(29.12682°N, 100.44236°W; WGS 84). 11 November 2018. Drew
R. Davis and Krista M. Ruppert. Verified by Travis J. LaDuc.
Biodiversity Collections, University of Texas at Austin (TNHC
112457 [DRD 5156]). Adult individual (249.5 g, 320 mm SVL, 149
96 GEOGRAPHIC DISTRIBUTION
mm tail length) trapped in a baited minnow trap at 0915 h. New
county record extending the western boundary of this species in
Texas (Dixon 2013. Amphibians and Reptiles of Texas: with Keys,
Taxonomic Synopses, Bibliography, and Distribution Maps.
Third Edition. Texas A&M University Press, College Station,
Texas. viii + 447 pp.) and across its known range (Petranka 1998.
Salamanders of the United States and Canada. Smithsonian
Institution Press, Washington, D.C. xvi + 587 pp.; Martof 1973.
Siren intermedia. Cat. Am. Amphib. Rept. 127:1–3). This site was
a shallow (ca. 1 m deep) pool with abundant aquatic vegetation
with limited connectivity to other nearby pools along Imperialist
Creek and is within the Rio Grande watershed. The nearest
known records to this individual are from ca. 28 km to the
southwest from specimens collected in 1880 near Normandy,
Maverick County, Texas (National Museum of Natural History,
Smithsonian Institution [USNM] 10856–10862; Goin 1957.
Herpetologica 13:37–42). We currently recognize this individual
as S. intermedia but acknowledge that the taxonomic status of
this species has not yet been fully resolved. This specimen was
collected under a Texas Parks and Wildlife Scientific Collecting
Permit (SPR-1018-294) issued to DRD.
DREW R. DAVIS (e-mail: drew.davis@utrgv.edu), KRISTA M.
RUPPERT, and RICHARD J. KLINE, School of Earth, Environmental, and
Marine Sciences, University of Texas Rio Grande Valley, 100 Marine Lab
Drive, South Padre Island, Texas 78597, USA.
TARICHA SIERRAE (Sierra Newt). USA: CALIFORNIA: sierra
Co.: Fiddle Creek, tributary to the North Fork Yuba River, ca.
150 m upstream of Brandy City Road crossing (39.52552°N,
120.99223°W; WGS 84), 2480 ft. elev. 5 July 2017. K. D. Wiseman.
Verified by Erica J. Ely. California Academy of Sciences (CAS
263152). Individual (71 mm SVL, 167 mm total, 10 g) found
underwater (6 cm depth; 14°C water temperature) in ca. 5 × 6 m
instream pool at 1330 h. This specimen represents first record of
species in Sierra County (Vindum and Koo 1999. Amphibians and
Reptiles of the Tahoe National Forest: Historical Collections and
Results of 1997–1999 California Academy of Sciences Surveys. 27
pp.), extending their range ca. 8.6 km SE (CAS 205601) and 8.3–
9.2 km NE of specimens (CAS 205227, 205240, 208701) collected
in Yuba County. Specimen was collected under CDFW permit
#006634.
KEVIN D. WISEMAN, Garcia and Associates, 2601 Mission St. Suite
600, San Francisco, California 94110, USA; Department of Herpetology,
California Academy of Sciences, 55 Music Concourse Drive, San Francisco,
California 94118, USA; e-mail: kwiseman@garciaandassociates.com.
ANURA — FROGS
ELEUTHERODACTYLUS PLANIROSTRIS (Greenhouse Frog).
MEXICO: QUINTANA ROO: MuniCipio baCalar: Bacalar, Street 30
between Av. 3 and Av. 5 (18.68240°N, 88.38804°W; WGS 84), 15 m
elev. 3 February 2018. Pablo M. Beutelspacher-García. Verified by
Fred Kraus. Herpetological Collection of El Colegio de la Frontera
Sur, Unidad Chetumal (ECO-CH-H4087–4088). First municipality
records and southernmost locality on the Caribbean versant of
Mexico for this exotic species. Previous reported populations in
Quintana Roo were from the northeastern portion of the state
from Playa del Carmen (Cedeño-Vázquez et al. 2014. Rev. Mex.
Biodiv. 85:650–653), Cozumel Island (Pavón-Vázquez et al. 2016.
Mesoamer. Herpetol. 3:531–533), and Cancún (García-Balderas
et al. 2016. Mesoamer. Herpetol. 3:173). The two frogs were found
in the morning during the removal of stones and tree branches
Herpetological Review 50(1), 2019
from an abandoned field surrounded by houses. The frogs were
collected under permit SGPA/DGVS/01205/17 issued to Fausto
R. Méndez de la Cruz by SEMARNAT, with an extension to JRCV.
J. ROGELIO CEDEÑO-VÁZQUEZ, El Colegio de la Frontera Sur.
Departamento de Sistemática y Ecología Acuática, Av. Centenario Km 5.5,
C.P. 77049 Chetumal, Quintana Roo, México (e-mail: rcedenov@ecosur.mx);
PABLO M. BEUTELSPACHER-GARCÍA, Martinica 342, Fracc. Caribe, C.P.
77086 Chetumal, Quintana Roo, México (e-mail: botrhopspacher@hotmail.
com).
ELEUTHERODACTYLUS PLANIROSTRIS (Greenhouse Frog).
USA: TEXAS: harris Co.: Houston, Hazard St, ca. 50 m S of the jct
with Vermont St (29.748721°N, 95.407088°W; WGS 84), 16 m elev.
23 September 2018. Suzanne Simpson. Verified by Carl Franklin.
Amphibian and Reptile Diversity Research Center, University
of Texas at Arlington (UTADC 9236, 9237; photo vouchers). An
individual was observed crossing the driveway of a residence
and was photographed then released. New county record (Dixon
2013. Amphibians and Reptiles of Texas: with Keys, Taxonomic
Synopses, Bibliography, and Distribution Maps. Texas A&M
University Press, College Station, Texas. 447 pp.). This represents
the fourth county in Texas from which this species has been
observed. The other records are from Tarrant County from 1991
(UTA 16617), Galveston County from 1999 (Biodiversity Research
and Teaching Collection, Texas A&M University [TCWC] 83067),
and Aransas County from 2018 (iNaturalist 15331253).
SUZANNE E. SIMPSON, Bayou Land Conservancy, 10330 Lake Road
Bldg J, Houston, Texas 77070, USA (e-mail: ssimpson@bayouland.org);
PAUL S. CRUMP, Texas Parks and Wildlife Department, Nongame and
Rare Species Program, 4200 Smith School Road, Austin, Texas 78744, USA
(e-mail: paul.crump@tpwd.texas.gov); TOBY J. HIBBITTS, Biodiversity
Research and Teaching Collection, Department of Wildlife and Fisheries
Sciences, Texas A&M University, 2258 Tamu, College Station, Texas 77843,
USA (e-mail: thibbitts@tamu.edu).
GASTROPHRYNE CAROLINENSIS (Eastern Narrow-mouthed
Toad). USA: TEXAS: Kleberg Co.: North Padre Island, within Padre
Island National Seashore, 0.37 km SE of the intersection of Bird
Island Basin Road and Park Road 22 (27.46202°N, 97.28760°W;
NAD 83), 1.5 m elev. 6 August 2015. Alicia Walker. Verified by
Travis J. LaDuc. Biodiversity Collections, University of Texas at
Austin (TNHC 96013, 96014). First specimens for this county
(Dixon 2013. Amphibians and Reptiles of Texas, with Keys,
Taxonomic Synopses, Bibliography, and Distribution Maps.
Texas A&M University Press, College Station, Texas. 447 pp.).
Although G. carolinensis has been audio-recorded on the island
for over a decade, live specimens were first collected on 6 August
2015. Specimen was collected under permit number PAIS-2015-
SCI-0009 issued by the National Park Service.
ALICIA WALKER, National Park Service, 20301 Park Rd 22, Corpus
Christi, Texas 78418, USA; e-mail: alicia_walker@nps.gov.
HYLA CHRYSOSCELIS (Cope’s Gray Treefrog). USA:
PENNSYLVANIA: washington Co.: ca. 3 km S of West Alexander,
near Old Brick Road (40.07790°N, 80.50171°W; WGS 84). 21 June
2013. Brandon M. Ruhe. Verified by Nathan H. Nazdrowicz and
Thomas C. LaDuke. East Stroudsburg University (ESUP A00257;
audio voucher) and Pennsylvania Amphibian and Reptile
Survey (PARS 10448; audio voucher). Chorus with a call pulse
repetition rates >50 p/s recorded from a floodplain wetland
complex along an unnamed tributary to the Wheeling Creek.
New state record for Pennsylvania (McCoy 1982. Amphibians

and reptiles in Pennsylvania: Checklist, Bibliography, and Atlas
of Distribution. Spec. Publ. Carnegie Mus. Nat. Hist. No. 6.
Pittsburgh, Pennsylvania. 91 pp.), and an extension of more than
60 km north-northwest of the nearest record in West Virginia
(Little and Wiley 1989. J. Herpetol. 22:299–303).
BRANDON M. RUHE, The Mid-Atlantic Center for Herpetology and
Conservation, P.O. Box 620, Oley, Pennsylvania, 19547, USA (e-mail: bruhe@
machac.org); JASON POSTON, The Mid-Atlantic Center for Herpetology
and Conservation, P.O. Box 620, Oley, Pennsylvania, 19547, USA;
CHRISTOPHER A. URBAN, The Pennsylvania Fish & Boat Commission,
Centre Region Office, 595 E. Rolling Ridge Drive, Bellefonte, Pennsylvania
16823, USA.
HYLA VERSICOLOR (Gray Treefrog). USA: MAINE: aroostooK Co.:
State Route 11, Mile 24, T8 R5 WELS (46.306017°N, 68.356893°W;
WGS 84), 320 m elev. 12 June 2018. Scott B. Lindemann. Verified
by Greg Watkins-Colwell. Yale Peabody Museum of Natural
History (YPM 23340).
penobsCot: Billfish Brook, T6 R8 WELS (46.135686°N,
68.798687°W; WGS 84), 190 m elev. 12 June 2018. Scott B.
Lindemann. Verified by Greg Watkins-Colwell. YPM 23341.
Both specimens collected at night from chorusing
populations in palustrine emergent wetland bordered by mixed
coniferous/deciduous forest. First verified Aroostook County
record, and northernmost record from adjacent Penobscot
County. An additional chorus was heard by SBL and Remi
Wagner from similar habitat near US Route 1 in Monticello,
Aroostook County (46.301752°N, 67.841723°W; WGS 84; 130
m elev.) on 17 June 2018. Together, these records extend the
known distribution 28 km north of the closest unverified report
in Hodgdon, Aroostook County and 45 km north-northeast of
the nearest verified (photo) record from T3 R7 WELS, Penobscot
County (Hunter et al. 1999. Maine Amphibians and Reptiles.
University of Maine Press, Orono, Maine. 252 pp.; Maine
Amphibian and Reptile Atlas Project data, Maine Department
of Inland Fisheries and Wildlife, Bangor, Maine). This extension
of the documented range of Gray Treefrog may represent recent
range expansion, similar to presumed range expansion reported
from northeastern New Brunswick, Canada (McAlpine et al.
2009. Can. Field Nat. 123:309–312). Work was performed under
a Maine Department of Inland Fisheries and Wildlife scientific
collection permit (#2018-184) issued to SBL.
SCOTT B. LINDEMANN (e-mail: lindemann.scott@gmail.com) and
AIDAN M. O’BRIEN, Department of Wildlife, Fisheries, and Conservation
Biology, University of Maine, 5755 Nutting Hall, Room 210, Orono, Maine
04469, USA (e-mail: aidan.obrien@maine.edu).
LITHOBATES PIPIENS (Northern Leopard Frog). USA: MAINE:
aroostooK Co.: mouth of Musquacook Stream on Allagash River,
T13 R12 WELS (46.81501°N, 69.23294°W; WGS 84), 230 m elev.
