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GREEN BELL PEPPER

Scientific Name: Capsicum annuum L.


Family: Solanaceae

Quality Characteristics green for 7–8 weeks before starting to change color from
green to red. Pericarp thickness also increased in ‘Domino’
G reen bell pepper is the most frequently used type of pepper.
The green bell pepper pod is an unripe fruit, fully
pepper fruit ripened on the plant and was positively corre-
lated with fruit firmness (Tadesse et al. 2002). At harvest,
developed in terms of size and seed viability. green peppers are generally firmer and have higher water
Although all bell peppers are green when unripe, different potential and insoluble pectins and lower water-soluble
varieties may develop red, yellow, orange, or purple color- pectins than red peppers (Lurie et al. 1986; Sethu et al.
ation during ripening. Green, red, and yellow peppers are the 1996). In addition, a high positive correlation was found
most common peppers sold at the market, followed by between the firmness of bell peppers at harvest and their
orange, purple, and other unusual colors (Frank et al. 2001). postharvest life (Janse 1989).
Bell peppers harvested before the ripe color stage (i.e., As green bell peppers mature on the plant, their sugar
green) are in general less expensive than ripe peppers content increases so they become sweeter (Sethu et al. 1996).
because on-the-plant ripening of colored peppers leads to They also develop higher nutrient content, primarily vita-
greater losses owing to decay and various other types of mins A and C (Howard et al. 2000; Luning et al. 1994;
damage. In addition, green bell peppers can better withstand Márkus et al. 1999; Osuna-García et al. 1998; Simonne et al.
transport and have a longer postharvest life than ripe colored 1997). The total soluble solids content of ‘Domino’ bell
fruit (Frank et al. 2001; Fox et al. 2005). peppers increased rapidly from 6 to 8% during fruit matura-
Although bell pepper is considered to be a non-climacteric tion and ripening (Tadesse et al. 2002). Glucose and fructose
fruit due to its very low endogenous rate of eth-ylene production contents increased significantly from the green to turning
(González-Aguilar 2004), when harvested at 10–30% stage, and subsequently to the red stage, but the highest
coloration (i.e., the “color-break” stage) and ripened off the amount of sucrose was observed at the turning stage (Luning
plant at 20°C, fruits of red and yellow bell pepper varieties et al. 1994). Along with the changes in coloration, texture,
developed similar appearance (i.e., red or yellow color) and and composition during ripening of bell peppers, sweetness,
compositional attributes (i.e., total soluble solids, acidity, and sourness, and red bell pepper aroma were reported to
pH) as those ripened on the plant (Molinari et al. 1999). increase during ripening from the green to the red stage,
Exposure of bell peppers with 10– 60% red color development whereas bitterness and grassy, green bell pepper, and
to 100 µL/L ethylene during storage at 20°C resulted in cucumber aromas decreased significantly (Luning et al.
accelerated chlorophyll break-down and increased carotenoid 1994). Overall, the best quality green bell peppers should
production after 3–6 days, compared with fruit stored in air at have firm flesh, bright skin color, and sweet flavor
the same temperature, without altering the flavor, ascorbic acid (González-Aguilar 2004).
content, or antioxi-dant capacity (Fox et al. 2005). Nonetheless, Green bell peppers contain on average 92% water, 5%
when harvested before the color-break stage and stored at 22°C, carbohydrates, 0.9% proteins, 2% fiber, 89 mg of vitamin C,
peppers did not develop a fully red color even when exogenous and 632 IU of vitamin A per 100 g of fresh fruit (USDA 2006).
ethylene was applied (Krajayklang et al. 2000). Red fruit is in general much more nutritious than green fruit
Surface color change was reported to be the major indica-tor because it contains three times more vitamin C and four times
of bell pepper maturity because of its close association with more vitamin E than green fruit. Vitamin E (α-tocopherol)
several other attributes, namely total soluble solids content and present in different pepper varieties increases during ripening,
fruit firmness (Tadesse et al. 2002). The time until full from the mature green to the fully red stages (Márkus et al.
coloration and rate of color changes on the plant are dependent 1999; Osuna-García et al. 1998). Likewise, ascorbic acid levels
on the environmental conditions during fruit development in the increase during ripening and peak at the red stage.
field, and on cultivar characteristics. For example, ‘Domino’ Consequently, ripe peppers have higher content of ascorbic acid
peppers ripened on the plant remained than green peppers do (Howard et al. 2000; Luning et al. 1994;
Márkus et al. 1999; Osuna-García et al.

