Professional Documents
Culture Documents
OF MALVAN
Kaustubh Warde & Trisha Gupta
Contents
3
Background
Trash fish, also known as low value bycatch, reduction fish or FMFO fish, refers
to fish that are used for the production of fishmeal and fish oil, used as high-
protein feed largely by the aquaculture and poultry industries[1]. These are
generally small-sized and/or damaged fish that captured as bycatch in
mechanized and non-selective fishing gear. Unchecked harvesting of these
species for reduction fisheries can put enormous pressure on marine habitats,
food webs and can endanger the sustainability of commercial fish populations[2].
This guide describes the common species sold as trash fish in Malvan,
Sindhudurg, Maharashtra, based on research[3] conducted in this region. Species
descriptions were obtained from sources such as FAO[4], FishBase[5], CCARI[6],
SeaLifeBase[7], and Finfishes of the Gulf of Mannar[8].
4
Fisheries Craft and Gear
A diverse range of fishing craft and gear operate out of Malvan. The main
fisheries contributing to the fishmeal industry are trawlers, gillnets and shore
seines.
5
Illustration reproduced from Sharks and Rays of Malvan [9]
Fisheries Craft and Gear
6
Illustration reproduced from Sharks and Rays of Malvan [9]
Fish Morphology and Technical Terms
7
Image adapted from FAO Species Identification Sheets [4]
Fish Morphology and Technical Terms
8
Image adapted from FAO Species Identification Sheets [4]
Trash Fish of Malvan: Common Families
At least 110 species across 50 different families were identified in the trash fish
in Malvan. This included fish (bony as well as cartilaginous) as well as
invertebrates such as squid, prawn, crab and cuttlefish. More than half of the
reduction fish species were small, pelagic and planktivorous (i.e., feeding on
plankton).
The most common trash fish species recorded in Malvan (52 species, belong to
29 families) are described in this book. A brief description of each family is
provided, followed by descriptions and photos of the trash fish species in the
family on individual pages. The common name of each species is provided, along
with the local Marathi name wherever possible.
This book is not intended to be a comprehensive guide to all trash fish species;
rather, it describes the common species found in this region.
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FAMILY: MURAENESOCIDAE
Characteristic Features:
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ORDER: ANGUILLIFORMES FAMILY: MURAENESOCIDAE
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FAMILY: SYNODONTIDAE
Characteristic Features:
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ORDER: AULOPIFORMES FAMILY: SYNODONTIDAE
• Body elongate and tubular. Mouth very oblique; snout short; eyes
small and set far forward near tip of upper jaw; a single row of
teeth on upper jaw visible even when mouth is closed.
• Anal fin rays 14 or 15; anal fin base much longer than dorsal fin
base; pelvic fin rays 8, inner rays much longer than outer rays.
• A series of longitudinal stripes, alternating blue and yellow, cover
back and sides; belly pale yellow or gold. A large, dark, oblique
spot at upper corner of gill cover. Distal parts of dorsal and caudal
fins dusky; other fins pale yellowish.
• Found on sandy bottom and feeds on small fishes
• Marketed fresh, but the damaged fish are sold as trash fish.
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FAMILY: CARANGIDAE
Characteristic Features:
• Body extremely variable in shape, ranging from elongated and fusiform to deep and strongly
compressed.
• Head varying from moderately long and rounded to short deep and very compressed.
• Snout pointed to blunt, lower jaw protruding to subtended, eye small to large, teeth in jaws in
rows or bands.
• Row of enlarged scutes present over the lateral line of the body.
• Anal fin with two anterior spines that are separated from rest of fin by the gap.
• Darker above (green or blue to blackish) and paler below (silvery to white or yellow-golden),
some species almost entirely silvery when alive, others with dark or coloured bars or stripes on
head, body or fins, some able to change patterns; young of many species with bars or spots.
• Largely continental distributed occurring from brackish environment to the oceanic waters.
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ORDER: CARANGIFORMES FAMILY: CARANGIDAE
• The adults are sold as fresh whereas the juveniles and the
damaged scads are sold as trash fish.
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ORDER: CARANGIFORMES FAMILY: CARANGIDAE
• The adults are sold fresh as well as dried salted whereas the
juveniles and damaged scads are sold as trash fish.
