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Fee HA EE)Editorial Committee of Palaeovertebrata Sinica PALAEOVERTEBRATA SINICA Volume II Amphibians, Reptilians, and Avians Editor-in-Chief: Li Jinling | Associate Editor-in-Chief: Zhou Zhonghe Fascicle 4 (Serial no. 8) Basal Archosauromorphs, Crocodylomorphs, and Pterosaurs By Wu Xiaochun, Li Jinling, Wang Xiaolin et al. Supported by the Special Research Program of Basic Science and Technology of the Ministry of Science and Technology (2006FY 120400) Science Press BeijingCHET APMED ) iF BAS ff: He WHE: KAS AH BB oh (Ununhtwe) : oe Gh FR] 2 Hh SER SH ke ERY REE Ee HAR Ew ER ELH 24K RH RE REG KE KLE ARM KR Editorial Committee of Palaeovertebrata Sinica Chairman; Qiu Zhanxiang Vice-Chairpersons: Zhang Miman and Wu Xinzhi Members; Deng Tao, Gao Keqin, Hu Yaoming, Jin Fan, Li Chuankui, Li Jinling, Meng Jin, Miao Desui, Ni Xijun, Qiu Zhanxiang, Qiu Zhuding, Wang Xiaoming, Wang Yuan, Wang Yuanging, Wu Xiaochun, Wu Xinzhi, Xu Xing, You Hailu, Zhang Miman, Zhang Zhaoqun, Zhou Zhonghe, and Zhu MinASMBES A BAYT. BFERD KW AR E-mail: xewu@mus-nature.ca SHABSS RHA SRABRRICE 4P44H E-mail: lijinling@ivppaccn RHA RASS 3£ MAK E-mail: wangxiaolin@ivppac.n 4ENLSS E-mail: jiangshunxing@ivpp.ac.cn “CRIB” 4% 3 E-mail: cheng_xin1982@126.com TK RFZ0B HM REA CRM ATEN FD TEAL Pie A aE EB 1 FL ttf eT aia tit i Sa Efe PR TE Acs ehh, HAE, EHR. TERK, FEM, DSC RR. HI AEB tt Mab ois SAE HP TLE BEAAE) Wy}Contributors to this Fascicle Basal Archosauromorphs ‘Wu Xiaochun E-mail: xewu@mus-nature.ca Introduction to Crocodylomorpha —- Wu Xiaochun Crocodylomorpha LiJinling E-mail: lijinling@ivpp.ac.cn Wu Xiaochun Introduction to Pterosauria Wang Xiaolin E-mail: wangxiaolin@ivppac.cn Jiang Shunxing E-mail: jiangshunxing@ivpp.acen Suborder “Rhamphorhynchoidea” Cheng Xin. E-mail: cheng_xin1982@126 4&*# jie (evolutionary taxonomy), L, Hennig 1 {t4ehy Ke FF AGCEIR [phylogenetic systematics, MPI KA (cladistic systematics, eft {9% cladistics) ] JL Sokal All Sneath Yy fC 2EHI Hel A AF (numerical taxonomy) fY “= feo.” AVI. 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HERE PE De AS ESE SE: RRR sve ; 1: Je//)44 Parvielass ARCHOSAUROMORPHA 10 SETA@MUH Basal archosauromorphs-- 10 \iJEH Order PROTOROSAURIA . un KHUEF Family Tanystropheidae - FRAG Genus Macrocnemus wed {SEI Genus Tanystropheus ~~ JOE FARE Protorosauria incertae familiae Jeli Genus Fuyuansaurus Heli, Genus Dinocephalosaurus JSF} Family Prolacertidae Parrington- ANGMAR Genus Prolacertoidess---- JE} YA Subparviclass ARCHOSAURIFORMES:- JEP IRIE Basal archosauriforms ii@6} Family Proterosuchidae fi) Genus Proterosuchus 41951 Family Erythrosuchidae~ 1758, Genus ShansisuchusHRERH IR Genus Vjushkovia~ sera 27 HB I Genus Guchengosuchus « URSEHIAL Family Euparkeriidae Mfr HeENRY I Genus Halachaisuchus- Ob 4 85 I? Genus Turfanosuchus? seen 31 JEP ADEEAAGE Basal archosauriforms incertae familiae 34 HE PEHS IB Genus Fugusuchus 5 | Genus Litorosuchus £ RWS IR ? Genus Wangisuchus? DHHS