16 June 2017. David E. Putnam, Caleb Ward, Alyssa Dougherty,
Gannon Pratt, and Samantha Boyce. Verified by Greg Watkins-
Colwell. Yale Peabody Museum of Natural History (YPM 23337;
photo voucher). One adult of unknown sex was photographed
in a Silver Maple (Acer saccharinum) swamp, adjacent to a
floodplain emergent wetland with grasses and sedges. This
record extends the known distribution 57 km WNW and 55
km WNW of the closest previous records, from T10 R7 WELS
township and Eagle Lake, Aroostook County, respectively, and
partially fills a distribution gap in northwestern Maine (Hunter
et al. 1999. Maine Amphibians and Reptiles. University of Maine
Press, Orono, Maine. 252 pp.; Maine Amphibian and Reptile Atlas
Herpetological Review 50(1), 2019
GEOGRAPHIC DISTRIBUTION 97
Project files, Maine Department of Inland Fisheries and Wildlife,
Bangor, Maine).
SCOTT B. LINDEMANN (e-mail: lindemann.scott@gmail.com) and
AIDAN M. O’BRIEN, Department of Wildlife, Fisheries, and Conservation
Biology, University of Maine, 5755 Nutting Hall, Room 210, Orono, Maine
04469, USA (e-mail: aidan.obrien@maine.edu); DAVID E. PUTNAM,
University of Maine at Presque Isle, 214 South Hall, Presque Isle, Maine
04769, USA (e-mail: david.putnam@maine.edu).
OSTEOPILUS SEPTENTRIONALIS (Cuban Treefrog). USA:
GEORGIA: glynn Co.: Jekyll Island State Park (see localities
below). 19 September–3 November 2017. Christian A. Jarboe,
Heather L. Platt, Yank Moore, and Joseph Colbert. Verified
by Coleman Sheehy. Florida Museum of Natural History (UF
181912–181915). On 17 June 2012, a visitor on the north end
of Jekyll Island photographed a single O. septentrionalis near
an urbanized restaurant-hotel complex. We first captured
O. septentrionalis on the island ca. 200 m away from the 2012
record on 24 May 2013 after observing one in a PVC pipe hung
on a Quercus hemisphaerica (Darlington Oak; 31.09837°N,
81.40607°W; WGS 84; this individual is now included in Jekyll
Island’s collection as specimen JI-Osep-02). A second individual
was captured on 18 October 2013 in another pipe within 10 m of
the initial capture. On 28 October 2013, a single O. septentrionalis
was heard calling near a golf course in the middle of the island
(31.01545°N, 81.41809°W; WGS 84). On 15 May 2017, a single O.
septentrionalis was recorded during a frog-call survey near the
complex noted above (31.09603°N, 81.40405°W; WGS 84). On
11 July 2017, an individual was photographed in an ephemeral
wetland located on the southern end of the island near a youth
education facility (31.01550°N, 81.42567°W; WGS 84). On 23 July
2017, two more individuals were detected during frog call surveys
by the restaurant-hotel complex and one was captured on 19
September 2017 while it was resting in the filter equipment of
an artificial water feature (UF 181912). Afterwards we increased
our sampling effort at this site and a total of 26 individuals were
collected. Finally, on 3 November 2017, a single individual (UF
181915) was located and collected from a building near the
center of the island (31.06439°N, 81.41994°W; WGS 84). A total
of 17 specimens collected from 19 September–3 November 2017
had an average snout-vent length of 58.6 mm, median length of
55.5 mm, and range of 42.1–83.1 mm. Given that three dozen
O. septentrionalis of varying size classes have been observed
throughout the 23 km2 island over the last few years, we believe it
is likely they are breeding and established.
The first documented record of O. septentrionalis from Georgia
is a specimen collected on 23 September 2004 in Chatham
County (Johnson 2007. Herpetol. Rev. 38:349). Jensen et al. (2008.
Amphibians and Reptiles of Georgia. University of Georgia Press,
Athens, Georgia. 575 pp.) added a 2006 record from Glynn County
and noted that Cuban Treefrogs “may eventually colonize parts of
coastal and southern Georgia.” Additional Georgia records were
later collected in 2007 (Spalding County: UF 151666), 2012 (Walton
County: confirmed by J. B. Jensen, unpubl.), 2013 (Wilkinson
County: USGS Nonindigenous Aquatic Species database,
specimen ID 616791), and 2017 (Dougherty County: UF 162559).
Although these individual records have been documented in the
state, we believe we have generated sufficient evidence to conclude
for the first time that a breeding population of O. septentrionalis is
established in the state on Jekyll Island, where we have collected
31 individuals to date and confirmed another five records either
through photographs or auditory surveys.
98 GEOGRAPHIC DISTRIBUTION
Individuals located at our urban site were associated with
ornamental plants, specifically Ravenala madagascariensis
(Traveler’s Palm). The spread of O. septentrionalis has been
documented via the transportation of ornamental plants
throughout Florida (Johnson 2013. Univeristy of Florida IFAS
Extension. WEC218. 7 pp.) and the Caribbean (Grenada:
Somma and Graham 2015. IRCF Rept. Amphib. 20:40–42; Virgin
Islands: Perry 2009. Appl. Herpetol. 6:191–192). It is uncertain
whether O. septentrionalis was transported to Jekyll Island
through landscape plants but we believe this is the most likely
explanation. All specimens were collected under Scientific
Collecting Permit #029, issued by Georgia DNR. Special thanks
to J. B. Jensen for assistance and feedback in the documentation
of this occurrence.
CHRISTIAN A. JARBOE (e-mail: ajarboe93@gmail.com), JOSEPH E.
COLBERT (e-mail: jcolbert@jekyllisland.com), YANK MOORE, Jekyll Island
Authority, Conservation Department, Jekyll Island, Georgia 31527, USA
(e-mail: ymoore@jekyllisland.com); DAVID STEEN, Jekyll Island Authority,
Georgia Sea Turtle Center, 100 James Road, Jekyll Island, Georgia 31527,
USA (e-mail: dsteen@jekyllisland.com); STEVE A. JOHNSON, University
of Florida, Department of Wildlife Ecology and Conservation, Gainesville,
Florida 32611, USA (e-mail: tadpole@ufl.edu).
OSTEOPILUS SEPTENTRIONALIS (Cuban Treefrog). USA:
MASSACHUSSETTS: haMpden Co.: Chicopee (42.135145°N,
72.618750°W; WGS 84). 2 February 2018. Brianne Zulkiewicz.
Verified by Lance McBrayer. Georgia Southern University –
Savannah Science Museum Herpetology Collection (GSU 26257;
photo voucher). We report the first state record for the exotic
Osteopilus septentrionalis in Massachusetts. One individual frog
was discovered in a potted plant purchased from a department
store garden section; presumably the plant originated from an
area where the species is established. Osteopilus septentrionalis
are known to spread through shipments of ornamental plants
(Krysko et al. 2011. Zootaxa 3028:1–64); however, we suggest
the relatively cold climate of Massachusetts will prohibit O.
septentrionalis from becoming invasive.
CHRISTIAN A. JARBOE (e-mail: ajarboe93@gmail.com) and JOSEPH
E. COLBERT, Jekyll Island Authority, Conservation Department, Jekyll
Island, Georgia 31527, USA (e-mail: jcolbert@jekyllisland.com); BRIANNE
ZULKIEWICZ, Pioneer Valley, Massachusetts, USA; DAVID STEEN, Georgia
Sea Turtle Center, Jekyll Island Authority, 100 James Road, Jekyll Island,
Georgia 31527, USA.
PRISTIMANTIS MORO (Honduran Robber Frog). REPUBLIC OF
PANAMA: VERAGUAS: Santa Fe distriCt: Alto Ortiga, Los Olivos
(8.61208°N, 81.17661°W; WGS 84), 383 m elev. 3 September 2016.
E. E. Flores. Verified by Abel Batista. Amphibian and Reptile
Diversity Research Center, University of Texas at Arlington
(UTADC 9176; photo voucher). This constitutes the second
known locality on the Caribbean versant of Panama, extending
the species’ range on that side 199 km west from Cerro Bruja,
Colon Province (Círculo Herpetológico de Panamá [CHP] 6109),
and 9 km northwest of the closest record on the Pacific versant
from Cerro Negro, Veraguas, Province (Carrizo 2010. Riqueza
y Abundancia de la herpetofauna de la Cuenca Alta del Río
Santa Maria, Santa Fe, Veraguas. Tesis de Maestría. Universidad
Autónoma de Chiriquí, Ciudad de David. 103 pp.). The juvenile
frog was found at 1130 h hidden inside a bromeliad on a tree
approximately 7 m above ground. This remote locality is also
at the lowest elevation reported for the species at 383 m; 167
m lower than 550 m (Köhler 2011. Reptiles of Central America.
Herpetological Review 50(1), 2019
Herpeton, Verlag Elke Köhler, Offenbach, Germany. 400 pp.).
Field work was supported with funds provided by Sistema
Nacional de Investigación de Panamá (SNI).
ERIC ENRIQUE FLORES, Sistema Nacional de Investigación de
Panamá (SNI) & Smithsonian Tropical Research Institute, Apartado 0923-
00126, Santiago de Veraguas, Panama (e-mail: sailax1@gmail.com);
RACHEL PAGE, Smithsonian Tropical Research Institute, Apartado 0843-
03092, Panama (e-mail: pager@si.edu); DINEILYS APARICIO, University of
Panama, Veraguas, Panamá (e-mail: apariciodineilys@hotmail.com); ENOC
BORDONES, Alto Ortiga community, Santa Fe, Panama.
PTYCHOHYLA HYPOMYKTER (Copan Stream Frog)
GUATEMALA: El QUICHÉ: MuniCipality oF Chajul: Batzchocolá
(15.5722°N, 91.1139°W; WGS 84), 1315 m elev. 10 June 2017.
Sergio A. González-Mollinedo and Guillermo A. Mármol-
Acatán. Verified by Jonathan A. Campbell. Reference Collections,
Universidad del Valle de Guatemala (UVGF 0006; photo voucher).
Another individual (UVGF 0007 [photo voucher]) was collected
from Ixlaj (15.4867°N, 91.9662°W; WGS 84; 1832 m elev.) on 15
August 2017. First records for El Quiché Department, with the
nearest records in Guatemala being from 32 km to the northwest
of the position reported for the Batzchocolá locality at Finca El
Valle, Barillas, Huehuetenango (Amphibian and Reptile Diversity
Research Center, University of Texas at Arlington [UTA] A-52606–
52609, A-55165).
MuniCipality oF san juan CotZal: Xeputul I. (15.4806°N,
90.9033°W; WGS 84), 1648 m elev. 17 August 2017. Sergio A.
González-Mollinedo and Guillermo A. Mármol-Acatán. Verified
by Jonathan A. Campbell. UVGF 0010 (photo voucher). First
record for the municipality, with the closest known locality
for this record being 63 km to the west at Finca Pueblo Nuevo,
Municipality of Panzos, Alta Verapaz (Duellman 2001. The Hylid
Frogs of Middle America, 2 Volumes. SSAR Contrib. Herpetol. 18.
1158 pp.).
The Batzchocolá site is a transition between cloud forest and
pine-oak forest, Ixlaj is within remnant cloud forest next to a
stream surrounded by cropland and pastures, and Xeputul I is
situated in cloud forest adjacent to agricultural fields. All three
localities are adjacent to the Visis Cabá Biosphere Reserve, which
suggest the species is present there and along other mountain
slopes in the area where these forest types occur. Other frogs
found coexisting with P. hypomykter included: Craugastor
lineatus, C. xucanebi, Hyalinobatrachium fleischmanni,
Plectrohyla hartwegi, P. ixil, and P. quecchi.