265
266 COLOR ATLAS OF POSTHARVEST QUALITY OF FRUITS AND VEGETABLES

1998; Simonne et al. 1997). Carotenoid content also of water vapor inside the package (i.e., 99–100% relative
increases as the peppers change from green to deep red humidity) plus exposure to fluctuating temperatures during
(Hornero-Méndez and Mínguez-Mosquera 2000; Howard et handling and distribution may lead to condensation inside
al. 2000; Márkus et al. 1999; Simonne et al. 1997). A the package and on the fruit surface, creating a favorable
dramatic increase in the carotenoid content from 23.5-fold in environment for pathogen development (Miller et al. 1986;
‘Numex’ to 38.0-fold in ‘Nana’ peppers was observed Mohamed 1990; Rodov et al. 1995).
during ripening (Hornero-Mández and Mínguez-Mosquera
2000). Although colored peppers tend to have higher
Temperature Effects on Quality
contents of vitamin C and carotenoids than green peppers do,
green, red, and orange peppers were reported to have much Water loss, tissue softening, shriveling, and CI are the prin-
higher contents of vitamin C and carotenoids than other cipal factors that limit the quality and postharvest life of bell
unusually colored peppers such as black, purple, or white peppers (Hampshire et al. 1987; Kissinger et al. 2005;
(Simonne et al. 1997). Lownds et al. 1994; Sethu et al. 1996; Smith et al. 2006).
Because the bell pepper fruit is hollow, with a thin wall of
approximately 5–8 mm thickness, it has a reduced capacity
Optimum Postharvest Handling Conditions
to store large volumes of water for long periods. Therefore,
To avoid CI and to reduce water loss and shriveling, bell the fruit is easily subjected to loss of moisture and conse-
peppers should be pre-cooled using forced-air cooling with quently to shriveling and loss of firmness (Kissinger et al.
high relative humidity air as soon as possible after harvest to 2005). Moreover, poor postharvest handling practices may
a temperature no lower than 7°C. Pre-cooling can also be increase the fruit’s susceptibility to water loss and quality
accomplished using hydro-cooling or vacuum-cooling deterioration. Ceponis et al. (1987) reported that 70% of the
(González-Aguilar 2004). Chlorinated water treatments are bell peppers that were inspected for conformity to grade
sometimes used to remove field dirt and sanitize green bell standards upon reaching the New York market exhibited a
peppers in order to make them more attractive in the market. variety of disorders. Softening, flaccidity, shriveling, and
However, the chlorine concentration and dipping time have wilting as a result of water loss during postharvest handling
a significant effect on the postharvest quality of the fruit. combined with decay are the major problems that reduce
Green bell peppers dipped for more than 20 minutes in postharvest marketability and consumer acceptance of bell
chlorine solutions with concentrations higher than 100 g/ peppers (Janse 1989).
mL followed by storage for 1 week at 10°C showed a Temperature during handling not only affects the texture
decrease in chlorophyll, soluble solids, and ascorbic acid of bell pepper fruit but also the rate of color development.
