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Macroeconomics 8th Edition Abel Test

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Macroeconomics, 8e (Abel/Bernanke/Croushore)
Chapter 7 The Asset Market, Money, and Prices

7.1 What Is Money?

1) A disadvantage of the barter system is that


A) no trade occurs.
B) people must produce all their own food, clothing, and shelter.
C) the opportunity to specialize is greatly reduced.
D) gold is the only unit of account.
Answer: C
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

2) The use of money is more efficient than barter because the introduction of money
A) reduces the need for economic specialization.
B) reduces the need to exchange goods.
C) reduces the need for other stores of value.
D) reduces transaction costs.
Answer: D
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

3) In economics, money refers to


A) income.
B) wealth.
C) assets used and accepted as payment.
D) currency.
Answer: C
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

4) moneys primary role in the economy comes from the benefits of lowering transactions costs
and allowing specialization. This function of money is called
A) store of value.
B) medium of exchange.
C) standard of deferred payment.
D) unit of account.
Answer: B
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

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5) For something to satisfy the medium-of-exchange function of money, it must be
A) backed by gold.
B) readily exchangeable for other goods.
C) issued by a central bank.
D) an inherently valuable commodity.
Answer: B
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

6) The following are all functions of money EXCEPT


A) medium of exchange.
B) store of value.
C) unit of account.
D) source of anxiety.
Answer: D
Diff: 1
Topic: Section: 7.1
Question Status: New

7) In some countries, prices in stores are listed in terms of U.S. dollars, rather than in units of the
local currency. That's most likely because
A) the country's political system is unstable.
B) interest rates are higher using U.S. dollars than using the local currency.
C) there is no other store of value.
D) the country has experienced high rates of inflation.
Answer: D
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

8) The number of units of one good that trade for one unit of alternative goods can be determined
most easily when
A) there is one unit of account.
B) the goods all weigh about the same.
C) the goods are all new.
D) the goods are actively traded through barter.
Answer: A
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

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9) A good that is used as a medium of exchange as well as being a consumption good is called
A) a barter money.
B) a commodity money.
C) a legal tender.
D) a debased money.
Answer: B
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

10) Why do people keep currency in their pockets when bank deposits pay interest?
A) Because banks might steal your money.
B) Because currency is more liquid.
C) Because bank deposits lose value due to inflation.
D) Because bank deposits lose value due to changes in interest rates.
Answer: B
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

11) One of moneys primary roles in the economy comes from the use of money to transfer
purchasing power to the future. This role of money is called
A) store of value.
B) unit of account.
C) medium of exchange.
D) standard of deferred payment.
Answer: A
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

12) Which of the following measures is the best measure of money as a medium of exchange?
A) M1
B) M2
C) M3
D) None of the above
Answer: A
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

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13) Suppose your bank raises its minimum-balance requirement for free checking on checking
accounts by $500. You take $500 out of your passbook savings account and put it in your
checking account. What is the overall effect on M1 and M2?
A) M1 rises by $500, M2 falls by $500.
B) M1 is unchanged, M2 is unchanged.
C) M1 rises by $500, M2 is unchanged.
D) M1 is unchanged, M2 falls by $500.
Answer: C
Diff: 3
Topic: Section: 7.1
Question Status: Previous Edition

14) M1 does not include


A) MMMFs.
B) travelers' checks.
C) currency.
D) transaction accounts.
Answer: A
Diff: 1
Topic: Section: 7.1
Question Status: Revised

15) Which of the following is not part of M1?


A) Transaction accounts
B) Checking accounts
C) Time deposits
D) Traveler's checks
Answer: C
Diff: 1
Topic: Section: 7.1
Question Status: Revised

16) Which of the following statements about M1 and M2 is true?


A) Demand deposits are not part of M1.
B) M2 is more liquid than M1.
C) M1 is larger than M2.
D) Savings deposits are part of M2.
Answer: D
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

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17) Which of the following statements about M1 and M2 is not true?
A) Transaction accounts are part of M1.
B) M2 is more liquid than M1.
C) M2 is larger than M1.
D) Transaction accounts are part of M2.
Answer: B
Diff: 2
Topic: Section: 7.1
Question Status: New

18) M2 includes
A) large-denomination time deposits.
B) institutional MMMFs.
C) commercial paper.
D) M1.
Answer: D
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

19) M2 does not include


A) Treasury bonds.
B) passbook savings accounts.
C) small-denomination time deposits.
D) M1.
Answer: A
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

20) Over half of U.S. currency is


A) held abroad.
B) used in the underground economy.
C) held by banks as reserves.
D) held by businesses, especially retailers, for making transactions.
Answer: A
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

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21) People in other countries want to hold U.S. dollars as a
A) medium of exchange.
B) store of value.
C) unit of account.
D) standard of deferred payment.
Answer: B
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

22) We shouldn't be concerned about U.S. currency held abroad because


A) the currency will never return to the United States.
B) foreigners use it to buy U.S. bonds.
C) it represents an interest-free loan to the United States.
D) foreigners can't spend it in their own countries.
Answer: C
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

23) What's the most common way for a central bank to reduce the money supply?
A) Collect higher taxes
B) Sell bonds to the public
C) Buy bonds from the government
D) Buy bonds from the public
Answer: B
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

24) Suppose you read in the paper that the Federal Reserve plans to expand the money supply.
The Fed is most likely to do this by
A) printing more currency and distributing it.
B) purchasing government bonds from the public.
C) selling government bonds to the public.
D) buying newly issued government bonds directly from the government itself.
Answer: B
Diff: 1
Topic: Section: 7.1
Question Status: New

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25) A developing country does not have enough taxes to cover its expenditures and is unable to
borrow. This government would be most likely to cover its deficit by
A) purchasing government bonds from the public.
B) selling government bonds to the public.
C) selling newly issued government bonds directly to the central bank.
D) buying newly issued government bonds directly from the central bank.
Answer: C
Diff: 2
Topic: Section: 7.1
Question Status: Previous Edition

26) What are the major components of M1? What are the major components of M2? Describe
each component.
Answer: The principal components of M1 are currency, transaction accounts, and traveler's
checks. Currency includes coins and Federal Reserve notes (i.e., paper money). Transaction
accounts are those against which checks may be drawn. Traveler's checks are a substitute for
currency that can be replaced if lost or stolen. The principal components of M2 are M1, savings
account deposits including money market deposit accounts (MMDAs), small time deposits, and
money market mutual funds (MMMFs). Small time deposits are certificates of deposit (CDs) of
less than $100,000 denomination. MMMFs invest their shareholders' funds in short-term
securities, pay market-based interest rates, and allow holders to write a limited number of
checks. MMDAs are like MMMFs, except they are offered by banks or thrift institutions such as
savings and loan associations.
Diff: 2
Topic: Section: 7.1
Question Status: New

27) What function is money playing in each of these situations:


a. You walk into a store in Germany and see that all the prices are in euros.
b. You buy a candy bar for $1.25.
c. Your Aunt Jane keeps $100 bills tucked into many books in her house.
Answer: a. unit of account; b. medium of exchange; c. store of value.
Diff: 2
Topic: Section: 7.1
Question Status: Previous Edition