SERGIO A. GONZÁLEZ-MOLLINEDO (e-mail: sergiogonmoll@gmail.
com) and GUILLERMO A. MÁRMOL-ACATÁN, Department of Biology,
Universidad del Valle de Guatemala, 18 Ave. 11-95 zone 15, Vista Hermosa
III, Guatemala, Guatemala (e-mail: gamarmol@gmail.com).
SCAPHIOPUS HOLBROOKII (Eastern Spadefoot). USA:
INDIANA: spenCer Co.: ca 1.0 km S of Sand Ridge, Luce Township
on CR 600W (37.89286°N, 87.16935°W; WGS 84). 8 September
2018. Michael J. Lodato and Noah M. Gordon. Verified by Dan
Wylie. Illinois Natural History Survey (INHS 31187). Additionally,
a breeding chorus (INHS 2018h; audio voucher) was recorded at
a site ca. 4.0 km E of Richland City, Luce Township on CR 425N
(37.94533°N, 87.13042°W; WGS 84) and was verified by Chris
Phillips. One of us (MJL), in conjunction with Scott Dugas and
David Ayer, noted spadefoots at the Richland City site following
heavy rains on 14 April 2002. No specimens or photographs were
taken. Thus, this report provides the first vouchered record of the
species in the county (Minton 2001. Amphibians and Reptiles of

Indiana. Indiana Academy of Science, Indianapolis, Indiana. 404
pp.). The appearance of Scaphiopus on 8 September 2018 was
triggered by rainfall in excess of 65 mm during the previous 24
h. Spadefoots were again active at each of these sites following
heavy rains on 25 September 2018, but chorusing was not
detected. Land use at each site is agricultural, primarily row
crop. In addition to the collected vouchers from the Sand Ridge
site, numerous individuals were observed at both sites on wet
roads and/or in adjacent fields. These localities lay ca. 61 km
west of the nearest record for the species in adjacent Perry
County, Indiana (Minton 2001, op. cit.) and ca. 65 km east of
known populations in Posey County, Indiana (Lodato et al. 2002.
Herpetol. Rev. 33:64; Museum of Zoology, University of Michigan
[UMMZ] 230111), thus filling the distribution gap for this species
in southwest Indiana. There are no records for the species from
the adjacent western coalfield of Kentucky (J. MacGregor, pers.
comm.). Specimens collected under Indiana Department of
Natural Resources Fish & Wildlife License No. 1804290185150.
MICHAEL J. LODATO, 8438 Pelican Pointe Drive, Evansville, Indiana
47725, USA (e-mail: mikelodato@wowway.com); NOAH M. GORDON,
Department of Biology, University of Evansville, 1800 Lincoln Avenue,
Evansville, Indiana 47722, USA (e-mail: ng62@evansville.edu).
TESTUDINES — TURTLES
CHELYDRA SERPENTINA (Snapping Turtle). USA: MAINE:
pisCataquis Co.: Churchill Dam on Allagash River, T10 R12 WELS
(46.49274°N, 69.28813°W; WGS 84). 280 m elev. 24 August 2016.
Robert Johnston. Verified by Greg Watkins-Colwell. Yale Peabody
Museum of Natural History (YPM 21969; photo voucher). One
adult turtle of unknown sex was photographed walking across
the top of the dam. Churchill Dam lies at the base of Churchill
Lake, one of many lakes on the Allagash River system. This
record extends the known distribution of C. serpentina into
northwestern Maine (Hunter et al. 1999. Maine Amphibians
and Reptiles. University of Maine Press, Orono, Maine. 252 pp.;
Maine Amphibian and Reptile Atlas Project [MARAP] files, Maine
Department of Inland Fisheries and Wildlife, Bangor, Maine).
This record extends the known distribution 32 km WNW of the
nearest photographic record from T8 R9 WELS township (2002),
and 37 km WSW from the next closest record, a 1960 unverified
sight report from T11 R9 WELS township (MARAP files). An
additional sight report by DEP from Henderson Brook in T13 R12
WELS township (46.76114°N, 69.30659°W; WGS 84; 250 m elev.)
is located 30 km N of Churchill Dam. These records suggest the
species may be widely distributed in the Allagash River system.
SCOTT B. LINDEMANN (e-mail: lindemann.scott@gmail.com) and
AIDAN M. O’BRIEN, Department of Wildlife, Fisheries, and Conservation
Biology, University of Maine, 5755 Nutting Hall, Room 210, Orono, Maine
04469, USA (e-mail: aidan.obrien@maine.edu); DAVID E. PUTNAM,
University of Maine at Presque Isle, 214 South Hall, Presque Isle, Maine
04769, USA (e-mail: david.putnam@maine.edu).
CHRYSEMYS PICTA (Painted Turtle). USA: GEORGIA: Baldwin
Co.: wooded wetland ca. 25 m below pond outflow (unnamed
tributary to Reedy Creek) at Bartram Forest Wildlife Management
Area, ca. 7.0 air km SSE of Milledgeville (33.01836°N, 83.20566°W;
NAD 83). 28 April 2009. Christopher E. Skelton and Mieko A.
Camp. Verified by John B. Jensen and Kenneth Krysko. Florida
Museum of Natural History (UF 164461; photo voucher). Single
adult captured and photographed in small wetland. Habitat
consisted of thin Liquidambar styraciflua (Sweetgum) overstory
Herpetological Review 50(1), 2019
GEOGRAPHIC DISTRIBUTION 99
with Ligustrum sinense (Chinese Privet) and Arundinaria
gigantea (Giant Cane) understory. This specimen represents a
new county record and fills part of a gap in the distribution of this
species along the southern edge of its range in Georgia (Jensen
et al. 2008. Amphibians and Reptiles of Georgia. University of
Georgia Press, Athens. 575 pp.). There are records from three
of the five adjacent counties, the closest of which is ca. 16.7 km
NNW in Putnam County (Georgia Herp Atlas 01872).
CHRISTOPHER E. SKELTON, Skelton Environmental Consulting
Services Inc., 510 Saint Barbaras Ln NW, Marietta, Georgia 30064, USA
(e-mail: skeltonenv@gmail.com); MIEKO A. CAMP, Nontidal Wetlands
Program, Maryland Department of the Environment, 1800 Washington
Blvd., Ste. 430, Baltimore, Maryland 21230, USA.
eMYdOideA BLAndinGii (Blanding’s Turtle). USA:
WISCONSIN: bayField Co.: Moquah Barrens Research Natural
Area (46.61663°N, 91.21296°W). 18 June 2018. Ryan Raschke.
Verified by Joshua M. Kapfer. Milwaukee Public Museum (MPM
VZP907; photo voucher). An adult individual was photographed
early evening, 0.5 km from a small waterbody. New county record
that extends the known range of this species in Wisconsin (Vogt
1981. Natural History of Amphibians and Reptiles in Wisconsin.
Milwaukee Public Museum, Milwaukee, Wisconsin. 205 pp.;
Casper 1996. Geographic Distributions of the Amphibians and
Reptiles of Wisconsin. Milwaukee Public Museum, Milwaukee,
Wisconsin. 87 pp.). This record is roughly 26 km to the east of
the boundary with Douglas County, where Blanding’s Turtles
are already confirmed. The observation site falls within the
Chequamegon-Nicolet National Forest, on the east side of the
Moquah Barrens Management Area that contains a mosaic of
open pine (Pinus banksiana, P. resinosa) barren and savanna with
native grass and “heath” shrubs (e.g., Vaccinium spp., Comptonia
peregrine), hardwood saplings (Populus spp., Quercus spp.),
pine plantations, and hardwood forest, with closed-canopy
plantations and hardwood forests outside the management area
to the east. Mixed pine-hardwood forest is located north of the
observation site, and open barrens are located to the south.
RYAN RASCHKE, USDA Forest Service, Washburn Ranger District,
Washburn, Wisconsin 54891, USA (e-mail: raschke.ryan@gmail.com);
BRIAN R. STURTEVANT, USFS Northern Research Station Institute for
Applied Ecosystem Studies, Rhinelander, Wisconsin 54501, USA (e-mail:
bsturtevant@fs.fed.us); KATHLEEN M. QUIGLEY, Department of Plant,
Soil, and Microbial Sciences, Michigan State University, East Lansing,
Michigan 48824, USA (e-mail: kathlquigl@gmail.com).
EMYDOIDEA BLANDINGII (Blanding’s Turtle). USA:
WISCONSIN: ashland Co.: specific locality information withheld
due to the sensitive nature of the site. 20 July 2018. Gary Mertig.
Verified by Joshua M. Kapfer. Milwaukee Public Museum (MPM
VZP909; photo voucher). New county record filling a gap in
the known range of this species in Wisconsin (Casper 1996.
Geographic Distributions of the Amphibians and Reptiles of
Wisconsin. Milwaukee Public Museum, Milwaukee, Wisconsin.
87 pp.). One large adult was observed and photographed.
The individual was first observed in the observer’s yard, then
monitored as it moved to nearest road where it was assisted
across. The property where the observation was made is forested
with area of open hay fields, and numerous small streams and
creeks with their associated wetlands, both forested and open.
This observation complements a record in the Chequamegon–
Nicolet National Forest, ca. 39.5 km from adjacent Sawyer
County (Wisconsin Natural Heritage Inventory; Nov 2018).
100 GEOGRAPHIC DISTRIBUTION
GARY MERTIG (e-mail: gmertig@fs.fed.us) and CARRIE NELSON, U.S.
Forest Service, P.O. Box 896, 10650 Nyman Ave, Hayward, Wisconsin 54843,
USA; RYAN J. MAGANA, Wisconsin Department of Natural Resources-
Bureau of Natural Heritage Conservation, 810 W Maple St, Spooner,
Wisconsin 53801, USA; RICHARD A. STAFFEN, Wisconsin Department of
Natural Resources-Bureau of Natural Heritage Conservation, 101 S Webster
St, P.O. Box 7921, Madison, Wisconsin 53707, USA.
GRAPTEMYS GEOGRAPHICA (Common Map Turtle). USA:
MICHIGAN: benZie Co.: Platte River, 0.91 river km downstream
of MI-22 (44.71543°N, 86.12689°W; WGS 84). 23 May 2016.
Kenneth Bosma. Verified by James Harding. Auburn University
Museum of Natural History (AHAP-D 2516; photo voucher).
This represents the first vouchered record for Benzie County,
confirming unvouchered records from an unpublished National
Park Service report (Casper and Anton 2008. Natural Resource
Technical Report NPS/GLKN/NRTR-2008/147; Phillips 2016.
JNAH 2016:45–69). Graptemys geographica have reportedly
been observed in the Betsie River in Benzie County and Grand
Traverse County (J. Harding, pers. comm.), and both counties
lie within the species’ expected range in Michigan (Harding
and Mifsud 2017. Amphibians and Reptiles of the Great Lakes
Region. Revised Edition. University of Michigan Press, Ann
Arbor, Michigan. 408 pp.).
MATTHEW WELC, Department of Biological Sciences, 331 Funchess
Hall, Auburn University, Auburn, Alabama 36849, USA (e-mail: welcmatt@
auburn.edu); KENNETH BOSMA.
GRAPTEMYS PSEUDOGEOGRAPHICA (False Map Turtle).
USA: TEXAS: palo pinto Co.: Brazos River, Hwy 180 crossing W
of Mineral Springs (32.79857°N, 98.18609°W; WGS 84). 19 August
2018. Peter V. Lindeman. Verified by Michael Forstner. Florida
Museum of Natural History (UF 185765; photo voucher). A
basking male and a swimming female. Brazos River, Hwy 281
crossing S of Mineral Springs (32.64101°N, 98.10079°W; WGS 84).
19 August 2018. Peter V. Lindeman. Verified by Michael Forstner.
UF 185766 (photo voucher). A basking male.
stephens Co.: Clear Fork of the Brazos, Hwy 183 crossing NW
of Breckenridge (32.92744°N, 99.01026°W; WGS 84). 19 August
2018. Peter V. Lindeman. Verified by Michael Forstner. UF 185767
(photo voucher). A swimming juvenile female. Clear Fork of
the Brazos, Hwy 578 crossing N of Breckenridge (32.90164°N,
98.89949°W; WGS 84). 19 August 2018. Peter V. Lindeman. Verified
by Michael Forstner. UF 185768 (photo voucher). A basking male.
