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George Paxinos Charles Watson

Prince of Wales Medical Research Institute Division of Health Sciences
The University of New South Wales Curtin University of Technology
Sydney, Australia Perth, Australia

g.paxinos@unsw.edu.au c.watson@curtin.edu.au
www.powmri.edu.au/staff/paxinos.htm

Amsterdam Boston Heidelberg London New York Oxford

Paris San Diego San Francisco Singapore Sydney Tokyo
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Elsevier Academic Press

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We dedicate this book to Kosta Theodore Paxinos and Anwen Angharad Williams.
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Contents
Preface . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . IX

Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . IX

Features of the Fifth Edition . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . X

Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . X

Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . XI

Stereotaxic reference system . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . XIII

Nomenclature and the construction of abbreviations . . . . . . . . . . . . . . . . . . . . XVI

The basis of delineation of structures . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . XVII

References. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . XXV

Index of structures. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . XXX

Index of abbreviations . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . XXXVI

Figures . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . XLIII

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Preface
In the first four editions of this atlas, we relied on a coronal section set that
had some significant limitations; the section frequency proved over time to
be too wide, sections did not always appear at regular intervals, and a few
damaged sections had been replaced with sections from another brain. The
fifth edition is based on a new coronal set which includes 161 sections from a
single brain at regular 120µm intervals. This edition of the atlas is not simply
an incremental improvement on the previous edition, but a completely new
and far more comprehensive map of the rat brain.
Although the fifth edition features a different coronal section set, readers can
be assured that the stereotaxic coordinates in the new atlas match those in
previous editions. We have increased the scope of the atlas by incorporating
new anatomical concepts where appropriate, and have once again delineated
and named some areas not previously recognised.
Over the past decade our efforts at mapping the brain have been greatly
enhanced by the availability of sections stained with a wide range of different
chemical markers. A further contribution to the accuracy of our maps has
been the knowledge we have we have gained from comparative
neuroanatomical studies. One of us (GP) has published atlases of the human
(Paxinos and Huang, 1995; Mai et al. 2004), monkey (Paxinos et al. 2000),
and mouse brain (Paxinos and Franklin, 2004), and both of us are part of
a team that is in the final stages of preparing an atlas of the chicken brain
(Puelles et al. in press). Each of these atlas projects has provided us with
new insights that have enhanced our ability to interpret the anatomy of
the rat brain.
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
Acknowledgements
We thank Hongqin Wang for outstanding technical assistance from the
inception to the completion of the project. We are indebted to Lewis Tsalis for
speed, accuracy and brilliance in construction of diagrams and designing this
book; to Paul Halasz for his imagination in construction of the CD-ROM; to
Julia Tsalis for accurate labelling of diagrams; to Hongmei Liu for technical
assistance, and to Yvette Paxinos for the cover design.
We acknowledge with gratitude the intellectual contribution to delineations
made by Yuri Koutcherov (hypothalamus), Konrad Talbot (hippocampus),
Nicola Palomero-Gallagher and Karl Zilles (cortex), Brent Vogt (cingulate
cortex), Jan Vogt (cerebellum, precerebellar nuclei and vestibular nuclei),
George Alheid (basal forbrain), Pascal Carrive (periaqueductal gray and
hypothalamus), Glenda Halliday (substantia nigra and VTA), Ellen Covey and
Manolo Malmierca (auditory system), Joel Elmquist (hypothalamus), Ann
Goodchild and David Hopkins (rostroventrolateral and caudoventrolateral
medulla), Jose DeOlmos (amygdala), Henk Groenewegen (thalamus), Joseph
Travers (orofacial motor nuclei), John Mitrofanis (zona incerta), Pierre-Yves
Risold (septum), Miklos Palkovits (paralemniscal nuclei), Harvey Karten
(pretectal area), Chip Gerfen (basal ganglia), Terry Furlong (hypothalamus),
Jean Buettner-Ennever (oculomotor nuclei), Marina Bentivoglio
(parafascicular nucleus).
George Paxinos acknowledges the support he has received from the Australian
National Health and Medical Research Council (he holds an NHMRC
Principal Research Fellowship), as well as assistance from the Clive and Vera
Ramaciotti Foundation, the Rebecca Cooper Foundation, and the Brennan
Foundation.
We have appreciated the intelligent and enthusiastic support from our
Elsevier editor Johannes Menzel. His patience and consideration have made
a real difference to the successful completion of this project. We also thank
Maureen Twaig and other Elsevier staff for their willingness to help in solving
production problems.
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Features of the Fifth Edition
• 161 coronal diagrams based on a single brain
• Diagrams spaced at constant 120 µm intervals giving scientists the most
comprehensive and convenient atlas of the rat brain
• The most accurate stereotaxic reference system available
• Outlines of figures and brain structures in blue, but labels and leader lines
in black for increased clarity of delineations
• All delineations re-examined in the light of recent findings
• Delineations of brain structures have been made with reference to sections
stained for Nissl substance, AChE, parvalbumin, calbindin, calretinin,
SMI-32, tyrosine hydroxylase, and NADPH diaphorase (Paxinos et al.
1999a,b)
• Extensive use was made of reference works, including the third edition
of The Rat Nervous System (Paxinos, 2004) and other recent
neuroanatomical literature
• Spinal cord drawings from the atlas of Molander and Grant(1995)
• Diagrams available on CD-ROM for printing.
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
Introduction
There are many reasons why the rat is the most commonly selected subject for
research in mammalian neuroscience. First, rats are the right size: neither too
small for accurate stereotaxic localization of discrete brain areas, nor too large
for cost-effective laboratory management. Second, rats are generally hardy
animals and are resistant to infections. Third, a number of inbred strains are
available commercially, so that animals of consistent size can be used for
stereotaxic procedures.
When the first edition of The Rat Brain in Stereotaxic Coordinates was
published in 1982, it was the first atlas to be based on the flat skull position.
It offered a choice of bregma, lambda, or the midpoint of the interaural line
as the reference point. Although the coordinates were developed from study
of adult male Wistar rats with weights ranging from 270 to 310 g, the atlas
can be successfully used with male or female rats, with weights ranging from
250 to 350 g (Paxinos et al., 1985).
With each new edition of the atlas, we have attempted to improve the
accuracy of our delineations and have incorporated new findings on brain
anatomy. However, our work has been hampered by the fact that our original
series of coronal sections suffered from a number of limitations. First of all,
our primary sections series showed sections at 0.5 mm intervals, which is
insufficient to adequately represent all major structures in the brain for
modern research purposes. Although we later attempted to better illustrate
some areas by using some intervening sections, we could never fully
compensate for the wide section interval in the primary series. In addition,
we lost some sections in some areas of the brain and were forced to interpolate
sections from another brain to compensate for the missing sections.
We were aware that the only real solution to these problems was to replace the
coronal section series with a new section set based on shorter intervals and
with all sections taken from the one brain. The new coronal section series is
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presented in the present (fifth) edition of the atlas. It shows diagrams of
sections taken at regular intervals of 0.12 mm. Having constant intervals
between the sections shown in the atlas diagrams eliminates one of the
annoying features of many brain atlases – the fact that when the reader turns
a page they do not know how far they have advanced along the prime axis.
All sections are from the one brain.
The sections in our new coronal series were stained with cresyl violet or
with methods to demonstrate AChE or NADPH diaphorase because we found
that these three methods were compatible with using fresh (unfixed) tissue,
a requirement for deriving an accurate stereotaxic grid. However, we
consistently used other markers to confirm our delineations (Paxinos et al.,
1999a, Paxinos et al., 1999b).
We have once again been greatly assisted by the suggestions of many
colleagues in the delineation of structures. We welcome further advice that
might improve the accuracy of our diagrams in the future. Please email us
on g.paxinos@unsw.edu.au or c.watson@curtin.edu.au.
The present book will be followed by a comprehensive publication, which will
include accompanying photographs and revised diagrams of sagittal and
horizontal sections.
Methods
A fresh brain from a male 290 g Wistar rat was frozen, and coronal sections
were cut at 40 mm thickness. The sections were cut at right angles to the
horizontal plane joining bregma and lambda.
Stereotaxic surgery
We placed an anesthetized rats in a Kopf small-animal stereotaxic instrument,
and the incisor bar was adjusted until the heights of lambda and bregma were
equal. This flat-skull position was achieved when the incisor bar was lowered
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
3.3 ± 0.4 mm below horizontal zero (Table 1). Because the point of
intersection of the lambdoid and sagittal sutures is variable, we have chosen
to define lambda as the midpoint of the curve of best fit along the lambdoid
suture (see skull diagram). This redefined reference point is considerably
more reliable than the true lambda (the point of intersection of the sagittal
and lambdoid sutures), and it is located 0.3 ± 0.3 mm anterior to the
interaural line. We defined bregma as the point of intersection of the sagittal
suture with the curve of best fit along the coronal suture. When the two sides
of the coronal suture meet the sagittal suture at different points, bregma
usually falls midway between the two junctions. The anteroposterior position
of bregma was 9.1 ± 0.3 mm anterior to the coronal plane passing through
the interaural line, but for the brain represented in this atlas bregma is
deemed to lie at 9.0 mm. The top of the skull at bregma and lambda was
10.0 ± 0.2 mm dorsal to the interaural zero plane. To confirm the stereotaxic
orientation of sections in the brain used for this atlas, reference needle tracks
were made perpendicular to the horizontal and coronal planes. One
horizontal needle insertions perpendicular to the coronal plane were made
from the posterior of the brain at 4.0 mm above the interaural line and was
2.0 mm lateral to the midline. The reference track from the horizontal needle
appears as a small hole in coronal sections.
Following surgery, the rat was decapitated and the whole head frozen on dry
ice. The frozen skull was then prised off the frozen brain, and the brain was
carefully mounted on the stage of microtome so that the sections would be
cut in the coronal stereotaxic plane.
Every third section was used for preparation of the atlas diagrams, so that the
interval between atlas diagrams is 0.12 mm. Exceptions to this rule are found
in the region rostral to the rostrum of the corpus collosum (Interaural AP
11.28) and in the region of the medulla caudal to the inferior olive (Interaural
AP -5.76 mm). In these two regions, sections were selected for presentation in
the atlas at 0.24 mm intervals. Finally, the olfactory bulb is depicted at only
three representative levels.
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Histological methods
The ‘atlas’ sections were stained with either cresyl violet or for the
demonstration of AChE on an alternate basis, so that cresyl violet sections are
0.24 mm apart and AChE sections are also 0.24 mm apart. The two sections
that intervene between each ‘atlas’ section were stained with cresyl violet or
for the presence or AChE or NADPH diphorase, according to the following
sequence:
1. Cresyl violet ‘atlas’ section
2. AChE intervening section
3. NADPH intervening section
4. AChE ‘atlas’ section
5. Cresyl violet intervening section
6. NADPH diaphorase intervening sections
This sequence was repeated throughout the series of coronal section. This
arrangement ensures that every ‘atlas’ section is accompanied by two adjacent
sections, each of a different stain. For example, the ‘atlas’ AChE section
described as number four above is preceded by an NADPH diaphorase
section and followed by a cresyl violet section. This arrangement gave us
maximum information for each ‘atlas’ section from the three stains. Staining
was carried out on the same day as section cutting.
All sections, whether ‘atlas’ or intervening, were photographed on 4"x5" black
and white negatives and printed on 36"x24" photographic paper. Each ‘atlas’
section was then covered with a sheet of ‘Mylar’ tracing film and outlines of
structures were drawn in pencil. The final pencil drawings were scanned and
then digitized using Adobe Illustrator.
Quality of Sections
In some cases, the sections were slightly stretched or compressed in the
process of cutting and mounting on slides. We have compensated for this by
constructing diagrams which represent, as best we can judge from the study
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
of adjacent sections, the original shape of the brain section. In the worst cases,
the ‘atlas’ section was so badly damaged that we have taken our drawing from
an adjacent section.
Cresyl Violet Staining
Slides were immersed for 5 min in each of the following: xylene, xylene, 100%
alcohol, 100% alcohol, 95% alcohol, and 70% alcohol. They were dipped in
distilled water and stained in 0.5% cresyl violet for 15-30 min. They were
differentiated in water for 3-5 min and then dehydrated through 70% alcohol,
95% alcohol, 100% alcohol, and 100% alcohol. They were then put in xylene
and coverslipped.
To make 500 mL of 0.5% cresyl violet of about pH 3.9, mix 2.5 g of cresylecht
violet (Chroma Gesellschaft, Postfach 11 10, D-73257, Kongen, Germany, Fax
number: 49-7024-82660), 300 mL of water, 30 mL of 1.0 M sodium acetate
(13.6 g of granular sodium acetate in 92 mL of water), and 170 mL of 1.0 M
acetic acid (29 mL of glacial acetic acid added to 471 mL of water). Mix this
solution for at least 7 days on a magnetic stirrer, then filter.
AChE Histochemistry
The method for the demonstration of AChE followed the procedures of
Koelle and Friedenwald (1949) and Lewis (1961). Slides were incubated for
15 h in 100 mL of stock solution (see below) to which had been added
116 mg of S-acetylthiocholine iodide and 3.0 mg ethopropazine (May &
Baker). The slides were rinsed with tap water and developed for 10 min in
1% sodium sulphide (1.0 g in 100 mL of water) at pH 7.5. They were then
rinsed with water and immersed in 4% paraformaldehyde in phosphate
buffer for 8 h, and then allowed to dry. Subsequently, they were dehydrated
for 5 min in 100% alcohol, then immersed in xylene and coverslipped with
Permount. The stock solution was a 50 mM sodium acetate buffer at pH 5.0
which was made 4.0 mM with respect to copper sulphate and 16 mM with
respect to glycine. This was done by adding 6.8 g of sodium acetate, 1.0 g of
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copper sulphate crystals, and 1.2 g of glycine to 1.0 L of water and lowering
the pH to 5.0 with HCl. We found that fresh, unfixed tissue from the frozen
brains showed a substantially stronger reaction for both stains than tissue
fixed with formalin, paraformaldehyde, glutaraldehyde, or alcohol.
NADPH diaphorase
The sections were washed in phosphate buffer for 10 minutes and incubated
in 10 ml of a phosphate buffer solution containing 0.0125% nitroblue
tetrazolium, 0.05% NADPH, 0.5% Triton X-100, and 1 mM magnesium
chloride. The pH of the solution was adjusted to 7.6. The sections were
incubated at 4ºC for 48 hours. The incubation was stopped with a wash in
phosphate buffer.
Photography and drawings
Photography
The photographs of stained brain sections were taken with a Nikon Multiplot
macrophotographic apparatus using 4"x5" Kodax Plus X film. High contrast
paper was used to print the photographs of Nissl sections, whereas lower
contrast paper was used to print the photographs of AChE and NADPH
sections.
Drawings
Drawings, which later formed the basis of the figures, were made by tracing
the photographs of sections. We drew only the right side of each section and
derived the outline of structures on the left side by mirror image construction
using Adobe Illustrator.
Fiber tracts in the drawings are outlined by solid lines, and nuclei and cell
groups are outlined by broken lines. In general, each abbreviation is placed in
the center of the structure to which it relates; where this is not possible, the
abbreviation is placed alongside the structure and a leader line is used. The
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
abbreviations for fiber tracts and fissures are almost always positioned on the
left side of the figure, and the abbreviations for nuclei and other cell groups
are generally positioned on the right side. The outlines of the ventricles and
aqueduct are filled in with solid color.
Stereotaxic Reference System
The stereotaxic reference system is based on the flat skull position, in which
bregma and lambda lie in the same coronal plane. Two coronal and two
horizontal zero-reference planes are referred to in these drawings. One
reference coronal plane cuts through bregma and the other cuts through the
interaural line. Similarly, one horizontal plane is at the level of bregma on the
top of the skull and the other is at the level of the interaural line. Lambda is
usually located 0.3 mm anterior to the interaural line, and it can be used as
an alternative reference point in conjunction with the dorsoventral
coordinate of bregma. The position of the stereotaxic reference points and
planes are indicated on the skull diagram. The stereotaxic reference grid
shows 0.2 mm intervals.
Drawings of coronal brain sections
In each of the coronal drawings, the large number at the bottom left shows
the anteroposterior distance of the section from the vertical coronal plane
passing through the interaural line. The large number at bottom right shows
the anteroposterior distance of the plate from a vertical coronal plane passing
through bregma. Note that these two coronal planes are 10 mm apart, so the
two numbers on any one plate add up to 10 mm. The small numbers on the
left margin show the dorsoventral distance from the horizontal plane passing
through the interaural line. The numbers on the right margin show the
dorsoventral distance from the horizontal plane passing through bregma and
lambda on the surface of the skull. The numbers on the top and bottom
margins show the distance of structures from the midline sagittal plane.
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290 g Male Wistar

Bregma Lambda

Interaural Line
9.0 mm

Bregma Lambda

10.0 mm

3.3 mm
Incisor Bar

Skull Diagram Dorsal and lateral views of the skull of a 290 g Wistar rat. The positions of bregma, lambda and the plane
of the interaural line are shown above the lateral view. The distance between the horizontal plane passing through the
interaural line is shown on the right of the lateral view. The distance between the incisor bar and the horizontal plane
passing through the interaural line is shown on the left of the lateral view. Lambda (midpoint of the curve of best fit
along the lambdoid suture) is 0.3 mm anterior to the coronal plane passing through the interaural line.

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Accuracy of the stereotaxic coordinates
In almost all cases, the potential error in defining the position of any point
in the brain is less than 0.5 mm. Although we used medium-sized (average
290 g) male Wistar rats in the construction of this atlas, we recognize that
researchers often use animals of different sex, strain, and weight. Because
of this, we have estimated the error that may occur if this atlas is used with
female Wistar rats, male hooded (Long Evans) rats, male Sprague Dawley rats
of 300-g weight, juvenile (180 g) Wistar rats, and mature (436 g) Wistar rats.
The results of these estimations are shown in Table 1 (reproduced from
Paxinos et al., 1985).
It is evident from these studies that no substantial stereotaxic error will occur
when rats of different sex and strain are chosen, provided that the rats are of
similar weight to those on which the atlas is based (290 g). For example, for
rats of different sex and strain but of similar weight, the anteroposterior
distance between the interaural line and bregma is between 9.0 and 9.4 mm.
Similarly, the dorsoventral distance between the interaural line and the
surface of the skull at bregma and lambda is very stable (9.8-10.1 mm).
By contrast, craniometric data for juvenile (180 g) and mature (436 g) Wistar
rats differ substantially from those of other groups. The anteroposterior
distance between the interaural line and bregma is 7.7 mm in the juvenile

Table 1 Craniometric and stereotaxic data (means + S.D.) for rats of different sex, strain and weight

Mean AP AP DV AP AP AP DV
weight I–B I–L I–B I – Acb B – ac I – 7n I – incisor bar
Subject (g)* (mm) (mm) (mm) (mm)** (mm)** (mm)** (mm)

‘Atlas’ Wistar 290 9.1 ± 0.3 0.3 ± 0.3 10.0 ± 0.2 11.7 0.0 -1.3 -3.3 ± 0.4
Coronal plates 300 9.2 0.2 10.1
Sagittal plates 270 8.9 0.0 10.0
Horizontal plates 290 9.1 0.2 10.1
Female Wistar 282 9.3 ± 0.2 0.5 ± 0.3 10.0 ± 0.1 11.6 0.1 -1.2 -3.2 ± 0.5
Hooded 290 9.4 ± 0.4 0.3 ± 0.6 9.8 ± 0.2 11.9 0.0 -1.2 -3.9 ± 0.6
Sprague 299 9.0 ± 0.2 0.7 ± 0.2 10.1 ± 0.1 11.7 0.1 -1.2 -3.9 ± 0.5
Juvenile Wistar 180 7.7 ± 0.4 -0.4 ± 0.3 9.9 ± 0.2 10.2 -0.1 -1.6 -2.0 ± 0.4
Mature Wistar 436 9.7 ± 0.3 0.6 ± 0.3 10.7 ± 0.4 12.4 -0.1 -0.8 -2.7 ± 0.3

* S.D.s ≤ 20g.
** S.D.s ≤ 0.4 mm.
ac, anterior commissure; Acb, accumbens nucleus; AP, anterior-posterior; B, bregma; DV, dorsal-ventral; 7n, facial nerve; I, interaural line; L, lambda.
Reprinted with permission from J. Neuroscience Methods. 13 (1985) 139-143.

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and 9.7 mm in the mature rats (9.0 mm in 290-g male rats). Lambda is
0.4 mm posterior to the interaural line in the juvenile rats and 0.6 mm
anterior to this line in the mature rats (0.3 mm anterior in 290-g rats).
Unexpectedly, the dorsoventral distance between the interaural line and
bregma for juvenile rats (9.9 mm) was almost the same as that of 290-g rats
(10.0 mm). In the mature rats, the interaural line to bregma vertical distance
was 10.7 mm. In female rats, as well as in hooded, juvenile (180 g), mature
(436 g) and 290-g Wistar rats, bregma was found to be above the most
forward crossing fibers of the anterior commissure. This is the point at which
the posterior limbs of the anterior commissure appear. These data confirm
the observation of Whishaw et al. (1977) that bregma is more stable than the
interaural line for positioning of electrodes in brain structures close to, or
anterior to, bregma. However, data from insertion of needles aimed at the
level where the facial nerve leaves the facial genu show that the interaural
reference point is more stable than bregma for localization of such posterior
structures. Therefore, if juvenile or mature rats are used, greater accuracy can
be achieved if bregma is used as the reference point for work with rostral
structures and the interaural line for work with caudal structures.
A further improvement in accuracy can be obtained by taking into account
the actual location of the accumbens nucleus and the genu of the facial nerve.
In agreement with Slotnick and Brown (1980), we noticed that coordinates of
structures were closer to target if the coordinates given by the interaural and
bregma reference systems were averaged.
No atlas or stereotaxic instrument will compensate for using bregma and
lambdoid points inappropriately. These reference skull marks for bregma is
the midpoint of the curve of best fit along the coronal suture, and the
reference skull mark for lambda is the midpoint of the curve of best fit along
the lambdoid suture. These two reference marks are not necessarily the points
of intersection of these sutures with the midline suture.
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
Nomenclature and the construction
of abbreviations
The need for a stable neuroanatomical nomenclature to accurately and
efficiently convey information between neuroscientists is obvious. However,
many terms or abbreviations are still used to describe a single structure, and,
in some cases, the same term or abbreviation is used for completely different
structures. We urge all researchers to consider the merits of our system of
nomenclature because it is systematic and derived after extensive
consultations with neuroanatomy experts.
In considering the merit of a particular name over synonyms, we have chosen
terms that have been ratified by modern usage, particularly usage by experts
in that field. We have used anglicized versions of terms rather than older
latinized versions wherever possible, and we have in all but a handful of cases
avoided the use of eponyms.
Neuroscience communities concerned with different systems have
developed identical abbreviations for completely different structures; for
example SO may stand for both supraoptic nucleus and superior olive,
SC for suprachiasmatic nucleus and superior colliculus, and IC for inferior
colliculus and internal capsule. In dealing with the entire nervous system
(as increasingly more researchers do) these parochial abbreviation schemes
are serious obstacles to communication of data. An additional complication
arises when homologous structures are nonetheless named or abbreviated
differently in different species. We have made an effort to establish
homologies and are using the same abbreviations for homologous structures
in atlases of the rat (Paxinos and Watson, 1986), mouse (Franklin and
Paxinos, 1996), monkey (Paxinos et al., 2000), human (Paxinos et al., 1990;
Mai et al., 2004), and chicken (Puelles et al., in press).
The importance of following a logical system of abbreviations is shown by
the fact that a term such as the accumbens nucleus can be (and probably has
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been) abbreviated about 20 different ways. We used the abbreviation Acb in
this as well as all our other atlases. We have adopted the same rule for other
all other structures, and so we have maintained the same abbreviation across
mammalian and avian species, homologies permitting.
Our abbreviations have been constructed on the basis of the following
principles:
1. The abbreviations represent the order of words as spoken in English
(e.g., DLG = dorsal lateral geniculate nucleus) rather than the order in
which they appear in older latinised terms.
2. The general principle used in the abbreviations of the names of elements
in the periodic table was followed: the capital letter representing the first
letter of a word in a nucleus is followed by a lower case letter most
characteristic of that word (not necessarily the second letter;
e.g., Mg = magnesium; Rt = reticular thalamic nucleus).
3. All nuclei and other cell groupings (such as cortical areas) begin with a
capital letter, except some cranial nerve nuclei which begin with a number
followed by capital N (for ‘nucleus’). All fibre bundles begin with lower
case letters except some cranial nerves which begin with a number
followed by lower case n (for ‘nerve’). Thus, there is no necessity in for
the letter “N” to been used to point out that a structure is a nucleus, except
in the case of some cranial nerves. Similarly, there is no need for the letter
“t” to be used to denote a fiber tracts.
4. Compound names of nuclei have a capital letter for each part
(e.g., LPGi = lateral paragigantocellular nucleus.
5. If a word occurs in the names of a number of structures, it is almost
always given the same abbreviation (e.g., Rt = reticular thalamic nucleus;
RtTg = reticulotegmental nucleus of the pons). Exceptions to this rule are
made for well-established abbreviations such as VTA.
6. Abbreviations of brain regions are omitted where the identity of the
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
region in question is clear from its position (CMn = centromedian
thalamic nucleus; not CMnTh).
7. Arabic numerals are used instead of Roman numerals in identifying
cranial nerves and nuclei (as in the Berman, 1968, atlas), layers of the
cortex, and layers of the spinal cord. While the spoken meaning is the
same, the detection threshold is lower, ambiguity is reduced, and they
are easier to position in small spaces available on diagrams.
To assist in the recognition of brain structures, the labels for cell groups are
placed on the right hand side of the diagram, and the labels for fibre bundles
and nerves are placed on the left hand side of the diagram. However, where
structures are crowded, we have placed some cell group labels on the left hand
side for reasons of clarity.
The basis of delineation of structures
For the fifth edition, we completely reviewed our delineations of all areas of
the brain. Our primary guide was an extensive collection of histochemically
stained sections (monoclonal antibodies and enzyme-based stains - Paxinos
et al., 1999a,b). We have also made extensive use of other publications from
our laboratory (Paxinos, 1995; Paxinos and Huang, 1995; Paxinos et al.,
1994), the atlas prepared by Swanson (2004), as well as many authoritative
studies published in major journals such as the Journal of Comparative
Neurology.
We present below a brief account of the basis of delineation of structures.
We have not repeated here the rationale for the delineation of structures
presented in the second edition (Paxinos and Watson, 1996). Readers will be
better able to judge the suitability of our delineations when our photographs
are published as part of the comprehensive (three cardinal planes) edition of
this atlas in 2005.
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Olfactory System
Refer to Shipley et al. (2004) and de Olmos et al. (1978) for a general
description of the olfactory system.
Intermediate endopiriform nucleus (IEn)
We have given this name to an area ventral to the dorsal endopiriform
nucleus (DEn). Both DEn and IEn are deep to the piriform cortex. We
previously included this area in DEn (RBSC4); Swanson sometimes calls
it DEn and sometimes includes it in the deep layer of the piriform cortex.
The cells in this area are relatively sparse and smaller than those in DEn.
Both DEn and IEn are NADPH positive (Fig 88 Paxinos et al., 1999a),
but IEn has parvalbumin positive elements (Fig 89 Paxinos et al., 1999a).
Basal Ganglia and Basal Forebrain
Refer to Heimer et al. (1995) and Gerfen (2004) for a general description
of the basal ganglia and to de Olmos et al. (2004) for a discussion of the
substantia innominata and extended amygdala. Immunoreactivity for
parvalbumin and the neurofilament protein SMI-32 identifies the ventral
pallidum (Paxinos et al.,1999a). We retained the term substantia innominata
and identified dorsal, ventral (as in Grove, 1988), and basal components with
the assistance of George Alheid. The basal component is marked by some
positivity in tyrosine hydroxylase but is negative for SMI-32 (although
surrounding areas are positive).
The concept of ventral pallidum, first proposed by Heimer and his associates,
has been the guiding principle for structure/function relations of the basal
forebrain (Barragan and Ferreyra-Moyano, 1995; Heimer et al., 1997).
The researchers at the University of Virginia and Universidad Nacional de
Cordoba have carved out of the substantia inominata another big territory,
the sublenticular extended amygdala (Alheid et al., 1995). Paxinos and
Franklin (2001) have used the new scheme for their mouse brain atlas. We
retained the name substantia inominata (for the part remaining after the
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
territory of the ventral pallidum has been defined) in keeping with earlier
editions of this atlas. The dorsal substantia inominata roughly corresponds to
the sublenticular extended amygdala, central part, while the ventral substantia
inominata corresponds to the sublenticular extended amygdala, medial part.
The area previously called fundus striati resembles the striatum proper in
some respects and the accumbens shell in others. Given that the use of the
term fundus striati creates problems with primate homologues, we followed
the advice of George Alheid and called it the lateral accumbens shell. The
remaining accumbens is delineated in accordance with Zaborszky et al.
(1985) and Heimer et al. (1991). We followed de Olmos et al. (1995) in the
identification of the interstitial nucleus of the posterior limb of the anterior
commissure (IPAC).
Substantia nigra
We used the primate terminology for the dorsal and ventral tiers of the
substantia nigra and also for the part of the VTA which is called the
parabrachial pigmented nucleus. The names of the primate SN subdivisions
were developed earlier than those subsequently used in rodent studies, and the
identified primate subdivisions are consistent with the degeneration patterns
seen in Parkinson’s disease (Glenda Halliday, personal communication, 2004).
The reticular part of the substantia nigra can be divided into a ventrolateral
and a dorsomedial component on the basis of parvalbumin and calbindin
distribution (Paxinos et al., 1999b). The remainder of the substantia nigra
and the ventral tegmental area were delineated according to the work of
McRitchie et al. (1996).
Navicular nucleus (Nv)
We have renamed the area in the basal forebrain which we previously called
the semilunar nucleus. The reason is that the name semilunar nucleus has
priority in the avian literature and it refers to a completely different structure
in birds. The existence of the semilunar nucleus was established on the
basis of NADPH-diaphorase histochemistry (Paxinos et al.,1999a).
We acknowledge assistance of R. Harlan and P-Y. Wang in the identification
of this structure (Ahima and Harlan, 1990; Wang and Zhang, 1995).
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Globus pallidus external and internal parts (GPE, GPI)
We have used the terms internal and external instead of medial and lateral
in relation to the parts of globus pallidus to be consistent with the primate
literature.
Lateral stripe of the striatum (LSS)
LSS is a dense band of cells in Nissl stained sections. The area is negative in
calbindin sections and lighter stained than the LAcbSh and striatum in TH
(the distinction is very clear in Fig. 87 of Paxinos et al., 1999a).
Dorsal and ventral parts of the claustrum (DCl and VCl)
We have identified distinct dorsal and ventral parts of the claustrum in AChE
stained sections. The dorsal part is positive for AChE and the ventral part is
negative.
Ventral Tegmental Area
We have identified the rat homologue of the human parapeduncular nucleus
(Paxinos & Huang, 1995) but have named it the parainterfascicular nucleus.
We suggest that the new term be also used for the human given its more
descriptive nature. With the identification of the parainterfascicular in the rat,
the entire VTA (in most levels) is represented by specifically name component
parts. This avoids the problem of previous editions of this atlas where the
label VTA was placed only on what we now call parainterfascicular nucleus,
giving the impression that it alone was the VTA. The VTA in our view consists
of the paranigral, the parainterfascicular, the parabrachial pigmented nuclei
and the VTAR.
Septum, Hypothalamus, and Neurosecretory Nuclei
Refer to Simerly (2004), Armstrong (2004), Risold (2004), and Oldfield
and McKinley (2004) for a general description of these structures. Jutting
ventrolaterally from the anterodorsal preoptic nucleus is a strip which is
negative for parvalbumin which we have called the alar nucleus. The alar
nucleus displays substance P positive cell bodies but little reactivity in its
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
neuropil (Larsen, 1992). In the preoptic area we followed Simerly (2004) and
Simerly et al. (1984) except for the identification of the ventromedial and
ventrolateral preoptic nuclei, for which we followed Elmquist et al. (1996)
and Sherin et al. (1996). The compact part of the medial preoptic nucleus is
negative for substance P (Harding et al., 2004).
In the lateral hypothalamus we identified a ventrolateral hypothalamic
nucleus on the basis of NADPH-diaphorase reactivity (Paxinos et al., 1999a).
This nucleus is caudal to the ventrolateral preoptic nucleus and dorsal to the
supraoptic nucleus. The ventral part of dorsomedial nucleus is marked by
densely stained cell bodies and terminals in NADPHdiaphorase preparations
(Paxinos et al., 1999a). The gemini nucleus is a conspicuous nest of a
NADPH-diaphorase cell bodies (Paxinos et al., 1999a). The parasubthalamic
nucleus is present in the rat (Wang and Zhang, 1995), but it is not as impressive
as the homologous structure seen in the mouse brain (Paxinos and Franklin,
2001). The arcuate nucleus was delineated according to the work of Magoul
et al. (1994). See Paxinos and Watson (1986) for the identification of the
striohypothalamic, magnocellular lateral hypothalamic, terete, and
subincertal nuclei.
Lateral hypothalamus (PLH, TuLH, PeFLH, JPLH)
We have given names to different regions that are now recognized as
comprising the lateral hypothalamus. The features characteristic of lateral
hypothalamus (particularly the population of large cells) are not limited to
the area lateral to the fornix. The orexin and hypocretin containing cells are
not confined to the classically defined LH area, but are also found medial to
the fornix. Swanson (2004) correctly extended the lateral hypothalamus
medial to the fornix, but still identified the fornix as another boundary from
where various areas emanate. The region both medial and lateral to the fornix
when stimulated electrically induces attack by a cat on a rat (Paxinos, Bandler
and Flynn, unpublished observations), indicating that the PeFLH behaves in a
unitary fashion as it concerns this behavior. The components of the lateral
hypothalamus in our scheme are as follows.
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PLH – ‘peduncular part of the lateral hypothalamus’
TuLH – ‘tuberal part of the lateral hypothalamus’
PeFLH – perifornical part of the lateral hypothalamus’
JPLH – justaparaventricular part of the lateral hypothalamus’
Posterior hypothalamus, dorsal area (PHD)
This is an area previously identified as PHA in our atlas.
Arcuate nucleus (ArcMP)
ArcMP is ACHE positive and this distinguishes it from DM
Episupraoptic nucleus (ESO)
We named this nucleus on the basis of its location. Its rostral pole begins at
the caudal pole of the ventrolateral preoptic nucleus; ESO can be found in
Figs 39-45.
Paraterate nucleus (PTe)
This nucleus is located within the ventrolateral hypothalamic tract (Swanson,
2004) and rostral, dorsal and lateral to the terate hypothalamic nucleus.
Amygdala and Bed Nucleus of Stria Terminalis
Refer to de Olmos et al. (2004) for a general description of the amygdala and
the bed nucleus of the stria terminalis. The anterodorsal part of the medial
nucleus of the amygdala and the basomedial nucleus are defined by the
presence of intense NADPH-diaphorase reactivity (Paxinos et al., 1999a). The
lateral part of the central nucleus of the amygdala is marked by the presence
of tyrosine hydroxylase fibers and AChE negativity (Paxinos et al., 1999a).
Reticulostrial nucleus (RtSt)
We have named this nucleus for its postion between the stria terminalis and
the reticular nucleus. In calretinin stained sections it has a densely positive
neuropil whereas the reticular nucleus has a pale neuropil. In parvalbumin
sections, RtSt is positive in neuropil while the reticular nucleus is positive for
cells and neuropil (streaky and spotty). RtSt is largest at anterior pole of the
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
thalamus. In calbindin sections, the stria terminalis is positive whereas the
RtSt is negative (Fig 160, Paxinos et al., 1999a). Its medial part is negative and
lateral is positive in parvalbumin (Fig 166, Paxinos et al., 1999a).
Rostral amygdalopiriform area (RAPir)
The rostral amygdalopiriform area is a distinct region between PLCo and Pir
has a dense layer 2 in lateral two thirds but much less dense in medial third.
We had outlined but not labeled this structure in previous editions of our
atlas. This is the area which Swanson (2004) calls the posterior amygdalo
cortical area, a term which we did not adopt because it does not fit well with
the names we had already given to surrounding areas.
Thalamus
Refer to Groenewegen and Witter (2004) for a general description of thalamic
nuclei. See Paxinos and Watson (1986) for the identification of the ethmoid,
retroethmoid, subgeniculate, and precommissural nuclei. We have reverted to
the use of the term ventral posterior nucleus, parvicellular part (Paxinos and
Watson, 1982) for the nucleus that we previously named the gustatory
nucleus of the thalamus (Paxinos and Watson, 1986). We made this change
on the advice of Clifford Saper that gustatory input is more medial in this
nucleus, and autonomic-related inputs can be found at more lateral parts
of this structure (Yasui et al., 1989).
Retroreuniens area (RRe)
We gave the name RRe to a region dorsal to the PH, ventral to CM, and medial
to VPPC and SPF. Caudally, RRe merges with the periventricular gray matter.
Paraxiphoid (PaXi)
The paraxiphoid nucleus (PaXi) lies between the xiphoid nucleus (Xi) of the
thalamus medially and the zona incerta laterally. It appears to be part of a belt
separating hypothalamus from thalamus. In Fig 264 of Paxinos et al., 1999a,
an NADPH positive belt can be seen to extend from the reticular nucleus to
zona incerta and paraxiphoid.
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Ventral limitans thalamic nucleus (Vli)
This nucleus is a thin sheet between subparafascicular, parvicellular part and
the medial lemniscus. We so named it to complement the posterior limitans
thalamic nucleus. Palkovits has observed CGRP positivity in this nucleus
(personal communications, 2004).
Hippocampal Region
Refer to Witter and Amaral (2004) for a general description of the
hippocampal region.
We now distinguish a dorsal and a ventral subiculum. We have labeled the
transition area of dorsal and ventral subiculum as STr. We have drawn the
borders of the presubiculum and parasubiculum so as to reach the white
matter as explained in Haug (1976) and Mulders et al. (1997).
The entorhinal parcellation scheme of Insausti et al. (1997) is appealing
because each of the cytoarchitecturally distinct divisions has a different
pattern of connections as they detail in their paper. The parcellation
recognizes that the medial and lateral sectors of the entorhinal area are
separated by two intermediate sectors obvious in our Nissl preparations.
Insausti et al. specifically identify six entorhinal fields: (1) an
amygdalopiriform cortex which they termed amygdalo-entorhinal transition
field; (2) a medial entorhinal field (MEnt) equivalent to the ventromedial
entorhinal area of Krettek and Price (1977); (3) a caudomedial entorhinal
field (CEnt), which is the classic medial entorhinal area; (4) a ventral
intermediate entorhinal field (VIEnt) equivalent to the caudal ventrolateral
entorhinal field of Krettek and Price (1977); (5) a dorsal intermediate
entorhinal field (DIEnt); and (6) a dorsolateral entorhinal field (DLEnt).
The last two fields together are equivalent to the dorsolateral entorhinal field
of Krettek and Price (1977).
The postsubicular area was identified on the basis of the work of Van Groen
et al. (1992).
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
Cerebral cortex
There have been two comprehensive cortical parcellation schemes in recent
decades. The first notable one was presented by Zilles (1985) and was
constructed on the original stained sections of the earlier editions of our
atlas. In the second edition of our atlas we used the cortical parcellations of
Zilles (1985). The second comprehensive cortical delineation scheme was
presented by Swanson (1992). The Zilles (1985) delineations differ
significantly from the Swanson (1992, 2004) scheme. The atlas of chemical
markers (Paxinos et al., 1999a,b) enabled us to make a decision on the
strengths of the two schemes. On this basis we have retained many of the
features of the sensory, motor, and insular areas proposed by Zilles (1985).
However, we have curtailed the rostral spread of Zilles’s occipital areas and
delineated the sensory representation of the trunk region and temporal
association area in line with Swanson (1992, 2004).
We have retained the perirhinal cortex at caudal levels (along with Zilles, 1985)
because there is a characteristic NADPH-diaphorase reactivity associated with
this area. Palomero-Gallagher and Zilles (2004) have recently completed a
substantial revision of the Zilles and Wree (1995) plan especially in the non-
sensory parietal regions and we have followed their lead. Strong parvalbumin
immunoreactivity is present in layer 4 of the primary somatosensory cortex.
SMI-32 immunoreactivity formed distinctive patches in layer 4 of the barrel
field and forelimb and hindlimb region.
The primary auditory area was identified on the basis of reduced calbindin
immunoreactivity in the deep layers. All the auditory areas were marked by
the presence of SMI- 32 positive cells in the superficial layers (Paxinos et al.,
1999a,b).
AChE marked the location of the prelimbic and agranular insular cortices.
NADPH-diaphorase assisted in defining the agranular insular, perirhinal,
and retrosplenial granular cortices. Additionally, NADPH-diaphorase
immunoreactivity indicated the ventral part of the medial entorhinal cortex.
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Calretinin immunoreactivity assisted in delineation of the lateral entorhinal
cortex where the outer part of layer one is densely stained.
The dorsolateral orbital cortex was delineated in accordance with the work
of Ray and Price (1992). We use the term frontal association cortex for the
frontal cortex that others allocated to the secondary motor cortex (Swanson,
1992, 2004; Zilles, 1985). This designation is in agreement with
microstimulation data (Neafsey et al., 1986).
Retrosplenial cortex (RSD and RSG)
The retrosplenial dysgranular cortex (RSD) was previously named by us the
retrosplenial agranular cortex, but we have changed it on the advice of Brent
Vogt. The retrosplenial granular cortex (RSG) is divided into three areas a, b,
and c. Some authors refer to the RSG as area 24, according to the original
scheme of Brodmann.
Reticular Formation
Refer to Jones (1995) for a general description of the reticular formation.
The intermediate reticular zone was first identified in the rat (Paxinos and
Watson, 1986), but is seen to advantage in the human brain (Paxinos and
Huang, 1995). The intermediate reticular zone at levels of the caudal pole of
the facial nerve nucleus is marked by NADPH-diaphorase positive cells. The
lateral paragigantocellular nucleus is conspicuous in NADPH-diaphorase
preparations (Paxinos et al., 1999b). We have identified the parapyramidal
nucleus as the cell group dorsolateral to the pyramidal tract, which is outlined
but not named in the second edition of this atlas (Paxinos and Watson, 1986).
The identification of the epifascicular nucleus is based on the description of
this nucleus in the human brain (Paxinos and Huang, 1995).
Conterminal nucleus (Ct)
We have identified the conterminal nucleus in the medulla close to the
inferior olive. This group was originally identified by Olzewski and Baxter
(1954) and is clearly shown in the human brain stem atlas (Paxinos and
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
Huang, 1995). The nucleus is seen as two separate AChE positive cell groups,
one lateral to the inferior olive (caudal pole of IOA) and a second group
medial to the IOA.
Periaqueductal and Periventricular Gray
Refer to Keay and Bandler (2004) for a general description of the
periaqueductal gray. The boundaries of periaqueductal gray cell columns
were drawn according to Carrive (1993), Carrive and Paxinos (1994), and
Paxinos and Huang (1995). We identify the rodent homologue of the human
pleoglial periaqueductal gray in Figs 77-80. Two nuclei lateral to the central
gray pars alpha were identified on the basis of SMI-32 immunoreactivity –
central gray pars beta and central gray pars gamma (Paxinos et al., 1999b).
Lithoid nucleus (Li)
We have named this nucleus for the Greek word for a stone. This word was
particularly applied to an elongated stone used in the ancient Olympics.
Li is a prominent group of large cells in the dorsal part of the rostral PVG.
While generally ovoid in cross section, the medial and lateral sides are in
Figures 71-73 parallel to each other. Li lies medial to MCPC caudally, and
medial to RPF rostrally. It is ventral to the PrC and dorsal to the fasciclus
retroflexus. More caudally, it is dorsal to Dk. It can be readily identified in
a horizontal section (Fig 105, Paxinos and Watson, 1998).
Tegmental Nuclei
For the identification of the anterior tegmental, microcellular tegmental,
subpeducular tegmental, rabdoid, and epirubrospinal nuclei, see Paxinos
and Watson (1986).
Epipeduncular nucleus (EpP)
A small but distinctive group of large cells below the peripeduncular nucleus
and above the cerebral peduncle (Fig 78-80) has no home in the surrounding
nuclei. We have named this group the epipedunclular nucleus.
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Reticular tegmental nucleus, L part (RtTgL)
In Figs 112-114 the rat homologue of the subnuclues L of the LtRt nucleus
Olzewski and Baxter (1954) can been identified (see also Paxinos and Huang,
1995).
Raphe Nuclei
We identified the raphe nuclei on the basis of 5-hydroxytryptamine
sections prepared by G. Halliday and I. Tork (see also Harding et al., 2004).
We identified the raphe interpositus nucleus on the basis of the work of
Buettner-Ennever et al. (1988).
Locus Coeruleus and Brainstem Catecholamine Cell Groups
Refer to Aston-Jones (2004) for the delineation of the locus coeruleus.
We delineated the catecholamine cell groups by following Hökfelt et al.
(1984) with assistance from our own tyrosine hydroxylase preparations
(Paxinos et al., 1999b).
Brainstem Nuclei Associated with Taste, Respiratory,
Cardiovascular and Other Autonomic Functions
Refer to Saper (1995) and Norgren (1995) for a general description of
these nuclei.
Nucleus of the solitary tract
The posterior part of the nucleus of the solitary tract was delineated in
accordance with the work of Whitehead (1990), Herbert et al. (1990),
McRitchie (1992), and Altschuler et al. (1989). The rostral part of the
nucleus of the solitary tract was difficult to delineate, but we recognize a
rostrolateral subnucleus on the basis of NADPH-diaphorase positivity.
Medullary respiratory groups and the Botzinger complex
These areas were delineated in accordance with Ellenberger et al. (1990),
Kanjhan et al. (1995), and Cox and Halliday (1993).
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
Parabrachial nucleus
The parabrachial nucleus is delineated in accordance with Fulwiler and Saper
(1984), Herbert et al. (1990), Whitehead (1990), and Herbert and Saper
(1990). The external part of the lateral parabrachial nucleus and medial
parabrachial nucleus are marked by NADPH-diaphorase positive cells and
fibers (Paxinos et al., 1999b).
Oromotor Nuclei
Refer to Travers (2004) for a description of the oromotor nuclei.
Precerebellar Nuclei and Red Nucleus
Refer to Ruigrok (2004) for a general description of these structures. Within
what has been previously called the pararubral area there is a circumscribed
cell group which we called the pararubral nucleus. We have named the large
cells above the lateral lemniscus the epilemniscal nucleus (Paxinos et al.,
1999b).
Cerebellum
The identification of lobules, fissures, and deep cerebellar nuclei is based on
the work of Voogd (2004) and Swanson (2004).
Somatosensory System
Refer to Tracey (2004b) for a general description of the somatosensory
system. The general basis of delineation of these structures is described in
Paxinos and Watson (1996). However, we followed Marfurt and Rajchert
(1991) for the borders of the spinal trigeminal nucleus.
Trigeminal transition zone (5Tr)
We identify the NADPH-diaphorase positive area medial to the principal
sensory trigeminal nucleus as the 5Tr given it juxtaposition between the
trigeminal and the parabrachial nuclei.
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The trigeminosolitary zone (5Sol)
The trigeminosolitary zone commences caudal to the trigeminal transition
zone, and extends as far caudal as the level of the area postrema. The rostral
part of this zone was previously identified by Paxinos and Huang (1995) in
the human and named the subsolitary nucleus. We note that in the medulla,
much like the in thalamus (VPM, VPPC – Lundy and Norgren, 2004), there
is a progression of functional areas from the trigeminal concerned with
somatosensory function to the solitary concerned with gustatory function
from receptors of the same peripheral structures.
Matrix (Mx)
Paxinos and Huang (1995) identified in the human the pericuneate and
peritrigeminal matrix. We observed a similar structure in the rat and
identified it as the residual region after the solitary, trigeminosolitary,
cuneate and parvicellular reticular nuclei are accounted for.
details on the basis of delineation of the components of the auditory system
refer to Paxinos and Watson (1986).
Nucleus of the commissure of the inferior colliculus (Com)
The commissure of the inferior colliculus is populated by many cells. On the
advice of Ellen Covey we have named this the nucleus of the commissure of
the inferior colliculus.
Dorsal cochlear nucleus
We have adopted new abbreviations for the layers of the dorsal cochlear
nucleus: DCDp is the dorsal cochlear deep core; DCFu is the dorsal cochlear
fusiform layer; DCMo is the dorsal cochlear molecular layer.
Periolivary horn (POH)
The periolivary region has a dorsolateral protrusion positive in parvalbumin
(Paxinos et al., 1999b), which we termed the periolivary horn (POH).

Cuneate nucleus, rotund part (CuR)

We note that as in primates, the rat has a rotund part in the cuneate nucleus
which almost certainly represents the forepaw area.

Visual System
Refer to Sefton et al. (2004) for a general description of the visual system. The
ventral tegmental visual relay zone was identified on the basis of the work of
Giolli et al. (1985). The intergeniculate leaf was delineated on the basis of the
work of Morin and Blanchard (1995).