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28) What happens to M1 and M2 due to each of the following changes?
(a) You take $500 out of your checking account and put it into a passbook savings account.
(b) You take $1000 out of your checking account and buy traveler's checks.
(c) You take $1500 out of your money-market mutual fund and deposit into your checking
account.
(d) You cash in $2000 in savings bonds and invest the money in a certificate of deposit.
Answer:
(a) M1 falls $500, M2 is unchanged (remember that M1 is part of M2).
(b) M1 and M2 are both unchanged.
(c) M1 rises $1500, M2 is unchanged.
(d) M1 is unchanged, M2 rises $2000.
Diff: 2
Topic: Section: 7.1
Question Status: Previous Edition

29) Why is per-capita U.S. currency demand so large? Who is holding large amounts of U.S.
currency and why are they doing so? Should U.S. policymakers be concerned about this? Why?
Answer: Currency demand is large mostly because foreigners hold many dollars. They do so
because of inflation or political instability in their countries. Policymakers shouldn't be very
concerned, since foreigners' dollar holdings represent an interest-free loan to the United States.
However, a cause for concern may be that fluctuations in our money supply may reflect
conditions abroad that are unrelated to the U.S. economy.
Diff: 1
Topic: Section: 7.1
Question Status: Previous Edition

7.2 Portfolio Allocation and the Demand for Assets

1) People's best guesses about returns on assets are called


A) expected returns.
B) liquidity.
C) risk.
D) the term structure of returns.
Answer: A
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

2) The set of assets that a holder of wealth chooses to own is called


A) an asset assortment.
B) a wealth strategy.
C) a portfolio.
D) an investment envelope.
Answer: C
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

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3) The uncertainty about the return an asset will earn is
A) liquidity.
B) risk.
C) time to maturity.
D) stochastic dominance.
Answer: B
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

4) The risk premium is


A) the amount by which the expected return on a risky asset exceeds the return on an otherwise
comparable safe asset.
B) a measure of the riskiness of the overall economy in a domestic country compared with a
foreign country.
C) the amount an investor must pay to insure his or her stock portfolio to protect against a fall in
value.
D) the amount an investment bank charges to guarantee an annuity that pays a fixed rate of
return in the future.
Answer: A
Diff: 1
Topic: Section: 7.2
Question Status: New

5) The amount by which the expected return on a risky asset exceeds the return on an otherwise
comparable safe asset is known as the
A) CDS spread.
B) risk premium.
C) VIX.
D) term spread.
Answer: B
Diff: 1
Topic: Section: 7.2
Question Status: New

6) The ease and quickness with which an asset can be exchanged for goods, services, or other
assets is its
A) risk.
B) time to maturity.
C) velocity.
D) liquidity.
Answer: D
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

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7) Time to maturity refers to the amount of time until
A) an asset repays the principal to an investor.
B) an asset pays interest for the first time.
C) a bond can be sold on the secondary market.
D) the yield curve shows an upward slope.
Answer: A
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

8) Compared with money, bonds have


A) less risk and less liquidity.
B) less risk and more liquidity.
C) more risk and less liquidity.
D) more risk and more liquidity.
Answer: C
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

9) AAA Company stock has a higher expected rate of return than ZZZ Company stock. All else
being equal, you would expect that relative to ZZZ, AAA company stock provides
A) less risk and less liquidity.
B) less risk and more liquidity.
C) more risk and less liquidity.
D) more risk and more liquidity.
Answer: C
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

10) The least liquid asset on this list is


A) money.
B) bonds.
C) houses.
D) stocks.
Answer: C
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

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11) In the early 2000s, lenders began issuing mortgage loans to people who would normally not
be qualified to take out loans because they did not meet lending standards. Those borrowers are
known as
A) alternative borrowers.
B) weak borrowers.
C) subprime borrowers.
D) credit risks.
Answer: C
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

12) The financial crisis occurred in 2008 in large part because of losses on securities consisting
of bundles of mortgage loans known as
A) home loan loss reserves.
B) credit default swaps.
C) mortgage-backed securities.
D) naked put options.
Answer: C
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

13) A one-year bond has an interest rate of 5% today. Investors expect that in one year, a one
year bond will have an interest rate equal to 7%. According to the expectations theory of the term
structure of interest rates, in equilibrium, a two-year bond today will have an interest rate equal
to
A) 3.0%.
B) 5.0%.
C) 5.5%.
D) 6.0%.
Answer: D
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

14) The idea that investors today compare the returns on bonds with differing times to maturity
to see which is expected to give them the highest return is the underlying principle behind the
________ of the term structure of interest rates.
A) expectations theory
B) investors' viewpoint analysis
C) segmented-markets theory
D) yield comparison theory
Answer: A
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

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15) The interest rate on long-term bonds is somewhat higher than suggested by the expectations
theory because
A) the expectations theory doesn't account for taxes.
B) a risk premium exists.
C) an inflation premium must be added to long-term bonds.
D) the Fed can only control short-term interest rates.
Answer: B
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

16) By spreading her investments out over many different assets, an investor achieves
A) a higher expected return.
B) increased risk.
C) diversification.
D) greater liquidity.
Answer: C
Diff: 1
Topic: Section: 7.2
Question Status: Previous Edition

17) Suppose that you could buy a one-year bond today, which has an interest rate of 3%. If you
wait a year and buy a one-year bond then, the interest rate will be 4%. Two years from now, a
one-year bond is expected to offer an interest rate of 5%. According to the expectations theory of
the term structure of interest rates, what is the interest rate on a two-year bond today? What is the
interest rate on a three-year bond today?
Answer: Two-year bond: (3% + 4%)/2 = 3.5%; Three-year bond: (3% + 4% + 5%)/3 = 4%.
Diff: 2
Topic: Section: 7.2
Question Status: Previous Edition

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18) Suppose that:
1) The interest on a one-year bond today is 3%;
2) The interest on a one-year bond starting one year from now is expected to be 4% per year;
3) The interest on a one-year bond starting two years from now is expected to be 5% per year;
4) The risk premium on a two-year bond is 0.5%; and
5) The risk premium on a three-year bond is 1.0%.

Use that information to answer the following questions.


a) According to the expectations theory, what is the interest rate today on a two-year bond?
Show your work.
b) According to the expectations theory, what is the interest rate today on a three-year bond?
Show your work.
c) Plot the yield curve.
Answer:
a) (3% + 4%)/2 + 0.5% = 4.0%
b) (3% + 4% + 5%)/3 + 1.0% = 5.0%
c) plot 3 points with 1, 2, and 3 years to maturity vs. yields of 3%, 4%, and 5%.
Diff: 2
Topic: Section: 7.2
Question Status: Previous Edition

7.3 The Demand for Money

1) A 10% decrease in real income usually leads to ________ in money demand.


A) an increase
B) no change
C) a decrease of less than 10%
D) a decrease of 10%
Answer: C
Diff: 1
Topic: Section: 7.3
Question Status: Previous Edition

2) A 5% increase in real income usually leads to ________ in money demand.