All turtles photographed using an 83× Nikon CoolPix camera.
These are new records for these two adjacent counties and fill a gap
in the range of Graptemys pseudogeographica in the upper Brazos
River drainage bounded by Parker County to the east, Shackelford
County to the west, and Young and Throckmorton counties to
the north (Lindeman 2013. The Map Turtle and Sawback Atlas:
Ecology, Evolution, Distribution, and Conservation. University
of Oklahoma Press, Norman, Oklahoma. 460 pp.). The Palo Pinto
specimen photographs voucher the county, which is shown
as having an unvouchered sight record (from lower Palo Pinto
Creek, dating from 1992) in Lindeman (2013, op. cit.). A Palo Pinto
County record was indicated for G. pseudogeographica by Dixon
(2013. Amphibians and Reptiles of Texas: with Keys, Taxonomic
Synopses, Bibliography, and Distribution Maps. Texas A&M
University Press, College Station, Texas. 447 pp.), dating to the
2000 second edition of his checklist. There is no indication in
Dixon’s notes regarding to which specimen the record refers,
but it may be a genetic sample taken from a specimen that was
Herpetological Review 50(1), 2019
catalogued with no locality information available other than the
county (M. Forstner, pers. comm.).
PETER V. LINDEMAN, Department of Biology and Health Sciences, 126
Cooper Hall, Edinboro University of Pennsylvania, Edinboro, Pennsylvania
16444, USA; e-mail: plindeman@edinboro.edu.
GRAPTEMYS PSEUDOGEOGRAPHICA KOHNII (Mississippi
Map Turtle). USA: MISSOURI: shannon Co.: Jacks Fork River
(tributary to the Current River), near Alley Spring (37.14875°N,
91.44361°W; WGS 84). 12 July 2017. Jeffrey E. Dawson. Verified
by Jeffrey T. Briggler and Richard E. Daniel. University of
Missouri–Columbia (UMC 3527P; photo voucher). One adult
male photographed basking on a partially submerged log. First
documentation of this species in the Jacks Fork River and new
county record (Daniel and Edmond 2018. Atlas of Missouri
Amphibians and Reptiles for 2017. http://atlas.moherp.org/
pubs/atlas17.pdf; 8 Nov 2018). The nearest known record is from
the Current River, ca. 150 river km downstream (ca. 80 straight-
line km to the southeast) in Ripley County, Missouri (Daniel et al.
2010. Missouri Herpetol. Assoc. Newsl. 23:6–8).
JEFFREY E. DAWSON, Charles H. Hoessle Herpetarium, Saint Louis
Zoo, 1 Government Drive, St. Louis, Missouri 63110, USA; e-mail: jdawson@
stlzoo.org.
MACROCHELYS TEMMINCKII (Alligator Snapping Turtle). USA:
MISSISSIPPI: stone Co.: ca. 30 m upstream of the New Hope Rd
bridge over Double Branch Creek (30.89162°N, 89.24423°W; WGS
84). 9 May 2018. Grover J. Brown and Sarah Kohoutek. Verified by
Luke S. Pearson. Florida Museum of Natural History (UF 185553;
photo voucher). One juvenile alligator snapping turtle (102.1
mm SCL) was caught in this small stream in a baited hoopnet.
Macrochelys temminckii has not been reported from Stone
County in past issues of Herpetological Review, the Mississippi
Museum of Natural Science collection records, Mississippi
Natural Heritage Program records, nor in museum records
available on VertNet.org. This record is 51 km south from the
nearest record (Mississippi Museum of Natural Science [MMNS]
179524) in Forrest County, Mississippi. This turtle was handled
under scientific collecting permit #0530181 (MMNS).
GROVER J. BROWN (e-mail: grover.brown@usm.edu), SARAH
KOHOUTEK (e-mail: sarah.kohoutek@usm.edu), and DE’ANDRE
DAVIS, Department of Biological Sciences, The University of Southern
Mississippi, 118 College Drive, Hattiesburg, Mississippi 39406, USA (e-mail:
deAndre.l.davis@usm.edu).
PSEUDEMYS CONCINNA SUWANNIENSIS (Suwannee Cooter).
USA: FLORIDA: sarasota Co.: Phillippi Creek (27.30633°N,
82.51164°W; WGS 84). 13 September 2018. George L. Heinrich
and Timothy J. Walsh. Verified by Coleman M. Sheehy III. Florida
Museum of Natural History (UF 185965, 185966; photo vouchers).
Two adult P. c. suwanniensis (male and female), along with an
adult P. floridana peninsularis (Peninsula Cooter) and an adult P.
nelsoni (Florida Red-bellied Cooter), were observed basking on
a concrete structure. Range extension, new county record, and
new river record (Heinrich et al. 2015. J. North Am. Herpetol.
1:53–59; Heinrich and Walsh 2016. Herpetol. Rev. 47:422). This
record extends the range 30.1 km SW from the nearest published
locality on the Manatee River (Manatee County; Heinrich
and Walsh 2016, op. cit.)
GEORGE L. HEINRICH, Heinrich Ecological Services, 1213 Alhambra
Way S., St. Petersburg, Florida 33705-4620, USA (e-mail: george@
heinrichecologicalservices.com); TIMOTHY J. WALSH, Bruce Museum, 1

Museum Drive, Greenwich, Connecticut 06830-7157, USA (e-mail: twalsh@
brucemuseum.org).
STERNOTHERUS ODORATUS (Eastern Musk Turtle). USA:
MAINE: penobsCot Co.: Leonard’s Mill on Blackman Stream,
Bradley (44.872401°N, 68.634296°W; WGS 84). 40 m elev. 24 July
2018. Aidan M. O’Brien, Scott B. Lindemann, and Elijah McGill.
Yale Peabody Museum of Natural History (YPM 21971). Verified by
Greg Watkins-Colwell. Empty shell found near historic mill dam
on Blackman Stream. Habitat was palustrine emergent wetland
with surrounding mixed coniferous-deciduous forest. This
occurrence represents the northeastern-most specimen and is
only the second report of the species east of the Penobscot River
(Hunter et al. 1999. Maine Amphibians and Reptiles. University
of Maine Press, Orono, Maine. 252 pp.; Maine Amphibian and
Reptile Atlas Project [MARAP] files, Maine Department of Inland
Fisheries and Wildlife, Bangor, Maine). The only previous record
east of the Penobscot River, from the town of Beddington,
Washington County, accurately described S. odoratus captured
in a Maine Sea-Run Salmon Commission weir trap on the
Narraguagus River, ca. 50 km east of this 2018 occurrence (Mairs
1962. Maine Field Naturalist 18:182). This 2018 occurrence is
located 11 km SE of the closest record, a 2011 photo record
from Old Town, Penobscot County, west of the Penobscot River
(MARAP files). Together, the 2018 and 1962 occurrences suggest
that additional populations of S. odoratus may occur between the
Penobscot and Narraguagus rivers. Work was performed under
a Maine Department of Inland Fisheries and Wildlife scientific
collection permit (#2018-184) issued to SBL.
SCOTT B. LINDEMANN (e-mail: lindemann.scott@gmail.com) and
AIDAN M. O’BRIEN, Department of Wildlife, Fisheries, and Conservation
Biology, University of Maine, 5755 Nutting Hall, Room 210, Orono, Maine
04469, USA (e-mail: aidan.obrien@maine.edu).
STERNOTHERUS ODORATUS (Common Musk Turtle). USA:
WISCONSIN: outagaMie Co.: Wolf River Bottoms Wildlife Area
on an Osprey Flowage dike (44.52056°N, 88.61556°W; WGS
84). 26 July 2018. Jordyn H. Ziesmer. Verified by Rori A. Paloski.
Milwaukee Public Museum Collections (MPM VZP 906, photo
voucher). New county record (Vogt 1981. Natural History of
Amphibians and Reptiles in Wisconsin. Milwaukee Public
Museum, Milwaukee, Wisconsin, 205 pp.). An adult musk
turtle was seen walking across a dike from one waterbody to
an adjacent waterbody. This record is ca. 51 km S of the nearest
published account (Milwaukee Public Museum [MPM] 836a-b).
JORDYN H. ZIESMER, N8648 Comorn Rd, Ripon, Wisconsin 54971,
USA; e-mail: jordynziesmer@gmail.com.
TRACHEMYS SCRIPTA ELEGANS (Red-eared Slider). USA:
OHIO: union Co.: Mill Creek at Swartzkopf Park, Marysville
(40.24308°N, 83.37309°W; WGS 84). 16 July 2008. Jeffrey E.
Dawson. Verified by Jeffrey G. Davis. Cincinnati Museum
Center, Geier Collections and Research Center (CMC HP 480;
photo voucher). One large adult photographed basking near a
Graptemys geographica on a fallen tree.
vinton Co.: Lake Hope in Lake Hope State Park (39.33718°N,
82.34789°W; WGS 84). 5 May 2009. Jeffrey E. Dawson and Cheri
L. Dawson. Verified by Jeffrey G. Davis. CMC HP 5263 (photo
voucher). A single large adult photographed basking on a branch
extending up from shallow water.
Both specimens represent new county records (Wynn and
Moody 2006. Ohio Turtle, Lizard, and Snake Atlas. Ohio Biol. Surv.
Herpetological Review 50(1), 2019
GEOGRAPHIC DISTRIBUTION 101
Misc. Contr. No. 10, Columbus. iv + 81 pp.). Trachemys scripta
elegans may be native to the Scioto River of central Ohio (Conant
1951. Reptiles of Ohio. Second edition. University of Notre Dame
Press, Notre Dame, Indiana. 284 pp.), but the species has now
been found throughout Ohio as a result of pet releases (Wynn
and Moody 2006, op. cit.), and introduced populations have
the potential to become established in the state (Dawson and
Hatcher 2009. Herpetol. Rev. 40:236).
JEFFREY E. DAWSON, Charles H. Hoessle Herpetarium, Saint Louis
Zoo, 1 Government Drive, St. Louis, Missouri 63110, USA; e-mail: jdawson@
stlzoo.org.
SQUAMATA — LIZARDS
AMEIVULA NIGRIGULA. BRAZIL: BAHIA: palMas de Monte alto:
Serra dos Montes Altos State Park (14.28556°S; 43.153639°W; WGS
84), 763 m elev. 18 May 2018. J. G. G Sousa, L. A. Cavalcante, and
C. Ferreira-Silva. Verified by R. W. Ávila. Coleção Herpetológica da
Universidade Regional do Cariri, Crato, Ceará (URCA-H 14142–
14144). Specimens found foraging on the edge of a white sand
dirt road. The known distribution of this species is restricted to
the states of Bahia (Bom Jesus da Lapa, Cafarnaum, Catinga do
Moura, Central, Gentio do Ouro, Itaguaçu da Bahia, Jacobina),
and Minas Gerais (Riacho dos Machados, Grão-Mogol). This
new municipality record fills the gap between Bom Jesus da
Lapa, Bahia (ca. 135 km north), and Riacho dos Machados,
Minas Gerais (ca. 176 km south; Pinto-Silva and Silva-Soares
2018. Herpetol. Notes 11:405–408). Specimens collected under
a permit issued by Instituto Chico Mendes de Conservação da
Biodiversidade-ICMBio 29613-1.