IR 2 Genus Fenhosuchus’ UEH Order PHYTOSAURIA BEAWEIR Genus Diandongosuchus rrr 39 +J€H. A Magnorder ARCHOSAURIA (283: 1 Superorder PSEUDOSUCHIA -- STHIGSEL Family Gracilisuchidae 851 Genus Turfanosuchus ACAIS hh Genus Yonghesuchus YEAH Grandorder POPOSAUROIDEA-~ W583 IR Genus Qianosuchus AURIS I, Genus Xilousuchus----- SERENE Genus Lotosaurus 49 “EEE ASIANA Archosauria gen. et sp. probl. An Mt HEIR 2 Genus Chasmatosaurus' 4W3)K 2 Genus Strigosuchus ‘Gii(6°3 J, 2. Genus Pachysuchus? Boa ww S2A8S5 ~~ ee 2, WEY EI RDG SLB REE TR ssceseeee SL Hiatt: SERA EIRP A DOS A EAC IDET prvi ee BRR: 66, KH Grandorder CROCODYLOMORPHA YEN Order SPHENOSUCHIA ~~ BYEMIFL Family Sphenosuchidae-BUM5 1K, Genus Dibothrosuchus~ 67 BOERS ALAS Sphenosuchia incertae familiae -----------68 "FEHR I Genus Phyllodontosuchus ~~ HENS 21583, Genus Junggarsuchus ------ @EWAH Subgrandorder CROCODYLIFORMES §H8H Order PROTOSUCHIA ~~ UGVIEL Family Edentosuchidae ~ RGGI Genus Edentosuchus SUEWAA RA Crocodyliformes incertae familiae -- i, Genus Dianosuchus it PY JIBS fa, Genus Sichuanosuchus-~ ES /\nff3 I 7 Genus Microchampsa’ E085") F] Mirorder MESOEUCROCODYLIA. "PY 5[ HAASE Mesoeucrocodylia incertae fa ihe Genus Dianchungosaurus 4854 Family Hsisosuchidae ~ PERS Genus Hsisosuchus 141831 Order NOTOSUCHIA- Wil9 HALAS Notosuchia incertae familiae- ‘#ifEW3 ll Genus Chimaerasuchus evsaneseanrneseneee BB MHA Submirorder NEOSUCHIA ~~ #7H3A} Family Atoposauridae-- SF Genus Theriosuchus EUHSTFL Family Teleosauridae: ARS) Genus Peipehsuchus SUBS} Family Goniopholi HERS IR Genus Sunosuchus APLUETAL Family Paralligatoridae~ SKEW JR Genus Rugosuchus-~ SU FHA Inframirorder EUSUCHIA 83H Order CROCODYLIA - 185A} Family Planocraniidae- °F (WS 1 Genus Planocrania + KIMMEL Suborder BREVIROSTRES ~~ fi Genus Platyognathus 5 Genus Shantungosuchus jaeWAFL Superfamily Crocodyloide: IEHH#S I, Genus Asiatosuchus « 8) Family Crocodylidae------------ ‘YIKGEAL Subfamily Tomistominae ~~ 0608 It, Genus Tomistomars-- WINS I Genus Penghusuchus S10} Subfamily Crocodylinae JAWS IR? Asiatosuchus? ~ ‘LUNHSFL Family Alligatoridae --~ ‘HKALE Subfamily Alligatorinae--- ‘SkWDHS I Genus Alligator Ji #3 i, Genus Protoalligator ul LUTTE MLB AE Alligatoridae gen, et sp. indet. -----112 SOS BA BAS: fii Jat, Genus Wanosuchus « Eusuchia incertae familiae ---------- 113 1B 14 114 14 iy RRR KRADE 7 DY, ROR HIE ASHE seseeneneesensrceneee] 25, ig AD RINIIIR DEAE BE US WEE errno i 129 BRR 31 HUE Order PTEROSAURIA, 131 “121.” Suborder “RHAMPHORHYNCHOIDEA” 131 KEHRUERL Family Anurognathidae- BESCILI Genus Dendrorhynchoide 132 PAu] HEI, Genus Jeholopterus-- woe 13S MEAL Family Rhamphorhynchidae oe 13D SLBA Genus Angustinaripterus 0139 F"%HNR Genus Prerorhynchus: 141 LITRE AEIR, Genus Sericipterus eer 142 WHERILIR Genus Qinglongopterus wees 142 ARH RIERL Family Scaphognathidae ~~ v5 It EA, Genus Jianchangnathus 146SiGHROEAL? Family Scaphognathidae? 