A) a decrease
B) no change
C) an increase of less than 5%
D) a decrease of 5%
Answer: C
Diff: 1
Topic: Section: 7.3
Question Status: New

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3) Which of the following is most likely to lead to a decrease of 10% in the nominal demand for
money?
A) An increase in real income of 5%
B) A decrease in real income of 5%
C) A decline of 10% in the price level
D) An increase of 10% in the price level
Answer: C
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

4) Which of the following is most likely to lead to an increase of 1% in the nominal demand for
money?
A) An increase in real income of 0.5%
B) A decrease in real income of 0.5%
C) A decline of 1% in the price level
D) An increase of 1% in the price level
Answer: D
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

5) The opportunity cost of holding currency decreases when


A) income decreases.
B) the interest rate on bonds decreases.
C) the interest rate on money decreases.
D) wealth decreases.
Answer: B
Diff: 1
Topic: Section: 7.3
Question Status: Previous Edition

6) An increase in the real interest rate would cause an increase in the real demand for money
A) no matter what the change in expected inflation.
B) if expected inflation fell by less than the rise in the real interest rate.
C) if expected inflation fell by the same amount as the rise in the real interest rate.
D) if expected inflation fell by more than the rise in the real interest rate.
Answer: D
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

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7) An increase in expected inflation is likely to cause
A) a decline in the demand for real balances.
B) an increase in the demand for real balances.
C) no change in the demand for real balances.
D) no change in the demand for real balances only if the income elasticity of real money demand
is zero.
Answer: A
Diff: 1
Topic: Section: 7.3
Question Status: Previous Edition

8) Mr. Pierpont has wealth of $200,000. He wants to keep at least $80,000 in bonds at all times,
and will shift $10,000 into bonds from his checking account for each percentage point that the
interest rate on bonds exceeds the interest rate on his checking account. If the interest rate on
checking accounts is 4% and the interest rate on bonds is 9%, how much does Mr. Pierpont keep
in his checking account?
A) $50,000
B) $70,000
C) $130,000
D) $150,000
Answer: B
Diff: 1
Topic: Section: 7.3
Question Status: Previous Edition

9) Mr. Pierpont has wealth of $200,000. He wants to keep at least $80,000 in bonds at all times,
and will shift $10,000 into bonds from his checking account for each percentage point that the
interest rate on bonds exceeds the interest rate on his checking account. Currently, he keeps
$100,000 in bonds, which pay him 7%. What is the current interest rate on checking accounts?
A) 5%
B) 7%
C) 9%
D) 10%
Answer: A
Diff: 1
Topic: Section: 7.3
Question Status: Previous Edition

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10) Money demand is given by
Md/P = 1000 + .2Y - 1000i.
Given that P = 200, Y = 2000, and i = .10, real money demand is equal to
A) 1,300.
B) 1,500.
C) 260,000.
D) 300,000.
Answer: A
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

11) Over time, the wealth of society increases and payments technologies get more efficient.
What is the effect on money demand of these two changes?
A) Money demand rises proportionately to the rise in wealth.
B) Money demand rises, but less than proportionately to the rise in wealth.
C) The overall effect is ambiguous.
D) Money demand declines.
Answer: C
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

12) If there is a financial panic and increased uncertainty about the returns in the stock market
and bond market, what is the likely effect on money demand?
A) Money demand declines first, then rises when inflation increases.
B) Money demand rises.
C) The overall effect is ambiguous.
D) Money demand declines.
Answer: B
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

13) Suppose a new law imposes a tax on all trades of bonds and stock. What is the likely effect
on money demand?
A) Money demand declines first, then rises when inflation increases.
B) Money demand rises.
C) The overall effect is ambiguous.
D) Money demand declines.
Answer: B
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

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14) If real income rises 4%, prices rise 1%, and nominal money demand rises 4%, what is the
income elasticity of real money demand?
A) 3/4
B) 4/5
C) 5/6
D) 1
Answer: A
Diff: 3
Topic: Section: 7.3
Question Status: Previous Edition

15) If real income rises 5%, prices rise 3%, and nominal money demand rises 7%, what is the
income elasticity of real money demand?
A) 3/4
B) 4/5
C) 5/6
D) 6/7
Answer: B
Diff: 3
Topic: Section: 7.3
Question Status: New

16) If the interest elasticity of money demand is -0.1, by what percent does money demand
change if the nominal interest rate rises from 2% to 3%?
A) -0.1%
B) 5%
C) 0%
D) -5%
Answer: D
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

17) If the income elasticity of money demand is 3/4 and the interest elasticity of money demand
is -1/4, by what percent does money demand rise if income rises 10% and the nominal interest
rate rises from 4% to 5%?
A) 7.50%
B) 6.25%
C) 5.00%
D) 1.25%
Answer: D
Diff: 3
Topic: Section: 7.3
Question Status: Previous Edition

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18) Velocity is defined as
A) nominal money stock/nominal GDP.
B) nominal GDP/nominal money stock.
C) real money stock/real GDP.
D) mc2.
Answer: B
Diff: 1
Topic: Section: 7.3
Question Status: Previous Edition

19) If real GDP is $4 billion, the price level is 1.25, and the nominal money stock is $500
million, then velocity is
A) 0.1.
B) 1.
C) 10.
D) 100.
Answer: C
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

20) Money demand is given by


Md/P = 1000 + .2Y - 1000i.
Given that P = 200, Y = 2000, and i = .10, velocity is equal to
A) 0.65.
B) 0.75.
C) 1.33.
D) 1.54.
Answer: D
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

21) Suppose velocity is 3, real output is 9000, and the price level is 1.5. What is the level of real
money demand in this economy?
A) 2000
B) 3000
C) 6000
D) 30,000
Answer: B
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

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22) Suppose real money demand is 1000, real output is 6000, and the price level is 200. What is
the level of velocity in this economy?
A) 2
B) 3
C) 6
D) 12
Answer: C
Diff: 2
Topic: Section: 7.3
Question Status: New

23) Suppose velocity is constant at 4, real output is 10, and the price level is 2. From this initial
situation, the government increases the nominal money supply to 6. If velocity and output remain
unchanged, by how much will the price level increase?
A) 2.4%
B) 20%
C) 24%
D) 50%
Answer: B
Diff: 3
Topic: Section: 7.3
Question Status: Previous Edition

24) What happens to real money demand (rise, fall, no change) due to a change in each of the
following factors?
(a) A tax on stock market transactions is introduced.
(b) Computerized bond trading reduces transactions costs.
(c) People's average level of wealth rises.
(d) The threat of a recession increases the riskiness of stocks and bonds.
(e) The interest rate paid on checking account balances declines.
(f) The price level falls in a one-time jump.
Answer:
(a) Rises
(b) Falls
(c) Rises
(d) Rises
(e) Falls
(f) Is unchanged
Diff: 1
Topic: Section: 7.3
Question Status: Previous Edition

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25) Give five examples of factors that could reduce the demand for money.
Answer: Lower price level, lower real income, higher real interest rate, higher expected
inflation, lower nominal interest rate on money, lower wealth, lower risk on alternative assets,
higher risk on money, increased liquidity of alternative assets, or increased efficiency of
payments technologies.
Diff: 2
Topic: Section: 7.3
Question Status: Previous Edition

26) Suppose the money demand function is


Md/P = 1000 + 0.2Y - 1000 (r + πe).
(a) Calculate velocity if Y = 2000, r = .06, and πe = .04.
(b) If the money supply (Ms) is 2600, what is the price level?
(c) Now suppose the real interest rate rises to 0.11, but Y and Ms are unchanged. What happens
to velocity and the price level? So if the nominal interest rate were to rise from 0.10 to 0.15 over
the course of a year, with Y remaining at 2000, what would the inflation rate be?
Answer:
(a) V = PY/M = Y/(M/P). From the money demand function, M/P = 1300. So V = 2000/1300 =
1.54.
(b) P = Ms/( Md/P) = 2600/1300 = 2.
(c) Now Md/P = 1250. So V = 2000/1250 = 1.6. P = Ms/(Md/P) = 2600/1250 = 2.08. The
inflation rate would be 4%.
Diff: 3
Topic: Section: 7.3
Question Status: Previous Edition