Auditory System
Refer to Malmierca and Merchan (2004) for a general description of the
auditory system. We used Faye-Lund and Osen (1985) as well as Malmierca
and Merchan (2004) for the identification of areas of the inferior colliculus.
The medial geniculate was delineated according to the work of LeDoux et al.
(1985). The nucleus of the central acoustic tract has been identified in the cat,
and Ellen Covey has delineated this structure in our atlas. For additional

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List of Structures accessory optic tract aopt

accumbens nucleus Acb
accumbens nucleus, core AcbC
accumbens nucleus, rostral pole AcbR
Names of the structures are listed in alphabetical
accumbens nucleus, shell AcbSh
order. Each name is followed by abbreviation of
acoustic radiation ar
the structure.
acoustic stria as
agranular insular cortex AI
1st cerebellar lobule (lingula) 1Cb agranular insular cortex, dorsal part AID
2b cerebellar lobule 2bCb agranular insular cortex, posterior part AIP
2nd and 3rd cerebellar lobules 2/3Cb alar nucleus Al
2nd cerebellar lobule 2Cb alveus of the hippocampus alv
3rd and 4th cerebellar lobules 3/4Cb ambiguus nucleus Amb
3rd ventricle 3V ambiguus nucleus, compact part AmbC
4th and 5th cerebellar lobules 4/5Cb ambiguus nucleus, loose part AmbL
4th cerebellar lobule 4Cb amygdalohippocampal area AHi
4th ventricle 4V amygdalohippocampal area, posteromedial part
5th cerebellar lobule 5Cb AHiPM
6a cerebellar lobule 6aCb amygdaloid fissure af
6b cerebellar lobule 6bCb amygdaloid intramedullary gray IMG
6c cerebellar lobule 6cCb amygdalostriatal transition area ASt
6th cerebellar lobule 6Cb anbiguus nucleus, subcompact part AmbSC
7th cerebellar lobule 7Cb angular thalamic nucleus AngT
8th cerebellar lobule 8Cb ansa lenticularis al
9th cerebellar lobule, a 9aCb ansoparamedian fissure apmf
9th cerebellar lobule, a and b 9a,bCb anterior amygdaloid area AA
9th cerebellar lobule, b 9bCb anterior amygdaloid area, dorsal part AAD
9th cerebellar lobule, c 9cCb anterior amygdaloid area, ventral part AAV
10th cerebellar lobule (nodule) 10Cb anterior cerebral artery acer
anterior commissural nucleus AC
anterior commissure ac
A anterior commissure, anterior part aca
A1 noradrenaline cells A1 anterior commissure, intrabulbar part aci
A1 noradrenaline cells/C1 adrenaline cells A1/C1 anterior commissure, posterior part acp
A11dopamine cells A11 anterior hypothalamic area AH
A12 dopamine cells A12 anterior hypothalamic area, central part AHC
A14 dopamine cells A14 anterior lobe of pituitary APit
A2 noradrenaline cells A2 anterior olfactory nucleus, dorsal part AOD
A4 noradrenaline cells A4 anterior olfactory nucleus, external part AOE
A5 noradrenaline cells A5 anterior olfactory nucleus, ventroposterior part
A7 noradrenaline cells A7 AOVP
abducens nucleus 6N anterior perifornical nucleus APF
accessory abducens nucleus 6Acs anterior pretectal nucleus APT
accessory abducens nucleus 6Acs anterior pretectal nucleus, dorsal part APTD
accessory abducens/facial nucleus Acs6/7 anterior pretectal nucleus, ventral part APTV
accessory nerve nucleus 11N anterior spinal artery asp
accessory neurosecretory nuclei ANS anterior tegmental nucleus ATg
accessory olfactory bulb AOB anterodorsal preoptic nucleus ADP
accessory olfactory tract aot anterodorsal thalamic nucleus AD
anterovent thalamic nucleus, dorsomedial part
AVDM
anteroventral preoptic nucleus AVPO
anteroventral thalamic nucleus, ventrolateral part
AVVL
aqueduct Aq
arcuate hypothalamic nucleus, dorsal part ArcD
arcuate hypothalamic nucleus, lateral part ArcL
arcuate hypothalamic nucleus, lateroposterior part
ArcLP
arcuate hypothalamic nucleus, medial part ArcM
area postrema AP
artery a
ascending fibers of the facial nerve asc7
azygous pericallosal artery azp
B brachium of the inferior colliculus bic
B9 serotonin cells B9
Barrington’s nucleus Bar
basal nucleus (Meynert) B
basilar artery bas
basolateral amygdaloid nucleus BL
basolateral amygdaloid nucleus, anterior part BLA
basolateral amygdaloid nucleus, posterior part BLP
basolateral amygdaloid nucleus, ventral part BLV
basomedial amygdaloid nucleus BM
basomedial amygdaloid nucleus, posterior part BMP
bed nucleus of stria terminalis, fusiform part Fu
bed nucleus of stria terminalis, supracapsular division
STS
bed nucleus of stria terminalis, supracapsular
division, lateralpart STSM
bed nucleus of stria terminalis, supracapsular
division, medial part STSL
bed nucleus of the anterior commissure BAC
bed nucleus of the stria terminalis BST
bed nucleus of the stria terminalis, intermediate
division STI
bed nucleus of the stria terminalis, intraamygdaloid
division STIA
bed nucleus of the stria terminalis, lateral division
STL
bed nucleus of the stria terminalis, lateral division,
dorsal part STLD
bed nucleus of the stria terminalis, lateral division,
juxtacapsular part STLJ
bed nucleus of the stria terminalis, lateral division,
ventral part STLV
bed nucleus of the stria terminalis, medial division
STM
bed nucleus of the stria terminalis, medial division,
anterior part STMA
bed nucleus of the stria terminalis, medial division,
anterolateral part STMAL
bed nucleus of the stria terminalis, medial division,
posterior part STMP
bed nucleus of the stria terminalis, medial division,
posterointermediate part STMPI
bed nucleus of the stria terminalis, medial division,
posterolateral part STMPL
bed nucleus of the stria terminalis, medial division,
posteromedial part STMPM
bed nucleus of the stria terminalis, medial division,
ventral part STMV
blood vessel bv
brachium of the superior colliculus bsc
brachium pontis (stem of middle cerebellar peduncle)
bp
C
C1 adrenaline cells C1
C1 adrenaline cells and A1 noradrenaline cells
C1/A1
C2 adrenaline cells C2
C3 adrenaline cells C3
caudal interstitial nucleus of the medial longitudinal
fasciculus CI
caudal linear nucleus of the raphe CLi
caudal periolivary nucleus CPO
caudomedial entothinal cortex CEnt
caudoventral respiratory group CVRG
caudoventrolateral reticular nucleus CVL
cell bridges of the ventral striatum CB
central amygdaloid nucleus, medial division,
anteroventral part CeMAV
central amygdaloid nucleus Ce
central amygdaloid nucleus, capsular part CeC
central amygdaloid nucleus, lateral division CeL
central amygdaloid nucleus, medial division CeM
central amygdaloid nucleus, medial posteroventral
part CeMPV
central canal CC
central cervical nucleus of the spinal cord CeCv
central gray CG
central gray of the pons CGPn

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central gray, alpha part CGA
central gray, gamma part CGG
central gray, nucleus O CGO
central medial thalamic nucleus CM
central tegmental tract ctg
cerebellar white matter cbw
cerebellum Cb
cerebral cortex Cx
cerebral peduncle cp
choroid plexus chp
cingulate cortex, area 1 Cg1
cingulate cortex, area 2 Cg2
cingulum cg
circular nucleus Cir
cochlear root of the vestibulocochlear nerve 8cn
commissural nucleus of the inferior colliculus Com
commissural stria terminalis cst
commissure of the inferior colliculus cic
commissure of the superior colliculus csc
conterminal nucleus Ct
copula of the pyramis Cop
corpus callosum cc
crus 2 of the ansiform lobule Crus2
cuneate fasciculus cu
cuneate nucleus Cu
cuneate nucleus, rotundus part CuR
cuneiform nucleus CnF
cuneiform nucleus, dorsal part CnFD
cuneiform nucleus, intermediate part CnFI
cuneiform nucleus, ventral part CnFV
D dorsal tuberomammillary nucleus DTM
decussation of the superior cerebellar peduncle dscp
decussation of the trapezoid body tzd
deep cerebral white matter dcw
deep gray layer of the superior colliculus DpG
deep mesencephalic nucleus DpMe
deep white layer of the superior colliculus DpWh
dentate gyrus DG
dorsal 3rd ventricle D3V
dorsal acoustic stria das
dorsal cochlear nucleus DC
dorsal cochlear nucleus, deep core DCDp
dorsal cochlear nucleus, fusiform layer DCFu
dorsal cochlear nucleus, molecular layer DCMo
dorsal cortex of the inferior colliculus DCIC
dorsal corticospinal tract dcs
dorsal endopiriform nucleus DEn
The Rat Brain in Stereotaxic Coordinates 5th Edition
dorsal fornix df
dorsal hippocampal commissure dhc
dorsal hypothalamic area DA
dorsal hypothalamic nucleus Do
dorsal intermediate entorhinal cortex DIEnt
dorsal lateral geniculate nucleus DLG
dorsal lateral olfactory tract dlo
dorsal longitudinal fasciculus dlf
dorsal motor nucleus of vagus 10N
dorsal nucleus (Clarke) D
dorsal nucleus of the lateral lemniscus DLL
dorsal paragigantocellular nucleus DPGi
dorsal part of claustrum DCl
dorsal peduncular cortex DP
dorsal peduncular pontine nucleus DPPn
dorsal raphe nucleus DR
dorsal raphe nucleus, dorsal part DRD
dorsal raphe nucleus, lateral part DRL
dorsal raphe nucleus, ventral part DRV
dorsal spinocerebellar fibres and olivocerebellar fibres
dsc/oc
dorsal subiculum DS
dorsal tegmental bundle dtg
dorsal tegmental decussation dtgd
dorsal tegmental nucleus, central part DTgC
dorsal tegmental nucleus, pericentral part DTgP
dorsal tenia tecta DTT
dorsal tenia tecta layer 1 DTT1
dorsal terminal nucleus of the accessory optic tract
DT
dorsal transition zone DTr
dorsolateral entorhinal cortex DLEnt
dorsolateral orbital cortex DLO
dorsolateral periaqueductal gray DLPAG
dorsolateral pontine nucleus DLPn
dorsomedial hypothalamic nucleus DM
dorsomedial hypothalamic nucleus, dorsal part
DMD
dorsomedial periaqueductal gray DMPAG
dorsomedial pontine nucleus DMPn
dorsomedial spinal trigeminal nucleus DMSp5
dorsomedial spinal trigeminal nucleus, dorsal part
DMSp5D
dorsomedial spinal trigeminal nucleus, ventral part
DMSp5V
dysgranular insular cortex DI
Paxinos & Watson
E gigantocellular reticular nucleus Gi
gigantocellular reticular nucleus, alpha part GiA
ectorhinal cortex Ect
gigantocellular reticular nucleus, ventral part GiV
ectotrigeminal nucleus E5
glomerular layer of the accessory olfactory bulb GlA
Edinger-Westphal nucleus EW
glomerular layer of the olfactory bulb Gl
epipeduncular nucleus EpP
gracile fasciculus gr
ependyma and subependymal layer E
granular cell layer of the olfactory bulb GrO
epilemniscal nucleus ELm
granular insular cortex GI
epimicrocellular nucleus EMi
granular layer of the dentate gyrus GrDG
episupraoptic nucleus ESO
granule cell layer of cochlear nuclei GrC
ethmoid thalamic nucleus Eth
granule cell layer of the accessory olfactory bulb
external capsule ec
GrA
external cortex of the inferior colliculus ECIC
granule cell layer of the cerebellum GrCb
external cortex of the inferior colliculus. layer 1
gustatory thalamic nucleus Gus
ECIC1
external cuneate nucleus ECu
external globus pallidus EGP
external medullary lamina eml
H
habenular commissure hbc
external plexiform layer of the accessory olfactory
hilus of the dentate gyrus Hil
bulb EPlA
hippocampal fissure hif
external plexiform layer of the olfactory bulb EPl
hypoglossal nucleus 12N
hypoglossal nucleus, geniohyoid part 12GH
F
facial motor nucleus, accessory part 7Acs
facial nerve 7n
I
indusium griseum IG
facial nucleus 7N
inferior cerebellar peduncle (restiform body) icp
facial nucleus, dorsal intermediate subnucleus 7DI
inferior cerebellar peduncle decussation icpd
facial nucleus, dorsolateral subnucleus 7DL
inferior colliculus IC
facial nucleus, dorsomedial subnucleus 7DM
inferior olive, beta subnucleus IOBe
facial nucleus, lateral subnucleus 7L
inferior olive, cap of Kooy of the medial nucleus
facial nucleus, ventromedial subnucleus 7VM
IOK
fasciculus retroflexus fr
fasciola cinereum FC
inferior olive, dorsal nucleus IOD
inferior olive, dorsomedial cell column IODMC
field CA1 of the hippocampus CA1
inferior olive, dorsomedial cell group IODM
field CA2 of the hippocampus CA2
inferior olive, medial nucleus IOM
field CA3 of the hippocampus CA3
inferior olive, principal nucleus IOPr
fimbria of the hippocampus fi
inferior olive, subnucleus A of medial nucleus IOA
flocculus Fl
inferior olive, subnucleus B of medial nucleus IOB
forceps major of the corpus callosum fmj
inferior olive, ventrolateral protrusion IOVL
forceps minor of the corpus callosum fmi
inferior salivatory nucleus IS
fornix f
infralimbic cortex IL
frontal assocn cortex FrA
infundibular reccess IRe
infundibular stem InfS
G interanterodorsal thalamic nucleus IAD
gelatinous layer of the caudal spinal trigeminal interanteromedial thalamic nucleus IAM
nucleus Ge5 intercalated amygdaloid nucleus, main part IM
gemini hypothalamic nucleus Gem intercalated nuclei of the amygdala I
genu of the corpus callosum gcc intercalated nucleus of the medulla In
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intercrural fissure icf
interfascicular nucleus IF
intermediate acoustic stria ias
intermediate endopiriform nucleus IEn
intermediate geniculate nucleus IntG
intermediate interstitial nucleus of the medial
longitudinal fasciculus II
intermediate reticular nucleus IRt
intermediate reticular nucleus, alpha part IRtA
intermediate white layer of the superior colliculus
InWh
intermediodorsal thalamic nucleus IMD
intermediomedial cell column of the spinal cord
IMM
intermedioventral thalamic commissure imvc
intermedius nucleus of the medulla InM
internal arcuate fibers ia
internal capsule ic
internal globus pallidus (intrapeduncular nucleus)
IGP
internal medullary lamina iml
internal plexiform layer of the olfactory bulb IPl
interoculomotor nucleus I3
interpeduncular fossa IPF
interpeduncular nucleus IP
interpeduncular nucleus, apical subnucleus IPA
interpeduncular nucleus, caudal subnucleus IPC
interpeduncular nucleus, dorsolateral subnucleus
IPDL
interpeduncular nucleus, dorsomedial subnucleus
IPDM
interpeduncular nucleus, intermediate subnucleus
IPI
interpeduncular nucleus, lateral subnucleus IPL
interpeduncular nucleus, rostral subnucleus IPR
interpedunculotegmental tract ipt
interposed cerebellar nucleus, anterior part IntA
interposed cerebellar nucleus, dorsolateral hump
IntDL
interposed cerebellar nucleus, posterior part IntP
interstitial basal nucleus of the medulla IB
interstitial nucleus of Cajal InC
interstitial nucleus of Cajal. shell region InCSh
interstitial nucleus of the posterior limb of the
anterior commissure IPAC
interstitial nucleus of the posterior limb of the
anterior commissure, central part IPACC
The Rat Brain in Stereotaxic Coordinates 5th Edition
interstitial nucleus of the posterior limb of the
anterior commissure, lateral part IPACL
interstitial nucleus of the posterior limb of the
anterior commissure, medial part IPACM
interstitial nucleus of the vestibulocochlear nerve I8
intertrigeminal nucleus I5
interventricular foramen IVF
intradecussational nucleus of the decussation of the
superior cerebellar peduncle ID
islands of Calleja ICj
islands of Calleja, major island ICjM
J lateral posterior thalamic nucleus, mediorostral part
juxtaolivary nucleus JxO
juxtaparaventricular part of lateral hypothalamus
JPLH
juxtarestiform body jx
K lateral septal nucleus, intermediate part LSI
Kîlliker-Fuse nucleus KF
L lateral stripe of the striatum LSS
lacunosum moleculare layer of the hippocampus
LMol
lambdoid septal zone Ld
lat amygdaloid nucleus La
lateral accumbens shell LAcbSh
lateral amygdaloid nucleus, dorsolateral part LaDL
lateral amygdaloid nucleus, ventrolateral part LaVL
lateral amygdaloid nucleus, ventromedial part
LaVM
lateral cerebellar nucleus, parvicellular part LatPC
lateral cervical nucleus of the spinal cord LatC
lateral entorhinal cortex LEnt
lateral habenular nucleus, lateral part LHbL
lateral habenular nucleus, medial part LHbM
lateral hypothalamic area LH
lateral lemniscus ll
lateral mammillary nucleus LM
lateral olfactory tract lo
lateral orbital cortex LO
lateral parabrachial nucleus, central part LPBC
lateral parabrachial nucleus, crescent part LPBCr
lateral parabrachial nucleus, dorsal part LPBD
lateral parabrachial nucleus, medial part LPBM
lateral parabrachial nucleus, superior part LPBS
Paxinos & Watson
lateral parabrachial nucleus, ventral part LPBV
lateral paragigantocellular nucleus LPGi
lateral paragigantocellular nucleus. alpha part
LPGiA
lateral parietal association cortex LPtA
lateral periaqueductal gray LPAG
lateral posterior thalamic nucleus LP
lateral posterior thalamic nucleus, laterocaudal part
LPLC
lateral posterior thalamic nucleus, laterorostral part
LPLR
lateral posterior thalamic nucleus, mediocaudal part
LPMC
LPMR
lateral recess of the 4th ventricle LR4V
lateral reticular nucleus LRt
lateral reticular nucleus, parvicellular part LRtPC
lateral reticular nucleus, subtrigeminal part LRtS5
lateral septal nucleus, dorsal part LSD
lateral septal nucleus, ventral part LSV
lateral spinal nucleus LSp
lateral superior olive LSO
lateral terminal nucleus of the accessory optic tract
LT
lateral vestibular nucleus LVe
lateroanterior hypothalamic nucleus LA
laterodorsal tegmental nucleus LDTg
laterodorsal tegmental nucleus, ventral part LDTgV
laterodorsal thalamic nucleus LD
laterodorsal thalamic nucleus, dorsomedial part
LDDM
laterodorsal thalamic nucleus, ventrolateral part
LDVL
lateroventral periolivary nucleus LVPO
layer 1 of cortex 1
layer 1a of cortex 1a
layer 1b of cortex 1b
layer 4 of cortex 4
layer 5 of cortex 5
layer 5a of cortex 5a
layer 5b of cortex 5b
layer 6 of cortex 6
layer 6a of cortex 6a
layer 6b of cortex 6b
layers 3 and 4 of cortex 3/4
lemina terminalis LTer
linear nucleus of the medulla Li
lithoid nucleus Lth
locus coeruleus LC
longitudinal fasciculus of the pons lfp
lucidum layer of the hippocampus Lu
M
magnocellular nucleus of the lateral hypothalamus
MCLH
magnocellular nucleus of the posterior commissure
MCPC
magnocellular preoptic nucleus MCPO
mammillary peduncle mp
mammillary recess of the 3rd ventricle MRe
mammillotegmental tract mtg
mammillothalamic tract mt
marginal zone of the medial geniculate MZMG
matrix region of the medulla Mx
medial accessory oculomotor nucleus MA3
medial amygdaloid nucleus Me
medial amygdaloid nucleus, ant dorsal MeAD
medial amygdaloid nucleus, anteroventral part
MeAV
medial amygdaloid nucleus, posterodorsal part
MePD
medial amygdaloid nucleus, posteroventral part
MePV
medial cerebellar nucleus, caudomedial part
MedCM
medial cerebellar nucleus, dorsolateral protuberance
MedDL
medial corticohypothalamic tract mch
medial eminence, external layer MEE
medial eminence, internal layer MEI
medial entorhinal cortex MEnt
medial entorhinal cortex, ventral part MEntV
medial forebrain bundle mfb
medial forebrain bundle, ‘a’ component mfba
medial forebrain bundle, ‘b’ component mfbb
medial geniculate nucleus MG
medial geniculate nucleus, dorsal part MGD
medial geniculate nucleus, medial part MGM
medial geniculate nucleus, ventral part MGV
medial habenular nucleus MHb
medial lemniscus ml
medial lemniscus decussation mld
medial longitudinal fasciculus mlf
medial mammillary nucleus, lateral part ML
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RBSC5 Book.qxp 16/9/04 16:36 Page XXXIII
medial mammillary nucleus, medial part MM
medial mammillary nucleus, median part MnM
medial orbital cortex MO
medial parabrachial nucleus MPB
medial paralemniscial nucleus MPL
medial parietal association cortex MPtA
medial preoptic nucleus MPO
medial preoptic nucleus, central part MPOC
medial preoptic nucleus, lateral part MPOL
medial preoptic nucleus, medial part MPOM
medial pretectal nucleus MPT
medial septal nucleus MS
medial superior olive MSO
medial terminal nucleus of the accessory optic tract
MT
medial tuberal nucleus MTu
medial vestibular nucleus MVe
medial vestibular nucleus, magnocellular part
MVeMC
medial vestibular nucleus, parvicellular part MVePC
median accessory nucleus of the medulla MnA
median eminence ME
median preoptic nucleus MnPO
mediodorsal thalamic nucleus MD
mediodorsal thalamic nucleus, central part MDC
mediodorsal thalamic nucleus, lateral part MDL
mediodorsal thalamic nucleus, medial part MDM
mediodorsal thalamic nucleus, paralaminar part
MDPL
medioventral periolivary nucleus MVPO
medullary reticular nucleus, dorsal part MdD
medullary reticular nucleus, ventral part MdV
mesencephalic trigeminal nucleus Me5
mesencephalic trigeminal tract me5
microcellular tegmental nucleus MiTg
middle cerebellar peduncle mcp
middle cerebral artery mcer
mitral cell layer of the accessory olfactory bulb MiA
mitral cell layer of the olfactory bulb Mi
molecular layer of the cerebellum MoCb
molecular layer of the dentate gyrus MoDG
molecular layer of the subiculum MoS
motor root of the trigeminal nerve m5
motor trigeminal nucleus 5N
motor trigeminal nucleus, accessory subnucleus 5Acs
motor trigeminal nucleus, masseter part 5Ma
motor trigeminal nucleus, mylohyoid part 5MHy
motor trigeminal nucleus, temporalis part 5Te
motor trigeminal nucleus, ventromedial part 5VM
The Rat Brain in Stereotaxic Coordinates 5th Edition
N olfactory bulb OB
olfactory nerve layer ON
paraventricular hypothalamic nucleus, dorsal cap
PaDC
navicular nucleus of the basal forebrain Nv
olfactory tubercle Tu paraventricular hypothalamic nucleus, lateral
nervus intermedius component of the facial nerve
olfactory tubercle layer 1 Tu1 magnocellular part PaLM
7ni
olfactory ventricle (olfactory part of lateral ventricle) paraventricular hypothalamic nucleus, medial
nigrostriatal bundle ns
OV magnocellular part PaMM
nucleus of Darkschewitsch Dk
olivary pretectal nucleus OPT paraventricular hypothalamic nucleus, medial
nucleus of origin of efferents of the vestibular nerve
olivocerebellar tract oc parvicellular part PaMP
EVe
olivocochlear bundle ocb paraventricular hypothalamic nucleus, posterior part
nucleus of Roller Ro
optic chiasm och PaPo
nucleus of the ansa lenticularis AL
optic nerve 2n paraventricular hypothalamic nucleus, ventral part
nucleus of the brachium of the inferior colliculus
optic nerve layer of the superior colliculus Op PaV
BIC
optic tract opt paraventricular thalamic nucleus PV
nucleus of the central acoustic tract CAT
oriens layer of the hippocampus Or paraventricular thalamic nucleus, anterior part PVA
nucleus of the commissural stria terminalis CST
oval paracentral thalamic nucleus OPC paraventricular thalamic nucleus, posterior part
nucleus of the dorsal hipp commissure DHC
PVP
nucleus of the fields of Forel F
nucleus of the horizontal limb of the diagonal band P paraxiphoid nucleus of thalamus PaXi
parietal cortex, posterior area, caudal part PtPC
HDB paraabducens nucleus Pa6
parietal cortex, posterior area, dorsal part PtPD
nucleus of the lateral olfactory tract LOT parabigeminal nucleus PBG
parietal cortex, posterior area, rostral part PtPR
nucleus of the lateral olfactory tract, dorsal part parabrachial pigmented nucleus of the VTA PBP
parvicellular motor trigeminal nucleus PC5
LOTD paracentral thalamic nucleus PC
parvicellular reticular nucleus PCRt
nucleus of the lateral olfactory tract, layer 1 LOT1 paracochlear glial substance PCGS
parvicellular reticular nucleus, alpha part PCRtA
nucleus of the optic tract OT paracommissural nucleus of the posterior
peduncular part of lateral hypothalamus PLH
nucleus of the posterior commissure PCom commissure PaC
parafascicular thalamic nucleus PF pedunculopontine tegmental nucleus PPTg
nucleus of the solitary tract Sol
parafloccular sulcus pfs periaqueductal gray PAG
nucleus of the solitary tract, central part SolCe
paraflocculus PFl perifacial zone P7
nucleus of the solitary tract, commissural part SolC
parainterfascicular nucleus of the VTA PIF perifornical nucleus PeF
nucleus of the solitary tract, gelatinous part SolG
paralambdoid septal nucleus PLd perifornical part of lateral hypothalamus PeFLH
nucleus of the solitary tract, intermediate part
paralemniscal nucleus PL perilemniscal nucleus, ventral part PLV
SolIM
paramedian lobule PM periolivary horn POH
nucleus of the solitary tract, interstitial part SolI
paramedian raphe nucleus PMnR peripeduncular nucleus PP
nucleus of the solitary tract, lateral part SolL
paramedian reticular nucleus PMn perirhinal cortex PRh
nucleus of the solitary tract, rostrolateral part SolRL
paramedian sulcus pms peritrigeminal zone P5
nucleus of the solitary tract, ventrolateral part
paranigral nucleus of the VTA PN periventricular gray PVG
SolVL
parapyramidal nucleus PPy periventricular hypothalamic nucleus Pe
nucleus of the stria medullaris SM
pararubral nucleus PaR periventricular hypothalamic nucleus, anterior
nucleus of the trapezoid body Tz
parasolitary nucleus PSol parvicellular part PeAP
nucleus of the vertical limb of the diagonal band
parastrial nucleus PS pineal gland Pi
VDB
parasubiculum PaS pineal stalk PiSt
nucleus X X
parasubthalamic nucleus PSTh piriform cortex Pir
nucleus Y Y
paratenial thalamic nucleus PT piriform cortex, layer 1 Pir1
nucleus Z Z
paraterete nucleus PTe piriform cortex, layer 1a Pir1a
paratrigeminal nucleus Pa5 plioglial part of periaqueductai gray PlPAG
O paratrochlear nucleus Pa4 polymorph layer of the dentate gyrus PoDG
obex Obex paraventricular hypoth nucleus Pa pontine nuclei Pn
oculomotor nerve 3n paraventricular hypothalamic nucleus, anterior pontine raphe nucleus PnR
oculomotor nucleus, parvicellular part 3PC parvicellular part PaAP pontine reticular nucleus, caudal part PnC
Paxinos & Watson XXXIII
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pontine reticular nucleus, oral part PnO
posterior cerebral artery pcer
posterior commissure pc
posterior hypothalamic area PHA
posterior hypothalamic area, dorsal part PHD
posterior hypothalamic nucleus PH
posterior intralaminar thalamic nucleus PIL
posterior limitans thalamic nucleus PLi
posterior lobe of pituitary PPit
posterior pretectal nucleus PPT
posterior superior fissure psf
posterior thalamic nuclear group Po
posterior thalamic nuclear group, triangular part
PoT
posterodorsal preoptic nucleus PDPO
posterodorsal raphe nucleus PDR
posterodorsal tegmental nucleus PDTg
posterolateral cortical amygdaloid nucleus PLCo
posterolateral cortical amygdaloid nucleus, layer 1
PLCo1
posterolateral fissure plf
posteromedial cortical amygdaloid nucleus PMCo
posteromedian thalamic nucleus PoMn
postsubiculum Post
pre-Botzinger complex PrBo
precommissural fornix pcf
precommissural nucleus PrC
preculminate fissure pcuf
prelimbic cortex PrL
premammillary nucleus, dorsal part PMD
premammillary nucleus, ventral part PMV
preoptic recess of the 3rd ventricle P3V
prepyramidal fissure ppf
prerubral field PR
presubiculum PrS
primary auditory cortex Au1
primary fissure prf
primary motor cortex M1
primary somatosensory cortex S1
primary somatosensory cortex, barrel field S1BF
primary somatosensory cortex, dysgranular zone
S1DZ
primary somatosensory cortex, forelimb region
S1FL
primary somatosensory cortex, hindlimb region
S1HL
primary somatosensory cortex, jaw region S1J
primary somatosensory cortex, jaw region, oral
surface S1JO
The Rat Brain in Stereotaxic Coordinates 5th Edition
primary somatosensory cortex, oral dysgranular zone
S1DZO
primary somatosensory cortex, shoulder region
S1Sh
primary somatosensory cortex, shoulder/neck region
S1ShNc
primary somatosensory cortex, trunk region S1Tr
primary somatosensory cortex, upper lip region
S1ULp
primary visual cortex V1
primary visual cortex, binocular area V1B
primary visual cortex, monocular area V1M
principal mammillary tract pm
principal sensory trigeminal nucleus Pr5
principal sensory trigeminal nucleus, dorsomedial
part Pr5DM
principal sensory trigeminal nucleus, ventrolateral
part Pr5VL
Purkinje cell layer of the cerebellum Pk
pyramidal cell layer of the hippocampus Py
pyramidal decussation pyd
pyramidal tract py
R secondary somatosensory cortex S2
radiatum layer of the hippocampus Rad
raphe interpositus nucleus RIP
raphe magnus nucleus RMg
raphe obscurus nucleus ROb
raphe pallidus nucleus RPa
recess of the inferior colliculus ReIC
red nucleus R
red nucleus, magnocellular part RMC
red nucleus, parvicellular part RPC
region where VA and VL overlap VA/VL
reticluostrial nucleus RtSt
reticular thalamic nucleus Rt
reticulotegmental nucleus of the pons RtTg
reticulotegmental nucleus of the pons, lateral part
RtTgL
retroambiguus nucleus RAmb
retrochiasmatic area, lateral part RChL
retroethmoid nucleus REth
retrolemniscal nucleus RL
retroparafascicular nucleus RPF
retrorubral field RRF
retrorubral nucleus RR
retrosplenial dysgranular cortex RSD
retrosplenial granular cortex RSG
Paxinos & Watson
retrosplenial granular cortex, c region RSGc
retrouniens area RRe
reuniens thalamic nucleus Re
rhabdoid nucleus Rbd
rhinal fissure rf
rhinal incisure ri
rhomboid thalamic nucleus Rh
root of abducens nerve 6n
root of accessory nerve 11n
root of hypoglossal nerve 12n
rostral interstitial nucleus of medial longitudinal
fasciculus RI
rostral ventral respiratory group RVRG
rostroventrolateral reticular nucleus RVL
rostrum of the corpus callosum rcc
rubrospinal tract rs
S submedius thalamic nucleus, dorsal part SubD
scaphoid thalamic nucleus Sc
secondary auditory cortex, dorsal area AuD
secondary auditory cortex, ventral area AuV
secondary fissure sf
secondary motor cortex M2
secondary visual cortex, lateral area V2L
secondary visual cortex, mediolateral area V2ML
secondary visual cortex, mediomedial area V2MM
sensory root of the trigeminal nerve s5
septofimbrial nucleus SFi
septohippocampal nucleus SHi
septohypothalamic nucleus SHy
simple lobule Sim
simple lobule A SimA
simple lobule B SimB
simplex fissure simf
solitary nucleus, dorsolateral part SolDL
solitary nucleus, ventral part SolV
solitary tract sol
sphenoid nucleus Sph
spinal trigeminal nucleus, caudal part Sp5C
spinal trigeminal nucleus, interpolar part Sp5I
spinal trigeminal nucleus, oral part Sp5O
spinal trigeminal tract sp5
spinal vestibular nucleus SpVe
splenium of the corpus callosum scc
stigmoid hypothalamic nucleus Stg
stratum lucidum of the hippocampus SLu
stria medullaris of the thalamus sm
stria terminalis st
strial part of the preoptic area StA
striohypothalamic nucleus StHy
subbrachial nucleus SubB
subcoeruleus nucleus, alpha part SubCA
subcoeruleus nucleus, dorsal part SubCD
subcoeruleus nucleus, ventral part SubCV
subcommissural organ SCO
subfornical organ SFO
subgeniculate nucleus SubG
subiculum S
subiculum, transition area STr
subincertal nucleus SubI
sublenticular extended amygdala EA
sublenticular extended amygdala, central part EAC
sublenticular extended amygdala, medial part EAM
submammillothalamic nucleus SMT
submedius thalamic nucleus Sub
submedius thalamic nucleus, ventral part SubV
subparafascicular thalamic nucleus, parvicellular part
SPFPC
subparaventricular zone of the hypothalamus SPa
subpeduncular tegmental nucleus SPTg
subpostrema area SubP
substantia innominata SI
substantia innominata, basal part SIB
substantia innominata, dorsal part SID
substantia innominata, ventral part SIV
substantia nigra, compact part, dorsal tier SNCD
substantia nigra, compact part, medial tier SNCM
substantia nigra, lateral part SNL
substantia nigra, reticular part SNR
subthalamic nucleus STh
superficial gray layer of the superior colliculus SuG
superior cerebellar peduncle (brachium
conjunctivum) scp
superior cerebellar peduncle, descending limb scpd
superior medullary velum SMV
superior paraolivary nucleus SPO
superior salivatory nucleus SuS
superior thalamic radiation str
superior vestibular nucleus SuVe
suprachiasmatic nucleus SCh
suprachiasmatic nucleus, ventromedial part SChVM
suprachiasmatic nucleus, dorsolateral part SChDL
suprageniculate thalamic nucleus SG
supragenual nucleus SGe
supramammillary decussation sumd
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RBSC5 Book.qxp 16/9/04 16:36 Page XXXV
supramammillary nucleus SuM
supramammillary nucleus, lateral part SuML
supramammillary nucleus, medial part SuMM
supraoculomotor cap Su3C
supraoculomotor periaqueductal gray Su3
supraoptic decussation sod
supraoptic nucleus SO
supraoptic nucleus, retrochiasmatic part SOR
supratrigeminal nucleus Su5
T ventral cochlear nucleus, anterior part VCA
tectospinal tract ts
temporal associatin cortex TeA
terete hypothalamic nucleus Te
transverse fibers of the pons tfp
trapezoid body tz
triangular nucleus Tr
triangular septal nucleus TS
trigeminal ganglion 5Gn
trigeminal transition zone 5Tr
trigeminal-solitary transition zone 5Sol
trigeminothalamic tract tth
trochlear nerve 4n
trochlear nerve decussation 4d
trochlear nucleus 4N
trochlear nucleus shell region 4Sh
tuberal region of lateral hypothalamus TuLH
The Rat Brain in Stereotaxic Coordinates
U ventral posterior nucleus of the thalamus,
parvicellular part VPPC
ventromedial preoptic nucleus VMPO
ventromedial thalamic nucleus VM
uncinate fasciculus decussation und
ventral posterolateral thalamic nucleus VPL vertebral artery vert
uncinate fasciculus of the cerebellum un ventral posteromedial thalamic nucleus VPM
vestibular root of the vestibulocochlear nerve 8vn
ventral reuniens thalamic nucleus VRe vestibulocerebellar nucleus VeCb
V ventral spinocerebellar tract vsc
ventral spinocerebellar tract decussation vscd
vestibulocochlear ganglion 8Gn
vestibulocochlear nerve 8n
vagus nerve 10n
ventral subiculum VS vestibulomesencephalic tract veme
vascular organ of the lamina terminalis VOLT
ventral tegmental area VTA vestibulospinal tract vesp
vein v ventral tegmental area, rostral part VTAR vomeronasal nerve vn
ventral anterior thalamic nucleus VA ventral tegmental decussation vtgd vomeronasal nerve layer VN
ventral tegmental nucleus VTg ventrolateral hypothalamic tract vlh
ventral cochlear nucleus, capsular part VCCap ventral tenia tecta VTT
ventral cochlear nucleus, granule cell layer VCAGr
ventral cochlear nucleus, posterior part VCP
ventral tenia tecta, layer 1 VTT1
ventral tuberomammillary nucleus VTM
X
xiphoid thalamic nucleus Xi
ventral cochlear nucleus, posterior part, octopus cell ventrolateral preoptic nucleus VLPO
ventrolateral thalamic nucleus VL
area VCPO
ventral endopiriform nucleus VEn ventromedial hypothalamic nucleus VMH Z
ventromedial hypothalamic nucleus, anterior part zona incerta ZI
ventral hippocampal commissure vhc
VMHA zona incerta, caudal part ZIC
ventral intermediate entorhinal cortex VIEnt
ventromedial hypothalamic nucleus, central part zona incerta, dorsal part ZID
ventral lateral geniculate nucleus VLG
VMHC zona incerta, rostral part ZIR
ventral lateral geniculate nucleus, layer 1 VLG1 zona incerta, ventral part ZIV
ventromedial hypothalamic nucleus, dorsomedial part
ventral linear nucleus of the thalamus VLi VMHDM zona layer of the superior colliculus Zo
ventral nucleus of the lat lemniscus VLL ventromedial hypothalamic nucleus, ventrolateral part zona limitans ZL
ventral orbital cortex VO VMHVL zonal layer of the superior colliculus GI
ventral pallidum VP ventromedial nucleus of the hypothalamus shell
ventral part of claustrum VCl VMHSh
5th Edition Paxinos & Watson XXXV
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Index of
Abbreviations
The abbreviations are listed in alphabetical order.
Each abbreviation is followed by the structure name
and the number of the figures on which the
abbreviation appears.
1 layer 1 of cortex 9-21, 30-34, 52, 54-55, 59-75,
94, 99-109
1b layer 1b of cortex 8
1Cb 1st cerebellar lobule (lingula) 112-128
2 layer 2 of cortex 7-75, 94, 99-109
2/3Cb 2nd and 3rd cerebellar lobules 106-109
2bCb 2b cerebellar lobule 110
2Cb 2nd cerebellar lobule 103-120
3 layer 3 of cortex 8-75, 99-109
3/4 layers 3 and 4 of cortex 94
3/4Cb 3rd and 4th cerebellar lobules 127-129
3Cb 3rd cerebellar lobule 106-120, 122, 124-126,
134
3N oculomotor nucleus 85-91
3n oculomotor nerve 79-82
3PC oculomotor nucleus, parvicellular part 84-91
3V 3rd ventricle 29-73
4 layer 4 of cortex 8-10, 30-33, 52
4/5Cb 4th and 5th cerebellar lobules 106-109
4Cb 4th cerebellar lobule 108-126
4N trochlear nucleus 92-95
4n trochlear nerve 94-112
4Sh trochlear nucleus shell region 92-95
4V 4th ventricle 104-146
5 layer 5 of cortex 52
5a layer 5a of cortex 52, 94
5Acs motor trigeminal nucleus, accessory
subnucleus 113-116
5b layer 5b of cortex 52, 94
5Cb 5th cerebellar lobule 109-120, 122-131
5Ma motor trigeminal nucleus, masseter part
109-115
5MHy motor trigeminal nucleus, mylohyoid part
109-110
5N motor trigeminal nucleus 108
5Sol trigeminal-solitary transition zone 122-148
5Te motor trigeminal nucleus, temporalis part
109-115
The Rat Brain in Stereotaxic Coordinates 5th Edition
5Tr trigeminal transition zone 113-117
5VM motor trigeminal nucleus, ventromedial part
111-113
6 layer 6 of cortex 94
6a layer 6a of cortex 52
6aCb 6a cerebellar lobule 121-127, 129-133
6Acs accessory abducens nucleus 117-120
6b layer 6b of cortex 52
6bCb 6b cerebellar lobule 134-138
6Cb 6th cerebellar lobule 128, 133
6cCb 6c cerebellar lobule 134-137, 139-143
6N abducens nucleus 117-119
6n root of abducens nerve 115-116
7Acs facial motor nucleus, accessory part 120-125,
128-132
7Cb 7th cerebellar lobule 137-150
7DI facial nucleus, dorsal intermediate subnucleus
123-131
7DL facial nucleus, dorsolateral subnucleus
122-132
7DM facial nucleus, dorsomedial subnucleus
121-131
7L facial nucleus, lateral subnucleus 122-134
7n facial nerve 111-119
7ni nervus intermedius component of the facial
nerve 121-123
7VI facial nucleus, ventral intermediate subnucleus
121-132
7VM facial nucleus, ventromedial subnucleus
121-130
8Cb 8th cerebellar lobule 135-155
8cn cochlear root of the vestibulocochlear nerve
112-117, 121-123, 126-127
8vn vestibular root of the vestibulocochlear nerve
115-125
9a,bCb 9th cerebellar lobule, a and b 140-159
9Cb 9th cerebellar lobule 131-133, 135-139
9cCb 9th cerebellar lobule, c 140-158
9n glossopharyngeal nerve 130-132
10Cb 10th cerebellar lobule (nodule) 129-149
10N dorsal motor nucleus of vagus 136-156
10n vagus nerve 132-133, 135-137, 139, 141-143,
147, 149-150
11N accessory nerve nucleus 157-161
12GH hypoglossal nucleus, geniohyoid part
147-156
12N hypoglossal nucleus 138-157
12n root of hypoglossal nerve 137, 141, 144,
149-155
Paxinos & Watson
A
artery 8-9, 38-39, 41, 43-45, 47-49, 52-63, 65-74,
76-79, 81, 83-85, 87, 91-105, 107, 109, 113-114,
118, 120-126, 128, 130-131, 148-149
A1 A1 noradrenaline cells 152-161
A1/C1 A1 noradrenaline cells/C1 adrenaline cells
151
A11 A11dopamine cells 59-70
A13 A13 dopamine cells 50-57
A2 A2 noradrenaline cells 155-161
A5 A5 noradrenaline cells 108-113, 115-125, 127
A7 A7 noradrenaline cells 102-108
AA anterior amygdaloid area 35-48
ac anterior commissure 34-37
aca anterior commissure, anterior part 8-33
AcbC accumbens nucleus, core 11-29
AcbR accumbens nucleus, rostral pole 10
AcbSh accumbens nucleus, shell 11-29
acer anterior cerebral artery 13-20, 29-30, 35-37
aci anterior commissure, intrabulbar part 2-7
ACo anterior cortical amygdaloid nucleus 37-55
acp anterior commissure, posterior part 34-43
AD anterodorsal thalamic nucleus 43-52
af amygdaloid fissure 46-47, 50, 54, 68-80
AHA anterior hypothalamic area, anterior part
41-45
AHC anterior hypothalamic area, central part
46-50
AHiAL amygdalohippocampal area, anterolateral
part 59-66
AHiPL amygdalohippocampal area, posterolateral
67-70
AHiPM amygdalohippocampal area, posteromedial
part 67-81
AHP anterior hypothalamic area, posterior part
47-52
AID agranular insular cortex, dorsal part 8-32
AIP agranular insular cortex, posterior part 33-57
AIV agranular insular cortex, ventral part 8-32
Al alar nucleus 33-34
alv alveus of the hippocampus 47-96
AM anteromedial thalamic nucleus 42-53
AmbC ambiguus nucleus, compact part 133-140
AmbL ambiguus nucleus, loose part 146-151
AmbSC anbiguus nucleus, subcompact part
141-145
AMV anteromedial thalamic nucleus, ventral part
45-49
AngT angular thalamic nucleus 53-55
ANS accessory neurosecretory nuclei 43-50
AOD anterior olfactory nucleus, dorsal part 4-7
AOE anterior olfactory nucleus, external part 3-5
AOL anterior olfactory nucleus, lateral part 3-8
AOM anterior olfactory nucleus, medial part 4-7
AOP anterior olfactory nucleus, posterior part
9-11
aot accessory olfactory tract 46
AOV anterior olfactory nucleus, ventral part 4-5
AOVP anterior olfactory nucleus, ventroposterior
part 6-9
AP area postrema 147-152
APF anterior perifornical nucleus 39-41
APir amygdalopiriform transition area 64-88
apmf ansoparamedian fissure 139-148, 150
APT anterior pretectal nucleus 80-82
APTD anterior pretectal nucleus, dorsal part
69-79
APTV anterior pretectal nucleus, ventral part
70-79
Aq aqueduct 74-103
Arc arcuate hypothalamic nucleus 47-48
ArcD arcuate hypothalamic nucleus, dorsal part
49-61
ArcL arcuate hypothalamic nucleus, lateral part
49-61
ArcLP arcuate hypothalamic nucleus,
lateroposterior part 62-68
ArcM arcuate hypothalamic nucleus, medial part
49-61
ArcMP arcuate hypothalamic nucleus, medial
posterior part 62-69
asc7 ascending fibers of the facial nerve 122-126
asp anterior spinal artery 137, 139-142, 144-145
ASt amygdalostriatal transition area 46-50, 54-61
ATg anterior tegmental nucleus 96-98
Au1 primary auditory cortex 60-90
AuD secondary auditory cortex, dorsal area 58-90
AuV secondary auditory cortex, ventral area
60-90
AV anteroventral thalamic nucleus 42
AVDM anterovent thalamic nucleus, dorsomedial
part 43-53
AVPe anteroventral periventricular nucleus 32-34
AVVL anteroventral thalamic nucleus, ventrolateral
part 43-51
azac azygous anterior cerebral artery 10-12
azp azygous pericallosal artery 10-18
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RBSC5 Book.qxp 16/9/04 16:36 Page XXXVII
B CeM
B basal nucleus (Meynert) 36-55, 57-59
B9 B9 serotonin cells 90-96, 100
BAC bed nucleus of the anterior commissure 37-40
BAOT bed nucleus of the accessory olfactory tract
46-51
Bar Barrington’s nucleus 108-112
bas basilar artery 85-96, 98-135
BIC nucleus of the brachium of the inferior
colliculus 88-98
bic brachium of the inferior colliculus 82-102
BL basolateral amygdaloid nucleus 74-75
BLA basolateral amygdaloid nucleus, anterior part
46-61
BLP basolateral amygdaloid nucleus, posterior part
53-73
BLV basolateral amygdaloid nucleus, ventral part
49-61
BMA basomedial amygdaloid nucleus, anterior part
43-56
BMP basomedial amygdaloid nucleus, posterior
part 54-67
Bo Botzinger complex 133-137
bsc brachium of the superior colliculus 69-71,
74-86
C Com commissural nucleus of the inferior colliculus
C1 C1 adrenaline cells 131-146
C1/A1 C1 adrenaline cells and A1 noradrenaline
cells 147-150
C2 C2 adrenaline cells 138-139, 141-143
C3 C3 adrenaline cells 135-137
CA1 field CA1 of the hippocampus 52-66, 68-90
CA2 field CA2 of the hippocampus 52-61, 63-77
CA3 field CA3 of the hippocampus 47-81, 83-84
CAT nucleus of the central acoustic tract 105-109
CB cell bridges of the ventral striatum 24-34
cbc cerebellar commissure 120-122, 124, 126
cbw cerebellar white matter 108-156
CC central canal 147-161
cc corpus callosum 21-74
CeC central amygdaloid nucleus, capsular part
46-61
CeCv central cervical nucleus of the spinal cord
149-161
CeL central amygdaloid nucleus, lateral division
49-60
The Rat Brain in Stereotaxic Coordinates 5th Edition
central amygdaloid nucleus, medial division
45-57
CEnt caudomedial entothinal cortex 89-112
CG central gray 106-108
cg cingulum 11-83
Cg1 cingulate cortex, area 1 8-46
Cg2 cingulate cortex, area 2 14-46
CGA central gray, alpha part 109-118
CGB central gray, beta part 109-115
CGG central gray, gamma part 115-118
CGO central gray, nucleus O 110-114
CGPn central gray of the pons 113-115
chp choroid plexus 34-41, 43-76, 118-119,
124-143, 145-146
CI caudal interstitial nucleus of the medial
longitudinal fasciculus 122-128
CIC central nucleus of the inferior colliculus
98-109
cic commissure of the inferior colliculus 98-102
Cir circular nucleus 45
CL centrolateral thalamic nucleus 51-69
Cl claustrum 8-14
CLi caudal linear nucleus of the raphe 85-94
cll commissure of the lateral lemniscus 100-103
CM central medial thalamic nucleus 44-66
CnFD cuneiform nucleus, dorsal part 101-106
CnFI cuneiform nucleus, intermediate part 98-105
CnFV cuneiform nucleus, ventral part 98-105
99-102
Cop copula of the pyramis 135-154
cp cerebral peduncle 61-93
CPO caudal periolivary nucleus 121
CPu caudate putamen (striatum) 12-66
Crus1 crus 1 of the ansiform lobule 117-141
Crus2 crus 2 of the ansiform lobule 132-150
csc commissure of the superior colliculus 76- 86
CST nucleus of the commissural stria terminalis
35
cst commissural stria terminalis 39, 41-64
Ct conterminal nucleus 152-155
Cu cuneate nucleus 140-161
cu cuneate fasciculus 143-162
CuR cuneate nucleus, rotundus part 147-154
CVL caudoventrolateral reticular nucleus 138-145,
150-151
Cx cerebral cortex 112-113
CxA1 cortex-amygdala transition zone, layer 1
34-47
Paxinos & Watson
D DpWh deep white layer of the superior colliculus
D dorsal nucleus (Clarke) 162c-d
D3V dorsal 3rd ventricle 42-76
DA dorsal hypothalamic area 52-60
das dorsal acoustic stria 126-129
DCDp dorsal cochlear nucleus, deep core 121-130
DCFu dorsal cochlear nucleus, fusiform layer
121-131
DCGr dorsal cochlear nucleus, granular layer 121
DCIC dorsal cortex of the inferior colliculus
101-110
DCl dorsal part of claustrum 15-49
DCMo dorsal cochlear nucleus, molecular layer
121-131
dcs dorsal corticospinal tract 160-162
dcw deep cerebral white matter 55-98
DEn dorsal endopiriform nucleus 8-70
df dorsal fornix 40-68
DG dentate gyrus 66
dhc dorsal hippocampal commissure 49-92
DI dysgranular insular cortex 10-57
DIEnt dorsal intermediate entorhinal cortex 85-95
Dk nucleus of Darkschewitsch 72-82
DLEnt dorsolateral entorhinal cortex 85-103
DLG dorsal lateral geniculate nucleus 60-78
DLL dorsal nucleus of the lateral lemniscus
100-106
DLO dorsolateral orbital cortex 6-7
dlo dorsal lateral olfactory tract 3-4
DLPAG dorsolateral periaqueductal gray 82-101
DM dorsomedial hypothalamic nucleus 63-64
DMC dorsomedial hypothalamic nucleus, compact
part 58-62
DMD dorsomedial hypothalamic nucleus, dorsal
part 52-62
DMPAG dorsomedial periaqueductal gray 77-106
DMSp5 dorsomedial spinal trigeminal nucleus
120-140
DMTg dorsomedial tegmental area 105-115
DMV dorsomedial hypothalamic nucleus, ventral
part 59-62
DP dorsal peduncular cortex 9-15
DpG deep gray layer of the superior colliculus
78-100
DPGi dorsal paragigantocellular nucleus 120-132
DpMe deep mesencephalic nucleus 81-100
DPO dorsal periolivary region 113-116, 118-119
DPPn dorsal peduncular pontine nucleus 102
79-100
DR dorsal raphe nucleus 90-91, 111
DRC dorsal raphe nucleus, caudal part 104-110
DRD dorsal raphe nucleus, dorsal part 92-103
DRL dorsal raphe nucleus, lateral part 93-100
DRV dorsal raphe nucleus, ventral part 93-103
DS dorsal subiculum 74-90
dsc dorsal spinocerebellar tract 150-161
dsc/oc dorsal spinocerebellar fibres and
olivocerebellar fibres 137-149
dscp decussation of the superior cerebellar peduncle
96-98
DT dorsal terminal nucleus of the accessory optic
tract 83-87
DTgC dorsal tegmental nucleus, central part
107-110
dtgd dorsal tegmental decussation 84-89
DTgP dorsal tegmental nucleus, pericentral part
104-110
DTM dorsal tuberomammillary nucleus 63-65
DTr dorsal transition zone 8-9
DTT dorsal tenia tecta 9-14
DTT1 dorsal tenia tecta layer 1 8
E
E ependyma and subependymal layer 1-5, 13-39
E/OV ependymal and subendymal layer/olfactory
ventricle 6-12
E5 ectotrigeminal nucleus 143
EA sublenticular extended amygdala 47-48
EAC sublenticular extended amygdala, central part
39-46
EAM sublenticular extended amygdala, medial part
43-46
ec external capsule 14-66
ECIC external cortex of the inferior colliculus
91-110
Ect ectorhinal cortex 58-111
ECu external cuneate nucleus 135-151
EF epifascicular nucleus 132-134
EGP external globus pallidus 35-59
ELm epilemniscal nucleus 82
eml external medullary lamina 60-65
EPl external plexiform layer of the olfactory bulb
1-5
EPlA external plexiform layer of the accessory
olfactory bulb 3-4
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EpP epipeduncular nucleus 78-80
ERS epirubrospinal nucleus 98-99
ESO episupraoptic nucleus 39-45
Eth ethmoid thalamic nucleus 70-73
EVe nucleus of origin of efferents of the vestibular
nerve 116-125
EW Edinger-Westphal nucleus 73, 76-90
F IntA interposed cerebellar nucleus, anterior part
F nucleus of the fields of Forel 67-71
f fornix 34-72
FC fasciola cinereum 57-75, 77
fi fimbria of the hippocampus 32-68
Fl flocculus 107-125
fmi forceps minor of the corpus callosum 8-13
fmj forceps major of the corpus callosum 77-101
fr fasciculus retroflexus 58-80
Fr3 frontal cortex, area 3 7-12
FrA frontal assocn cortex 4-5
Fu bed nucleus of stria terminalis, fusiform part
32-36
FVe F cell group of the vestibular complex
137-140
G icp inferior cerebellar peduncle (restiform body)
g7 genu of the facial nerve 118-124
gcc genu of the corpus callosum 14-20
Ge5 gelatinous layer of the caudal spinal trigeminal
nucleus 158-161
Gem gemini hypothalamic nucleus 66-70
GI granular insular cortex 12-57
Gi gigantocellular reticular nucleus 119-148
GiA gigantocellular reticular nucleus, alpha part
119-133
GiV gigantocellular reticular nucleus, ventral part
134-143
Gl glomerular layer of the olfactory bulb 1-5
GlA glomerular layer of the accessory olfactory bulb
4-5
Gr gracile nucleus 144-161
gr gracile fasciculus 153-162
GrA granule cell layer of the accessory olfactory
bulb 2-5
GrC granule cell layer of cochlear nuclei 109-128
GrCb granule cell layer of the cerebellum 120,
122, 125, 131, 134
GrDG granular layer of the dentate gyrus 47-89
The Rat Brain in Stereotaxic Coordinates 5th Edition
GrO granular cell layer of the olfactory bulb
1-6
H 75-84
hbc habenular commissure 67-72
HDB nucleus of the horizontal limb of the
diagonal band 24-46
hif hippocampal fissure 54-85
I 122-129
I intercalated nuclei of the amygdala 39-40, 42-48,
52-55, 57-60
I8 interstitial nucleus of the vestibulocochlear nerve
118-123
ia internal arcuate fibers 151-156, 160-161
IAD interanterodorsal thalamic nucleus 44-48
IAM interanteromedial thalamic nucleus 47-51
IB interstitial basal nucleus of the medulla
155-161
IC inferior colliculus 111
ic internal capsule 33-71
icf intercrural fissure 132-140
ICj islands of Calleja 10, 12-32
ICjM islands of Calleja, major island 15-16, 18-24
116-143
icpd inferior cerebellar peduncle decussation
122-124
ID intradecussational nucleus of the decussation
of the superior cerebellar peduncle 89-92
IEn intermediate endopiriform nucleus 8-39
IF interfascicular nucleus 75-87
IG indusium griseum 14-76
IGL intergeniculate leaf 65-75
IGP internal globus pallidus (intrapeduncular
nucleus) 49-56
II intermediate interstitial nucleus of the medial
longitudinal fasciculus 115-116
IL infralimbic cortex 9-13
ILL intermediate nucleus of the lateral lemniscus
99-105
IM intercalated amygdaloid nucleus, main part
49-51
IMA intramedullary thalamic area 65-78
IMD intermediodorsal thalamic nucleus 53- 64
IMG amygdaloid intramedullary gray 54-56
iml internal medullary lamina 65
Paxinos & Watson
imvc intermedioventral thalamic commissure 60
In intercalated nucleus of the medulla 140-141
InC interstitial nucleus of Cajal 75-90
InCSh interstitial nucleus of Cajal. shell region
InfS infundibular stem 62-65
InG intermediate gray layer of the superior
colliculus 77-99
InM intermedius nucleus of the medulla 142-146
IntDL interposed cerebellar nucleus, dorsolateral
hump 123-132
IntDM interposed cerebellar nucleus, dorsomedial
crest 124-127, 130-131
IntP interposed cerebellar nucleus, posterior part
128-133
IntPPC interposed cerebellar nucleus, posterior
parvicellular part 128-130
InWh intermediate white layer of the superior
colliculus 76-99
IOA inferior olive, subnucleus A of medial nucleus
148-154
IOB inferior olive, subnucleus B of medial nucleus
146-154
IOBe inferior olive, beta subnucleus 146-153
IOC inferior olive, subnucleus C of medial nucleus
146-154
IOD inferior olive, dorsal nucleus 134-149
IODM inferior olive, dorsomedial cell group
142-143
IOK inferior olive, cap of Kooy of the medial
nucleus 147-151
IOM inferior olive, medial nucleus 134-145,
155-156
IOPr inferior olive, principal nucleus 133-147
IOVL inferior olive, ventrolateral protrusion
144-146
IP interpeduncular nucleus 79
IPA interpeduncular nucleus, apical subnucleus
89-92
IPAC interstitial nucleus of the posterior limb
of the anterior commissure 28-34, 44-45
IPACL interstitial nucleus of the posterior limb
of the anterior commissure, lateral part 35-43
IPACM interstitial nucleus of the posterior limb
of the anterior commissure, medial part 35-43
IPC interpeduncular nucleus, caudal subnucleus
80-93
IPDL interpeduncular nucleus, dorsolateral
subnucleus 85-90
IPDM interpeduncular nucleus, dorsomedial
subnucleus 84-87
IPF interpeduncular fossa 75-80
IPI interpeduncular nucleus, intermediate
subnucleus 87-91
IPL interpeduncular nucleus, lateral subnucleus
80-93
IPl internal plexiform layer of the olfactory bulb
1-5
IPR interpeduncular nucleus, rostral subnucleus
80-88
IRt intermediate reticular nucleus 117-119,
133-161
IRtA intermediate reticular nucleus, alpha part
120-132
IS inferior salivatory nucleus 128-135
IVF interventricular foramen 39-42
J
JPLH juxtaparaventricular part of lateral
hypothalamus 45-48
JxO juxtaolivary nucleus 134-141
K
KF Kîlliker-Fuse nucleus 105-110
L
LA lateroanterior hypothalamic nucleus 40-45
La lat amygdaloid nucleus 68
LAcbSh lateral accumbens shell 15-26
LaDL lateral amygdaloid nucleus, dorsolateral part
50-67
Lat lateral (dentate) cerebellar nucleus 122-130
LatPC lateral cerebellar nucleus, parvicellular part
124-128
LaVL lateral amygdaloid nucleus, ventrolateral part
55-61
LaVM lateral amygdaloid nucleus, ventromedial
part 55-67
LC locus coeruleus 112-119
Ld lambdoid septal zone 23-30
LDDM laterodorsal thalamic nucleus, dorsomedial
part 50-59
LDTg laterodorsal tegmental nucleus 101-112
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LDTgV laterodorsal tegmental nucleus, ventral part LPMC lateral posterior thalamic nucleus, MDM mediodorsal thalamic nucleus, medial part MnA median accessory nucleus of the medulla
104-108 mediocaudal part 70-81 51-66 155-161
LDVL laterodorsal thalamic nucleus, ventrolateral LPMR lateral posterior thalamic nucleus, MdV medullary reticular nucleus, ventral part MnM medial mammillary nucleus, median part
part 48-61 mediorostral part 59-72 149-161 68-70
LEnt lateral entorhinal cortex 59-84, 104-107 LPO lateral preoptic area 28-42 ME median eminence 48 MnPO median preoptic nucleus 29-35
lfp longitudinal fasciculus of the pons 94-107 LPtA lateral parietal association cortex 59-67 Me5 mesencephalic trigeminal nucleus 86-107, MnR median raphe nucleus 93-106
LHb lateral habenular nucleus 52-56, 69 LR4V lateral recess of the 4th ventricle 113-139 109-116 MO medial orbital cortex 5-8
LHbL lateral habenular nucleus, lateral part LRt lateral reticular nucleus 142-157 me5 mesencephalic trigeminal tract 86-115
MoCb molecular layer of the cerebellum 120, 122,
57-68 LRtPC lateral reticular nucleus, parvicellular part MeAD medial amygdaloid nucleus, ant dorsal
125, 131, 134
LHbM lateral habenular nucleus, medial part 148-157 42-55
MoDG molecular layer of the dentate gyrus 46-90
57-68 LRtS5 lateral reticular nucleus, subtrigeminal part MeAV medial amygdaloid nucleus, anteroventral
MoS molecular layer of the subiculum 90-92
Li linear nucleus of the medulla 135-143 143-150 part 47-54
mp mammillary peduncle 70-83
ll lateral lemniscus 92-108 LSD lateral septal nucleus, dorsal part 16-41 Med medial (fastigial) cerebellar nucleus 124-133
MPA medial preoptic area 29-44
LM lateral mammillary nucleus 68-73 MedCM medial cerebellar nucleus, caudomedial
LSI lateral septal nucleus, intermediate part MPB medial parabrachial nucleus 105-116
LMol lacunosum moleculare layer of the part 133
14-37 MPBE medial parabrachial nucleus external part
hippocampus 54-89 MedDL medial cerebellar nucleus, dorsolateral
LSO lateral superior olive 112-122
LO lateral orbital cortex 5-13 protuberance 127-132 108-111
LSS lateral stripe of the striatum 14-38
lo lateral olfactory tract 3-44 MedL medial cerebellar nucleus, lateral part MPL medial paralemniscial nucleus 99-105
LSV lateral septal nucleus, ventral part 16-36
LOT nucleus of the lateral olfactory tract 37 127-131 MPO medial preoptic nucleus 43-44
LT lateral terminal nucleus of the accessory optic
LOT1 nucleus of the lateral olfactory tract, layer 1 MEE medial eminence, external layer 49-61 MPOC medial preoptic nucleus, central part
tract 71-78
38-45 MEI medial eminence, internal layer 49-61 37-39
LTer lemina terminalis 35-37
LPAG lateral periaqueductal gray 77-105 MEnt medial entorhinal cortex 85-105 MPOL medial preoptic nucleus, lateral part 34-40
Lth lithoid nucleus 71-76
LPB lateral parabrachial nucleus 105, 111-114 MEntV medial entorhinal cortex, ventral part MPOM medial preoptic nucleus, medial part
LV lateral ventricle 13-97
LPBC lateral parabrachial nucleus, central part 82-83 35-42
106-111 LVe lateral vestibular nucleus 122-127 MePD medial amygdaloid nucleus, posterodorsal MPT medial pretectal nucleus 70-77
LPBCr lateral parabrachial nucleus, crescent part LVPO lateroventral periolivary nucleus 109-119 part 53-63 MPtA medial parietal association cortex 60-67
107-110 MePV medial amygdaloid nucleus, posteroventral MRe mammillary recess of the 3rd ventricle 66-70
LPBD lateral parabrachial nucleus, dorsal part
107-110
M part 55-63
mfb medial forebrain bundle 12-31, 40-70
MS medial septal nucleus 20-33
M1 primary motor cortex 7-59 MSO medial superior olive 109-120
LPBE lateral parabrachial nucleus, external part mfba medial forebrain bundle, ‘a’ component
M2 secondary motor cortex 6-60 MT medial terminal nucleus of the accessory optic
106-111 32-39
m5 motor root of the trigeminal nerve 88-115 tract 74-79
LPBI lateral parabrachial nucleus, internal part mfbb medial forebrain bundle, ‘b’ component
MA3 medial accessory oculomotor nucleus 75-83 mt mammillothalamic tract 47-68
107-113 32-39
mcer middle cerebral artery 23, 25-27, 29, 32, mtg mammillotegmental tract 69-83
LPBM lateral parabrachial nucleus, medial part MGD medial geniculate nucleus, dorsal part 73-87
35-38 MTu medial tuberal nucleus 55-62
110 MGM medial geniculate nucleus, medial part
mch medial corticohypothalamic tract 38, 40-41 MVe medial vestibular nucleus 139-142
LPBS lateral parabrachial nucleus, superior part 74-87
105-106 MCLH magnocellular nucleus of the lateral MGV medial geniculate nucleus, ventral part MVeMC medial vestibular nucleus, magnocellular
LPBV lateral parabrachial nucleus, ventral part hypothalamus 54-59 73-88 part 115-138
107-113 mcp middle cerebellar peduncle 92-118 MHb medial habenular nucleus 48-70 MVePC medial vestibular nucleus, parvicellular part
LPGi lateral paragigantocellular nucleus 119-120, MCPC magnocellular nucleus of the posterior Mi mitral cell layer of the olfactory bulb 1-5 117-138
132-142 commissure 71-78 MiA mitral cell layer of the accessory olfactory bulb MVPO medioventral periolivary nucleus 105-121
LPGiA lateral paragigantocellular nucleus. alpha MCPO magnocellular preoptic nucleus 32-46 3-4 Mx matrix region of the medulla 129-158
part 121-131 MD mediodorsal thalamic nucleus 47-50 MiTg microcellular tegmental nucleus 90-101 MZMG marginal zone of the medial geniculate
LPGiE lateral paragigantocellular nucleus. external MDC mediodorsal thalamic nucleus, central part ML medial mammillary nucleus, lateral part 68-78 73-88
part 121-136 55-61 ml medial lemniscus 55-153
LPLC lateral posterior thalamic nucleus, MdD medullary reticular nucleus, dorsal part mld medial lemniscus decussation 148-155
laterocaudal part 71-78 149-161 mlf medial longitudinal fasciculus 78-161 N
LPLR lateral posterior thalamic nucleus, MDL mediodorsal thalamic nucleus, lateral part MM medial mammillary nucleus, medial part ns nigrostriatal bundle 47-72
laterorostral part 62-71 50-66 68-73 Nv navicular nucleus of the basal forebrain 10-17