148 AI; HEHE NB, Genus Orientognathus -----------e- sees 14 MEF} Family Wukongopteridae-- anseeitSA 47 RAEI Genus Wukongopterus sees 1ST S78 MILI Genus Darwinopterus 153 ‘AS RE I, Genus Kunpengopterus 159 MEAL? Family Wukongopteridae? 163 KMRL EIR Genus Changchengopterus ---rsennm 163 ROGIER? Genus Jianchangopterus “164 “URUK JEIEA” PEASE “Rhamphorhynchoidea” incertae familiae oe 166 UROL, Genus Fenghuangopterus =~ 166 SER Genus Daohugoupterus HLA? Genus Archaeoistiodactyl 3 FEM] Suborder PTERODACTYLOIDEA 79 F-J2K} Superfamily Archacopterodactyloidea: ‘Hi KOEAL Family Ctenochasmatidae Hof WL Genus Huanheprerus- 47 WEIR Genus Eosipterus ALSER ICI Genus Beipiaopterus FE HRIEIR Genus Cathayopterus Het RLM Genus Gegepterus 598 HEM) Genus Elanodactylus WEARER Genus Prerofiltrus vsenesseserseenee] BG ROE Family Gallodactylidae--- 186, JR HN, Genus Gladocephaloideus 187 fis ROEM? Family Gallodactylidae? 89 EI Genus Feilongus 189 AbJ7 REF} Family Boreopteridae “191 ACA WHEN Genus Boreopterus- HEEKOLIR, Genus Zhenyuanopterus- FA JERI BERL AS familiae SF REIR Genus Moganopterus AGERE Superfamily Pteranodontoidea WERVER Family Anhangueridae Archacopterodactyloidea incertaeLF RIEIR Genus Liaoningopterus-s-sesensenee 199 WA ZEAL Family Istiodactylidae ~ WURIEIR Genus Istiodactylus--- 5 ARMS ASI AR, Genus Nurhachius ALPE WLI Genus Liaoxipterus- HSK Genus Longchengpierus £7 L)98 AER Genus Hongshanopterus ~~ FAROE PAGE Pteranodontoidea incertae familiae 209 FE RILIR Genus Nemicolopterus SILI, Genus Guidraco 89H |, Genus Hamipterus {PSR IR Genus Ikrandraco ~~ HAR EIE Genus Linlongopterus HER IERIA, Superfamily Azhdarchoidea-~ HERA JER Family Dsungaripteridae E85 4.3 Ia, Genus Dsungaripterus ~~ #8) Genus Noripterus APRA, Genus Lonchognathosaurus-~ TTPNSIERL Family Tapejaridae "PEI IG IRE Genus Sinopterus ~~ FRNA} 2 Family Tapejaridae? Sere RIK, Genus Eoazhdarcho WIPS EAE Family Chaoyangopteridae IPRA Genus Chaoyangopterus-~ HAIER Genus Shenzhoupterus HESIEAL Family Azhdarchidae-~ WRI Genus Zhejiangopterus ~ FEW AR AGE Pterodactyloidea incertae familiae ----242 ABECRIER Genus Haopterus ~ 2) Genus Kryptodrakon- SLELROEIG? Genus Yirianopterus?-- “9238 EIR? Genus Ningchengopterus?Bia SEP AER JEP ERBRS —, BFERBRELRARGAKK (—) BB RAAT STR, MEP AVA ALT RE, EMV SIE AE, AR AE, BRAS SARE RHE HAN | Airs. ATR RIL, A BERR EEE FEMIAS, MSIEBR EER A SLAF aE, LAR TE YS LAR AT ETE | IE FAWIE TARE, AVE AME BER (BEL) EASE AE PE, tht 45 7 ARDY Ae EAB AL /44_ (Subparviclass Archosauriformes) 1, 3JZELA (Magnorder * ve VF fs < we f rc) ang siperorder Averetataralia ERE MagoorderArchosauria oe Ad ParviclassArchosaurmorpha RAL FEE Two crown groups of Infraclass Diapsida PL BVA Ae 8d Se PL ME AAP ALAS CSL), SC FH i A ASHE A (4B Gauthier et al., 1988 1 Laurin, 1991 BE). RARArchosauria) +1{f28§ A (Superorder Pseudosuchia) 41, #iljf"$%!4 (Paracrocodylomorphs ) "PEA (Grandorder Poposauroidea) HUBS HLA EHR AYA FAME CLI 2) 6 #13 PEAT BIGOEAL (Order Protorosauria), JsiitrFt (Family Prolacertidae), 6} (Family Proterosuchidae) , 51834} (Family Erythrosuchidae) °, #i#F1 (Order Phytosauria) . ii ya83® (Family Euparkeriidae) , 2F 4185 (Family Gracilisuchidae) #JL4> As la] FR HSS HE Mat Parvielass Archosauromorpha_4:1.