7.4 Asset Market Equilibrium

1) Under a situation of asset market equilibrium,


A) the quantity of money supplied equals the quantity of money demanded.
B) the quantity of money supplied equals the quantity of nonmonetary assets demanded.
C) the quantity of nonmonetary assets supplied equals the quantity of monetary assets demanded.
D) the quantity of money supplied equals the quantity of nonmonetary assets supplied.
Answer: A
Diff: 1
Topic: Section: 7.4
Question Status: Previous Edition

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2) When the real quantity of money supplied equals the real quantity of money demanded, there
is said to be
A) goods market equilibrium.
B) asset market equilibrium.
C) monetary neutrality.
D) money illusion.
Answer: B
Diff: 1
Topic: Section: 7.4
Question Status: Previous Edition

3) Which of these variables is not a variable in the equation for the asset market equilibrium
condition?
A) Nominal money supply
B) Price level
C) Real income
D) Investment
Answer: D
Diff: 1
Topic: Section: 7.4
Question Status: New

4) Which of these variables is not a variable in the equation for the asset market equilibrium
condition?
A) Saving
B) Expected rate of inflation
C) Real interest rate
D) Real income
Answer: A
Diff: 1
Topic: Section: 7.4
Question Status: New

5) If the quantity of money demanded exceeds the quantity of money supplied, then
A) the quantity of nonmonetary assets demanded exceeds the quantity supplied.
B) the quantity of nonmonetary assets supplied exceeds the quantity demanded.
C) the quantity of nonmonetary assets demanded will still equal the quantity supplied, all else
being equal.
D) you can make no conclusions about the relative supply and demand of nonmonetary assets.
Answer: B
Diff: 2
Topic: Section: 7.4
Question Status: Previous Edition

21
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6) Suppose the real money demand function is
Md//P = 2400 + 0.2 Y - 10,000 (r + πe).
Assume M = 4000, P = 2.0, πe = .03, and Y = 5000. The real interest rate that clears the asset
market is
A) 3%.
B) 6%.
C) 11%.
D) 14%.
Answer: C
Diff: 3
Topic: Section: 7.4
Question Status: Previous Edition

7) Suppose the real money demand function is


Md/P = 2400 + 0.2 Y - 10,000 (r + πe).
Assume M = 5000, πe = .03, and Y = 5000. If the price level were to decrease from 2.5 to 2.0,
then the real interest rate would decrease by how many percentage points (assuming Md/, πe, and
Y are unchanged)?
A) 4
B) 5
C) 9
D) 14
Answer: B
Diff: 3
Topic: Section: 7.4
Question Status: Previous Edition

8) Suppose the real money demand function is


Md/P = 2400 + 0.2 Y - 10,000 (r + πe).
Assume M = 5000, P = 2.0, and πe = .03. If Y were to increase from 4000 to 5000, then the real
interest rate would increase by how many percentage points?
A) 2
B) 4
C) 5
D) 7
Answer: A
Diff: 3
Topic: Section: 7.4
Question Status: Previous Edition

22
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9) Suppose real money demand is
L = 0.8 Y - 100,000 (r + πe).
If the nominal money supply is 12,000, real output is 15,000, the real interest rate is .02, and the
expected inflation rate is .01, then the price level is
A) 3/4.
B) 1.
C) 4/3.
D) 3.
Answer: C
Diff: 3
Topic: Section: 7.4
Question Status: Previous Edition

10) Suppose the real interest rate is 4% and the expected inflation rate is 3%. If the money
supply increases by 10% and output, the real interest rate, and the expected inflation rate are
unchanged, then the price level increases by
A) 3%.
B) 4%.
C) 7%.
D) 10%.
Answer: D
Diff: 3
Topic: Section: 7.4
Question Status: Previous Edition

11) Define asset market equilibrium and state the asset market equilibrium condition.
Answer: Asset market equilibrium exists when the quantity of assets supplied equals the
quantity of assets demanded in a national economy in some time period. The asset market
equilibrium equation is M/P = L(Y, r + πe).
Diff: 1
Topic: Section 7.4
Question Status: New

23
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12) Suppose the money demand function is given by
Md/P = 640 + 0.1Y - 5000 (r + πe).
Suppose the central bank changes the nominal money supply depending on income and inflation:
Ms = 1000 + 0.1Y - 4000π.
(a) If expected inflation equals actual inflation = 0.03, Y = 1000, and r = 0.02, calculate the price
level.
(b) If inflation rises to 0.04 while the other variables remain as in part a, calculate the price
level.
(c) If expected inflation rises to 0.04 while the other variables remain as in part a, calculate the
price level.
(d) If the real interest rate rises to 0.03 while the other variables remain as in part a, calculate the
price level.
Answer:
Plug in the value of Y and use text Eq. (7.10) to get P = [1100 - 4000π]/[740 - 5000(r + )].
When r = 0.02, this becomes P = [1100 - 4000π]/[640 - 5000π].
(a) P = 980/490 = 2.
(b) P = 940/490 = 1.92.
(c) P = 980/440 = 2.23.
(d) P = 980/440 = 2.23.
Diff: 2
Topic: Section: 7.4
Question Status: Previous Edition

13) Assume that prices and wages adjust rapidly so that the markets for labor, goods, and assets
are always in equilibrium. What are the effects of each of the following on output, the expected
real interest rate, and the current price level?
(a) a temporary increase in taxes
(b) a reduction in the effective tax rate on capital
(c) an increase in expected inflation
Answer:
(a) Under Ricardian equivalence, no effect on output, real interest rate, or price level. Without
Ricardian equivalence, higher national saving means a lower expected real interest rate. Output
is unchanged because of no change in labor supply or demand. The lower expected real interest
rate increases real money demand, thus reducing the price level.
(b) The lower tax rate on capital increases desired investment, thus raising the expected real
interest rate. No effect on the labor market, so output is not changed. The higher expected real
interest rate reduces real money demand, thus increasing the price level.
(c) The increase in the expected inflation rate has no effect on the labor market or goods market,
so output and the expected real interest rate do not change. The higher expected inflation rate
reduces real money demand, thus increasing the price level.
Diff: 2
Topic: Section: 7.4
Question Status: Previous Edition

24
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7.5 Money Growth and Inflation

1) If the nominal money supply doubles while real money demand is unchanged, what happens
to the price level?
A) The price level increases by a factor of four.
B) The price level doubles.
C) The price level is unchanged.
D) The price level falls by one-half.
Answer: B
Diff: 1
Topic: Section: 7.5
Question Status: Previous Edition

2) If real money demand doubles while the nominal money supply is unchanged, what happens
to the price level?
A) The price level increases by a factor of four.
B) The price level doubles.
C) The price level is unchanged.
D) The price level falls by one-half.
Answer: D
Diff: 1
Topic: Section: 7.5
Question Status: Previous Edition

3) If nominal money supply grows 3% and real money demand grows 8%, the inflation rate is
A) -5%.
B) 8/3%.
C) 5%.
D) 11%.
Answer: A
Diff: 2
Topic: Section: 7.5
Question Status: Previous Edition

