of the interactions between species in communities on many spatial and temporal scales, including the distribution, structure, abundance, demography, and interactions between coexisting populations. Chapter 17 Community Structure • A community has attributes that differ from those of its components • Number of species • Relative abundance of species • Nature of species interactions • Physical structure 17.1 The Number of Species and Their Relative Abundance Define Diversity • Species richness (S) is the count of the number of species occurring within the community • Relative abundance represents the percentage each species contributes to the total number of individuals of all species Species Rank - Abundance Curves 17.1 The Number of Species and Their Relative Abundance Define Diversity • Diversity indexes provide a way to quantify the relationship between species number and relative abundance • Simpson's index (D) = (ni/N)2 • = summation for all species • ni = number of individuals of species i • N = total number of individuals of all species • D ranges between 0 and 1 and as both species' richness and evenness increase, the value approaches 0 17.1 The Number of Species and Their Relative Abundance Define Diversity • Simpson's index (D) = (ni/N)2 • Simpson's reciprocal index or Simpson's diversity index = 1/D • The lowest possible value is 1, representing a community containing only one species • The maximum value is the number of species in the community (species richness: S) • Maximum value with perfect even-ness of relative abundance 17.1 The Number of Species and Their Relative Abundance Define Diversity • Shannon (or Shannon-Weiner) index = H = -(pi)(loge pi) • Relative abundance of each species = p i = ni / N • pi = proportion of species i • In the absence of diversity, where only one species is present, H = 0 • Hmax = ln S, occurs when all species are present in equal numbers Not all species within a community are equivalent