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USATF TF Level II Coaching Education CVPhysiologyNotes
USATF TF Level II Coaching Education CVPhysiologyNotes
Click the Goals Button: These are the goals for this
module. These topics are all very important background knowledge for a coach.
Go to the next screen and Click the Button labeled #1: Diffusion. Diffusion relies on the uneven
distribution of a gas or fluid between two different compartments. In the alveoli of the lung which is one compartment - there is a high concentration of oxygen and in the blood vessel which is another compartment - there is a low concentration of oxygen. The driving force for oxygen from the alveoli to the capillary is the size of the concentration gradient for oxygen. This is referred to as the difference between the partial pressures for oxygen. Just think of partial pressure as an indication of the concentration of a gas it actually could be oxygen or carbon dioxide. Note that carbon dioxide is moving from the blood capillary into the alveoli. This is because the carbon dioxide concentration in the blood when it returns to the lungs is much higher that the concentration of carbon dioxide in the alveoli so carbon dioxide will diffuse down its concentration gradient into the alveoli of the lung. The principle of diffusion explains why you smell the perfume when the lid of the bottle is removed. The molecules of perfume are in very high concentration inside the bottle. If you remove the lid the perfume molecules will move down their concentration gradient into the air around you. The molecules will continue to move down their concentration gradient away from the bottle. When blood is moving through the tissues, as you seen in the movie clip at the bottom of the screen note how carbon dioxide is moving down its concentration gradient from the cell into the blood vessel. Oxygen is moving from the blood vessel into the cell.
Click the Return Button Click the Button labeled #2: Transport of oxygen. Oxygen does not dissolve very well in water. One
liter of blood can only hold about 3 mls of oxygen and the heart would have to pump over 1000 liters of blood per minute to deliver the 3 liters of oxygen that the muscles of an athlete may need to cover the demands of their muscles during a strenuous run. The strategy used to improve the situation is to add an oxygen carrier to the blood that appreciably increases its oxygen carrying capacity that reduces the blood flow requirements. Lets take a brief look at how the oxygen carrier works.
Click the Return Button Click the Button labeled #3: Transport of carbon dioxide. Carbon dioxide or CO2 is the gaseous
waste produced by the Krebs cycle. As fast as the CO2 is produced the gas diffuses out of the cell, and into the capillary. As the blood is passing through the tissues it is dumping off the oxygen the cells need and at the same time it will pick up the carbon dioxide. The PCO2 or partial pressure of carbon dioxide in the cells is higher than it is in the blood that is entering the tissue bed. At rest, the PCO2 of cells is around 46 mmHg but it can get a lot higher than this during exercise. The PCO2 entering the tissue capillaries is around 40mmHg. As a result, the CO2 diffuses down its
concentration gradient out of the cell and into the capillary. Now we will take brief look at how CO2 is transported back to the lungs where it is expelled into the atmosphere.
Click the Return Button Click the Important Terminology Button: The normal heart is a muscle about the size of a fist. When the athlete is at rest every beat it drives about 3 ounces of blood through its chambers. Inside it is divided into two parts. Blood enters the heart on the right side which contracts sending it from the upper right chamber the right atrium into the lower chamber the right ventricle. Another beat pushes the blood again into the lungs where it picks up oxygen before moving into the left atrium which pumps it into the left ventricle. The left ventricle has to pump blood to the brain and to the most distant cells in the toes. When an athlete starts running the working muscle cells need more oxygen. To meet this demand two things happen 1) the heart speeds up its delivery of blood and 2) there is a redistribution of blood flow from inactive organs to the active skeletal muscles. In this section we will overview the important terminology used to describe amount of blood leaving the heart such as cardiac output, stroke volume and heart rate and how exercise and training affects these in the immediate and long term. Getting the blood back to the heart is the job of the venous system and we have three pumping mechanisms that helps accomplish this task. We will briefly overview these three mechanisms. And, we will finish this section by taking a look at how the body redistributes blood flow from organs that are not essential to the exercise to the working muscles that are in desperate need of the oxygen.
Click the Cardiac Output, stroke volume and heart rate button: Cardiac output is the amount of blood
pumped per minute by the heart. Increases in cardiac output are closely related to how hard the athlete is working which, in turn is related to how much oxygen the working muscles are demanding. The cardiac output is the most important descriptor of heart function because it determines the quantity of oxygen that can be transported by the blood. At rest cardiac output is around 5 liters per minute and increase to around 30 liters per minute during exercise. Blood flowing into the right atrium and ventricle must equal the flow through the lung and flow into the left atrium and ventricle. If you roll the cursor along the red bar under the graph you will see how cardiac output increases. Cardiac output depends on two variables the stroke volume and the heart rate
Go to the next screen: Note that at rest there is hardly any difference in cardiac output between the trained and untrained. However, a trained runner has a higher cardiac output capacity. The question is how is this higher cardiac output accomplished. Remember that cardiac output depends on two things the heart rate and the stroke volume. Stroke volume is the amount of blood the heart can pump per beat. Lets take a look at stroke volume and see what happens to that under exercising conditions.