4) If real money demand increases 5% and real money supply increases 10%, by about how
much does the price level change?
A) Falls by 5%
B) Unchanged
C) Rises by 2%
D) Rises by 5%
Answer: D
Diff: 1
Topic: Section: 7.5
Question Status: Previous Edition

25
Copyright © 2014 Pearson Education, Inc.
5) If the income elasticity of money demand is 3/4 and income increases 8%, by about how much
does the price level change?
A) Falls by 6%
B) Unchanged
C) Rises by 6%
D) Rises by 8%
Answer: A
Diff: 1
Topic: Section: 7.5
Question Status: Previous Edition

6) If the nominal money supply grows 5%, real income falls 2%, and the income elasticity of
money demand is 0.8, then the inflation rate is
A) 3.0%.
B) 3.4%.
C) 6.6%.
D) 7.0%.
Answer: C
Diff: 2
Topic: Section: 7.5
Question Status: Previous Edition

7) If the nominal money supply grows 6%, real income rises 2%, and the inflation rate is 5%,
then the income elasticity of money demand is
A) 0.5.
B) 0.75.
C) 1.0.
D) 1.5.
Answer: A
Diff: 2
Topic: Section: 7.5
Question Status: Previous Edition

8) If the nominal money supply grows 10%, the inflation rate is 6%, and the income elasticity of
money demand is 1.0, then real income growth equals
A) 1%.
B) 2%.
C) 3%.
D) 4%.
Answer: D
Diff: 2
Topic: Section: 7.5
Question Status: Previous Edition

26
Copyright © 2014 Pearson Education, Inc.
9) Large differences in inflation rates among countries are almost always the result of large
differences in
A) productivity.
B) real income growth.
C) the growth rates of real money demand.
D) the growth rates of nominal money supplies.
Answer: D
Diff: 1
Topic: Section: 7.5
Question Status: Previous Edition

10) When a government prints money to finance its expenditures, it is likely to cause
A) unemployment.
B) inflation.
C) deflation.
D) reductions in the use of barter.
Answer: B
Diff: 1
Topic: Section: 7.5
Question Status: Previous Edition

11) The most likely explanation for the high inflation rates that countries like Russia and the
Ukraine have suffered is that
A) large inflows of foreign funds increase the money supply, causing inflation.
B) without inflation, these countries would be unable to achieve high rates of growth.
C) borrowing from the central bank is the most expedient method of funding the government's
expenditures.
D) the flood of financial innovations has increased liquidity in these nations' economies.
Answer: C
Diff: 2
Topic: Section: 7.5
Question Status: Previous Edition

12) Bonds sold by the U.S. government that offer a certain real interest rate are known as
A) zero-coupon bonds.
B) Treasury Inflation-Protected Securities.
C) denominalized securities.
D) savings bonds.
Answer: B
Diff: 2
Topic: Section: 7.5
Question Status: New

27
Copyright © 2014 Pearson Education, Inc.
13) Calculate the change in the price level for each of the following events, taken one at a time,
with other variables unchanged.
(a) Money supply increases 10%.
(b) Money demand increases 5%.
(c) Money supply decreases 5% while money demand increases 5%.
(d) Money supply increases 15% while money demand increases 5%.
Answer:
(a) 10%
(b) -5%
(c) -10%
(d) 10%
Diff: 1
Topic: Section: 7.5
Question Status: Previous Edition

14) Why did some of the formerly Communist countries of Eastern Europe have inflation rates
over 100%, while others didn't? Which factor was more important in explaining the differing
inflation rates, real money demand or nominal money supply? Why did the countries with high
inflation rates allow inflation to get so high?
Answer: Some countries had high inflation while others didn't because of differences in rates of
money growth. Real money demand didn't vary enough to explain the differences in inflation
rates; instead, nominal money supply growth was strongly correlated with inflation. The
countries allowed inflation to get so high because they were trying to finance government
expenditures by printing money.
Diff: 1
Topic: Section: 7.5
Question Status: Previous Edition

28
Copyright © 2014 Pearson Education, Inc.
Another random document with
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absent in Myxinoids, but the gastric branches of the Vagus are
continued, united as a single nerve, along the intestine to the anus.
No fish possesses a Nervus accessorius. Also a separate Nervus
hypoglossus (twelfth pair)[13] is absent, but elements from the first
spinal nerve are distributed in the area normally supplied by this
nerve in higher vertebrates.

The number of Spinal nerves corresponds to that of the vertebræ,


through or between which they pass out. Each nerve has two roots,
an anterior and posterior, the former of which has no ganglion, and
exclusively contains motor elements. The posterior or dorsal has a
ganglionic enlargement, and contains sensory elements only. After
leaving the vertebral canal each spinal nerve usually divides into a
dorsal and ventral branch. The Gadoids show that peculiarity that
each of the posterior roots of some or many of the spinal nerves
possesses two separate threads, each of which has a ganglion of its
own; the one of these threads joins the dorsal and the other the
ventral branch. In fishes in which the spinal chord is very short, as in
Plectognaths, Lophius, the roots of the nerves are extremely long,
forming a thick Cauda equina. The additional function which the
(five) anterior spinal nerves of Trigla have to perform in supplying the
sensitive pectoral appendages and their muscles has caused the
development of a paired series of globular swellings of the
corresponding portion of the spinal chord. A similar structure is found
in Polynemus.
Fig. 46.
Brain and anterior portion of the
spinal chord of Trigla
(Gurnard), showing the
globular swellings at the
base of the anterior spinal
nerves.
A Sympathic nervous system appears to be absent in
Branchiostoma, and has not yet been clearly made out in
Cyclostomes. It is well developed in the Palæichthyes, but without
cephalic portion. This latter is present in all Osseous fishes, in which
communication of the Sympathic has been found to exist with all
cerebral nerves, except the olfactory, optic, and acustic. The
sympathic trunks run along each side of the aorta and the back of
the abdomen into the hæmal canal; communicate in their course with
the ventral branches of each of the spinal nerves; and, finally, often
blend together into a common trunk beneath the tail. At the points of
communication with the cerebral and spinal nerves frequently
ganglia are developed, from which nerves emerge which are
distributed to the various viscera.
CHAPTER VII.

THE ORGANS OF SENSE.

Characteristic of the Organ of Smell in Fishes is that it has no


relation whatever to the respiratory function, with the exception of the
Dipnoi, in which possibly part of the water received for respiration
passes through the nasal sac.
The olfactory organ is single in Branchiostoma and the
Cyclostomes. In the former a small depression on the front end of
the body, clothed with a ciliated epithelium, is regarded as a
rudimentary organ of smell. In the adult Petromyzon a membranous
tube leads from the single opening on the top of the head into the
cartilaginous olfactory capsule, the inside of which is clothed by
membranes prolonged into a posterior blind tube (Fig. 30, s), which
penetrates the cartilaginous roof of the palate, but not the mucous
membrane of the buccal cavity. In the Myxinoids the outer tube is
strengthened by cartilaginous rings like a trachea; the capsule is
lined by a longitudinally folded pituitary membrane, and the posterior
tube opens backwards on the roof of the mouth; the opening is
provided with a valve.
In all other Fishes the organ of smell is double, one being on
each side; it consists of a sac lined with a pituitary membrane, and
without, or with one or two, openings. The position of these openings
is very different in the various orders or suborders of Fishes.
In the Dipnoi the nasal sac opens downwards by two wide
openings which are within the boundaries of the cavity of the mouth.
The pituitary membrane is transversely folded, the transverse folds
being divided by one longitudinal fold. The walls of the sac are
strengthened by sundry small cartilages.
Also in Chondropterygians the openings, of which there is one to
each sac, are on the lower part of the snout, and in the Rays,
Holocephali, and some Sharks, each extends into the cleft of the
mouth. The openings are protected by valvular flaps, supported by
small cartilages, and moved by muscles, whence it may be
concluded that these fishes are able to scent (actively) as well as to
smell (passively).