¥/)¥id Subparviclass Archosauriformes ALU /|it lw Magnorder Archosauria EAH Superorder Pseudosuchia fA Basal archosaromrphs Paracrosodylomorphs aerate anne 4 asl archosriforms Licatns "haa Pacudosuchians AEP EAR fer ee aa + Gaal - g os = 3 : es = = = He 5 . ashe ef - § & Z a te* 3,255 a4 ‘i ef" aa 3 baa 24 & + 3 3 : 2 gi id i pa 3 zs 5 2g z Suchians 2 eA KH, LRA EAE RAO, 04, Nesbitt, 2011, Lit al, 2012, ra Cth SBT AT REBATE (HLAE Sues, et al, 2014 Ail Li et al, 2016 246165) a ic 98 2003, Modesto et Su (2) FE SCATIE BR, METAS RA Bak LA IEA LER NE ay AH TE PY CE Ae FERENTIAL AAEM — PE RRA, Fy tL © SIA FF CErythrosuchidae) #2 ft 38 3183 (i (Exvuirosuchus) iM Att. BE AE A Me BH Erythrosuchus Al Erythrosuchidae (RUF 183 RAISING RHE AHHH» “Exytheo-” Ih LA". HE *S1A9" RM PME, Som PAST ARIELAL, RAOGUCLA AER, ARE “SRB ATE AE. ezFEBSED IGS. AR BET FE EET AMIE BNE AMA Vd Be SSFP IIE TBE BBE SEAT ETTE, MPR, A PIR PARE TINY Se HE ATA GE Fi LASTER ENS. TEAS , RNS +! (Archosauromorpha) {172 3 AURFHEE. Nesbitt (2011) 4zihi, iPANKFIE TIEAIZ (Archosauriformes) Me Az Ayal) FRR A, AM Ele TIME FEY AE — AG VISE (hn Li et al., 2012, Butler etal., 2014, Sookias et al. 2014), HUH Nesbitt AGMA T(E, AMES YAY ERI BSA RHEE CULE 2) NE SAIS AAPA TB A. = 2 F% WE \ ® (Subparviclass Archosauriformes) % * Gauthier % (1988) i SLAY Archosauriformes, (£5x ttf #3 Ft (Proterosuchidae), 3 [04+ (Erythrosuchidae), i ASIR (Litorosuchus) , IRSE85F+ (Euparkeriidae) | Hi dA (Phytosauria) #422 A (Archosauria) HUE ASE IAI ALSE SEP ETAT a. SR TE EEE: ok AA EL, POF ROE, WP Ae RT ak AE He (AS SUSE A BEASLEY LEZ we (A SET 3 BR He KFA) AUF A BRD. E2%E B (Magnorder Archosauria) “J Benton (1999) fty.%#5 3 (Avesuchia) , Archosauria ff] Cope 4 1869 4: AK EE FILES ASL ACHE AN JE Ld HL FB PONE ASITAT ai. SO APE ONY REE SEAT «PME AUCPAS98 CE AH TAY ERE (lagenar/cochlea process) PAC, FAR, SMR ZAIL ACT A, MEAG BAT LA ALAS (WERT ILID ER) AMER, CRRA ea SES — EE RASTER MEET CREME). BORA Re RETO OE Ke, IRE ELAS Yi BEAL EASES V5 mB Aa Te TT ST AAR BA Be / DEED Keadittiy 50° $3] 90° Fy Tha, {28542 B (Superorder Pseudosuchia) Ff 12 QRS AIHA TS RRR AS AEA EA. AGATE LEE RA: ARR. Ree a, AEE Se UR ES OD UIP Sg — HY Td Bh, AAAS AC HY Zitted 4E 1887-1890 46 ft XC HEH ir He, ME GLH Ae UT (BS AS HOY PA 7 A: J (aetosaurs) Fl Dyoplax, 7é 20 te2cdn+, (FSR TALL Ese HH ATL (Thecodontia Owen, 1859) ‘PIE (Suborder Pseudosuchia) , fj Zittel YEH ACW ZEM AT FL nel (Suborder Actosuria) , JE9>, MIAH NUAT HALE (Suborder Phytosauria) #ildi#5IE A (Suborder Proterosuchia) , 7 “HYG AS” MEAT Ay HOM WES Hh, (RSA HN Ea A EA — 4 “BE AOR”, TAG ACH AY HTS” ah ABB A TBS AS, (9) Ante Romer (1966) fy (Vertebrate Paleontology) —43+1, Gauthier #{] Padian (1985) J Gauthier (1986) HELAASE BARRETT GE TREE, [alIME Wi HE Ee Bf 5 — XH HER [Superorder SHEAR (synapomorphy) . [Alte (1) SEIT BAR AS wT REST AE sae