JOSÉ GUILHERME GONÇALVES SOUSA, Programa de Pós-
graduação em Ecologia e Recursos Naturais, Universidade Federal do
Ceará (UFC), Fortaleza, Ceará, Brazil (e-mail: sousajgg@gmail.com);
LEONIDES AZEVEDO CAVALCANTE, Graduação em Ciências Biológicas,
Universidade Regional do Cariri (URCA) Crato, Ceará, Brazil (e-mail:
azevedoleonides@gmail.com); CRISTIANA FERREIRA-SILVA, Programa
de Pós-graduação em Ciências Biológicas (Zoologia), Universidade
Estadual Paulista Júlio de Mesquita Filho, Botucatu, São Paulo, Brazil
(e-mail: cristianasilva006@gmail.com).
CALUMMA JULIAE. MADAGASCAR: MoraManga: Mangabe Forest
(19.0256°S, 48.16415°E; WGS 84). 18 October 2018. Raphali R.
Andriantsimanarilafy. Verified by David Prötzel. Zoologische
Staatssammlung München, Germany (ARR MV 001–003; photo
voucher). Two adult females were observed within 3 m of each other,
both on vegetation, 2–2.5 m above ground. The first individual was
observed at 1540 h on a small branch of Psiadia altissima near the
Sahamarirana River; the second at 1925 h roosting on a branch
of an unknown plant species. Calumma juliae is known from
fragmented and degraded primary rainforest (Prötzel et al. 2017.
Zool. J. Linn. Soc. 184:471–498). This new county record is ca. 13.5
km SW of the type locality, which is the only site known for this
species (Prötzel et al. 2017, op. cit.). Fieldwork was done under the
research permit number 254/17/MEEF/SG/DGF/DSAPSCB.Re
issued on 18 October 2017 by the Ministry of the Environment,
the Ecology and the Forest, Madagascar.
RAPHALI R. ANDRIANTSIMANARILAFY (e-mail: arraphali@voakajy.
mg), PIERRE RAZAFINDRAIBE (e-mail: ttpierot@yahoo.fr), EDDIE F.
RAKONTONDRASOA (e-mail: rakotoedds1@voakajy.mg) and EMILE
RAZANAKOTO, Madagasikara Voakajy, Lot II F 14 P Bis A – Andraisoro,
BP 5181, 101 Antananarivo, Madagascar (e-mail: emile.razanakoto@gmail.
com).
102 GEOGRAPHIC DISTRIBUTION
CORYTOPAHANES HERNANDESII (Hernandez’s Helmeted
Basilisk). MEXICO: QUINTANA ROO: MuniCipio solidaridad: 8.7
km NW Akumal (20.44703°N, 87.38042°W; WGS 84), 21 m elev.
16 September 2018. Diego F. Campos-Moreno, and César R.
Lucio-Palacio. Verified by Jonathan A. Campbell. Amphibian
and Reptile Diversity Research Center, University of Texas at
Arlington (UTADC 9222, 9223; photo vouchers). First record
for the municipality, and fifth for the state, extending the range
ca. 37 airline km east from the closest known locality at Cobá
(archaeological site and village), Municipality of Tulum (Lee
1996. The Amphibians and Reptiles of the Yucatan Peninsula.
Comstock Publishing Associates, Cornell University Press,
Ithaca, New York. 500 pp.; Cedeño-Vázquez et al. 2003. Herpetol.
Rev. 34:393–395). The adult lizard was observed basking at 1000
h on the ground among fallen leaves within the understory of
secondary tropical semideciduous forest. After a few minutes the
lizard ran to a nearby tree and climbed the trunk up to a height
of 90 cm. We thank the inhabitants of Uxuxubi hamlet, especially
Miguel Pani, Ramón, and Nereida for their hospitality and field
assistance.
J. ROGELIO CEDEÑO-VÁZQUEZ (e-mail: rcedenov@ecosur.mx),
DIEGO FERNANDO CAMPOS-MORENO (e-mail: dfcampos@ecosur.edu.
mx), and CÉSAR R. LUCIO-PALACIO, El Colegio de la Frontera Sur Campus
Chetumal, Av. Centenario Km 5.5, C.P. 77049 Chetumal, Quintana Roo,
México (e-mail: raz.lucio@gmail.com).
ELGARIA VELAZQUEZI (Central Baja California Alligator
Lizard). MEXICO: BAJA CALIFORNIA SUR: MuniCipality oF la paZ:
Rancho Los Queleles, ca. 10 m N of Arroyo La Soledad, Sierra
de La Giganta (24.81147°N, 110.8419°W; WGS 84), 377 m elev.
29 June 2018. Policarpio Amador Espinoza, Rosa Maria Bibo
Amador, Jesus Guadalupe Amador Bibo, and Shane J. MacFarlan.
Verified by L. Lee Grismer. La Sierra University Herpetological
Collection (LSUHC - LSUDPC 10759; photo voucher). First record
for the Sierra de La Giganta (Grismer and Hollingsworth 2001.
Herpetologica 57:488–496), extending the geographic range of E.
velazquezi in Baja California Sur ca. 30 km to the south of Mission
Los Dolores. The juvenile lizard was observed at about 1800 h
within a structure constructed for shading livestock. A second,
larger individual was observed two days later as it crawled
among rocks beneath the desert scrub canopy immediately
outside Rancho Los Queleles. Local ranchers remarked that this
was the first time they had ever noticed this species in the area.
Both lizards were discovered during an ecological survey in the
Sierra de La Giganta that was funded by a grant from the National
Geographic Society (HJ-099R-17).
SHANE J. MACFARLAN, Department of Anthropology, University of
Utah, Salt Lake City, Utah 84112, USA (e-mail: shane.macfarlan@anthro.
utah.edu); JUAN JOSE GARCIA, Department of Education, Culture,
and Society, University of Utah, Salt Lake City, Utah 84112, USA (e-mail:
folklorejay@gmail.com); ERIC SCHNITER Economic Science Institute,
Chapman University, Orange, California 92866, USA (e-mail: eschniter@
gmail.com); DIEGO GUEVARA BELTRAN, Department of Psychology,
Arizona State University, Tempe, Arizona 85287, USA (e-mail: dguevar3@
asu.edu); JESUS GUADALUPE AMADOR BIBO Departamento de Ciencia
Sociales, Universidad Autónoma de Baja California Sur, La Paz, BCS,
Mexico (e-mail: jea2_16@alu.uabcs.mx); GORGONIO RUIZ-CAMPOS
Departmento de Biología, Universidad Autónoma de Baja California,
Ensenada, BC, Mexico (e-mail: gruiz@uabc.edu.mx).
HEMIDACTYLUS AFF. PARVIMACULATUS (Sri Lankan House
Gecko). USA: TEXAS: ChaMbers Co.: Chambers County Safety
Herpetological Review 50(1), 2019
Rest Area Westbound, Interstate 10, ca. 3.9 rd km E jct TX Hwy
61 (29.84009°N, 94.60810°W). 13 September 2018. Drew R. Davis.
Verified by Aaron M. Bauer. Biodiversity Collections, University
of Texas at Austin (TNHC 112136 [DRD 5004]). A juvenile was
found on the exterior wall of a structure at 2147 h.
orange Co.: Texas Travel Information Center at Orange,
Interstate 10, ca. 1.3 rd km W of Sabine River (30.12354°N,
93.71206°W). 13 September 2018. Drew R. Davis. Verified by
Aaron M. Bauer. TNHC 112133 (DRD 5001). A juvenile was found
on a support column of a pavilion at 2011 h. Two additional
individuals, a juvenile (TNHC 112134 [DRD 5002]) and an adult
female (TNHC 112135 [DRD 5003]), were found nearby from
2020–2030 h.
Though morphologically similar to Hemidactylus
parvimaculatus, these geckos may represent a related form (A.
Bauer, pers. comm.). These four records of H. aff. parvimaculatus
represent the first documented occurrence of this species in
Texas (Dixon 2013. Amphibians and Reptiles of Texas: with Keys,
Taxonomic Synopses, Bibliography, and Distribution Maps.
Third Edition. Texas A&M University Press, College Station,
Texas. viii + 447 pp.; Hibbitts and Hibbitts 2015. Texas Lizards:
A Field Guide. University of Texas Press, Austin, Texas. xvi +
333 pp.). Within the United States, this introduced species of
gecko is known only from Louisiana (Boundy and Carr 2017.
Amphibians and Reptiles of Louisiana: An Identification and
Reference Guide. Louisiana State University Press, Baton Rouge,
Louisiana. xi + 386 pp.), where it is believed to have been first
introduced to New Orleans in 2010 (Heckard et al. 2013. IRCF
Reptile and Amphibians 20:192–196). Since the introduction
of H. aff. parvimaculatus to New Orleans, records of additional
populations have been reported, primarily from localities
surrounding Lake Pontchartrain (Borgardt 2015. Herpetol. Rev.
46:217; Borgardt 2016. Herpetol. Rev. 47:258; Glorioso 2016.
Herpetol. Rev. 47:81; Erdmann 2017. Herpetol. Rev. 48:125).
The nearest known locality where H. aff. parvimaculatus has
been reported from these Texas records is from ca. 319 airline
km to the east in Tangipahoa Parish, Louisiana (Southeastern
Louisiana University Vertebrate Museum [SLU] 6631–6633;
Ermann 2017, op. cit.). Both of these new localities in Texas occur
along Interstate 10, a major transportation corridor connecting
Houston, Texas to New Orleans, Louisiana, which may have
assisted in the spread of this species outside of the New Orleans
region. It seems likely that this species will continue to disperse
along the Interstate 10 corridor throughout both Texas and
Louisiana, and efforts should be made to monitor this spread.
All specimens were collected under a Texas Parks and Wildlife
Scientific Collecting Permit (SPR-1018-294) issued to DRD.
DREW R. DAVIS, School of Earth, Environmental, and Marine Sciences,
University of Texas Rio Grande Valley, 100 Marine Lab Drive, South Padre
Island, Texas 78597, USA (e-mail: drew.davis@utrgv.edu); TRAVIS J.
LADUC, Biodiversity Collections, Department of Integrative Biology, The
University of Texas at Austin, 10100 Burnet Road, PRC 176 – R4000, Austin,
Texas 78758, USA (e-mail: travieso@austin.utexas.edu).
HEMIDACTYLUS TURCICUS (Mediterranean Gecko). MEXICO:
BAJA CALIFORNIA: MuniCipality oF MexiCali: Pete’s Camp, Playa
Paraíso, San Felipe (31.1348°N, 114.888405°W; WGS 84), 4 m
elev. 6 August 2017. A. Hinsley. Verified by Clark R. Mahrdt. San
Diego Natural History Museum (SDSNH HerpPC 05391; photo
voucher). First record for the municipality and a range extension
of ca.146 airline km SW from the nearest published locality at
Golfo de Santa Clara, Sonora (Rorabaugh and Lemos-Espinal

2016. A Field Guide to the Amphibians and Reptiles of Sonora,
Mexico. ECO Herpetological Publishing and Distribution, Rodeo,
New Mexico. 688 pp.). It is also the third municipality record
within Baja California (Martínez-Isac and Valdez-Villavicencio
2000. Herpetol. Rev. 31:254; Valdez-Villavicencio et al. 2017.
Mesoam. Herpetol. 4:458–459). A population is now established
at this reported locality; apparently the species arrived about six
years ago (C. Burns, pers. observ.).
JORGE H. VALDEZ-VILLAVECENCIO, Conservación de Fauna del
Noroeste, Ensenada, Baja California, C.P. 22785, México (e-mail: j_h_
valdez@yahoo.com.mx); CAROLANNE BURNS, 315 W 2nd Street, Calexico,
California 92231, USA; AMY HINSLEY, Department of Zoology, University
of Oxford, Oxford, OX1 3PS, UK (e-mail: amy.hinsley@zoo.ox.ac.uk).
