Trevor D.

Keyler

TKEYLER001@CSBSJU.EDU PERMANENT:
WEBSITE: HTTP://TREVORKEYLER.WEEBLY.COM/

2014 Forest Ridge Lane

EDUCATION Buffalo, MN 55313
612-940-0303
University of Minnesota, Minneapolis, Minn.
• PhD: Integrated Biosciences 2018
• MS: Integrated Biosciences 2014

St. Olaf College, Northfield, Minn.

• Bachelor of Arts in Biology, May 2007

TEACHING EXPERIENCE

College of St. Benedict/St. John’s University

Visiting Assistant Professor
- Introductory Biology: Fall semester 2018
- Fundamentals of Human Physiology: Spring semester 2019
- Human Anatomy and Physiology 325: Fall semester 2019
- Human Anatomy and Physiology 326: Spring semester 2020
- Student Registration Advising, Fall semester 2019
University of Minnesota Duluth
Graduate Teaching Assistant
- Ecology lab: Fall semester 2017
- Animal Physiology lab: Fall semester 2012, 2013, 2014 & 2015
- Biology in Society lab: Spring semester 2016, 2017 & online course
- General Biology 2 lab and discussion: Fall semester 2014
- General Biology 1 lab and discussion: Spring semester 2013
Course Chaperone and Assistant
- Woods Hole Marine Biological Laboratory – May 20-June 3, 2014
- Friday Harbor Laboratories – May 22-June 1, 2013 and May 19- June 3, 2015
• Teaching Assistant in field studies in marine biology course: assist students in
field, lab and class work. Instruct ichthyology lecture and lab.
Creation and approval of IACUC protocol 1504-32496A, May 15, 2015
Boreas Leadership Program
- Building better presentations. April 25, 2014
Guest Lectures
- Ichthyology: Schooling and Shoaling of fishes. February 8, 2016
- Ichthyology: Thermoregulation and Buoyancy. February 3, 2016
- Freshwater Ecology: Trophic dynamics. November 2, 2015
- Animal Physiology: Pattern recognition in spitting cobras. September 14, 2015
- Ichthyology: Blood & Circulation. February 3, 2015
- Ichthyology Lab: Bony fish dissection. February 3, 2015
- Ichthyology: Siscowet Foraging. February 5, 2015
- Freshwater Ecology: Trophic dynamics. October 16, 2013
- Animal Physiology: Pattern recognition in spitting cobras. October 17, 2013
- Animal Physiology: Ionic and osmotic balance. November 11, 2013
Semester Course on Teaching
- Teaching and learning in the life sciences. Spring semester 2014
St. Olaf College Biology Department
Teacher’s Assistant, 2007
• Collaborate with professors to help teach student’s basic biological concepts.
• Set up labs and assist students with completion of lab work.
PROFESSIONAL EXPERIENCE

Learning Enhancement Services New Faculty Mentor- Fall 2019-Present

College of St. Benedict/St. John’s University, Minn.
• Mentoring resource for new faculty member

Writing consultant–August 2017–May 2018

University of Minnesota Duluth Academic Writing and Learning Center, Duluth, Minn.
• Provide one-to-one writing consultation to undergraduate/graduate students and faculty

Fisheries Technician–May–September 2016

Wisconsin Department of Natural Resources, Escanaba Station, Vilas County, Wisc.
• Limnology: D.O. reading, Chlorophyll A data, zooplankton collection, Secchi readings
• Monitoring of research lakes including limnological data collection and pop. assessment
• Fish assessment: Fyke netting, electrofishing, gillnetting and hydroacoustics
• Creel survey and demographic data analysis

Associate Researcher–Pre-clinical in vivo neuropathy studies, July 2011–March 2012

Algos Preclinical Services, St. Paul, Minnesota
• Coordinate contracted preclinical pharmacology/physiology testing to validate compounds
• Behavioral research in rodent models to assess pain states
• Surgery procedures on rodent models to induce/relieve pain states