The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson XXXIX
RBSC5 Book.qxp 16/9/04 16:36 Page XL
O PC5
Obex obex 153
oc olivocerebellar tract 129-136
ocb olivocochlear bundle 115-121
och optic chiasm 29-40
Op optic nerve layer of the superior colliculus
77-98
OPC oval paracentral thalamic nucleus 60-66
OPT olivary pretectal nucleus 70-78
opt optic tract 41-79
Or oriens layer of the hippocampus 47-89
OT nucleus of the optic tract 72-81
OV olfactory ventricle (olfactory part of lateral
ventricle) 1-5
P anterior parvicellular part 38-39
P3V preoptic recess of the 3rd ventricle 31
P5 peritrigeminal zone 107-116
P7 perifacial zone 120-131, 133-134
Pa4 paratrochlear nucleus 94-98
Pa5 paratrigeminal nucleus 141-150
Pa6 paraabducens nucleus 117-120
PaAP paraventricular hypothalamic nucleus,
anterior parvicellular part 40-44
PaC paracommissural nucleus of the posterior
commissure 76-80
PaDC paraventricular hypothalamic nucleus,
dorsal cap 47-49
PAG periaqueductal gray 74-76
PaLM paraventricular hypothalamic nucleus, lateral
magnocellular part 47-49
PaMM paraventricular hypothalamic nucleus,
medial magnocellular part 45-46
PaMP paraventricular hypothalamic nucleus,
medial parvicellular part 45-50
PaPo paraventricular hypothalamic nucleus,
posterior part 50-52
PaR pararubral nucleus 79-88
PaS parasubiculum 87-110
PaV paraventricular hypothalamic nucleus, ventral
part 45-49
PaXi paraxiphoid nucleus of thalamus 46-56
PBG parabigeminal nucleus 91-99
PBP parabrachial pigmented nucleus of the VTA
71-87
PC paracentral thalamic nucleus 44-66
pc posterior commissure 71-79
The Rat Brain in Stereotaxic Coordinates 5th Edition
parvicellular motor trigeminal nucleus
108-112
pcer posterior cerebral artery 85-87
PCGS paracochlear glial substance 111-118
PCom nucleus of the posterior commissure
73-76
PCRt parvicellular reticular nucleus 115-116,
128-148
PCRtA parvicellular reticular nucleus, alpha part
117-127
pcuf preculminate fissure 109-125
PDPO posterodorsal preoptic nucleus 38
PDR posterodorsal raphe nucleus 93-101
PDTg posterodorsal tegmental nucleus 111-115
Pe periventricular hypothalamic nucleus 35-62
PeAP periventricular hypothalamic nucleus,
PeF perifornical nucleus 55-63
PeFLH perifornical part of lateral hypothalamus
51-65
PF parafascicular thalamic nucleus 64-70
PFl paraflocculus 113-137
pfs parafloccular sulcus 119-121, 123-131,
133-135
PH posterior hypothalamic nucleus 61-69
PHA posterior hypothalamic area 70-71
PHD posterior hypothalamic area, dorsal part
57-64
Pi pineal gland 94-106
PIF parainterfascicular nucleus of the VTA 79-88
PIL posterior intralaminar thalamic nucleus 73-84
Pir piriform cortex 7-74
PiSt pineal stalk 92-93
Pk Purkinje cell layer of the cerebellum 120, 122,
125, 131, 134
PL paralemniscal nucleus 97-99
PLCo posterolateral cortical amygdaloid nucleus
51-53, 68-74
PLCo1 posterolateral cortical amygdaloid nucleus,
layer 1 54-67
PLd paralambdoid septal nucleus 22-29
plf posterolateral fissure 115-121, 123-124, 129,
132-148
PLH peduncular part of lateral hypothalamus
43- 71
PLi posterior limitans thalamic nucleus 70-81
PlPAG plioglial part of periaqueductai gray 77-80
PLV perilemniscal nucleus, ventral part 103-106
PM paramedian lobule 132-153
Paxinos & Watson
pm principal mammillary tract 69
PMCo posteromedial cortical amygdaloid nucleus
56-80
PMD premammillary nucleus, dorsal part 66-67
PMn paramedian reticular nucleus 139-144
PMnR paramedian raphe nucleus 92-106
pms paramedian sulcus 136-142, 144-146, 148
PMV premammillary nucleus, ventral part 63-66
PN paranigral nucleus of the VTA 76-87
Pn pontine nuclei 89-105
PnC pontine reticular nucleus, caudal part
108-118
PnO pontine reticular nucleus, oral part 93-107
PnR pontine raphe nucleus 107-108
PnV pontine reticular nucleus, ventral part
113-118
Po posterior thalamic nuclear group 54-81
PoDG polymorph layer of the dentate gyrus 51-87
POH periolivary horn 112-116
PoMn posteromedian thalamic nucleus 65-66
Post postsubiculum 83-103
PoT posterior thalamic nuclear group, triangular
part 74-84
PP peripeduncular nucleus 75-82
ppf prepyramidal fissure 137-150
PPT posterior pretectal nucleus 75-79
PPTg pedunculopontine tegmental nucleus
89-106
PPy parapyramidal nucleus 122-132
PR prerubral field 65-75
Pr prepositus nucleus 119-139
Pr5DM principal sensory trigeminal nucleus,
dorsomedial part 111-119
Pr5VL principal sensory trigeminal nucleus,
ventrolateral part 107-120
PrBo pre-Botzinger complex 138-140
PrC precommissural nucleus 67-72
PrCnF precuneiform area 91-100
prf primary fissure 111-131
PRh perirhinal cortex 58-111
PrL prelimbic cortex 6-13
PrS presubiculum 87-95
PS parastrial nucleus 32-37
psf posterior superior fissure 115-135, 140-143
PSol parasolitary nucleus 142-146
PSTh parasubthalamic nucleus 64-66
PT paratenial thalamic nucleus 41-51
PTe paraterete nucleus 51-59
PtPC parietal cortex, posterior area, caudal part
81-84
PtPD parietal cortex, posterior area, dorsal part
65-80
PtPR parietal cortex, posterior area, rostral part
68-80
PV paraventricular thalamic nucleus 51-57
PVA paraventricular thalamic nucleus, anterior part
40-50
PVG periventricular gray 68-73
PVP paraventricular thalamic nucleus, posterior
part 58-67
Py pyramidal cell layer of the hippocampus 48-89
py pyramidal tract 108-157
pyd pyramidal decussation 155-161
R
Rad radiatum layer of the hippocampus 49-89
RAmb retroambiguus nucleus 152-160
RAPir rostral amygdalopiriform area 57-63
Rbd rhabdoid nucleus 91-103
RCh retrochiasmatic area 42-47
RChL retrochiasmatic area, lateral part 45-54
Re reuniens thalamic nucleus 42-62
ReIC recess of the inferior colliculus 104-108
REth retroethmoid nucleus 72-77
rf rhinal fissure 4-111
Rh rhomboid thalamic nucleus 48-60
RI rostral interstitial nucleus of medial longitudinal
fasciculus 69-72
ri rhinal incisure 4-9
RIP raphe interpositus nucleus 113-119
RL retrolemniscal nucleus 105, 107
RLi rostral linear nucleus of the raphe 74-84, 95
RMC red nucleus, magnocellular part 78-88
RMg raphe magnus nucleus 113-134
Ro nucleus of Roller 135-150
ROb raphe obscurus nucleus 130-152
RPa raphe pallidus nucleus 114-152
RPC red nucleus, parvicellular part 76-86
RPF retroparafascicular nucleus 72-74
RR retrorubral nucleus 93-97
RRe retrouniens area 63-66
RRF retrorubral field 86-93
rs rubrospinal tract 86-161
RSD retrosplenial dysgranular cortex 47-111
RSG retrosplenial granular cortex 93
XL
RBSC5 Book.qxp 16/9/04 16:36 Page XLI
RSGa retrosplenial granular cortex, a region
79-104
RSGb retrosplenial granular cortex, b region
66-96
RSGc retrosplenial granular cortex, c region
47-90
Rt reticular thalamic nucleus 42-67
RtSt reticluostrial nucleus 42-52
RtTg reticulotegmental nucleus of the pons
98-113
RtTgL reticulotegmental nucleus of the pons, lateral
part 112-114
RtTgP reticulotegmental nucleus of the pons,
pericentral part 99-104
RVL rostroventrolateral reticular nucleus 133-137
RVRG rostral ventral respiratory group 141-151
S sm stria medullaris of the thalamus 38-69
S1 primary somatosensory cortex 65-71
S1BF primary somatosensory cortex, barrel field
36-69
S1DZ primary somatosensory cortex, dysgranular
zone 13-61
S1DZO primary somatosensory cortex, oral
dysgranular zone 14-23
S1FL primary somatosensory cortex, forelimb
region 13-45
S1HL primary somatosensory cortex, hindlimb
region 30-52
S1J primary somatosensory cortex, jaw region
10-23
S1Sh primary somatosensory cortex, shoulder
region 46-52
S1Tr primary somatosensory cortex, trunk region
53-61
S1ULp primary somatosensory cortex, upper lip
region 14-59
S2 secondary somatosensory cortex 23-65
s5 sensory root of the trigeminal nerve 88-117
Sag sagulum nucleus 102-106
Sc scaphoid thalamic nucleus 70-72
scc splenium of the corpus callosum 75-76
SCh suprachiasmatic nucleus 37-38
SChDL suprachiasmatic nucleus, dorsolateral part
40-41
SChVM suprachiasmatic nucleus, ventromedial part
40-41
SCO subcommissural organ 70-75
The Rat Brain in Stereotaxic Coordinates 5th Edition
scp superior cerebellar peduncle (brachium
conjunctivum) 61-72, 74-82, 87, 96, 98-129
scpd superior cerebellar peduncle, descending limb
86-95, 112-115
sf secondary fissure 136-155
SFi septofimbrial nucleus 32-42
SFO subfornical organ 38-48
SG suprageniculate thalamic nucleus 76-85
SGe supragenual nucleus 116-119
SHi septohippocampal nucleus 14-35
SHy septohypothalamic nucleus 28-37
SIB substantia innominata, basal part 24-40
Sim simple lobule 111-113, 124-135
SimA simple lobule A 114-123
SimB simple lobule B 114-123
simf simplex fissure 121-130
SLu stratum lucidum of the hippocampus 49-82
SM nucleus of the stria medullaris 42-43, 45
SMT submammillothalamic nucleus 66-68
SMV superior medullary velum 112, 114, 116,
122-123, 126, 128
SNCD substantia nigra, compact part, dorsal tier
71-88
SNCM substantia nigra, compact part, medial tier
81-88
SNCV substantia nigra, compacta part, ventral tier
81-88
SNL substantia nigra, lateral part 71-87
SNR substantia nigra, reticular part 69-89
SO supraoptic nucleus 33-48
sod supraoptic decussation 38-71
Sol nucleus of the solitary tract 125
sol solitary tract 128-157
SolC nucleus of the solitary tract, commissural part
147-161
SolCe nucleus of the solitary tract, central part
143-147
SolDL solitary nucleus, dorsolateral part 141-156
SolDM nucleus of the solitary tract, dorsomedial
part 131-140
SolG nucleus of the solitary tract, gelatinous part
144-146
SolI nucleus of the solitary tract, interstitial part
141-150
SolIM nucleus of the solitary tract, intermediate
part 126-151
SolL nucleus of the solitary tract, lateral part
130-143
Paxinos & Watson
SolM nucleus of the solitary tract, medial part
129-161
SolRL nucleus of the solitary tract, rostrolateral part
127-130
SolV solitary nucleus, ventral part 129-157
SolVL nucleus of the solitary tract, ventrolateral
part 135-157
SOR supraoptic nucleus, retrochiasmatic part
49-54
sp5 spinal trigeminal tract 118-161
Sp5C spinal trigeminal nucleus, caudal part
149-161
Sp5I spinal trigeminal nucleus, interpolar part
129-151
Sp5O spinal trigeminal nucleus, oral part 120-130
SPa subparaventricular zone of the hypothalamus
41-50
SPF subparafascicular thalamic nucleus 65-68
SPFPC subparafascicular thalamic nucleus,
parvicellular part 68-76
Sph sphenoid nucleus 111-113
SPO superior paraolivary nucleus 110-120
SPTg subpeduncular tegmental nucleus 96-104
SpVe spinal vestibular nucleus 125-142
st stria terminalis 37-67
StA strial part of the preoptic area 33-35
Stg stigmoid hypothalamic nucleus 50-52
STh subthalamic nucleus 59-68
StHy striohypothalamic nucleus 37-42
STI bed nucleus of the stria terminalis, intermediate
division 35
STIA bed nucleus of the stria terminalis,
intraamygdaloid division 52-63
STLD bed nucleus of the stria terminalis, lateral
division, dorsal part 31-35
STLI bed nucleus of the stria terminalis, lateral
division, intermediate part 36-38
STLJ bed nucleus of the stria terminalis, lateral
division, juxtacapsular part 34-36
STLP bed nucleus of the stria terminalis, lateral
division, posterior part 30-40
STLV bed nucleus of the stria terminalis, lateral
division, ventral part 30-37
STMA bed nucleus of the stria terminalis, medial
division, anterior part 26-34
STMAL bed nucleus of the stria terminalis, medial
division, anterolateral part 34-37
STMAM bed nucleus of the stria terminalis, medial
division, anteromedial part 34-37
STMP bed nucleus of the stria terminalis, medial
division, posterior part 42
STMPI bed nucleus of the stria terminalis, medial
division, posterointermediate part 38-42
STMPL bed nucleus of the stria terminalis, medial
division, posterolateral part 38-44
STMPM bed nucleus of the stria terminalis, medial
division, posteromedial part 38-45
STMV bed nucleus of the stria terminalis, medial
division, ventral part 30-36
STr subiculum, transition area 91-96
str superior thalamic radiation 66-73
STSL bed nucleus of stria terminalis, supracapsular
division, medial part 45-46
STSM bed nucleus of stria terminalis, supracapsular
division, lateralpart 45-46
Su3 supraoculomotor periaqueductal gray 83-92
Su3C supraoculomotor cap 84-92
Su5 supratrigeminal nucleus 106-113
Sub submedius thalamic nucleus 49-51, 57-60
SubB subbrachial nucleus 83-92
SubCA subcoeruleus nucleus, alpha part 109-112
SubCD subcoeruleus nucleus, dorsal part 107-116
SubCV subcoeruleus nucleus, ventral part 107-
116
SubD submedius thalamic nucleus, dorsal part
52-56
SubG subgeniculate nucleus 67-72
SubI subincertal nucleus 56-60
SubP subpostrema area 146-152
SubV submedius thalamic nucleus, ventral part
52-56
SuG superficial gray layer of the superior colliculus
77-98
SuM supramammillary nucleus 74
sumd supramammillary decussation 69-73
SuML supramammillary nucleus, lateral part
67-72
SuMM supramammillary nucleus, medial part
68-73
SuS superior salivatory nucleus 118-127
SuVe superior vestibular nucleus 114-123
T
Te terete hypothalamic nucleus 58-66
TeA temporal associatin cortex 71-106
tfp transverse fibers of the pons 88-91, 93-107
Tr triangular nucleus 98-105
XLI
RBSC5 Book.qxp 16/9/04 16:36 Page XLII

TS triangular septal nucleus 35-45 VCA ventral cochlear nucleus, anterior part VLL ventral nucleus of the lat lemniscus 97-108 vsc ventral spinocerebellar tract 105-161
ts tectospinal tract 85-161 109-121 VLPAG ventrolateral periaqueductal gray 88-106 vscd ventral spinocerebellar tract decussation 123
tth trigeminothalamic tract 83-117, 120-128 VCAGr ventral cochlear nucleus, granule cell layer VLPO ventrolateral preoptic nucleus 33-38 VTA ventral tegmental area 89-90
Tu olfactory tubercle 10-35 115-118, 120 VM ventromedial thalamic nucleus 48-64 VTAR ventral tegmental area, rostral part 72-75
TuLH tuberal region of lateral hypothalamus VCCap ventral cochlear nucleus, capsular part VMH ventromedial hypothalamic nucleus 48-49, VTg ventral tegmental nucleus 100-105
46-62 122-125 60-61 vtgd ventral tegmental decussation 78-86
Tz nucleus of the trapezoid body 107-120 VCl ventral part of claustrum 15-49 VMHA ventromedial hypothalamic nucleus, VTM ventral tuberomammillary nucleus 64-74
tz trapezoid body 105-125 VCP ventral cochlear nucleus, posterior part anterior part 47 VTT ventral tenia tecta 7, 10
tzd decussation of the trapezoid body 118 120-122, 125-127 VMHC ventromedial hypothalamic nucleus, central
VTT1 ventral tenia tecta, layer 1 8-9
VCPO ventral cochlear nucleus, posterior part, part 50-59
octopus cell area 123-125 VMHDM ventromedial hypothalamic nucleus,
U VDB nucleus of the vertical limb of the diagonal dorsomedial part 50-59 X
un uncinate fasciculus of the cerebellum 115-122
band 18-29 VMHSh ventromedial nucleus of the hypothalamus X nucleus X 125-139
und uncinate fasciculus decussation 123-128
VeCb vestibulocerebellar nucleus 122-126 shell 47-61 Xi xiphoid thalamic nucleus 45-48, 51-53
veme vestibulomesencephalic tract 116-122 VMHVL ventromedial hypothalamic nucleus,
V VEn ventral endopiriform nucleus 40-58 ventrolateral part 50-59
v vein 92-93, 95, 98, 101-102, 105-107 vert vertebral artery 137, 139-145, 147-161 VMPO ventromedial preoptic nucleus 31-36, Y
V1 primary visual cortex 70-81 vesp vestibulospinal tract 123-126 107-109 Y nucleus Y 124-127
V1B primary visual cortex, binocular area vhc ventral hippocampal commissure 38-48 VO ventral orbital cortex 5-11
82-111 VIEnt ventral intermediate entorhinal cortex VOLT vascular organ of the lamina terminalis
V1M primary visual cortex, monocular area 85-100 28-33
Z
VL ventrolateral thalamic nucleus 48-59 VP ventral pallidum 11-42 Z nucleus Z 140-142
82-111
VLG ventral lateral geniculate nucleus 60-63, VPL ventral posterolateral thalamic nucleus 51-70 ZI zona incerta 54-55
V2L secondary visual cortex, lateral area 79-111
V2ML secondary visual cortex, mediolateral area 65-71, 73-77 VPM ventral posteromedial thalamic nucleus ZIC zona incerta, caudal part 70-76
68-98 VLG1 ventral lateral geniculate nucleus, layer 1 54-73 ZID zona incerta, dorsal part 56-69
V2MM secondary visual cortex, mediomedial area 64, 72 VPPC ventral posterior nucleus of the thalamus, ZIR zona incerta, rostral part 49-53
68-107 VLH ventrolateral hypothalamic nucleus 39-46 parvicellular part 62-69 ZIV zona incerta, ventral part 56-69
VA ventral anterior thalamic nucleus 46, 48-52 vlh ventrolateral hypothalamic tract 47-56 VRe ventral reuniens thalamic nucleus 43-61 ZL zona limitans 30-32
VA/VL region where VA and VL overlap 47 VLi ventral linear nucleus of the thalamus 69-74 VS ventral subiculum 71-90 Zo zona layer of the superior colliculus 77-99

The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson XLII
RBSC5 Book.qxp 16/9/04 16:36 Page XLIII

Figures

Coronal sections of the brain Figures 1-161

Transverse sections of the spinal cord Figures 162a, b, c, d, e

The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson XLIII
RBSC5 Book.qxp 16/9/04 16:36 Page 1

4 3 2 1 0 1 2 3 4

+5 0 -5 -10 -15
10 Bregma 0
Figure 1
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
7 3

The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
EPl Gl
6 Mi 4
E
IPl
GrO

OV
5 5

4 6

3 7

2 8

Interaural 16.56 mm Bregma 7.56 mm

4 3 2 1 0 1 2 3 4

E ependyma/subepen IPl int plexi olf bulb

EPl ext plex olf bulf Mi mitral olf bulb
Gl glomerular olf b OV olfact ventricle
GrO granular olf bulb
RBSC5 Book.qxp 16/9/04 16:36 Page 2

aci ac intrabulbar GrO granular olf bulb

E ependyma/subepen IPl int plexi olf bulb
EPl ext plex olf bulf Mi mitral olf bulb
Gl glomerular olf b OV olfact ventricle
GrA gran acc olf bulb

4 3 2 1 0 1 2 3 4

+5 0 -5 -10 -15
10 Bregma 0
Figure 2
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

Mi
EPl
IPl
Gl
6 GrA 4
aci

E
GrO
OV
5 5

4 6

3 7

2 8

Interaural 16.08 mm Bregma 7.08 mm

4 3 2 1 0 1 2 3 4
RBSC5 Book.qxp 16/9/04 16:36 Page 3

4 3 2 1 0 1 2 3 4

+5 0 -5 -10 -15
10 Bregma 0
Figure 3
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
7 3
Mi EPl
IPl
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson
Gl
EPlA
6 4
dlo
MiA
aci
GrA
GrO E
5 5
OV AOL
AOE
lo

4 6

3 7

2 8

Interaural 15.60 mm Bregma 6.60 mm

4 3 2 1 0 1 2 3 4

aci ac intrabulbar GrA gran acc olf bulb

AOE ant olfact ext GrO granular olf bulb
AOL ant olfact lat IPl int plexi olf bulb
dlo dorsal lat olf tr lo lat olfactory tr
E ependyma/subepen Mi mitral olf bulb
EPl ext plex olf bulf MiA mitral acc olf bulb
EPlA ext plex acc olf b OV olfact ventricle
Gl glomerular olf b
RBSC5 Book.qxp 16/9/04 16:36 Page 4

aci ac intrabulbar
AOD ant olf dorsal
AOE ant olfact ext
AOL ant olfact lat
AOM ant olfact medial
AOV ant olfact vent
dlo dorsal lat olf tr GlA glom acc olf bulb MiA mitral acc olf bulb
E ependyma/subepen GrO granular olf bulb OV olfact ventricle
EPl ext plex olf bulf GrA gran acc olf bulb rf rhinal fissure
EPlA ext plex acc olf b IPl int plexi olf bulb ri rhinal incisure
FrA frontal assocn cx lo lat olfactory tr
Gl glomerular olf b Mi mitral olf bulb

4 3 2 1 0 1 2 3 4

+5 0 -5 -10 -15
10 Bregma 0
Figure 4
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
7 3
FrA
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
rf
GlA
ri
EPlA
MiA
AOE
5 dlo Gl AOD 5
GrA
IPl AOM
Mi GrO AOL
aci EPl OV E
lo
4 6
AOV

3 7

2 8

Interaural 15.12 mm Bregma 6.12 mm

4 3 2 1 0 1 2 3 4
RBSC5 Book.qxp 16/9/04 16:36 Page 5

4 3 2 1 0 1 2 3 4

+5 0 -5 -10 -15
10 Bregma 0
Figure 5
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
FrA
7 3

The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
MO LO

GlA
VO
rf
5 AOE 5
ri AOD
GrA
IPl GrO
aci AOL
E OV AOM
4 6
lo
AOV
Gl
AOE
3 Mi EPl 7

2 8

Interaural 14.64 mm Bregma 5.64 mm

4 3 2 1 0 1 2 3 4

aci ac intrabulbar E ependyma/subepen GrO granular olf bulb OV olfact ventricle

AOD ant olf dorsal EPl ext plex olf bulf IPl int plexi olf bulb rf rhinal fissure
AOE ant olfact ext FrA frontal assocn cx LO lat orbital cx ri rhinal incisure
AOL ant olfact lat Gl glomerular olf b lo lat olfactory tr VO ventral orbital cx
AOM ant olfact medial GlA glom acc olf bulb Mi mitral olf bulb
AOV ant olfact vent GrA gran acc olf bulb MO medial orbital cx
RBSC5 Book.qxp 16/9/04 16:36 Page 6

aci ac intrabulbar AOVP ant olf ventropost LO lat orbital cx PrL prelimbic cx
AOD ant olf dorsal DLO dorsolat orbital cx lo lat olfactory tr rf rhinal fissure
AOL ant olfact lat E/OV epend/olf ventr M2 2ary motor cx ri rhinal incisure
AOM ant olfact medial GrO granular olf bulb MO medial orbital cx VO ventral orbital cx

6 5 4 3 2 1 0 1 2 3 4 5 6

+5 0 -5 -10 -15
10 Bregma 0
Figure 6
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

M2

7 3
PrL
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
DLO
MO LO

VO
5 5

rf AOD

ri
4 aci
E/OV 6
AOM AOL
GrO
lo AOVP

3 7

2 8

1 9

Interaural 14.16 mm Bregma 5.16 mm

6 5 4 3 2 1 0 1 2 3 4 5 6
RBSC5 Book.qxp 16/9/04 16:36 Page 7

6 5 4 3 2 1 0 1 2 3 4 5 6

+5 0 -5 -10 -15
10 Bregma 0
Figure 7
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 M2
2

7 3
M1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

PrL

Fr3
6 4

LO DLO
VO
MO
5 5

rf AOD
ri
4 6
aci AOM AOL 2
lo E/OV Pir1
AOVP
3 7
VTT

2 8

1 9

Interaural 13.68 mm Bregma 4.68 mm

6 5 4 3 2 1 0 1 2 3 4 5 6

2 layer 2 cortex AOVP ant olf ventropost lo lat olfactory tr PrL prelimbic cx
aci ac intrabulbar DLO dorsolat orbital cx M1 primary motor cx rf rhinal fissure
AOD ant olf dorsal E/OV epend/olf ventr M2 2ary motor cx ri rhinal incisure
AOL ant olfact lat Fr3 frontal area 3 MO medial orbital cx VO ventral orbital cx
AOM ant olfact medial LO lat orbital cx Pir1 piriform layer 1 VTT ventral tenia tecta
RBSC5 Book.qxp 16/9/04 16:36 Page 8

1b layer 1b cortex AOL ant olfact lat fmi forceps minor Pir1 piriform layer 1
2 layer 2 cortex AOVP ant olf ventropost Fr3 frontal area 3 PrL prelimbic cx
3 layer 3 cortex Cg1 cingulate area 1 IEn intermed endopir ri rhinal incisure
4 layer 4 cortex Cl claustrum LO lat orbital cx VO ventral orbital cx
a artery DEn dorsal endopirif lo lat olfactory tr VTT1 vent TT layer 1
aca ant comm, ant DTr dorsal trans zone M1 primary motor cx
AID ant insular dorsal DTT1 dors TT layer 1 M2 2ary motor cx
AIV ant insular ventral E/OV epend/olf ventr MO medial orbital cx

6 5 4 3 2 1 0 1 2 3 4 5 6

+5 0 -5 -10 -15
10 Bregma 0
Figure 8
5 5
9 1

0 Interaural 10
M2
+15 +10 +5 0 -5
8 2
Cg1
M1

7 3
fmi
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

PrL
Cl Fr3
6 4

AID
LO
5 VO 5
MO
AIV

a
4 ri 6
DTr DTT1 DEn
rf E/OV
3 2 1b Pir1a
aca 4
2 3 IEn

AOVP
3 lo AOL 7
3
2
VTT1

2 8

1 9

Interaural 13.20 mm Bregma 4.20 mm

6 5 4 3 2 1 0 1 2 3 4 5 6
RBSC5 Book.qxp 16/9/04 16:36 Page 9

6 5 4 3 2 1 0 1 2 3 4 5 6

+5 0 -5 -10 -15
10 Bregma 0
Figure 9
5 5
9 1

0 Interaural 10 M2
+15 +10 +5 0 -5
8 2
Cg1
M1

7 3

fmi
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

PrL
6 4
Cl Fr3

AID
5 IL 5
VO AIV
LO
DP
a DTr
4 6
ri E/OV
DTT DEn
1 2 3 4
rf
aca
AOVP IEn
3 3 7
lo 2
3 AOP Pir1
2
VTT1

2 8

1 9

Interaural 12.72 mm Bregma 3.72 mm

6 5 4 3 2 1 0 1 2 3 4 5 6

2 layer 2 cortex AOVP ant olf ventropost fmi forceps minor Pir1 piriform layer 1
3 layer 3 cortex Cg1 cingulate area 1 Fr3 frontal area 3 PrL prelimbic cx
4 layer 4 cortex Cl claustrum IEn intermed endopir rf rhinal fissure
a artery DEn dorsal endopirif IL infralimbic cx ri rhinal incisure
aca ant comm, ant DP dorsal pedunc cx LO lat orbital cx VO ventral orbital cx
AID ant insular dorsal DTr dorsal trans zone lo lat olfactory tr VTT1 vent TT layer 1
AIV ant insular ventral DTT dorsal tenia tecta M1 primary motor cx
AOP ant olfact post E/OV epend/olf ventr M2 2ary motor cx
RBSC5 Book.qxp 16/9/04 16:36 Page 10

2 layer 2 cortex azp azyg perical art Fr3 frontal area 3 Pir1 piriform layer 1
3 layer 3 cortex Cg1 cingulate area 1 ICj islands of Calleja PrL prelimbic cx
4 layer 4 cortex Cl claustrum IEn intermed endopir rf rhinal fissure
aca ant comm, ant DEn dorsal endopirif IL infralimbic cx S1J S1 cx, jaw region
AcbR accumbens rost DI dysgran insular LO lat orbital cx Tu1 olf tub layer 1
AID ant insular dorsal DP dorsal pedunc cx lo lat olfactory tr VO ventral orbital cx
AIV ant insular ventral DTT dorsal tenia tecta M1 primary motor cx VTT ventral tenia tecta
AOP ant olfact post E/OV epend/olf ventr M2 2ary motor cx
azac azyg ant cer art fmi forceps minor Nv navicular nu

6 5 4 3 2 1 0 1 2 3 4 5 6

+5 0 -5 -10 -15
10 Bregma 0
Figure 10
5 5
9 1

0 Interaural 10
M2
+15 +10 +5 0 -5
M1
8 Cg1 2

Fr3

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

PrL
fmi S1J

6 4

Cl
DI
5 IL 5
AID
LO AIV
VO
azp DP
E/OV
4 azac 6
DTT 4
1 2 3
AcbR DEn
rf aca
3 VTT AOP IEn 7
3 2
Pir1
lo Nv

3
ICj 2
2 Tu1 8

1 9

Interaural 12.24 mm Bregma 3.24 mm

6 5 4 3 2 1 0 1 2 3 4 5 6
RBSC5 Book.qxp 16/9/04 16:36 Page 11

6 5 4 3 2 1 0 1 2 3 4 5 6

+5 0 -5 -10 -15
10 Bregma 0
Figure 11
5 5
9 1

0 Interaural 10
M2
+15 +10 +5 0 -5 M1
8 2
Cg1
Fr3

7 cg 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

PrL S1J
fmi
6 4

Cl
5 IL DI 5

azp
AID
VO LO
DP AIV
4 6
DTT E/OV
1 2 3 AcbC
AcbSh DEn
rf
aca
3 azac AOP IEn 7
3 2
Nv Pir1
lo
VP VP VP
3
2 2 8
Tu1

1 9

Interaural 12.00 mm Bregma 3.00 mm

6 5 4 3 2 1 0 1 2 3 4 5 6

2 layer 2 cortex azp azyg perical art Fr3 frontal area 3 PrL prelimbic cx
3 layer 3 cortex cg cingulum IEn intermed endopir rf rhinal fissure
aca ant comm, ant Cl claustrum IL infralimbic cx S1J S1 cx, jaw region
AcbC accumbens core DEn dorsal endopirif LO lat orbital cx Tu1 olf tub layer 1
AcbSh accumbens shell DI dysgran insular lo lat olfactory tr VO ventral orbital cx
AID ant insular dorsal DP dorsal pedunc cx M1 primary motor cx VP ventral pallidum
AIV ant insular ventral DTT dorsal tenia tecta M2 2ary motor cx
AOP ant olfact post E/OV epend/olf ventr Nv navicular nu
azac azyg ant cer art fmi forceps minor Pir1 piriform layer 1
RBSC5 Book.qxp 16/9/04 16:36 Page 12

2 layer 2 cortex cg cingulum fmi forceps minor M2 2ary motor cx

3 layer 3 cortex Cg1 cingulate area 1 Fr3 frontal area 3 mfb med forebr bundle
aca ant comm, ant Cl claustrum GI granular insular Nv navicular nu
AcbC accumbens core CPu caudate putamen ICj islands of Calleja Pir1 piriform layer 1
AcbSh accumbens shell DEn dorsal endopirif IEn intermed endopir PrL prelimbic cx
AID ant insular dorsal DI dysgran insular IL infralimbic cx S1J S1 cx, jaw region
AIV ant insular ventral DP dorsal pedunc cx LO lat orbital cx Tu1 olf tub layer 1
azac azyg ant cer art DTT dorsal tenia tecta lo lat olfactory tr VP ventral pallidum
azp azyg perical art E/OV epend/olf ventr M1 primary motor cx

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 12
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1

8 Fr3 2
Cg1

cg
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 3

fmi S1J

PrL
6 CPu 4

5 azp Cl 5
IL
GI

E/OV
DI
DP
DTT AID
4 LO AIV 6
1 2 3
AcbC
DEn
rf
aca
Nv 3
3 AcbSh 7
IEn 2
azac
Pir1
ICj
lo
mfb VP
3
2 2 8
Tu1

1 9

Interaural 11.76 mm Bregma 2.76 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 13

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 13
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1
8 2
Cg1

S1FL
cg
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 S1DZ 3
fmi
PrL

6 S1J 4
CPu
azp

IL
5 5
E GI
DP Cl
DI
LV
4 DTT 12 3 AID 6
AIV
LO
AcbC

Nv
3 aca DEn 7
rf
acer AcbSh IEn 3
2
Pir1
ICj
lo mfb
2 ICj VP 8
3
2
Tu1

1 9

Interaural 11.52 mm Bregma 2.52 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex Cg1 cingulate area 1 GI granular insular mfb med forebr bundle
3 layer 3 cortex Cl claustrum ICj islands of Calleja Nv navicular nu
aca ant comm, ant CPu caudate putamen IEn intermed endopir Pir1 piriform layer 1
AcbC accumbens core DEn dorsal endopirif IL infralimbic cx PrL prelimbic cx
AcbSh accumbens shell DI dysgran insular LO lat orbital cx S1DZ S1 dysgranular zn
AID ant insular dorsal DP dorsal pedunc cx lo lat olfactory tr S1FL S1 forelimb region
AIV ant insular ventral DTT dorsal tenia tecta LV lat ventricle S1J S1 cx, jaw region
azp azyg perical art E ependyma/subepen M1 primary motor cx Tu1 olf tub layer 1
cg cingulum fmi forceps minor M2 2ary motor cx VP ventral pallidum
RBSC5 Book.qxp 16/9/04 16:36 Page 14

2 layer 2 cortex
3 layer 3 cortex
aca ant comm, ant
AcbC accumbens core
AcbSh accumbens shell
AID ant insular dorsal
AIV ant insular ventral
azp azyg perical art DI dysgran insular ICj islands of Calleja M1 primary motor cx S1DZO S1 oral dysgran zn
cg cingulum DP dorsal pedunc cx IEn intermed endopir M2 2ary motor cx S1FL S1 forelimb region
Cg1 cingulate area 1 DTT dorsal tenia tecta IG indusium griseum mfb med forebr bundle S1J S1 cx, jaw region
Cg2 cingulate area 2 E ependyma/subepen lo lat olfactory tr Nv navicular nu S1ULp S1 upper lip region
Cl claustrum ec external capsule LSI lat septal intermed Pir1 piriform layer 1 SHi septohipp nu
CPu caudate putamen gcc genu of corp call LSS lat stripe of str rf rhinal fissure Tu1 olf tub layer 1
DEn dorsal endopirif GI granular insular LV lat ventricle S1DZ S1 dysgranular zn VP ventral pallidum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 14
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1
8 Cg1 2

S1FL

cg S1DZ
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 3
Cg2
S1J

6 IG S1DZO 4

azp
ec gcc
E S1ULp
5 CPu 5
DP LV
DTT
GI
1 2 Cl
LSI
4 DI 6
SHi AID

AcbC AIV
Nv
3 DEn 7
aca
rf
AcbSh
IEn 3
2
VP LSS Pir1
acer
VP
2 lo mfb VP 8
3
ICj
2
Tu1

1 9

Interaural 11.28 mm Bregma 2.28 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 15

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 15
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1

8 Cg1 2
S1FL

S1DZ
cg
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 3
Cg2
S1J

S1DZO
6 IG 4
ec azp
gcc
E S1ULp
5 CPu 5
LV
DP
12 3 DCl
LSI
4 GI 6
VCl
SHi DI

AcbC AID
Nv AcbSh
AIV
3 DEn 7
aca
ICjM
rf ICj
3
IEn 2
LAcbSh LSS Pir1
VP
2 acer VP 8
mfb
lo mfb ICj
3
2
Tu1
1 9

Interaural 11.16 mm Bregma 2.16 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex
3 layer 3 cortex
aca ant comm, ant
AcbC accumbens core
AcbSh accumbens shell
AID ant insular dorsal
AIV ant insular ventral
azp azyg perical art
cg cingulum E ependyma/subepen LAcbSh lat accumb shell Nv navicular nu SHi septohipp nu
Cg1 cingulate area 1 ec external capsule lo lat olfactory tr Pir1 piriform layer 1 Tu1 olf tub layer 1
Cg2 cingulate area 2 gcc genu of corp call LSI lat septal intermed rf rhinal fissure VCl ventral claustrum
CPu caudate putamen GI granular insular LSS lat stripe of str S1DZ S1 dysgranular zn VP ventral pallidum
DCl dorsal claustrum ICj islands of Calleja LV lat ventricle S1DZO S1 oral dysgran zn
DEn dorsal endopirif ICjM major is Calleja M1 primary motor cx S1FL S1 forelimb region
DI dysgran insular IEn intermed endopir M2 2ary motor cx S1J S1 cx, jaw region
DP dorsal pedunc cx IG indusium griseum mfb med forebr bundle S1ULp S1 upper lip region
RBSC5 Book.qxp 16/9/04 16:36 Page 16

2 layer 2 cortex
3 layer 3 cortex
aca ant comm, ant
AcbC accumbens core
AcbSh accumbens shell
AID ant insular dorsal
AIV ant insular ventral
azp azyg perical art
cg cingulum ec external capsule lo lat olfactory tr mfb med forebr bundle S1ULp S1 upper lip region
Cg1 cingulate area 1 gcc genu of corp call LSD lat septal dors Nv navicular nu SHi septohipp nu
Cg2 cingulate area 2 GI granular insular LSI lat septal intermed Pir1 piriform layer 1 Tu1 olf tub layer 1
CPu caudate putamen ICj islands of Calleja LSS lat stripe of str rf rhinal fissure VCl ventral claustrum
DCl dorsal claustrum ICjM major is Calleja LSV lat septal vent S1DZ S1 dysgranular zn VP ventral pallidum
DEn dorsal endopirif IEn intermed endopir LV lat ventricle S1DZO S1 oral dysgran zn
DI dysgran insular IG indusium griseum M1 primary motor cx S1FL S1 forelimb region
E ependyma/subepen LAcbSh lat accumb shell M2 2ary motor cx S1J S1 cx, jaw region

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 16
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1
M2
M1

8 Cg1 2
S1FL

cg Z
D
S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 S1J
3

O
S1DZ
6 IG 4
azp
gcc E
ec
CPu S1ULp
LV
5 5
1 3 LSD
IG
2 DCl
LSV
GI
4 LSI VCl 6
SHi DI

AID
AcbSh AcbC
Nv AIV
3 rf ICjM 7
aca DEn
Tu LSS
ICj 3
LAcbSh 2
IEn Pir1
2 acer 8
mfb VP VP
lo mfb

3
2
Tu1
1 9
ICj

Interaural 11.04 mm Bregma 2.04 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 17

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 17
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1
Cg1
8 2
S1FL

cg
DZ
S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3
S1J

IG
O
6 S1DZ 4
azp

gcc E
ec LSD
S1ULp
5 LV 5
IG 1 3 CPu
2

DCl
SHi LSV GI
4 6
LSI VCl
DI

AID
AcbC
Nv
3 AIV 7
aca
DEn
VP LSS
rf ICj
AcbSh 3
LAcbSh IEn 2
ICj
2 Pir1 8
acer
mfb VP
lo
ICj 3
2
1 Tu1 9

Interaural 10.92 mm Bregma 1.92 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex
3 layer 3 cortex
aca ant comm, ant
AcbC accumbens core
AcbSh accumbens shell
AID ant insular dorsal
AIV ant insular ventral
azp azyg perical art
cg cingulum ec external capsule LSD lat septal dors Nv navicular nu SHi septohipp nu
Cg1 cingulate area 1 gcc genu of corp call LSI lat septal intermed Pir1 piriform layer 1 Tu1 olf tub layer 1
Cg2 cingulate area 2 GI granular insular LSS lat stripe of str rf rhinal fissure VCl ventral claustrum
CPu caudate putamen ICj islands of Calleja LSV lat septal vent S1DZ S1 dysgranular zn VP ventral pallidum
DCl dorsal claustrum IEn intermed endopir LV lat ventricle S1DZO S1 oral dysgran zn
DEn dorsal endopirif IG indusium griseum M1 primary motor cx S1FL S1 forelimb region
DI dysgran insular LAcbSh lat accumb shell M2 2ary motor cx S1J S1 cx, jaw region
E ependyma/subepen lo lat olfactory tr mfb med forebr bundle S1ULp S1 upper lip region
RBSC5 Book.qxp 16/9/04 16:36 Page 18

2 layer 2 cortex
3 layer 3 cortex
aca ant comm, ant
AcbC accumbens core
AcbSh accumbens shell
AID ant insular dorsal
AIV ant insular ventral
azp azyg perical art
cg cingulum ec external capsule lo lat olfactory tr mfb med forebr bundle SHi septohipp nu
Cg1 cingulate area 1 gcc genu of corp call LSD lat septal dors Pir piriform cx Tu1 olf tub layer 1
Cg2 cingulate area 2 GI granular insular LSI lat septal intermed rf rhinal fissure VCl ventral claustrum
CPu caudate putamen ICj islands of Calleja LSS lat stripe of str S1DZ S1 dysgranular zn VDB nu vert limb diag b
DCl dorsal claustrum ICjM major is Calleja LSV lat septal vent S1DZO S1 oral dysgran zn VP ventral pallidum
DEn dorsal endopirif IEn intermed endopir LV lat ventricle S1FL S1 forelimb region
DI dysgran insular IG indusium griseum M1 primary motor cx S1J S1 cx, jaw region
E ependyma/subepen LAcbSh lat accumb shell M2 2ary motor cx S1ULp S1 upper lip region

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 18
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1
Cg1
8 2
S1FL

cg
DZ
S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 3
Cg2 S1J

ZO
IG S1D
6 4
azp
E
gcc
S1ULp
LV CPu
5 ec LSD 5
IG 1 3
2
DCl
LSV
GI
SHi
4 LSI DI
6
VCl
AID
AcbC AIV
VDB ICjM DEn
3 7
aca AcbSh
rf
ICj VP
LSS IEn 3
2
1
LAcbSh Pir
2 acer
8
lo VP
mfb
ICj

1 9

Interaural 10.80 mm Bregma 1.80 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 19

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 19
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1
Cg1
8 2
S1FL

cg
DZ
S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3
S1J

IG ZO
ec S1D
6 4
E
gcc

LV S1ULp
5 LSD 5
CPu
SHi
LSV
DCl GI
4 LSI 6
VCl DI

AID
AcbC
3 VDB ICjM AIV 7
aca DEn
rf AcbSh

ICj
LSS IEn 3
2
2 LAcbSh 1 8
VP Pir
VP
lo mfb acer VP
3 ICj
2
1 Tu1 9

Interaural 10.68 mm Bregma 1.68 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex Cg1 cingulate area 1 gcc genu of corp call LSD lat septal dors Pir piriform cx Tu1 olf tub layer 1
3 layer 3 cortex Cg2 cingulate area 2 GI granular insular LSI lat septal intermed rf rhinal fissure VCl ventral claustrum
aca ant comm, ant CPu caudate putamen ICj islands of Calleja LSS lat stripe of str S1DZ S1 dysgranular zn VDB nu vert limb diag b
AcbC accumbens core DCl dorsal claustrum ICjM major is Calleja LSV lat septal vent S1DZO S1 oral dysgran zn VP ventral pallidum
AcbSh accumbens shell DEn dorsal endopirif IEn intermed endopir LV lat ventricle S1FL S1 forelimb region
AID ant insular dorsal DI dysgran insular IG indusium griseum M1 primary motor cx S1J S1 cx, jaw region
AIV ant insular ventral E ependyma/subepen LAcbSh lat accumb shell M2 2ary motor cx S1ULp S1 upper lip region
cg cingulum ec external capsule lo lat olfactory tr mfb med forebr bundle SHi septohipp nu
RBSC5 Book.qxp 16/9/04 16:36 Page 20

2 layer 2 cortex
3 layer 3 cortex
aca ant comm, ant
AcbC accumbens core
AcbSh accumbens shell
AID ant insular dorsal
AIV ant insular ventral
Cg1 cingulate area 1
Cg2 cingulate area 2 GI granular insular LSI lat septal intermed Pir piriform cx Tu1 olf tub layer 1
CPu caudate putamen ICj islands of Calleja LSS lat stripe of str rf rhinal fissure VCl ventral claustrum
DCl dorsal claustrum ICjM major is Calleja LSV lat septal vent S1DZ S1 dysgranular zn VDB nu vert limb diag b
DEn dorsal endopirif IEn intermed endopir LV lat ventricle S1DZO S1 oral dysgran zn VP ventral pallidum
DI dysgran insular IG indusium griseum M1 primary motor cx S1FL S1 forelimb region
E ependyma/subepen LAcbSh lat accumb shell M2 2ary motor cx S1J S1 cx, jaw region
ec external capsule lo lat olfactory tr mfb med forebr bundle S1ULp S1 upper lip region
gcc genu of corp call LSD lat septal dors MS medial septal nu SHi septohipp nu

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 20
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1
Cg1
8 2
S1FL

cg DZ
S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3
S1J

IG ZO
S1D
6 4
ec E
gcc

LV S1ULp
LSD
5 5
SHi CPu

LSV

4 LSI GI 6
DCl
DI
VCl
MS
AID
3 7
aca ICjM AIV
VDB DEn
rf AcbC
AcbSh LSS
IEn 3
LAcbSh
2 2 8
1
ICj VP Pir
mfb acer VP
lo
ICj
3
2
1 Tu1 9

Interaural 10.56 mm Bregma 1.56 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 21

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 21
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1
8 Cg1 2
S1FL

Z
D
cg

S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 S1J 3

ZO
IG S1D

6 E 4
cc

ec LSD S1ULp
LV
5 SHi 5
CPu

LSI LSV
4 GI 6
DCl
DI
VCl
MS AID
ICjM
3 7
aca AcbSh DEn AIV
VDB

rf AcbC
LSS 3
IEn
2 2 8
ICj LAcbSh
1
Pir
VP
lo mfb
ICj
1 3 9
2
Tu1

Interaural 10.44 mm Bregma 1.44 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex
3 layer 3 cortex
aca ant comm, ant
AcbC accumbens core
AcbSh accumbens shell
AID ant insular dorsal
AIV ant insular ventral
cc corpus callosum
cg cingulum ec external capsule LSD lat septal dors MS medial septal nu SHi septohipp nu
Cg1 cingulate area 1 GI granular insular LSI lat septal intermed Pir piriform cx Tu1 olf tub layer 1
Cg2 cingulate area 2 ICj islands of Calleja LSS lat stripe of str rf rhinal fissure VCl ventral claustrum
CPu caudate putamen ICjM major is Calleja LSV lat septal vent S1DZ S1 dysgranular zn VDB nu vert limb diag b
DCl dorsal claustrum IEn intermed endopir LV lat ventricle S1DZO S1 oral dysgran zn VP ventral pallidum
DEn dorsal endopirif IG indusium griseum M1 primary motor cx S1FL S1 forelimb region
DI dysgran insular LAcbSh lat accumb shell M2 2ary motor cx S1J S1 cx, jaw region
E ependyma/subepen lo lat olfactory tr mfb med forebr bundle S1ULp S1 upper lip region
RBSC5 Book.qxp 16/9/04 16:36 Page 22

2 layer 2 cortex
3 layer 3 cortex
aca ant comm, ant
AcbC accumbens core
AcbSh accumbens shell
AID ant insular dorsal
AIV ant insular ventral
cc corpus callosum
cg cingulum ec external capsule LSD lat septal dors MS medial septal nu S1ULp S1 upper lip region
Cg1 cingulate area 1 GI granular insular LSI lat septal intermed Pir1 piriform layer 1 SHi septohipp nu
Cg2 cingulate area 2 ICj islands of Calleja LSS lat stripe of str PLd paralambdoid sept Tu1 olf tub layer 1
CPu caudate putamen ICjM major is Calleja LSV lat septal vent rf rhinal fissure VCl ventral claustrum
DCl dorsal claustrum IEn intermed endopir LV lat ventricle S1DZ S1 dysgranular zn VDB nu vert limb diag b
DEn dorsal endopirif IG indusium griseum M1 primary motor cx S1DZO S1 oral dysgran zn VP ventral pallidum
DI dysgran insular LAcbSh lat accumb shell M2 2ary motor cx S1FL S1 forelimb region
E ependyma/subepen lo lat olfactory tr mfb med forebr bundle S1J S1 cx, jaw region