HEMIDACTYLUS TURCICUS (Mediterranean Gecko). USA:
TEXAS: palo pinto Co.: 13 km N of Brad (32.86433°N, 98.47548°W;
WGS 84). 21 September 2017. Jeffrey T. Jenkerson. Verified by
Travis J. LaDuc. Biodiversity Collections, University of Texas at
Austin (TNHC 109800). Individual captured at 2115 h along a
bunkhouse wall near a flood light at Camp Grady Spruce. This
observation represents a new county record, adding to the rapidly
expanding range of H. turcicus (Dixon 2013. Amphibians and
Reptiles of Texas: with Keys, Taxonomic Synopses, Bibliography,
and Distribution Maps. Texas A&M University Press, College
Station, Texas. 447 pp.). The nearest collected specimen is from
ca. 75.6 km to the southeast from near Stephenville, Erath County
(Biodiversity Research and Teaching Collections, Texas A&M
University [TCWC] 87834). Specimen collected under Scientific
Research Permit (SPR-0416-108) from Texas Parks and Wildlife
Department.
JEFFREY T. JENKERSON (e-mail: jjenkerson@bio-west.com),
CATHLEEN M. JENKERSON, BIO-WEST Inc., 1812 Central Commerce
Court, Round Rock, Texas 78664 USA.
OPHIOMORUS PUNCTATISSIMUS (Limbless Skink). GREECE:
PELOPONNESE: aChaia preFeCture: Platanovrysi, northwest from
the town of Halandritsa (38.11965°N, 21.76258°E; WGS 84), 337
m elev. 6 April 2013. Elias Tzoras, Philippos Katsiyiannis, and
Marios Vergetopoulos. Verified by P. Lymberakis. Natural History
Museum of Crete-University of Crete (NHMC 80.3.83.22; photo
voucher). This record represents the first observation of this
species in the wider region of Achaia and fills a distributional gap
between the prefectures of Corinthia in the northeast and Ilia in
West Peloponnese. In the Peloponnese Peninsula, Ophiomorus
punctatissimus is known from the prefectures of Ilia, Corinthia,
Argolida, Arcadia, Lakonia and Messinia (Chondropoulos 1986.
Amphibia-Reptilia 7:217–235; Sillero et al. 2014. Amphibia-
Reptilia 35:1–31) and from the neighboring islands, Kythera
and Elafonisos (Broggi 2016. Herpetozoa 29:37–46). The nearest
previous record is from Feneos Mountain, 52 km to the southeast
(Mayer et al. 1990. Herpetozoa 2:87–106). We found one adult
individual under a stone in a field of olive groves (Olea europaea).
Multiple additional individuals have since been found in the
same area.
ELIAS TZORAS, Patra, 26442 Achaia, Greece (e-mail: eliastzoras@
outlook.com); PHILIPPOS KATSIGIANNIS, Kato Platanovrysi, 25008 Achaia,
Greece (e-mail: greeknature@gmail.com); MARIOS VERGETOPOULOS,
Sepolia, 10443, Attica, Greece (e-mail: mariosvergetopoulos@hotmail.
com).
TRACHYLEPIS QUINQUETAENIATA (African Five-lined Skink).
USA: CALIFORNIA: los angeles Co.: Glendora (detailed locality
Herpetological Review 50(1), 2019
GEOGRAPHIC DISTRIBUTION 103
information available upon request to the Section of Herpetology,
Natural History Museum of Los Angeles County). 20 October 2018.
Gregory B. Pauly, Patrick D. Gavit, and Adam G. Clause. Verified
by Aaron M. Bauer. Natural History Museum of Los Angeles
County (LACM 190261–190265). Nine additional specimens
were collected 11 November 2018 (LACM 190266–190274). New
county and state record. Collected individuals ranged from recent
hatchlings to adult females and males, and at least 20 additional
individuals of various size classes were observed during walking
surveys of this residential neighborhood. As of fall 2018, the
inhabited area is approximately 150 m north to south and 160
m east to west, encompassing at least 15 house lots (1.64 ha).
The first free-ranging skink documented in this neighborhood
was a subadult observed 20 March 2016 by PDG. A photograph
of this skink was uploaded to the Reptiles and Amphibians of
Southern California (RASCals) project (iNaturalist 2810014) at
that time, but it was incorrectly identified as the native Western
Skink (Plestiodon skiltonianus). Subsequent observations in
August 2018 of an adult male (iNaturalist 15330968) and juvenile
(iNaturalist 16442860) made it clear this was a nonnative
species, first identified on iNaturalist as T. quinquetaeniata by W.
Flaxington. Surveys of this neighborhood then demonstrated an
established population. Collected specimens were compared to
the results of Broadley and Bauer (1998. Afr. J. Herpetol. 47:43–
58), and color pattern and scale counts confirmed these skinks
as T. quinquetaeniata.
Interviews with residents revealed a history of nonnative
reptiles occasionally showing up in this neighborhood as well
as a reptile dealer residing in the neighborhood. For example,
one of us (PDG) observed a Scheltopusik (Pseudopus apodus) in
this neighborhood on 20 September 2015 (iNaturalist 2352998),
and a local homeowner also reported an Asian Water Monitor
(Varanus salvator) found swimming in her backyard pool in
the mid to late 2000s (LACM-Photo Collection 2408). Multiple
individuals of both species had been previously imported by the
reptile dealer. The dealer also imported adult T. quinquetaeniata
from Egypt in September 2014. By March 2016, at least one T.
quinquetaeniata had crossed the street to the south of the
likely introduction site (iNaturalist 2810014), and there were
enough adult individuals throughout the neighborhood actively
breeding that there was a rapid population increase in the
summer and fall of 2018 as reported during interviews with
area residents. By this time, the population was up to the edge
of a second street south of the likely introduction site with no
documented individuals across that street. The population has
expanded largely in a southward direction, with adults and
recent hatchlings up to 225 m away from the dealer’s residence,
suggesting there was no or minimal lag time prior to the rapid
expansion of this population.
Native lizards including Western Fence Lizards (Sceloporus
occidentalis) and Southern Alligator Lizards (Elgaria
multicarinata) also occur in this neighborhood. Trachylepis
quinquetaeniata and S. occidentalis were observed using the
same basking habitats and retreat sites, and both are dietary
generalists that likely have broadly overlapping diets. One
neighborhood resident reported a decline in S. occidentalis
with the increasing occurrence of T. quinquetaeniata in his
backyard. Repeated surveys at other Southern California
localities where nonnative Anolis carolinensis, Anolis sagrei,
and Podarcis siculus occur have shown displacement of S.
occidentalis as nonnative populations expand (Pauly, unpubl.
data). We will continue monitoring this population to assess
104 GEOGRAPHIC DISTRIBUTION
changes in the native and nonnative lizard populations, and we
will work to eradicate the population while it is still early in the
introduction process.
Trachylepis quinquetaeniata is native to eastern, central,
and western Africa with populations extending north into Egypt
along the Nile Valley (Spawls et al. 2018. Field Guide to East
African Reptiles. Bloomsbury Publishing, London, UK. 624 pp.).
There is also an introduced population near the southern tip of
the Sinai Peninsula in Egypt that likely arrived there as stowaways
in nursery shipments (Necas and Vigasova 2004. Sauria 26[3]:13–
14.) and another introduced population in St. Lucie County,
Florida, USA, which also resulted from escapees from a reptile
dealer (Krysko et al. 2010. IRCF Reptiles Amphib. 17:183–184).
Thus, the Los Angeles County population reported here is the
first for California and the second for the United States.
The early detection of this nonnative population highlights
the value of citizen science projects in documenting and tracking
the spread of nonnative species. Just in Southern California
alone, reptile-focused citizen science projects led by the Natural
History Museum of Los Angeles County have documented
other established nonnative lizard populations in recent years
including Anolis carolinensis, Hemidactylus turcicus, and H.
platyurus (Larson et al. 2015. Herpetol. Rev. 46:59; Pauly and
Borthwick 2015. Herpetol. Rev. 46:567; Pauly et al. 2015. Herpetol.
Rev. 46:59; Pauly et al. 2015. Herpetol. Rev. 46:569).
Funding for the field surveys was provided by the Urban
Nature Research Center at the Natural History Museum of
Los Angeles County. We thank A. G. Clause for assistance with
field work, W. Flaxington for his efforts to identify this species
on iNaturalist, and the reptile dealer who kindly provided
information on importation history and is working with us to
prevent the further spread of this nonnative species.
GREGORY B. PAULY (e-mail: gpauly@nhm.org) and PATRICK D.
GAVIT, Natural History Museum of Los Angeles County, 900 Exposition
Boulevard, Los Angeles, California 90007, USA (e-mail: ebiner.gavit@gmail.
com).
XENOSAURUS AGRENON (Mountain Knob-scaled Lizard).
MEXICO: OAXACA: MuniCipality oF san pedro el alto: 1.8 km
NE Crucero Copalita, Hwy 175 Oaxaca-Puerto Angel (15.972°N,
95.5577°W; WGS 84), 1192 m elev. 24 November 2017. Víctor
H. Jiménez-Arcos, R. Alejandro Calzada-Arciniega, and Luis
E. Lozano-Aguilar. Verified by Guillermo A. Woolrich-Piña.
Amphibian and Reptile Diversity Research Center, University of
Texas at Arlington (UTADC 9179; photo voucher). First record for
the municipality, extending the known distribution 20 km west
of the nearest reported locality at Cafetal Alemania, Municipality
of Santa María Huatulco (Nieto-Montes de Oca et al. 2017. Mol.
Phylogen. Evol. 106:241–253). The lizard was found within a
rock crevice ca. 1.8 m above ground level in a pine-oak forest.
A second individual was observed about 5 m from the first one,
subsequently captured and photographed, and then released.
Our fieldwork was supported by the Mohamed Bin Zayed Species
Conservation Fund (grant #172516436) awarded to VHJA. We
thank Dr. Guillermo A. Woolrich-Piña for verifying the species
identity. The lizards were photographed under permit (FAUT-
0322) issued to VHJA by the Secretaría de Medio Ambiente y
Recursos Naturales.
VÍCTOR H. JIMÉNEZ-ARCOS, Naturam Sequi A.C. 16 de Septiembre
#43, Col. Cd. de los Niños, Naucalpan de Juárez, México, C.P. 53450, and
Laboratorio de Ecología, UBIPRO, FES Iztacala, Universidad Nacional
Autónoma de México, Av. De los Barrios 1, Tlalnepantla, México, C.P.
Herpetological Review 50(1), 2019
54090 (e-mail: vjimenezarcos@gmail.com); EUGUI R. MARTÍNEZ-PÉREZ,
Rancho Magnolia domicilio conocido, carretera Buenavista-Tierra Blanca,
C.P. 70920, San Agustín Loxicha, Oaxaca, México; LUIS E. LOZANO-
AGUILAR, Laboratorio de Herpetología, Departamento de Zoología,
Instituto de Biología, Universidad Nacional Autónoma de México, Apartado
postal 70515, 04510 México, Cd. de México, México; RAFAEL ALEJANDRO
CALZADA-ARCINIEGA, Departamento de Zoología, Instituto de Biología,
Universidad Nacional Autónoma de México, Apartado postal 70515, 04510
Cd. de México, México.
SQUAMATA — SNAKES
BOIGA QUINCUNCIATA (Assamese Cat Snake). INDIA: WEST
BENGAL: Alipurduar District, Buxa Tiger Reserve (26.7575°N,
89.58972°E; WGS 84), 744 m elevation. 5 June 2017. A. Ghosh and
N. Mukherjee. Verified by Gernot Vogel. Lee Kong Chian Natural
History Museum, National University of Singapore (ZRC[IMG]
2.401a–e; photo voucher). New record for West Bengal State,
extending range ca. 140 km E of nearest locality in Manas Tiger
Reserve, Assam State (Das et al. 2010. Russian J. Herpetol.
17:161–178). Adult individual of total length ca.126 cm found
perched on dry branches, 4 m above ground along cliff wall at
1930 h. Habitat was broadleaf forest in Himalayan foothills.