contents (Nunes and Emond 1999). Color developed faster when fruit was stored at 15.5°C
Storage of bell peppers at 7.5°C and 90–95% relative compared with storage at 7°C (Miller et al. 1986). Bell
humidity is recommended for maximum postharvest life (i.e., pepper fruit harvested at different color stages from green to
3–5 weeks) and to reduce water loss and shriveling. When green-orange developed orange coloration after 10–15 days
stored at higher temperatures and lower relative humidity at 22°C. Conversely, the color of peppers stored at 7°C for
levels, peppers may lose water rapidly, becoming flaccid, 20 days did not change until the fruit was trans-ferred to
shriveled, and dry (González-Aguilar 2004; Lownds et al. 22°C (González et al. 2005). Storage of green bell peppers
1994). Because some cultivars may be susceptible to CI at 7°C, at temperatures between 20 and 25°C led to the development
they should be stored at temperatures between 7 and 13°C. of fully orange-red color (Meir et al. 1995). Red and yellow
However, if stored at temperatures above 13°C, peppers may bell pepper varieties harvested at the onset of color change
show accelerated ripening or senescence and decay. Green bell developed full-red or full-yellow color after approximately
peppers can also be stored at 5°C for approximately 2 weeks, 5 days at 20°C (Molinari et al. 1999). Lownds et al. (1994)
and although the low storage temperature reduces water loss, CI also reported that the rate of color changes in several
symptoms may begin to develop after that period. Symptoms of different pepper cultivars was higher during storage at 14 or
CI may develop after a few days at 0°C or a few weeks at 5°C. 20°C than at 8°C. Furthermore, the advanced color
Ripe or colored peppers are less sensitive to CI than green development at 14 or 20°C relative to 8°C paralleled differ-
peppers are (González-Aguilar 2004). ences in water loss rates and suggested a direct relationship
To delay ripening and compositional changes associated between these variables. This relationship between changes
with ripening, reduce CI, and retard softening, the use of an in color and water loss should be considered to be cultivar
adequate temperature can be supplemented by packaging the and/or season dependent rather than assumed to be a gener-
pepper in polymeric films (González and Tiznado 1993; alization for all pepper cultivars. In fact, after 14 days of
González-Aguilar et al. 1999; Meir et al. 1995; Miller et al. storage at 20°C, ‘Bell Boy’ peppers from a first harvest had
1986). If bell peppers are packed in a film with a low perme- higher rates of water loss (25%) and yellow color develop-
ability to water vapor at a temperature between 7 and 10°C, their ment than fruit from the second harvest of the same plants,
postharvest life can be extended by about a week (González- which had only 5% water loss and were not completely
Aguilar 2004). However, excessive high levels yellow until after 20 days of storage (Nunes and Emond
2002).
SOLANACEOUS AND OTHER FRUIT VEGETABLES 267

The L* value (i.e., lightness) of ‘Bell Boy’ peppers Wrapped peppers stored for 4 weeks at 7 or 15.5°C and 90–
showed a slight tendency to decrease throughout storage, 95% relative humidity softened at a rate of −0.06 and −0.18
although the changes were small. ‘Bell Boy’ green bell units per week, respectively, whereas the softening rate for
peppers stored at 20°C showed a significant increase in L* nonwrapped fruit stored at the same temperatures was −0.17
value after 20 days, reflecting the development of yellow and −0.38 units per week, respectively (Miller et al. 1986).
coloration. The hue and chroma of bell pepper fruit stored at When exposed to temperatures lower than 7.5°C, green bell
0.5, 10, and 15°C did not change significantly during peppers may develop symptoms of CI such as pitting, water-
storage, in contrast to the hue and chroma of peppers stored soaked areas, decay, discoloration of the seed cavity, and
at 20°C. Bell peppers stored at 20°C showed a hue angle of softening without water loss (Cantwell 2006; González-Aguilar
approximately 108 after 20 days, which corresponds to a 2004). Symptoms of CI appeared as surface pitting followed by
yellowish color (a hue value of 90 degrees represents pure a combination of both surface and sheet pitting after 2–3 days
yellow). Although some of the peppers stored at 20°C at 0°C (Smith et al. 2006). Likewise, holding green bell peppers
showed a marked shift from green to yellow, other fruit at temperatures in the range of 0–3°C for a 10-day period
remained green, even after 20 days at 20°C (Nunes and resulted in surface pitting and, subsequently, pathogen invasion
Emond 2002), probably reflecting variations in initial (Burzo et al. 1994). Usually, symptoms of CI are aggravated
maturity. when bell peppers are transferred to ambient temperature.
