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a8lelrte 66te umll 414 f,asto?ll ees {t8?Psl1sa 18t$f n la ropsu$rprssrs gi *urt6r'94 ailsl gf grsttmgt'rg* nolltll.Xl eststrllrt't el0ffirluL lrl ib sepmrs oe tnregtstr*gs$ 'ldprurrl sggslrts*i tlf'o I}|sA r pleatsr;latesa lul neccslt$ tgopsr*url B)aqturs asEsutA stlgrrloslt' Srssscolo sgsl,satlec care partlnb )otw$Erattre $o d* a.l,t.Ssrt asadarlgc sqn oo flttrer I lrlrlmn tratoasltEtEa rogplratle*lreduo$ud eEusmul r.b 6luolJor I$ ts$wnoa sondllll ,mr6rrr1l !.n*E is faee dP iilrrLaltmo F6&P* J la ofurtpea aldonrlulr loDalilt a$ o r*eslvE suffut.grta dP ilrbstante nrt*ttltrsrillnpotrlre'r E)6satr$a rlditsst6 etiDnloaE& btrdroltsa mldm$lut et t'ntaf${.cl &e la otanrnaroa insL blss pa$i rll,a ressrne' r.;llratisreoo*lo r nrtrltlwptuelusiv rr lntr{ I <b eutststa srbs@nto prot91c6.In oooet eecpnugunlt pLsr reiluo|troaal* cm o reretr irrmftslar ln fEca illvlsludr

nutrlttv(iifuaamv -rl(1?99)r

La unsLe cEpurt ropmmnrl eote laeeltt ssu Dreootiat do re&rdi* rfllbsl&rf dtu ,. aontlsru.rlut 5,n ry$.r1c plsrrb rts pltm$ riafunpa ohl,nleorln f,elul E* Itoplam[ri]scL ape Sn e{$seeu 106 sostlll's Irta q@tatollnmuf, iet lasetlspgts r*tncul fll I @ rep{Rlc (riie,bktlo!, Bt celabr(196$}r i?tvitoc sluloXa trro trl Etlr

la ef,r&Elo aase &ltrsln[ ropcusul I'a plaatr6{bfftr! ISJ|.&Itrg ;:.strtJltl( f g64)91 colabr ) loonefil,srs sH *optttsul srtr I9tZ)en tenrlolpni. lteml.nst ersqluetv & fetoll loutonml l:';l,ebs( eatosal catltrnlnnl cE *opmrm$ deplaile estrustrv ila fstolll aln Eo1 otar$erost8'rlln ultsrlo*e rldltaloarteEnatattaarhsra lnc u arEtst eS EtErso do ro3rzue osts o xamrgt$o ilobtildlt& fi fix*t6 s po&to nodlflqn Nilb I'h rrdltf,! gl eil sa catt rel,attY$.futnnlt lrtrta f*totllo! ds esdluf btooblal'ae S'n or6#ole plaatol,oltl'n aolnnulnr L:odtfla[rtle

*69lll|'ate !.tstg f* dt norodr,B 1165,71*a;n8lirchLtltll,gl fL solabr(lgEnhotsf cS tra ooatsacEas miibrytl so trdnsfoilrl[ 5a turDul to* vogatalcllar Farlto1$la(tFf)r r ob*rr' srtr)c ulalr*ul, ln aab6r, la rbJa{6rp[ e.taft

11 Oi. Xerstl fn ulolurl n trrrdormma

rslp!,ratlrr i,ia:dew(l$a)-qrnu li.shlrilol Ia cri ,riwltc tl )fauf964hr ad,ttt oa lcrna osruta r6t| E slnA.trt slrtn l9(l :8s*d lc or6aaele in repeucrun aoltlrrut f.rts nldr rtB iglo8&tlrll* lcr eu llFi.gt sil alc tn sp{ libcr6r d.la smrf, aS ta ci.toptcs$ cololsl,l. &vin na!. iildm'foblr r e'lcmalcaa ar lobagigeeh 3n f, qlto$sw slaforeseg pornoebLlttetrl l* n sueulul r.suo-

laoslubltro rcw$o lrratf aiorlnr Gsl vfaooase al proplraoa omotla! 1l o*e8tarittn , llF{ *earta in reparer