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ORDER: CARANGIFORMES FAMILY: CARANGIDAE
• Easily recognized by its long pelvic fin into a groove along the
midline of the belly.
Juvenile
• Strongly compressed body almost ovate in shape, the eyes
lack adipose eye lid.
• The adults are sold fresh whereas the juveniles and damaged
fishes are sold as trash fish.
Adult
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FAMILY: MENIDAE
Characteristic Features:
• Commonly called Moonfish.
• Body very deep, almost triangular, extremely compressed; breast sharp edged; mouth small,
almost vertical, protrusible; teeth limited to villiform bands in jaws.
• Dorsal fin single, long-based, and low, except for a short peak anteriorly; spines thin and flexible,
anal fin very long based, commencing at point of insertion of pelvic fins; 2 spines in juveniles, but
these and some leading soft rays become overgrown with skin with increasing age; free tips of
visible rays splayed out, caudal fin deeply forked.
• Pectoral fins shorter than head length, pelvic fins inserted slightly in front of pectoral fins, with 1
short spine followed by 5 soft rays, the first 2 soft rays fused and greatly elongated.
• Upper sides deep metallic blue, remainder silvery, with a row of round to ovoid, dark slaty-blue
spots above and below lateral line.
• Inhabits coastal waters near the bottom on both continental shelves and round major island
groups to depths of 200 m; seems to prefer deep water but occasionally enters river estuaries.
Schooling; feeds on small invertebrates.
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ORDER: CARANGIFORMES FAMILY: MENIDAE
• Mouth inferior
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ORDER: CLUPEIFORMES FAMILY: CLUPEIDAE
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ORDER: CLUPEIFORMES FAMILY: CLUPEIDAE
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ORDER: CLUPEIFORMES FAMILY: CLUPEIDAE
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ORDER: CLUPEIFORMES FAMILY: CLUPEIDAE
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ORDER: CLUPEIFORMES FAMILY: CLUPEIDAE
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ORDER: CLUPEIFORMES FAMILY: CLUPEIDAE
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ORDER: CLUPEIFORMES FAMILY: CLUPEIDAE
Common name: Indian illisha
Marathi name: Bhing/Kanta
Latin name: Illisha melastoma
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FAMILY: ENGRUALIDAE
Characteristic Features:
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ORDER: CLUPEIFORMES FAMILY: ENGRAULIDAE
Common name: Commerson’s anchovy
Marathi name: Salshin
Latin name: Stolepherous commersonnii
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ORDER: CLUPEIFORMES FAMILY: ENGRAULIDAE
• 23-26 keeled scutes on belly, inner part of anal fin deep yellow
with whitish margin.
• Presence of black spot on the upper part of the gill opening, as
well as spots on cheeks and paired fins.
• Found in Indian ocean, inshore waters, entering into the
estuaries.
• Feeds on small fishes and larvae of crustaceans and mollusc.
• The damaged as well as the juvenile malabar thryssa are sold
as trash fish.
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ORDER: CLUPEIFORMES FAMILY: ENGRAULIDAE
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ORDER: CLUPEIFORMES FAMILY: ENGRAULIDAE
• Body fusiform, compressed, belly with sharp keel scutes from isthmus
to anus, maxillia reaching to or almost to the base of pectoral fin, anal
fin long, tip of the snout level with eye.
• Back blue or green, flanks silvery, a black venulose area behind gill
opening, with a golden area before it.
• Dorsal fin lemon yellow with a dusky tip, caudal fin lemon yellow with
upper, lower and hind edges black, anal fin clear.
• Found in coast pelagic waters often enters mangroves and adjacent
brackish waters. Feeds on small organisms in coastal waters.
• They have little market value and therefore they are sold as trash fish.
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ORDER: CLUPEIFORMES FAMILY: ENGRAULIDAE
Common name: Dussumier's thryssa
Marathi name: Khavale/Salali
Latin name: Thryssa dussumieri
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ORDER: KURTIFORMIS FAMILY: APOGONIDAE
36
FAMILY: CYNOGLOSSIDAE
Characteristic Features:
37
ORDER: PLEURONECTIFORMES FAMILY: CYNOGLOSSIDAE
• Body flat and elongate, with dorsal and anal fins joined to caudal
fin. Eyes on left side of the body with no scale between them,
snout short and obtusely pointed.