Junior Scientist–Pediatrics, October 2008–April 2011

University of Minnesota, Minneapolis, Minnesota
• Detection of proteins post bone marrow transplant associated with genetic disorders by
method of Western blotting
• Culture and production of retrovirus via transfection of kidney cells to be used in the
production of induced pluripotent stem cells
• Culturing of transplant patient’s fibroblast cells to be used in immunoblotting. Tasks
included making growth media, counting cells, maintaining cell lines, harvesting cells, and
preparing cells for experiments
• Learned skills: PCR, Western blots, cell culture, transfections, gel electrophoresis, UV/Vis
spectroscopy, protein assays, densitometry, genotyping and biological sample processing
and handling

Research Coordinator–Endocrinology, July 2006–April 2008

Hennepin County Medical Center, Minneapolis, Minnesota
• Collated medical data on Somali immigrants in correlation to diabetes prevalence
• Blood collection and glucose monitoring of patients in diabetic ketoacidosis
• Retrospective determination of gestational diabetes in Somali immigrants

RESEARCH GRANTS

Principle Investigator
2018 St. John’s University FDRC Grant
Deep Water Sculpin Behavior. $4980

Co-Principle Investigator
2016 Wisconsin Department of Natural Resources, $31,218
Optimal Walleye Foraging and Survival.
Co-PI’s, T. Hrabik and G. Sass.
SCHOLARSHIPS, FELLOWSHIPS & AWARDS

2019 Becoming Community Practitioner, College of St. Benedict/St. John’s University

2019 Academic Travel Award, College of St. Benedict/St. John’s University $1022
2018 Joan Duffy Student Travel Award, Minnesota American Fisheries Society $200
2018 UMD Graduate Office Student Travel Award $400
2017 Integrated Biosciences Graduate Program (IBS) Travel Grant $500
2016 Brule River Sportsmen’s Club Scholarship $5000
2016 International Association of Great Lakes Research Scholarship $2000
2016 Best Student Presentation Award, Minnesota American Fisheries Society $200
2016 Sander Award, North Central Division of the American Fisheries Society $200
2015 IBS Graduate Fellowship, University of Minnesota Duluth $8667
2015 Great Lakes Fishery Commission Travel Award $1500
2015 Swenson College of Science and Engineering Travel Award $500
2014 Minnesota Sea Grant and Lake Superior Chapter, Muskies Inc. Scholarship $1000
2008-2011 Regents Scholar, University of Minnesota
2006-2007 Honor Student, St. Olaf College
2005 Dean’s List, St. Olaf College

MEMBERSHIPS & COMMITTEES

2019 LES New Faculty Mentor, College of St. Benedict/St. John’s University

2017 Co-founder of Café Scientifique, Duluth, Minn. https://cafescitwinports.org/

2017 Co-founder of Twin Ports Early Career Scientist group

2016 Advisory Board for the Center for Excellence in Teaching and Learning: Graduate
Student Representative. http://d.umn.edu/center-excellence-teaching-and-learning

2015 Founder and President of the American Fisheries Society Student Subunit at the
University of Minnesota Duluth. President: May, 2015- June, 2016.

2001 Boy Scouts of America: Eagle Scout

LABORATORY/FIELD SKILLS

Fisheries-Related Science

Behavioral Conduct and analyze foraging experiments relating to reaction distance, angle of
attack, burst attack speed, capture rates and handling time. Competent in infrared
digital recording, light intensity measurement and ImageJ analysis.

Physiological Electroretinography (ERG) for deepwater and littoral fishes for collection of
visual sensitivity data. Light intensity data collection and analysis.

Collection Deepwater trawl collection and successful captive husbandry of deepwater fishes.
Experienced with gillnetting, fyke netting, seining and electrofishing. Long-term
mark-recapture studies.

Laboratory RT-PCR, PCR, Western blots, transfections, gel electrophoresis, UV/Vis

spectroscopy, protein assays, densitometry, cell culture, genotyping, plasmid
 purification, titer preparation, detailed protocol preparation, colony maintenance,
necropsy, SigmaPlot, Excel and Prism software competency

Other Sciences

Behavioral Pain Models: Neuropathic- Spinal nerve ligation, Chronic constriction injury,
Streptozotocin induced diabetic neuropathy. Inflammatory – Formalin, Complete
Freund’s Adjuvant, Carrageenan and Capsaicin injections, UV burn.
Implant/incision- Bone cancer, Post-surgical pain paw incision.