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 22
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1
Cg1
8 2
S1FL

cg Z
D
Cg2 S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 3
S1J

IG ZO
S1D
6 ec E 4
cc
LV
LSD
S1ULp
SHi
5 CPu 5

LSI

LSV
PLd
4 6
GI
MS DCl
DI

ICjM VCl
3 AID 7
aca AcbSh
AIV
rf VDB DEn
AcbC LSS
2 LAcbSh IEn 8
3
ICj 2
mfb VP Pir1
lo ICj
1 3 9
2
Tu1

Interaural 10.32 mm Bregma 1.32 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 23

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 23
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
M1
Cg1
8 2
S1FL
DZ
S1
cg
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 S1J 3
O
S1 DZ
IG
E
6 4
cc
S1ULp
ec LV
LSD
SHi
5 mcer CPu 5

LSI
S2
Ld LSV
4 PLd 6
DCl GI

MS VCl DI
ICjM
AID
3 7
aca DEn
AcbSh AIV
rf AcbC LSS
IEn 3
2 VDB LAcbSh 2 8
Pir1

lo mfb VP
ICj
1 ICj 9
3
2
Tu1
Interaural 10.20 mm Bregma 1.20 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex Cg1 cingulate area 1 ICj islands of Calleja LSS lat stripe of str PLd paralambdoid sept Tu1 olf tub layer 1
3 layer 3 cortex Cg2 cingulate area 2 ICjM major is Calleja LSV lat septal vent rf rhinal fissure VCl ventral claustrum
aca ant comm, ant CPu caudate putamen IEn intermed endopir LV lat ventricle S1DZ S1 dysgranular zn VDB nu vert limb diag b
AcbC accumbens core DCl dorsal claustrum IG indusium griseum M1 primary motor cx S1DZO S1 oral dysgran zn VP ventral pallidum
AcbSh accumbens shell DEn dorsal endopirif LAcbSh lat accumb shell M2 2ary motor cx S1FL S1 forelimb region
AID ant insular dorsal DI dysgran insular Ld lambdoid septal zn mcer mid cerebral art S1J S1 cx, jaw region
AIV ant insular ventral E ependyma/subepen lo lat olfactory tr mfb med forebr bundle S1ULp S1 upper lip region
cc corpus callosum ec external capsule LSD lat septal dors MS medial septal nu S2 2ary somatosens
cg cingulum GI granular insular LSI lat septal intermed Pir1 piriform layer 1 SHi septohipp nu
RBSC5 Book.qxp 16/9/04 16:36 Page 24

2 layer 2 cortex cc corpus callosum E ependyma/subepen LAcbSh lat accumb shell M1 primary motor cx S1FL S1 forelimb region
3 layer 3 cortex cg cingulum ec external capsule Ld lambdoid septal zn M2 2ary motor cx S2 2ary somatosens
aca ant comm, ant Cg1 cingulate area 1 GI granular insular lo lat olfactory tr mfb med forebr bundle SHi septohipp nu
AcbC accumbens core Cg2 cingulate area 2 HDB nu horiz limb DB LSD lat septal dors MS medial septal nu SIB subst innom basal
AcbSh accumbens shell CPu caudate putamen ICj islands of Calleja LSI lat septal intermed Pir1 piriform layer 1 Tu1 olf tub layer 1
AID ant insular dorsal DCl dorsal claustrum ICjM major is Calleja LSS lat stripe of str PLd paralambdoid sept VCl ventral claustrum
AIV ant insular ventral DEn dorsal endopirif IEn intermed endopir LSV lat septal vent rf rhinal fissure VDB nu vert limb diag b
CB cell bridges DI dysgran insular IG indusium griseum LV lat ventricle S1DZ S1 dysgranular zn VP ventral pallidum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 24
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2
Cg1 M1
8 2
S1FL

cg
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3
DZ
S1

IG E
6 4
cc S1ULp
ec LV
LSD

SHi CPu
5 5
LSI
S2
Ld LSV
4 PLd 6
DCl GI

MS VCl DI
ICjM
3 AID 7
rf aca AcbSh DEn AIV

VDB AcbC LSS

SIB
2 LAcbSh 3 8
IEn
2
HDB Pir1
CB ICj
lo mfb VP

1 ICj 9
3
2
Tu1
Interaural 10.08 mm Bregma 1.08 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 25

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 25
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1
8 Cg1 2
S1FL

cg
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3
DZ
S1

IG
ec E
6 S1ULp 4
cc
LV
LSD
SHi
5 CPu 5
LSI
S2
Ld
4 PLd LSV 6
DCl GI
mcer
MS DI
VCl

3 AID 7
aca AcbSh
rf
AIV
DEn
AcbC LSS
SIB
2 LAcbSh 3 8
VDB
2
IEn Pir1
VP
mfb HDB CB
lo
ICj
1 ICj 3 9
2
Tu1
Interaural 9.96 mm Bregma 0.96 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex cg cingulum GI granular insular LSI lat septal intermed Pir1 piriform layer 1 Tu1 olf tub layer 1
3 layer 3 cortex Cg1 cingulate area 1 HDB nu horiz limb DB LSS lat stripe of str PLd paralambdoid sept VCl ventral claustrum
aca ant comm, ant Cg2 cingulate area 2 ICj islands of Calleja LSV lat septal vent rf rhinal fissure VDB nu vert limb diag b
AcbC accumbens core CPu caudate putamen IEn intermed endopir LV lat ventricle S1DZ S1 dysgranular zn VP ventral pallidum
AcbSh accumbens shell DCl dorsal claustrum IG indusium griseum M1 primary motor cx S1FL S1 forelimb region
AID ant insular dorsal DEn dorsal endopirif LAcbSh lat accumb shell M2 2ary motor cx S1ULp S1 upper lip region
AIV ant insular ventral DI dysgran insular Ld lambdoid septal zn mcer mid cerebral art S2 2ary somatosens
CB cell bridges E ependyma/subepen lo lat olfactory tr mfb med forebr bundle SHi septohipp nu
cc corpus callosum ec external capsule LSD lat septal dors MS medial septal nu SIB subst innom basal
RBSC5 Book.qxp 16/9/04 16:36 Page 26

2 layer 2 cortex cg cingulum GI granular insular LSI lat septal intermed Pir1 piriform layer 1 STMA ST med div, ant
3 layer 3 cortex Cg1 cingulate area 1 HDB nu horiz limb DB LSS lat stripe of str PLd paralambdoid sept Tu1 olf tub layer 1
aca ant comm, ant Cg2 cingulate area 2 ICj islands of Calleja LSV lat septal vent rf rhinal fissure VCl ventral claustrum
AcbC accumbens core CPu caudate putamen IEn intermed endopir LV lat ventricle S1DZ S1 dysgranular zn VDB nu vert limb diag b
AcbSh accumbens shell DCl dorsal claustrum IG indusium griseum M1 primary motor cx S1FL S1 forelimb region VP ventral pallidum
AID ant insular dorsal DEn dorsal endopirif LAcbSh lat accumb shell M2 2ary motor cx S1ULp S1 upper lip region
AIV ant insular ventral DI dysgran insular Ld lambdoid septal zn mcer mid cerebral art S2 2ary somatosens
CB cell bridges E ependyma/subepen lo lat olfactory tr mfb med forebr bundle SHi septohipp nu
cc corpus callosum ec external capsule LSD lat septal dors MS medial septal nu SIB subst innom basal

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 26
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1
Cg1
8 S1FL 2

cg

DZ
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

Cg2
7 S1 3

IG E
S1ULp
6 cc 4
ec LV
LSD
SHi

5 5
LSI CPu
Ld S2

PLd
4 LSV 6
MS DCl GI

DI
AcbSh VCl
mcer
3 STMA
AID 7
aca
AcbC AIV
VDB DEn
rf LSS
SIB IEn
2 3 8
LAcbSh
2
mcer CB
Pir1
HDB VP
mfb ICj
lo
1 3 9
ICj 2
Tu1
Interaural 9.84 mm Bregma 0.84 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 27

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 27
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1

8 Cg1 2
S1FL

cg
DZ
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3
S1

IG
ec
S1ULp
6 cc E 4
LV
LSD
SHi

5 5
LSI CPu
Ld S2

PLd
4 LSV 6
GI
MS DCl
AcbSh DI
STMA VCl
3 AID 7
aca
AcbC AIV
rf DEn
LSS
VDB SIB
2 IEn 8
3
2
VP CB
Pir1
mcer HDB
mfb
lo
1 3 9
ICj 2
Tu1
Interaural 9.72 mm Bregma 0.72 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex cc corpus callosum E ependyma/subepen lo lat olfactory tr mfb med forebr bundle S2 2ary somatosens
3 layer 3 cortex cg cingulum ec external capsule LSD lat septal dors MS medial septal nu SHi septohipp nu
aca ant comm, ant Cg1 cingulate area 1 GI granular insular LSI lat septal intermed Pir1 piriform layer 1 SIB subst innom basal
AcbC accumbens core Cg2 cingulate area 2 HDB nu horiz limb DB LSV lat septal vent PLd paralambdoid sept STMA ST med div, ant
AcbSh accumbens shell CPu caudate putamen ICj islands of Calleja LV lat ventricle rf rhinal fissure Tu1 olf tub layer 1
AID ant insular dorsal DCl dorsal claustrum IEn intermed endopir M1 primary motor cx S1DZ S1 dysgranular zn VCl ventral claustrum
AIV ant insular ventral DEn dorsal endopirif IG indusium griseum M2 2ary motor cx S1FL S1 forelimb region VDB nu vert limb diag b
CB cell bridges DI dysgran insular Ld lambdoid septal zn mcer mid cerebral art S1ULp S1 upper lip region VP ventral pallidum
RBSC5 Book.qxp 16/9/04 16:36 Page 28

2 layer 2 cortex cg cingulum GI granular insular LSD lat septal dors PLd paralambdoid sept STMA ST med div, ant
3 layer 3 cortex Cg1 cingulate area 1 HDB nu horiz limb DB LSI lat septal intermed rf rhinal fissure Tu1 olf tub layer 1
aca ant comm, ant Cg2 cingulate area 2 ICj islands of Calleja LSV lat septal vent S1DZ S1 dysgranular zn VCl ventral claustrum
AcbC accumbens core CPu caudate putamen IEn intermed endopir LV lat ventricle S1FL S1 forelimb region VDB nu vert limb diag b
AcbSh accumbens shell DCl dorsal claustrum IG indusium griseum M1 primary motor cx S1ULp S1 upper lip region VOLT vasc org lam term
AID ant insular dorsal DEn dorsal endopirif IPAC interstitial nu acp M2 2ary motor cx S2 2ary somatosens VP ventral pallidum
AIV ant insular ventral DI dysgran insular Ld lambdoid septal zn mfb med forebr bundle SHi septohipp nu
CB cell bridges E ependyma/subepen lo lat olfactory tr MS medial septal nu SHy septohypothal nu
cc corpus callosum ec external capsule LPO lat preoptic area Pir1 piriform layer 1 SIB subst innom basal

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 28
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1

Cg1
8 S1FL 2

cg DZ
Cg2 S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 3

IG E S1ULp
ec
6 cc 4
LSD LV
SHi

5 LSI 5
CPu S2
Ld

PLd LSV
4 GI 6
MS DCl
AcbSh
SHy DI
VCl AID
rf
3 aca 7
LPO AIV
AcbC

STMA IPAC DEn

VDB LSS
VP
2 VOLT 3 8
SIB IEn
2
CB
Pir1
HDB
mfb
lo
1 3 ICj
9
ICj 2
Tu1
Interaural 9.60 mm Bregma 0.60 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 29

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 29
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1

8 Cg1 S1FL 2

cg S1DZ
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3

E
IG
S1ULp
6 ec 4
cc LV
LSD
SHi

5 5
Ld LSI
CPu S2

PLd
4 LSV
6
GI
MS AcbSh
DCl
DI
AcbC
3 SHy VCl AID 7
aca
rf LPO IPAC AIV
STMA DEn
VDB LSS
2 MnPO
VP 3 8
SIB
3V IEn 2
VOLT
CB CB Pir1
HDB
mfb
1 MPA 9
mcer lo och
acer ICj 3
Tu1 2
Interaural 9.48 mm Bregma 0.48 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex AIV ant insular ventral DCl dorsal claustrum ICj islands of Calleja LSD lat septal dors mfb med forebr bundle rf rhinal fissure SIB subst innom basal
3 layer 3 cortex CB cell bridges DEn dorsal endopirif IEn intermed endopir LSI lat septal intermed MnPO median preoptic nu S1DZ S1 dysgranular zn STMA ST med div, ant
3V 3rd ventricle cc corpus callosum DI dysgran insular IG indusium griseum LSV lat septal vent MPA med preoptic area S1FL S1 forelimb region Tu1 olf tub layer 1
aca ant comm, ant cg cingulum E ependyma/subepen IPAC interstitial nu acp LV lat ventricle MS medial septal nu S1ULp S1 upper lip region VCl ventral claustrum
AcbC accumbens core Cg1 cingulate area 1 ec external capsule Ld lambdoid septal zn M1 primary motor cx och optic chiasm S2 2ary somatosens VDB nu vert limb diag b
AcbSh accumbens shell Cg2 cingulate area 2 GI granular insular lo lat olfactory tr M2 2ary motor cx Pir1 piriform layer 1 SHi septohipp nu VOLT vasc org lam term
AID ant insular dorsal CPu caudate putamen HDB nu horiz limb DB LPO lat preoptic area mcer mid cerebral art PLd paralambdoid sept SHy septohypothal nu VP ventral pallidum
RBSC5 Book.qxp 16/9/04 16:36 Page 30

2 layer 2 cortex cc corpus callosum DI dysgran insular IG indusium griseum LSS lat stripe of str MPA med preoptic area S1HL S1 hindlimb region STLV ST lat div, vent
3 layer 3 cortex cg cingulum E ependyma/subepen IPAC interstitial nu acp LSV lat septal vent MS medial septal nu S1ULp S1 upper lip region STMA ST med div, ant
3V 3rd ventricle Cg1 cingulate area 1 ec external capsule Ld lambdoid septal zn LV lat ventricle och optic chiasm S2 2ary somatosens STMV ST med div, vent
4 layer 4 cortex Cg2 cingulate area 2 GI granular insular lo lat olfactory tr M1 primary motor cx Pir1 piriform layer 1 SHi septohipp nu Tu1 olf tub layer 1
aca ant comm, ant CPu caudate putamen HDB nu horiz limb DB LPO lat preoptic area M2 2ary motor cx rf rhinal fissure SHy septohypothal nu VCl ventral claustrum
AID ant insular dorsal DCl dorsal claustrum ICj islands of Calleja LSD lat septal dors mfb med forebr bundle S1DZ S1 dysgranular zn SIB subst innom basal VOLT vasc org lam term
AIV ant insular ventral DEn dorsal endopirif IEn intermed endopir LSI lat septal intermed MnPO median preoptic nu S1FL S1 forelimb region STLP ST lat div, post VP ventral pallidum
ZL zona limitans

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 30
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1
S1HL
8 Cg1 2
S1FL

cg Z
D
S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3

E
IG
S1ULp
6 cc LV 4
ec LSD
SHi

Ld LSI
5 1 5
2
3
4
CPu S2

4 LSV 6
GI
ZL DCl
MS
STLP DI
SHy STMA
VCl
3 aca AID 7

rf AIV
STMV DEn
MnPO LPO STLV IPAC
2 VOLT SIB 8
LSS
IEn 3
3V VP CB
HDB 2
mfb MPA Pir1
1 lo och 9
acer
ICj
3
2
Interaural 9.36 mm Tu1
Bregma 0.36 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 31

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 31
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1
S1HL
8 Cg1 S1FL 2

Z
D
cg

S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3

IG
E S1ULp
6 cc LV 4
ec LSD
SHi
LSI
1
5 2 5
3
4
S2
CPu

4 LSV LV 6
MS STMA GI
ZL DCl
STLD
STLP DI
SHy
VCl
3 aca AID 7

rf STMV AIV
STLV DEn
MnPO
LPO
2 VOLT IPAC LSS 8
SIB IEn
VP 3
CB 2
3V
MPA HDB Pir1
mfb
1 lo VMPO ICj 9
och
3
2
Interaural 9.24 mm Tu1
Bregma 0.24 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex cg cingulum ec external capsule LPO lat preoptic area mfb med forebr bundle S1DZ S1 dysgranular zn STLD STL dorsal VOLT vasc org lam term
3 layer 3 cortex Cg1 cingulate area 1 GI granular insular LSD lat septal dors MnPO median preoptic nu S1FL S1 forelimb region STLP ST lat div, post VP ventral pallidum
4 layer 4 cortex Cg2 cingulate area 2 HDB nu horiz limb DB LSI lat septal intermed MPA med preoptic area S1HL S1 hindlimb region STLV ST lat div, vent
aca ant comm, ant CPu caudate putamen ICj islands of Calleja LSS lat stripe of str MS medial septal nu S1ULp S1 upper lip region STMA ST med div, ant
AID ant insular dorsal DCl dorsal claustrum IEn intermed endopir LSV lat septal vent och optic chiasm S2 2ary somatosens STMV ST med div, vent
AIV ant insular ventral DEn dorsal endopirif IG indusium griseum LV lat ventricle P3V preopt recess 3V SHi septohipp nu Tu1 olf tub layer 1
CB cell bridges DI dysgran insular IPAC interstitial nu acp M1 primary motor cx Pir1 piriform layer 1 SHy septohypothal nu VCl ventral claustrum
cc corpus callosum E ependyma/subepen lo lat olfactory tr M2 2ary motor cx rf rhinal fissure SIB subst innom basal VMPO ventromed preopt
RBSC5 Book.qxp 16/9/04 16:36 Page 32

2 layer 2 cortex cc corpus callosum ec external capsule lo lat olfactory tr mcer mid cerebral art PS parastrial nu SHy septohypothal nu VMPO ventromed preopt
3 layer 3 cortex cg cingulum fi fimbria of hipp LPO lat preoptic area MCPO magnocell preopt rf rhinal fissure SIB subst innom basal VOLT vasc org lam term
3V 3rd ventricle Cg1 cingulate area 1 Fu ST fusiform part LSD lat septal dors mfba mfb ‘a’ component S1DZ S1 dysgranular zn STLD STL dorsal VP ventral pallidum
4 layer 4 cortex Cg2 cingulate area 2 GI granular insular LSI lat septal intermed mfbb mfb ‘b’ componet S1FL S1 forelimb region STLP ST lat div, post
aca ant comm, ant CPu caudate putamen HDB nu horiz limb DB LSS lat stripe of str MnPO median preoptic nu S1HL S1 hindlimb region STLV ST lat div, vent
AID ant insular dorsal DCl dorsal claustrum ICj islands of Calleja LSV lat septal vent MPA med preoptic area S1ULp S1 upper lip region STMA ST med div, ant
AIV ant insular ventral DEn dorsal endopirif IEn intermed endopir LV lat ventricle MS medial septal nu S2 2ary somatosens STMV ST med div, vent
AVPe anterovent perive DI dysgran insular IG indusium griseum M1 primary motor cx och optic chiasm SFi septofimbrial nu Tu1 olf tub layer 1
CB cell bridges E ependyma/subepen IPAC interstitial nu acp M2 2ary motor cx Pir1 piriform layer 1 SHi septohipp nu VCl ventral claustrum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 32
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1
S1HL
8 Cg1
2
S1FL

cg

Z
D
Cg2

S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 3
IG
E
S1ULp
cc
6 LV 4
LSD
ec
SHi

LSI
2 1
5 3 5
fi SFi 4
CPu
S2
MS
LSV LV
4 6
ZL
STLD
GI
SHy STMA
STLP DCl DI
3 aca 7
VCl AID
STMV
rf MnPO PS Fu STLV
VOLT AIV
LPO DEn
2 LSS 8
IPAC
AVPe MPA SIB VP IEn
3V 3
VMPO HDB CB
mfbb 2
1 mfba MCPO 9
och Pir1
ICj
lo 3
2
Interaural 9.12 mm mcer Tu1 Bregma 0.12 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 33

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 33
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1 S1HL
8 Cg1 S1FL 2

Z
cg

D
S1
Cg2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 3
IG E
S1ULp
cc
LV
6 SHi LSD 4
ec

LSI
5 1 5
3 2
4
S2
fi SFi CPu

LSV STLP
4 6
MS
STLD
STMA GI
SHy
DCl
ic
3 aca DI 7
MnPO STMV
StA VCl
rf VOLT PS Fu STLV
Al AIP

LPO LSS
2 IPAC DEn 8
3V
MPA VP
SIB IEn
AVPe
VMPO 3
mfbb HDB CB
mfba VLPO MCPO 2
1 och
9
SO Pir1
3
2
Interaural 9.00 mm lo Tu1
Bregma 0.00 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex cc corpus callosum E ependyma/subepen IG indusium griseum LV lat ventricle MS medial septal nu S2 2ary somatosens STLV ST lat div, vent
3 layer 3 cortex cg cingulum ec external capsule IPAC interstitial nu acp M1 primary motor cx och optic chiasm SFi septofimbrial nu STMA ST med div, ant
3V 3rd ventricle Cg1 cingulate area 1 fi fimbria of hipp lo lat olfactory tr M2 2ary motor cx Pir1 piriform layer 1 SHy septohypothal nu STMV ST med div, vent
4 layer 4 cortex Cg2 cingulate area 2 Fu ST fusiform part LPO lat preoptic area MCPO magnocell preopt PS parastrial nu SIB subst innom basal Tu1 olf tub layer 1
aca ant comm, ant CPu caudate putamen GI granular insular LSD lat septal dors mfba mfb ‘a’ component rf rhinal fissure SO supraoptic nu VCl ventral claustrum
AIP agran insular post DCl dorsal claustrum HDB nu horiz limb DB LSI lat septal intermed mfbb mfb ‘b’ componet S1DZ S1 dysgranular zn StA strial preopt area VLPO ventrolat preoptic
Al alar nu DEn dorsal endopirif ic internal capsule LSS lat stripe of str MnPO median preoptic nu S1FL S1 forelimb region STLD STL dorsal VMPO ventromed preopt
AVPe anterovent perive DI dysgran insular IEn intermed endopir LSV lat septal vent MPA med preoptic area S1HL S1 hindlimb region STLP ST lat div, post VOLT vasc org lam term
VP ventral pallidum
RBSC5 Book.qxp 16/9/04 16:36 Page 34

2 layer 2 cortex cc corpus callosum DI dysgran insular IG indusium griseum M1 primary motor cx Pir1 piriform layer 1 SHi septohipp nu STMA ST med div, ant
3 layer 3 cortex cg cingulum E ependyma/subepen IPAC interstitial nu acp M2 2ary motor cx PS parastrial nu SHy septohypothal nu STMAL ST med anterolat
3V 3rd ventricle Cg1 cingulate area 1 ec external capsule lo lat olfactory tr MCPO magnocell preopt rf rhinal fissure SIB subst innom basal STMAM ST med ant med
ac ant commissure Cg2 cingulate area 2 f fornix LPO lat preoptic area mfba mfb ‘a’ component S1DZ S1 dysgranular zn SO supraoptic nu STMV ST med div, vent
acp ant comm post chp choroid plexus Fu ST fusiform part LSD lat septal dors mfbb mfb ‘b’ componet S1FL S1 forelimb region StA strial preopt area Tu1 olf tub layer 1
AIP agran insular post CPu caudate putamen GI granular insular LSI lat septal intermed MnPO median preoptic nu S1HL S1 hindlimb region STLD STL dorsal VCl ventral claustrum
Al alar nu CxA1 cx-amyg trans 1 HDB nu horiz limb DB LSS lat stripe of str MPA med preoptic area S1ULp S1 upper lip region STLJ STL juxtacaps VLPO ventrolat preoptic
AVPe anterovent perive DCl dorsal claustrum ic internal capsule LSV lat septal vent MPOL medial preopt lat S2 2ary somatosens STLP ST lat div, post VMPO ventromed preopt
CB cell bridges DEn dorsal endopirif IEn intermed endopir LV lat ventricle och optic chiasm SFi septofimbrial nu STLV ST lat div, vent VP ventral pallidum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 34
5 5
10 0

0 Interaural 10

+15 +10 +5 0 -5
9 1

M2 M1 S1HL
8 2
Cg1 S1FL

cg Z
D
S1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

7 Cg2 3

IG E
S1ULp
cc
6 LSD 4
chp SHi LV
ec

LSI
fi
2 1
5 3 5

SFi S2
CPu
LSV
4 f 6
STMA STLD
SHy
ic GI
MnPO
STLP DCl
ac STMAM STLJ
3 MnPO STMAL DI 7
STMV
SHy STLV VCl
rf acp StA AIP
PS Fu
Al
MPOL
2 LSS DEn 8
LPO IPAC
MPA VP IEn
SIB
AVPe 3 3
CB 2
1 mfba mfbb VLPO HDB MCPO 2 9
3V VMPO Pir1
och SO CxA1
3
Interaural 8.88 mm lo 2
Bregma -0.12 mm
Tu1
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 35

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 35
5 5
9 1

0 Interaural 10
M2 M1
S1HL
+15 +10 +5 0 -5
8 Cg1 S1FL 2

DZ
cg

S1
7 Cg2 3

IG E
S1ULp
cc
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 ec chp LV 4
LSD
SHi

fi
5 LSI 5
TS
CPu S2

LSV STD
4 6
ic STLD
MnPO SFi SHy
f GI
STLJ
LTer CST DCl
STMAM DI
STMAL ac EGP
3 STMV 7
MnPO STI STLP VCl
rf AIP
acp SHy PS Fu STLV
StA
3V MPOM LSS DEn
2 LPO 8
VP IPACL
Pe MPA IPACM
SIB 3
IEn 2
MPOL
Pir1
mfba mfbb VMPO HDB MCPO AA 3
1 VLPO
9
2
och
SO Tu CxA1
lo acer
mcer
0 10

Interaural 8.76 mm Bregma -0.24 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex AIP agran insular post CPu caudate putamen ec external capsule ic internal capsule LSD lat septal dors
3 layer 3 cortex B basal nu CST nu comm st term EGP ext globus pallidus IEn intermed endopir LSI lat septal intermed
3V 3rd ventricle cc corpus callosum CxA1 cx-amyg trans 1 f fornix IG indusium griseum LSS lat stripe of str
AA ant amyg area cg cingulum DCl dorsal claustrum fi fimbria of hipp IPACL IPAC, lateral LSV lat septal vent
ac ant commissure Cg1 cingulate area 1 DEn dorsal endopirif Fu ST fusiform part IPACM IPAC, medial LTer lemina terminalis
acer ant cerebral art Cg2 cingulate area 2 DI dysgran insular GI granular insular lo lat olfactory tr LV lat ventricle
acp ant comm post chp choroid plexus E ependyma/subepen HDB nu horiz limb DB LPO lat preoptic area
RBSC5 Book.qxp 16/9/04 16:36 Page 36

M1 primary motor cx
M2 2ary motor cx
mcer mid cerebral art
MCPO magnocell preopt
mfba mfb ‘a’ component
mfbb mfb ‘b’ componet
MnPO median preoptic nu
MPA med preoptic area rf rhinal fissure SFi septofimbrial nu STLD STL dorsal STMV ST med div, vent
MPOL medial preopt lat S1BF S1 cx, barrel field SHi septohipp nu STLI STL intermediate TS triangular septal
MPOM medial preopt med S1DZ S1 dysgranular zn SHy septohypothal nu STLJ STL juxtacaps Tu olfactory tubercle
och optic chiasm S1FL S1 forelimb region SIB subst innom basal STLP ST lat div, post VCl ventral claustrum
Pe periventric hy nu S1HL S1 hindlimb region SO supraoptic nu STLV ST lat div, vent VLPO ventrolat preoptic
Pir1 piriform layer 1 S1ULp S1 upper lip region StA strial preopt area STMAL ST med anterolat VMPO ventromed preopt
PS parastrial nu S2 2ary somatosens STI ST intermediat STMAM ST med ant med VP ventral pallidum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 36
5 5
9 1

0 Interaural 10
M2 M1
+15 +10 +5 0 -5 S1HL
8 2
Cg1 S1FL

DZ
cg

S1
7 Cg2 S1BF 3
E
IG

cc S1ULp
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 LSD 4
ec chp LV

fi
5 LSI 5
SFi CPu
TS S2

ic LSV STD
4 6
LTer SHy STLI
f GI
EGP
3V STMAM STLJ DCl
STMAL DI
ac
3 STMV 7
rf STLP
STLI VCl
3V STLV AIP
SHy PS
acp
Fu B IPACL
Pe LSS
VP DEn
2 MPOL 8
MPA LPO SIB IPACM
IEn 3
MPOM B 2

HDB MCPO Pir1

mfba mfbb AA 3
1 VMPO VLPO 9
2
och SO
CxA1
lo acer
mcer
0 10

Interaural 8.64 mm Bregma -0.36 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 37

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 37
5 5
9 1

0 Interaural 10 M2 M1
S1HL
+15 +10 +5 0 -5
Cg1
8 S1FL
2

cg

Z
D
S1
Cg2
7 3
S1BF
IG E

cc
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 LSD 4
ec S1ULp
chp LV
fi LSI

5 TS 5
SFi
LTer
CPu S2
ic
st
4 SHy 6
AM
f 3V
S TM AL STLP
GI
M
BAC ST
DCl
ac EGP DI
3 STLI STLV 7
3V
PS VCl
rf SHy AIP
StHy B IPACL
Pe MPOC
acp DEn
2 MPA VP 8
LPO
SIB IPACM
MPOM IEn
LSS
B
VLPO 3
mfbb HDB MCPO 2
1 mfba 9
MPOL 3
SCh Pir1
ACo 2
och AA
acer SO CxA1
lo mcer LOT
0 10

Interaural 8.52 mm Bregma -0.48 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex acp ant comm post chp choroid plexus EGP ext globus pallidus IPACL IPAC, lateral LSS lat stripe of str
3 layer 3 cortex AIP agran insular post CPu caudate putamen f fornix IPACM IPAC, medial LTer lemina terminalis
3V 3rd ventricle B basal nu CxA1 cx-amyg trans 1 fi fimbria of hipp lo lat olfactory tr LV lat ventricle
a artery BAC bed nu ant comm DCl dorsal claustrum GI granular insular LOT nu of lat olf tr
AA ant amyg area cc corpus callosum DEn dorsal endopirif HDB nu horiz limb DB LOT1 LOT layer 1
ac ant commissure cg cingulum DI dysgran insular ic internal capsule LPO lat preoptic area
acer ant cerebral art Cg1 cingulate area 1 E ependyma/subepen IEn intermed endopir LSD lat septal dors
ACo ant cortical amyg Cg2 cingulate area 2 ec external capsule IG indusium griseum LSI lat septal intermed
RBSC5 Book.qxp 16/9/04 16:36 Page 38

M1 primary motor cx
M2 2ary motor cx
mcer mid cerebral art
mch med corthypo tr
MCPO magnocell preopt
mfba mfb ‘a’ component
mfbb mfb ‘b’ componet
MPA med preoptic area
MPOC medial preopt cent
MPOL medial preopt lat
MPOM medial preopt med
och optic chiasm
PDPO postdors preopt
Pe periventric hy nu
PeAP periv ant parvic
Pir1 piriform layer 1
PS parastrial nu SCh suprachiasmatic nu st stria terminalis STMPL STM posterolat
rf rhinal fissure SFi septofimbrial nu StHy striohypothal nu STMPM STM posteromed
S1BF S1 cx, barrel field SFO subfornical organ STLI STL intermediate TS triangular septal
S1DZ S1 dysgranular zn SHy septohypothal nu STLP ST lat div, post VCl ventral claustrum
S1FL S1 forelimb region SIB subst innom basal STLV ST lat div, vent vhc ventral hipp comm
S1HL S1 hindlimb region sm stria medullaris STMAL ST med anterolat VLPO ventrolat preoptic
S1ULp S1 upper lip region SO supraoptic nu STMAM ST med ant med VP ventral pallidum
S2 2ary somatosens sod supraoptic decussn STMPI STM postintermed

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 38
5 5
9 1

0 Interaural 10 M2 M1
S1HL
+15 +10 +5 0 -5 Cg1
8 S1FL 2

cg

Z
D
S1
Cg2
7 3
IG S1BF
E

cc
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 ec LSD 4
chp LV S1ULp
fi

TS
5 SFi 5
vhc
st SFO CPu S2
ic
4 STMPM 6
sm
f PI GI
3V M
BAC ST PL EGP
M
ST STLP DCl DI
mch
3 PeAP STLI 7
StHy
rf VCl AIP
PDPO

Pe MPOM VP IPACL
acp LSS
2 IPACM DEn
8
MPA LPO SIB
MPOC IPACL
B
IEn
3
mfba mfbb HDB MCPO
MPOL 2
1 VLPO AA 9
SCh 3
SO ACo Pir1
och 2 2
lo mcer sod CxA1
LOT1
a
0 10

Interaural 8.40 mm Bregma -0.60 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 39

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 39
5 5
9 1
M1
0 Interaural 10 M2 S1HL
+15 +10 +5 0 -5
8 Cg1 S1FL 2

DZ
cg

S1
Cg2
7 3
IG S1BF
E
cc
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 LSD 4
ec LV
chp

SFi S1ULp
TS
fi
5 5
vhc
SFO
st CPu
IVF S2
ic

4 STMPM 6
sm
cst 3V BAC EGP GI
f

PI

L
STMP
STM
APF DCl DI
3 7
rf StHy
PeAP STLP VCl
AIP
PDPO
EAC IPACL
acp DEn
2 Pe MPOM MPA 8
LPO B VP
SIB I
IPACM IEn
MPOC
3
mfba MPOL VLH MCPO AA 2
mfbb HDB
1 ESO
9
SChDL ACo 3 Pir1
SO 2
SChVM 2
och
a LOT1 CxA1
lo sod
0 10

Interaural 8.28 mm Bregma -0.72 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex APF ant perifornical nu CPu caudate putamen EAC sublentic EA cent I intercalated nu lo lat olfactory tr
3 layer 3 cortex B basal nu cst commiss st term ec external capsule ic internal capsule LOT1 LOT layer 1
3V 3rd ventricle BAC bed nu ant comm CxA1 cx-amyg trans 1 EGP ext globus pallidus IEn intermed endopir LPO lat preoptic area
a artery cc corpus callosum DCl dorsal claustrum ESO episupraoptic nu IG indusium griseum LSD lat septal dors
AA ant amyg area cg cingulum DEn dorsal endopirif f fornix IPACL IPAC, lateral LV lat ventricle
ACo ant cortical amyg Cg1 cingulate area 1 df dorsal fornix fi fimbria of hipp IPACM IPAC, medial
acp ant comm post Cg2 cingulate area 2 DI dysgran insular GI granular insular IVF interventric for
AIP agran insular post chp choroid plexus E ependyma/subepen HDB nu horiz limb DB LA lateroant hy nu
RBSC5 Book.qxp 16/9/04 16:36 Page 40

M1 primary motor cx
M2 2ary motor cx
mch med corthypo tr
MCPO magnocell preopt
mfb med forebr bundle
mfba mfb ‘a’ component
mfbb mfb ‘b’ componet
MPA med preoptic area
MPOC medial preopt cent PVA paraventric th ant S1ULp S1 upper lip region st stria terminalis VEn ventral endopir
MPOL medial preopt lat rf rhinal fissure S2 2ary somatosens StHy striohypothal nu vhc ventral hipp comm
MPOM medial preopt med SChDL suprach dorsolat SFi septofimbrial nu STLP ST lat div, post VLH ventrolat hy nu
och optic chiasm SChVM suprach ventromed SFO subfornical organ STMPI STM postintermed VP ventral pallidum
PaAP Pa ant parvicell S1BF S1 cx, barrel field SIB subst innom basal STMPL STM posterolat
Pe periventric hy nu S1DZ S1 dysgranular zn sm stria medullaris STMPM STM posteromed
PeAP periv ant parvic S1FL S1 forelimb region SO supraoptic nu TS triangular septal
Pir1 piriform layer 1 S1HL S1 hindlimb region sod supraoptic decussn VCl ventral claustrum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 40
5 5
9 1

0 Interaural 10 M2 M1
S1HL
+15 +10 +5 0 -5 Cg1
8 S1FL
2

cg

DZ
Cg2

S1
7 3
IG S1BF
cc
chp df LSD LV
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
TS
SFi
ec S1ULp
vhc
fi
5 5
SFO
st IVF
S2
CPu
ic sm PVA PM
4 M 6
ST
BAC
EGP GI
mch
3V

I
STMP

L
f DCl

STMP
DI
3 PaAP 7
B
APF
rf StHy STLP VCl AIP
EAC
MPOL B IPACL
2 Pe VP 8
acp MPA DEn
SIB
LPO I
MPOM MCPO
mfb IPACM VEn 3
LA VLH HDB AA 2
1 SO ESO 3 9
SChVM SChDL ACo Pir1
3
och 2
sod 2
lo LOT1
CxA1
0 10

Interaural 8.16 mm Bregma -0.84 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 41

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 41
5 5
9 1

10 M2 M1
0 Interaural S1HL
+15 +10 +5 0 -5 Cg1
8 S1FL 2

S1DZ
cg
Cg2
7 3
IG
S1BF
cc
chp LV
df LSD
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
ec TS

vhc SFi
S1ULp
fi
5 SFO 5
st IVF

cst CPu S2
ic

M
3V PVA
4 sm 6

STMP
PT
EGP GI

I
STMP
STMPL
f mch
3 APF B DCl DI 7

PaAP VCl
rf StHy AIP
Pe EAC B
IPACL
2 DEn
8
MPOM MPA
LPO VP
acp IPACM
AHA
MCPO VEn
SChDL SPa VLH AA
mfb LA
1 HDB 3 9
ESO 3 ACo
SChVM 3 2
SO
opt 2
sod 2 Pir1
a LOT1
lo CxA1
0 10

Interaural 8.04 mm Bregma -0.96 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex AIP agran insular post Cg2 cingulate area 2 df dorsal fornix GI granular insular LA lateroant hy nu
3 layer 3 cortex AM anteromed th nu chp choroid plexus DI dysgran insular HDB nu horiz limb DB lo lat olfactory tr
3V 3rd ventricle APF ant perifornical nu CPu caudate putamen EAC sublentic EA cent I intercalated nu LOT1 LOT layer 1
a artery AV anterovent th nu cst commiss st term ec external capsule ic internal capsule LPO lat preoptic area
AA ant amyg area B basal nu CxA1 cx-amyg trans 1 EGP ext globus pallidus IG indusium griseum LSD lat septal dors
ACo ant cortical amyg cc corpus callosum D3V dorsal 3rd vent ESO episupraoptic nu IPACL IPAC, lateral LV lat ventricle
acp ant comm post cg cingulum DCl dorsal claustrum f fornix IPACM IPAC, medial
AHA ant hypothal ant Cg1 cingulate area 1 DEn dorsal endopirif fi fimbria of hipp IVF interventric for
RBSC5 Book.qxp 16/9/04 16:36 Page 42

M1 primary motor cx
M2 2ary motor cx
mch med corthypo tr
MCPO magnocell preopt
MeAD med anterodorsal
mfb med forebr bundle
MPA med preoptic area
MPOM medial preopt med
opt optic tract
PaAP Pa ant parvicell
Pe periventric hy nu
Pir1 piriform layer 1
PT paratenial th nu
PVA paraventric th ant
RCh retrochiasm area
Re reuniens th nu
rf rhinal fissure
Rt reticular th nu
RtSt reticluostrial nu
S1BF S1 cx, barrel field
S1DZ S1 dysgranular zn
S1FL S1 forelimb region
S1HL S1 hindlimb region
S1ULp S1 upper lip region
S2 2ary somatosens st stria terminalis VEn ventral endopir
SFi septofimbrial nu StHy striohypothal nu vhc ventral hipp comm
SFO subfornical organ STMP ST med div, post VLH ventrolat hy nu
SM nu stria medull STMPI STM postintermed VP ventral pallidum
sm stria medullaris STMPL STM posterolat
SO supraoptic nu STMPM STM posteromed
sod supraoptic decussn TS triangular septal
SPa subparaventric zn VCl ventral claustrum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 42
5 5
9 1

0 Interaural 10 M1
M2 S1HL
+15 +10 +5 0 -5 Cg1
8 S1FL 2
S1DZ
cg
Cg2
7 3
IG S1BF

cc
chp LV
df
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 TS 4
SFi
vhc IVF

fi S1ULp
ec
5 st SFO 5
RtSt
D3V AV
cst
STMP CPu
S2
4 ic PT 6
sm
PVA Rt
AM EGP
STMPI B
GI
3V Re
3 f
DCl 7
PaAP DI
SM
STMPM B VCl
rf StHy STMPL EAC IPACM AIP
acp IPACL
2 8
MPOM MPA LPO VP I DEn
Pe
AHA MCPO I I
AA
SPa VLH HDB VEn 3
mfb LA
1 ESO 3 2 9
RCh SO ACo 3
MeAD 2 Pir1
opt
2
sod
lo LOT1 CxA1
0 10

Interaural 7.92 mm Bregma -1.08 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 43

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 43
5 5
9 1

0 Interaural 10 M2 M1
S1HL
+15 +10 +5 0 -5 Cg1
8 2
S1FL

DZ
S1
cg
Cg2
7 3
IG
S1BF
cc
chp LV
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 df 4
TS

vhc
fi SFO
ec
5 AVDM 5
st S1ULp
RtSt
cst D3V AD
sm AVVL

4 ic 6
PT CPu S2
PVA Rt B
AM

EGP
B
3 sm Re VRe GI 7
DCl
f ANS DI
3V
PaAP STMPL SM VCl
rf B IPACM
EAC AIP
2 MPA
STMPM
EAM
8
DEn
acp MPO AHA PLH IPACL
Pe MCPO I
AA
1 LA VLH HDB VEn 3 9
mfb SPa 3 BMA
ESO
2
RCh SO MeAD
opt 2 ACo
3 Pir1
a sod LOT1
0 2 10
lo a CxA1

Interaural 7.80 mm Bregma -1.20 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex AIP agran insular post Cg1 cingulate area 1 DEn dorsal endopirif fi fimbria of hipp IPACM IPAC, medial
3 layer 3 cortex AM anteromed th nu Cg2 cingulate area 2 df dorsal fornix GI granular insular LA lateroant hy nu
3V 3rd ventricle ANS acc neurosecret chp choroid plexus DI dysgran insular HDB nu horiz limb DB lo lat olfactory tr
a artery AVDM AV th dorsomed CM centr med th nu EAC sublentic EA cent I intercalated nu LOT1 LOT layer 1
AA ant amyg area AVVL AV th ventrolat CPu caudate putamen EAM sublentic EA med IAD interanterodors nu LV lat ventricle
ACo ant cortical amyg B basal nu cst commiss st term ec external capsule ic internal capsule
acp ant comm post BMA basomed amyg ant CxA1 cx-amyg trans 1 EGP ext globus pallidus IG indusium griseum
AD anterodors th nu cc corpus callosum D3V dorsal 3rd vent ESO episupraoptic nu IPAC interstitial nu acp
AHA ant hypothal ant cg cingulum DCl dorsal claustrum f fornix IPACL IPAC, lateral
RBSC5 Book.qxp 16/9/04 16:36 Page 44

M1 primary motor cx
M2 2ary motor cx
MCPO magnocell preopt
MeAD med anterodorsal
mfb med forebr bundle
MPA med preoptic area
MPO medial preoptic nu
opt optic tract
PaAP Pa ant parvicell
PC paracentral th nu
Pe periventric hy nu
Pir1 piriform layer 1
PLH peduncular lat hy
PT paratenial th nu
PVA paraventric th ant
RCh retrochiasm area
Re reuniens th nu
rf rhinal fissure
Rt reticular th nu
RtSt reticluostrial nu
S1BF S1 cx, barrel field
S1DZ S1 dysgranular zn sm stria medullaris TS triangular septal
S1FL S1 forelimb region SO supraoptic nu VCl ventral claustrum
S1HL S1 hindlimb region sod supraoptic decussn VEn ventral endopir
S1ULp S1 upper lip region SPa subparaventric zn vhc ventral hipp comm
S2 2ary somatosens st stria terminalis VLH ventrolat hy nu
SFO subfornical organ STMPL STM posterolat VRe vent reuniens nu
SM nu stria medull STMPM STM posteromed

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 44
5 5
9 1

0 Interaural 10 M1
M2 S1HL
+15 +10 +5 0 -5
Cg1 S1FL
8 2

DZ
S1
cg
Cg2
7 3

IG S1BF

chp cc LV
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 df 4
TS

vhc
ec fi SFO
AD
5 RtSt S1ULp 5
st chp
cst D3V AVDM
sm
PC
AVVL
PVA PT
4 ic CPu S2 6
IAD Rt
AM
CM EGP
GI
PVA
3 Re VRe 7
DCl DI
STMPL B B
rf f ANS
3V B VCl AIP
sm EAC
2 PaAP STMPM EAM 8
MPA I DEn
AHA PLH IPAC
MCPO I
Pe HDB 3
VLH AA VEn
1 mfb 2 9
LA ESO
MPO 3 BMA
SPa SO Pir1
opt 2 3
ACo
RCh
MeAD LOT1 2
0 lo a sod 10
CxA1

Interaural 7.68 mm Bregma -1.32 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 45

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 45
5 5
9 1

0 Interaural 10 M2 M1
S1HL
+15 +10 +5 0 -5 Cg1
8 S1FL 2

DZ
S1
cg
Cg2
7 3
IG S1BF

chp df cc LV
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 TS 4

vhc
SFO
fi
ec AVDM
chp RtSt STSL S1ULp
5 st AD 5
cst D3V
sm
STSM
AVVL
PVA PT
PC S2
4 IAD CPu 6
ic B
CM AM Rt
EGP
Re AMV GI
3 7
Xi
VRe DI
B DCl
sm PaMP JPLH ANS
rf VCl AIP
f SM
2 8
PaMM EAM EAC
STMPM IPAC
SPa CeM DEn
PaV I
3V AHA PLH
Cir AA I 3
mfb MCPO VEn
1 Pe VLH HDB 9
LA 2
ESO 3 BMA
SO MeAD Pir1
opt 2 3
ACo
RCh RChL LOT1
0 sod 2 10
a CxA1

Interaural 7.56 mm Bregma -1.44 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex af amygdal fissure aot access olfact tr BMA basomed amyg ant chp choroid plexus DCl dorsal claustrum EGP ext globus pallidus IAD interanterodors nu
3 layer 3 cortex AHA ant hypothal ant ASt amygdalostr trans cc corpus callosum Cir circular nu DEn dorsal endopirif ESO episupraoptic nu ic internal capsule
3V 3rd ventricle AHC ant hypothal cent AVDM AV th dorsomed CeC central amyg caps CM centr med th nu df dorsal fornix f fornix IG indusium griseum
a artery AIP agran insular post AVVL AV th ventrolat CeM central amyg med CPu caudate putamen DI dysgran insular fi fimbria of hipp IPAC interstitial nu acp
AA ant amyg area AM anteromed th nu B basal nu cg cingulum cst commiss st term EAC sublentic EA cent GI granular insular JPLH juxtaparav lat hy
ACo ant cortical amyg AMV anteromed th vent BAOT bed nu acc olf tr Cg1 cingulate area 1 CxA1 cx-amyg trans 1 EAM sublentic EA med HDB nu horiz limb DB LA lateroant hy nu
AD anterodors th nu ANS acc neurosecret BLA basolat amyg ant Cg2 cingulate area 2 D3V dorsal 3rd vent ec external capsule I intercalated nu LOT1 LOT layer 1
LV lat ventricle
RBSC5 Book.qxp 16/9/04 16:36 Page 46

M1 primary motor cx
M2 2ary motor cx
MCPO magnocell preopt
MeAD med anterodorsal
mfb med forebr bundle
MoDG molecular dent gy
opt optic tract
PaMM Pa med magno
PaMP Pa med parvicell
PaV Pa, ventral part
PaXi paraxiphoid nu
PC paracentral th nu
Pe periventric hy nu
Pir1 piriform layer 1
PLH peduncular lat hy
PT paratenial th nu
PV paraventric th nu
PVA paraventric th ant
RCh retrochiasm area
RChL retrochiasm lat
Re reuniens th nu
rf rhinal fissure
Rt reticular th nu
RtSt reticluostrial nu
S1BF S1 cx, barrel field
S1DZ S1 dysgranular zn
S1FL S1 forelimb region
S1HL S1 hindlimb region SPa subparaventric zn VEn ventral endopir
S1Sh S1 shoulder region st stria terminalis vhc ventral hipp comm
S1ULp S1 upper lip region STMPM STM posteromed VLH ventrolat hy nu
S2 2ary somatosens STSL ST supracaps lat VRe vent reuniens nu
SFO subfornical organ STSM ST supracaps med Xi xiphoid th nu
SM nu stria medull TS triangular septal
sm stria medullaris TuLH tuberal lat hy
SO supraoptic nu VA ventral ant th nu
sod supraoptic decussn VCl ventral claustrum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 46
5 5
9 1

0 Interaural 10 M1
M2 S1HL
+15 +10 +5 0 -5 Cg1
8 S1Sh 2

DZ
S1
cg
Cg2
7 3
S1BF
IG

chp cc LV
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 df 4
vhc
SFO MoDG
fi AVDM
STSM STSL
st chp
5 ec AD S1ULp 5
cst sm D3V
RtSt

PVA PT AVVL
PC
VA CPu S2
4 6
CM
IAD Rt
AM

EGP
ic Re AMV GI
3 7
Xi VRe B DCl DI
PaXi
rf ANS VCl
PaMP AIP
2 3V JPLH 8
f PaMM
ASt
EAM EAC CeM DEn
PaV SPa CeC
PLH MCPO BLA
mfb AHC 3
HDB AA I
1 VLH VEn 2 9
Pe
cst
MeAD BMA Pir1
SO
opt TuLH 3
BAOT ACo
aot sod 2
0 RCh RChL 10
CxA1
af
Interaural 7.44 mm Bregma -1.56 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 47

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 47
5 5
9 1

M2 M1
0 Interaural 10 S1HL
RSD
+15 +10 +5 0 -5 S1Sh

Z
8 2

S1D
cg
RSGc
7 a S1BF 3
alv IG

df cc Or
chp LV
vhc CA3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 SFO
4
GrDG
fi S1ULp
MoDG RtSt
st chp
AD
ec cst sm D3V
5 5
MD AVDM
PVA AVVL
PT S2
PC
CM CPu
4 VA/VL 6
IAD
ic AM EGP
IAM Rt
DCl GI
AMV
mt DI
3 7
Re B
VRe VCl
rf ANS
PaXi Xi AIP

ns JPLH
PaDC PaLM ASt
2 PaMP 3V EA
CeM DEn 8
f CeC
PaV BLA
SPa AHP PLH AA
cst vlh mfb I 3
AHC I VEn
SO 2
1 Pe MeAD 9
BMA Pir1
VMHA VMHSh
opt TuLH BAOT ACo 3
sod MeAV
Arc RChL 2
a RCh
0 af
CxA1 10

Interaural 7.28 mm Bregma -1.72 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex AHP ant hypothal post AVVL AV th ventrolat cg cingulum df dorsal fornix I intercalated nu
3 layer 3 cortex AIP agran insular post B basal nu chp choroid plexus DI dysgran insular IAD interanterodors nu
3V 3rd ventricle alv alveus of hipp BAOT bed nu acc olf tr CM centr med th nu EA sublentic ext am IAM interanteromed nu
a artery AM anteromed th nu BLA basolat amyg ant CPu caudate putamen ec external capsule ic internal capsule
AA ant amyg area AMV anteromed th vent BMA basomed amyg ant cst commiss st term EGP ext globus pallidus IG indusium griseum
ACo ant cortical amyg ANS acc neurosecret CA3 field CA3 hipp CxA1 cx-amyg trans 1 f fornix JPLH juxtaparav lat hy
AD anterodors th nu Arc arcuate hy nu cc corpus callosum D3V dorsal 3rd vent fi fimbria of hipp LDVL LD ventrolat
af amygdal fissure ASt amygdalostr trans CeC central amyg caps DCl dorsal claustrum GI granular insular LV lat ventricle
AHC ant hypothal cent AVDM AV th dorsomed CeM central amyg med DEn dorsal endopirif GrDG granular dent gy
RBSC5 Book.qxp 16/9/04 16:36 Page 48

M1 primary motor cx MoDG molecular dent gy PaV Pa, ventral part Py pyramidal cells Rt reticular th nu SFO subfornical organ VA/VL VA and VL VMHA VMH, anterior
M2 2ary motor cx mt mammillothal tr PaXi paraxiphoid nu RCh retrochiasm area RtSt reticluostrial nu sm stria medullaris VCl ventral claustrum VMHSh VMH shell
MD mediodorsal th nu ns nigrostriat bundle PC paracentral th nu RChL retrochiasm lat S1BF S1 cx, barrel field SO supraoptic nu VEn ventral endopir VRe vent reuniens nu
ME median eminence opt optic tract Pe periventric hy nu Re reuniens th nu S1DZ S1 dysgranular zn sod supraoptic decussn vhc ventral hipp comm Xi xiphoid th nu
MeAD med anterodorsal Or oriens layer hipp Pir1 piriform layer 1 rf rhinal fissure S1HL S1 hindlimb region SPa subparaventric zn VL ventrolat th nu
MeAV med anterovent PaDC Pa, dorsal cap PLH peduncular lat hy Rh rhomboid thal nu S1Sh S1 shoulder region st stria terminalis vlh vent lat hy tr
mfb med forebr bundle PaLM Pa lat magnocell PT paratenial th nu RSD retrosple dysgran S1ULp S1 upper lip region TuLH tuberal lat hy VM ventromed nu
MHb med habenular nu PaMP Pa med parvicell PVA paraventric th ant RSGc RSG, c region S2 2ary somatosens VA ventral ant th nu VMH ventromed hy nu

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 48
5 5
9 1
M2 M1
0 Interaural 10 S1HL
RSD
+15 +10 +5 0 -5 S1Sh
8 2

Z
S1D
cg
RSGc

7 S1BF 3
alv IG
Or
CA3 LV
chp df cc
Py
vhc
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
SFO
fi GrDG LDVL S1ULp
st RtSt
ec chp MoDG
AD AVVL
cst sm D3V MHb
5 5
MD PC AVDM
PVA S2
PT
VL CPu
IAD
4 CM 6
AM
ic IAM Rt
AMV EGP GI
VA
mt
Rh DCl
VM DI
3 7
Re
VRe ANS VCl
rf B
AIP
ns Xi PaXi
PaDC JPLH
PaMP ASt
2 SPa PaV PaLM 8
CeM DEn
f EA CeC
mfb AHP BLA
PLH
cst AHC MeAD AA
vlh I 3
1 3V Pe MeAV I VEn 9
2
SO BMA Pir1
opt VMH TuLH
sod VMHSh
a BAOT ACo
Arc
0 RChL 10
ME