• Light brown on eyed side with dark brown mottling forming
diffuse, irregular crossbands, dorsal and anal fins grey/black.
• Found in sandy and sandy-muddy waters of the continental shelf
down to 25m, also enter into estuaries.
• Feeds mostly on bottom living invertebrates, especially worms.
• Sold fresh as well dried in the market. Economically important fish.
• The damaged and juveniles tonguesole are sold as trash fish.
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FAMILY: PARALICHTHYIDAE
Characteristic Features:
41
FAMILY: DREPANIDAE
Characteristic Features:
42
ORDER: PERCIFORMES FAMILY: DREPANIDAE
Common name: Spotted sickle fish
Marathi name: -
Latin name: Drepane punctata
44
ORDER: PERCIFORMES FAMILY: HAEMULIDAE
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ORDER: PERCIFORMES FAMILY: HAEMULIDAE
46
FAMILY: LEIOGNATHIDAE
Characteristic Features:
48
ORDER: PERCIFORMES FAMILY: LEIOGNATHIDAE
Common name: Orangefin ponyfish
Marathi name: Kap
Latin name: Photopectoralis bindus
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ORDER: PERCIFORMES FAMILY: LEIOGNATHIDAE
Common name: Splendid ponyfish
Marathi name: Kengi
Latin name: Eubleekeria splendens
50
ORDER: PERCIFORMES FAMILY: LEIOGNATHIDAE
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FAMILY: SILLAGINIDAE
Characteristic Features:
• Moderate-sized, elongate perciform fishes. Opercle with small sharp spine; lower part
of preopercle horizontal.
• Snout long and conical. Mouth small, terminal; end of upper jaw sliding below
preorbital bone, teeth villiform, in broad bands.
• Two separate dorsal fins, the first with 9 to 12 slender spines; the second with 1 spine
and 16 to 27 soft rays; anal fin long, with 2 weak spines and 14 to 27 soft rays.
• Scales small, ctenoid; lateral line slightly arched. Swimbladder frequently complex.
• Body silvery to sandy grey or green, sometimes with black spots on body and pectoral-
fin base.
• Inshore, bottom-dwelling, schooling fishes with a few species found in deeper water to
about 180m
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ORDER: PERCIFORMES FAMILY: SILLAGINIDAE
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FAMILY: TERAPONIDAE
Characteristic Features:
54
ORDER: PERCIFORMES FAMILY: TERAPONTIDAE
Common name: Jarbua tarpon/Crescent grunter
Marathi name: Karkanda
Latin name: Terapon jarbua
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ORDER: PERCIFORMES FAMILY: TERAPONTIDAE
Common name: Small scaled terapon
Marathi name: Bani/Karkanda
Latin name: Terapon puta
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FAMILY: TRICHIURIDAE
Characteristic Features:
• Commonly called Ribbonfish
• Body extremely elongate, compressed and ribbon-like, large mouth which is not protusible, lower
jaw projecting, jaws with string canine teeth, and those in the front of upper jaw are fang-like.
• Dorsal fin low and long and begins shortly behind eye, the anterior spinous part is shorter than the
posterior soft portion and these two parts are separated by a distinct notch in somes species; anal
fin low or reduced to short spinules; pectoral fins are short and low on the body; pelvic fins reduced
to a scale-like spine or completely absent in some species.
• Caudal fin either small and forked or absent, the body tapering to a point.
• Lateral line single. Scales absent, body generally silvery, a little darker on back; usually no distinct
marks or blotches; dorsal and anal fins sometimes tinged with pale yellow; pectoral fins
semitransparent.
• Voracious predators distributed in tropical and temperate seas and generally inhabit deeper waters
over the continental shelf and the slope, some species are common in shallow coastal waters.
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ORDER: PERCIFORMES FAMILY: TRICHIURIDAE
Common name: Largehead hairtail
Marathi name: Waakti/Baga
Latin name:Trichuris lepturus
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FAMILY: PLATYCEPHALIDAE
Characteristic Features:
• Elongate fishes with head moderately to strongly depressed. Mouth large; lower jaw
longer than upper; fine teeth on jaws, canines present in a few species.
• Two dorsal fins, well separated, pelvic fins thoracic in position, set far apart toward
sides of body.
• Lateral line complete, scales ctenoid, usually embedded on head and breast.