Wrapping green bell peppers in plastic films may con- Whitaker (1995) reported that approx-imately 30% of the ‘Bell
tribute to better green-color retention compared with non- Tower’ green bell peppers stored for 2 weeks at 2°C exhibited
wrapped fruit (González-Aguilar et al. 1999; Srinivasa et al. pitting on about 25% of their surface. Three days after return of
2006). For example, the hue angle of green bell peppers the chilled fruit to 20°C, 60% showed severe pitting over about
packaged in different materials gradually decreased (became 30–40% of the surface. ‘Bell Boy’ peppers showed moderate
less green and more yellow) during storage, but showed the signs of CI such as surface pitting and darkening of the seeds
lowest value in unpackaged fruits (Srinivasa et al. 2006). after approximately 13–14 days of storage at 0°C plus 1 day at
Higher L* and hue values were reported for green bell 20°C and were considered unacceptable for sale. ‘Bell Boy’
pepper wrapped in a plastic film compared to nonwrapped green bell peppers stored at 5°C showed moderate signs of CI
fruit, meaning that wrapped fruit was more bright and green after approximately 17 days plus 1 additional day at 20°C
after 28 days at 8°C than nonwrapped fruit (González- (Nunes and Emond 2002). Green bell peppers stored at 1, 4, or
Aguilar et al. 1999). 7°C for 14 days also developed CI symptoms such as pitting
A detailed assessment of bell peppers in retail stores shows and seed discoloration after transfer to 15°C for 5 days (Miller
that very often the fruit feels soft, flaccid, and shriv-eled without and Risse 1986). ‘Bell Boy’ green bell peppers stored at 10, 15,
tissue breakage. Peppers are often displayed without any kind or 20°C showed no CI symptoms during 14- or 20-day storage
of protective packaging and are thus sub-jected to adverse periods (Nunes and Emond 2002). Lin et al. (1993) reported that
conditions, which may cause shriveling and increased softening ‘Bison’ and ‘Doria’ mature-green peppers showed surface
as a result of loss of moisture. As peppers lose water they pitting after 3 days to 1 week at 1°C, but fruit stored at 13°C did
become less turgid and less firm (Showalter 1973). Losses in not show any signs of CI symptoms.
firmness of 19–21% were observed for fruit stored for 20 days
at room temperature, whereas for fruit stored for the same Although a study by Miller and Risse (1986) showed that
period at 8°C, the loss in firmness was only 1–3% (Sethu et al. film-wrapping would not alleviate CI symptoms in green bell
1996). A pro-nounced increase in fruit softening was associated peppers stored at chilling temperatures, others have shown that
with increases in weight loss during storage for 12 days at storage of bell peppers in polyethylene bags for 14 days at 3°C
approximately 15°C (Showalter 1973). A direct relationship or for 18 days at 1 or 4°C delayed the devel-opment and
was found between softening and water loss, as softening alleviated the symptoms of CI (Kosson 2003; Meir et al. 1995).
followed a pattern similar to that of water loss in peppers stored Furthermore, wrapping in a plastic film may have delayed the
at 8, 14, or 20°C (Lownds et al. 1994). Although the firmness development of CI symptoms in bell peppers stored for 28 days
of ‘Bell Boy’ green bell peppers decreased for fruit stored at 0, at 8°C compared to nonwrapped fruit (González-Aguilar et al.
5, 10, or 15°C, softening never reached unac-ceptable levels. 1999, 2000). Mature-green peppers held at 4°C in perforated or
Conversely, the firmness of the peppers stored at 20°C attained nonperforated polyeth-ylene bags did not develop CI symptoms
unacceptable levels after approxi-mately 5–18 days of storage, on the fruit surface after 18 days. In contrast, peppers stored at
depending on the harvest date. The reason for the relatively low 1°C plus 3 addi-tional days at room temperature showed initial
loss of firmness in ‘Bell Boy’ peppers stored at temperatures CI symptoms after 3, 6, and 9 days when stored without
below 20°C may be related to the high relative humidity (90– packaging, in nonperforated packaging, or in perforated
95%) that was maintained during storage (Nunes and Emond packaging, respec-tively, with the peppers packaged in
2002). In fact, when wrapped in a plastic film, green bell perforated polyethylene bags showing the lowest severity of CI
peppers stored for 28 days at 8°C maintained significantly (Kosson 2003). An increase in injury due to exposure to chilling
greater firmness than nonwrapped fruit (González-Aguilar et al. temperatures
1999).