Interaural 7.20 mm Bregma -1.80 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 49

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 49
5 5
9 1
M2 M1 S1HL
0 Interaural 10 RSD
+15 +10 +5 0 -5 S1Sh

Z
S1D
8 2
cg
RSGc

7 alv Or S1BF
3
IG
SLu CA3 LV
chp df cc Rad Py

dhc
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
ec fi GrDG
LDVL RtSt
st
MoDG S1ULp
D3V AD AVVL
cst MHb
sm
5 5
MD AVDM
PC
PVA
PT VL S2
CM
4 Rt CPu 6
B
ic AM
IAM VA
AMV EGP
GI
mt Rh
DCl
3 Sub VM DI 7
Re VCl
rf VRe B
ns ZIR IGP
PaXi AIP
PaDC CeL ASt
2 PaMP PaLM ANS 8
DEn
mfb f 3V PaV AHP PLH
CeM
CeC BLA
cst SPa
vlh AHC IM
Pe MeAD VEn
1 3 9
VMHSh BLV 2
BMA
opt VMH TuLH Pir1
ArcD MeAV
sod
a SOR ACo
ArcM BAOT
0 RChL 10
MEI ArcL
MEE
Interaural 7.08 mm Bregma -1.92 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex af amygdal fissure ANS acc neurosecret B basal nu CeC central amyg caps cst commiss st term ec external capsule ic internal capsule
3 layer 3 cortex AHC ant hypothal cent ArcD arcuate hy dors BAOT bed nu acc olf tr CeL central amyg lat D3V dorsal 3rd vent EGP ext globus pallidus IG indusium griseum
3V 3rd ventricle AHP ant hypothal post ArcL arcuate hy lat BLA basolat amyg ant CeM central amyg med DCl dorsal claustrum f fornix IGP int globus pallidus
a artery AIP agran insular post ArcM arcuate hy med BLV basolat amyg vent cg cingulum DEn dorsal endopirif fi fimbria of hipp IM main ntercalated
A13 A13 dopamine alv alveus of hipp ASt amygdalostr trans BMA basomed amyg ant chp choroid plexus df dorsal fornix GI granular insular LaDL lat amyg dorsolat
ACo ant cortical amyg AM anteromed th nu AVDM AV th dorsomed CA3 field CA3 hipp CM centr med th nu dhc dorsal hipp comm GrDG granular dent gy LDDM LD dorsomed
AD anterodors th nu AMV anteromed th vent AVVL AV th ventrolat cc corpus callosum CPu caudate putamen DI dysgran insular IAM interanteromed nu LDVL LD ventrolat
LV lat ventricle
RBSC5 Book.qxp 16/9/04 16:36 Page 50

M1 primary motor cx MHb med habenular nu PaPo Pa, post part Py pyramidal cells RtSt reticluostrial nu sod supraoptic decussn VEn ventral endopir VRe vent reuniens nu
M2 2ary motor cx MoDG molecular dent gy PaV Pa, ventral part Rad radiatum layer S1BF S1 cx, barrel field SOR SO retrochiasm VL ventrolat th nu ZIR zona incerta rost
MD mediodorsal th nu mt mammillothal tr PaXi paraxiphoid nu RChL retrochiasm lat S1DZ S1 dysgranular zn SPa subparaventric zn vlh vent lat hy tr
MDL mediodorsal lat ns nigrostriat bundle PC paracentral th nu Re reuniens th nu S1HL S1 hindlimb region st stria terminalis VM ventromed nu
MeAD med anterodorsal opt optic tract Pe periventric hy nu rf rhinal fissure S1Sh S1 shoulder region Stg stigmoid hy nu VMH ventromed hy nu
MeAV med anterovent Or oriens layer hipp Pir1 piriform layer 1 Rh rhomboid thal nu S1ULp S1 upper lip region Sub submedius th nu VMHC VMH, central
MEE med eminence ext PaDC Pa, dorsal cap PLH peduncular lat hy RSD retrosple dysgran S2 2ary somatosens TuLH tuberal lat hy VMHDM VMH, dorsomed
MEI med eminence int PaLM Pa lat magnocell PT paratenial th nu RSGc RSG, c region SLu stratum lucidum VA ventral ant th nu VMHSh VMH shell
mfb med forebr bundle PaMP Pa med parvicell PVA paraventric th ant Rt reticular th nu sm stria medullaris VCl ventral claustrum VMHVL VMH, ventrolat

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 50
5 5
9 1

10 M2 M1
0 Interaural S1HL
RSD
+15 +10 +5 0 -5 S1Sh
8 2

DZ
S1
cg
RSGc

7 alv IG
Or 3
Py
SLu S1BF
df cc Rad
chp LV
ec dhc CA3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

fi
LDDM RtSt
st MHb MoDG LDVL
GrDG D3V AD
cst sm S1ULp
5 5
MDL MD AVDM
PVA
AVVL
PT
VL S2
CM PC CPu
4 6
Rt
AM VA
ic IAM
EGP GI
Rh
3 mt 7
Sub DI
Re VM
rf
VRe B
ns ZIR AIP
PaXi A13 IGP ASt LaDL
2 PaMP PaPo CeL CeC 8
SPa Stg ANS CeM
Pe DEn
cst AHP
f PLH
mfb BLA
vlh AHC MeAD
1 IM 9
VMHDM VEn
VMHSh
opt BLV
sod TuLH BMA 3
MeAV
VMHC 3V VMHVL 2
ArcM ArcD ACo Pir1
0 ArcL RChL BAOT 10
SOR
MEE MEI
af
Interaural 6.96 mm Bregma -2.04 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 51

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 51
5 5
9 1
M2 M1
0 Interaural 10 S1HL
RSD
+15 +10 +5 0 -5 S1Sh
8 S1DZ 2

cg
RSGc
alv
7 Or 3
Py S1BF
IG
df cc CA3
chp LV
Rad
dhc
MoDG SLu
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
PoDG
GrDG
fi
RtSt S1ULp
st MoDG LDDM LDVL
ec D3V
sm MHb AD
5 cst 5
AVDM
MDL PV MDM
CL S2
PT VL
AVVL
PC Rt
4 CM CPu 6
VPL
GI
ic AM
IAM
VA
EGP DI
mt Rh
3 7
Sub
rf VM B AIP
Re
PaXi VRe IGP LaDL
ns ZIR
Xi
2 A13 PaPo CeL 8
CeM CeC
Stg DEn
cst mfb Pe PLH
f 3V AHP BLA
vlh PTe
1 PeFLH MeAD IM VEn 9
VMHDM
VMHC
opt VMHSh BMA BLV
TuLH
sod MeAV 3
VMHVL
ACo 2
0 SOR BAOT PLCo Pir1 10
ArcL RChL
ArcD ArcM
MEI
Interaural 6.84 mm MEE Bregma -2.16 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex 6a layer 6a cortex alv alveus of hipp BAOT bed nu acc olf tr CeC central amyg caps cst commiss st term ec external capsule ic internal capsule
2 layer 2 cortex 6b layer 6b cortex AM anteromed th nu BLA basolat amyg ant CeL central amyg lat D3V dorsal 3rd vent EGP ext globus pallidus IG indusium griseum
3 layer 3 cortex a artery ArcD arcuate hy dors BLV basolat amyg vent CeM central amyg med DA dorsal hy area f fornix IGP int globus pallidus
3V 3rd ventricle A13 A13 dopamine ArcL arcuate hy lat BMA basomed amyg ant cg cingulum DEn dorsal endopirif fi fimbria of hipp IM main ntercalated
4 layer 4 cortex ACo ant cortical amyg ArcM arcuate hy med CA1 field CA1 hipp chp choroid plexus df dorsal fornix GI granular insular LaDL lat amyg dorsolat
5 layer 5 cortex AD anterodors th nu AVDM AV th dorsomed CA2 field CA2 hipp CL centrolat th nu dhc dorsal hipp comm GrDG granular dent gy LDDM LD dorsomed
5a layer 5a cortex AHP ant hypothal post AVVL AV th ventrolat CA3 field CA3 hipp CM centr med th nu DI dysgran insular I intercalated nu LDVL LD ventrolat
5b layer 5b cortex AIP agran insular post B basal nu cc corpus callosum CPu caudate putamen DMD dorsomedial dors IAM interanteromed nu LHb lat habenular nu
LV lat ventricle
RBSC5 Book.qxp 16/9/04 16:36 Page 52

M1 primary motor cx MHb med habenular nu Pe periventric hy nu Py pyramidal cells RtSt reticluostrial nu sod supraoptic decussn VA ventral ant th nu VPL vent posterolat nu
M2 2ary motor cx MoDG molecular dent gy PeFLH perifornical lat hy Rad radiatum layer S1BF S1 cx, barrel field SOR SO retrochiasm VEn ventral endopir VRe vent reuniens nu
MDL mediodorsal lat mt mammillothal tr Pir1 piriform layer 1 RChL retrochiasm lat S1DZ S1 dysgranular zn st stria terminalis VL ventrolat th nu Xi xiphoid th nu
MDM mediodorsal med ns nigrostriat bundle PLCo postlat cort amyg Re reuniens th nu S1HL S1 hindlimb region Stg stigmoid hy nu vlh vent lat hy tr ZIR zona incerta rost
MeAD med anterodorsal opt optic tract PLH peduncular lat hy rf rhinal fissure S1Sh S1 shoulder region STIA ST intraamygdal VM ventromed nu
MeAV med anterovent Or oriens layer hipp PoDG polymorph dent gy Rh rhomboid thal nu S1ULp S1 upper lip region Sub submedius th nu VMHC VMH, central
MEE med eminence ext PaPo Pa, post part PT paratenial th nu RSD retrosple dysgran S2 2ary somatosens SubD submedius dors VMHDM VMH, dorsomed
MEI med eminence int PaXi paraxiphoid nu PTe paraterete nu RSGc RSG, c region SLu stratum lucidum SubV submedius vent VMHSh VMH shell
mfb med forebr bundle PC paracentral th nu PV paraventric th nu Rt reticular th nu sm stria medullaris TuLH tuberal lat hy VMHVL VMH, ventrolat

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 52
5 5
9 1
M2 M1
RSD S1HL
0 Interaural 10
S1Sh
+15 +10 +5 0 -5

Z
S1D
8 2
2 4 1
cg 2
RSGc 1 3 5a 5b 6a 6b S1BF 3
alv
4
Or
7 Py 5 3
IG CA1 CA2
Rad SLu 6a
chp df cc LV
CA3 6b
dhc CA3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 PoDG MoDG 4
fi
RtSt S1ULp
st GrDG MoDG LDVL
D3V LDDM
cst sm AD
5 5
ec LHb MHb AVDM
PV MDM MDL S2
CL VL Rt

4 PC CPu GI
6
CM VPL
VA
ic AM
DI
Rh SubD
EGP
3 rf 7
mt Re VM AIP
SubV
Xi B
VRe IGP
ns PaXi ZIR CeL LaDL
2 cst A13 CeM 8
3V DA DEn
Stg PaPo CeC
mfb AHP
Pe PLH
f VMHDM DMD BLA
PeFLH STIA
vlh PTe MeAD I 3
1 2
9
opt VMHC VMHSh VEn Pir1
sod TuLH BMA BLV
VMHSh MeAV
VMHVL
a ArcD ArcM ACo
0 PLCo 10
RChL SOR
ArcL
MEE MEI
Interaural 6.72 mm Bregma -2.28 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 53

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 53
5 5
9 1
M2 M1
RSD S1Tr
0 Interaural 10

+15 +10 +5 0 -5

Z
S1D
8 2
cg
alv RSGc
Or CA2
S1BF
7 IG Py Rad 3
CA1
df cc SLu
chp MoDG LV
dhc GrDG
CA3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 GrDG 4
fi PoDG
ec S1ULp
st MoDG LDVL
sm D3V LDDM
cst
5 LHb MHb 5
PV CL
MDM AVDM S2
IMD MDL AngT
Rt

4 VL 6
PC GI
CM VPL CPu
ic
AM
DI
SubD EGP
Rh
3 rf 7
SubV AIP
mt VM
Re B
Xi VRe
ns ZIR IGP LaDL
PaXi A13 CeL
2 cst 8
DA CeC
mfb MePD CeM DEn
PeFLH
DMD PLH
Pe BLA BLP
f VMHC STIA
vlh PTe
1 opt I 9
VMHDM MeAD
3V VMHSh VEn
BMA 3
sod TuLH BLV 2
MeAV Pir1
a ArcD VMHVL ACo
0 SOR 10
RChL
PLCo
ArcL
ArcM MEE MEI
Interaural 6.60 mm Bregma -2.40 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex AIP agran insular post AVDM AV th dorsomed CA2 field CA2 hipp CL centrolat th nu dhc dorsal hipp comm GrDG granular dent gy LaDL lat amyg dorsolat
2 layer 2 cortex alv alveus of hipp B basal nu CA3 field CA3 hipp CM centr med th nu DI dysgran insular hif hipp fissure LDDM LD dorsomed
3 layer 3 cortex AM anteromed th nu BLA basolat amyg ant cc corpus callosum CPu caudate putamen DMD dorsomedial dors I intercalated nu LDVL LD ventrolat
3V 3rd ventricle AngT angular th nu BLP basolat amyg post CeC central amyg caps cst commiss st term ec external capsule ic internal capsule LHb lat habenular nu
a artery ArcD arcuate hy dors BLV basolat amyg vent CeL central amyg lat D3V dorsal 3rd vent EGP ext globus pallidus IG indusium griseum LMol lacunosum molec
A13 A13 dopamine ArcL arcuate hy lat BMA basomed amyg ant CeM central amyg med DA dorsal hy area f fornix IGP int globus pallidus LV lat ventricle
ACo ant cortical amyg ArcM arcuate hy med BMP basomed am post cg cingulum DEn dorsal endopirif fi fimbria of hipp IMD intermediodors nu
af amygdal fissure ASt amygdalostr trans CA1 field CA1 hipp chp choroid plexus df dorsal fornix GI granular insular IMG intramedull gray
RBSC5 Book.qxp 16/9/04 16:36 Page 54

M1 primary motor cx MePD med posterodors PC paracentral th nu PoDG polymorph dent gy RSD retrosple dysgran sm stria medullaris VL ventrolat th nu VRe vent reuniens nu
M2 2ary motor cx mfb med forebr bundle Pe periventric hy nu PTe paraterete nu RSGc RSG, c region sod supraoptic decussn vlh vent lat hy tr Xi xiphoid th nu
MCLH magnocell lat hy MHb med habenular nu PeFLH perifornical lat hy PV paraventric th nu Rt reticular th nu SOR SO retrochiasm VM ventromed nu ZI zona incerta
MDL mediodorsal lat MoDG molecular dent gy Pir piriform cx Py pyramidal cells S1BF S1 cx, barrel field st stria terminalis VMHC VMH, central ZIR zona incerta rost
MDM mediodorsal med mt mammillothal tr Pir1 piriform layer 1 Rad radiatum layer S1DZ S1 dysgranular zn STIA ST intraamygdal VMHDM VMH, dorsomed
MeAD med anterodorsal ns nigrostriat bundle PLCo postlat cort amyg RChL retrochiasm lat S1Tr S1 trunk region SubD submedius dors VMHSh VMH shell
MeAV med anterovent opt optic tract PLCo1 postlat cort am 1 Re reuniens th nu S1ULp S1 upper lip region SubV submedius vent VMHVL VMH, ventrolat
MEE med eminence ext Or oriens layer hipp PLH peduncular lat hy rf rhinal fissure S2 2ary somatosens TuLH tuberal lat hy VPL vent posterolat nu
MEI med eminence int PaXi paraxiphoid nu Po post thalamic nu Rh rhomboid thal nu SLu stratum lucidum VEn ventral endopir VPM vent posteromed

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 54
5 5
9 1
M2 M1
0 Interaural 10 RSD S1Tr
+15 +10 +5 0 -5

Z
S1D
8 2
cg
alv RSGc
Or
Py S1BF
7 CA2 IG Rad 3
CA1
df cc LMol SLu
chp dhc GrDG LV
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 hif 4
CA3 PoDG S1ULp
fi
st LDDM LDVL
sm D3V
cst LHb
5 ec CL 5
MHb
PV S2
MDM MDL AngT
IMD Po
4 Rt 6
PC VPL CPu GI
ic CM
VPM
VL DI
rf Rh SubD EGP
3 7
SubV VM AIP
mt Re B
VRe ASt
ZI IGP CeL
PaXi LaDL
2 cst A13 MePD
8
ns CeC
DA MCLH CeM IMG DEn
mfb Pe
DMD PeFLH
3V STIA
PLH BLA BLP Pir
vlh f VEn 3 2 1
1 opt I 9
PTe
MeAD
sod VMHDM VMHSh BMP
VMHC BMA BLV
TuLH MeAV
VMHVL
a ArcD ArcM ACo 3
0 ArcL
SOR 10
RChL 2
PLCo1
af
Interaural 6.48 mm MEE
MEI
Bregma -2.52 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 55

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 55
5 5
9 M1
1
M2
RSD S1Tr
0 Interaural 10

Z
+15 +10 +5 0 -5

S1D
8 cg 2
alv RSGc
Or
Py S1BF
IG Rad
7 CA2 3
dcw CA1
df cc MoDG SLu
dhc LMol GrDG CA3
chp LV
hif CA3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 S1ULp 4
fi PoDG

st LDVL
sm D3V LDDM
cst LHb
5 MHb S2 5
ec PV MDC
CL
MDM AngT
MDL Po Rt
IMD PC
4 GI 6
VPL CPu
CM VPM
VL DI
ic
ml Rh SubD EGP
3 7
rf SubV VM LaDL AIP
B
Re
mt ASt
VRe IGP
cst ZI CeL
ns PaXi A13 LaVM
2 CeM CeC 8
mfb DA IMG DEn
PeFLH
3V MCLH LaVL
DMD PeF PLH STIA
opt Pe I BLA BLP
MePD I
f VEn
vlh PTe
1 MeAD 9
sod 3
VMHDM VMHSh BMA BMP 2 Pir
VMHC TuLH BLV 1
a MTu MePV
a VMHVL ACo 3

0 3V
2 10
PLCo1
ArcDArcM MEI ArcL
MEE
Interaural 6.36 mm Bregma -2.64 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex alv alveus of hipp BLP basolat amyg post CeC central amyg caps cst commiss st term DMD dorsomedial dors I intercalated nu LaVM lat am ventromed
2 layer 2 cortex AngT angular th nu BLV basolat amyg vent CeL central amyg lat D3V dorsal 3rd vent ec external capsule ic internal capsule LDDM LD dorsomed
3 layer 3 cortex ArcD arcuate hy dors BMA basomed amyg ant CeM central amyg med DA dorsal hy area EGP ext globus pallidus IG indusium griseum LDVL LD ventrolat
3V 3rd ventricle ArcL arcuate hy lat BMP basomed am post cg cingulum dcw deep cerebr white f fornix IGP int globus pallidus LHb lat habenular nu
a artery ArcM arcuate hy med CA1 field CA1 hipp chp choroid plexus DEn dorsal endopirif fi fimbria of hipp IMD intermediodors nu LHbL lat habenular lat
A13 A13 dopamine ASt amygdalostr trans CA2 field CA2 hipp CL centrolat th nu df dorsal fornix GI granular insular IMG intramedull gray LMol lacunosum molec
ACo ant cortical amyg B basal nu CA3 field CA3 hipp CM centr med th nu dhc dorsal hipp comm GrDG granular dent gy LaDL lat amyg dorsolat LV lat ventricle
AIP agran insular post BLA basolat amyg ant cc corpus callosum CPu caudate putamen DI dysgran insular hif hipp fissure LaVL lat am ventrolat
RBSC5 Book.qxp 16/9/04 16:36 Page 56

M1 primary motor cx MePD med posterodors opt optic tract PLCo1 postlat cort am 1 Re reuniens th nu S1ULp S1 upper lip region SubV submedius vent VMHVL VMH, ventrolat
M2 2ary motor cx MePV med posterovent Or oriens layer hipp PLH peduncular lat hy rf rhinal fissure S2 2ary somatosens TuLH tuberal lat hy VPL vent posterolat nu
MCLH magnocell lat hy mfb med forebr bundle PaXi paraxiphoid nu PMCo postmed cort am Rh rhomboid thal nu SLu stratum lucidum VEn ventral endopir VPM vent posteromed
MDC mediodorsal cent MHb med habenular nu PC paracentral th nu Po post thalamic nu RSD retrosple dysgran sm stria medullaris VL ventrolat th nu VRe vent reuniens nu
MDL mediodorsal lat ml medial lemniscus Pe periventric hy nu PoDG polymorph dent gy RSGc RSG, c region sod supraoptic decussn vlh vent lat hy tr ZI zona incerta
MDM mediodorsal med MoDG molecular dent gy PeF perifornical nu PTe paraterete nu Rt reticular th nu st stria terminalis VM ventromed nu ZID zona incerta does
MeAD med anterodorsal mt mammillothal tr PeFLH perifornical lat hy PV paraventric th nu S1BF S1 cx, barrel field STIA ST intraamygdal VMHC VMH, central ZIV zona incerta vent
MEE med eminence ext MTu medial tuberal nu Pir piriform cx Py pyramidal cells S1DZ S1 dysgranular zn SubD submedius dors VMHDM VMH, dorsomed
MEI med eminence int ns nigrostriat bundle Pir1 piriform layer 1 Rad radiatum layer S1Tr S1 trunk region SubI subincertal nu VMHSh VMH shell

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 56
5 5
9 1
M1
M2
10 RSD S1Tr
0 Interaural
+15 +10 +5 0 -5

Z
S1D
8 2
cg
alv
RSGc Or
dcw
Py
CA1 Rad S1BF
7 IG 3
LMol CA2
df cc MoDG
SLu
dhc CA3
chp CA3 LV
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 hif 4
PoDG
fi GrDG S1ULp
LDDM LDVL
st sm D3V MHb
LHb
5 ec cst 5
PV MDC CL
MDM S2
MDL Po
IMD Rt
4 PC VPM CPu 6
VPL
CM GI
ic VL

Rh SubD DI
ml EGP
3 7
rf SubV VM
cst Re
mt ASt LaDL AIP
VRe ZID
PaXi CeL
A13 IGP IMG
2 ZIV 8
SubI CeC LaVM
ns DA CeM
sod mfb Pe PeFLH
opt LaVL DEn
DMD MePD STIA
3V PeF MCLH BLA
BLP
1 f PLH 9
vlh BMA VEn
VMHDM
VMHSh TuLH PTe 3
MePV BMP BLV 2
VMHC
a MTu Pir1
VMHVL PMCo 3
a
0 2 10
PLCo1
ArcD ArcM MEE MEI ArcL

Interaural 6.24 mm Bregma -2.76 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 57

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 57
5 5
9 1

0 Interaural 10 M2 M1
S1Tr
+15 +10 +5 0 -5 RSD

Z
S1D
8 2

cg
alv
RSGc Or
Py S1BF
7 Rad 3
IG LMol CA2
CA1 MoDG
df cc
dcw SLu
CA3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 chp dhc LV 4
hif FC S1ULp
fi
GrDG PoDG MoDG LDVL
sm D3V
st MHb LDDM
5 LHbL 5
cst LHbM
CL
PV S2
ec MDC MDL
MDM Rt
Po
IMD
4 6
VPM GI
PC VPL CPu
CM
ic VL
EGP DI
3 Rh 7
ml Sub B
rf
cst VM CeM AIP
Re ZID
VRe ASt LaDL
mt CeL
PHD ZIV LaVM
2 A13 CeC 8
DA
ns SubI LaVL
opt sod DEn
mfb PeFLH
Pe PLH MePD STIA
DMD PeF BLA
3V MCLH BLP 3
f I
1 VEn 2 9
I Pir1
VMHDM TuLH PTe BLV
MePV BMP
a VMHSh VMHC MTu RAPir
a PMCo 3
VMHVL
2
0 PLCo1 10
ArcDArcM MEI ArcL
MEE
Interaural 6.08 mm Bregma -2.92 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

2 layer 2 cortex ArcL arcuate hy lat BMP basomed am post cg cingulum dcw deep cerebr white ec external capsule GrDG granular dent gy LaVM lat am ventromed
3 layer 3 cortex ArcM arcuate hy med CA1 field CA1 hipp chp choroid plexus DEn dorsal endopirif Ect ectorhinal cx hif hipp fissure LDDM LD dorsomed
3V 3rd ventricle ASt amygdalostr trans CA2 field CA2 hipp CL centrolat th nu df dorsal fornix EGP ext globus pallidus I intercalated nu LDVL LD ventrolat
a artery AuD 2ary aud cx, dors CA3 field CA3 hipp CM centr med th nu dhc dorsal hipp comm f fornix ic internal capsule LHbL lat habenular lat
A13 A13 dopamine B basal nu cc corpus callosum CPu caudate putamen DI dysgran insular FC fasciola cinereum IG indusium griseum LHbM lat habenular med
AIP agran insular post BLA basolat amyg ant CeC central amyg caps cst commiss st term DM dorsomedial hy nu fi fimbria of hipp IMD intermediodors nu LMol lacunosum molec
alv alveus of hipp BLP basolat amyg post CeL central amyg lat D3V dorsal 3rd vent DMC dorsomed compact fr fasc retroflexus LaDL lat amyg dorsolat LV lat ventricle
ArcD arcuate hy dors BLV basolat amyg vent CeM central amyg med DA dorsal hy area DMD dorsomedial dors GI granular insular LaVL lat am ventrolat
RBSC5 Book.qxp 16/9/04 16:36 Page 58

M1 primary motor cx MePV med posterovent Or oriens layer hipp PMCo postmed cort am RAPir rost amygdalopir S1Tr S1 trunk region SubI subincertal nu VMHVL VMH, ventrolat
M2 2ary motor cx mfb med forebr bundle PC paracentral th nu Po post thalamic nu Re reuniens th nu S1ULp S1 upper lip region Te terete hypothal nu VPL vent posterolat nu
MCLH magnocell lat hy MHb med habenular nu Pe periventric hy nu PoDG polymorph dent gy rf rhinal fissure S2 2ary somatosens TuLH tuberal lat hy VPM vent posteromed
MDC mediodorsal cent ml medial lemniscus PeF perifornical nu PRh perirhinal cx Rh rhomboid thal nu SLu stratum lucidum VEn ventral endopir VRe vent reuniens nu
MDL mediodorsal lat MoDG molecular dent gy PeFLH perifornical lat hy PTe paraterete nu RSD retrosple dysgran sm stria medullaris VL ventrolat th nu ZID zona incerta does
MDM mediodorsal med mt mammillothal tr PHD post hy area dors PV paraventric th nu RSGb RSG, b region sod supraoptic decussn VM ventromed nu ZIV zona incerta vent
MEE med eminence ext MTu medial tuberal nu Pir1 piriform layer 1 PVP paraventr th post Rt reticular th nu st stria terminalis VMHC VMH, central
MEI med eminence int ns nigrostriat bundle PLCo1 postlat cort am 1 Py pyramidal cells S1BF S1 cx, barrel field STIA ST intraamygdal VMHDM VMH, dorsomed
MePD med posterodors opt optic tract PLH peduncular lat hy Rad radiatum layer S1DZ S1 dysgranular zn Sub submedius th nu VMHSh VMH shell

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 58
5 5
9 1
M2 M1
0 Interaural 10 RSD S1Tr
+15 +10 +5 0 -5

Z
S1D
8 2
cg

alv RSGc Or
Py S1BF
dcw Rad
7 alv IG CA1 LMol 3
CA2
df MoDG
cc
dhc SLu
CA3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 FC 4
chp LV
hif PoDG S1ULp
MoDG
fi
D3V GrDG LDVL
ec sm MHb LDDM
st S2
5 LHbL 5
cst CL
fr LHbM
PVP
MDC MDL Po AuD
MDM Rt
4 IMD 6
VPM
PC
VPL
CM VL Ect
ic EGP CPu
3 cst Rh 7
ml Sub
VM B
rf
st Re ZID PRh
VRe ASt LaDL
mt CeL
ZIV
2 ns PHD LaVM 8
opt DA SubI CeC
mfb
LaVL DEn
sod 3V PeFLH PLH
DMD MePD STIA
Pe BLA
PeF MCLH 3
DMC I BLP
1 f 2 9
PTe I VEn Pir1
VMHSh TuLH BLV
VMHDM BMP
ArcD Te MePV
a VMHC MTu
ArcM RAPir
a 3
VMHVL PMCo
0 2
10
ArcL PLCo1
MEE MEI
Interaural 6.00 mm Bregma -3.00 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 59

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 59
5 5
9 1
M2 M1 LPtA
RSD S1Tr

Z
0 Interaural 10

S1D
+15 +10 +5 0 -5
8 cg 2
alv
RSGc
Or S1BF
Py
Rad
CA1
dcw IG LMol
7 MoDG CA2 3
df cc
dhc SLu
FC CA3 CA3 S1ULp
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 chp MoDG LV 4
hif PoDG
chp S2
fi LPMR GrDG LDVL
sm LDDM
D3V
MHb LHbL
5 ec 5
cst fr CL
LHbM PVP AuD
MDL Po
MDC
MDM Rt
IMD
4 6
VPM
PC VPL CPu Ect
CM
ic VL EGP
3 Rh B 7
ml Sub
rf
VM PRh
Re VRe
st ZID ASt LaDL
CeL
ZIV LEnt
opt mt
2 sod STh CeC LaVM LaVL 8
ns PHD DA A11
SubI
mfb DMD STIA
Pe PeFLH MePD
MCLH BLA DEn
DMC PeF I BLP
1 3V DMV PLH
9
f
PTe BMP BLV 3
MePV 2
a VMHDM TuLH 1
Te AHiAL Pir
VMHSh MTu RAPir
a VMHC VMHVL PMCo 3
0 ArcD ArcM 10
2
ArcL PLCo1

Interaural 5.88 mm MEE MEI

Bregma -3.12 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex ArcL arcuate hy lat BLV basolat amyg vent chp choroid plexus df dorsal fornix eml ext medullary lam IG indusium griseum LHbL lat habenular lat
2 layer 2 cortex ArcM arcuate hy med BMP basomed am post CL centrolat th nu dhc dorsal hipp comm f fornix IMD intermediodors nu LHbM lat habenular med
3 layer 3 cortex ASt amygdalostr trans CA1 field CA1 hipp CM centr med th nu DLG dors lat geniculate FC fasciola cinereum imvc intmedvent comm LMol lacunosum molec
3V 3rd ventricle Au1 primary aud cx CA2 field CA2 hipp CPu caudate putamen DMC dorsomed compact fi fimbria of hipp LaDL lat amyg dorsolat LPMR LP mediorostral
a artery AuD 2ary aud cx, dors CA3 field CA3 hipp cst commiss st term DMD dorsomedial dors fr fasc retroflexus LaVL lat am ventrolat LPtA lat parietal assocn
A11 A11 dopamine AuV 2ary aud cx, vent cc corpus callosum D3V dorsal 3rd vent DMV dorsomed hy vent GrDG granular dent gy LaVM lat am ventromed LV lat ventricle
AHiAL AHi anterolat B basal nu CeC central amyg caps DA dorsal hy area ec external capsule hif hipp fissure LDDM LD dorsomed
alv alveus of hipp BLA basolat amyg ant CeL central amyg lat dcw deep cerebr white Ect ectorhinal cx I intercalated nu LDVL LD ventrolat
ArcD arcuate hy dors BLP basolat amyg post cg cingulum DEn dorsal endopirif EGP ext globus pallidus ic internal capsule LEnt lat entorhinal cx
RBSC5 Book.qxp 16/9/04 16:36 Page 60

M1 primary motor cx MePV med posterovent OPC oval paracent nu PLCo1 postlat cort am 1 Rad radiatum layer S1DZ S1 dysgranular zn STIA ST intraamygdal VMHC VMH, central
M2 2ary motor cx mfb med forebr bundle opt optic tract PLH peduncular lat hy RAPir rost amygdalopir S1Tr S1 trunk region Sub submedius th nu VMHDM VMH, dorsomed
MCLH magnocell lat hy MHb med habenular nu Or oriens layer hipp PMCo postmed cort am Re reuniens th nu S1ULp S1 upper lip region SubI subincertal nu VMHSh VMH shell
MDC mediodorsal cent ml medial lemniscus PC paracentral th nu Po post thalamic nu rf rhinal fissure S2 2ary somatosens Te terete hypothal nu VMHVL VMH, ventrolat
MDL mediodorsal lat MoDG molecular dent gy Pe periventric hy nu PoDG polymorph dent gy Rh rhomboid thal nu SLu stratum lucidum TuLH tuberal lat hy VPL vent posterolat nu
MDM mediodorsal med MPtA med parietal assn PeF perifornical nu PRh perirhinal cx RSD retrosple dysgran sm stria medullaris VL ventrolat th nu VPM vent posteromed
MEE med eminence ext mt mammillothal tr PeFLH perifornical lat hy PTe paraterete nu RSGc RSG, c region sod supraoptic decussn VLG vent lat genic nu VRe vent reuniens nu
MEI med eminence int MTu medial tuberal nu PHD post hy area dors PVP paraventr th post Rt reticular th nu st stria terminalis VM ventromed nu ZID zona incerta does
MePD med posterodors ns nigrostriat bundle Pir piriform cx Py pyramidal cells S1BF S1 cx, barrel field STh subthalamic nu VMH ventromed hy nu ZIV zona incerta vent

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 60
5 5
9 M2
1
MPtA
RSD LPtA
0 Interaural 10 S1Tr

Z
S1D
+15 +10 +5 0 -5
8 alv cg 2
RSGc
Or
S1BF
Py
Rad
CA1 LMol
IG
7 CA2
3
MoDG
df cc dhc
CA3 SLu
S2
FC
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 MoDG 4
chp hif PoDG LV
chp DLG AuD
fi D3V LDVL
GrDG
sm LPMR VLG
MHb
LHbL
5 st Au1 5
cst fr CL
PVP
ec LHbM MDL
eml MDC Rt
MDM Po
IMD AuV
4 6
VPM
PC VPL
CM Ect
imvc OPC CPu
ic
rf Rh
3 ml 7
Sub VM PRh
Re
st VRe ZID
opt CeL LaDL
ZIV LEnt
mt A11 ASt
sod PHD
2 STh 8
DA SubI CeC
ns LaVM
mfb PeFLH LaVL
Pe DMD DEn
MePD STIA
DMC PeF PLH
3V BLA
1 f I BLP 9
DMV
VMHSh TuLH Te 3
MePV BMP BLV 2 Pir
a AHiAL 1
VMH MTu
a ArcD PMCo RAPir
0 ArcM 3 10
ArcL
2
MEE MEI PLCo1
Interaural 5.76 mm Bregma -3.24 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 61

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 61
5 5
9 MPtA 1
RSD LPtA

0 Interaural 10 S1Tr

Z
S1D
+15 +10 +5 0 -5 cg
8 alv 2
RSGc
Or
Py S1BF
Rad
IG CA1 LMol
7 hif MoDG CA2 3
df cc
dcw
dhc CA3
FC SLu
PoDG S2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

MoDG
6 4
chp chp DLG LV
AuD
sm D3V GrDG LDVL VLG
fi LPMR
LHbL
5 st MHb LHbM Au1 5
fr CL
ec cst PVP

eml MDC MDL Po

AuV
MDM Rt
IMD
4 VPM 6
VPL
PC
CM CPu Ect
ic OPC
ml scp
3 7
rf Re
scp VM PRh
st VRe ZID LaDL
opt PHD ZIV ASt
LV LEnt
sod mt A11 STh
2 ns CeC LaVM 8
PH
mfb Pe LaVL
cp PeFLH MePD DEn
3V DMD BLA
PeF PLH STIA
BLP
1 f 3 9
MePV 2
DMC DMV BMP BLV 1
VMHSh TuLH AHiAL Pir
a Te
ArcD MTu PMCo RAPir
VMH 3
a
0 2 10
PLCo1
ArcM ArcL
MEE MEI
Interaural 5.64 mm Bregma -3.36 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex ArcL arcuate hy lat BLP basolat amyg post chp choroid plexus df dorsal fornix f fornix InfS infundibular stem LPLR LP laterorostral
2 layer 2 cortex ArcLP arcuate lateropost BLV basolat amyg vent CL centrolat th nu dhc dorsal hipp comm FC fasciola cinereum LaDL lat amyg dorsolat LPMR LP mediorostral
3 layer 3 cortex ArcM arcuate hy med BMP basomed am post CM centr med th nu DLG dors lat geniculate fi fimbria of hipp LaVL lat am ventrolat LPtA lat parietal assocn
3V 3rd ventricle ArcMP arcuate med post CA1 field CA1 hipp cp cerebral peduncle DMC dorsomed compact fr fasc retroflexus LaVM lat am ventromed LV lat ventricle
a artery ASt amygdalostr trans CA2 field CA2 hipp CPu caudate putamen DMD dorsomedial dors GrDG granular dent gy LDVL LD ventrolat
A11 A11 dopamine Au1 primary aud cx CA3 field CA3 hipp cst commiss st term DMV dorsomed hy vent hif hipp fissure LEnt lat entorhinal cx
AHiAL AHi anterolat AuD 2ary aud cx, dors cc corpus callosum D3V dorsal 3rd vent ec external capsule ic internal capsule LHbL lat habenular lat
alv alveus of hipp AuV 2ary aud cx, vent CeC central amyg caps dcw deep cerebr white Ect ectorhinal cx IG indusium griseum LHbM lat habenular med
ArcD arcuate hy dors BLA basolat amyg ant cg cingulum DEn dorsal endopirif eml ext medullary lam IMD intermediodors nu LMol lacunosum molec
RBSC5 Book.qxp 16/9/04 16:36 Page 62

MDC mediodorsal cent MHb med habenular nu opt optic tract Pir piriform cx Py pyramidal cells S1BF S1 cx, barrel field st stria terminalis VMHSh VMH shell
MDL mediodorsal lat ml medial lemniscus Or oriens layer hipp PLCo1 postlat cort am 1 Rad radiatum layer S1DZ S1 dysgranular zn STh subthalamic nu VPL vent posterolat nu
MDM mediodorsal med MoDG molecular dent gy PC paracentral th nu PLH peduncular lat hy RAPir rost amygdalopir S1Tr S1 trunk region STIA ST intraamygdal VPM vent posteromed
MEE med eminence ext MPtA med parietal assn Pe periventric hy nu PMCo postmed cort am Re reuniens th nu S2 2ary somatosens Te terete hypothal nu VPPC vent post parvic
MEI med eminence int mt mammillothal tr PeF perifornical nu Po post thalamic nu rf rhinal fissure scp sup cerebellar ped TuLH tuberal lat hy VRe vent reuniens nu
MePD med posterodors MTu medial tuberal nu PeFLH perifornical lat hy PoDG polymorph dent gy RSD retrosple dysgran SLu stratum lucidum VLG vent lat genic nu ZID zona incerta does
MePV med posterovent ns nigrostriat bundle PH post hy nu PRh perirhinal cx RSGc RSG, c region sm stria medullaris VM ventromed nu ZIV zona incerta vent
mfb med forebr bundle OPC oval paracent nu PHD post hy area dors PVP paraventr th post Rt reticular th nu sod supraoptic decussn VMH ventromed hy nu

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 62
5 5
9 MPtA
1
RSD
LPtA
0 Interaural 10

+15 +10 +5 0 -5
8 alv cg 2
RSGc
Or
Py S1BF
Rad
IG LMol
7 MoDG 3
df
cc
CA1 CA2
dhc CA3
FC
dcw PoDG SLu
S2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 hif MoDG 4
chp
chp D3V GrDG DLG LV AuD
sm LPLR
MHb LPMR VLG
LHbM LHbL
5 5
eml
fr CL Au1
st PVP
ec MDL
cst IMD Po Rt
fi MDM
4 AuV 6
cst VPM VPL
PC
OPC Ect
CM CPu
st
3 ml VPPC 7
rf ic Re
scp VM
ZID PRh
PHD LaDL
opt ZIV
LV LV LEnt
mt A11
2 STh 8
sod ns LaVM
3V PH
st mfb MePD
Pe DEn
PeFLH STIA
cp PLH
PeF BLP
1 DMD 3 2 9
f BMP 1
MePV Pir
DMC AHiAL
a TuLH Te
DMV MTu PMCo 3 RAPir
a
0 ArcMP ArcLP 2 10
PLCo1
InfS
Interaural 5.52 mm Bregma -3.48 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 63

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 63
5 5
9 MPtA 1
RSD
LPtA
0 Interaural 10

+15 +10 +5 0 -5
8 alv 2
cg RSGc
Or
Py S1BF
Rad
IG CA1 ol
7 LM MoDG 3
hif df cc
dcw CA2
dhc CA3
FC PoDG S2
CA3 SLu
MoDG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
AuD
chp
chp GrDG LPLR DLG LV
sm D3V LPMR
fi
MHb VLG
LHbM
5 LHbL Au1 5
ec
fr PVP CL
ic cst MDL Po Rt
IMD eml AuV
4 MDM VPL 6
st
VPM CPu
PC
CM Ect
OPC
3 ic ml VPPC 7
rf RRe
ZID PRh
scp
VM ZIV
PHD LaDL
opt
LV LEnt
sod A11
2 mt STh 8
ns 3V LaVM
PH
mfb MePD
st DEn Pir
STIA 3 2 1
cp PeF BLP
1 DTM PLH 9
MePV BMP
f PeFLH AHiAL
a DM Te RAPir
PMCo
PMV 3
3V 2
0 ArcLP 10
PLCo1
ArcMP
InfS
Interaural 5.40 mm Bregma -3.60 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex alv alveus of hipp BLP basolat amyg post chp choroid plexus dcw deep cerebr white ec external capsule GrDG granular dent gy LEnt lat entorhinal cx
2 layer 2 cortex APir amygdalopir trans BMP basomed am post CL centrolat th nu DEn dorsal endopirif Ect ectorhinal cx hif hipp fissure LHbL lat habenular lat
3 layer 3 cortex ArcLP arcuate lateropost CA1 field CA1 hipp CM centr med th nu df dorsal fornix eml ext medullary lam ic internal capsule LHbM lat habenular med
3V 3rd ventricle ArcMP arcuate med post CA2 field CA2 hipp cp cerebral peduncle dhc dorsal hipp comm f fornix IG indusium griseum LMol lacunosum molec
a artery Au1 primary aud cx CA3 field CA3 hipp CPu caudate putamen DLG dors lat geniculate FC fasciola cinereum IMD intermediodors nu LPLR LP laterorostral
A11 A11 dopamine AuD 2ary aud cx, dors cc corpus callosum cst commiss st term DM dorsomedial hy nu fi fimbria of hipp InfS infundibular stem LPMR LP mediorostral
AHiAL AHi anterolat AuV 2ary aud cx, vent cg cingulum D3V dorsal 3rd vent DTM dors tuberomamm fr fasc retroflexus LaDL lat amyg dorsolat LPtA lat parietal assocn
LaVM lat am ventromed LV lat ventricle
RBSC5 Book.qxp 16/9/04 16:36 Page 64

MDL mediodorsal lat MoDG molecular dent gy PC paracentral th nu PLCo1 postlat cort am 1 PSTh parasubthal nu RSD retrosple dysgran sm stria medullaris VLG1 VLG layer 1
MDM mediodorsal med MPtA med parietal assn PeF perifornical nu PLH peduncular lat hy PVP paraventr th post RSGc RSG, c region sod supraoptic decussn VM ventromed nu
MePD med posterodors mt mammillothal tr PeFLH perifornical lat hy PMCo postmed cort am Py pyramidal cells Rt reticular th nu st stria terminalis VPL vent posterolat nu
MePV med posterovent ns nigrostriat bundle PF parafascic th nu PMV premamm nu vent Rad radiatum layer S1BF S1 cx, barrel field STh subthalamic nu VPM vent posteromed
mfb med forebr bundle OPC oval paracent nu PH post hy nu Po post thalamic nu RAPir rost amygdalopir S2 2ary somatosens STIA ST intraamygdal VPPC vent post parvic
MHb med habenular nu opt optic tract PHD post hy area dors PoDG polymorph dent gy rf rhinal fissure scp sup cerebellar ped Te terete hypothal nu VTM vent tuberomamm
ml medial lemniscus Or oriens layer hipp Pir piriform cx PRh perirhinal cx RRe retroreuniens SLu stratum lucidum VLG vent lat genic nu ZID zona incerta does
ZIV zona incerta vent

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 64
5 5
9 1
MPtA
RSD
0 Interaural 10 LPtA
+15 +10 +5 0 -5
8 alv cg RSGc 2
Or
Py S1BF
Rad
IG LMol
7 CA1 3
cc MoDG
df dhc CA2
FC S2
hif PoDG
dcw CA3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 MoDG SLu 4
chp AuD
DG
Gr
chp sm D3V LPLR DLG
LPMR
fi MHb
5 LHbL 5
LHbM VLG1 Au1
CL
fr
ec PVP Po
PF
eml IMD MDL
st Rt AuV
4 6
MDM PC VPL
cst VPM CPu
CM OPC Ect
ic LV
3 RRe VPPC 7
rf ml ZID
opt PRh
scp PHD LaDL
A11 VM ZIV
3V LEnt
2 sod 8
ns mt chp LaVM
mfb PH
STh
PLH DEn
cp BLP
st DTM PeFLH
1 PSTh 9
AHiAL BMP
f 32
3V Te APir 1
DM Pir
PMV PMCo
VTM 3
ArcMP
0 ArcLP 2 10
PLCo1

Interaural 5.28 mm InfS

Bregma -3.72 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 65

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 65
5 5
9 RSD MPtA 1

0 Interaural 10 LPtA

+15 +10 +5 0 -5 cg
8 alv RSGc PtPD 2
Or
Py
Rad S1BF
IG CA1 LMol
7 MoDG 3
hif df cc
CA2
dhc CA3 S1
dcw FC
PoDG
S2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 MoDG 4
SLu
chp AuD
IMA
sm GrDG LPLR
D3V
LPMR DLG
MHb LHbM IGL
5 LHbL 5
fi Au1
CL VLG
ic Po
fr PVP
iml
PoMn MDL
ec eml PF AuV
4 PC VPM 6
MDM VPL Rt
CM CPu
OPC
opt Ect
RRe
3 VPPC 7
chp ml LV
SPF ZID
rf A11
ic PRh
scp ZIV
PR
LaDL LEnt
2 sod 3V PH 8
ns mt PLH
mfb LaVM
STh
DEn
PSTh
1 st cp PeFLH BLP 9
DTM
f AHiAL 3
BMP 2
a Te 1
PMV APir Pir
3V VTM PMCo
0 3 10
ArcLP
2
ArcMP
PLCo1
Interaural 5.16 mm InfS
Bregma -3.84 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex APir amygdalopir trans CA1 field CA1 hipp cp cerebral peduncle DLG dors lat geniculate fr fasc retroflexus iml int medullary lam LPLR LP laterorostral
2 layer 2 cortex ArcLP arcuate lateropost CA2 field CA2 hipp CPu caudate putamen DTM dors tuberomamm Gem gemini hy nu InfS infundibular stem LPMR LP mediorostral
3 layer 3 cortex ArcMP arcuate med post CA3 field CA3 hipp D3V dorsal 3rd vent ec external capsule GrDG granular dent gy LaDL lat amyg dorsolat LPtA lat parietal assocn
3V 3rd ventricle Au1 primary aud cx cc corpus callosum dcw deep cerebr white Ect ectorhinal cx hif hipp fissure LaVM lat am ventromed LV lat ventricle
a artery AuD 2ary aud cx, dors cg cingulum DEn dorsal endopirif eml ext medullary lam ic internal capsule LEnt lat entorhinal cx
A11 A11 dopamine AuV 2ary aud cx, vent chp choroid plexus df dorsal fornix f fornix IG indusium griseum LHbL lat habenular lat
AHiAL AHi anterolat BLP basolat amyg post CL centrolat th nu DG dentate gyrus FC fasciola cinereum IGL intergenic leaf LHbM lat habenular med
alv alveus of hipp BMP basomed am post CM centr med th nu dhc dorsal hipp comm fi fimbria of hipp IMA intramedul th area LMol lacunosum molec
RBSC5 Book.qxp 16/9/04 16:36 Page 66

MDL mediodorsal lat mt mammillothal tr PH post hy nu PoDG polymorph dent gy Rad radiatum layer S1BF S1 cx, barrel field st stria terminalis VTM vent tuberomamm
MDM mediodorsal med ns nigrostriat bundle Pir piriform cx PoMn posteromedian nu rf rhinal fissure S2 2ary somatosens STh subthalamic nu ZID zona incerta does
mfb med forebr bundle OPC oval paracent nu PLCo1 postlat cort am 1 PR prerubral field RRe retroreuniens scp sup cerebellar ped str sup thal radiation ZIV zona incerta vent
MHb med habenular nu opt optic tract PLH peduncular lat hy PRh perirhinal cx RSD retrosple dysgran SLu stratum lucidum Te terete hypothal nu
ml medial lemniscus Or oriens layer hipp PMCo postmed cort am PSTh parasubthal nu RSGb RSG, b region sm stria medullaris VLG vent lat genic nu
MoDG molecular dent gy PC paracentral th nu PMD premamm nu dors PtPD parietal postdors RSGc RSG, c region SMT submammilloth nu VPL vent posterolat nu
MPtA med parietal assn PeFLH perifornical lat hy PMV premamm nu vent PVP paraventr th post Rt reticular th nu sod supraoptic decussn VPM vent posteromed
MRe mamm recess 3V PF parafascic th nu Po post thalamic nu Py pyramidal cells S1 1ary somatosens SPF subparafascic nu VPPC vent post parvic

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 66
5 5
9 MPtA
1
RSD
LPtA
0 Interaural 10

+15 +10 +5 0 -5 PtPD

cg RSGc
8 alv 2
Or
Py
Rad S1BF
RSGb
LMol
7 IG MoDG 3
hif df cc
CA1
dcw dhc FC CA3 S1
PoDG CA2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 MoDG 4
SLu AuD
chp GrDG
IMA
sm LPLR
D3V DLG
LPMR
MHb
5 LHbL IGL Au1 5
LHbM CL
fi
VLG CPu
str PVP
ec fr Po
PF AuV
PF MDL
4 PoMn VPM 6
PC VPL
MDM Rt
CM OPC Ect
RRe
3 chp VPPC LV 7
opt ml A11 SPF
rf ZID PRh
ic ZIV
scp PR
sod 3V LaDL LEnt
2 PH 8
ns mt Gem STh
mfb SMT LaVM
SMT DEn
PLH
st
1 cp PSTh BLP 9
f PMD DG AHiAL 32
a Te BMP 1
APir Pir
MRe PMV
3V PMCo
VTM 3
0 ArcLP 10
2
ArcMP PLCo1

Interaural 5.04 mm Bregma -3.96 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 67

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 67
5 5
9 1
MPtA
RSD
LPtA
0 Interaural 10

+15 +10 +5 0 -5 cg PtPD

8 alv RSGc 2
Or
Py
Rad S1BF
RSGb LMol
7 MoDG 3
hif IG
df cc
CA2 S1
alv dhc CA3
PoDG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

FC MoDG AuD
6 4
GrDG SLu
chp IMA
LPLR
sm DLG
hbc LPMR
D3V Au1
5 IGL 5
MHb CL
LHbM VLG
32 1
LHbL
fi PVP PrC
str Po AuV
fr
4 VPM 6
PF VPL
dcw Rt
Ect
opt
chp SubG
3 VPPC LV 7
ic ml PRh
rf A11 SPF ZID
scp PR ZIV
LEnt
alv 3V F
sod LaDL
2 PH 8
ns
LaVM
Gem
mfb mt STh DEn
SMT PLH 32
1 st cp BLP 9
1
SuML BMP Pir
f AHiPM AHiPL
a
PMD
GrDG APir
PMCo 3
0 MRe ArcLP VTM 2 10
PLCo1
ArcMP
Interaural 4.92 mm Bregma -4.08 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex AHiPL AHi posterlat AuD 2ary aud cx, dors cc corpus callosum DEn dorsal endopirif FC fasciola cinereum ic internal capsule LHbL lat habenular lat
2 layer 2 cortex AHiPM AHi posteromed AuV 2ary aud cx, vent cg cingulum df dorsal fornix fi fimbria of hipp IG indusium griseum LHbM lat habenular med
3 layer 3 cortex alv alveus of hipp BLP basolat amyg post chp choroid plexus dhc dorsal hipp comm fr fasc retroflexus IGL intergenic leaf LM lat mammillary nu
3V 3rd ventricle APir amygdalopir trans BMP basomed am post CL centrolat th nu DLG dors lat geniculate Gem gemini hy nu IMA intramedul th area LMol lacunosum molec
a artery ArcLP arcuate lateropost CA1 field CA1 hipp cp cerebral peduncle Ect ectorhinal cx GrDG granular dent gy La lat amyg nu LPLR LP laterorostral
A11 A11 dopamine ArcMP arcuate med post CA2 field CA2 hipp D3V dorsal 3rd vent F nu fields of Forel hbc habenular comm LaDL lat amyg dorsolat LPMR LP mediorostral
af amygdal fissure Au1 primary aud cx CA3 field CA3 hipp dcw deep cerebr white f fornix hif hipp fissure LaVM lat am ventromed LPtA lat parietal assocn
LEnt lat entorhinal cx LV lat ventricle
RBSC5 Book.qxp 16/9/04 16:36 Page 68

mfb med forebr bundle MRe mamm recess 3V PLCo postlat cort amyg PrC precommiss nu rf rhinal fissure SLu stratum lucidum str sup thal radiation VPM vent posteromed
MHb med habenular nu mt mammillothal tr PLCo1 postlat cort am 1 PRh perirhinal cx RSD retrosple dysgran sm stria medullaris SubG subgeniculate nu VPPC vent post parvic
ML med mammill lat ns nigrostriat bundle PLH peduncular lat hy PtPD parietal postdors RSGb RSG, b region SMT submammilloth nu SuML supramamm lat VTM vent tuberomamm
ml medial lemniscus opt optic tract PMCo postmed cort am PtPR parietal postrost RSGc RSG, c region sod supraoptic decussn SuMM supramamm med ZID zona incerta does
MM med mammill med Or oriens layer hipp PMD premamm nu dors PVG periventric gray Rt reticular th nu SPF subparafascic nu V2ML V2 cx, mediolat ZIV zona incerta vent
MnM med mamm median PF parafascic th nu Po post thalamic nu PVP paraventr th post S1 1ary somatosens SPFPC SPF parvicell V2MM V2 cx, mediomed
MoDG molecular dent gy PH post hy nu PoDG polymorph dent gy Py pyramidal cells S1BF S1 cx, barrel field st stria terminalis VLG vent lat genic nu
MPtA med parietal assn Pir piriform cx PR prerubral field Rad radiatum layer scp sup cerebellar ped STh subthalamic nu VPL vent posterolat nu

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 68
5 5
9 1
RSD V2MM
V2ML
0 Interaural 10