AVRAJJAL GHOSH, Department of Ecology and Environmental
Sciences, School of Life Sciences, Pondicherry University, Puducherry
605 104, India (e-mail: pitviper18@gmail.com); NILANJAN MUKHERJEE,
6/1/1/A Swinhoe Street, Ballygunge, Kolkata 700 019, West Bengal, India
(e-mail: mukherjee.nilanjan91@gmail.com).
CEMOPHORA COCCINEA COPEI (Northern Scarlet Snake).
USA: TEXAS: hunt Co.: Holly Creek Road, 0.45 km NE jct FM 1565
(32.87765°N, 96.25467°W; WGS 84). 29 May 2011. David Harkins.
Verified by Wade A. Ryberg. Biodiversity Research and Teaching
Collections, Texas A&M University (TCWC 100351). New county
record (Dixon 2013. Amphibians and Reptiles of Texas: with Keys,
Taxonomic Synopses, Bibliography, and Distribution Maps. Texas
A&M University Press, College Station, Texas. 447 pp.) extending
the known range in Texas west and northwest from Van Zandt
and Wood counties. The nearest records are 77 km east near the
ghost town of Coke, Wood County, Texas (American Museum
of Natural History [AMNH] R-138471) and 78 km southeast in
southeastern Van Zandt County, Texas (Amphibian and Reptile
Diversity Research Center, University of Texas at Arlington [UTA]
12793).
laMar Co.: Camp Maxey Military Reservation (33.79098°N,
95.60551°W; WGS 84). 17 May 2018. Drew E. Dittmer. Verified by
Wade A. Ryberg. TCWC 104040. New county record (Dixon 2013.
op. cit.) filling a distributional gap between Wood County, Texas
to the south and Atoka County, Oklahoma to the north. The
nearest known record to this location is 60 km north-northwest
at McGee Creek State Park in Atoka County, Oklahoma (Sam
Noble Oklahoma Museum of Natural History [OMNH] 41852).
The specimen was captured in a funnel trap placed along a
three-wing drift fence. There was a Gastrophryne olivacea in
the same funnel trap. The drift fence array was in the Post-Oak
Savannah eco region. The Camp Maxey Texas Army National
Guard training facility has many areas with sandy soils and pine
woodlands that closely match the described habitat associations
for the species. However, this snake was captured in a habitat
that is best described as a relatively low lying seasonally flooded
wetland and atypical for the habitat that the species is normally
associated.

Specimens collected under Texas Parks and Wildlife
Department Scientific Research Permit (SPR-0506-662) issued
to TJH.
DREW E. DITTMER, Department of Natural Resources Management,
Goddard Building, Texas Tech University, Box 42125, Lubbock Texas
79409, USA (e-mail: dittmer.drew.e@gmail.com); SARAH FRITTS, Texas
State University, 601 University Drive, San Marcos, Texas 78666, USA;
DAVID HARKINS, 10007 CR 2440, Royse City, Texas 75189, USA; TOBY J.
HIBBITTS, Biodiversity Research and Teaching Collections, Department
of Wildlife and Fisheries Sciences, Texas A&M University, College Station,
Texas 77843-2258, USA (e-mail: thibbitts@tamu.edu).
CLONOPHIS KIRTLANDII (Kirtland’s Snake). USA: OHIO:
Fulton Co.: specific locality withheld due to conservation
concerns. 4 September 2017. Matthew D. Cross and Joshua C.
Morrison. Verified by Jeff Davis. Cincinnati Museum of Natural
History (CMC HP 9739–9742; photo voucher). Four individuals
were found and photographed under artificial cover objects
placed by Toledo Zoo staff as part of statewide survey efforts for
C. kirtlandii. This location is a new county record for this species
in Ohio (Wynn and Moody 2006. Ohio Turtle, Snake, and Lizard
Atlas. The Ohio Biological Survey, Columbus, Ohio. 80 pp.).
Our observation is 20.8 airline km W of the nearest previously
documented locality in the northwestern region of Ohio (Wynn
and Moody 2006, op. cit.).
MATTHEW D. CROSS, Conservation Department, Toledo Zoo and
Aquarium, Toledo, Ohio 43614, USA (e-mail: matt.cross@toledozoo.org);
JOSHUA C. MORRISON, Whitehouse, Ohio 43571, USA.
CROTALUS ATROX (Western Diamond-backed Rattlesnake)
USA: TEXAS: ellis Co.: FM 66, ca. 1.4 mi W of County Road 4418
(32.2477°N, 97.0625°W; WGS 84), 184 m. 30 June 2018. Viviana
Ricardez and Brett Bartek. Verified by Carl J. Franklin. Amphibian
and Reptile Diversity and Research Center, University of Texas at
Arlington (UTA R-65034). Juvenile specimen found dead on road
at 2232 h. This record fills a gap within Hill, Johnson, Tarrant,
and Navarro counties (Dixon 2013. Amphibians and Reptiles
of Texas: with Keys, Taxonomic Synopses, Bibliography, and
Distribution Maps. Texas A&M University Press, College Station,
Texas. 447 pp.). The specimen reported herein was collected ca.
5.14 road mi NE of the next nearest county record along Hwy 66
in Hill County (Biodiversity Research and Teaching Collections,
Texas A&M University [TCWC] 88226). Specimen collected under
Scientific Research Permit SPR-1017-201 issued to Andrew
Brinker by Texas Parks and Wildlife Department.
VIVIANA RICARDEZ, Texasturtles.org, 326 Choctaw Trail, Waxahachie,
Texas, 75165 USA; e-mail: vricardez@gmail.com.
DIPSAS VARIEGATA VARIEGATA. BRAZIL: PERNAMBUCO:
são ViCente Férrer: Refúgio de Vida Silvestre Mata do Siriji
(7.61541°S, 35.50559°W; WGS 84), 591 m elev. 27 June 2018. E. G.
Dias. Verified by F. G. Rodrigues França. Coleção Herpetológica
and Paleoherpetológica of the Universidade Federal Rural of
Pernambuco, Dois Irmãos, Pernambuco, Brazil (CHP-UFRPE
5273–5275). One female (CHP-UFRPE 5273: 508 mm SVL, 518
mm TL, 22 g) and two males (CHP-UFRPE 5274: 608 mm SVL,
615 mm TL, 28 g; CHP-UFRPE 5275: 610 mm SVL, 619 mm TL,
31 g) were collected. Dipsas variegata variegata is known from
northwestern and western Venezuela, French Guiana, and the
Brazilian states of Pará and Bahia (Test et al. 1966. Mus. Zool.,
Univ. Michigan 128:1–63; Markezich 2002. Herpetol. Rev. 33:69–
74; Gasc and Rodrigues 1980. Bull. Mus. Nat. Hist. Nat. Paris
Herpetological Review 50(1), 2019
GEOGRAPHIC DISTRIBUTION 105
2:559–598; Chippaux 1986. Les serpents de la Guyane française,
ORSTOM Faune Trop., Paris, France, 165 pp.; Cunha and
Nascimento 1993. Bol. do Mus. Paraense Emílio Goeldi 9:44–46;
Hamdan and Lira-da-Silva 2012. Salamandra 48:31–50; Freitas
2014. Check List 10:1020–1030). First state record, extending the
geographic distribution ca. 740 km NNE from the municipality
of Elísio Medrado, Bahia, Brazil (Freitas 2014, op. cit.), the
closest known record. Collection license granted by the Instituto
Brasileiro do Meio Ambiente e dos Recursos Naturais Renováveis
– IBAMA, SISBIO nº 11218-1.
EMERSON GONÇALVES DIAS (e-mail: emersonherpeto@gmail.com),
IGOR YURI GONÇALVES SILVA DOS SANTOS, FLÁVIO JOSÉ SILVA,
ERICA SUZAN MARTINS LIMA, ERIC BERNARDINO GADELHA ROCHA,
RAYSSA LIMA DOS SANTOS, and EDNILZA MARANHÃO DOS SANTOS,
Laboratório Interdisciplinar de Anfíbios e Répteis, Departamento de
Biologia, Universidade Federal Rural de Pernambuco, Rua Dom Manoel de
Medeiros, s/n, Dois Irmãos - CEP: 52171-900 - Recife/PE.
HALDEA STRIATULA (Rough Earth Snake). USA: FLORIDA:
duval Co.: Pope Duval Park (30.30456°N, 81.90300°W; WGS
84). 6 May 2018. Daniel Durden and Bailey Peacock. Verified
by Coleman Sheehy. Florida Museum of Natural History (UF
184112). One adult specimen was found under a piece of plastic
debris before the first fishing pond at ca. 1558 h. New county
record (Krysko et al. 2011. Atlas of Amphibians and Reptiles in
Florida. Final report, Florida Fish and Wildlife Conservation
Commission, Tallahassee, Florida. 524 pp.). This record fills a
distributional area under Nassau County. The nearest records to
this specimen are two in Baker County (UF 119269, 119285) and
one in Nassau County (UF 117669).
BAILEY PEACOCK, 1365 Sunset View Ln, Jacksonville, Florida 32207,
USA; e-mail: baileypeacock@gmail.com.
IALTRIS DORSALIS (Hispaniolan W-headed Racer).
DOMINICAN REPUBLIC: LA ALTAGRACIA PROVINCE: 5.5 km
N of La Otra Banda, adjacent to Río Anamuya (18.69963°N,
68.66422°W; WGS 84), 117 m elev. 27 April 2016. Stesha A.
Pasachnik, Christopher M. Pellecchia, and Miguel A. Landestoy.
Verified by Robert Powell. Milwaukee Public Museum (MPM 835
a–d; photo voucher). First record for this species on the Cordillera
Oriental and a range extension of ca. 34 km N of the closest known
locality, 3 km NW of Boca de Yuma (UF/FSM 21569; F. Thompson
1975 field notes accessed by Leroy Nuñez), based on distribution
maps presented in Schwartz and Henderson (1991. Amphibians
and Reptiles of the West Indies: Descriptions, Distributions and
Natural History. University of Florida Press, Gainesville, Florida.
720 pp.). Zipple et al. (1994. Cat. Amer. Amphib. Rept. 592:1–3)
marked the location of UF/FSM 21569 on their maps, but the
dot appeared to be much further north than the actual location.
The adult male was photographed at 0900 h after a day and
night of continuous rainfall while crossing a dirt road headed to
Anamuyita Cacao Plantation.
STESHA ANN PASACHNIK, San Diego Zoo Institute for Conservation
Research, 15600 San Pasqual Valley Rd., Escondido, California 92025,
USA (e-mail: sapasachnik@gmail.com); CHRISTOPHER MICHAEL
PELLECCHIA, International Reptile Conservation Foundation, Tucson,
Arizona 85745, USA (e-mail: pellecchiacm@ircf.org); MIGUEL ANGEL
LANDESTOY, Escuela de Biología, Universidad Autónoma de Santo
Domingo, Dominican Republic (e-mail: hispanioland@gmail.com).
INDOTYPHLOPS BRAMINUS (Brahminy Blind Snake). USA:
CALIFORNIA: riverside Co.: Norco, Bluff Street (33.91270°N,
106 GEOGRAPHIC DISTRIBUTION
117.60380°W; WGS 84), 186 m elev. 11 April 2018. Bonnie
Johnson. Verified by Brad Hollingsworth. San Diego Natural
History Museum (SDNHM 76335). Another individual was
collected on 15 September 2018 (SDNHM 76336). This is the
first record for Riverside County. The suburban collection site
is within a Mediterranean (semi-arid) climatic region. The
upper layer of soil in this residential garden has been enriched
with commercial organic soil mix. The garden and adjacent
ornamental landscaping is maintained by supplemental
watering. Indotyphlops braminus feeds on small invertebrates,
mainly on ant and termite larvae and pupae. This non-native
population is sympatric at a fine scale (under stepping stones
and rocks) with the invasive and ecologically disruptive
Argentine Ant (Linepithema humile). The property owner had
lived at this residence since February 2005, with the first I.
braminus being observed in the fall of 2010. Since 2016, snakes
have been observed on seven different occasions. In California,
I. braminus has been reported from a site in San Diego County
located 149 km to the southeast (Palmer and Fisher 2010.