Endpoints: Thermal Pain- Hargreave’s, Laser, Hot plate, Warm water immersion
Tail flick, Radiant heat tail flick. Mechanical Pain- Von Frey filaments, Digital
Randall-Selitto paw pressure, Weight bearing.

Field Environmental studies in Australia:

Primary and secondary succession of grasslands and temperate forests, fire
regime implementation, tree and weed growth transect studies, tidal zone quadrat
studies, bird banding, mammal trapping, population estimates, gap dynamics,
niche occupation, otolith age estimates, insect predation.

Timber rattlesnake research, Mississippi river valley, Minnesota:

Protection/scientific oriented research on Crotalus horridus in collaboration, and
funded by the Minnesota Dept. of Natural Resources: Assist with capture/re-
capture population estimates, telemetry, movement and distribution, longevity,
sexual dimorphisms, DNA sample collections, and loss of habitat and disturbance
effects.

PEER REVIEWER

2019 Aquatic Ecosystem Health & Management

2017 River Research and Applications
2017 Transactions of the American Fisheries Society
2016 Journal of Photochemistry and Photobiology B: Biology
2016 Bulletin of Marine Science
2015 North American Journal of Fisheries Management
2015, 2017 Duluth Journal of Undergraduate Research

PUBLICATIONS
Keyler, T*., T.R. Hrabik, A.F. Mensinger. L.S. Rogers, and O.T. Gorman. 2019.
Effect of light intensity and substrate type on siscowet lake trout (Salvelinus
namaycush siscowet) predation on deepwater sculpin (Myoxocephalus
thompsonii). Hydrobilogia, 840(1), 77-88.

Keyler, T*., B. G. Matthias, and T. R Hrabik. 2019. Siscowet lake charr

(Salvelinus namaycush siscowet) visual foraging habitat in relation to
daily and seasonal light cycles. Hydrobiologia, 840(1), 63-76.

Keyler, T*., T.R. Hrabik, C.L. Austin, O.T. Gorman and A.F. Mensinger. 2015.
Foraging mechanisms of siscowet (Salvelinus namaycush siscowet) on pelagic
prey. Journal of Great Lakes Research, 41(4), 1162-1711.
doi:10.1016/j.jglr.2015.09.016
 Keyler, T*., T.R. Hrabik, L. Roger, K. Harrington and A. Mensinger. Visual
sensitivity of Walleye (Sander vitreus). In review. Journal of Great Lakes
Research

Keyler, T*., T.R. Hrabik, L.J.Rogers, and O.T. Gorman. Foraging mechanisms of
siscowet lake trout (Salvelinus namaycush siscowet) on the benthic deepwater
sculpin (myoxocephalus thompsonii): effect of light intensity and substrate
composition on predator-prey response. In Prep.

Keyler, T*., T.R. Hrabik, and B. Matthias. A Lake Superior light model for siscowet
lake trout visual foraging habitat. In Prep.

Umpierrez, G. E., Jones, S., Smiley, D., Mulligan, P., Keyler, T., Temponi, A., &
Robalino, G. (2009). Insulin Analogs Versus Human Insulin in the Treatment of
Patients With Diabetic Ketoacidosis A randomized controlled trial. Diabetes
Care, 32(7), 1164-1169.

REPORTS

Great Lakes Fishery Commission, 2016 Completion Report. A Hyberbaric Holding &
Transport Vessel for Collection of Deepwater Fishes for Research and
Broodstock Development. Owen Gorman & Trevor Keyler

Wisconsin Department of Natural Resources 2017 annual SFR report. The Effects of
Water Clarity and Light Intensity on Jevenile Walleye Foraging and Growth.
Greg Sass, Tom Hrabik & Trevor Keyler

NON-PEER REVIWED PUBLICATIONS

University of Minnesota Duluth – Swenson College of Science and Engineering and University home
page article entitled Fishing for Results. Published March 14, 2016.

Minnesota Sea Grant – Article in The Seiche featuring lake trout research entitled Lake Trout and the
Tao of Research, December issue, 2014.