268 COLOR ATLAS OF POSTHARVEST QUALITY OF FRUITS AND VEGETABLES

generally corresponded with increased fruit water loss. A nificantly lower (1.3 and 1.4%, respectively) than the weight
significant correlation was found between CI and percentage loss of nonwrapped fruit (9.2 and 15.4%, respectively)
water loss for green bell peppers stored for 14 days at 0°C (González-Aguilar et al. 1999, 2000). Wrapped peppers stored
followed by 2 days at 20°C (Smith et al. 2006). for 4 weeks at 7 or 15.5°C and 90–95% relative humidity lost
Softening, increased flabbiness, loss of turgidity, wilting weight at a rate of 0.06 and 0.12% per week, respectively,
and shriveling, and dryness are symptoms generally associ- whereas nonwrapped fruit lost 0.74 and 1.29% per week,
ated with loss of moisture. Lownds et al. (1994) reported that respectively (Miller et al. 1986); wrapping bell peppers
development of flaccidity in bell peppers appears to be provided a net reduction of 2.56% in weight loss, and no
directly associated with water loss. Hruschka (1977) shriveling was observed. Compared to nonwrapped fruit,
observed that when extremely severe shrivel was noted in polyethylene bag packaging reduced weight loss by 40–50% for
green bell peppers, percentage weight loss averaged 15% bell peppers stored for 2 weeks at 7.5°C fol-lowed by 3
(ranging from 8 to 27%). Furthermore, deterioration in com- additional days at 17°C (Meir et al. 1995). The weight loss in
mercial appearance accompanied moderate shriveling symp- unpackaged bell peppers stored for 16 days at approximately
toms and was noted when weight loss averaged 12% 27°C was significantly higher (14–15%) compared with fruit
(ranging from 3 to 22%) (Hruschka 1977). According to packaged using different packaging materials. Bell peppers
Robinson et al. (1975), a weight loss of about 7% should be packed in a chitosan film lost 10–11% of their initial weight
considered to be the maximum amount that is acceptable in during 16 days at room tem-perature (27°C and 65% relative
terms of saleability. Lownds et al. (1994) reported that New humidity), whereas fruit packaged in low-density polyethylene
Mexican type of peppers become flaccid in 3–5 days at film lost 2–2.5% of its weight during the same period (Srinivasa
20°C, which corresponded to a weight loss of 7–10%. et al. 2006). Bell peppers dipped in a solution of mineral-based
González and Tiznado (1993) also observed that when green oil and stored for 18 days at 10°C and 80–85% relative humidity
bell peppers lose 5% of their initial weight, signs of lost less moisture than nontreated fruit. After 18 days, coated
shriveling become evident. Green and red bell peppers held fruit had a weight loss of only 3.3% and was free from wilting
for 3 weeks at 17°C and 85% relative humidity had a weight and shriveling, whereas nontreated fruit had lost approximately
loss rate of 4% per week, which was accompanied by 10% of its weight and showed severe wilting and shriveling
decreased firmness of the fruit. Decreasing the storage (Lerdthanangkul and Krochta 1996).