Fig. 47.—Nostrils of Raia lemprieri,


with nasal flaps reverted.
In the majority of Teleostei the olfactory capsules are lateral or
superior on the snout, covered externally by the skin, each usually
pierced by two openings, which are either close together, or more or
less remote from each other; the posterior is generally open, the
anterior provided with a valve or tube. In the Chromides and
Labroidei ctenoidei a single opening only exists for each sac. In the
Murænidæ the two openings of each side are either superior, or
lateral, or labial, that is, they are continued downwards and pierce
the margin of the upper lip. In many Tetrodonts nasal openings are
absent, and replaced by a conical papilla, in which the olfactory
nerve terminates.
It is certain that fishes possess the faculty of perceiving odours,
and that various scents attract or repel them. A mangled carcase or
fresh blood attracts Sharks as well as the voracious Serrasal
monoids of the South American rivers. There is no reason to doubt
that the seat of that perception is in the olfactory sac; and it may be
reasonably conjectured that its strength depends mainly on the
degree of development indicated by the number and extent of the
interior folds of the pituitary membrane.
Organ of Sight.—The position, direction, and dimensions of the
eyes of fishes vary greatly. In some they have an upward aspect,
and are often very close together; in others they are lateral, and in a
few they are even directed downwards. The Flat-fishes represent the
extraordinary anomaly that both eyes are on the same side of the
head, and rarely on the same level, one being generally placed more
forward than the other. In certain species of marine fishes the eyes
are of an extraordinary size, a peculiarity indicating that the fish
either lives at a great depth, to which only a small proportion of the
rays of light penetrate, or that it is of nocturnal habits. In fishes which
have descended to such great depths that no rays whatever can
reach them, or in freshwater fishes living in caves, or in species
which grovel and live constantly in mud, the eyes are more or less
aborted, sometimes quite rudimentary, and covered by the skin. In
very few this organ appears to be entirely absent. In some Gobioids
and Trachinoids (Periophthalmus, Boleophthalmus, Uronoscopus,
etc.) the eyes, which are on the upper side of the head, can be
elevated and depressed at the will of the fish. In the range of their
vision and acuteness of sight, Fishes are very inferior to the higher
classes of Vertebrates, yet at the same time it is evident that they
perceive their prey or approaching danger from a considerable
distance; and it would appear that the visual powers of a
Periophthalmus, when hunting insects on mud-flats of the tropical
coasts, are quite equal to that of a frog. Again, the discrimination
with which fishes sometimes prefer one colour or kind of artificial fly
to another affords sufficient evidence that the vision, at least of
certain species is by no means devoid of clearness and precision.
The eye of Branchiostoma is of the most rudimentary condition. It
is simply a minute speck coated by dark pigment, and receiving the
end of a short nerve. In Myxinoids the minute rudiment of the eye is
covered by the skin and muscles. This is also the case in many of
the blind Teleosteous fishes; however, whilst in the former fishes the
organ of sight has not attained to any degree of development, the
rudimentary eye of blind Teleostei is a retrogressive formation, in
which often a lens and other portions of the eye can be recognised.
In fishes with a well-developed eye it is imbedded in a layer of
gelatinous and adipose substance, which covers the cavity of the
orbit. A lacrymal gland is absent. In the orbit of one fish only,
Chorismodentex, an organ has been found which can be compared
to a saccus lacrymalis. It is a round, blind, wide sac, of the size of a
pea, situated below the anterior corner of the orbit, between the
maxillary bone and the muscles of the cheek, communicating by a
rather wide foramen with the orbital cavity. The membrane by which
it is formed is continuous with that coating the orbita. In the
Chondropterygians the eyeball is supported by and moves on a
cartilaginous peduncle of the orbital wall. In the majority of
Teleosteans, and in Acipenser, a fibrous ligament attaches the
sclerotic to the wall of the orbit. The proper muscles of the eyeball
exist in all fishes, and consist of the four Musculi recti and the two M.
obliqui. In many Teleostei the former rise from a subcranial canal, the
origin of the M. rectus externus being prolonged farthest backwards.
The Recti muscles are extraordinarily long in the Hammerheaded
Sharks, in which they extend from the basis cranii along the lateral
prolongations of the head to the eyes, which are situated at the
extremities of the hammer.
In all fishes the general integument of the head passes over the
eye, and becomes transparent where it enters the orbit; sometimes it
simply passes over the orbit, sometimes it forms a circular fold. The
anterior and posterior portions may be especially broad and the seat
of an adipose deposit (adipose eyelids), as in Scomber, Caranx,
Mugil, etc. In many of these fishes the extent of these eyelids varies
with the seasons; during the spawning season they are so much
loaded with fat as nearly to hide the whole eye. Many Sharks
possess a nictitating membrane, developed from the lower part of
the palpebral fold, and moved by a proper set of muscles.
Fig. 48.
Vertical section through eye of
Xiphias. (After Owen.)
co, Cornea; sc, sclerotica; o, nervus
opticus; c, sclerotic capsule; a,
membrana argentea; v,
membrana vasculosa; u,
membrana uvea; ch, choroid
gland; r, retina; f, processus
falciformis; h, humor vitreus; l,
lens; i, iris.

The form of the bulbus (Fig. 48) is subhemispherical, the cornea


(co) being flat. If it were convex, as in higher Vertebrates, it would be
more liable to injury; but being level with the side of the head the
chances of injury by friction are diminished. The sclerotica (sc) is
cartilaginous in Chondropterygians and Acipensers, fibrous and of
varying thickness in Teleosteans, in the majority of which it is
supported by a pair of cartilaginous or ossified hemispheroid cups
(c). In a few fishes, as in Ceratodus, Xiphias, the cups are confluent
into one cup, which possesses a foramen behind to allow the
passage of the optic nerve (o). The cornea of Anableps shows an
unique peculiarity. It is crossed by a dark horizontal stripe of the
conjunctiva, dividing it into an upper and lower portion; also the iris is
perforated by two pupils. This fish is observed to swim frequently
with half of its head out of the water, and it is a fact that it can see
out of the water as well as in it.
The membranes situated between the sclerotica and retina are
collectively called choroidea, and three in number. The one in
immediate contact with the sclerotic, and continued upon the iris, is
by no means constantly present; it is the membrana argentea (a),
and composed of microscopical crystals reflecting a silvery or
sometimes golden lustre. The middle layer is the membrana
vasculosa s. halleri (v), the chief seat of the ramifications of the
choroid vessels; the innermost layer is the membrana ruyscheana or
uvea (u), which is composed of hexagonal pigment-cells, usually of a
deep brown or black colour.
In many Teleostei a rete mirabile surrounds the entry of the optic
nerve; it is situated between the membrana argentea and vasculosa,
and called the choroid gland (ch). It receives its arterial blood from
the artery issuing from the pseudobranchia; the presence of a
choroid gland always being combined with that of a pseudobranchia.
Teleosteans without pseudobranchia lack a choroid gland. In the
Palæichthyes, on the other hand, the pseudobranchia is present and
a choroid gland absent.
The iris (i) is merely the continuation of the choroid membrane; its
capability of contracting and expanding is much more limited than in
higher Vertebrates. The pupil is generally round, sometimes
horizontally or vertically elliptical, sometimes fringed. In the Rays and
Pleuronectidæ a lobe descends from the upper margin of the pupil,
and the outer integument overlying this lobe is coloured and non-
transparent; a structure evidently preventing light from entering the
eye from above.
In most Teleostei a fold of the Choroidea, called the Processus
falciformis (f), extends from the vicinity of the entrance of the optic
nerve to the lens. It seems to be constantly absent in Ganoids.
The retina (r) is the membrane into which the optic nerve
penetrates, and in which its terminal filaments are distributed. It
consists of several layers (Fig. 49). The outermost is an extremely
delicate membrane (a), followed by a layer of nerve-cells (b), from
which the terminal filaments issue, passing through several granular
strata (c, d, e), on which the innermost stratum rests. This stratum is
composed of cylindrical rods (f) vertically arranged, between which
twin fusiform corpuscles (g) are intercalated. This last layer is thickly
covered with a dark pigment. The retina extends over a portion of the
iris, and a well-defined raised rim runs along its anterior margin.