University of Minnesota Duluth, Alumni News - Alumni web page article entitled Fish Finders: Research
Discovers Siscowet’s Spectacular Vision. Released June, 2014.

University of Minnesota Duluth – University web page article entitled: Research Discovers
Siscowet Have Spectacular Vision. Published April, 2014.

INVITED ORAL PRESENTATIONS

Trevor Keyler, What Does a Fish See? Central Minnesota Audubon Society. St.
Cloud, MN, October 16, 2019.

Trevor Keyler, Visual Properties, Behavior, and Habitat of Select North American
Fishes. GRAD TALK. University of Minnesota, Duluth. March 13, 2018.

Trevor Keyler, Thomas Hrabik, Loranzie Rogers, Owen Gorman, Kelly Harrington,
Allen Mensinger. Foraging Behavior and Visual Properties of Select Midwest Fishes.
Trout Lake Station, University of Madison, WI. Boulder Junction, WI, June 29, 2016.
 Trevor Keyler, Allen Mensinger, Owen Gorman, Thomas Hrabik. Study of foraging
behavior of siscowet lake trout on pelagic fish. Muskies Inc. Lake Superior Chapter
guest speaker. Duluth, MN, October 14, 2014.

Trevor Keyler, Allen Mensinger, Owen Gorman, Thomas Hrabik. Foraging

mechanisms of siscowet lake trout (Salvelinus namaycush siscowet) on pelagic prey.
Great Lakes Fishery Commission, Lake Superior Technical Committee Meeting, Red
Cliff, WI, July 28-31, 2014.

PRESENTATIONS / PROFESSIONAL MEETING PROCEEDINGS

Presenting Author

Trevor Keyler, Bryan Matthias, Thomas Hrabik. Siscowet lake charr

(Salvelinus namaycush siscowet) visual foraging habitat in relation to daily
and seasonal light cycles. International Charr Symposium, Duluth, MN, June 18-
21, 2018.

Trevor Keyler, Thomas Hrabik, Allen Mensinger, Loranzie Rogers, Owen Gorman.
Effect of light intensity and substrate type on siscowet lake trout (Salvelinus
namaycush siscowet) predation on deepwater sculpin (Myoxocephalus
thompsonii). International Charr Symposium, Duluth, MN, June 18-21, 2018.

Trevor Keyler, Thomas Hrabik, Allen Mensinger, Loranzie Rogers, Owen Gorman.
Foraging mechanisms of siscowet lake trout on the benthic deepwater sculpin: effect
of light intensity and substrate composition on native predator-prey interaction.
Midwest Fish and Wildlife Meeting, Milwaukee, WI, January 29-31, 2018

Trevor Keyler, Thomas Hrabik, Loranzie Rogers, Kelly Harrington and Allen
Mensinger. Visual sensitivity of Walleye (Sander vitreus). Twin Ports Early Career
Scientist Symposium, Duluth, MN, March 30, 2017.

Trevor Keyler, Thomas Hrabik, Loranzie Rogers, Kelly Harrington, Allen Mensinger.
Visual sensitivity of walleye (Sander vitreus): A comparison of freshwater fishes.
Minnesota American Fisheries Society Meeting, Duluth, MN, February 1-3, 2016.

Trevor Keyler, Thomas Hrabik, Loranzie Rogers, Kelly Harrington, Allen Mensinger.
Visual sensitivity of walleye (Sander vitreus): A comparison of freshwater fishes.
Midwest Fish and Wildlife Meeting, Grand Rapids, MI, January 24-27, 2016.

Trevor Keyler, Allen Mensinger, Owen Gorman, Thomas Hrabik. Foraging

mechanisms of siscowet lake trout (Salvelinus namaycush siscowet) on pelagic prey.
National American Fisheries Society Meeting, Portland, OR, August 16-20, 2015.

Trevor Keyler, Allen Mensinger, Owen Gorman, Thomas Hrabik. Foraging

mechanisms of siscowet lake trout. 2015 MN Chapter of the American Fisheries
Society Meeting, Brainerd, MN, March 2-4, 2015.