+15 +10 +5 0 -5 PtPD

8 alv cg RSGc 2
Or PtPR
Py
RSGb Rad
CA1 LMol S1BF
7 IG MoDG 3
df cc
hif
alv dhc CA3 S1
FC PoDG CA2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 GrDG AuD 4
chp
IMA
SLu
sm hbc LPLR
LPMR DLG
D3V
LHbM MHb LHbL
5 Au1 5
IGL
CL
VLG
dcw PrC 32 1
str Po

4 AuV 6
fr PF VPM VPL
fi
PVG
chp opt LV
VPPC SubG Ect
A11
3 SPFPC 7
ml SPF
ic ZID
rf scp 3V PR
ZIV PRh
F
sod
2 PH STh Or LEnt 8
ns SMT
mfb Gem
La
mt
SuML PLH DEn
cp SuMM GrDG
1 3 2 1 9
SuML BLP
f
a MM AHiPM AHiPL Pir
MnM
ML LM
APir
VTM PMCo PLCo
MRe
0 ArcLP
10
ArcMP
af
Interaural 4.80 mm Bregma -4.20 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 69

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 69
5 5
9 V2MM
1
RSD V2ML
10 PtPD
0 Interaural
+15 +10 +5 0 -5 alv PtPR
8 cg RSGc 2
Or S1BF
Rad
RSGb Py
LMol
IG CA1 MoDG
7 S1 3
cc

dhc PoDG AuD

hif
CA2
FC
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 GrDG CA3 4
chp IMA
bsc LPLR SLu
APTD LPMR Au1
sm hbc DLG
D3V LHb IGL
5 Or 5
dcw
MHb
PrC CL VLG
3 2 1 AuV
alv str Po
chp
4 VPM LV 6
PF VPL
PVG Ect
fr SubG
opt VPPC
rf SPFPC
3 A11 PRh 7
ml VLi ZID
RI ZIV
scp 3V PR LEnt
sod F

ns PH CA3
2 mtg 8
mfb SNR DEn
Gem
sumd Or
PLH 32 1
GrDG Pir
pm SuMM BLP
cp SuML CA3
1 AHiPL
9
MnM APir
f AHiPM
a MM VTM
ML LM PLCo
PMCo
MRe VTM
0 af 10
ArcMP

Interaural 4.64 mm Bregma -4.36 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex AHiAL AHi anterolat ArcMP arcuate med post CA2 field CA2 hipp D3V dorsal 3rd vent F nu fields of Forel hif hipp fissure LM lat mammillary nu
2 layer 2 cortex AHiPL AHi posterlat Au1 primary aud cx CA3 field CA3 hipp dcw deep cerebr white f fornix ic internal capsule LMol lacunosum molec
3 layer 3 cortex AHiPM AHi posteromed AuD 2ary aud cx, dors cc corpus callosum DEn dorsal endopirif FC fasciola cinereum IG indusium griseum LPLR LP laterorostral
3V 3rd ventricle alv alveus of hipp AuV 2ary aud cx, vent cg cingulum dhc dorsal hipp comm fr fasc retroflexus IGL intergenic leaf LPMC LP mediocaudal
a artery APir amygdalopir trans BLP basolat amyg post chp choroid plexus DLG dors lat geniculate Gem gemini hy nu IMA intramedul th area LPMR LP mediorostral
A11 A11 dopamine APTD ant pretectal dors bsc brachium sup coll CL centrolat th nu Ect ectorhinal cx GrDG granular dent gy LEnt lat entorhinal cx LV lat ventricle
af amygdal fissure APTV ant pretectal vent CA1 field CA1 hipp cp cerebral peduncle Eth ethmoid th nu hbc habenular comm LHb lat habenular nu
RBSC5 Book.qxp 16/9/04 16:36 Page 70

mfb med forebr bundle MRe mamm recess 3V Pir piriform cx PrC precommiss nu RSD retrosple dysgran sm stria medullaris V1 primary visual ZIC zona incerta caud
MHb med habenular nu mtg mammilloteg tr PLCo postlat cort amyg PRh perirhinal cx RSGb RSG, b region SNR subst nigra retic V2ML V2 cx, mediolat ZID zona incerta does
ML med mammill lat ns nigrostriat bundle PLH peduncular lat hy PtPD parietal postdors RSGc RSG, c region sod supraoptic decussn V2MM V2 cx, mediomed ZIV zona incerta vent
ml medial lemniscus OPT olivary pretect nu PLi post limitans th nu PtPR parietal postrost S1 1ary somatosens SPFPC SPF parvicell VLG vent lat genic nu
MM med mammill med opt optic tract pm principal mamm tr PVG periventric gray S1BF S1 cx, barrel field str sup thal radiation VLi ventral linear nu
MnM med mamm median Or oriens layer hipp PMCo postmed cort am Py pyramidal cells Sc scaphoid th nu SubG subgeniculate nu VPL vent posterolat nu
MoDG molecular dent gy PF parafascic th nu Po post thalamic nu Rad radiatum layer SCO subcomm organ sumd supramamm decusn VPM vent posteromed
mp mammillary ped PH post hy nu PoDG polymorph dent gy rf rhinal fissure scp sup cerebellar ped SuML supramamm lat VPPC vent post parvic
MPT med pretectal nu PHA post hy area PR prerubral field RI rost interst mlf SLu stratum lucidum SuMM supramamm med VTM vent tuberomamm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 70
5 5
9 1
RSD V2MM
V2ML
0 Interaural 10 V1
+15 +10 +5 0 -5 PtPD
8 alv cg RSGc 2
PtPR
Or
Py
Rad
RSGb
LMol S1
7 IG 3
cc CA1 MoDG
hif
dhc
PoDG
FC
AuD
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 bsc CA3 CA2 4

GrDG
chp
LPMC
D3V IMA
OPT hbc APTD LPLR
MHb MPT LPMR DLG
PLi SLu Au1
5 dcw 5
SCO 3V IGL
str PrC APTV

Sc Po VPL VLG
alv 3 2 1 AuV
4 PVG Or 6
PF VPM
chp alv
A11 Eth
Ect
fr LV
opt SubG
SPFPC
3 VLi
7
rf ml ZIC PRh
ic
scp RI PR
sod F CA3
PHA LEnt
2 ns 8
mtg SLu
mfb PLH
sumd SNR
Rad
Gem DEn
SuMM
SuML GrDG 3 2 1
cp mp
1 MnM BLP Pir 9
f
a AHiPM AHiPL APir
MM ML
LM
PLCo
PMCo
0 VTM 10
MRe
af

Interaural 4.56 mm Bregma -4.44 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 71

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 71
5 5
9 V2MM
1
RSD
V2ML
0 Interaural 10
V1
+15 +10 +5 0 -5 cg
8 RSGc PtPD 2
alv
PtPR
Or
RSGb CA1
Rad Py
IG LMol S1
7 cc 3
hif MoDG

alv dhc
PoDG AuD
dcw FC
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 bsc GrDG 4
chp CA3
CA2
D3V LPMC
OPT IMA Au1
hbc APTD LPLC
pc MPT LPMR SLu
5 DLG 5
SCO PLi
PrC CA3
3V IGL
LPLR AuV
MCPC APTV LV
PVG VLG Or TeA
4 opt str Sc Po 6
3 2 1
Lth VPM
Eth Ect
CA3

SPFPC SubG SLu

3 VLi 7
fr PRh
rf ic
ml PR ZIC CA2
sod LT LEnt
scp PHA RI
F
SNL
2 ns mtg Rad 8
SNCD
sumd PLH Py
PBP SNR 3 2 1
CA1 Or
SuMM Pir
SuML BLP
1 mp 9
a cp
f MM APir
ML LM VTM GrDG AHiPM
VS
PLCo
PMCo
0 10
af

Interaural 4.44 mm Bregma -4.56 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex APTD ant pretectal dors CA3 field CA3 hipp DLG dors lat geniculate hif hipp fissure LPLR LP laterorostral
2 layer 2 cortex APTV ant pretectal vent cc corpus callosum Ect ectorhinal cx ic internal capsule LPMC LP mediocaudal
3 layer 3 cortex Au1 primary aud cx cg cingulum Eth ethmoid th nu IG indusium griseum LPMR LP mediorostral
3V 3rd ventricle AuD 2ary aud cx, dors chp choroid plexus F nu fields of Forel IGL intergenic leaf LT lat terminal nu
a artery AuV 2ary aud cx, vent cp cerebral peduncle f fornix IMA intramedul th area Lth lithoid nu
af amygdal fissure BLP basolat amyg post D3V dorsal 3rd vent FC fasciola cinereum LEnt lat entorhinal cx LV lat ventricle
AHiPM AHi posteromed bsc brachium sup coll dcw deep cerebr white fr fasc retroflexus LM lat mammillary nu
alv alveus of hipp CA1 field CA1 hipp dhc dorsal hipp comm GrDG granular dent gy LMol lacunosum molec
APir amygdalopir trans CA2 field CA2 hipp Dk nu Darkschewitsch hbc habenular comm LPLC LP laterocaudal
RBSC5 Book.qxp 16/9/04 16:36 Page 72

MCPC magnocell nu pc OPT olivary pretect nu PLH peduncular lat hy PtPR parietal postrost RSGb RSG, b region SNR subst nigra retic V1 primary visual VTM vent tuberomamm
ML med mammill lat opt optic tract PLi post limitans th nu PVG periventric gray RSGc RSG, c region sod supraoptic decussn V2ML V2 cx, mediolat ZIC zona incerta caud
ml medial lemniscus Or oriens layer hipp PMCo postmed cort am Py pyramidal cells S1 1ary somatosens SPFPC SPF parvicell V2MM V2 cx, mediomed
MM med mammill med OT nu of optic tract Po post thalamic nu Rad radiatum layer Sc scaphoid th nu str sup thal radiation VLG vent lat genic nu
MoDG molecular dent gy PBP parabrach pigment PoDG polymorph dent gy REth retroethmoid nu SCO subcomm organ SubG subgeniculate nu VLG1 VLG layer 1
mp mammillary ped pc posterior comm PR prerubral field rf rhinal fissure scp sup cerebellar ped sumd supramamm decusn VLi ventral linear nu
MPT med pretectal nu PHA post hy area PrC precommiss nu RI rost interst mlf SLu stratum lucidum SuML supramamm lat VPM vent posteromed
mtg mammilloteg tr Pir piriform cx PRh perirhinal cx RPF retroparafasc nu SNCD SN compact dors SuMM supramamm med VS ventral subiculum
ns nigrostriat bundle PLCo postlat cort amyg PtPD parietal postdors RSD retrosple dysgran SNL subst nigra lat TeA temporal assocn VTAR rostral VTA

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 72
5 5
9 RSD V2MM 1
V2ML
0 Interaural 10
V1
+15 +10 +5 0 -5
alv cg RSGc
8 PtPD 2
Or
RSGb PtPR
IG Py
Rad
7 LMol 3
hif cc MoDG AuD
dhc
PoDG
FC
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 GrDG 4
chp CA3
CA2
D3V LPMC Au1
OPT OT IMA
hbc MPT APTD LPLC
5 pc PLi DLG 5
MCPC LPMR
IGL SLu
PrC
SCO APTV
3V AuV
VLG1
str LV
4 opt RPF Sc Po 3 2 CA3 6
VPM Or
Lth Eth
TeA
SLu
PVG REth
Dk
SPFPC ZIC Ect
3 CA3 CA2 7
VLi SubG
scp fr PR
rf ml PRh
LT
RI SNL
mtg
2 LEnt 8
ns Rad

CD
sumd VTAR PBP
SNR
SN CA1
Py
SuMM Or
SuML 3 2
1 mp 1 9
cp f BLP Pir
a MM
VTM
LM GrDG VS AHiPM
ML APir
PLCo

0 PMCo 10
af

Interaural 4.32 mm Bregma -4.68 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 73

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 73
5 5
9 RSD V2MM 1
V2ML

0 Interaural 10 V1
cg RSGc
+15 +10 +5 0 -5
alv PtPD
8 2
Or
Py PtPR
RSGb Rad
CA1
LMol
7 cc IG MoDG AuD 3
hif
dhc
PoDG
FC GrDG
chp
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
LPMC Au1
D3V OT IMA
OPT
MPT APTD PLi
PCom LPLC DLG Py
5 5
MCPC CA3
pc AuV
SCO IGL CA2
str Po
3V LV
MGD
APTV VPM TeA
4 RPF 32 1 6
Lth MGV Or SLu
Or

VLG
opt Eth
PIL Ect
PVG REth
MZMG Py
EW SPFPC
3 rf ZIC LT PRh 7
Dk Rad
scp PR VLi
SNL
ml LEnt
fr

2 SNCD 8
mtg
SNR CA1
VTAR PBP
3 2
sumd 1
BLP Pir
cp SuMM
1 mp GrDG 9
a APir
MM VS
hif VTM
LM PLCo
ML AHiPM
PMCo
0 af 10

Interaural 4.20 mm Bregma -4.80 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex APTD ant pretectal dors CA1 field CA1 hipp dhc dorsal hipp comm GrDG granular dent gy LPMC LP mediocaudal
2 layer 2 cortex APTV ant pretectal vent CA2 field CA2 hipp Dk nu Darkschewitsch hif hipp fissure LT lat terminal nu
3 layer 3 cortex Aq aqueduct CA3 field CA3 hipp DLG dors lat geniculate IG indusium griseum Lth lithoid nu
3V 3rd ventricle Au1 primary aud cx cc corpus callosum DS dorsal subiculum IGL intergenic leaf LV lat ventricle
a artery AuD 2ary aud cx, dors cg cingulum Ect ectorhinal cx IMA intramedul th area
af amygdal fissure AuV 2ary aud cx, vent chp choroid plexus Eth ethmoid th nu LEnt lat entorhinal cx
AHiPM AHi posteromed BL basolat amyg nu cp cerebral peduncle EW Edinger-Westphal LM lat mammillary nu
alv alveus of hipp BLP basolat amyg post D3V dorsal 3rd vent FC fasciola cinereum LMol lacunosum molec
APir amygdalopir trans bsc brachium sup coll dcw deep cerebr white fr fasc retroflexus LPLC LP laterocaudal
RBSC5 Book.qxp 16/9/04 16:36 Page 74

MCPC magnocell nu pc MPT med pretectal nu PBP parabrach pigment PoDG polymorph dent gy REth retroethmoid nu SLu stratum lucidum TeA temporal assocn VTM vent tuberomamm
MGD med genic dorsal MT med terminal nu pc posterior comm PoT post th nu triang rf rhinal fissure SNCD SN compact dors V1 primary visual ZIC zona incerta caud
MGM med genic medial mtg mammilloteg tr PCom nu of post comm PR prerubral field RLi rost linear raphe SNL subst nigra lat V2ML V2 cx, mediolat
MGV med genic ventral MZMG marg zn med genic PIL post intralam th nu PRh perirhinal cx RPF retroparafasc nu SNR subst nigra retic V2MM V2 cx, mediomed
ML med mammill lat OPT olivary pretect nu Pir piriform cx PtPD parietal postdors RSD retrosple dysgran SPFPC SPF parvicell VLG vent lat genic nu
ml medial lemniscus opt optic tract PLCo postlat cort amyg PtPR parietal postrost RSGb RSG, b region str sup thal radiation VLi ventral linear nu
MM med mammill med Or oriens layer hipp PLi post limitans th nu PVG periventric gray RSGc RSG, c region SuM supramammill nu VPM vent posteromed
MoDG molecular dent gy OT nu of optic tract PMCo postmed cort am Py pyramidal cells SCO subcomm organ sumd supramamm decusn VS ventral subiculum
mp mammillary ped PAG periaqueduct gray Po post thalamic nu Rad radiatum layer scp sup cerebellar ped SuMM supramamm med VTAR rostral VTA

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 74
5 5
9 1
RSD V2MM
V2ML
0 Interaural 10
V1
+15 +10 +5 0 -5 cg RSGc
8 alv PtPD 2

Or
CA1 PtPR
RSGb Py
Rad
7 LMol 3
cc IG MoDG AuD
hif
dhc DS
PoDG
FC
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 GrDG 4
bsc chp
OT LPMC Au1
D3V OPT IMA
dcw
MPT PLi
APTD LPLC
5 PCom DLG 5
LV
pc Po
AuV
Aq SCO MGD IGL CA2
MCPC APTV Py
PAG Or
MGM Rad
4 Lth RPF PoT 32 TeA 6
opt MGV 1 SLu

VLG
MZMG CA3
PIL Ect
Dk
REth SPFPC
3 rf ZIC LT 7
PRh
VLi SNL MoDG Py
scp PR
ml LEnt
fr
2 RLi CD 8
SN
mtg SNR
hif VTAR PBP 3 2 1
CA1

Pir
cp BL
mp SuM
1 a MT 9
GrDG VS
VTM APir
ML
PLCo
AHiPM
PMCo
0 af 10

Interaural 4.08 mm Bregma -4.92 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 75

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 75
5 5
9 1
RSD V2MM
V2ML
0 Interaural 10
V1
+15 +10 +5 0 -5 RSGc
8 cg 2
PtPD
Or
RSGb Py PtPR
Rad
7 3
alv IG LMol CA1
scc AuD
hif dhc MoDG
DS
FC PoDG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 CA1 GrDG 4
chp OT
PPT Au1
bsc D3V OPT LPMC
MPT LV
IMA
APTD LPLC
5 5
pc SCO PCom DLG
dcw
Po
Aq PLi CA2 AuV
PAG MGD IGL
4 MCPC Or 6
APTV MGM Py
Lth SLu Rad TeA
opt MGV 3
PoT 21
MZMG

VLG
Dk PIL Ect
InC InCSh C
3 REth SPFP PP 7
rf LT CA3
ZIC PRh
scp PR
ml SNL MoDG
mtg MA3 LMol CA1
2 RLi SNCD 8
fr
Rad LEnt
hif SNR
IF PBP
VTAR
mp
IPF BL
1 PoDG 9
cp
GrDG VS APir
ML MT AHiPM

PMCo
0 af 10

Interaural 3.96 mm Bregma -5.04 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex
2 layer 2 cortex
3 layer 3 cortex
a artery
af amygdal fissure
AHiPM AHi posteromed
alv alveus of hipp
APir amygdalopir trans
APTD ant pretectal dors
APTV ant pretectal vent CA3 field CA3 hipp DLG dors lat geniculate IG indusium griseum LPLC LP laterocaudal
Aq aqueduct cg cingulum DS dorsal subiculum IGL intergenic leaf LPMC LP mediocaudal
Au1 primary aud cx chp choroid plexus Ect ectorhinal cx IMA intramedul th area LT lat terminal nu
AuD 2ary aud cx, dors cp cerebral peduncle EW Edinger-Westphal InC interstit nu Cajal Lth lithoid nu
AuV 2ary aud cx, vent csc comm of sup coll FC fasciola cinereum InCSh interstit nu shell LV lat ventricle
BL basolat amyg nu D3V dorsal 3rd vent fr fasc retroflexus InWh intermed white SC
bsc brachium sup coll dcw deep cerebr white GrDG granular dent gy IPF interpedunc fossa
CA1 field CA1 hipp dhc dorsal hipp comm hif hipp fissure LEnt lat entorhinal cx
CA2 field CA2 hipp Dk nu Darkschewitsch IF interfascicular nu LMol lacunosum molec
RBSC5 Book.qxp 16/9/04 16:36 Page 76

MA3 medial acc oculom mp mammillary ped OT nu of optic tract PMCo postmed cort am PRh perirhinal cx RPC red nu parvicell SLu stratum lucidum V2MM V2 cx, mediomed
MCPC magnocell nu pc MPT med pretectal nu PaC paracommissural PN paranigral nu PtPD parietal postdors RSD retrosple dysgran SNCD SN compact dors VLG vent lat genic nu
MGD med genic dorsal MT med terminal nu PAG periaqueduct gray Po post thalamic nu PtPR parietal postrost RSGb RSG, b region SNL subst nigra lat VS ventral subiculum
MGM med genic medial mtg mammilloteg tr PBP parabrach pigment PoDG polymorph dent gy Py pyramidal cells RSGc RSG, c region SNR subst nigra retic VTAR rostral VTA
MGV med genic ventral MZMG marg zn med genic pc posterior comm PoT post th nu triang Rad radiatum layer scc splenium corp call SPFPC SPF parvicell ZIC zona incerta caud
ML med mammill lat OPT olivary pretect nu PCom nu of post comm PP peripeduncular nu REth retroethmoid nu SCO subcomm organ TeA temporal assocn
ml medial lemniscus opt optic tract PIL post intralam th nu PPT post pretectal nu rf rhinal fissure scp sup cerebellar ped V1 primary visual
MoDG molecular dent gy Or oriens layer hipp PLi post limitans th nu PR prerubral field RLi rost linear raphe SG supragenic th nu V2ML V2 cx, mediolat

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 76
5 5
9 1
RSD V2MM V2ML
0 Interaural 10
V1
+15 +10 +5 0 -5
8 RSGc 2
cg PtPD
alv
Or

RSGb Py PtPR
Rad
7 3
LMol
scc IG
CA1 AuD
hif dhc MoDG
CA1 DS
PoDG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 chp GrDG
4
bsc LV
D3V OT
MPT PPT Au1
dcw
OPT LPMC
PaC IMA
csc InWh
5 5
APTD LPLC
pc DLG
PCom LMol
opt Po SLu
PAG PLi SG Py CA2
Aq AuV
MCPC MGD Rad
4 Lth 6
MGM
Or TeA

VLG
APTV MGV
PoT CA3
EW Dk MZMG
InC InCSh Ect
SPFPC PIL MoDG Rad
3 7
PP LT
MA3 REth ZIC PRh
rf
scp RPC
SNL
ml
mtg RLi
2 8
SNCD Py
LEnt
fr IF PBP SNR
SNCD
hif PN
IPF
1 mp MT 9
PoDG
a cp
GrDG APir
ML VS
AHiPM

0 PMCo 10
af

Interaural 3.84 mm Bregma -5.16 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 77

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 77
5 5
9 RSD V2MM V2ML 1
V1
0 Interaural 10

+15 +10 +5 0 -5 cg RSGc

8 PtPD 2

PtPR
alv RSGb
dhc
7 hif CA1 Or 3
fmj FC AuD
DS MoDG

a SuG PoDG
GrDG
Zo
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 Op 4
bsc InG OT Py
Au1
MPT PPT OPT LPMC
dcw DMPAG PaC IMA Rad
InWh LPLC
csc LV
5 DLG 5
PCom Po SG
APTD AuV
MCPC CA2
pc PlPAG PLi MGD CA3
Aq
LPAG Rad
opt MGM Or TeA
4 LMol 6
PoT MGV SLu

VLG
APTV MZMG Ect
Dk
InCSh PIL
InC PP LT
3 rf REth PRh 7
EW
SNL MoDG CA1
ml scp MA3 RPC
hif RLi

mtg CD
2 SN LEnt 8
IF PBP SNR
a fr
MT PoDG
PN
IPF GrDG
mp SNCD
1 a VS APir 9
cp
AHiPM
ML
PMCo
af
0 10

Interaural 3.72 mm Bregma -5.28 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

a artery
af amygdal fissure
AHiPM AHi posteromed
alv alveus of hipp
APir amygdalopir trans
APTD ant pretectal dors
APTV ant pretectal vent
Aq aqueduct
Au1 primary aud cx
AuD 2ary aud cx, dors
AuV 2ary aud cx, vent
bsc brachium sup coll
CA1 field CA1 hipp
CA2 field CA2 hipp
CA3 field CA3 hipp
cg cingulum
cp cerebral peduncle DS dorsal subiculum hif hipp fissure LEnt lat entorhinal cx
csc comm of sup coll Ect ectorhinal cx IF interfascicular nu LMol lacunosum molec
dcw deep cerebr white EpP epipeduncular nu IMA intramedul th area LPAG lat periaqu gray
dhc dorsal hipp comm EW Edinger-Westphal InC interstit nu Cajal LPLC LP laterocaudal
Dk nu Darkschewitsch FC fasciola cinereum InCSh interstit nu shell LPMC LP mediocaudal
DLG dors lat geniculate fmj forceps major InG intermed gray SC LT lat terminal nu
DMPAG dorsomedial PAG fr fasc retroflexus InWh intermed white SC LV lat ventricle
DpG deep gray SC GrDG granular dent gy IPF interpedunc fossa
RBSC5 Book.qxp 16/9/04 16:36 Page 78

MA3 medial acc oculom MoDG molecular dent gy opt optic tract PlPAG plioglial PAG PRh perirhinal cx RLi rost linear raphe SLu stratum lucidum V2MM V2 cx, mediomed
MCPC magnocell nu pc mp mammillary ped Or oriens layer hipp PMCo postmed cort am PtPD parietal postdors RMC red nu magnocell SNCD SN compact dors VLG vent lat genic nu
MGD med genic dorsal MPT med pretectal nu OT nu of optic tract PN paranigral nu PtPR parietal postrost RPC red nu parvicell SNL subst nigra lat VS ventral subiculum
MGM med genic medial MT med terminal nu PaC paracommissural Po post thalamic nu Py pyramidal cells RSD retrosple dysgran SNR subst nigra retic vtgd vent teg decussn
MGV med genic ventral mtg mammilloteg tr PBP parabrach pigment PoDG polymorph dent gy Rad radiatum layer RSGb RSG, b region SuG superfic gray SC Zo zonal layer SC
ML med mammill lat MZMG marg zn med genic pc posterior comm PoT post th nu triang REth retroethmoid nu RSGc RSG, c region TeA temporal assocn
ml medial lemniscus Op optic n layer SC PIL post intralam th nu PP peripeduncular nu rf rhinal fissure scp sup cerebellar ped V1 primary visual
mlf med long fasc OPT olivary pretect nu PLi post limitans th nu PPT post pretectal nu RL retrolemniscal nu SG supragenic th nu V2ML V2 cx, mediolat

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 78
5 5
9 1
V2MM V2ML
RSD
0 Interaural 10 V1
+15 +10 +5 0 -5
cg
8 2
fmj RSGc
PtPD

dhc
7 alv PtPR 3
RSGb LMol
DS
dcw MoDG AuD

PoDG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 Zo SuG 4
Op GrDG
bsc OT
InG IMA Py
LV Au1
OPT
InWh LPMC
csc PaC
PPT LPLC CA1
5 Py
5
DLG Or
PlPAG DpG Po SG
DMPAG Rad
PCom APTD MGD AuV
opt pc
CA3
Aq PLi LMol
LPAG MGM TeA
4 MCPC 6
hif MGV Rad
EW APTV
Dk PoT MZMG Ect
InCSh SLu
InC LT
mlf PIL PP
3 rf PRh 7
MA3 RPC EpP
scp SNL
ml GrDG
mtg RLi RMC SNCD
2 vtgd 8
G LEnt
PN PBP MT
SNR MoD
IF
PoDG
fr SNCD VS
mp IPF
1 a cp 9
APir
a MT ML AHiPM

PMCo
af
0 10

Interaural 3.60 mm Bregma -5.40 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 79

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 79
5 5
9 V2MM V2ML
1
RSD
V1
0 Interaural 10

+15 +10 +5 0 -5 cg RSGc V2L

8 2
fmj
PtPD
dhc
PtPR
RSGb
7 3
DS LMol AuD
RSGa
Zo MoDG Or
dcw
a SuG
GrDG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 Op PoDG 4
Py
alv OT Au1
bsc PaC InG LV
PPT
csc InWh LPMC Rad
5 opt PlPAG Rad 5
DMPAG DpWh Po CA1 AuV
SG MGD
DpG APTD
pc
Aq LPAG PLi MGM Or
SLu TeA
4 6
MGV Py
MA3
APTV PoT MZMG CA3 MoDG Ect
EW Dk

mlf InC InCSh PIL PP

rf PaR PRh
3 EpP 7
SNL
RPC
mtg RLi
a ml SNCD
scp RMC LMol LEnt
MT
2 vtgd PoDG 8
a PBP GrDG
SNR
hif IF SNCD
PN PIF
IP VS
cp mp fr
1 APir 9
3n IPF
AHiPM
PMCo
af
0 10

Interaural 3.48 mm Bregma -5.52 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

3n oculomotor nerve APTV ant pretectal vent cg cingulum DpWh deep white SC hif hipp fissure IPF interpedunc fossa
a artery Aq aqueduct cp cerebral peduncle DS dorsal subiculum IF interfascicular nu IPL interpedunc lat
af amygdal fissure Au1 primary aud cx csc comm of sup coll Ect ectorhinal cx InC interstit nu Cajal IPR interpedunc rost
AHiPM AHi posteromed AuD 2ary aud cx, dors dcw deep cerebr white EpP epipeduncular nu InCSh interstit nu shell LEnt lat entorhinal cx
alv alveus of hipp AuV 2ary aud cx, vent dhc dorsal hipp comm EW Edinger-Westphal InG intermed gray SC LMol lacunosum molec
APir amygdalopir trans bsc brachium sup coll Dk nu Darkschewitsch fmj forceps major InWh intermed white SC LPAG lat periaqu gray
APT ant pretectal nu CA1 field CA1 hipp DMPAG dorsomedial PAG fr fasc retroflexus IP interpeduncular nu LPMC LP mediocaudal
APTD ant pretectal dors CA3 field CA3 hipp DpG deep gray SC GrDG granular dent gy IPC interpedunc caudal LV lat ventricle
RBSC5 Book.qxp 16/9/04 16:36 Page 80

MA3 medial acc oculom MT med terminal nu PaR pararubral nu PN paranigral nu PtPR parietal postrost RSGa RSG, a region SNR subst nigra retic vtgd vent teg decussn
MGD med genic dorsal mtg mammilloteg tr PBP parabrach pigment Po post thalamic nu Py pyramidal cells RSGb RSG, b region SuG superfic gray SC Zo zonal layer SC
MGM med genic medial MZMG marg zn med genic pc posterior comm PoDG polymorph dent gy Rad radiatum layer RSGc RSG, c region TeA temporal assocn
MGV med genic ventral Op optic n layer SC PIF parainterfascic nu PoT post th nu triang rf rhinal fissure scp sup cerebellar ped V1 primary visual
ml medial lemniscus opt optic tract PIL post intralam th nu PP peripeduncular nu RLi rost linear raphe SG supragenic th nu V2L V2, lateral
mlf med long fasc Or oriens layer hipp PLi post limitans th nu PPT post pretectal nu RMC red nu magnocell SLu stratum lucidum V2ML V2 cx, mediolat
MoDG molecular dent gy OT nu of optic tract PlPAG plioglial PAG PRh perirhinal cx RPC red nu parvicell SNCD SN compact dors V2MM V2 cx, mediomed
mp mammillary ped PaC paracommissural PMCo postmed cort am PtPD parietal postdors RSD retrosple dysgran SNL subst nigra lat VS ventral subiculum

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 80
5 5
9 V2MM
1
RSD V2ML
0 Interaural 10 V1
+15 +10 +5 0 -5
8 RSGc 2
V2L
fmj
cg PtPD
dhc
alv
7 RSGb PtPR 3
DS
RSGa LMol Or
Zo AuD
SuG GrDG MoDG
alv Py
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 Op 4
PoDG
Rad
dcw bsc InG
PaC LV Au1
OT
csc
InWh CA1
5 DMPAG Po 5
LPMC
PlPAG DpWh DpG SG MGD SLu AuV
LPAG PLi Py
Aq MGM
4 Rad TeA 6
Or
APT MGV
EW Dk Ect
PoT MZMG
InCSh
rf mlf InC
PIL PP PRh
3 PaR 7
CA3
EpP
RPC SNL
mtg LMol
RLi RMC SNCD MoDG
ml scp
PoDG LEnt
2 vtgd
PBP MoDG 8
SNR
hif IF PIF
PN SNCD
IPR IPL GrDG
cp mp VS
1 fr IPC 9
APir
3n IPF
AHiPM
PMCo
af
0 10

Interaural 3.36 mm Bregma -5.64 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 81

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 81
5 5
9 1
RSD V2MM V2ML
0 Interaural 10 V1
+15 +10 +5 0 -5 RSGc
8 cg 2
fmj V2L
dhc
RSGb
7 alv LV PtPC 3
RSGa DS
Zo LMol AuD
SuG MoDG
a
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 Op 4
PoDG
dcw InG Py Au1
GrDG
Py
bsc csc OT
alv InWh Rad
5 DMPAG Or 5
LPMC
DpG Po SG AuV
DpWh MGD
SLu CA1
Aq LPAG Rad
PLi MGM TeA
4 APT MGV 6
EW Ect
Dk DpMe PoT MZMG
MA3 InCSh Or
rf InC PIL PP PRh
mlf
3 a PaR 7
SNL CA3
scp mtg RLi RPC
RMC SNCD PoDG LEnt
ml PBP
a
2 vtgd PBP SNR 8
hif SNCD
IF
PN PIF SNCV
VS
IPR SNCM
cp IPL APir
1 mp IPC 9
3n a AHiPM

0 10

Interaural 3.24 mm Bregma -5.76 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

3n oculomotor nerve AuD 2ary aud cx, dors csc comm of sup coll DpWh deep white SC IF interfascicular nu LEnt lat entorhinal cx
a artery AuV 2ary aud cx, vent dcw deep cerebr white DS dorsal subiculum InC interstit nu Cajal LMol lacunosum molec
AHiPM AHi posteromed bic brachium inf coll dhc dorsal hipp comm Ect ectorhinal cx InCSh interstit nu shell LPAG lat periaqu gray
alv alveus of hipp bsc brachium sup coll Dk nu Darkschewitsch ELm epilemniscal nu InG intermed gray SC LPMC LP mediocaudal
APir amygdalopir trans CA1 field CA1 hipp DLPAG dorsolat PAG EW Edinger-Westphal InWh intermed white SC LV lat ventricle
APT ant pretectal nu CA3 field CA3 hipp DMPAG dorsomedial PAG fmj forceps major IPC interpedunc caudal
Aq aqueduct cg cingulum DpG deep gray SC GrDG granular dent gy IPL interpedunc lat
Au1 primary aud cx cp cerebral peduncle DpMe deep mesenceph hif hipp fissure IPR interpedunc rost
RBSC5 Book.qxp 16/9/04 16:36 Page 82

MA3 medial acc oculom MoDG molecular dent gy PaR pararubral nu PoDG polymorph dent gy rf rhinal fissure RSGc RSG, c region SNL subst nigra lat V2L V2, lateral
MEntV med entorh vent mp mammillary ped PBP parabrach pigment PoT post th nu triang RLi rost linear raphe scp sup cerebellar ped SNR subst nigra retic V2ML V2 cx, mediolat
MGD med genic dorsal mtg mammilloteg tr PIF parainterfascic nu PP peripeduncular nu RMC red nu magnocell SG supragenic th nu SuG superfic gray SC V2MM V2 cx, mediomed
MGM med genic medial MZMG marg zn med genic PIL post intralam th nu PRh perirhinal cx RPC red nu parvicell SLu stratum lucidum TeA temporal assocn VS ventral subiculum
MGV med genic ventral Op optic n layer SC PLi post limitans th nu PtPC parietal postcaud RSD retrosple dysgran SNCD SN compact dors V1 primary visual vtgd vent teg decussn
ml medial lemniscus Or oriens layer hipp PN paranigral nu Py pyramidal cells RSGa RSG, a region SNCM SN compact med V1B V1, binocular Zo zonal layer SC
mlf med long fasc OT nu of optic tract Po post thalamic nu Rad radiatum layer RSGb RSG, b region SNCV SN compact ventr V1M V1, monocular

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 82
5 5
9 RSD V2MM 1
V2ML
V1M
0 Interaural 10

+15 +10 +5 0 -5 cg RSGc V1B

8 fmj 2
LV
V2L
dhc
RSGb
alv
7 RSGa PtPC 3
VS
Zo
AuD
SuG LMol
Op MoDG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
InG PoDG
Py Au1
alv
InWh
csc Or Rad
dcw bsc
5 DMPAG DpG 5
SG SLu CA1
DLPAG MGD AuV
Rad
DpWh Or
Aq LPAG
bic
TeA
4 MGV 6
EW APT
Dk MGM
Py Ect
DpMe MZMG
MA3 PoT
InCSh PP
mlf InC PIL
3 rf PRh 7
PaR GrDG
mtg SNL
ELm RLi PoDG
RPC PBP
RMC SNCD
2 SNR 8
scp vtgd LEnt
PBP
hif ml
IF PIF SNCV
PN DS
1 IPR 9
cp mp
3n IPL SNCM APir
IPC MEnt

0 10

Interaural 3.12 mm Bregma -5.88 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 83

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 83
5 5
9 V2MM 1
RSD V2ML
V1M
0 Interaural 10
V1B
+15 +10 +5 0 -5 cg RSGc
8 fmj 2
LV
dhc V2L
RSGb
alv
RSGa Post
7 Zo DS PtPC 3

SuG AuD
Op LMol
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 dcw a InG GrDG MoDG Py 4

Or Au1
alv PoDG Rad
Rad
bsc csc InWh
DT
CA1
5 DMPAG DpG 5
DLPAG MGD AuV
Py
SG
Aq LPAG DpWh MZMG
bic TeA
4 CA3 6
MGV
EW Su3 MGM Ect
InCSh PoT Or
InC DpMe
MA3 PIL SubB
mlf PRh
3 rf 7
PaR
mtg RPC SNL
RLi
hif vtgd SNCD
RMC
SNR
2 LEnt 8
tth PBP
ml IF PIF SNCV
VS
PN
SNCM
mp IPR
1 cp MEnt 9
IPL
IPC APir

0 10

Interaural 3.00 mm Bregma -6.00 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

3PC 3 nu, parvicell AuV 2ary aud cx, vent csc comm of sup coll DpWh deep white SC GrDG granular dent gy IPC interpedunc caudal
a artery bic brachium inf coll dcw deep cerebr white DS dorsal subiculum hif hipp fissure IPDM interped dorsomed
alv alveus of hipp bsc brachium sup coll dhc dorsal hipp comm DT dors terminal nu IF interfascicular nu IPL interpedunc lat
APir amygdalopir trans CA1 field CA1 hipp DLPAG dorsolat PAG dtgd dorsal teg decussn InC interstit nu Cajal IPR interpedunc rost
Aq aqueduct CA3 field CA3 hipp DMPAG dorsomedial PAG Ect ectorhinal cx InCSh interstit nu shell LEnt lat entorhinal cx
Au1 primary aud cx cg cingulum DpG deep gray SC EW Edinger-Westphal InG intermed gray SC LMol lacunosum molec
AuD 2ary aud cx, dors cp cerebral peduncle DpMe deep mesenceph fmj forceps major InWh intermed white SC LPAG lat periaqu gray
LV lat ventricle
RBSC5 Book.qxp 16/9/04 16:36 Page 84

MA3 medial acc oculom
MEntV med entorh vent
MGD med genic dorsal
MGM med genic medial
MGV med genic ventral
ml medial lemniscus
mlf med long fasc
MoDG molecular dent gy
mp mammillary ped
mtg mammilloteg tr
MZMG marg zn med genic
Op optic n layer SC
Or oriens layer hipp
PaR pararubral nu
PBP parabrach pigment
PIF parainterfascic nu
PIL post intralam th nu
PN paranigral nu
PoDG polymorph dent gy RMC red nu magnocell SNCV SN compact ventr V1B V1, binocular
Post postsubiculum RPC red nu parvicell SNL subst nigra lat V1M V1, monocular
PoT post th nu triang RSD retrosple dysgran SNR subst nigra retic V2L V2, lateral
PRh perirhinal cx RSGa RSG, a region Su3 supra3 periaq gray V2ML V2 cx, mediolat
PtPC parietal postcaud RSGb RSG, b region Su3C supra3 cap V2MM V2 cx, mediomed
Py pyramidal cells RSGc RSG, c region SubB subbrachial nu VS ventral subiculum
Rad radiatum layer SG supragenic th nu SuG superfic gray SC vtgd vent teg decussn
rf rhinal fissure SNCD SN compact dors TeA temporal assocn Zo zonal layer SC
RLi rost linear raphe SNCM SN compact med tth trigeminothal tr

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 84
5 5
9 1
RSD V2MM V2ML

0 Interaural 10 V1M

+15 +10 +5 0 -5
RSGc V1B
8 2
fmj

dhc V2L
RSGb LV
alv
7 Zo RSGa 3
Post Or
DS PtPC
SuG
AuD
Op
LMol
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
InG
Py
dcw a Au1
csc InWh
bsc Rad
5 DMPAG DpG DT 5

Dp
CA3
DLPAG

W
alv AuV

h
MGD
Aq LPAG SG MZMG CA1
bic TeA
4 Su3C MGV 6

EW Su3
a DpMe PoT MGM Or Ect
3PC InCSh
a InC
mlf PIL SubB
3 PoDG PRh 7
rf
dtgd RPC PaR SNL
P
a PB
RMC SNR
RLi GrDG
2 SNCD 8
vtgd
hif PBP LEnt
ml tth IF SNCV MoDG
cp PIF

SNR VS
IPDM
1 IPR IPL SNCM 9
IPC PN MEnt APir

0 10

Interaural 2.88 mm Bregma -6.12 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 85

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 85
5 5 V2MM
9 1
RSD V2ML
V1M
0 Interaural 10
V1B
+15 +10 +5 0 -5 RSGc
8 fmj 2
LV V2L
dhc RSGb
alv RSGa AuD
Zo Or
7 Post DS 3
SuG
Op LMol Au1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

MoDG
6 dcw InG Py 4
csc InWh
Rad AuV
a
DpG
bsc
5 alv DMPAG CA1 5
DT MGD GrDG TeA
DLPAG
MZMG PoDG
Aq SG
LPAG DpWh Ect
Su3C MGV
4 bic 6
EW MGM PRh
Su3 DpMe
rf
3PC InC
3N SubB
ts DLEnt
mlf
3 SNL 7
PaR SNCD
a RPC
dtgd SNR
RMC
hif vtgd DIEnt
SNCV VS
2 CLi PBP 8
IPDM IF PIF
PN SNCM
ml tth VIEnt
cp IPR MEnt
1 IPDL IPL APir 9
IPC
pcer
bas

0 10

Interaural 2.76 mm Bregma -6.24 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

3N oculomotor nu AuV 2ary aud cx, vent csc comm of sup coll DpMe deep mesenceph GrDG granular dent gy IPDM interped dorsomed
3PC 3 nu, parvicell bas basilar artery dcw deep cerebr white DpWh deep white SC hif hipp fissure IPL interpedunc lat
a artery bic brachium inf coll dhc dorsal hipp comm DS dorsal subiculum IF interfascicular nu IPR interpedunc rost
alv alveus of hipp bsc brachium sup coll DIEnt dors int entorh cx DT dors terminal nu InC interstit nu Cajal LMol lacunosum molec
APir amygdalopir trans CA1 field CA1 hipp DLEnt dorsolat entorh cx dtgd dorsal teg decussn InG intermed gray SC LPAG lat periaqu gray
Aq aqueduct cg cingulum DLPAG dorsolat PAG Ect ectorhinal cx InWh intermed white SC LV lat ventricle
Au1 primary aud cx CLi caud linear raphe DMPAG dorsomedial PAG EW Edinger-Westphal IPC interpedunc caudal
AuD 2ary aud cx, dors cp cerebral peduncle DpG deep gray SC fmj forceps major IPDL interped dorsolat
RBSC5 Book.qxp 16/9/04 16:36 Page 86

Me5 mesenceph 5 nu MZMG marg zn med genic Post postsubiculum RSD retrosple dysgran SNL subst nigra lat V1B V1, binocular
me5 mesenceph 5 tr Op optic n layer SC PRh perirhinal cx RSGa RSG, a region SNR subst nigra retic V1M V1, monocular
MEnt medial entorhinal Or oriens layer hipp Py pyramidal cells RSGb RSG, b region Su3 supra3 periaq gray V2L V2, lateral
MGD med genic dorsal PaR pararubral nu Rad radiatum layer RSGc RSG, c region Su3C supra3 cap V2ML V2 cx, mediolat
MGM med genic medial PBP parabrach pigment rf rhinal fissure scpd scp descend limb SubB subbrachial nu V2MM V2 cx, mediomed
MGV med genic ventral pcer post cerebral a RMC red nu magnocell SG supragenic th nu SuG superfic gray SC VIEnt vent int entorh
ml medial lemniscus PIF parainterfascic nu RPC red nu parvicell SNCD SN compact dors TeA temporal assocn VS ventral subiculum
mlf med long fasc PN paranigral nu RRF retrorubral field SNCM SN compact med ts tectospinal tract vtgd vent teg decussn
MoDG molecular dent gy PoDG polymorph dent gy rs rubrospinal tract SNCV SN compact ventr tth trigeminothal tr Zo zonal layer SC

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 86
5 5
9 V2MM 1
RSD V2ML
V1M
0 Interaural 10
RSGc
+15 +10 +5 0 -5 V1B
8 fmj 2

RSGb V2L
LV
dhc
Zo RSGa
dcw alv
7 AuD 3
Post
SuG DS
Or
Op Au1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 InG 4
bsc LMol
Py
csc InWh AuV

DMPAG DT Rad
5 DpG 5
DLPAG
alv
DpWh MGD TeA
Aq LPAG PoDG CA1
me5 Me5 MZMG
Su3C
bic MGV Ect
4 MGM 6
EW Su3
ts 3PC InC DpMe PRh
3N SubB
rf GrDG
3 mlf 7
RPC PaR SNL
dtgd MoDG DLEnt
PBP
RMC SNR
PBP SNCD
2 vtgd RRF 8
rs CLi VS DIEnt
scpd IF PIF SNCV
tth PN
IPR VIEnt
cp ml MEnt
1 SNCM 9
IPL APir
IPC
pcer
IPDL bas IPDM

0 10

Interaural 2.64 mm Bregma -6.36 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 87

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 87
5 5
9 V2MM V2ML 1
RSD
V1M
0 Interaural 10

+15 +10 +5 0 -5 RSGc V1B

8 2
fmj
RSGb
V2L
dhc RSGa LV
Zo
7 alv 3
SuG Post AuD
DS
Op
dcw Or
InG Au1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 Py 4
LMol
InWh MoDG
AuV

DpG GrDG Rad

DMPAG
5 DT 5
a DLPAG TeA
CA1
me5 Aq Me5 DpWh MGD
LPAG PoDG
Su3C MZMG
Ect
4 MGV 6
EW Su3 MGM
bic
3PC PRh
DpMe
ts 3N
rf InC SubB
a
3 a mlf 7
PaR DLEnt
dtgd SNCD SNL
RMC
scp SNR PrS
PBP SNCV VS
RRF
2 CLi PaS 8
rs DIEnt
scpd IF PIF
cp tth IPR PN
SNCM MEnt VIEnt
ml
1 IPL APir 9
IPC
IPI
pcer IPDL
IPDM bas

0 10

Interaural 2.52 mm Bregma -6.48 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

3N oculomotor nu
3PC 3 nu, parvicell
a artery
alv alveus of hipp
APir amygdalopir trans
Aq aqueduct
Au1 primary aud cx
AuD 2ary aud cx, dors
AuV 2ary aud cx, vent
bas basilar artery
BIC nu brach inf coll
bic brachium inf coll
CA1 field CA1 hipp
CLi caud linear raphe
cp cerebral peduncle
dcw deep cerebr white
dhc dorsal hipp comm DS dorsal subiculum InC interstit nu Cajal IPR interpedunc rost
DIEnt dors int entorh cx DT dors terminal nu InG intermed gray SC LMol lacunosum molec
DLEnt dorsolat entorh cx dtgd dorsal teg decussn InWh intermed white SC LPAG lat periaqu gray
DLPAG dorsolat PAG Ect ectorhinal cx IPC interpedunc caudal LV lat ventricle
DMPAG dorsomedial PAG EW Edinger-Westphal IPDL interped dorsolat
DpG deep gray SC fmj forceps major IPDM interped dorsomed
DpMe deep mesenceph GrDG granular dent gy IPI interped intermed
DpWh deep white SC IF interfascicular nu IPL interpedunc lat
RBSC5 Book.qxp 16/9/04 16:36 Page 88

m5 motor root 5n mlf med long fasc pcer post cerebral a Rad radiatum layer RSGc RSG, c region SNR subst nigra retic tth trigeminothal tr VS ventral subiculum
Me5 mesenceph 5 nu MoDG molecular dent gy PIF parainterfascic nu rf rhinal fissure s5 sensory root of 5n Su3 supra3 periaq gray V1B V1, binocular Zo zonal layer SC
me5 mesenceph 5 tr MZMG marg zn med genic PN paranigral nu RMC red nu magnocell scp sup cerebellar ped Su3C supra3 cap V1M V1, monocular
MEnt medial entorhinal Op optic n layer SC PoDG polymorph dent gy RRF retrorubral field scpd scp descend limb SubB subbrachial nu V2L V2, lateral
MGD med genic dorsal Or oriens layer hipp Post postsubiculum rs rubrospinal tract SNCD SN compact dors SuG superfic gray SC V2ML V2 cx, mediolat
MGM med genic medial PaR pararubral nu PRh perirhinal cx RSD retrosple dysgran SNCM SN compact med TeA temporal assocn V2MM V2 cx, mediomed
MGV med genic ventral PaS parasubiculum PrS presubiculum RSGa RSG, a region SNCV SN compact ventr tfp trans fibers pons VIEnt vent int entorh
ml medial lemniscus PBP parabrach pigment Py pyramidal cells RSGb RSG, b region SNL subst nigra lat ts tectospinal tract VLPAG ventrolat PAG

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 88
5 5 V2MM
9 1
RSD V2ML

0 Interaural 10 V1M
RSGc
+15 +10 +5 0 -5
V1B
8 fmj 2

RSGb
Zo RSGa V2L
dhc LV
7 3
alv SuG Post
Op DS AuD

InG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 Or Au1 4
dcw
InWh
Rad
DpG AuV
DMPAG
5 DLPAG DpWh
5
MGV CA1
LMol TeA
me5 Aq
LPAG
Me5 GrDG
VLPAG
4 bic Su3C BIC Ect 6
ts EW Su3
3PC DpMe Py
MZMG
3N SubB PRh
InC MoDG
rf
alv
3 mlf 7
dtgd PaR DLEnt
RMC PrS
RRF SNR
CLi PaS VS
SNCD
2 SNCV DIEnt 8
rs SNCM
scpd
cp PIF
tth IPR IPDL MEnt
s5 VIEnt
ml
1 m5 APir 9
IPL
IPI
IPC
tfp

0 bas 10

Interaural 2.40 mm Bregma -6.60 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 89

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 89
5 5
9 V2MM 1
RSD V2ML

0 Interaural 10 V1M
RSGc
+15 +10 +5 0 -5
V1B
8 2
fmj RSGb

Zo RSGa LV V2L
dhc
7 SuG 3
alv
AuD
Op Post DS
InG
Or Au1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 dcw 4
InWh
PrS
AuV
DpG CA1
DMPAG
DLPAG Rad
5 DpWh Py TeA 5
LMol
Me5 Aq LPAG GrDG
me5
bic BIC
VLPAG EW
VLPAG Ect
Su3C MoDG
4 6
alv ts Su3
PRh
3PC InC DpMe SubB
rf 3N
PrS
mlf DLEnt
3 7
dtgd
RRF PaS VS
CLi
PPTg
DIEnt
rs
2 SNR
8
scpd ID
CEnt
cp IPA VTA VIEnt
tth IPDL MEnt
s5 ml
1 m5 IPC IPL 9
IPI
tfp Pn

0 bas 10

Interaural 2.28 mm Bregma -6.72 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

3N oculomotor nu
3PC 3 nu, parvicell
alv alveus of hipp
Aq aqueduct
Au1 primary aud cx
AuD 2ary aud cx, dors
AuV 2ary aud cx, vent
B9 B9 serotonin cells
bas basilar artery
BIC nu brach inf coll
bic brachium inf coll
CA1 field CA1 hipp
CEnt coudomed ent cx
CLi caud linear raphe
cp cerebral peduncle DpG deep gray SC EW Edinger-Westphal IPA interpedunc apical
dcw deep cerebr white DpMe deep mesenceph fmj forceps major IPC interpedunc caudal
dhc dorsal hipp comm DpWh deep white SC GrDG granular dent gy IPDL interped dorsolat
DIEnt dors int entorh cx DR dorsal raphe nu ID intradecussat nu IPI interped intermed
DLEnt dorsolat entorh cx DS dorsal subiculum InC interstit nu Cajal IPL interpedunc lat
DLPAG dorsolat PAG dtgd dorsal teg decussn InG intermed gray SC LMol lacunosum molec
DMPAG dorsomedial PAG Ect ectorhinal cx InWh intermed white SC LPAG lat periaqu gray
LV lat ventricle
RBSC5 Book.qxp 16/9/04 16:36 Page 90

m5 motor root 5n
Me5 mesenceph 5 nu
me5 mesenceph 5 tr
MEnt medial entorhinal
MiTg microcell teg nu
ml medial lemniscus
mlf med long fasc
MoDG molecular dent gy
MoS molec layer subic
Op optic n layer SC
Or oriens layer hipp
PaS parasubiculum
Pn pontine nuclei
Post postsubiculum
PPTg pedunculopont teg
PRh perirhinal cx
PrS presubiculum RSGb RSG, b region SuG superfic gray SC V2ML V2 cx, mediolat
Py pyramidal cells RSGc RSG, c region TeA temporal assocn V2MM V2 cx, mediomed
Rad radiatum layer s5 sensory root of 5n tfp trans fibers pons VIEnt vent int entorh
rf rhinal fissure scpd scp descend limb ts tectospinal tract VS ventral subiculum
RRF retrorubral field SNR subst nigra retic tth trigeminothal tr VTA vent tegment area
rs rubrospinal tract Su3 supra3 periaq gray V1B V1, binocular Zo zonal layer SC
RSD retrosple dysgran Su3C supra3 cap V1M V1, monocular
RSGa RSG, a region SubB subbrachial nu V2L V2, lateral

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 90
5 5
9 1
V2MM
V2ML
RSD
10 V1M
0 Interaural
+15 +10 +5 0 -5 RSGc
V1B
8 2
fmj RSGb

Zo RSGa V2L
LV
7 SuG 3
dhc
alv Op AuD
Post
InG DS
dcw
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 InWh Au1 4

DpG
PrS
AuV
alv DMPAG
5 DLPAG DpWh 5
CA1

Aq TeA
LPAG MoS
me5 Me5
bic VLPAG EW BIC MoDG
Su3C
4 Ect 6
DR Su3
ts 3PC
PRh
rf 3N InC DpMe SubB
PrS
3 mlf MiTg VS 7
DLEnt
RRF PaS
ts CLi

2 PPTg DIEnt 8
rs
ID
scpd
cp VIEnt
IPA VTA B9 CEnt
s5 MEnt
1 tth IPI IPDL 9
ml
m5 IPC
IPL
Pn
0 10
tfp

bas
Interaural 2.16 mm Bregma -6.84 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 91

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 91
5 5
9 1
V2MM V2ML
0 Interaural 10 RSD
V1M
+15 +10 +5 0 -5
8 V1B 2
RSGb
fmj
Zo RSGa

7 LV 3
SuG V2L
dhc Op
alv
InG Post
dcw
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 InWh 4
alv DpG

DMPAG TeA
5 DLPAG
5
DpWh
Aq STr
bic LPAG PrS MoS
Me5 BIC
ECIC Ect
4 me5 VLPAG PrCnF 6
DR
ts Su3C PRh
3PC Su3
rf SubB
3N
DpMe
DLEnt
3 mlf MiTg 7
PaS
RRF PBG
ts
CLi
DIEnt
2 PPTg 8
scpd ID
rs VIEnt
Rbd B9 CEnt
IPA MEnt
a cp
s5 tth IPI
1 9
ml DPPn
m5
IPL
IPC
Pn
0 10
tfp