• Dark above, pale below; the dark colours with various shades of brown, grey or
black; brighter hues of reddish, purplish or greenish in some.
• Benthic fishes found on mud or sand bottoms of shelf areas at depths of about 75m
or less; a second group of species are associated with rocky shores or coral reefs.
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ORDER: SCORPAENIFORMES FAMILY: PLATYCEPHALIDAE
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FAMILY: SCORPAENIDAE
Characteristic Features:
• Moderately compressed to robust fishes, usually bass like in appearance, with large spiny heads.
Mouth moderate to large, terminal, oblique, protractile; teeth usually villiform arranged in bands.
• Eyes moderate to large; preopercular margin with 3 to 5 spines, the uppermost 3 better developed;
opercle with 2 divergent spines or a single spine; other spines scattered on head.
• A single dorsal fin, usually notched at posterior end of spinous part, with 8 to 18 spines and 4 to 14
segmented rays; anal fin with 2 to 4 spines and 5 to 14 segmented rays; posterior most segmented
rays of dorsal and anal fins split to base in most species, appearing as two rays but counted as one;
pectoral fin broad-based, large, fanlike, with 11 to 23 rays; pelvic fins thoracic in position, with 1
spine and 3 to 5 branched or simple rays.
• Caudal fin rounded to square-cut, never forked. Venom glands associated with fin spines.
• Inshore scorpionfishes are mostly brown or variously mottled and barred with dark pigment on a
lighter background, often with a pale or reddish belly. Those from deeper water are mostly red, often
with spots of darker red, brown or black and with white skin filaments.
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ORDER: SCORPAENIFORMES FAMILY: SCORPAENIDAE
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FAMILY: ARIIDAE
Characteristic Features:
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ORDER: SILURIFORMES FAMILY: ARIIDAE
• Pectoral spine longer than the dorsal spine. First dorsal fin ray
prolonged into a long filament
• The median longitudinal groove becomes narrow and deep posteriorly
and reaches nearly to the base of the occipital process.
• Palate teeth globular, in a single large irregularly ovate patch, not fully
formed, with loosely packed teeth, inner margins of patches not
straight.
• Body greyish-silver, becoming white on the sides and below.
• Found in marine coastal and estuarine waters on the muddy or sandy
substratum. Feed on small invertebrates and small fishes.
• They are sold fresh but the damaged and juveniles are sold as trash
fish.
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FAMILY: PLOTOSIDAE
Characteristic Features:
66
ORDER: SILURIFORMES FAMILY: PLOTOSIDAE
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ORDER: TETRADONTIFORMES FAMILY: BALISTIDAE
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ORDER: TETRADONTIFORMES FAMILY: OSTRACIIDAE
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ORDER: TETRADONTIFORMES FAMILY: TETRAODONTIDAE
Common name: Milk-spotted puffer
Marathi name: Kand/English gaend
Latin name: Chelonodon patoca
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FAMILY: TRIACANTHIDAE
Characteristic Features:
• Deep compressed bodies; skin moderately thick with numerous scales not individually easily
discernible to the unaided eye, each scale bearing upright spinules and having a rough,
shagreen-like appearance.
• Mouth small and usually terminal and molariform teeth.
• Dorsal fin has 6 spines, dorsal fin rays 20 to 26, anal fin rays 13 to 22; caudal fin deeply forked
and caudal peduncle distinctly tapered; pelvic fin represented by a large spine which is movably
articulated with anterior end of pelvis and capable of being locked erect by a flange on they
pelvis; most dorsal, anal and pectoral fin rays branched.
• Lateral line inconspicuous. Generally silvery, with upper half of body dusky, with or without
darker blotches.
• Tripod fishes are benthic, occurring usually on flat, sandy or weed-covered bottoms. They feed
on bottom living invertebrates.
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ORDER: TETRADONTIFORMES FAMILY: TRIACANTHIDAE
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FAMILY: MATUTIDAE
Characteristic Features:
• Front narrow, antennae rudimentary, chelipes subequal, walking legs natatorial, P2-
• Male abdomen with 5 segments, female opening is called sternal and male coxal.
• G1 elongated and columnar, distal tip rounded whereas G2 is long and narrow,
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ORDER: DECAPODA FAMILY: MATUTIDAE
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FAMILY: PORTUNIDAE
Characteristic Features:
• Carapace usually broader than long and broadest between last pair of antero-lateral
teeth, transversely hexagonal to transversely ovate.
