Featuring the HUMAN EVOLUTION Repoted by Group 1 AUSTRALOPITHECUS THE FIRST DEFINITE HOMINID
Australopithecus was originally defined by Raymond Dart in
1925 on the basis of a small child’s skull from Taung, South Africa, that was 2-3 million years old. This fossil had an ape- sized brain but a short face with smaller front teeth, including a small, human-like canine tooth much smaller than the projecting ones of apes, and larger back teeth (molars) than those of apes. Perhaps most importantly, the place where the spinal cord exited the skull was positioned underneath it, showing that these creatures stood fully upright with their head positioned over their vertebral column. Dart named this fossil species Australopithecus africanus (Dart 1925). Dartdescribed Australopithecus as “intermediate between living anthropoids and man” (1925: 574), but no formal definition of the genus was given beyond that. Nearly a century later, we understand australopiths to be intermediate in time and in anatomy between earlier hominins (Ardipithecus, Orrorin, Sahelanthropus; See Su’s article)and Homo, and the genus Australopithecus is defined not only by similarities among its species, but also by differences from the groups which pre- and post-date it. Strictly speaking, Australopithecus is probably not a true genus, because some species may be more closely related to humans than others, and anthropologists are not certain about the precise relationships among them. Australopiths were terrestrial bipedal ape-like animals that had large chewing teeth with thick enamel caps, but whose brains were only very slightly larger than those of great apes. They are the closest known relatives of our genus Homo, and we most likely evolved from a species that was part of this adaptive radiation. hominin species that span the time period from 4.18 to about 2 million years ago. WHAT IS AUSTRALOPITHECUS ? AUSTRALOPITHECUS BODY FEATURES Australopiths were fully upright bipeds whose skeletons display evidence of a history of selection for travelling bipedally on the ground, and that had lost features seen in most primates that would have made them good tree- climbers, such as a grasping foot. evidence for upright posture include an upright position of the skull and a spine with curvatures allowing vertical posture, a short, broad pelvis providing effective leverage for propulsion and balance over the two lower limbs, a femoral carrying angle and a tibia oriented orthogonally to the ankle joint, which together position the feet directly under the knees as in humans today, and stiff feet with longitudinal and transverse arches that lacked opposable big toes. There are also footprints of Australopithecus at Laetoli, Tanzania, 3.66 Ma that clearly indicate bipedal locomotion. Australopiths were roughly 1.2-1.5 m tall and probably weighed about 30- 50 kg Males were almost twice the size of females, a level of difference, or sexual dimorphism, greater than modern chimpanzees or humans but less than gorillas or orangutans .Their cranial capacity was 420-550 cc3, making their brains slightly larger for their body size than are those of modern apes. Australopithecus species lack canine tooth size sexual dimorphism, and have canines much reduced in size compared with extant apes, only very slightly larger than those of females. This indicates that males were not using their teeth to bite each other during fighting for access to mates. However, because males were so much larger than females, they still probably competed heavily for access to females, which possibly signifies a novel means of male-male competition in these hominins. Australopith front teeth are smaller than those of extant apes, but the premolars and molars are expanded and thickly enameled The jaws are robust, as is cranial evidence of the chewing apparatus, namely large cheekbones and generally well developed crests for attachment of neck and chewing muscles. The upper face is fairly vertical, but the lower face projects forward more than in humans (Figure 3). The face and dental anatomy suggests that australopiths were adapted to eating tough, hard-to-process foods such as tubers, nuts, seeds or roots, at least during times of food scarcity. Studies of isotopes within teeth, microscopic enamel wear, and of plant particles recovered from dental plaque all seem to confirm a tough diet AUSTRALOPITHECUS ARTIFACTS FOUND… Rareartifacts attributed to australopiths suggest that there is more to their behavior than what anatomical clues indicate. For quite some time paleoanthropologists have known about bone digging and probing tools from australopith sites in South Africa, but it is now apparent that at least some australopiths also made and used stone tools. Cutmarks have been found on animal bones at the 3.4 Ma site of Dikika, Ethiopia, which