Interaural 2.04 mm bas

Bregma -6.96 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

3N oculomotor nu B9 B9 serotonin cells dcw deep cerebr white DpMe deep mesenceph fmj forceps major IPL interpedunc lat
3PC 3 nu, parvicell bas basilar artery dhc dorsal hipp comm DPPn dors ped pont nu ID intradecussat nu ll lat lemniscus
4N trochlear nu BIC nu brach inf coll DIEnt dors int entorh cx DpWh deep white SC InG intermed gray SC LPAG lat periaqu gray
4Sh trochlear shell bic brachium inf coll DLEnt dorsolat entorh cx DR dorsal raphe nu InWh intermed white SC LV lat ventricle
a artery CEnt coudomed ent cx DLPAG dorsolat PAG DRD dorsal raphe dors IPA interpedunc apical
alv alveus of hipp CLi caud linear raphe DMPAG dorsomedial PAG ECIC ext cx inf coll IPC interpedunc caudal
Aq aqueduct cp cerebral peduncle DpG deep gray SC Ect ectorhinal cx IPI interped intermed
RBSC5 Book.qxp 16/9/04 16:36 Page 92

m5 motor root 5n MoS molec layer subic PPTg pedunculopont teg RSD retrosple dysgran SubB subbrachial nu V1M V1, monocular
mcp mid cerebellar ped Op optic n layer SC PrCnF precuneiform area RSGa RSG, a region SuG superfic gray SC V2L V2, lateral
Me5 mesenceph 5 nu PaS parasubiculum PRh perirhinal cx RSGb RSG, b region TeA temporal assocn V2ML V2 cx, mediolat
me5 mesenceph 5 tr PBG parabigeminal nu PrS presubiculum s5 sensory root of 5n tfp trans fibers pons V2MM V2 cx, mediomed
MEnt medial entorhinal PiSt pineal stalk Rbd rhabdoid nu scpd scp descend limb ts tectospinal tract VIEnt vent int entorh
MiTg microcell teg nu PMnR paramedian raphe rf rhinal fissure STr subiculum trans tth trigeminothal tr VLPAG ventrolat PAG
ml medial lemniscus Pn pontine nuclei RRF retrorubral field Su3 supra3 periaq gray v vein Zo zonal layer SC
mlf med long fasc Post postsubiculum rs rubrospinal tract Su3C supra3 cap V1B V1, binocular

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 92
5 5
9 1
V2MM V2ML
RSD V1M
0 Interaural 10

+15 +10 +5 0 -5
8 V1B 2
fmj RSGb

v Zo RSGa

PiSt
7 SuG 3
LV V2L
Op
dhc Post
InG
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 InWh 4

a
DpG
dcw TeA
DMPAG
5 DLPAG 5
DpWh
PrS
Aq LPAG BIC STr
bic Me5 ECIC MoS
me5 Ect
4 VLPAG 6
ll DRD PrCnF
Su3C
alv 4Sh Su3 PRh
SubB
rf
4N DpMe PBG
3 mlf MiTg PaS 7
DLEnt
RRF
ts
CLi

2 ID PPTg DIEnt 8
scpd
Rbd
rs VIEnt
PMnR CEnt
B9 MEnt
s5 IPA
1 cp tth 9
m5
ml IPC
IPL DPPn
mcp

0 Pn 10

Interaural 1.92 mm bas

Bregma -7.08 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 93

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 93
5 5
9 V2MM V2ML 1

RSD V1M
0 Interaural 10

+15 +10 +5 0 -5 V1B

8 2
RSGb

v RSGa
Zo
PiSt
SuG V2L
7 3
Op LV
Post
dcw InG
InWh
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
a
DpG
TeA
DMPAG
5 DLPAG DpWh PrS 5
STr
bic Aq LPAG BIC
ECIC
alv Ect
VLPAG
4 ll 6
me5 DRD DRL Me5 PrCnF
PRh
PDR
4Sh DRV
rf PaS
DpMe PBG
4N DLEnt
3 MiTg 7
mlf RRF
ts CLi DIEnt
RR

scpd PPTg
2 VIEnt
8
ts
rs
Rbd
a PnO CEnt MEnt
MnR PMnR
B9
1 s5 cp 9
m5 tth
ml IPC
mcp IPL

0 10
Pn

Interaural 1.80 mm tfp Bregma -7.20 mm

bas
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex 5b layer 5b cortex BIC nu brach inf coll DLEnt dorsolat entorh cx DRL dorsal raphe lat IPC interpedunc caudal
2 layer 2 cortex 6 layer 6 cortex bic brachium inf coll DLPAG dorsolat PAG DRV dorsal raphe vent IPL interpedunc lat
3/4 layers 3&4 cx a artery CEnt coudomed ent cx DMPAG dorsomedial PAG ECIC ext cx inf coll lfp long fasc pons
4N trochlear nu alv alveus of hipp CLi caud linear raphe DpG deep gray SC Ect ectorhinal cx ll lat lemniscus
4n trochlear nerve Aq aqueduct cp cerebral peduncle DpMe deep mesenceph fmj forceps major LPAG lat periaqu gray
4Sh trochlear shell B9 B9 serotonin cells dcw deep cerebr white DpWh deep white SC InG intermed gray SC LV lat ventricle
5a layer 5a cortex bas basilar artery DIEnt dors int entorh cx DRD dorsal raphe dors InWh intermed white SC
RBSC5 Book.qxp 16/9/04 16:36 Page 94

m5 motor root 5n Op optic n layer SC PnO pontine retic oral RRF retrorubral field STr subiculum trans V2L V2, lateral
mcp mid cerebellar ped Pa4 paratrochlear nu Post postsubiculum rs rubrospinal tract SuG superfic gray SC V2ML V2 cx, mediolat
Me5 mesenceph 5 nu PaS parasubiculum PPTg pedunculopont teg RSD retrosple dysgran TeA temporal assocn V2MM V2 cx, mediomed
me5 mesenceph 5 tr PBG parabigeminal nu PrCnF precuneiform area RSG retrosplenial gran tfp trans fibers pons VIEnt vent int entorh
MEnt medial entorhinal PDR postdors raphe PRh perirhinal cx RSGa RSG, a region ts tectospinal tract VLPAG ventrolat PAG
MiTg microcell teg nu Pi ;ineal gland PrS presubiculum RSGb RSG, b region tth trigeminothal tr Zo zonal layer SC
ml medial lemniscus PiSt pineal stalk Rbd rhabdoid nu S subiculum v vein
mlf med long fasc PMnR paramedian raphe rf rhinal fissure s5 sensory root of 5n V1B V1, binocular
MnR median raphe nu Pn pontine nuclei RR retrorubral nu scpd scp descend limb V1M V1, monocular

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 94
5 5
9 1
V2MM V2ML
0 Interaural 10 V1M
RSD
+15 +10 +5 0 -5
8 V1B
2
RSGb 6
fmj 5b
RSGa 5a
Zo 1
Pi 2
3/4
7 SuG 3
V2L
Op
InG LV
dcw Post
InWh
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

DpG
TeA
DMPAG
5 DLPAG 5
DpWh PrS
BIC
bic Aq LPAG ECIC
alv STr
me5 VLPAG Ect
DRD Me5 PrCnF
4 DRL 6
ll PRh
4Sh DRV PDR
PBG PaS
rf
4N MiTg
Pa4 DpMe DLEnt
3 mlf 7
ts CLi DIEnt
4n
scpd PPTg RR
Rbd VIEnt
2 8
a ts
rs CEnt
PnO MEnt
MnR
PMnR B9
s5 m5
1 9
lfp tth
mcp
ml

0 Pn 10

Interaural 1.68 mm tfp

bas
Bregma -7.32 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 95

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0

v
Figure 95
5 5
9 V2ML 1
V2MM
V1M
0 Interaural 10 RSD
+15 +10 +5 0 -5
V1B
8 RSGb 2

Pi RSGa
Zo
V2L
SuG
7 3
Op
Post
dcw InG LV
InWh
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 TeA 4
DpG

DMPAG PrS
DLPAG DpWh
5 BIC 5
bic Aq LPAG STr Ect
Me5 ECIC
alv
DRD VLPAG PrCnF PaS
me5
4 DRL PRh 6
ll
rf 4Sh PDR
DRV PBG
MiTg DLEnt
4N DpMe
Pa4
4n mlf
3 7
ts DIEnt
CLi
RR
scpd PPTg
VIEnt
a Rbd
2 CEnt 8
MEnt
ll rs ts
PnO
m5
MnR PMnR
s5
mcp B9
1 9
lfp
m5
tth
ml

0 10
Pn

Interaural 1.56 mm tfp

bas
Bregma -7.44 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4N trochlear nu BIC nu brach inf coll DpMe deep mesenceph InG intermed gray SC
4n trochlear nerve bic brachium inf coll DpWh deep white SC InWh intermed white SC
4Sh trochlear shell CEnt coudomed ent cx DRD dorsal raphe dors lfp long fasc pons
a artery dcw deep cerebr white DRL dorsal raphe lat ll lat lemniscus
alv alveus of hipp DIEnt dors int entorh cx DRV dorsal raphe vent LPAG lat periaqu gray
Aq aqueduct DLEnt dorsolat entorh cx dscp decussation scp LV lat ventricle
ATg ant tegmental nu DLPAG dorsolat PAG ECIC ext cx inf coll
B9 B9 serotonin cells DMPAG dorsomedial PAG Ect ectorhinal cx
bas basilar artery DpG deep gray SC fmj forceps major
RBSC5 Book.qxp 16/9/04 16:36 Page 96

m5 motor root 5n Op optic n layer SC Post postsubiculum rs rubrospinal tract SuG superfic gray SC V2ML V2 cx, mediolat
mcp mid cerebellar ped Pa4 paratrochlear nu PPTg pedunculopont teg RSD retrosple dysgran TeA temporal assocn V2MM V2 cx, mediomed
Me5 mesenceph 5 nu PaS parasubiculum PrCnF precuneiform area RSGa RSG, a region tfp trans fibers pons VIEnt vent int entorh
me5 mesenceph 5 tr PBG parabigeminal nu PRh perirhinal cx RSGb RSG, b region ts tectospinal tract VLPAG ventrolat PAG
MEnt medial entorhinal PDR postdors raphe PrS presubiculum s5 sensory root of 5n tth trigeminothal tr Zo zonal layer SC
MiTg microcell teg nu Pi ;ineal gland Rbd rhabdoid nu scp sup cerebellar ped v vein
ml medial lemniscus PMnR paramedian raphe rf rhinal fissure scpd scp descend limb V1B V1, binocular
mlf med long fasc Pn pontine nuclei RLi rost linear raphe SPTg subpedunc teg V1M V1, monocular
MnR median raphe nu PnO pontine retic oral RR retrorubral nu STr subiculum trans V2L V2, lateral

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 96
5 5
9 V2ML 1
V2MM
V1M
0 Interaural 10
RSD
+15 +10 +5 0 -5 V1B
8 2
RSGb
fmj
RSGa
Pi
Zo V2L
SuG
7 3
Op
InG
InWh Post
dcw
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 TeA 4
DpG LV
DpWh
DMPAG
bic
DLPAG ECIC
5 BIC 5
STr Ect
alv Aq LPAG
PaS
me5
Me5 PrCnF
4 ll VLPAG PRh 6
rf DRD
DRL
PDR PBG
DRV DpMe MiTg
4n 4n Pa4 DLEnt
3 mlf 7

scp PPTg
dscp RR VIEnt
a ATg
ll SPTg
2 a CEnt 8
Rbd
MEnt
m5 ts PnO
rs
MnR
mcp
s5
1 B9 9
PMnR
lfp
m5 tth
ml

0 10
Pn

Interaural 1.44 mm tfp Bregma -7.56 mm

bas
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 97

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 97
5 5
9 V2ML 1
V2MM V1M
0 Interaural 10

+15 +10 +5 0 -5 RSD

V1B
8 2
RSGa
Pi
Zo
fmj V2L
7 SuG 3
Op
InG
Post
dcw InWh
bic
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 TeA 4
DpG

DpWh LV
DMPAG ECIC
DLPAG BIC
5 5
PaS Ect
Aq LPAG

me5 PrCnF PRh

VLPAG Me5
4 rf ll 6
DRD
DRL PBG
4n MiTg DLEnt
4n DRV PDR DpMe
Pa4
3 mlf 7
PL
PPTg VIEnt
a RR
dscp
a
Rbd CEnt
ATg SPTg
2 VLL MEnt 8
rs PnO
m5 ts
ll MnR
s5 mcp

1 PMnR 9
tth
m5 lfp ml

0 10
Pn

Interaural 1.32 mm tfp Bregma -7.68 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4n trochlear nerve cic comm of inf coll DpWh deep white SC InG intermed gray SC
a artery CnFI cuneiform interm DRD dorsal raphe dors InWh intermed white SC
Aq aqueduct CnFV cuneiform vent DRL dorsal raphe lat lfp long fasc pons
ATg ant tegmental nu dcw deep cerebr white DRV dorsal raphe vent ll lat lemniscus
bas basilar artery DLEnt dorsolat entorh cx dscp decussation scp LPAG lat periaqu gray
BIC nu brach inf coll DLPAG dorsolat PAG ECIC ext cx inf coll LV lat ventricle
bic brachium inf coll DMPAG dorsomedial PAG Ect ectorhinal cx
CEnt coudomed ent cx DpG deep gray SC ERS epirubrospinal nu
CIC central nu inf coll DpMe deep mesenceph fmj forceps major
RBSC5 Book.qxp 16/9/04 16:36 Page 98

m5 motor root 5n
mcp mid cerebellar ped
Me5 mesenceph 5 nu
me5 mesenceph 5 tr
MEnt medial entorhinal
MiTg microcell teg nu
ml medial lemniscus
mlf med long fasc
MnR median raphe nu
Op optic n layer SC
Pa4 paratrochlear nu
PaS parasubiculum
PBG parabigeminal nu
PDR postdors raphe
Pi ;ineal gland
PL paralemniscal nu
PMnR paramedian raphe
Pn pontine nuclei
PnO pontine retic oral RSD retrosple dysgran Tr triangular nu VIEnt vent int entorh
Post postsubiculum RSGa RSG, a region ts tectospinal tract VLL ventral nu lat lem
PPTg pedunculopont teg RtTg reticulotegmental tth trigeminothal tr VLPAG ventrolat PAG
PrCnF precuneiform area s5 sensory root of 5n v vein Zo zonal layer SC
PRh perirhinal cx scp sup cerebellar ped V1B V1, binocular
Rbd rhabdoid nu SPTg subpedunc teg V1M V1, monocular
rf rhinal fissure SuG superfic gray SC V2L V2, lateral
RR retrorubral nu TeA temporal assocn V2ML V2 cx, mediolat
rs rubrospinal tract tfp trans fibers pons V2MM V2 cx, mediomed

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 98
5 5
9 1
V2ML
10 V2MM V1M
0 Interaural
+15 +10 +5 0 -5 RSD
8 V1B 2
fmj
Pi RSGa
a
V2L
a a Zo
7 3
SuG
Op
InG Post
dcw
InWh
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 TeA 4
cic DpG
ECIC
DpWh
DMPAG
DLPAG
5 PaS 5
bic Ect
LPAG BIC
Aq
CIC
me5 PRh
rf Me5
4 ll VLPAG PrCnF 6
DRD PBG
DRL CnFI
4n MiTg DLEnt
PDR CnFV
4n DRV
DpMe
Pa4 VIEnt
3 v mlf PL CEnt 7
a PPTg
dscp Tr
scp Rbd
SPTg
ATg ERS
2 VLL MEnt 8
m5 PMnR
rs PnO
mcp
ts MnR
ll s5

1 9
RtTg
tth
m5
lfp ml

0 10
Pn

Interaural 1.20 mm tfp

Bregma -7.80 mm
bas
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 99

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 99
5 5
9 1
V1M
V2MM
0 Interaural 10
RSD
+15 +10 +5 0 -5 V1B
8 2
Pi RSGa
a
V2L
Zo
7 3
fmj InG Post
InWh
Com TeA
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 cic ECIC 4
DpG 3 2 1

DMPAG
PaS Ect
5 DLPAG DpWh CIC 5
Aq LPAG
bic PRh
PrCnF
rf VLPAG
4 ll me5 Me5 6
DRD CnFI MiTg DLEnt
4n 4n DRL
CnFV
DRV PDR DpMe PBG

PPTg PL VIEnt
mlf
3 scp 7
CEnt
Tr
Rbd MPL
SPTg
ILL
PL MEnt
2 ERS 8
rs ts PnO VLL
mcp
MnR
m5 PMnR
s5
1 9
ll
RtTg RtTgP
tth
m5
lfp ml

0 10

Pn
Interaural 1.08 mm tfp
bas
Bregma -7.92 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex CEnt coudomed ent cx DLPAG dorsolat PAG Ect ectorhinal cx
2 layer 2 cortex CIC central nu inf coll DMPAG dorsomedial PAG ERS epirubrospinal nu
3 layer 3 cortex cic comm of inf coll DpG deep gray SC fmj forceps major
4n trochlear nerve cll comm of lat lem DpMe deep mesenceph ILL intermediate nu ll
a artery CnFI cuneiform interm DpWh deep white SC InG intermed gray SC
Aq aqueduct CnFV cuneiform vent DRD dorsal raphe dors InWh intermed white SC
B9 B9 serotonin cells Com commiss nu IC DRL dorsal raphe lat lfp long fasc pons
bas basilar artery DLEnt dorsolat entorh cx DRV dorsal raphe vent ll lat lemniscus
bic brachium inf coll DLL dorsal nu lat lem ECIC ext cx inf coll LPAG lat periaqu gray
RBSC5 Book.qxp 16/9/04 16:36 Page 100

m5 motor root 5n
mcp mid cerebellar ped
Me5 mesenceph 5 nu
me5 mesenceph 5 tr
MEnt medial entorhinal
MiTg microcell teg nu
ml medial lemniscus
mlf med long fasc
MnR median raphe nu Pn pontine nuclei rs rubrospinal tract TeA temporal assocn V2MM V2 cx, mediomed
MPL med paralemniscal PnO pontine retic oral RSD retrosple dysgran tfp trans fibers pons VIEnt vent int entorh
PaS parasubiculum Post postsubiculum RSGa RSG, a region Tr triangular nu VLL ventral nu lat lem
PBG parabigeminal nu PPTg pedunculopont teg RtTg reticulotegmental ts tectospinal tract VLPAG ventrolat PAG
PDR postdors raphe PrCnF precuneiform area RtTgP reticuloteg pericen tth trigeminothal tr VTg vent tegment nu
Pi ;ineal gland PRh perirhinal cx s5 sensory root of 5n V1B V1, binocular Zo zonal layer SC
PL paralemniscal nu Rbd rhabdoid nu scp sup cerebellar ped V1M V1, monocular
PMnR paramedian raphe rf rhinal fissure SPTg subpedunc teg V2L V2, lateral

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 100
5 5
9 1

10 V2MM V1M
0 Interaural
+15 +10 +5 0 -5 RSD
8 V1B 2
Pi
RSGa
V2L
7 3
fmj Post

Com
TeA
ECIC
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 cic 3 21 4
DpWh DpG
DMPAG PaS
ll Ect
5 DLPAG CIC 5
LPAG
Aq
Me5
bic PRh
me5
ll VLPAG PrCnF
4 CnFI MiTg 6
rf DRD
4n DRL CnFV
4n DLEnt
DpMe
DRV PDR PPTg
DLL
cll scp mlf CEnt
3 a 7
Rbd VTg
Tr
SPTg
MPL ILL
MnR VIEnt
PMnR MEnt
2 ts
8
rs PnO
VLL
m5

s5 mcp
1 RtTgP 9
ll RtTg
B9
tth
lfp ml
m5
0 10

Pn
Interaural 0.96 mm tfp
bas Bregma -8.04 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 101

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 101
5 5
9 1

0 Interaural 10 V2MM V1M

+15 +10 +5 0 -5
RSD
8 V1B
2

Pi RSGa
fmj V2L
7 3
DCIC Post
Com TeA
cic
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
PaS
DMPAG ECIC
3 21 Ect
CIC
DLPAG
5 5
LPAG
bic Aq
PRh

Me5 CnFD MiTg

rf ll VLPAG
4 4n me5 6
CnFI
4n LDTg CnFV
DRD
v PDR
DRV DLL
cll PPTg CEnt
3 a scp mlf 7
Rbd DLEnt
SPTg Tr
VTg MEnt
MPL ILL

2 mcp MnR 8
PMnR
rs ts
PnO
VLL
m5
s5
1 9
RtTgP
ll RtTg
m5 tth
ml
lfp lfp
0 10

Pn
Interaural 0.84 mm bas
Bregma -8.16 mm
tfp
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex CEnt coudomed ent cx DLEnt dorsolat entorh cx fmj forceps major
2 layer 2 cortex CIC central nu inf coll DLL dorsal nu lat lem ILL intermediate nu ll
3 layer 3 cortex cic comm of inf coll DLPAG dorsolat PAG LDTg laterodorsal teg
4n trochlear nerve cll comm of lat lem DMPAG dorsomedial PAG lfp long fasc pons
a artery CnFD cuneiform dors DPPn dorsal pedunc pont ll lat lemniscus
A7 A7 noradr cells CnFI cuneiform interm DRD dorsal raphe dors LPAG lat periaqu gray
Aq aqueduct CnFV cuneiform vent DRV dorsal raphe vent
bas basilar artery Com commiss nu IC ECIC ext cx inf coll
bic brachium inf coll DCIC dorsal cx IC Ect ectorhinal cx
RBSC5 Book.qxp 16/9/04 16:36 Page 102

m5 motor root 5n MnR median raphe nu Post postsubiculum RtTg reticulotegmental Tr triangular nu VLL ventral nu lat lem
mcp mid cerebellar ped MPL med paralemniscal PPTg pedunculopont teg RtTgP reticuloteg pericen ts tectospinal tract VLPAG ventrolat PAG
Me5 mesenceph 5 nu PaS parasubiculum PRh perirhinal cx s5 sensory root of 5n tth trigeminothal tr VTg vent tegment nu
me5 mesenceph 5 tr PDR postdors raphe Rbd rhabdoid nu Sag sagulum nu v vein
MEnt medial entorhinal Pi ;ineal gland rf rhinal fissure scp sup cerebellar ped V1B V1, binocular
MiTg microcell teg nu PMnR paramedian raphe rs rubrospinal tract SPTg subpedunc teg V1M V1, monocular
ml medial lemniscus Pn pontine nuclei RSD retrosple dysgran TeA temporal assocn V2L V2, lateral
mlf med long fasc PnO pontine retic oral RSGa RSG, a region tfp trans fibers pons V2MM V2 cx, mediomed

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 102
5 v
9 5 1

10 V2MM V1M
0 Interaural
+15 +10 +5 0 -5
RSD
8 V1B 2
Pi
RSGa
V2L
7 3
Post
DCIC
Com
TeA
cic
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 PaS 4
ECIC
DMPAG 3 21

CIC Ect

5 5
LPAG
Aq
PRh
Me5
bic me5
rf CnFD
4 VLPAG 6
4n CnFI Sag
ll 4n CnFV
LDTg
DRD
DRV PPTg DLL
cll Rbd CEnt DLEnt
a
3 a scp 7
mlf
Tr MEnt
cll SPTg
VTg
MPL ILL
mcp
2 MnR A7 8
ts PMnR
rs PnO
m5
s5
VLL
1 9
RtTgP
RtTg DPPn
ll
tth
ml
0 m5 lfp 10

Interaural 0.72 mm tfp

bas
Pn
Bregma -8.28 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 103

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
V2MM
V1M
Figure 103
5 5
8 RSD 2
V1B

0 Interaural 10
Pi RSGa
+15 +10 +5 0 -5 V2L
7 3
DCIC Post

TeA
6 DMPAG 4
ECIC PaS
3 21

CIC
Ect
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 LPAG
5
2Cb
PRh

4n Aq
ll VLPAG CnFD
4 me5 Me5
6
rf CnFI Sag
4n CnFV
DRD LDTg
DLEnt
cll PPTg DLL
a DRV CEnt
3 scp 7
a
mlf Rbd MEnt
SPTg
DRI VTg Tr
PMnR ILL
MPL
2 mcp MnR 8
A7
ts
rs PnO
m5
1 s5 RtTgP 9
VLL
RtTg PLV

m5 ll tth
0 10
lfp ml

Pn
tfp
-1 11
bas

Interaural 0.60 mm Bregma -8.40 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex bas basilar artery DLL dorsal nu lat lem ILL intermediate nu ll
2 layer 2 cortex CEnt coudomed ent cx DMPAG dorsomedial PAG LDTg laterodorsal teg
2Cb 2nd Cb lobule CIC central nu inf coll DRC dorsal raphe caud LDTgV latdors teg vent
3 layer 3 cortex cll comm of lat lem DRD dorsal raphe dors LEnt lat entorhinal cx
4n trochlear nerve CnFD cuneiform dors DRI dorsal raphe intermed lfp long fasc pons
4V 4th ventricle CnFI cuneiform interm DRV dorsal raphe vent ll lat lemniscus
a artery CnFV cuneiform vent DTgP dors teg pericent LPAG lat periaqu gray
A7 A7 noradr cells DCIC dorsal cx IC ECIC ext cx inf coll
Aq aqueduct DLEnt dorsolat entorh cx Ect ectorhinal cx
RBSC5 Book.qxp 16/9/04 16:36 Page 104

m5 motor root 5n
mcp mid cerebellar ped
Me5 mesenceph 5 nu
me5 mesenceph 5 tr
MEnt medial entorhinal
ml medial lemniscus
mlf med long fasc
MnR median raphe nu
MPL med paralemniscal
PaS parasubiculum
Pi ;ineal gland
PLV perilemnisc vent
PMnR paramedian raphe
Pn pontine nuclei
PnO pontine retic oral
Post postsubiculum
PPTg pedunculopont teg RtTg reticulotegmental Tr triangular nu VLPAG ventrolat PAG
PRh perirhinal cx RtTgP reticuloteg pericen ts tectospinal tract VTg vent tegment nu
Rbd rhabdoid nu s5 sensory root of 5n tth trigeminothal tr
ReIC recess of inf coll Sag sagulum nu V1B V1, binocular
rf rhinal fissure scp sup cerebellar ped V1M V1, monocular
rs rubrospinal tract SPTg subpedunc teg V2L V2, lateral
RSD retrosple dysgran TeA temporal assocn V2MM V2 cx, mediomed
RSGa RSG, a region tfp trans fibers pons VLL ventral nu lat lem

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
V2MM
V1M
Figure 104
5 5
8 V1B 2
RSD
0 Interaural 10
Pi V2L
+15 +10 +5 0 -5
7 3
RSGa TeA
DCIC

6 DMPAG 4
ECIC PaS Ect
3 2 1
CIC
ReIC
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 PRh 5
LPAG

2Cb
rf 4n

VLPAG LEnt
4 CnFD 6
me5 4V Me5 CnFI
Sag
CEnt
4n CnFV
a DRC DTgP LDTg PPTg DLL
a LDTgV MEnt
3 scp 7
mlf Tr
SPTg
PPTg ILL
VTg
MPL
mcp
2 MnR PMnR A7 8
ll ts
rs
m5 PnO
s5
1 RtTgP 9
PLV
VLL
RtTg

tth
0 m5 10
ll ml
tfp lfp

Pn
tfp
-1 11
bas

Interaural 0.48 mm Bregma -8.52 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 105

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
V2MM
V1M Figure 105
5 5
8 V1B 2

10 Pi
0 Interaural
V2L
+15 +10 +5 0 -5
7 RSD 3
DCIC TeA

DMPAG

6 PaS 4
CIC ECIC Ect
ReIC 3 2 1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 PRh 5
LPAG
v
4n 2Cb
rf
4 VLPAG CnFD CnFI Sag 6
CnFV LEnt
4n 4V Me5
RL CEnt
me5 LPB LPBS
aa DRC LDTg DLL
a scp DTgP PPTg
MEnt
3 MPL
Tr
7
LDTgV
mlf ILL
VTg MPB
mcp vsc DMTg
MnR PMnR KF
2 A7 8

rs ts
m5
ll PnO
1 s5 9
tz
RtTg PLV VLL
m5
Pn CAT
tth
0 MVPO 10
ml
lfp

tfp Pn

-1 11
bas

Interaural 0.36 mm Bregma -8.64 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex 4n trochlear nerve CG central gray DMPAG dorsomedial PAG KF Kîlliker-Fuse nu LPB lat parabrach
2 layer 2 cortex 4V 4th ventricle CIC central nu inf coll DMTg dorsomed teg area LDTg laterodorsal teg LPBC lat parabrach cent
2/3Cb 2nd&3rd Cb lobs a artery CnFD cuneiform dors DRC dorsal raphe caud LDTgV latdors teg vent LPBE lat parabrach ext
2Cb 2nd Cb lobule A7 A7 noradr cells CnFI cuneiform interm DTgP dors teg pericent LEnt lat entorhinal cx LPBS lat parabrach sup
3 layer 3 cortex bas basilar artery CnFV cuneiform vent ECIC ext cx inf coll lfp long fasc pons
3Cb 3rd Cb lobule CAT nu cent acoust tr DCIC dorsal cx IC Ect ectorhinal cx ll lat lemniscus
4/5Cb 4th&5th Cb lob CEnt coudomed ent cx DLL dorsal nu lat lem ILL intermediate nu ll LPAG lat periaqu gray
RBSC5 Book.qxp 16/9/04 16:36 Page 106

m5 motor root 5n
mcp mid cerebellar ped
Me5 mesenceph 5 nu
me5 mesenceph 5 tr
MEnt medial entorhinal
ml medial lemniscus
mlf med long fasc
MnR median raphe nu
MPB medial parabrach
MPL med paralemniscal
MVPO mediovent periol
PaS parasubiculum
Pi ;ineal gland
PLV perilemnisc vent
PMnR paramedian raphe
Pn pontine nuclei
PnO pontine retic oral RtTg reticulotegmental ts tectospinal tract VLL ventral nu lat lem
PPTg pedunculopont teg s5 sensory root of 5n tth trigeminothal tr VLPAG ventrolat PAG
PRh perirhinal cx Sag sagulum nu tz trapezoid body vsc vent spinocer tr
ReIC recess of inf coll scp sup cerebellar ped v vein VTg vent tegment nu
rf rhinal fissure Su5 supratrigem nu V1B V1, binocular
RL retrolemniscal nu TeA temporal assocn V1M V1, monocular
rs rubrospinal tract tfp trans fibers pons V2L V2, lateral
RSD retrosple dysgran Tr triangular nu V2MM V2 cx, mediomed

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0

V1M
Figure 106
5 5 V2MM
8 2
V1B

0 Interaural 10 Pi

+15 +10 +5 0 -5 V2L

7 RSD 3
DCIC
TeA
DMPAG

6 ReIC ECIC PaS 4

CIC 3 2 1 Ect
3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
2Cb PRh

4n 4/5Cb
rf v 4n LEnt
4 CnFD 6
CEnt
4V VLPAG
2/3Cb
LPBS
LPBC
me5 LDTg Me5 Sag
DRC PPTg DLL
DTgP
scp
3 LPBE
7
CG LDTgV
vsc MPB
mlf
mcp DMTg
MnR Su5 KF

2 8
ts A7 A7

m5 rs
PnO
s5 PMnR
1 tz vsc 9
ll
PLV
RtTg VLL

CAT
0 tth 10
MVPO
ml
lfp

tfp
-1 11
bas

Interaural 0.24 mm Bregma -8.76 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 107

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 107
5 5
8 V1M 2
V2MM
0 Interaural 10 V1B
+15 +10 +5 0 -5
7 3
RSD V2L
DCIC

6 4
ReIC 3 2 1 Ect
ECIC PaS
CIC
3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
PRh
2Cb

4n 4/5Cb
4 a LEnt v
6
rf Me5 LPBD
4n LPBCr
LPBI RL
me5 2/3Cb
vsc
a 4V CEnt
m5 DRC DTgP LPBV
3 scp LDTg Fl 7
DTgC LPBC
LPBE
CG LDTgV MPB
mcp
mlf KF
DMTg Su5
2 A7
8
PnR SubCD
P5 Pr5VL
ts
m5 PnO
1 9
tz rs SubCV
s5
ll vsc
RtTg

VLL
0 tth
10
CAT
Tz
ml
tz MVPO
lfp
-1 tfp 11
bas
Interaural 0.12 mm Bregma -8.88 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex 4Cb 4th Cb lobule Bar Barrington’s nu DCIC dorsal cx IC Fl flocculus LPBC lat parabrach cent
2 layer 2 cortex 4n trochlear nerve bas basilar artery DMTg dorsomed teg area KF Kîlliker-Fuse nu LPBCr lat parab crescent
2/3Cb 2nd&3rd Cb lobs 4V 4th ventricle CAT nu cent acoust tr DRC dorsal raphe caud LDTg laterodorsal teg LPBD lat parabrach dors
2Cb 2nd Cb lobule 5N motor trigeminal cbw cereb white mat DTgC dorsal teg nu, cent LDTgV latdors teg vent LPBE lat parabrach ext
3 layer 3 cortex a artery CEnt coudomed ent cx DTgP dors teg pericent LEnt lat entorhinal cx LPBI lat parabrach int
3Cb 3rd Cb lobule A5 A5 noradr cells CG central gray ECIC ext cx inf coll lfp long fasc pons LPBV lat parabrach vent
4/5Cb 4th&5th Cb lob A7 A7 noradr cells CIC central nu inf coll Ect ectorhinal cx ll lat lemniscus
RBSC5 Book.qxp 16/9/04 16:36 Page 108

m5 motor root 5n MPBE med parabrach ext Pr5VL princ 5 ventrolat RSD retrosple dysgran tfp trans fibers pons V1M V1, monocular
mcp mid cerebellar ped P5 peritrigeminal zn PRh perirhinal cx RtTg reticulotegmental ts tectospinal tract V2L V2, lateral
Me5 mesenceph 5 nu PaS parasubiculum py pyramidal tract s5 sensory root of 5n tth trigeminothal tr V2MM V2 cx, mediomed
me5 mesenceph 5 tr PC5 parvicell motor 5 ReIC recess of inf coll scp sup cerebellar ped Tz nu trapezoid body VLL ventral nu lat lem
ml medial lemniscus PnC pontine retic caud rf rhinal fissure Su5 supratrigem nu tz trapezoid body VMPO ventromed preopt
mlf med long fasc PnO pontine retic oral RL retrolemniscal nu SubCD SubC dorsal v vein vsc vent spinocer tr
MPB medial parabrach PnR pontine raphe nu rs rubrospinal tract SubCV SubC ventral V1B V1, binocular

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 108
5 5
8 2
V1M
0 Interaural 10 V1B

+15 +10 +5 0 -5
7 4Cb 3
V2L

RSD
DCIC
6 ECIC Ect 4
3 2 1
ReIC
PaS
CIC
3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 PRh 5

cbw

4 rf
2Cb 4n 6
4/5Cb
CEnt
4n
4V LPBI LPBCr
LPBD 2/3Cb
3 DRC Bar 7
DTgP LPBV Fl
scp LDTg
me5 DTgC
MPB LPBC
CG MPBE LPBE
mcp
2 mlf LDTgV Su5 KF 8
DMTg
A7
SubCD
PnR 5N
P5 PC5
Pr5VL
1 9
ts PnC
s5
SubCV
m5 rs RtTg

0 ll VLL 10
A5
tth CAT
vsc
tz ml Tz
tz MVPO
-1 11
py

Interaural 0.00 mm bas Bregma -9.00 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 109

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 109
5 5 5Cb
8 2
V1B
V1M
0 Interaural 10
4Cb
+15 +10 +5 0 -5
V2L
7 3
pcuf
ECIC RSD
DCIC 3 2 1
Ect
3Cb
6 4
PaS
CIC
PRh
3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5

CEnt
rf 2Cb
a cbw 4/5Cb
4 6
vsc LPBD
a 2Cb
LPBV LPBI LPBCr
4V 4n 2/3Cb

3 scp DRC LDTg Bar LPBC 7

DTgC DTgP
cbw Me5 LPBE Fl
me5
mcp MPB
CGB MPBE KF
CGA
mlf SubCA Su5 GrC
2 DMTg SubCD 8
5Te
VCA
P5 5Ma
ts 5MHy Pr5VL

1 s5 RtTg 9
PC5
PnC
tz m5 SubCV

A5
rs
0 MSO 10
vsc LVPO
tth CAT
ml Tz
tz MVPO
-1 py 11

Interaural -0.12 mm bas

Bregma -9.12 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1 layer 1 cortex 4Cb 4th Cb lobule A5 A5 noradr cells CGO cent gray nu O Ect ectorhinal cx LPBE lat parabrach ext
2 layer 2 cortex 4n trochlear nerve Bar Barrington’s nu CIC central nu inf coll Fl flocculus LPBI lat parabrach int
2/3Cb 2nd&3rd Cb lobs 4V 4th ventricle bas basilar artery DCIC dorsal cx IC GrC granule cochlear LPBM lat parabrach med
2bCb 2b cerbellar lob 5Cb 5th Cb lobule CAT nu cent acoust tr DMTg dorsomed teg area KF Kîlliker-Fuse nu LPBV lat parabrach vent
2Cb 2nd Cb lobule 5Ma mo 5 masseter cbw cereb white mat DRC dorsal raphe caud LDTg laterodorsal teg LVPO laterovent periol
3 layer 3 cortex 5MHy mo 5 mylohyoid CEnt coudomed ent cx DTgC dorsal teg nu, cent LPBC lat parabrach cent
3Cb 3rd Cb lobule 5Te mo 5 temporalis CGA central gray alpha DTgP dors teg pericent LPBCr lat parab crescent
4/5Cb 4th&5th Cb lob a artery CGB central gray bets ECIC ext cx inf coll LPBD lat parabrach dors
RBSC5 Book.qxp 16/9/04 16:36 Page 110

m5 motor root 5n MPBE med parabrach ext PnC pontine retic caud RSD retrosple dysgran SubCD SubC dorsal V1M V1, monocular
mcp mid cerebellar ped MSO med superior olive Pr5DL princ 5 dorsolat RtTg reticulotegmental SubCV SubC ventral V2L V2, lateral
Me5 mesenceph 5 nu MVPO mediovent periol Pr5VL princ 5 ventrolat s5 sensory root of 5n ts tectospinal tract VCA vent cochlear ant
me5 mesenceph 5 tr P5 peritrigeminal zn PRh perirhinal cx scp sup cerebellar ped tth trigeminothal tr VMPO ventromed preopt
ml medial lemniscus PaS parasubiculum py pyramidal tract SPO superior paraoliv Tz nu trapezoid body vsc vent spinocer tr
mlf med long fasc PC5 parvicell motor 5 rf rhinal fissure Su5 supratrigem nu tz trapezoid body
MPB medial parabrach pcuf preculminate fiss rs rubrospinal tract SubCA SubC alpha V1B V1, binocular

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 110
5 5 5Cb
8 V1B 2
V1M
0 Interaural 10 V2L
4Cb
+15 +10 +5 0 -5
7 Ect 3
pcuf
RSD
DCIC
PRh
rf
6 PaS 4
3Cb ECIC

3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
5Cb CEnt

cbw 4Cb

4 6
2bCb
LPBC vsc 2Cb LPBV LPBD
4n LPBI
4V Me5
Fl
cbw scp LPBCr
3 LPBE DRC DTgP Bar 7
mcp me5 DTgC LDTg MPB
MPBE KF
CGB
CGO CGA Su5 GrC
SubCA mlf
SubCD VCA
2 DMTg 5Te Pr5DM 8
5Ma
P5
5MHy Pr5VL
tz RtTg s5
ts
1 PC5 9
m5
PnC SubCV
A5
rs MSO

0 tth
SPO 10
vsc
ml Tz LVPO
tz

py MVPO
-1 11
bas
Interaural -0.24 mm Bregma -9.24 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 111

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 111
5 5 5Cb
8 2
V1B
V1M
0 Interaural 10
4Cb V2L
+15 +10 +5 0 -5
7 Ect 3
pcuf RSD
rf PRh
3Cb IC
6 prf
4
Sim

5Cb
3Cb CEnt
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
cbw

4 cbw 6
2Cb LPBI
vsc
LPBV 4n LPB
4V LPBE
scp Me5 Fl
mcp Sph DR LPBC
3 cbw 7
LDTgBar MPB MPBE
Pr5DM me5
PDTg
CGB Su5 PCGS
CGO CGA
GrC
mlf 5Te VCA
2 SubCA 8
DMTg
P5
5Ma
RtTg Pr5VL
s5 SubCD
m5 5VM
tz
1 PC5 9
ts PnC SubCV
A5
A5
rs
7n
0 vsc SPO 10
LVPO
tth
ml Tz MSO
tz
tz MVPO
py
-1 11
bas

Interaural -0.36 mm Bregma -9.36 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 5VM mo 5 ventromed CGB central gray bets LC locus coeruleus
2Cb 2nd Cb lobule 7n facial nerve CGO cent gray nu O LDTg laterodorsal teg
3Cb 3rd Cb lobule 8cn cochlear root 8 n Cx cerebral cortex LPB lat parabrach
4Cb 4th Cb lobule A5 A5 noradr cells DMTg dorsomed teg area LPBC lat parabrach cent
4n trochlear nerve Bar Barrington’s nu DR dorsal raphe nu LPBE lat parabrach ext
4V 4th ventricle bas basilar artery Ect ectorhinal cx LPBI lat parabrach int
5Cb 5th Cb lobule cbw cereb white mat Fl flocculus LPBV lat parabrach vent
5Ma mo 5 masseter CEnt coudomed ent cx GrC granule cochlear LSO lat superior olive
5Te mo 5 temporalis CGA central gray alpha IC inferior colliculus LVPO laterovent periol
RBSC5 Book.qxp 16/9/04 16:36 Page 112

m5 motor root 5n
mcp mid cerebellar ped
Me5 mesenceph 5 nu
me5 mesenceph 5 tr
ml medial lemniscus
mlf med long fasc
MPB medial parabrach
MPBE med parabrach ext
MSO med superior olive
MVPO mediovent periol
P5 peritrigeminal zn
PC5 parvicell motor 5
PCGS paracoch glial
pcuf preculminate fiss
PDTg posterodorsal teg
PnC pontine retic caud
POH periolivary horn RSD retrosple dysgran Sph sphenoid nu Tz nu trapezoid body
Pr5DM princ 5 dorsomed RtTg reticulotegmental SPO superior paraoliv tz trapezoid body
Pr5VL princ 5 ventrolat RtTgL reticuloteg lat Su5 supratrigem nu V1B V1, binocular
prf primary fissure s5 sensory root of 5n SubCA SubC alpha V1M V1, monocular
PRh perirhinal cx scp sup cerebellar ped SubCD SubC dorsal V2L V2, lateral
py pyramidal tract scpd scp descend limb SubCV SubC ventral VCA vent cochlear ant
rf rhinal fissure Sim simple lobule ts tectospinal tract vsc vent spinocer tr
rs rubrospinal tract SMV sup medull velum tth trigeminothal tr

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 112
5 5Cb
8 5 2

0 Interaural 10
cbw
Cx
+15 +10 +5 0 -5
4Cb
7 3

pcuf
prf CEnt
6 4
3Cb
Sim

cbw
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 cbw 5

cbw
4 6
vsc LPBI
2Cb
SMV LPBV
4n LC
scp LPB
mcp 1Cb Fl
3 cbw 4V LDTg Me5 7
Sph MPB
Pr5DM
PDTg CGB me5 PCGS
Bar Su5
CGO CGA GrC
scpd
SubCA mlf SubCD 5Te VCA
2 DMTg
8
RtTgL
P5 5Ma
s5 PC5 RtTg Pr5VL
5VM
tz m5
8cn
1 ts SubCV 9
PnC POH A5

7n rs LSO
A5
7n
0 vsc MSO SPO LVPO 10
tth
ml Tz
tz
MVPO

-1 py 11
bas
Interaural -0.48 mm Bregma -9.48 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 113

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 113
5 5 5Cb
8 2

0 Interaural 10

+15 +10 +5 0 -5 Cx
cbw
7 4Cb 3

prf
pcuf

6 4
Sim
3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 cbw 5

a
cbw
4 PFl
6
2Cb LPBI LPBV
LPB
vsc 4V
1Cb Fl
mcp scp 4V MPB
3 cbw
Sph LC PCGS 7
CGPn Me5 Pr5DM
GrC
me5 PDTg
CGB Su5 5Tr
CGO CGA SubCD LR4V
scpd mlf P5
2 DMTg 5Te VCA 8
RtTgL
RtTg 5Ma
5Acs
s5 Pr5VL
5VM
m5
1 8cn ts POH 9
PnC SubCV
RIP
DPO
7n
rs PnV A5
7n LSO
0 RMg 10
vsc MSO SPO LVPO
tz tth
ml Tz
tz MVPO

py
-1 11
bas

Interaural -0.60 mm Bregma -9.60 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 5Tr trigem trans zone CGB central gray bets LPB lat parabrach
2Cb 2nd Cb lobule 5VM mo 5 ventromed CGO cent gray nu O LPBI lat parabrach int
3Cb 3rd Cb lobule 7n facial nerve CGPn central gray pons LPBV lat parabrach vent
4Cb 4th Cb lobule 8cn cochlear root 8 n Cx cerebral cortex LR4V lat recess 4V
4V 4th ventricle a artery DMTg dorsomed teg area LSO lat superior olive
5Acs mo 5 accessory A5 A5 noradr cells DPO dorsal periolivary LVPO laterovent periol
5Cb 5th Cb lobule bas basilar artery Fl flocculus
5Ma mo 5 masseter cbw cereb white mat GrC granule cochlear
5Te mo 5 temporalis CGA central gray alpha LC locus coeruleus
RBSC5 Book.qxp 16/9/04 16:36 Page 114

m5 motor root 5n MVPO mediovent periol POH periolivary horn rs rubrospinal tract SimB simple lobule B ts tectospinal tract
mcp mid cerebellar ped P5 peritrigeminal zn Pr5DM princ 5 dorsomed RtTg reticulotegmental SMV sup medull velum tth trigeminothal tr
Me5 mesenceph 5 nu PCGS paracoch glial Pr5VL princ 5 ventrolat RtTgL reticuloteg lat Sph sphenoid nu Tz nu trapezoid body
me5 mesenceph 5 tr pcuf preculminate fiss prf primary fissure s5 sensory root of 5n SPO superior paraoliv tz trapezoid body
ml medial lemniscus PDTg posterodorsal teg py pyramidal tract scp sup cerebellar ped Su5 supratrigem nu VCA vent cochlear ant
mlf med long fasc PFl paraflocculus RIP raphe interpos nu scpd scp descend limb SubCD SubC dorsal vsc vent spinocer tr
MPB medial parabrach PnC pontine retic caud RMg raphe magnus nu Sim simple lobule SubCV SubC ventral
MSO med superior olive PnV pontine retic vent RPa raphe pallidus nu SimA simple lobule A SuVe superior vestib nu

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 114
5 5 5Cb
8 2

0 Interaural 10

+15 +10 +5 0 -5
cbw 4Cb
7 3

pcuf

SimA
6 3Cb 4
cbw
SimB
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 prf 5
2Cb
cbw

PFl
a 2Cb
4 cbw 6

vsc LPB
mcp scp
1Cb Fl
3 SMV SuVe PCGS 7
cbw 4V LC Me5 MPB
4V GrC
PDTg Pr5DM
scpd me5 CGO CGB 5Tr
CGA LR4V
2 4V 8
SubCD P5
RtTgL DMTg 5Te VCA
mlf
s5 5Acs 5Ma
m5 Pr5VL
1 9
POH
8cn tz ts PnC SubCV
RIP
DPO
7n PnV
0 rs A5 10
RMg LSO
MSO SPO
tth LVPO
vsc RPa
ml Tz
tz MVPO
-1 py 11

bas
Interaural -0.72 mm Bregma -9.72 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 115

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 115
5Cb
5 5
8 cbw
2

0 Interaural 10

+15 +10 +5 0 -5
7 4Cb 3
cbw
pcuf SimA

6 4
3Cb

prf cbw SimB

The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 cbw 5
psf

PFl
cbw 2Cb
4 6

vsc
un
mcp
plf scp 1Cb
3 7
4V
LC MPB SuVe Fl
cbw Me5
ocb 4V PDTg CGPn PCGS
5Tr Pr5DM GrC
me5 CGB CGG
scpd MVeMC LR4V VCAGr
8vn 4V CGA
2 ocb SubCD 8
DMTg P5
mlf II 5Te VCA

s5 5Acs 5Ma
m5
tz
1 RIP PCRt Pr5VL 9
8cn ts PnC POH
ocb SubCV
DPO
7n 6n
0 RMg 10
LSO
rs PnV A5
vsc tth SPO
MSO
ml RPa LVPO
Tz
tz MVPO
-1 py 11

bas
Interaural -0.84 mm Bregma -9.84 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 5Tr trigem trans zone CGB central gray bets II int interstit mlf
2Cb 2nd Cb lobule 6n root of abducens CGG cent gray gamma LC locus coeruleus
3Cb 3rd Cb lobule 7n facial nerve CGPn central gray pons LR4V lat recess 4V
4Cb 4th Cb lobule 8cn cochlear root 8 n DMTg dorsomed teg area LSO lat superior olive
4V 4th ventricle 8vn vestib root 8 n DPO dorsal periolivary LVPO laterovent periol
5Acs mo 5 accessory A5 A5 noradr cells EVe nu efferents 8vn
5Cb 5th Cb lobule bas basilar artery Fl flocculus
5Ma mo 5 masseter cbw cereb white mat GrC granule cochlear
5Te mo 5 temporalis CGA central gray alpha icp inf cerebellar ped
RBSC5 Book.qxp 16/9/04 16:36 Page 116

m5 motor root 5n MVPO mediovent periol PnC pontine retic caud RMg raphe magnus nu SMV sup medull velum un uncinate fascic
mcp mid cerebellar ped ocb olivocochl bundle PnV pontine retic vent RPa raphe pallidus nu SPO superior paraoliv VCA vent cochlear ant
Me5 mesenceph 5 nu P5 peritrigeminal zn POH periolivary horn rs rubrospinal tract SubCD SubC dorsal VCAGr VCA granule layer
me5 mesenceph 5 tr PCGS paracoch glial Pr5DM princ 5 dorsomed s5 sensory root of 5n SubCV SubC ventral veme vestibulomes tr
ml medial lemniscus PCRt parvicell ret nu Pr5VL princ 5 ventrolat scp sup cerebellar ped SuVe superior vestib nu vsc vent spinocer tr
mlf med long fasc pcuf preculminate fiss prf primary fissure scpd scp descend limb ts tectospinal tract
MPB medial parabrach PDTg posterodorsal teg psf post superior fiss SGe supragenual nu tth trigeminothal tr
MSO med superior olive PFl paraflocculus py pyramidal tract SimA simple lobule A Tz nu trapezoid body
MVeMC med vestib magno plf posterolat fissure RIP raphe interpos nu SimB simple lobule B tz trapezoid body

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0

5Cb
Figure 116
5 5
8 2
cbw

0 Interaural 10

+15 +10 +5 0 -5
7 4Cb 3
cbw SimA
pcuf

6 4
SimB
prf 3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 cbw 5
psf

4 cbw 2Cb 6
PFl
vsc
cbw un
mcp
scp
plf icp MPB
1Cb
3 GrC 7
SMV Me5 SuVe Fl
4V LC PCGS LR4V
MVeMC
SGe EVe Me5
4V
CGG 5Tr VCAGr
2 veme 8
8vn 7n ocb mlf II CGA Pr5DM VCA
P5
s5
5Acs PCRt
1 Pr5VL 9
ocb ts RIP PnC SubCD
8cn tz SubCV
7n POH DPO
A5
0 6n PnV 10
rs A5
RMg LSO
vsc SPO
MSO tth
LVPO
ml RPa Tz
tz MVPO
-1 11
py

Interaural -0.96 mm bas

Bregma -9.96 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 117

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 117
5
5Cb
8 5 2
cbw
0 Interaural 10

+15 +10 +5 0 -5
7 4Cb SimA 3

cbw

SimB
6 4
pcuf
prf

psf Crus1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 3Cb 5
cbw

2Cb

4 6
cbw vsc PFl
un
mcp icp scp
3 1Cb LC 7
plf SuVe GrC
LR4V
MVePC PCGS
EVe Fl
SGe MVeMC
4V CGA CGG VCAGr
2 veme ocb 5Tr 8
mlf 6N VCA
8vn Pr5DM
Pa6
s5 PCRtA
ocb Pr5VL
1 9
8cn 7n RIP IRt
ts PnC
6Acs
A5
0 tz 10
rs LSO A5
PnV
vsc RMg
MSO tth SPO
ml RPa Tz
LVPO
-1 11
py tz
MVPO

Interaural -1.08 mm bas

Bregma -10.08 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 7n facial nerve chp choroid plexus IRt intermed ret nu
2Cb 2nd Cb lobule 8cn cochlear root 8 n Crus1 crus 1 ansiform LC locus coeruleus
3Cb 3rd Cb lobule 8vn vestib root 8 n DPO dorsal periolivary LR4V lat recess 4V
4Cb 4th Cb lobule a artery EVe nu efferents 8vn LSO lat superior olive
4V 4th ventricle A5 A5 noradr cells Fl flocculus LVPO laterovent periol
5Cb 5th Cb lobule bas basilar artery g7 genu of 7 n
5Tr trigem trans zone cbw cereb white mat GrC granule cochlear
6Acs access abducens CGA central gray alpha I8 interstitial nu 8n
6N abducens nu CGG cent gray gamma icp inf cerebellar ped
RBSC5 Book.qxp 16/9/04 16:36 Page 118

mcp mid cerebellar ped Pa6 paraabducens nu Pr5DM princ 5 dorsomed rs rubrospinal tract SuS sup salivatory nu VCA vent cochlear ant
ml medial lemniscus PCGS paracoch glial Pr5VL princ 5 ventrolat s5 sensory root of 5n SuVe superior vestib nu VCAGr VCA granule layer
mlf med long fasc PCRtA parvicell ret alpha prf primary fissure scp sup cerebellar ped ts tectospinal tract veme vestibulomes tr
MSO med superior olive pcuf preculminate fiss psf post superior fiss SGe supragenual nu tth trigeminothal tr vsc vent spinocer tr
MVeMC med vestib magno PFl paraflocculus py pyramidal tract SimA simple lobule A Tz nu trapezoid body
MVePC med vestib parvi plf posterolat fissure RIP raphe interpos nu SimB simple lobule B tz trapezoid body
MVPO mediovent periol PnC pontine retic caud RMg raphe magnus nu sp5 sp trigeminal tr tzd trapezoid decuss
ocb olivocochl bundle PnV pontine retic vent RPa raphe pallidus nu SPO superior paraoliv un uncinate fascic

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 118
5Cb
5 5
8 2
cbw
0 Interaural 10