Go to the next screen: Maximal stroke volume can exceed 200 mL per beat during exercise at rest it typically ranges from 50 to 110 mL/beat. Slide the cursor along the red bar and watch what happens to both cardiac output and stroke volume. Note how stroke volume reaches its highest value about half way along the red bar. It turns out that stroke volume reaches its highest value at around 40 to 60% of VO2max. Also note, that cardiac output continues to increase despite the fact that stroke volume does not change any more as the athlete runs faster. When the athlete reaches anaerobic threshold or the point at which the aerobic energy system cant keep up with the energy demands and the muscle cells cant clear the hydrogen ions or lactate. The athlete can still run a bit faster but the build up of hydrogen ions will eventually cause the athlete to stop and recover. Theres only one variable left the heart rate. This variable is responsible for the increasing cardiac output even though stroke volume does not increase after about 40 to 60% of VO2max.
Go to the next screen: Maximum stroke volume can be double in the athlete. Women generally have a lower stroke volume than men do because they have smaller hearts. The larger stroke volume is the result of the increase in heart size that can be achieved through training. However, there is also a genetic factor involved here elite athletes appear to have the genetically larger heart and this, in addition to the training effect, provides them with an athletic advantage. Keep in mind that any adjustment in structure of the heart is a slow process that typically takes months even years to be completed and there is a genetic limit to its stroke volume capacity.
Go to the next screen. Now lets talk about heart rate or the beating frequency of the heart. Heart rate increases linearly with increasing workload or the volume of oxygen consumed. Slide the cursor along the red bar to see the relationship of heart rate to stroke volume and cardiac output. Note that while stroke volume reaches its maximum about half way along the red bar the heart rate keeps increasing up to the anaerobic threshold. Cardiac output stops increasing at this point as well. So, after about 40 to 60% of VO2 max the increase in cardiac output is due to the increasing heart rate. Stroke volume is related to the size of the heart and this is fixed depending on genetics and its structural changes that can occur with training over a long period of time. But heart rate is capable of instantly changing over a wide range according to the oxygen needs of the muscle cell. The beating frequency depends on the autonomic nervous system and hormone level especially the amount of adrenalin or epinephrine in the blood. Go to the next screen: Heart rate is what is known as its functional capacity. It allows rapid adjustment to the needs of the moment and these adjustments occur in milliseconds to seconds. The structural capacity of the heart sets the maximal amount of blood it can pump in one beat. The body can adjust this capacity to higher levels if needed and does this by enlarging the heart size but this adjustment occurs very, very slowly. This structural adjustment is an important feature because it allows the heart to be kept at the smaller size until the need for enlargement arises. Indeed the heart will shrink back to its normal size once the athlete stops high performance training.
However, the maximal beating frequency of the heart does not change with training. This is fixed and depends on the age of the athlete the older the athlete the lower the maximal heart rate. Training does reduce the heart rate at sub maximal workloads, though, because the heart becomes stronger and can pump out more blood per beat. Indeed, a slow heart rate in a healthy individual indicates an efficient circulatory system. I want to make one more comment about the max heart rate while there is no training an athlete can do that will increase the max heart rate, there is a gradually lowering of the max heart rate with advancing age. The max heart rate loses about 1 beat per min each year. This becomes a relevant issue for you if you are training older runners and you are using heart rate as an estimate of training stress.
Go to the next screen: Lets just quickly review what we just discussed. The amount of blood the heart pumps in one minute is called the cardiac output. Cardiac output depends on two variables the heart rate and the stroke volume. Stroke volume is the amount of blood pumped out per beat and the heart rate is the number of beats the heart makes in 1 minute. Both stroke volume and heart rate increase during exercise increasing the cardiac output. Stroke volume increases up to about 40 60% of VO2max. After that it plateaus. Heart rate varies in direct relationship to the workload and plays a larger role in increasing cardiac output.
The stroke volume can increase with training but there is a genetic limit and it takes a long time for this to occur because it involves a structural change. There is an upper limit to the maximum heart rate and this does not change with training. However, during rest and exercise the trained athlete has a lower heart rate for the same workload than an untrained athlete. And finally, the athlete has a higher cardiac output than the untrained. Some of this is due to genetics that is the elite athlete has a naturally larger heart and some of it is due to the structural changes that occur with training.
Click the Return Button Click the Venous Return Button: The left side of
the heart can only pump out as much blood as it receives from the right side. The venous system is responsible for keeping pace with the cardiac output needs. The blood in the venous system is under very low pressure and the walls of the veins have a very low compliance which simply means that they easily expand to depending on the amount of blood they need to hold. If the cells need a higher cardiac output the blood must be sent back to the heart at a higher rate of speed. There are three mechanisms that increase the venous return to the heart. One mechanism is a muscle pump and a respiratory pump and the veins will constrict. This reduces the amount of blood that is in the veins forcing it out of the veins towards the heart. As the muscle contract their veins are compressed and the blood within them is forced toward the heart. Note the valves in the veins that stop the backflow of blood until the next muscle contraction. The second mechanism is the respiratory pump, As the chest wall expands due to the brain signals to the respiratory muscles to contract this lowers the pressure in the chest cavity allowing the veins in the chest cavity to fill with blood. When the athlete breathes out the pressure in the chest cavity increases sending the venous blood towards the heart. And finally, the veins will constrict and this reduces the volume of the venous system squeezing the blood toward the heart.
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Click the Return Button Click the Home Menu Button You have now completed this module
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