aohl,nh*L de elbstasta sr loihrso ql aolub*

,Iitaroa oltoplamsd Sa tl.llpsf Benloadol il* r,rpmro a foEt gtr dc She*ol Ef 0hntnn(f9{rf$q8}-epfi. li'Sbl.r[at lntgtlt qr son$tatat cl fu tta:"ql lr*l.oadel ila na5}sr*11,a lF4)rAu&rtl fl n$lcu s stebttcgtr ru lntre msl"scr pl sltopl,amA Fs tt$sfoErto st|gtsrrllnlta It*t ) fl er$l6ul nu nsl efe o f,orug resnlat6rlll surttn sE Art! s.st alt *otLs rs f,$a BrttBlr elat firod"rtbos8trrlcrprln rtst trp fulr ml&rtutr rLtonrlslsla&r 6o fonoeasl ta cl,toplsnfr,e Iot fa prtocdo dr rcpann a*E gblrtratr'lndlrrl4rdltnr*r " cltrp [or1 prln pLsmottsaroa lorreeo$o ducs le pLmdcras s$$tlp?ulut btnc nen}rnna qi cXtnplsmAr lnattrrl, Iteadnov* trr&ir gtr ssoln {irT*(1961) ofst6 s6 rtn contllflll* 01n 891 ta tficaxrtr reps$nrlut pno&mrl dl,aparo E1 le$twE rtl,nFr st* flt prtn lrrtsnssattuL plannodemolorr tdulfl *utnrlr lF5) n nfguarlut Iri$ Srlntre cese gl i;or$t*tr i,rlr(lgftlri{g'tso? g.rr*onrldar{ e& eaa nsi W trdl,raturd fogtru pcrcufgshibornstr cetc tsaperst;l.lrs latra oo-&Oo0.dr rosaro{ uorl,atsn*tlcsriatfrr olteplaml

*70* gtul c& nn[lerl thrt Jsuf *llElsr unor aetfel flr teoparatrrl[ cudlltlilor ce a*ls lB dscurnrl ls*atl' ds la r*tl

ila vertebiLln

ternl.ls cgnd {btate nfuanlut

m& trundr ta "IO Aa tins]l

to* l0oe sata ah*ssaru

I uarsrgdJaa d.tr.forengetilc ls u$ es Ia sltu.l atrtorlS * tstertBtlstrla l{llaad, ds !16diu rLe rmrll.cf.lrd

ml}to poectpl,tatl,t r srE lec t crotto3e h'ul; anmrlu* ) suhstangr dr raa*r.rl1 tnr tsamng ealdi gl" uacc*& d*ert!,ug o f,lordfert in stsrs lllle buas' Sl o trsears anrrial& s fl{$rt.lor ) topans.'.lo*te {pl$tos &ro la sr!e8aE8urnrl. r'e9anr prrfuad, 91 ti*
Eat,jr

ctnd. ?Ertu rqscslts este bogEt6 9L n dsut Jis$tat* a rmrll ile plqtsarA gt reseple f,el gt toesnc* spsste condtltt ur f*rpl* lo6 trroc8sa toemturl-lor 3n EtEro ds !'opf,ls t8gfuEil el transfoBr jra cubstantelos ds *ogdtlt8r solmt*? L.tr(1961) rnonllanoaaEEE nEpmrnrl &,t'oernql ?l&*tli co.re glautolo Feo *&cutasotalle h sea& sr Lso no{ltfto$rlJr stEp6 s* ) ifsui etapsrr@sta Fload& do tsspsrstntl bobt$ige nebtlsneto 91 @g*srs$ prin mrea oA pls$atca rutoll,lcasA r tsperahui

$oaaalba shls s dsri.rp sepsblJl os te$lt FDI chr,Elfel sl solabr(f$a]rd*n cotrdftufe ou.rnatlsc-,tn DrlrEtpst rtqeJtosstu:te uel r1* I rhtBll bat& l,rnsto lcsi$l biolog*srprF$e gl trun$lree al'1,st-4eseats i& iG6

91 I'natato *fi$3as sonslbt'lE iatr.clll 6rr{ nol 4tslto(rtotra&o(19qE)

te astivf,tats gt 6e'01 rsl{ grrosasgL ds e*g$ter. In sts)o s dfiI| a ro?dr$rlul n-a *o,astatat c& plontotrf Laana&. t .lnt foaft s*ieteate l& sBrr*iuiiu?ll lor pot realsto 1 ln iirsr$t, ordr lulolddr ptnd la - af c($*r$ipstr -,.ir(1953)rtttl&" lltaonmst 1964 dist.gou&rnu(L965)*tsrl.etnnta la *er eats assul* ) rlrf unrt Procss flstoLogl nst* o8l.be(spti.eort i$r *l *;:1s-1ll,pva( 195a)ri*$iir.!lLe!( 19F )r

$ rl'

:nc$rtlrl c$llril Brlrro

prln procoeole bl,$ohtrs&s & Eo aE{"f,68t6

q mldonuLul tn or$snele plaateLl cw corerpudr lte+ r1 nopau*ulut adlm le ponlrla o*eira a d{ruara repauodul' fglt*tr ilbea rtinsc ptrntst *ulnlnsatlcsnf,slad
3llofr*ilil;ortsf, r-steslal aL g,eetel

na*furunul *o retlatonfE'
o soru:*ti::s t?cnf

reatstoa|s

raaroo e$liletnd.ul de lEeewl6 tn aainnrll :!