temperature from 17 to 8°C resulted in a reduction of about
50% in the rate of weight loss in green and red peppers. In The increase in water loss by bell peppers during storage
peppers stored at 7°C, weight loss increased slightly from may be associated with cellular breakdown, loss of mem-
0.39% after 1 week to 1.7% after 20 days (González et al. brane integrity, and removal of epicuticular waxes, which
2005). Weight loss of ‘Bell Boy’ peppers from two different play an important role in water loss through the skin
harvests showed sig-nificant variation. Peppers from the first (González-Aguilar et al. 2000; Kissinger et al. 2005; Lownds
harvest stored at 20°C attained the maximum acceptable et al. 1993). In addition, differences in water loss when dif-
weight loss (i.e., 7%) after approximately 3 days of storage, ferent varieties of bell peppers are packed or maintained
whereas after 14 days, peppers stored at 0°C had a weight under the same storage conditions may suggest differences
loss of 8%, which was similar to those stored at 10°C (9%). in cuticle thickness, the presence of pores and/or cracks, and
After 14 days of storage, peppers stored at 5°C still had an epicuticular wax quality, which may vary from harvest to
acceptable weight loss (5%), whereas after 14 days peppers harvest (Lownds et al. 1994). The cuticle and its components
stored at 20°C had approximately 25% weight loss. (i.e., cuticle thickness, cuticular chemistry, and epicuticular
Conversely, in a second harvest, weight loss of ‘Bell Boy’ wax chemistry and distribution) are the main barriers to
green peppers stored under the same storage conditions water loss, and thus significantly affect the rate of water loss
never reached the maximum acceptable weight loss of 7%, (Kissinger et al. 2005; Lownds et al. 1993; Smith et al.
and after 14 days the fruit had only lost between 2 and 5% 2006). Water loss during storage of bell peppers at 8 or 20°C
of its weight (Nunes and Emond 2002). In addition to was significantly correlated with cultivar, initial water
decreasing the storage temperature to recommended levels, content, cell membrane ion leakage, lipoxygenase activity,
increasing the ambient humidity from 85% to a water- ratio of surface area to volume, and amount of epicuticular
saturated atmosphere reduced the weight loss of green and wax (Kissinger et al. 2005; Lownds et al. 1993). Further-
red peppers by 90%, resulting in delayed senescence more, membrane lipid content was lower in bell pepper
processes, better firmness retention, and inhibition of cell- genotypes that were more susceptible to water loss during
wall breakdown (Lurie et al. 1986). storage, whereas peppers that had lower water loss main-
Water loss may be significantly reduced by packing bell tained high levels of membrane lipid contents (Maalekuu et
peppers in plastic films (González and Tiznado 1993; al. 2006).
González-Aguilar et al. 1999, 2000; Meir et al. 1995; Miller et Concurrently with changes in the sensory attributes of bell
al. 1986; Mohamed 1990; Srinivasa et al. 2006) or by using pepper fruit, compositional changes also take place and are
edible, mineral-based oil coatings (Lerdthanangkul and Krochta greatly dependent on postharvest handling conditions. When
1996). For example, water loss in film-wrapped green bell bell pepper fruit was exposed to chilling temperatures in the
peppers stored for 14 or 28 days at 8°C was sig- range of 0–3°C, acidity increased by approximately
SOLANACEOUS AND OTHER FRUIT VEGETABLES 269

17% compared to storage at 10°C (Burzo et al. 1994). Like- postharvest life of the peppers. After a 4-day exposure to
wise, fruit acidity increased during storage at 7°C but, after 5°C, fruit transferred to 20°C for 1 day develops small pits
35 days, acidity began to decrease (González et al. 2005). on the skin. The symptoms of CI become more severe with
Ascorbic acid content of bell peppers did not show a signifi- increased exposure to 5°C, and the peppers show large pits,
cant change during 20 days at 7°C but decreased after 35 water-soaked areas, and browning of the seeds after 18 days
days (González et al. 2005). Total soluble solids concentra- at 5°C plus 1 day at 20°C (Figures 5.22 and 5.23). In some
tion did not change substantially in green bell peppers stored of the fruit the symptoms of CI are even more serious, as
in perforated polyethylene bags at 5 or 10°C for 28 days and severe water-soaking of the tissues and complete fruit col-
did not differ according to the storage temperatures lapse become evident after 10 days of storage at 5°C fol-
(Martinez et al. 2003). Titratable acidity in green bell lowed by 1 day at 20°C. After 20 days at 5°C plus 1 day at
peppers decreased by 75% between 14 and 28 days of 20°C, some fruit is also affected by decay that develops on
storage at 8°C, whereas pH and total soluble solids did not the necrotic tissue in the pits caused by exposure to low
change (González-Aguilar et al. 1999). temperature (Figure 5.24).
