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Read Online Textbook Travis Mia Sheridan Ebook All Chapter PDF
Read Online Textbook Travis Mia Sheridan Ebook All Chapter PDF
Internal Anatomy.
There are no capillaries, but the blood is delivered into the tissues
and finds its way, by irregular spaces or “lacunae,” into certain main
venous channels or “sinuses.” There are three such in the
cephalothorax, one median and the others lateral, considerably
dilated in front, in the region of the eyes, and connected by
transverse passages. By these the blood is brought back through the
pedicle to the lung-books. In the abdomen also there are three main
sinuses, two parallel to one another near the lower surface, and one
beneath the pericardium. These likewise bring the blood to the lung-
books, whence it is conducted finally by pulmonary veins (Fig. 186)
back to the pericardial chamber, and thus, by the ostia, to the heart.
The Spider’s blood is colourless, and the majority of the corpuscles
are “amoeboid,” or capable of changing their shape.
Generative System.—The internal generative organs present no
great complexity, consisting, in the male, of a pair of testes lying
beneath the liver, and connected by convoluted tubes, the “vasa
deferentia,” with a simple aperture under the abdomen, between the
anterior stigmata.
The ovaries are hollow sacs with short oviducts which presently
dilate to form chambers called “spermathecae,” which open to the
exterior by distinct ducts, thus forming a double orifice, fortified by
an external structure already alluded to as the “epigyne.” The eggs
project from the outer surface of the ovary like beads, connected with
the gland by narrow stalks, and it was not at first clear how they
found their way into the interior cavity, but it has been ascertained
that, when ripe, they pass through these stalks, the empty capsules
never presenting any external rupture.
The palpal organs have already been described. The spermatozoa,
when received by them, are not perfectly elaborated, but are
contained in little globular packets known as “spermatophores.”
Nervous System.—The Spider’s central nervous system is
entirely concentrated in the cephalothorax, near its floor, and
presents the appearance of a single mass, penetrated by the
oesophagus. It may, however, be divided into a pre-oesophageal
portion or brain, and a post-oesophageal or thoracic portion.
The brain supplies nerves to the eyes and chelicerae, while from
the thoracic mass nerves proceed to the other appendages, and
through the pedicle to the abdomen. The walls of the oesophagus are
closely invested on all sides by the nerve-sheath or neurilemma.
Sense-Organs.—Spiders possess the senses of sight, smell, and
touch. Whether or not they have a true auditory sense is still a matter
of doubt. Since sounds are conveyed by vibrations of the air, it is
never very easy to determine whether responses to sounds produced
near the animal experimented upon are proofs of the existence of an
auditory organ, or whether they are only perceived through the
ordinary channels of touch. In any case, the organs of hearing and of
smell have not yet been located in the Spider. M‘Cook considers
various hairs scattered over the body of the spider to be olfactory, but
from Gaskell’s researches upon allied Arachnid groups it would seem
that the true smelling organ is to be sought for in the rostrum.
Eyes.—Spiders possess from two to eight simple eyes, the external
appearance and arrangement of which have already been briefly
explained. They are sessile and immovable, though often so placed as
to command a view in several directions. In structure they are
essentially like the ocelli of Insects. Externally there is a lens,
succeeded by a mass of transparent cells, behind which is a layer of
pigment. Then come the rods and cones of the retina, to which the
optic nerve is distributed. A comparison of this with the arrangement
in the Vertebrate eye will show a reversal of the positions of the
retina and the pigment-layer. The lens is part of the outside covering
of the animal, and is cast at the time of moulting, when the spider is
temporarily blind. It is stated, however, that the eyes do not all moult
simultaneously. There is often a considerable difference between the
various eyes of the same spider, especially with regard to the
convexity of the lens and the number of rods and cones.
Though most spiders possess eight eyes, the number is sometimes
smaller, and in some groups of eight-eyed spiders two of the eyes are
sometimes so reduced and degenerate as to be practically
rudimentary. As might be expected, Cave-spiders (e.g. Anthrobia
mammouthia) may be entirely sightless.
Touch.—The sense of touch would appear to be extremely well
developed in some spiders, and there is reason for believing that the
Orb-weavers, at all events, depend far more upon it than upon that of
sight.
Among the hairs which are distributed over the spider’s body and
limbs, several different forms may be distinguished, and some of
them are undoubtedly very delicate sense-organs of probably tactile
function.
Spinning Glands.—Spiders vary greatly in their spinning
powers. Some only use their silk for spinning a cocoon to protect
their eggs, while others employ it to make snares and retreats, to
bind up their prey, and to anchor themselves to spots to which they
may wish to return, and whence they “drag at each remove a
lengthening chain.”