Herpetol. Rev. 41:518). Specimens were collected under a
California Department of Fish and Wildlife, Scientific Permit No.
5399, issued to ELE.
EDWARD L. ERVIN, Merkel & Associates, Inc., 5434 Ruffin Road,
San Diego, California 92123, USA (e-mail: eervin@merkelinc.com);
CLARK R. MAHRDT, San Diego Natural History Museum, Department
of Herpetology, P.O. Box 121390, San Diego, California 92112-1390, USA;
BONNIE JOHNSON, Orange County Water District, 14980 River Rd,
Corona, California 92880, USA.
INDOTYPHLOPS BRAMINUS (Brahminy Blind Snake). USA:
MASSACHUSETTS: Middlesex Co.: Watertown; 14 Mount
Auburn St (42.36562°N, 71.18622°W; WGS 84), near sea level. 18
October 2018. Sarah Jacobs. Verified by Tom French. Museum
of Comparative Zoology, Harvard University (MCZ R-195218).
Neonate (71 mm total length) with 338 total middorsal scales
and 192 vertebrae collected at 1715 h from a carpeted basement
of Thomson Safaris. This represents the second known record
in the state. The first record (Wallach et al. 1991. Herpetol. Rev.
22:68) originated from Boston’s North End, which is located 15
km due east from Watertown. The two localities are separated
by the Charles River, insuring that they represent separate
populations. This species is currently known from 13 states in
the USA (Alabama, Arizona, California, Florida, Georgia, Hawaii,
Louisiana, Massachusetts, Minnesota, North Carolina, Ohio,
Texas, and Virginia).
VAN WALLACH, 4 Potter Park, Cambridge, Massachusetts 02138,
USA (e-mail serpentes1@comcast.net); JOE MARTINEZ, Museum of
Comparative Zoology, Harvard University, 26 Oxford St., Cambridge,
Massachusetts 02162, USA (e-mail jmartinez@oeb.harvard.edu).
LAMPROPELTIS TRIANGULUM (Eastern Milksnake). USA:
TENNESSEE: CuMberland Co.: Catoosa Wildlife Management
Area (36.05993°N, 84.90082°W; WGS 84). 20 May 2017. Beau C.
Rapier and Lindsey P. Smith. Verified by A. Floyd Scott. David H.
Snyder Museum of Zoology, Austin Peay State University (APSU
19832; photo voucher). Adult found crossing gravel road through
closed canopy forest heading north. New county record filling a
gap in the known distribution of this species in the Cumberland
Plateau (Scott and Redmond 2017. Atlas of Reptiles in Tennessee.
http://www.apsubiology.org/tnreptileatlas/; accessed 4 Oct
2017). This locality is 20.9 km from the closest record (Tennessee
Tech University, uncatalogued) in Morgan County.
Herpetological Review 50(1), 2019
BEAU C. RAPIER (e-mail: brapier@my.apsu.edu), LINDSEY P. SMITH,
STEVEN J. HROMADA, and C. M. GIENGER, Center of Excellence for Field
Biology, Department of Biology, Austin Peay State University, Clarksville,
Tennessee 37040, USA.
LEPTODEIRA SPLENDIDA (Splendid Cat-eyed Snake).
MEXICO: CHIHUAHUA: MuniCipality oF urique: along road
paralleling the Rio Urique, 10 km downstream from Urique,
Chihuahua, Mexico (27.14813°N, 107.87907°W; WGS 84). 7
January 2017. Mark W. Herr, Sean P. Graham, Tomas Hernandez,
and Laine A. Giovanetto. Verified by Toby Hibbitts and Chris
Grünwald. James F. Scudday Vertebrate Collections at Sul Ross
State University, Alpine, Texas (SRSU-D 107; photo voucher).
Specimen found DOR. First validated record for the municipality
and second report from Chihuahua in over 50 years (Tanner
1985. Great Basin Nat. 45:615–676; Monte L. Bean Life Science
Museum, Brigham Young University [BYU] 22658), although
conflicting information has confused the actual locality of the
original Chihuahuan specimen. Lemos-Espinal and Smith
(2007. Anfibios y Reptiles del Estado de Chihuahua, México/
Amphibians and Reptiles of the State of Chihuahua, Mexico.
UNAM-CONABIO. México, D.F. 613 pp.) and Lemos-Espinal et
al. (2014. Amphibians and Reptiles of the Sierra Tarahumara
of Chihuahua, Mexico. ECO Herpetological Publishing and
Distribution, Rodeo, New Mexico. 405 pp.) depict on their range
maps a single record for this species in the Sierra Tarahumara of
southwestern Chihuahua along the borders of the municipalities
of Bocoyna and Guachochi. However, the contradicting written
localities provided in both publications say the record is
from the Municipality of Urique. The single specimen (BYU
22658) previously known for L. splendida from Chihuahua was
collected 12 December 1963 during the second Cross expedition,
which successfully descended the Rio Urique through Copper
Canyon after a disastrous first Cross expedition in October of
the same year. Locality information provided by Tanner (1985,
op. cit.) reads “approximately 15 mi. upstream Divisadero Trail,
Rio Urique.” The lower Divisadero Trail runs along Arroyo
Rurahuachi (Bjornsen 2009. Backpacker 37:33) and intercepts
the Rio Urique at their confluence (27.52602°N, 107.76755°W;
WGS 84). Therefore, we restrict the collecting locality for BYU
22658 to about 27.44504°N, 107.60027°W, which is ca. 6.2 km E
of the Municipality of Urique in the Municipality of Bocoyna. We
dedicate this note to the late John Cross, adventurer and leader
of the first expedition to conquer Copper Canyon by boat.
LAINE A. GIOVANETTO, Biology Department, New Jersey City
University, Jersey City, New Jersey 07305, USA (e-mail: lgiovanetto@njcu.
edu); MARK W. HERR, Biodiversity Institute and Natural History Museum,
University of Kansas, Lawrence, Kansas 66045, USA (e-mail: mwherr@gmail.
com); TOMAS HERNANDEZ, Department of Biology, Midland College,
Midland, Texas 79705, USA (e-mail: tohernandez@midland.edu); SEAN
P. GRAHAM, Department of Biology, Geology, and Physical Sciences, Sul
Ross State University, Alpine, Texas 79830, USA (e-mail: sean.graham@
sulross.edu).
MASTICOPHIS BILINEATUS (Sonoran Whipsnake). MEXICO:
NAYARIT: MuniCipio de teCuala: urban area of the city of Tecuala
(22.401331°N, 105.460383°W: WGS 84), 7 m elev. 31 May 2018.
Jesús Alberto Loc-Barragán. Verified by David Lazcano. Digital
Herpetology Collection, Universidad Autónoma de Nuevo
León, San Nicolas de los Garza (UANL 8241; photo voucher).
First municipality record and a range extension of ca. 12 airline
km SW of the nearest reported locality, ca. 0.40 km N of Rio

Acaponeta- Hwy 15, Municipality Acaponeta, Nayarit, based
on a specimen collected by A. E. Greer and F. W. Burley in
1963 (California Academy of Sciences [CAS] 23957). The adult
female was found during the morning, photographed, and then
immediately released. Thanks to David Lazcano for verifying
the species and accessioning its photo voucher into the UANL
collection.
JESÚS ALBERTO LOC-BARRAGÁN, Asociación para la Investigación
y Conservación de Anfibios y Reptiles A.C., Región Noroeste, Calle Bella
Italia, No. 170, Col. Lomas del Valle, C.P. 63066, Tepic, Nayarit, México;
e-mail: biolocbarragan@gmail.com.
MASTICOPHIS SCHOTTI (Schott’s Whipsnake). USA: TEXAS:
guadalupe Co.: Co Rd 232, ca. 1.45 km NE of Kingsbury (29.6576°N,
97.8179°W; WGS 84), 186 m elev. 30 October 2018. Haylea Rockett
and Austin Bohannon. Verified by Carl J. Franklin. Amphibian
and Reptile Diversity Research Center, University of Texas at
Arlington (UTADC 9234–9235; photo voucher). One adult male
specimen found DOR (775 mm SVL, 1155 mm TL, 122 g). New
county record (Dixon 2013. Amphibians and Reptiles of Texas:
with Keys, Taxonomic Synopses, Bibliography, and Distribution
Maps. Texas A&M University Press, College Station, Texas. 447
pp.). This record provides a northeastern distribution from Bexar
and Wilson counties where it is known to occur (Dixon 2013, op.
cit.). The nearest known record is ca. 70 km to the southwest in
Wilson County (Biodiversity Research and Teaching Collections,
Texas A&M University [TCWC] 89404). Specimen collected
under Scientific Permit SPR-0102-191 issued to MRJF by Texas
Parks and Wildlife Department.
AUSTIN M. A. BOHANNON (e-mail: amb200@txstate.edu), HAYLEA
D. ROCKETT, and MICHAEL R. J. FORSTNER, Department of Biology,
Texas State University, 601 University Drive, San Marcos, Texas 78666, USA.
OPHEODRYS AESTIVUS (Rough Green Snake). USA:
TENNESSEE: henry Co.: West Sandy Wildlife Managment Area
(36.30609°N, 88.16569°W). 20 October 2018. Lee J. Barton and
Cory S. Sawyers. Verified by A. Floyd Scott and C. M. Gienger.
Herpetological Review 50(1), 2019
GEOGRAPHIC DISTRIBUTION 107
David H. Synder Museum of Zoology, Austin Peay State
University (APSU 19898; photo voucher). First record for Henry
County (Scott and Redmond 2008. Atlas of Reptiles in Tennessee.
http://www.apsubiology.org/tnreptileatlas/; 2 Nov 2018). The
snake was found among vines that were removed from an oak
tree (Quercus sp.) and placed in a boat. The tree was located in
flooded hardwoods ca. 100 m from shore. The nearest known
records are ca. 75 km to the south in Decatur County (Biodiversity
Institute, University of Kansas [KU] 2469) and ca. 75 km to the
south-southeast in Carroll County (Louisiana State University
Museum of Natural Science [LSUMZ] 34958).
LEE J. BARTON, Freed-Hardeman University, 151 E. Main Street,
Henderson, Tennessee 38340, USA (e-mail: lbarton@fhu.edu); CORY S.
SAWYERS, Kuykendall Road, Puryear, Tennessee 38251, USA.
TYPHLOPS LEPTOLEPIS. CUBA: GUANTÁNAMO: baraCoa:
El Naranjo del Toa (20.38459°N, 74.56448°W; NAD 27 Cuba).
31 July 2008. Carlos A. Martínez Muñoz. Verified by Manuel
Iturriaga Monsisbay. Instituto de Ecología y Sistemática, Cuba
(CZACC 4.12963). First province record and a geographic range
extension of 34 km southeast of the nearest published locality
at Altiplanicie del Toldo, Holguín Province, Municipality of
Moa (20.45000°N, 74.88333°W; WGS 84; Domínguez et al. 2013.
Zootaxa 3681:136–146). The snake was found under a rock
during the day within submontane mesophyll evergreen forest.
The specimen was collected under the auspices of the project
“Colecciones zoológicas, su conservación y manejo II,” by the
Institute of Ecology and Systematics, Cuba.
JAVIER TORRES, Department of Ecology and Evolutionary Biology,
The University of Kansas, Lawrence, Kansas 66045, USA (e-mail:
javiertorres@ku.edu); CARLOS A. MARTÍNEZ-MUÑOZ, Zoologisches
Institut und Museum, Cytologie und Evolutionsbiologie, Ernst-Moritz-
Arndt-Universität Greifswald, Soldmannstrasse 23, D-17487 Greifswald,
Germany; ANAISA CAJIGAS GANDIA, Senior student, Biology major,
Faculty of Biology, University of Havana, Cuba (e-mail: acajigasgandia27@
gmail.com).