Visual quality of ‘Bell Boy’ peppers stored at 10°C
remains acceptable for 14–16 days (Figure 5.25). Although
Time and Temperature Effects on the Visual
after 14 days slight shriveling develops on the skin of the
Quality of ‘Bell Boy’ Green Bell Peppers
pepper, it does not increase further during the subsequent
‘Bell Boy’ green bell peppers shown in Figures 5.18–5.28 storage period. Darkening and dryness of the stem also
were harvested fully developed, green, and firm from a become evident after 14 days.
commercial operation in the Orleans Island, Quebec, ‘Bell Boy’ green bell peppers stored at 15°C maintain
Canada, during the summer season (i.e., August–Septem- acceptable visual appearance for 18 days (Figure 5.26).
ber). Promptly after harvest fresh peppers were stored at five However, after 16 days, changes in the color, such as devel-
different temperatures (0.5  0.5°C, 5.0  0.2°C, 10.0  opment of browning on the stem and slight surface yellow-
0.4°C, 15.0  0.2°C, and 20.0  0.2°C) and with 95–98% ing, render the fruit unmarketable. At this temperature, other
relative humidity. nonvisual quality factors also contribute to quality deteriora-
Changes in the visual quality of ‘Bell Boy’ green bell tion. Loss of firmness and development of an unpleasant
peppers are to a great extent dependent on the storage tem- aroma occurred after 18 days of storage.
perature. Major visual changes in peppers stored at tempera- Although ‘Bell Boy’ green bell peppers stored at 20°C
tures lower than 10°C are attributed to CI, and they increase maintain an acceptable visual appearance for 16 days (Figure
when the fruit is transferred to ambient temperatures. For fruit 5.27), after 12 days detrimental changes in the color such as
stored at temperatures above 5°C, major changes in the visual browning on the stems and slight skin shrivel-ing develop.
quality result from shriveling, darkening of the stem, or changes At this temperature, other nonvisual quality factors also
in fruit coloration. The color of ‘Bell Boy’ green bell peppers contribute to quality deterioration. Loss of firmness and
stored at 0, 5, 10, or 15°C changes from a bright green color to development of an unpleasant aroma occurred after 18 days
a more dull green color during storage (Figures 5.18, 5.21, 5.25, of storage. Remarkable changes in the color of some ‘Bell
5.26, and 5.27). However, some of the fruit, but not all, stored Boy’ green peppers from green to fully yellow are observed
at 20°C shows a marked shift in color from green to yellow after between 12 and 20 days of storage at 20°C. However, the
approximately 10 days of storage. After 20 days, some of the stem appears extremely dry and some brownish
peppers stored at 20°C are com-pletely yellow (Figure 5.28). discoloration develops on the shoulder (Figure 5.28).
Although ‘Bell Boy’ green bell peppers stored at 0°C Overall, changes in fruit and stem coloration, skin shriv-
maintain acceptable visual appearance for about 8–10 days eling, and symptoms of CI caused by exposure to cold
(Figure 5.18), after just 2–4 days at this temperature small temperatures, such as pitting, darkening of the seeds, water-
pits are visible, and after 12 days, CI symptoms render the soaking of the tissues, and decay are the most important
fruit unacceptable for sale. When transferred to 20°C for 1 visual factors that limit the postharvest life of ‘Bell Boy’
day, the symptoms of CI worsen; the pits increase in size and green bell peppers. Peppers stored at 10 or 15°C maintain
number and the seeds develop brownish coloration (Figures good quality for longer periods (16–18 days, respectively)
5.19 and 5.20). than peppers stored at either lower or higher temperatures.
‘Bell Boy’ peppers stored at 5°C maintain acceptable Peppers stored at 0, 5, or 20°C retain acceptable visual
visual appearance for approximately 16 days (Figure 5.21). quality for 2, 4, and 12 days, respectively, but quality dete-
However, after that period, development of CI limits the riorates very quickly afterward.

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