Trevor Keyler, Allen Mensinger, Owen Gorman, Thomas Hrabik. Study of foraging
behavior of siscowet lake trout (Salvelinus namaycush siscowet) on pelagic fish.
Midwest Fish and Wildlife Conference, Indianapolis, IN, February 8-11, 2015.

Trevor Keyler, Allen Mensinger, Owen Gorman, Thomas Hrabik. Foraging

mechanisms of siscowet lake trout (Salvelinus namaycush siscowet) on pelagic prey.
 University of Minnesota Duluth Biology Department Friday Science Seminar,
Duluth, MN, November 14, 2014.

Trevor Keyler, Allen Mensinger, Owen Gorman, Thomas Hrabik. Foraging

mechanisms of siscowet lake trout. University of Minnesota, Duluth. 2014 MN
Chapter of the American Fisheries Society Meeting, Mankato, MN, March 3-5,
2014.

Contributor

Trevor Keyler, Thomas Hrabik, Loranzie Rogers, Kelly Harrington and Allen
Mensinger. Visual sensitivity of Walleye (Sander vitreus). University of Minnesota,
Duluth. Great Lakes Fishery Commission, Ann Arbor, MI, March 27, 2017.
Presented by Owen Gorman.

Trevor Keyler, Allen Mensinger, Owen Gorman, Thomas Hrabik. Study of foraging
behavior of siscowet lake trout on pelagic fish. University of Minnesota, Duluth.
Lake Superior Technical Committee, Marquette, MI, January 7, 2015. Presented by
Owen Gorman.

Amir Moheet, Ameer Khowaja, Sidney A Jones, Shaban Nazarian, Trevor Keyler,
John V St Peter, Abdi Madar. Vitamin D receptor single nucleotide polymorphisms
in Somali immigrants. Division of Endocrinology, University of Minnesota,
Division of Endocrinology, Hennepin County Medical Center, College of Pharmacy,
University of Minnesota. The Endocrine Society’s 91st Annual Meeting,
Washington, DC, June 10-13, 2009.

Sidney A Jones, Shaban Nazarian, Crispin Semakula, Trevor Keyler, John V

St. Peter. Relationship Between Parathyroid Hormone and 25-OH Vitamin D
Concentrations in Somali Immigrants in Minneapolis, Minnesota. College of
Pharmacy, University of Minnesota, Minneapolis, Minnesota. American
Diabetes Association 68th Scientific Sessions, San Francisco, California, June
6 - 10, 2008. P2-307

Dawn Smiley, Sidney Jones, Denise Umpierrez, Angel Temponi, Crispin Semekula,
Trevor Keyler, Sheban Nezarian, Nola Tesfaye, Miguel Ceron, Guillermo E.
Umpierrez. The Benefits of Basal Bolus Insulin Regimen in the Management of
Patients with Diabetic Ketoacidosis (DKA).; American Diabetes Association 68th
Scientific Sessions, San Francisco, California, June 6 - 10, 2008. 547-P

Crispin J Semakula, Trevor Keyler, Sidney A Jones, John V St Peter. Prevalence of

Gestational Diabetes in Somalian Immigrants in Minneapolis, Minnesota. Hennepin
County Medical Center, Minneapolis, Minnesota and College of Pharmacy,
University of Minnesota, Minneapolis, Minnesota. American Diabetes Association
68th Scientific Sessions, San Francisco, California, June 6 - 10, 2008. P2-516

RESEARCH OUTREACH

Personal research blog: http://char-sighted.blogspot.com/

Minnesota Sea Grant- Guest on The Sea Grant Files radio show and podcast episode Burbot and the Lota
lota love song. Aired September 15, 2015. http://www.seagrant.umn.edu/audio/2015.09.15_burbot.mp3

Northland News Center- Interviewed for channel 3, KBJR news story on Lake Superior fishes. Aired
 September 9, 2015. https://www.youtube.com/watch?v=tf5tdb6Xr_w