All these functions are performed by the silk-glands of the Orb-
weavers, and hence it is with them that the organs have attained
their greatest perfection. We may conveniently take the case of the
common large Garden-spider, Epeira diademata. The glands occupy
the entire floor of the abdomen. They have been very thoroughly
investigated by Apstein,[266] and may be divided into five kinds.
On either side of the abdomen
there are two large “ampullaceal”
glands debouching on “spigots,”
one on the anterior, and one on
the middle spinneret; there are
three large “aggregate” glands
which all terminate on spigots on
the posterior spinneret; and three
“tubuliform” glands, two of which
have their orifices on the
posterior, and one on the middle
spinneret. Thus, in the entire
abdomen there are sixteen large
glands, terminating in the large
fusulae known as spigots. In
addition to this there are about
200 “piriform” glands whose
openings are on the short conical
fusulae of the posterior and
anterior spinnerets, and about Fig. 187.—Spinning glands. A,
400 “aciniform” glands which Aciniform; B, tubuliform; C, piriform
debouch, by cylindrical fusulae, gland.
on the middle and posterior
spinnerets. Thus there are, in all,
about 600 glands with their separate fusulae in the case of Epeira
diademata.
The great number of orifices from which silk may be emitted has
given rise to the widespread belief that, fine as the Spider’s line is, it
is woven of hundreds of strands. This is an entire misconception, as
we shall have occasion to show when we deal with the various
spinning operations.
A few families are, as has already been stated, characterised by the
possession of an extra spinning organ, the cribellum, and the orifices
on this sieve-like plate lead to a large number of small glands, the
“cribellum glands.”
Respiratory Organs.—Spiders possess two kinds of breathing
organs, very different in form, though essentially much alike. They
are called respectively “lung-books” and “tracheae.” The
Theraphosae (and Hypochilus) have four lung-books, while all other
spiders, except Nops, have two. Tracheae appear to be present
almost universally, but they have not been found in the Pholcidae.
The pulmonary stigmata lead into chambers which extend
forwards, and which are practically filled with horizontal shelves, so
to speak, attached at the front and sides, but having their posterior
edges free. These shelves are the leaves of the lung-book. Each leaf is
hollow, and its cavity is continuous, anteriorly and laterally, with the
blood-sinus into which the blood from the various parts of the
Spider’s body is poured.
The minute structure of the leaf is curious. Its under surface is
covered with smooth chitin, but from its upper surface rise vast
numbers of minute chitinous points whose summits are connected to
form a kind of trellis-work. The roof and floor of the flattened
chamber within are connected at intervals by columns. The
pulmonary chamber usually contains from fifteen to twenty of these
leaves, and the two chambers are always connected internally
between the stigmata.
The tracheae are either two or four (Dysderidae, Oonopidae,
Filistatidae) in number, and their stigmata may be separate or fused
in the middle line. Each consists of a large trunk, projecting
forwards, and giving off tufts of small tubes which lose themselves
among the organs of the abdomen, but do not ramify. In the tracheae
of Argyroneta[267] a lateral tuft is given off immediately after leaving
the stigma, and another tuft proceeds from the anterior end.
Histologically the main trunk of the trachea is precisely like the
general chamber of the pulmonary sac, and differs greatly from the
trachea of an insect.
Cephalothoracic Glands.—In addition to the generative glands
and the so-called “liver” which occupy so large a portion of the
abdomen, there are, in Spiders, certain glandular organs situated in
the cephalothorax which call for some notice. These are the coxal
glands and the poison-glands.
The COXAL GLANDS are two elongated brownish-yellow bodies,
situated beneath the lateral diverticula of the stomach, and between
it and the endosternite. They present four slight protuberances which
project a short distance into the coxae of the legs. The glands appear
to be ductless, but their function is thought to be excretory. They
were first observed in the Theraphosae.
All Spiders possess a pair of POISON-GLANDS, connected by a narrow
duct with a small opening near the extremity of the fang of the
chelicerae. The glands are sac-like bodies, usually situated in the
cephalothorax, but sometimes partially (Clubiona) or even entirely
(Mygale) in the patura, or basal joints of the chelicerae. Each sac has
a thin outer layer of spirally-arranged muscular and connective
tissue fibres, and a deep inner epithelial layer of glandular cells. The
cavity of the gland acts as a reservoir for the fluid it secretes. The
virulence of the poison secreted by these glands has been the subject
of much discussion, and the most diverse opinions have been held
with regard to it. The matter is again referred to on p. 360.
CHAPTER XIV
ARACHNIDA EMBOLOBRANCHIATA
(CONTINUED)—ARANEAE (CONTINUED)
HABITS—ECDYSIS—TREATMENT OF YOUNG—MIGRATION—
WEBS—NESTS—EGG-COCOONS—POISON—FERTILITY—
ENEMIES—PROTECTIVE COLORATION—MIMICRY—SENSES
—INTELLIGENCE—MATING HABITS—FOSSIL SPIDERS