• Regions of the carapace not well defined, antero-lateral teeth varying from 5-9.
• Antennules and antennae short, orbits broad, eyes conspicuous, buccal frame
quadrate, carpus of third maxillipeds articulated at inner dorsal margin of merus.
• Dactylus of P5 suboval or lanceolate, rarely styliform and always dorso-ventrally
flattened.
• P5 generally modified for swimming with at least propodus and dactylus paddle-like.
• Male abdomen with segments 3-5 fused to varying degrees, male openings coxal,
female openings sternal (vulvae).
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ORDER: DECAPODA FAMILY: PORTUNIDAE
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FAMILY: CALAPPIDAE
Characteristic Features:
• Carapace more or less circular or transversely ovate. Often a postero-lateral vault-like expansion
over the ambulatory legs.
• Antennules folding obliquely or nearly vertically. Antennae generally small. Third maxilliped not
elongate or acute, not completely closing the buccal cavern and not concealing the palp in
repose; exopodite broad.
• Afferent branchial openings having the normal position in front of the bases of third maxilliped.
Efferent branchial channels together forming a deep channel in the endostome, covered by a
pair of lamellar process from the first maxillipeds.
• Chelipeds practically symmetrical in size but characterized by a strong heterodonty; chela of a
great size and so curved as to shut closely against the pterygostomial region.
• Male abdomen with some segments fused. Female openings sternal. Male
openings coxal. G2 long and narrow, exceeding the length of G1
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ORDER: DECAPODA FAMILY: CALAPPIDAE
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FAMILY: SQUILLIDAE
Characteristic Features:
85
ORDER: STOMATOPODA FAMILY: SQULLIDAE
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FAMILY: LOLIGINIDAE
Characteristic Features:
• Adults have tentacular club with four rows of suckers on the manus. Arm suckers in
two rows. The body is fusiform of variable shape.
• The fins are terminal-lateral and united posteriorly. The buccal membrane has seven
lappets, usually beset with small suckers. Photophores occur rarely.
• The body form of hatchlings is always bullet shaped with well developed
terminal fins. Tentacles and arms are well developed; the fourth arm is larger than the
first.
• Generally, no great changes in body form from hatchling to adult. Young forms of the
Subgenera Loligo and Alloteuthis are very similar and therefore difficult to distinguish
at the species level.
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ORDER: TEUTHIDA FAMILY: LOLIGINIDAE
88
FAMILY: SEPIIDAE
Characteristic Features:
• The eyes are of myopside type. The arms are short and the rather long tentacles are
retractable: they can be withdrawn into the head cavities.
• The 4th pair of arms is longer than the others and the left one is hectocotylized .
The mantle is oval, its width is about half of its length and it is not fused with the head,
which is much narrower.
• The body is flattened dorso-ventrally, the fins run like two narrow ribbons along the
lateral side of the mantle.
• The eggs of Sepia look like blue grapes. The internal shell, made of calcareous plate, is
present and is situated under the skin of the dorsum.
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ORDER: SEPIIDA FAMILY: SEPIIDAE
90
References
[1] Funge-Smith, S., Lindebo, E., & Staples, D. (2005). Asian fisheries today: The production
and use of low value / trash fish from marine fisheries in the Asia-Pacific region. Rap
Publication 2005/16, 47.
[2] Lobo, A. S., Balmford, A., Arthur, R., & Manica, A. (2010). Commercializing bycatch can
push a fishery beyond economic extinction. Conservation Letters, 3(4), 277–285.
[4] FAO Species identification sheets for fishery purposes: Western Indian Ocean. Fishing
Area 51. http://www.fao.org/3/ad468e/ad468e00.htm
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Photo Credits:
The following images have been used under the Creative Commons license or with
permission from the owner:
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This guide has been created by Kaustubh Warde and Trisha Gupta as part
of research work on reduction fisheries conducted by Dakshin Foundation.
We acknowledge Chetan Rao, Murali M., Kartik Shanker and Naveen
Namboothri for their guidance on this study, and thank the Duleep Matthai
Nature Conservation Trust for supporting our work.
This work is free to copy, distribute and use for non-commercial purposes,
along with attribution to the original authors.
Contact:
www.dakshin.org
dakshinfoundation.india@gmail.com
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