VOLUNTEER EXPERIENCE

Professional Service
• Volunteer- Session moderator for the 9th International Charr Symposium, June 19, 2018
• Volunteer- Sea Grant/AFS beach clean-up for Park Point, MN. October 8, 2017
• Volunteer- Boyceville elementary school, fish anatomy. MN Sea Grant. May 31, 2017
• Volunteer- Sea Grant/AFS beach clean-up for Park Point, MN September 17, 2016
• Volunteer- University of WI, Trout Lake Station open house, August, 2016
• Volunteer- Minnesota Sea Grant info booth at Duluth Boat Show, February 20, 2016
• Coordinator- MN American Fisheries Society raffle, >$2000 raised, February 1, 2016
• Coordinator- MN American Fisheries Society T-shirt design and sales, >$600 raised
• Volunteer- Electrofishing White River, Iron River, WI. WI DNR, November 3, 2015
• Coordinator- Beach clean-up for Park Point, MN September 19, 2015
• Volunteer - Trout Unlimited/MNDNR coaster brook trout research. November 1, 2013

Outreach
• Volunteer- Invasive Buckthorn removal, Bagley Nature Center, November 15, 2014
• Volunteer- Guest Lecturer on reptiles, Hartley Nature Center, July 9, 2014
• Mentor - Cloquet High School students – First place: Regional Science Fair, 2013 & 2014
• Brain Awareness week - presented to 5th grade, Hermantown Middle school. April 4, 2013
• Judge - Lakeview Academy science fair, grades 6-12. December 9, 2013
• Volunteer - Northfield Hospital in the long-term care center, 2005-2007
• Judge - Northeast Minnesota Regional Science Fair, Biology. 2013, 2014, 2016, 2017
• Volunteer- Harbor City School, Careers in Science Day. October 2016

PANELS
• Advising Panel, McNair Scholars, June 10, 2014
• Advising Panel, University of Minnesota Bio Club Graduate Panel, March 26, 2015

PROFESSIONAL MEMBERSHIPS
Society for the Advancement of Biology Education Research
International Association of Great Lakes Research
American Fisheries Society
Minnesota Herpetological Society

UNDERGRADUATE MENTORING

Undergraduate Summer Research, CSB/SJU -2019

• Conducted Age-Length analysis for fish species on campus lakes
• Student presentation at the Midwest Fish and Wildlife Conference, January 27, 2020

Undergraduate Research Opportunities Program, UMD – 2012-2017

• Directed 6 undergraduate students in lab to assist with research, conduct individual research
projects and formally present results.
• Advisees: Projects
-Loranzie Rogers, McNair Scholar: Visual sensitivity of lean lake trout
-Julia Narow: Treatment of barotrauma in deepwater fishes of Lake Superior
-Sarah Muellner: Effect of light intensity on walleye growth
-Lee Austin, McNair Scholar: Foraging mechanisms of lake trout
(received the MN AFS scholarship, March 26, 2015 for his research)
-Sydney Mauch, McNair Scholar: Effect of salinity on round goby survival
-Marc Sizer: Mortality rates of deepwater fishes post-barotrauma
References:
Owen T. Gorman, Ph.D.
Bill Lamberts, Ph.D. U.S. Geological Survey
Professor and Department Chair Great Lakes Science Center
Biology Department Lake Superior Biological Station
College of St. Benedict/St. John’s University 2800 Lake Shore Drive East
Collegeville, MN 56321 Ashland, WI 54806
320-363-3160 715-682-6163
wlamberts@csbsju.edu otgorman@usgs.gov

Jennifer Schaefer, Ph.D. Jill D. Jensen, Ed.D.

Associate Professor of Biology Director, Academic Writing & Learning Center
Integrative Science Program Chair Associate Professor
Biology Department University of Minnesota, Duluth Campus
College of St. Benedict/St. John’s University 416 Library Drive
Collegeville, MN 56321 Duluth, MN 55812
320-363-3275 218-726-8242
jschaefer@csbsju.edu jjenson@d.umn.edu

Thomas R. Hrabik, Ph.D.

Sharon Moen, M.S. Professor
Senior Science Communicator Department of Biology
Minnesota Sea Grant University of Minnesota, Duluth Campus
31 West College Street 207 Swenson Science Building
Duluth, MN 55812 1035 Kirby Drive
218-726-6195 Duluth, MN 55812
smoen@d.umn.edu 218-726-7626
thrabik@umn.edu