is a site where A. afarensis is known .Furthermore, a novel stone tool industry predating even the Oldowan has been discovered at Lomekwi, Kenya, a site dated to 3.3 Ma .The makers of these tools remains unknown. Kenyanthropus platyops is known from other localities at Lomekwi, as are a number of fossils not attributed to any taxon. No hominins are definitively associated with thee tools, but the Lomekwian archeology demonstrate that australopiths were making and using stone tools in the mid-Pliocene, and probably consuming meat when available. The Taung skull was the first evidence showing that walking upright on two feet and a different diet (as inferred from the teeth) were the adaptations that initially set hominins apart from apes, and that these changes long preceded great expansion of the brain and the many complex behaviors that accompany it. Since the discovery of the Taung specimen, many hundreds of specimens from roughly eight species of Australopithecus have been discovered in South Africa (A. africanus, A. sediba), eastern Africa (Ethiopia, Kenya, Tanzania; A. anamensis, A. afarensis, A. deyiremeda, Kenyanthropus platyops) and central Africa (Chad; A. bahrelghazali). The oldest and most primitive australopiths are found in eastern Africa, particularly Ethiopia and Kenya, with more derived australopiths appearing later in South Africa. South African sites yield some very well-preserved fossils and associated skeletons, but the complexities of the formation of the cave assemblages in which they are found can lead to uncertainty in dating. However, it appears that Australopithecus spans more than 2 million years of time and occupied a variety of habitats. HOMO HABILIS One of the earliest members of the genus homo one HOMO HABILIS of the earliest members of the genus Homo, has a slightly larger braincase and smaller face and teeth than in Australopithecus or older hominin species. But it still retains some ape-like features, including long arms and a moderately- prognathic face. Its name, which means ‘handy man’, was given in 1964 because this species was thought to represent the first maker of stone tools. Currently, the oldest stone tools are dated slightly older than the oldest evidence of the genus Homo. Itwas discovered in 1960. The history of discovery was led by scientists Louis and Mary Leakey uncovered the fossilized remains of a unique early human between 1960 and 1963 at Olduvai Gorge in Tanzania. The type speciman, OH 7, was found by Jonathan Leakey, so was nicknamed "Jonny's child". Because this early human had a combination of features different from those seen in Australopithecus, Louis Leakey, South African scientist Philip Tobias, and British scientist John Napier declared these fossils a new species, and called them Homo habilis (meaning 'handy man'), because they suspected that it was this slightly larger-brained early human that made the thousands of stone tools also found at Olduvai Gorge. Homo habilis height average is at 3ft – 4ft 5 in cm its 100-135. And its average weigth is at 70 lbs or 32 kg. Early Homo had smaller teeth than Australopithecus, but their tooth enamel was still thick and their jaws were still strong, indicating their teeth were still adapted chewing some hard foods (possibly only seasonally when their preferred foods became less available). Dental microwear studies suggest that the diet of H. habilis was flexible and versatile and that they were capable of eating a broad range of foods, including some tougher foods like leaves, woody plants, and some animal tissues, but that they did not routinely consume or specialize in eating hard foods like brittle nuts or seeds, dried meat, or very hard tubers. Another line of evidence for the diet of H. habilis comes from some of the earliest cut- and percussion-marked bones, found back to 2.6 million years ago. Scientists usually associate these traces of butchery of large animals, direct evidence of meat and marrow eating, with the earliest appearance of the genus Homo, including H. habilis. Many scientists think early Homo, including H. habilis, made and used the first stone tools found in the archaeological record—these also date back to about 2.6 million years ago; however, this hypothesis is difficult to test because several other species of early human lived at the same time, and in the same geographic area, as where traces of the earliest tool use have been found. EXAMPLES OF HOMO’S SKULLS KNM-ER 1813 AGE: ABOUT 1.9 MILLION YEARS OLD SPECIES: HOMO HABILIS OH 24 AGE: ABOUT 1.8 MILLION YEARS OLDSPECIES: HOMO HABILIS OH 8 AGE: ABOUT 1.8 MILLION YEARS OLDSPECIES: HOMO HABILIS THANK YOU !!! REFERENCES • https://www.nature.com/scitable/knowledge/library/austr alopithecus-and-kin-145077614/ • http://humanorigins.si.edu/evidence/human- fossils/species/homo-habilis