Fig. 49.—Vertical section of


the Retina of the Perch, magn. X
350.
The vitreous humour (Fig. 48, h) which fills the posterior cavity of
the eyeball, is of a firmer consistency than in the higher Vertebrates.
The lens is spherical, or nearly so; firm, denser towards the centre,
and lies in a hollow of the vitreous humour. When a falciform process
is present, it is with one end attached to the lens, which is thus
steadied in its position. It consists of concentric layers consisting of
fibres, which in the nucleus of the body have marginal teeth, by
which they are interlocked together. In Petromyzon this serrature is
absent, or but faintly indicated.

Fig. 50.—Interlocking fibres of lens,


highly magnified.
The anterior cavity of the eye is very small in Fishes, in
consequence of the small degree of convexity of the cornea; the
quantity of the aqueous humour, therefore, is very small, just
sufficient to float the free border of the iris; and the lessened
refractive power of the aqueous humour is compensated by the
greater convexity of the lens.

Organ of Hearing.—No trace of an organ of hearing has been


found in Branchiostoma. In the Cyclostomes the labyrinth is
enclosed in externally visible cartilaginous capsules laterally
attached to the skull; it consists of a single semicircular canal in the
Myxinoids, whilst the Petromyzontes possess two semicircular
canals with a vestibulum.
In all other fishes the labyrinth consists of a vestibule and three
semicircular canals, the vestibule dilating into one or more sacs
which contain the otoliths. A tympanum, tympanic cavity, and
external parts, are entirely absent in the class of fishes.
In the Chondropterygians and Dipnoi, the labyrinth is enclosed in
the cartilaginous substance of the skull. In the former the excavation
in the cartilage is larger than the membranous labyrinth, but nearly
corresponds to it in form; the part which receives the membranous
vestibulum is called Vestibulum cartilagineum, from which a canal
issues and penetrates to the surface of the skull, where it is closed
by the skin in Sharks, but opens by a minute foramen in Rays. The
otolithic contents are soft and chalklike.
In the Holocephali part of the labyrinth is enclosed in the cartilage
of the skull, another part being in the cranial cavity, as in Ganoids
and Teleosteans. The membranous vestibulum is continued by a
canal to a single opening in the roof of the skull, from which two
smaller canals are continued to two small foramina in the skin
covering the occipital region.
In the Teleosteans the sac which contains the otoliths lies on
each side of the base of the cranial cavity and is often divided by a
septum into two compartments of unequal size, each containing a
firm and solid otolith; these bodies (Fig. 51), possess indented
margins, frequently other impressions and grooves, in which nerves
from the N. acusticus are lodged; they vary much in size and form,
but in both respects show a remarkable constancy in the same kind
of fishes. The vestibule is outwards in contact with the osseous side
wall of the skull, inwards with the metencephalon and medulla
oblongata; it contains another firm concretion, and opens by five
foramina into the three semicircular canals. The terminations of the
acustic nerve are distributed over the vestibular concretion and the
ampulliform ends (Fig. 52 p) of the semicircular canals, without being
continued into the latter, which are filled with fluid. The semicircular
canals (Fig. 52 g), are sometimes lodged in the cranial bones,
sometimes partly free in the cranial cavity. Many Teleostei have
fontanelles in the roof of the skull, closed by skin or very thin bone
only at the place where the auditory organ approaches the surface,
by which means sonorous undulations must be conducted with
greater ease to the ear.

Fig. 51.—Otolith of Haddock (Gadus


æglefinus). I. Outer, II. Inner aspect.
In many Teleostei a most remarkable relation obtains between
the organ of hearing and the air-bladder. In the most simple form this
connection is established in Percoids and the allied families, in which
the two anterior horns of the air-bladder are attached to fontanelles
of the occipital region of the skull, the vestibulum occupying the
opposite side of the membrane by which the fontanelle is closed.
The condition is similar, but more complicated in many Clupeoids.
The anterior narrow end of the air-bladder is produced into a canal at
the base of the skull, and divided into two very narrow branches,
which again bifurcate and terminate in a globular swelling. An
appendage of the vestibulum meets the anterior of these swellings,
and comes into close contact with it. Besides, the two vestibules
communicate with each other by a transverse canal, crossing the
cranial cavity below the brain.
Fig. 52.—Communication between auditory organ and air-bladder in the Carp.
(After E. H. Weber.)
a, Basisphenoid; b, Occipital; c, Supraoccipital; d, Exoccipital; e, Paroccipital; f,
Alisphenoid; g, Neural arch of first vertebra; h, i, k, second, third, and fourth
vertebra; h’, i’, Parapophyses of second and third vertebra; i", process of the
third vertebra for the attachment of the air-bladder; k, l, m, Chain of ossicles;
n, Air-bladder; o, vestibulum; p, p, Ampullæ; q, q, Canales semicirculares; r,
Sinus impar.
The connection is effected by means of a chain of ossicles in
Siluridæ, Characinidæ, Cyprinidæ and Gymnotidæ. A canal issues
from the communication between vestibule and its sac, and meeting
that from the other side forms with it a common sinus impar (Fig. 52,
r), lodged in the substance of the basi-occipital; this communicates
on each side by a small orifice with two subspherical atria, on the
body of the atlas, close to the foramen magnum. Each atrium is
supported externally by a small bone (m); a third larger bone (k)
completes the communication with the anterior part of the air-
bladder. From the sinus impar a bifid canal penetrates into the
alisphenoids, in which it terminates. In Cobitis and several Loach-like
Siluroids the small air-bladder consists of two globular portions
placed side by side, and wholly included within two bullæ, formed by
the modified parapophyses of the second and third vertebræ. The
three ossicles on each side are present, but concealed by the fore
part of the osseous bulla.