eslubglE(s$,eref,9$dtt

?.e.)le unolo spoctrfl lar ls altols f,n ulsL in'".lttr,a vle:lercersst&toeli ar: r.nl ar$tnt c& l"a r'1grwltar. n,rEgls*onfet ;:lantetol lennpaso la i;er contrtbr*e gt d*iltral' l$u :ra& tlzl, rcaco ;E geupqrla X$ coluLe ( fl'ontils!.nfi 1te.r!rurtlplgnestl f.nil,lbeprocosolo ds s".gts!6tltrghtrt !|"onL,ct gt gltwlsntclhgse rl sl eolabr(tr9?0)r :rln Loq!.*oa*ln porlosda ds rop*ra1illa twu se seetablls;:s E[t1rx.lte dtnt$B coluls prln ltltelnmdtul placmodnsroloardt"tlarl loat6 4 ltpo' Blvl&rslfss,fsa slto:,lemcl," are loc dseomtrtrla$$oa forr. lclorl e tasrqrrtlor 81 e Eomb,in.olgttor llpo.ttrotslcsrrrln Ee do mlilon ss 6$.Fegto gra&rl ds psmsabl.lltato gl hi4r'stsra r Itoptesslroo r$rogto aontlautnrl X$ ap& llborf1ee oi:*6 1* hlts* lflcalon trmeestrsr f,talolostrcer tn lera iloaswg setsusul bl"tsrnal a Sost attr&i*t r1cc{tre i;1cr reloi(l999)faer a sdslsrLaontat tnf,l'uan$andourttor :)::rotelmtos twrs{r duretoi ropaunrluL et a reststsntel lfer! la tm.,Le soiurL rb cal,sr la fsr 6 nu"il$ltoa f,lor

r,urata nspEugtrlut gl anBeatolo anatmi.*q oe nelrcileqa$.tlBcQsdf repeumilul, eR:f,s$t EnAtrtrntft latFe stup& gn altolamr dttgttul !.l!.astl 1re4 ooirrl lllordotrenr (19&lh Itolt bt i:lo calsl d.o erltrFi un solcttv esntlrg d8

.,atallo doNonomr(1961,) u,rta&r!.tde asssnda ta nugurlt {Lor e ds cnLs repqwul. lrlboraalloonolln'rlrl' a g#eetorar c{ rosLststlta la ln

*fA* .itcatrduahea(fg66) s sguiitsq rFcusul l.l;rsrnal la 1- eolurt t'{ lrralslcorelt$drFl cu tonJrsssturLlo Joaee gl ou ualdli{rto*r forcoeslrqB strlril.at p6:)susul r; Jer8al l.! | trucont qti,n r,fiuilrrl lwcrra gtlr( t 9ru ),stabLl.ln*i ilur:*rs sesfirl*p lr r ta corxillllle'tla rltls!fiXa{i eelo dstr& stapq pe bsaa fu}dl,etlon ecstsrlst $ts}r..xttl I i'tl,lrldeiamr ql aoleib.(19i1) Sf. egectrrl,ad, aaleulo catelatt{rar 9t ctrrlar reaaltat oonffusA d*ve:llle{ lsdLett)Esatmlco,

tl:sg!ete_sblig.tg'
I:3 toaIrn*e$t'1sr Lgt){.Fa$t?*g6'jt+1g68r:.g6e.195gfir gurnt r$ir bl &patn f,l Eofft d* dfid[fu$ar 6 rcAsrsrlul btbonml la ml I teepnl lusft, f,u etnrdlufin ooaui,f{,![e ,{r, llveds el Cf$dl.nn h, nLe& e fnEH,t*tr.r,1uL egrennril*,'itlrs.Loe*u|rr a{a ate EpBtlor}i}sE& ansesogi Eutorl($e*gl*Ev a.i. (lg}1 rlgg}) r fu&s'" ur& (1g59)rluplf 3a iif r. ( 196L r8t ltsoaava(lgg ) rsl,rs.r Ir ) l(1963)riindlsaenlrop*rlrrrirars$orrho tilri'r{l9tr6}#prst itlsiien$rrtl Xr |1968) rtar tn |ara &$astrsf :lrl*adelanu gt eslsbr(lg$l ) r-$st*ilrtr p (19&i)g.erodst& $u otn&," {f, nstg allfsrg&tL$ss gl sFectsllss$r cdtcfurlu!,pccrsls I r er,lisoceulut f i ou]ef allt la Brlnrrdltlor Ireo $shsts !,rosseul do d^lilerenfts?rsg uugurlLor ]ilorl.ftrl., eptta iatrtnd fu portoerl* dE lstnt{ fl:.at$ esesr.tsr*${rsi,t$ 'rtp It'r S1 $nrEc gB pfula sn etald1 Fs{a & r.*pnuxu,l,ul ad.isl stablli,.;$a aosent*tu,l rta !.nh.ar$ ia ;srl,Esri,o )reunrf$f edlmlrJatelo ssslrrss cprfl"rE[ corclusll-tc le caro eu fus t.&!dRt*nr gt colabr(lgbl)1aaro an etablltt cS 1tn osndiF illo dln jleu$stlrporilt tntnli f.a r"pelr$ nr 5,a norr*ntr:J, tu*e.t[!1 art gtarll ! oldor$ t$Etsrl.lor rtla sue sfr,xn& i:uJav(lg5?] rUsanl*ske f lbtEo-f,tirarl oltstl d. &cet*,tuerba#(tr966hE1 egocn +4s#lb$d,rtl !n 3s$&selsrrln oosAtfll!$ cnl,los snsllestt1n#$satul tnp llr11 t$ atqps ropsusrtul adi,nolcorelat& au lrcepu:nl c&dnrll mrclory a.rn dssf,&.gurnt la sf&ryttut lut ocptonkl.* *l oG*ssbf s Ia oc prlvogto

-71 5 tl 196?r ps elsd ts 1964 r-t llF$Lr DcafAeur6rll stpa$utrut *Elor 1o talr?at ssto lsrd.* graftstrl $rr2r urdo oe Soate obr tn tabslul rsrgr Fttclm lt $.rs fte gl. iefluatsto fqstorlloc toupomhrra qt mLdttrta 65upra G&Arlifr*j, seahrl&r Sntl,l,a| trbolulul gl, Ercf'.oulul rrtrcvd I, arilt rgF{etct gi ffiitr*t a/ lgryAealuceo@nlnh Sfl efootnnsrr ropauant ln aorlagisr frpt care sr Er9llst nl l il*,frronftenet rlusuairor flarifrrtl sN nat suatle& fef6 de trm &s|str[r daol. stt cct*trti risptt latm Oa- trfg [1r $.'$t d' rtl ds tetrleratnrDl ast rtil*c*fi a sro lBt at.rortr"Eltilr 8n rona teulieratdl tar 9t ds ol't6 partr its orietJl'a tls f,*tortrl amrell cliPatlofr tgt ln*ep rtpsuelll la raptenbrto 6ot b/ 'rutt- eqlJ.altt tmnl nugu.r'ilor flortSr* dla mul sgenopli3tatoc dlfoleafterll Itsbrtr b dr glrl tolnlr5. abla lu fcbruaElc mu aartls rrul ulnltcsr gr/ Set esi. uL& m86r dn stls ue rtilaus ael,lr tl g33 in 9!ln ll riaf, .trygililu. *oanunlcl otriol' tsruage domrtl$r '*o$41 Agc$ t Ssqoslnoa wl$ssls |'.t& nlll I rgle.rzutui *aolul actn tLnfu;Etl &c 4r*d1 lrs sor esf tua8[ EI'ot ]e'ilr. l-s rr|rt carrsl' a|brl.bl 691611 avird o posltte t"r atl*latte dcdAsrrat lrcoplnA ou lum

ci locelc* }lrlssrlrr

lesillsrlr # ln eoudltitlo dta l|.muregtlp 3a ost , anN'de ccssrtrrsl 'roirrl Slctrocd trr il aal lur6 repsJt tr-ru svrit 0*rslt "stis gl, 0ora$rs nrtgarXs -dotnl e'qrl '$,$1llr81l,r lrs lspnrrrrl Ilry l! ari $su!t lr* trut &fEltal'u smrnl'*r laa o polltte lntqlncdis$r o.lt*ltt tffini e*ut

Age ibrp& oun $eusreil er8tr. autorl - 8r 8or*tLsst 91 ookb' sogsl'st Irr. (1961)r L t* llt!* sqt*go$sno{sr tlFl}r lSl)r arx&ri Elssoanbo arEbl tlgGllr aupltln lrtr* (1961)r aadtra*snBo totr (1S6)r ltonol 8rr slrtraNt & (trse) lrrrr dbrpf,latlrmr in

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teii'E t* csa &s a {cua EtaFS I sflar E l0qcr rugrutt florlferl C.sogeblt8d* pri.laats:;qps*lpaum$ul f,ortitt sau Itrl1 c*ltt'ltlv tltctlvl$n pletr*lla rttt oa$q susurit flsrtf,ert s$nt sal sonstbtll f,a${ ds : sc[ruter'.:sro*l soagln is sisre do latoapiil{ta! eXat

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