Organ of Taste.—Some fishes, especially vegetable feeders, or


those provided with broad molar-like teeth, masticate their food; and
it may be observed in Carps and other Cyprinoid fish, that this
process of mastication frequently takes some time. But the majority
of fish swallow their food rapidly, and without mastication, and
therefore we may conclude that the sense of taste cannot be acute.
The tongue is often entirely absent, and even when it exists in its
most distinct state, it consists merely of ligamentous or cellular
substance, and is never furnished with muscles capable of producing
the movements of extension or retraction as in most higher
Vertebrates. A peculiar organ on the roof of the palate of Cyprinoids,
is perhaps an organ adapted for perception of this sense; in these
fishes the palate between and below the upper pharyngeal bones is
cushioned with a thick, soft contractile substance, richly supplied
with nerves from the Nervi vagus and glossopharyngeus.
Organs of Touch.—The faculty of touch is more developed than
that of taste, and there are numerous fishes which possess special
organs of touch. Most fishes are very sensitive to external touch,
although their body may be protected by hard horny scales. They
perceive impressions even on those parts which are covered by
osseous scutes, in the same manner as a tortoise perceives the
slightest touch of its carapace. The seat of the greatest
sensitiveness, however, appears to be the snout and the labial folds
surrounding the mouth. Many species possess soft and delicate
appendages, called barbels, which are almost constantly in action,
and clearly used as organs of touch. Among the Triglidæ and allied
families, there are many species which have one or more rays of the
pectoral fin detached from the membrane, and supplied with strong
nerves. Such detached rays (also found in the Polynemidæ,
Bathypterois) are used partly for locomotion, partly for the purpose of
exploring the ground over which the fish moves.
Some fish appear to be much less sensitive than others, or at
least lose their sensitiveness under peculiar circumstances. It is well
known that a Pike, whose mouth has been lacerated and torn by the
hook, continues to yield to the temptation of a bait immediately
afterwards. The Greenland Shark when feeding on the carcass of a
whale allows itself to be repeatedly stabbed in the head without
abandoning its prey. A pair of Congers are so dead to external
impression at the time of copulation, and so automatically, as it were,
engaged, that they have been taken by the hand together out of the
water.
CHAPTER VIII.

THE ORGANS OF NUTRITION AND DIGESTION.

Fishes are either exclusively carnivorous or herbivorous, but not


a few feed on vegetable substances as well as animal, or on mud
containing alimentary substance in a living or decomposing state.
Generally they are very voracious, especially the carnivorous kinds,
and the rule of “eat or be eaten” applies to them with unusual force.
They are almost constantly engaged in the pursuit and capture of
their prey, the degree of their power in these respects depending on
the dimensions of the mouth and gullet and the strength of the teeth
and jaws. If the teeth are sharp and hooked, they are capable of
securing the most slender and agile animals; if this kind of teeth is
combined with a wide gullet and distensible stomach, they are able
to overpower and swallow other fish larger than themselves; if the
teeth are broad, strong molars, they are able to crush the hardest
aliments; if they are feeble, they are only serviceable in procuring
some small or inert and unresisting prey. Teeth may be wanting
altogether. Whatever the prey, in the majority of cases it is swallowed
whole; but some of the most voracious fishes, like some Sharks and
Characinidæ, are provided with cutting teeth, which enable them to
tear their prey to pieces if too large to be swallowed whole. Auxiliary
organs for the purpose of overpowering their prey, which afterwards
is seized or torn by the teeth, like the claws of some carnivorous
mammals and birds, are not found in this class; but in a few fishes
the jaws themselves are modified for that purpose. In the Sword-
fishes the bones of the upper jaw form a long dagger-shaped
weapon, with which they not only attack large animals, but also
frequently kill fishes on which they feed. The Saw-fishes are armed
with a similar but still more complicated weapon, the saw, which is
armed on each side with large teeth implanted in deep sockets,
specially adapted for killing and tearing the prey before it is seized
and masticated by the small teeth within the mouth. Fishes show but
little choice in the selection of their food, and some devour their own
offspring indiscriminately with other fishes. Their digestive powers
are strong and rapid, but subject in some degree to the temperature,
which, when sinking below a certain point, lowers the vital powers of
these cold-blooded animals. On the whole, marine fishes are more
voracious than those inhabiting fresh waters; and whilst the latter
may survive total abstinence from food for weeks or months, the
marine species succumb to hunger within a few days. The growth of
fishes depends greatly on the nature and supply of food, and
different individuals of the same species may exhibit a great disparity
in their respective dimensions. They grow less rapidly and to smaller
dimensions in small ponds or shallow streams than in large lakes
and deep rivers. The young of coast fishes, when driven out to sea,
where they find a much smaller supply of food, remain in an
undeveloped condition, assuming an hydropic appearance. The
growth itself seems to continue in most fishes for a great length of
time, and we can scarcely set bounds to—certainly we know not with
precision—the utmost range of the specific size of fishes. Even
among species in no way remarkable for their dimensions we
sometimes meet with old individuals, favourably situated, which
more or less exceed the ordinary weight and measurement of their
kind. However, there are certain evidently short-lived species of
fishes which attain a remarkably uniform size within a very short
time; for instance, the Stickleback, many species of Gobius and
Clupea.

The organs of nutrition, manducation, and deglutition, are lodged


in two large cavities—an anterior (the mouth or buccal cavity), and a
posterior (the abdominal cavity). In the former the alimentary organs
are associated with those fulfilling the respiratory functions, the
transmission of food to the stomach and of water to the gills being
performed by similar acts of deglutition. The abdominal cavity
commences immediately behind the head, so, however, that an
extremely short thoracic cavity for the heart is partitioned off in front.
Beside the alimentary organs it contains also those of the urogenital
system and the air-bladder. The abdominal cavity is generally
situated in the trunk only, but in numerous fishes it extends into the
tail, being continued for some distance along each side of the hæmal
apophyses.
In numerous fishes the abdominal cavity opens outwards by one
or two openings. A single porus abdominalis in front of the vent is
found in Lepidosiren and some Sturgeons; a paired one, one on
each side of the vent, in Ceratodus, some species of Sturgeon,
Lepidosteus, Polypterus, Amia, and all Chondropterygians. As in
these fishes semen and ova are discharged by proper ducts, the
abdominal openings may serve for the expulsion of semen, and
those ova only which, having lost their way to the abdominal
aperture of the oviduct, would be retained in the abdominal cavity. In
those Teleosteans which lack an oviduct a single porus genitalis
opens behind the vent.
The mouth of fishes shows extreme variation with regard to form,
extent, and position. Generally opening in front, it may be turned
upwards, or may lie at the lower side of the snout, as in most
Chondropterygians, Sturgeons, and some Teleosteans. Vogt regards
this position as a persistent fœtal condition. In most fishes the jaws
are covered by the skin, which, before passing over the jaws, is often
folded, forming more or less fleshy lips. In the Sharks the skin retains
its external character even within the teeth, but in other fishes it
changes into a mucous membrane. A tongue may exist as a more or
less free and short projection, formed by the glosso*-hyal and a soft
covering, or may be entirely absent. Salivary glands and a velum
palati are absent in fishes.
With regard to the dentition, the class of Fishes offers an amount
of variation such as is not found in any of the other classes of
Vertebrates. As the teeth form one of the most important elements in
the classification of fishes, their special arrangement and form will be
referred to in the account of the various families and genera. Whilst
not a few fishes are entirely edentulous, in others most of the bones
of the buccal cavity, or some of them, may be toothed, as the bones
of the jaws, the palatines, pterygoids, vomers, basisphenoid,
glossohyal, branchial arches, upper and lower pharyngeals. In others
teeth may be found fixed in some portion of the buccal membrane
without being supported by underlying bone or cartilage; or the teeth

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