+15 +10 +5 0 -5 SimA

7 4Cb 3

cbw
SimB

6 prf 4
pcuf
3Cb
Crus1
psf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5

cbw

2Cb
4 6
a vsc
un
mcp scp
icp LC
3 cbw 7
1Cb SuVe GrC
chp MVePC LR4V PFl
cbw
EVe PCGS Fl
plf 8vn 4V SGe MVeMC
g7 CGA CGG VCAGr
2 veme
8
mlf Pr5DM
6N VCA
ocb Pa6 PCRtA
7n
sp5
1 IRt Pr5VL
9
RIP I8
ts PnC
8vn 6Acs
SuS
A5
0 tz rs PnV DPO LSO 10

vsc RMg

RPa SPO MSO

ml
Tz LVPO
-1 tzd 11
tz
py MVPO

Interaural -1.20 mm bas

Bregma 10.20 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 119

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 119
5Cb
5 5
8 2

0 Interaural 10
cbw
+15 +10 +5 0 -5 SimA
7 3
4Cb

SimB
cbw
prf
6 4
pcuf Crus1
psf 3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
pfs

cbw
4 2Cb 6

un
scp
vsc 1Cb
LC PFl
3 7
MVePC SuVe
chp GrC
cbw LR4V
plf Pr EVe Fl
4V SGe MVeMC PCGS
8vn
g7
2 veme mlf 6N Pr5DM 8
7n
ocb veme Pa6 VCA
PCRtA
7n

1 IRt 9
sp5
SuS Pr5VL I8
ts 6Acs
Gi
8vn SuS
RIP
A5
0 DPO LSO
10
rs GiA
vsc
tz RMg LPGi
SPO MSO
ml RPa
tz Tz MVPO LVPO
-1 11
py

bas
Interaural -1.32 mm Bregma 10.32 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 6Acs access abducens A5 A5 noradr cells DMSp5 dorsomed sp 5 nu Gi gigantocell ret nu IRt intermed ret nu
2Cb 2nd Cb lobule 6N abducens nu bas basilar artery DPGi dors paragigantoc GiA gigantocell alpha IRtA intermed rt alpha
3Cb 3rd Cb lobule 7Acs mo 7 accessory cbc cerebellar comm DPO dorsal periolivary GrC granule cochlear LC locus coeruleus
4Cb 4th Cb lobule 7n facial nerve cbw cereb white mat EVe nu efferents 8vn GrCb granule layer LPGi lat paragig cell
4V 4th ventricle 8vn vestib root 8 n chp choroid plexus Fl flocculus I8 interstitial nu 8n LR4V lat recess 4V
5Cb 5th Cb lobule a artery Crus1 crus 1 ansiform g7 genu of 7 n icp inf cerebellar ped LSO lat superior olive
LVPO laterovent periol
RBSC5 Book.qxp 16/9/04 16:36 Page 120

ml medial lemniscus P7 perifacial zone Pr prepositus nu RPa raphe pallidus nu SPO superior paraoliv VCA vent cochlear ant
mlf med long fasc Pa6 paraabducens nu Pr5DM princ 5 dorsomed rs rubrospinal tract SuS sup salivatory nu VCAGr VCA granule layer
MoCb molecul layer Cb PCRtA parvicell ret alpha Pr5VL princ 5 ventrolat scp sup cerebellar ped SuVe superior vestib nu VCP vent cochlear post
MSO med superior olive pcuf preculminate fiss prf primary fissure SGe supragenual nu ts tectospinal tract veme vestibulomes tr
MVeMC med vestib magno PFl paraflocculus psf post superior fiss SimA simple lobule A tth trigeminothal tr vsc vent spinocer tr
MVePC med vestib parvi pfs paraflocc sulcus py pyramidal tract SimB simple lobule B Tz nu trapezoid body
MVPO mediovent periol Pk Purkinje cells RIP raphe interpos nu sp5 sp trigeminal tr tz trapezoid body
ocb olivocochl bundle plf posterolat fissure RMg raphe magnus nu Sp5O spinal 5 oral un uncinate fascic

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 120
5 5 5Cb
8 2

0 Interaural 10
cbw SimA
+15 +10 +5 0 -5 GrCb
7 4Cb Pk 3
MoCb
SimB

cbw
prf
6 4

3Cb Crus1
psf
pcuf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5

pfs PFl

2Cb
4 6
a vsc cbc

scp un
cbw icp
3 plf 7
1Cb SuVe PFl
MVePC LR4V
GrC
cbw EVe Fl
4V Pr MVeMC
ocb
2 g7 mlf VCA 8
8vn Pa6 VCAGr
veme DMSp5 VCP
DPGi
sp5 Sp5O
PCRtA I8
1 9
IRtA
ocb ts
8vn 6Acs SuS Pr5VL
Gi 7Acs

0 rs A5 10
tz P7 SPO
vsc GiA
RMg LPGi LSO
MSO
tth RPa SPO
ml
-1 Tz 11
py MVPO

Interaural -1.44 mm bas Bregma 10.44 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 121

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 121
5 6aCb
8 5 2

10 simf
0 Interaural
prf SimA
+15 +10 +5 0 -5 cbw
7 3
SimB

cbw
6 4

psf Crus1
4Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 pcuf 5

pfs

PFl
4 cbw 6
a cbc
un
icp scp
vsc
3 PFl 7
1Cb SuVe GrC
plf
MVePC DCFu LR4V
Fl
cbw Pr DCDp
4V EVe MVeMC
DCMo
2 8vn ocb g7 mlf VCA 8
DCGr
veme asc7
DMSp5 VCP
DPGi

sp5 IRtA PCRtA I8

1 9
ts
7Acs Sp5O
SuS
8cn Gi
7ni
0 P7 A5 10
tz rs GiA 7DM 7VI
LPGiA
vsc RMg 7VM
tth LPGiE LSO
RPa
ml CPO
-1 MVPO 11
py

Interaural -1.56 mm bas

Bregma 10.56 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 7DL 7 nu dorsolat a artery Crus1 crus 1 ansiform Fl flocculus icpd icp decussation
3Cb 3rd Cb lobule 7DM 7 dorsomedial A5 A5 noradr cells DCDp dors coch dp core g7 genu of 7 n IntA interposed ant
4Cb 4th Cb lobule 7L 7 nu lateral asc7 asc fibers of 7 n DCFu dors coch fusiform Gi gigantocell ret nu IRtA intermed rt alpha
4V 4th ventricle 7ni nervus intermed bas basilar artery DCGr dors coch granular GiA gigantocell alpha Lat lat cerebellar nu
5Cb 5th Cb lobule 7VI 7 nu vent intermed cbc cerebellar comm DCMo dors coch molec GrC granule cochlear LPGiA lat paragigcel alph
5Sol trig sol trans zn 7VM 7 nu ventromedial cbw cereb white mat DMSp5 dorsomed sp 5 nu GrCb granule layer LPGiE lat paragigcel ext
6aCb 6a Cb lobule 8cn cochlear root 8 n CI caud interstit mlf DPGi dors paragigantoc I8 interstitial nu 8n LR4V lat recess 4V
7Acs mo 7 accessory 8vn vestib root 8 n CPO caudal perioliv nu EVe nu efferents 8vn icp inf cerebellar ped LSO lat superior olive
LVe lat vestibular nu
RBSC5 Book.qxp 16/9/04 16:36 Page 122

ml medial lemniscus
mlf med long fasc
MoCb molecul layer Cb
MVeMC med vestib magno
MVePC med vestib parvi
MVPO mediovent periol
ocb olivocochl bundle
P7 perifacial zone PPy parapyramidal nu rs rubrospinal tract Sp5O spinal 5 oral VCA vent cochlear ant
PCRtA parvicell ret alpha Pr prepositus nu scp sup cerebellar ped SuS sup salivatory nu VCCap vent cochlear cap
pcuf preculminate fiss prf primary fissure SimA simple lobule A SuVe superior vestib nu VCP vent cochlear post
PFl paraflocculus psf post superior fiss SimB simple lobule B ts tectospinal tract VeCb vestibulocereb nu
pfs paraflocc sulcus py pyramidal tract simf simplex fissure tth trigeminothal tr veme vestibulomes tr
Pk Purkinje cells RMg raphe magnus nu SMV sup medull velum tz trapezoid body vsc vent spinocer tr
plf posterolat fissure RPa raphe pallidus nu sp5 sp trigeminal tr un uncinate fascic

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 122
5 5 6aCb
8 2
simf SimA
prf 5Cb MoCb
0 Interaural 10
Pk
+15 +10 +5 0 -5
GrCb SimB
7 3

cbw
6 psf 4
cbw
Crus1
4Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 pcuf 5
pfs
3Cb
PFl

icpd
4 a IntA 6
cbc Lat
un
scp
VeCb
cbw LR4V
SuVe PFl
3 GrC 7
icp 1Cb MVePC
SMV LVe DCDp
EVe
VCCap
4V Pr Fl
8vn MVeMC
2 g7 CI GrC 8
DCMo
mlf DCFu
veme
5Sol DMSp5 VCP
asc7 DPGi
7ni I8
sp5 PCRtA
1 IRtA 9
Sp5O
7Acs ts Gi SuS
7DL
8cn
P7
rs A5
0 tz 7L 10
GiA 7DM 7VI
LPGiA
RMg
vsc 7VM
tth
ml PPy LPGiE
-1 RPa LSO 11
py

Interaural -1.68 mm bas

Bregma 10.68 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 123

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0

6aCb
Figure 123
5 5 simf SimA
8 2

0 Interaural 10
SimB
+15 +10 +5 0 -5
Crus1
7 3
5Cb
prf

psf

6 cbw 4

4Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 pcuf
5
pfs
IntDL
IntA
a icpd
4 6
vscd
scp Lat
und VeCb
SuVe LR4V
GrC PFl
3 1Cb
7
icp
MVePC LVe
DCFu SMV DCDp
plf VCCap Fl
4V Pr MVeMC
8vn EVe
2 DCMo g7 CI GrC 8
vesp mlf VCPO
DMSp5
asc7 DPGi 5Sol

8cn sp5 Sp5O

IRtA
1 PCRtA I8 9
7ni
ts Gi
7Acs SuS
tz 7DL
rs 7DI A5
0 7L 10
GiA LPGiA 7DM 7VI
vsc RMg 7VM P7
tth
PPy ml RPa
-1 11
py LPGiE

bas
Interaural -1.80 mm Bregma 10.80 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 7DL 7 nu dorsolat A5 A5 noradr cells DCFu dors coch fusiform GrC granule cochlear LatPC lat Cb parvicell
3Cb 3rd Cb lobule 7DM 7 dorsomedial asc7 asc fibers of 7 n DCMo dors coch molec I8 interstitial nu 8n LPGiA lat paragigcel alph
4Cb 4th Cb lobule 7L 7 nu lateral bas basilar artery DMSp5 dorsomed sp 5 nu icp inf cerebellar ped LPGiE lat paragigcel ext
4V 4th ventricle 7ni nervus intermed cbc cerebellar comm DPGi dors paragigantoc icpd icp decussation LR4V lat recess 4V
5Cb 5th Cb lobule 7VI 7 nu vent intermed cbw cereb white mat EVe nu efferents 8vn IntA interposed ant LVe lat vestibular nu
5Sol trig sol trans zn 7VM 7 nu ventromedial chp choroid plexus Fl flocculus IntDL interpos dorsolat
6aCb 6a Cb lobule 8cn cochlear root 8 n CI caud interstit mlf g7 genu of 7 n IntDM interpos dorsomed
7Acs mo 7 accessory 8vn vestib root 8 n Crus1 crus 1 ansiform Gi gigantocell ret nu IRtA intermed rt alpha
7DI 7 dors intermed a artery DCDp dors coch dp core GiA gigantocell alpha Lat lat cerebellar nu
RBSC5 Book.qxp 16/9/04 16:36 Page 124

Med med cerebellar nu
ml medial lemniscus
mlf med long fasc
MVeMC med vestib magno
MVePC med vestib parvi
P7 perifacial zone
PCRtA parvicell ret alpha
pcuf preculminate fiss
PFl paraflocculus
pfs paraflocc sulcus
plf posterolat fissure
PPy parapyramidal nu
Pr prepositus nu
prf primary fissure
psf post superior fiss SimA simple lobule A SuVe superior vestib nu VeCb vestibulocereb nu
py pyramidal tract SimB simple lobule B ts tectospinal tract vesp vest spinal tr
RMg raphe magnus nu simf simplex fissure tth trigeminothal tr vsc vent spinocer tr
RPa raphe pallidus nu SMV sup medull velum tz trapezoid body vscd vsc decussation
rs rubrospinal tract sp5 sp trigeminal tr und uncinate fasc dec Y nu Y
scp sup cerebellar ped Sp5O spinal 5 oral VCCap vent cochlear cap
Sim simple lobule SuS sup salivatory nu VCPO VCP octopus cells

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 124
5 5
8 simf
2
6aCb
0 Interaural 10
Sim
+15 +10 +5 0 -5
7 3
prf 5Cb
Crus1
psf
6 4

cbw
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
pfs
pcuf 4Cb
IntDL
IntDM PFl

icpd 3Cb IntA

4 a 6
Med Lat
scp cbc
und VeCb LatPC
cbw Y
DCMo PFl
3 LVe GrC 7
MVePC DCFu
chp icp 1Cb LR4V
plf DCDp
4V Pr EVe MVeMC
VCCap Fl
2 mlf 8
g7 CI
vesp GrC
asc7 8vn
DPGi
sp5 5Sol DMSp5 VCPO

1 IRtA PCRtA 9
ts Sp5O
Gi
7Acs SuS
tz 7DL
7DI A5
0 rs 10
GiA 7L
LPGiA 7DM 7VI
RMg
vsc PPy 7VM P7
tth
ml
-1 RPa 11
py LPGiE

Interaural -1.92 mm bas

Bregma 10.92 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 125

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
MoCb Figure 125
6aCb Pk
5 5
8 simf GrCb 2

0 Interaural 10 Sim
+15 +10 +5 0 -5
7 3
prf Crus1
psf

6 4
5Cb

Crus1
cbw
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 4Cb 5

pcuf IntDM
pfs
IntA IntDL PFl
4 3Cb 6
Med
scp VeCb Lat
a und LatPC
Y
cbw DCMo
LVe
3 GrC 7
icp 1Cb MVePC
chp LR4V
PFl
4V DCDp
Pr Fl
EVe mlf MVeMC SpVe X VCCap
2 DCFu 8
vesp CI
Sol GrC
DPGi
asc7 5Sol DMSp5 8vn VCPO
sp5
VCP
1 PCRtA 9
IRtA Sp5O
ts 7Acs SuS
tz Gi
7DL
P7 7DI A5
0 rs GiA 10
7VI 7L
LPGiA 7DM
vsc RMg
tth 7VM
PPy ml RPa LPGiE
-1 11
py

bas
Interaural -2.04 mm Bregma 11.04 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 7DI 7 dors intermed a artery Crus1 crus 1 ansiform Fl flocculus IntDM interpos dorsomed
3Cb 3rd Cb lobule 7DL 7 nu dorsolat A5 A5 noradr cells das dors acoustic str Gi gigantocell ret nu IRtA intermed rt alpha
4Cb 4th Cb lobule 7DM 7 dorsomedial asc7 asc fibers of 7 n DCDp dors coch dp core GiA gigantocell alpha Lat lat cerebellar nu
4V 4th ventricle 7L 7 nu lateral bas basilar artery DCFu dors coch fusiform GrC granule cochlear LatPC lat Cb parvicell
5Cb 5th Cb lobule 7VI 7 nu vent intermed cbc cerebellar comm DCMo dors coch molec GrCb granule layer LPGiA lat paragigcel alph
5Sol trig sol trans zn 7VM 7 nu ventromedial cbw cereb white mat DMSp5 dorsomed sp 5 nu icp inf cerebellar ped LPGiE lat paragigcel ext
6aCb 6a Cb lobule 8cn cochlear root 8 n chp choroid plexus DPGi dors paragigantoc IntA interposed ant LR4V lat recess 4V
7Acs mo 7 accessory 8vn vestib root 8 n CI caud interstit mlf EVe nu efferents 8vn IntDL interpos dorsolat LVe lat vestibular nu
RBSC5 Book.qxp 16/9/04 16:36 Page 126

Med med cerebellar nu
ml medial lemniscus
mlf med long fasc
MoCb molecul layer Cb
MVeMC med vestib magno
MVePC med vestib parvi
P7 perifacial zone
PCRtA parvicell ret alpha
pcuf preculminate fiss
PFl paraflocculus
pfs paraflocc sulcus
Pk Purkinje cells
PPy parapyramidal nu
Pr prepositus nu
prf primary fissure Sim simple lobule SpVe spinal vestib nu VCP vent cochlear post
psf post superior fiss simf simplex fissure SuS sup salivatory nu VCPO VCP octopus cells
py pyramidal tract SMV sup medull velum ts tectospinal tract VeCb vestibulocereb nu
RMg raphe magnus nu Sol nu of solitary tr tth trigeminothal tr vesp vest spinal tr
RPa raphe pallidus nu SolIM Sol intermediate tz trapezoid body vsc vent spinocer tr
rs rubrospinal tract sp5 sp trigeminal tr und uncinate fasc dec X nu X
scp sup cerebellar ped Sp5O spinal 5 oral VCCap vent cochlear cap Y nu Y

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 126
5 5 simf
8 2

10 6aCb
0 Interaural Sim
+15 +10 +5 0 -5
7 3
cbw Crus1
prf
psf
5Cb
6 4
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 cbw 5
4Cb
IntDM
pfs
IntDL PFl
IntA
4 3Cb 6
cbw Med Lat
a scp
und VeCb LatPC
GrC cbc
cbw LVe Y DCFu
4V 1Cb DCMo
3 das das
7
chp SMV MVePC
icp LR4V
DCDp
4V
Pr MVeMC SpVe X
8cn mlf
2 SolIM PFl 8
CI
vesp GrC
DPGi asc7 DMSp5
sp5 VCP
5Sol

1 ts PCRtA 9
IRtA Sp5O
Gi SuS

7DL
rs GiA 7DI
0 10
LPGiA 7DM 7L
PPy RMg 7VI
vsc 7VM P7
tth
ml RPa LPGiE
-1 11
py
bas
Interaural -2.16 mm Bregma 11.16 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 127

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
6aCb Figure 127
5 5 simf
8 2

0 Interaural 10 Sim

+15 +10 +5 0 -5
7 3
5Cb Crus1

psf
6 4

5Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
cbw IntDM
MedDL IntDL
pfs
3/4Cb IntA PFl
4 6
scp Med Lat
MedL LVe LatPC
GrC
Y
cbw 1Cb DCFu
3 das 7
und
chp MVePC LR4V
4V DCDp
icp SpVe
Pr MVeMC
mlf X PFl
2 8cn SolRL SolIM DCMo 8
CI
DPGi DMSp5
5Sol VCP

sp5
1 IRtA PCRtA 9
ts Sp5O
Gi
SuS

7DL A5
0 GiA 7DI 10
rs
LPGiA 7DM 7L
RMg 7VI
PPy
vsc tth 7VM P7
ml RPa LPGiE
-1 11
py
bas
Interaural -2.28 mm Bregma 11.28 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

1Cb 1st Cb lobule 7L 7 nu lateral Crus1 crus 1 ansiform GrC granule cochlear LPGiE lat paragigcel ext P7 perifacial zone RPa raphe pallidus nu SpVe spinal vestib nu
3/4Cb 3rd&4th Cb lobs 7VI 7 nu vent intermed das dors acoustic str icp inf cerebellar ped LR4V lat recess 4V PCRtA parvicell ret alpha rs rubrospinal tract SuS sup salivatory nu
4V 4th ventricle 7VM 7 nu ventromedial DCDp dors coch dp core IntA interposed ant LVe lat vestibular nu PFl paraflocculus scp sup cerebellar ped ts tectospinal tract
5Cb 5th Cb lobule 8cn cochlear root 8 n DCFu dors coch fusiform IntDL interpos dorsolat Med med cerebellar nu pfs paraflocc sulcus Sim simple lobule tth trigeminothal tr
5Sol trig sol trans zn A5 A5 noradr cells DCMo dors coch molec IntDM interpos dorsomed MedDL med Cb dorsolat PPy parapyramidal nu simf simplex fissure und uncinate fasc dec
6aCb 6a Cb lobule bas basilar artery DMSp5 dorsomed sp 5 nu IRtA intermed rt alpha ml medial lemniscus Pr prepositus nu SolIM Sol intermediate VCP vent cochlear post
7DI 7 dors intermed cbw cereb white mat DPGi dors paragigantoc Lat lat cerebellar nu mlf med long fasc psf post superior fiss SolRL Sol rolstrolat vsc vent spinocer tr
7DL 7 nu dorsolat chp choroid plexus Gi gigantocell ret nu LatPC lat Cb parvicell MVeMC med vestib magno py pyramidal tract sp5 sp trigeminal tr X nu X
7DM 7 dorsomedial CI caud interstit mlf GiA gigantocell alpha LPGiA lat paragigcel alph MVePC med vestib parvi RMg raphe magnus nu Sp5O spinal 5 oral Y nu Y
RBSC5 Book.qxp 16/9/04 16:36 Page 128

1Cb 1st Cb lobule 7DM 7 dorsomedial das dors acoustic str icp inf cerebellar ped LPGiA lat paragigcel alph MVePC med vestib parvi RMg raphe magnus nu SolRL Sol rolstrolat
3/4Cb 3rd&4th Cb lobs 7L 7 nu lateral DCDp dors coch dp core IntA interposed ant LPGiE lat paragigcel ext PCRt parvicell ret nu RPa raphe pallidus nu sp5 sp trigeminal tr
4V 4th ventricle 7VI 7 nu vent intermed DCFu dors coch fusiform IntDL interpos dorsolat LR4V lat recess 4V PFl paraflocculus rs rubrospinal tract Sp5O spinal 5 oral
5Cb 5th Cb lobule 7VM 7 nu ventromedial DCMo dors coch molec IntP interpos post Med med cerebellar nu pfs paraflocc sulcus scp sup cerebellar ped SpVe spinal vestib nu
5Sol trig sol trans zn bas basilar artery DMSp5 dorsomed sp 5 nu IntPPC interpos pos parv MedDL med Cb dorsolat PPy parapyramidal nu Sim simple lobule ts tectospinal tract
6Cb 6th Cb lobule cbw cereb white mat DPGi dors paragigantoc IRtA intermed rt alpha MedL med Cb lateral Pr prepositus nu simf simplex fissure tth trigeminothal tr
7Acs mo t accessory chp choroid plexus Gi gigantocell ret nu IS inf salivatory nu ml medial lemniscus prf primary fissure SMV sup medull velum und uncinate fasc dec
7DI 7 dors intermed CI caud interstit mlf GiA gigantocell alpha Lat lat cerebellar nu mlf med long fasc psf post superior fiss sol solitary tract vsc vent spinocer tr
7DL 7 nu dorsolat Crus1 crus 1 ansiform GrC granule cochlear LatPC lat Cb parvicell MVeMC med vestib magno py pyramidal tract SolIM Sol intermediate X nu X

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 128
5 5 6Cb
8 simf 2
cbw
0 Interaural 10
Sim
+15 +10 +5 0 -5
7 3
Crus1
prf

psf
6 4
prf
Crus1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
5Cb
cbw

pfs MedDL
3/4Cb IntA IntDL
4 PFl 6
scp und Med IntP Lat
a
MedL IntPPC LatPC
1Cb
chp DCFu
3 SMV 7
GrC
das DCDp
MVePC LR4V
4V SpVe
icp
Pr MVeMC X
2 SolRL SolIM sol DCMo PFl 8
CI
IS
DPGi DMSp5 GrC
5Sol mlf
sp5
1 PCRt 9
ts IRtA
Gi Sp5O

7Acs
7DL
0 GiA 7DI 10
rs 7L
LPGiA 7DM 7VI
RMg
PPy
tth
vsc ml P7
7VM LPGiE
-1 RPa 11
py
bas
Interaural -2.40 mm Bregma 11.40 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 129

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 129
5 5 simf 6aCb
8 2
cbw

0 Interaural 10 Sim

+15 +10 +5 0 -5
7 Crus1 3
6aCb
psf

6 Crus1 4

prf

5Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
cbw
pfs
MedDL

3/4Cb IntA PFl

4 Med IntP IntDL 6
scp plf MedL
10Cb Lat
IntPPC
chp
3 cbw DCFu 7
DCDp DCMo
4V
das MVePC
LR4V PFl
icp SpVe
Pr MVeMC X
2 SolRL sol SolM 8
mlf Mx
DPGi IS
oc DMSp5
SolIM SolV
sp5 5Sol
Sp5O
1 IRtA PCRt 9
ts
Gi

7DI 7Acs Sp5I

7DL
0 GiA
7DM 7VI
10
rs LPGiA 7L
RMg
PPy 7VM
vsc P7
ml RPa LPGiE
-1 11
py
bas

Interaural -2.52 mm Bregma 11.52 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

3/4Cb 3rd&4th Cb lobs 7L 7 nu lateral DCDp dors coch dp core IntDL interpos dorsolat Med med cerebellar nu PCRt parvicell ret nu RMg raphe magnus nu SolRL Sol rolstrolat
4V 4th ventricle 7VI 7 nu vent intermed DCFu dors coch fusiform IntP interpos post MedDL med Cb dorsolat PFl paraflocculus RPa raphe pallidus nu SolV Sol ventral
5Cb 5th Cb lobule 7VM 7 nu ventromedial DCMo dors coch molec IntPPC interpos pos parv ml medial lemniscus pfs paraflocc sulcus rs rubrospinal tract sp5 sp trigeminal tr
5Sol trig sol trans zn 10Cb 10th Cb lobule DMSp5 dorsomed sp 5 nu IRtA intermed rt alpha mlf med long fasc plf posterolat fissure scp sup cerebellar ped Sp5I spinal 5 interpolar
6aCb 6a Cb lobule bas basilar artery DPGi dors paragigantoc IS inf salivatory nu MVeMC med vestib magno PPy parapyramidal nu Sim simple lobule Sp5O spinal 5 oral
7Acs mo t accessory cbw cereb white mat Gi gigantocell ret nu Lat lat cerebellar nu MVePC med vestib parvi Pr prepositus nu simf simplex fissure SpVe spinal vestib nu
7DI 7 dors intermed chp choroid plexus GiA gigantocell alpha LPGiA lat paragigcel alph Mx matrix region prf primary fissure sol solitary tract ts tectospinal tract
7DL 7 nu dorsolat Crus1 crus 1 ansiform icp inf cerebellar ped LPGiE lat paragigcel ext oc olivocerebellar tr psf post superior fiss SolIM Sol intermediate vsc vent spinocer tr
7DM 7 dorsomedial das dors acoustic str IntA interposed ant LR4V lat recess 4V P7 perifacial zone py pyramidal tract SolM Sol medial X nu X
RBSC5 Book.qxp 16/9/04 16:36 Page 130

4V 4th ventricle 7VI 7 nu vent intermed DCDp dors coch dp core IntDM interpos dorsomed Med med cerebellar nu PCRt parvicell ret nu ROb raphe obscurus nu SolRL Sol rolstrolat
5Cb 5th Cb lobule 7VM 7 nu ventromedial DCFu dors coch fusiform IntP interpos post MedDL med Cb dorsolat PFl paraflocculus RPa raphe pallidus nu SolV Sol ventral
5Sol trig sol trans zn 9n glossopharyngeal n DCMo dors coch molec IntPPC interpos pos parv MedL med Cb lateral pfs paraflocc sulcus rs rubrospinal tract sp5 sp trigeminal tr
6aCb 6a Cb lobule 10Cb 10th Cb lobule DMSp5 dorsomed sp 5 nu IRtA intermed rt alpha ml medial lemniscus PPy parapyramidal nu Sim simple lobule Sp5I spinal 5 interpolar
7Acs mo t accessory a artery DPGi dors paragigantoc IS inf salivatory nu mlf med long fasc Pr prepositus nu simf simplex fissure Sp5O spinal 5 oral
7DI 7 dors intermed bas basilar artery Gi gigantocell ret nu Lat lat cerebellar nu MVeMC med vestib magno prf primary fissure sol solitary tract SpVe spinal vestib nu
7DL 7 nu dorsolat cbw cereb white mat GiA gigantocell alpha LPGiA lat paragigcel alph MVePC med vestib parvi psf post superior fiss SolIM Sol intermediate ts tectospinal tract
7DM 7 dorsomedial chp choroid plexus icp inf cerebellar ped LPGiE lat paragigcel ext Mx matrix region py pyramidal tract SolL Sol lateral vsc vent spinocer tr
7L 7 nu lateral Crus1 crus 1 ansiform IntDL interpos dorsolat LR4V lat recess 4V P7 perifacial zone RMg raphe magnus nu SolM Sol medial X nu X

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 130
5 5 simf
8 6aCb 2

10 cbw Sim
0 Interaural
+15 +10 +5 0 -5
7 3
Crus1

psf
6aCb
6 4

Crus1
prf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
5Cb
cbw
MedDL IntDM
pfs
Med IntDL
4 PFl 6
IntP
MedL
Lat
a IntPPC
10Cb
chp DCFu
3 cbw DCMo 7
MVePC DCDp
chp 4V SpVe
icp LR4V
X PFl
sol SolIM Pr MVeMC
2 SolM Mx 8
mlf
9n
SolRL DPGi IS SolL DMSp5
SolV 5Sol
oc Sp5O
sp5
1 ts 9
IRtA PCRt
Gi
7Acs Sp5I
ROb P7
7DL
0 7DM 7DI 10
rs
GiA
RMg 7VI 7L
LPGiA
vsc PPy 7VM P7
ml RPa
-1 LPGiE 11
py
bas
Interaural -2.64 mm Bregma 11.64 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 131

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 131
5 MoCb
8 5 2
Pk
6aCb
0 Interaural 10 GrCb Sim

+15 +10 +5 0 -5 cbw

7 3

psf

6 4
Crus1

prf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
5Cb MedDL

pfs IntDM
Med
4 9Cb IntDL 6
MedL IntP PFl

IntPPC
a 10Cb
DCMo chp 4V
3 cbw 7
DCFu
MVePC SpVe
4V PFl
chp icp X
SolDM Pr MVeMC sol LR4V
SolL
2 mlf 8
Mx
C1
9n IS
sp5 SolM DPGi SolIM DMSp5
5Sol SolV
oc
1 ts PCRt 9
IRtA
Gi Sp5I
7Acs ROb 7DL
7DI
0 7DM 10
rs GiA 7L
RMg LPGiA 7VI
vsc PPy P7
ml RPa LPGiE
-1 11
py
bas

Interaural -2.76 mm Bregma -11.76 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4V 4th ventricle 7VI 7 nu vent intermed Crus1 crus 1 ansiform IntDM interpos dorsomed ml medial lemniscus PFl paraflocculus ROb raphe obscurus nu SolV Sol ventral
5Cb 5th Cb lobule 9Cb 9th Cb lobule DCFu dors coch fusiform IntP interpos post mlf med long fasc pfs paraflocc sulcus RPa raphe pallidus nu sp5 sp trigeminal tr
5Sol trig sol trans zn 9n glossopharyngeal n DCMo dors coch molec IRtA intermed rt alpha MoCb molecul layer Cb Pk Purkinje cells rs rubrospinal tract Sp5I spinal 5 interpolar
6aCb 6a Cb lobule 10Cb 10th Cb lobule DMSp5 dorsomed sp 5 nu IS inf salivatory nu MVeMC med vestib magno PPy parapyramidal nu Sim simple lobule SpVe spinal vestib nu
7Acs mo t accessory a artery DPGi dors paragigantoc LPGiA lat paragigcel alph MVePC med vestib parvi Pr prepositus nu sol solitary tract ts tectospinal tract
7DI 7 dors intermed bas basilar artery Gi gigantocell ret nu LPGiE lat paragigcel ext Mx matrix region prf primary fissure SolDM Sol dorsomedial vsc vent spinocer tr
7DL 7 nu dorsolat C1 C1 adren cells GiA gigantocell alpha LR4V lat recess 4V oc olivocerebellar tr psf post superior fiss SolIM Sol intermediate X nu X
7DM 7 dorsomedial cbw cereb white mat icp inf cerebellar ped Med med cerebellar nu P7 perifacial zone py pyramidal tract SolL Sol lateral
7L 7 nu lateral chp choroid plexus IntDL interpos dorsolat MedDL med Cb dorsolat PCRt parvicell ret nu RMg raphe magnus nu SolM Sol medial
RBSC5 Book.qxp 16/9/04 16:36 Page 132

4V 4th ventricle 9n glossopharyngeal n Crus2 crus 2 ansiform IntDL interpos dorsolat MedDL med Cb dorsolat PFl paraflocculus ROb raphe obscurus nu SolM Sol medial
5Sol trig sol trans zn 10Cb 10th Cb lobule DMSp5 dorsomed sp 5 nu IntP interpos post ml medial lemniscus plf posterolat fissure RPa raphe pallidus nu SolV Sol ventral
6aCb 6a Cb lobule 10n vagus n DPGi dors paragigantoc IRtA intermed rt alpha mlf med long fasc PM paramedian lobule rs rubrospinal tract sp5 sp trigeminal tr
7Acs mo t accessory bas basilar artery EF epifascicular nu IS inf salivatory nu MVeMC med vestib magno PPy parapyramidal nu Sim simple lobule Sp5I spinal 5 interpolar
7DL 7 nu dorsolat C1 C1 adren cells Gi gigantocell ret nu LPGi lat paragig cell MVePC med vestib parvi Pr prepositus nu sol solitary tract SpVe spinal vestib nu
7L 7 nu lateral cbw cereb white mat GiA gigantocell alpha LPGiE lat paragigcel ext Mx matrix region psf post superior fiss SolDM Sol dorsomedial ts tectospinal tract
7VI 7 nu vent intermed chp choroid plexus icf intercrural fissure LR4V lat recess 4V oc olivocerebellar tr py pyramidal tract SolIM Sol intermediate vsc vent spinocer tr
9Cb 9th Cb lobule Crus1 crus 1 ansiform icp inf cerebellar ped Med med cerebellar nu PCRt parvicell ret nu RMg raphe magnus nu SolL Sol lateral X nu X

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 132
5 5
8 2
6aCb
10 Sim
0 Interaural
+15 +10 +5 0 -5
7 psf 3
cbw

Crus1
6aCb

6 4

6aCb
cbw
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
Crus1

icf MedDL

cbw Crus2
Med
4 IntDL 6
plf PFl
9Cb IntP
PM

10Cb
3 7

cbw PFl
chp icp MVePC SpVe
4V X LR4V
sol SolDM MVeMC
9n SolM Pr
2 EF IS Mx 8
mlf SolL
oc DPGi 5Sol DMSp5
SolIM SolV
sp5
C1
PCRt
1 10n IRtA 9
Sp5I
ts Gi
7Acs
ROb
7DL
0 10
rs
GiA 7VI 7L
RMg LPGi
vsc PPy
ml LPGiE
-1 RPa 11
py
bas
Interaural -2.88 mm Bregma -11.88 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 133

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 133
5 5
8 2
psf
6aCb
0 Interaural 10
Sim
+15 +10 +5 0 -5 Crus1
7 3

cbw
6 4
cbw

6Cb Crus1
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5

icf MedCM
Med
9Cb Crus2
4 6
9Cb PFl
pfs IntP
plf PM

10Cb
3 7

cbw MVePC SpVe

icp MVeMC X PFl
chp SolDM 4V SolIM LR4V
sol
Pr SolM
2 EF Mx 8
mlf
IS
sp5 SolL C1 DMSp5
5Sol SolV
oc
1 ts PCRt 9
10n IRt Sp5I
Gi
AmbC
ROb
P7
0 Bo 10
GiA 7L
rs RMg
C1 LPGi RVL
RPa
vsc
ml LPGiE
-1 11
py IOPr
bas

Interaural -3.00 mm Bregma -12.00 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4V 4th ventricle AmbC ambig compact DMSp5 dorsomed sp 5 nu IRt intermed ret nu mlf med long fasc pfs paraflocc sulcus RPa raphe pallidus nu SolRL Sol rolstrolat
5Sol trig sol trans zn bas basilar artery EF epifascicular nu IS inf salivatory nu MVeMC med vestib magno plf posterolat fissure rs rubrospinal tract SolV Sol ventral
6aCb 6a Cb lobule Bo Botzinger complex Gi gigantocell ret nu LPGi lat paragig cell MVePC med vestib parvi PM paramedian lobule RVL rostroventrolat rt sp5 sp trigeminal tr
6Cb 6th Cb lobule C1 C1 adren cells GiA gigantocell alpha LPGiE lat paragigcel ext Mx matrix region Pr prepositus nu Sim simple lobule Sp5I spinal 5 interpolar
7L 7 nu lateral cbw cereb white mat icf intercrural fissure LR4V lat recess 4V oc olivocerebellar tr psf post superior fiss sol solitary tract SpVe spinal vestib nu
9Cb 9th Cb lobule chp choroid plexus icp inf cerebellar ped Med med cerebellar nu P7 perifacial zone py pyramidal tract SolDM Sol dorsomedial ts tectospinal tract
10Cb 10th Cb lobule Crus1 crus 1 ansiform IntP interpos post MedCM med Cb caudomed PCRt parvicell ret nu RMg raphe magnus nu SolIM Sol intermediate vsc vent spinocer tr
10n vagus n Crus2 crus 2 ansiform IOPr IO principal nu ml medial lemniscus PFl paraflocculus ROb raphe obscurus nu SolM Sol medial X nu X
RBSC5 Book.qxp 16/9/04 16:36 Page 134

3Cb 3rd Cb lobule Bo Botzinger complex GiV gigantocell vent LPGi lat paragig cell oc olivocerebellar tr psf post superior fiss SolDM Sol dorsomedial ts tectospinal tract
4V 4th ventricle C1 C1 adren cells icf intercrural fissure LPGiE lat paragigcel ext P7 perifacial zone py pyramidal tract SolIM Sol intermediate vsc vent spinocer tr
5Sol trig sol trans zn cbw cereb white mat icp inf cerebellar ped LR4V lat recess 4V PCRt parvicell ret nu RMg raphe magnus nu SolL Sol lateral X nu X
6bCb 6b Cb lobule chp choroid plexus IOD IO dorsal nu ml medial lemniscus PFl paraflocculus ROb raphe obscurus nu SolM Sol medial
6cCb 6c Cb lobule Crus1 crus 1 ansiform IOM IO medial nu mlf med long fasc pfs paraflocc sulcus RPa raphe pallidus nu SolRL Sol rostrolat
7L 7 nu lateral Crus2 crus 2 ansiform IOPr IO principal nu MoCb molecul layer Cb Pk Purkinje cells rs rubrospinal tract SolV Sol ventral
10Cb 10th Cb lobule DMSp5 dorsomed sp 5 nu IRt intermed ret nu MVeMC med vestib magno plf posterolat fissure RVL rostroventrolat rt sp5 sp trigeminal tr
AmbC ambig compact EF epifascicular nu IS inf salivatory nu MVePC med vestib parvi PM paramedian lobule Sim simple lobule Sp5I spinal 5 interpolar
bas basilar artery Gi gigantocell ret nu JxO juxtaolivary nu Mx matrix region Pr prepositus nu sol solitary tract SpVe spinal vestib nu

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 134
5 5 6bCb
8 2
Sim
0 Interaural 10 Crus1
+15 +10 +5 0 -5 psf
6cCb MoCb
7 3
Pk
GrCb

6 4
cbw
Crus1

icf 3Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
Crus2

pfs
4 6
PFl
plf PM

cbw
10Cb
3 cbw 7
chp SpVe X
icp MVePC LR4V
SolIM SolDM SolM 4V MVeMC
sol Mx
Pr SolL
IS
2 SolV mlf EF SolRL 8
5Sol
oc DMSp5
sp5
PCRt
IRt Sp5I
10n
1 ts 9
Gi AmbC
ROb

rs Bo
GiV 7L
0 RVL
P7 10
IOM RMg LPGi
vsc C1
JxO
LPGiE
ml RPa
IOD
-1 py IOPr 11
bas

Interaural -3.12 mm Bregma -12.12 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 135

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 135
5 5
8 6bCb Sim 2
psf
0 Interaural 10

+15 +10 +5 0 -5
6cCb Crus1
7 3

cbw
cbw
6 4
8Cb

icf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
8Cb Crus2

pfs
4 cbw 6
cbw plf 9Cb PM
9Cb Cop PFl

3 10Cb 7
chp SpVe ECu
icp MVePC
SolIM SolDM MVeMC X LR4V
4V
sol Pr Mx
IS SolL
sp5 C3
2 8
mlf Ro
SolM 5Sol DMSp5
SolRL SolV
SolVL
oc IRt
Li Sp5I
ts Li PCRt
1 10n 9
Gi
AmbC
ROb

rs GiV Bo
0 RPa 10
C1 LPGi RVL
JxO
vsc
LPGiE

py ml IOM
-1 IOPr IOD 11
bas

Interaural -3.24 mm Bregma -12.24 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

V 4th ventricle bas basilar artery DMSp5 dorsomed sp 5 nu IRt intermed ret nu MVeMC med vestib magno Pr prepositus nu sol solitary tract SpVe spinal vestib nu
5Sol trig sol trans zn Bo Botzinger complex ECu ext cuneate nu IS inf salivatory nu MVePC med vestib parvi psf post superior fiss SolDM Sol dorsomedial ts tectospinal tract
6bCb 6b Cb lobule C1 C1 adren cells Gi gigantocell ret nu JxO juxtaolivary nu Mx matrix region py pyramidal tract SolIM Sol intermediate vsc vent spinocer tr
6cCb 6c Cb lobule C3 C3 adren cells GiV gigantocell vent Li linear nu oc olivocerebellar tr Ro nu of Roller SolL Sol lateral X nu X
8Cb 8th Cb lobule cbw cereb white mat icf intercrural fissure LPGi lat paragig cell PCRt parvicell ret nu ROb raphe obscurus nu SolM Sol medial
9Cb 9th Cb lobule chp choroid plexus icp inf cerebellar ped LPGiE lat paragigcel ext PFl paraflocculus RPa raphe pallidus nu SolV Sol ventral
10Cb 10th Cb lobule Cop copula of pyramis IOD IO dorsal nu LR4V lat recess 4V pfs paraflocc sulcus rs rubrospinal tract SolVL Sol ventrolat
10n vagus n Crus1 crus 1 ansiform IOM IO medial nu ml medial lemniscus plf posterolat fissure RVL rostroventrolat rt sp5 sp trigeminal tr
AmbC ambig compact Crus2 crus 2 ansiform IOPr IO principal nu mlf med long fasc PM paramedian lobule Sim simple lobule Sp5I spinal 5 interpolar
RBSC5 Book.qxp 16/9/04 16:36 Page 136

4V 4th ventricle AmbC ambig compact DMSp5 dorsomed sp 5 nu IRt intermed ret nu MVePC med vestib parvi py pyramidal tract SolIM Sol intermediate ts tectospinal tract
5Sol trig sol trans zn Bo Botzinger complex ECu ext cuneate nu JxO juxtaolivary nu Mx matrix region Ro nu of Roller SolL Sol lateral vsc vent spinocer tr
6bCb 6b Cb lobule C1 C1 adren cells Gi gigantocell ret nu Li linear nu oc olivocerebellar tr ROb raphe obscurus nu SolM Sol medial X nu X
6cCb 6c Cb lobule C3 C3 adren cells GiV gigantocell vent LPGi lat paragig cell PCRt parvicell ret nu RPa raphe pallidus nu SolRL Sol rostrolat
8Cb 8th Cb lobule cbw cereb white mat icf intercrural fissure LPGiE lat paragigcel ext PFl paraflocculus rs rubrospinal tract SolV Sol ventral
9Cb 9th Cb lobule chp choroid plexus icp inf cerebellar ped LR4V lat recess 4V plf posterolat fissure RVL rostroventrolat rt SolVL Sol ventrolat
10Cb 10th Cb lobule Cop copula of pyramis IOD IO dorsal nu ml medial lemniscus PM paramedian lobule sf 2ary fissure sp5 sp trigeminal tr
10N dorsal mo nu 10 Crus1 crus 1 ansiform IOM IO medial nu mlf med long fasc pms paramedian sulcus sol solitary tract Sp5I spinal 5 interpolar
10n vagus n Crus2 crus 2 ansiform IOPr IO principal nu MVeMC med vestib magno Pr prepositus nu SolDM Sol dorsomedial SpVe spinal vestib nu

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 136
5 5
8 pms 2
6bCb

0 Interaural 10

+15 +10 +5 0 -5
Crus1
7 6cCb
3

cbw

6 4
Crus1

icf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 8Cb 5
Crus2
sf
cbw
cbw cbw
4 9Cb 6

plf PM PFl
Cop

10Cb MVeMC
3 7
SpVe ECu
chp chp MVePC X
icp LR4V
sol 4V Mx
Pr SolM SolIM SolL
SolDM
2 SolV C3 SolRL SolVL DMSp5 8
Ro 10N 5Sol
mlf
sp5
IRt
ts PCRt
Li Sp5I
oc ROb Li
1 9
10n
Gi AmbC

rs Bo
GiV
0 10
RVL
IOD JxO LPGi C1
vsc LPGiE
RPa IOM
py ml IOPr
-1 11

Interaural -3.36 mm Bregma -12.36 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 137

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 137
5 5
8 6bCb 2
pms cbw
0 Interaural 10

+15 +10 +5 0 -5
7 6cCb 3

cbw
7Cb Crus1
6 icf 4

8Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
Crus2
sf 9Cb

ppf cbw
4 6
plf

10Cb PM
Cop PFl
3 7
SpVe ECu

C
PC

M
icp SolIM SolDM chp FVe X

Ve

Ve
chp LR4V

M
sol 4V Mx
Pr SolL
SolV C3 DMSp5
2 sp5 5Sol 8
mlf Ro 10N SolRL
dsc/oc SolM SolVL
IRt PCRt
Li Sp5I
1 ts 9
Gi AmbC
ROb
12n
10n
rs GiV Bo
0 RVL 10
LPGi C1
vsc JxO

IOD
ml py RPa IOM
-1 asp IOPr 11
vert

Interaural -3.48 mm Bregma -12.48 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4V 4th ventricle 10n vagus n Cop copula of pyramis icf intercrural fissure LR4V lat recess 4V pms paramedian sulcus sol solitary tract vert vertebral artery
5Sol trig sol trans zn 12n root of 12n Crus1 crus 1 ansiform icp inf cerebellar ped ml medial lemniscus ppf prepyramidal fiss SolDM Sol dorsomedial vsc vent spinocer tr
6bCb 6b Cb lobule AmbC ambig compact Crus2 crus 2 ansiform IOD IO dorsal nu mlf med long fasc Pr prepositus nu SolIM Sol intermediate X nu X
6cCb 6c Cb lobule asp ant spinal artery DMSp5 dorsomed sp 5 nu IOM IO medial nu MVePC med vestib parvi py pyramidal tract SolL Sol lateral
7Cb 7th Cb lobule Bo Botzinger complex dsc/oc dors sp cer/ol cer IOPr IO principal nu Mx matrix region Ro nu of Roller SolM Sol medial
8Cb 8th Cb lobule C1 C1 adren cells ECu ext cuneate nu IRt intermed ret nu PCRt parvicell ret nu ROb raphe obscurus nu sp5 sp trigeminal tr
9Cb 9th Cb lobule C3 C3 adren cells FVe F vestib group JxO juxtaolivary nu PFl paraflocculus RPa raphe pallidus nu Sp5I spinal 5 interpolar
10Cb 10th Cb lobule cbw cereb white mat Gi gigantocell ret nu Li linear nu plf posterolat fissure rs rubrospinal tract SpVe spinal vestib nu
10N dorsal mo nu 10 chp choroid plexus GiV gigantocell vent LPGi lat paragig cell PM paramedian lobule RVL rostroventrolat rt ts tectospinal tract
RBSC5 Book.qxp 16/9/04 16:36 Page 138

4V 4th ventricle AmbC ambig compact DMSp5 dorsomed sp 5 nu IOM IO medial nu MVeMC med vestib magno PrBo pre-Botzinger com SolIM Sol intermediate vsc vent spinocer tr
5Sol trig sol trans zn C1 C1 adren cells dsc/oc dors sp cer/ol cer IOPr IO principal nu MVePC med vestib parvi py pyramidal tract SolL Sol lateral X nu X
6bCb 6b Cb lobule C2 C2 adren cells ECu ext cuneate nu IRt intermed ret nu Mx matrix region Ro nu of Roller SolM Sol medial
7Cb 7th Cb lobule cbw cereb white mat FVe F vestib group JxO juxtaolivary nu PCRt parvicell ret nu ROb raphe obscurus nu SolV Sol ventral
8Cb 8th Cb lobule chp choroid plexus Gi gigantocell ret nu Li linear nu plf posterolat fissure RPa raphe pallidus nu SolVL Sol ventrolat
9Cb 9th Cb lobule Cop copula of pyramis GiV gigantocell vent LPGi lat paragig cell PM paramedian lobule rs rubrospinal tract sp5 sp trigeminal tr
10Cb 10th Cb lobule Crus1 crus 1 ansiform icf intercrural fissure LR4V lat recess 4V pms paramedian sulcus sf 2ary fissure Sp5I spinal 5 interpolar
10N dorsal mo nu 10 Crus2 crus 2 ansiform icp inf cerebellar ped ml medial lemniscus ppf prepyramidal fiss sol solitary tract SpVe spinal vestib nu
12N hypoglossal nu CVL caudoventrolat rt IOD IO dorsal nu mlf med long fasc Pr prepositus nu SolDM Sol dorsomedial ts tectospinal tract

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 138
5 5
8 6bCb
2
pms
0 Interaural 10

+15 +10 +5 0 -5 cbw

7 3
Crus1

icf
6 4
7Cb
ppf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 Crus2 5
Crus2
8Cb
sf
cbw
cbw
4 6
9Cb
plf
PM

cbw 10Cb
3 MVeMC Cop 7
ECu
MVePC SpVe FVe
chp X
chp icp LR4V
SolL sol 4V Mx
SolDM Pr C2 SolM
10N SolIM DMSp5
2 SolV 12N 5Sol 8
SolVL mlf Ro
sp5
dsc/oc 12N
IRt
Li PCRt Sp5I
1 ts 9
ROb Gi AmbC
Li
rs PrBo
0 GiV 10
RPa
C1 LPGi CVL
vsc JxO
IOM
IOD
ml
-1 py IOPr 11

Interaural -3.60 mm Bregma -12.60 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 139

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 139
5 5
8 2
pms
0 Interaural 10

+15 +10 +5 0 -5 6cCb

7 Crus1 3
psf
icf
cbw
7Cb
6 4
ppf

Crus2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
8Cb
sf
cbw

ppf cbw
4 apmf 6
9Cb
plf
PM

3 10Cb Cop 7
SolM SpVe ECu
icp MVe FVe X
sol SolDM
chp LR4V
SolL 4V 10N Mx
SolIM Pr DMSp5
sp5 SolV
2 12N 5Sol 8
C2
SolVL
dsc/oc Ro
mlf PCRt Sp5I
IRt
10n Li
1 PMn 9
Li

ts ROb Gi AmbC
PrBo
rs GiV
0 JxO CVL 10
vsc C1
LPGi
IOM
py ml RPa IOD
asp IOPr
-1 11
vert

Interaural -3.72 mm Bregma -12.72 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4V 4th ventricle 12N hypoglossal nu Crus1 crus 1 ansiform icf intercrural fissure LR4V lat recess 4V pms paramedian sulcus sf 2ary fissure Sp5I spinal 5 interpolar
5Sol trig sol trans zn AmbC ambig compact Crus2 crus 2 ansiform icp inf cerebellar ped ml medial lemniscus ppf prepyramidal fiss sol solitary tract SpVe spinal vestib nu
6cCb 6c Cb lobule apmf ansoparamed fiss CVL caudoventrolat rt IOD IO dorsal nu mlf med long fasc Pr prepositus nu SolDM Sol dorsomedial vert vertebral artery
7Cb 7th Cb lobule asp ant spinal artery DMSp5 dorsomed sp 5 nu IOM IO medial nu MVe med vestibular nu PrBo pre-Botzinger com SolIM Sol intermediate vsc vent spinocer tr
8Cb 8th Cb lobule C1 C1 adren cells dsc/oc dors sp cer/ol cer IOPr IO principal nu Mx matrix region py pyramidal tract SolL Sol lateral X nu X
9Cb 9th Cb lobule C2 C2 adren cells ECu ext cuneate nu IRt intermed ret nu PCRt parvicell ret nu Ro nu of Roller SolM Sol medial
10Cb 10th Cb lobule cbw cereb white mat FVe F vestib group JxO juxtaolivary nu plf posterolat fissure ROb raphe obscurus nu SolV Sol ventral
10N dorsal mo nu 10 chp choroid plexus Gi gigantocell ret nu Li linear nu PM paramedian lobule RPa raphe pallidus nu SolVL Sol ventrolat
10n vagus n Cop copula of pyramis GiV gigantocell vent LPGi lat paragig cell PMn paramedian ret nu rs rubrospinal tract sp5 sp trigeminal tr
RBSC5 Book.qxp 16/9/04 16:36 Page 140

4V 4th ventricle 12N hypoglossal nu Crus2 crus 2 ansiform icf intercrural fissure LPGi lat paragig cell pms paramedian sulcus rs rubrospinal tract sp5 sp trigeminal tr
5Sol trig sol trans zn AmbC ambig compact Cu cuneate nu icp inf cerebellar ped ml medial lemniscus ppf prepyramidal fiss sf 2ary fissure Sp5I spinal 5 interpolar
6cCb 6c Cb lobule apmf ansoparamed fiss CVL caudoventrolat rt In intercalated nu mlf med long fasc Pr prepositus nu sol solitary tract SpVe spinal vestib nu
7Cb 7th Cb lobule asp ant spinal artery DMSp5 dorsomed sp 5 nu IOD IO dorsal nu MVe med vestibular nu PrBo pre-Botzinger com SolDM Sol dorsomedial ts tectospinal tract
8Cb 8th Cb lobule C1 C1 adren cells dsc/oc dors sp cer/ol cer IOM IO medial nu Mx matrix region psf post superior fiss SolIM Sol intermediate vert vertebral artery
9a,bCb 9th Cb lobule, a&b cbw cereb white mat ECu ext cuneate nu IOPr IO principal nu PCRt parvicell ret nu py pyramidal tract SolL Sol lateral vsc vent spinocer tr
9cCb 9th Cb lobule, c chp choroid plexus FVe F vestib group IRt intermed ret nu plf posterolat fissure Ro nu of Roller SolM Sol medial Z nu Z
10Cb 10th Cb lobule Cop copula of pyramis Gi gigantocell ret nu JxO juxtaolivary nu PM paramedian lobule ROb raphe obscurus nu SolV Sol ventral
10N dorsal mo nu 10 Crus1 crus 1 ansiform GiV gigantocell vent Li linear nu PMn paramedian ret nu RPa raphe pallidus nu SolVL Sol ventrolat

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 140
5 5
8 2

10 pms cbw
0 Interaural 6cCb
+15 +10 +5 0 -5 Crus1
7 icf 3
psf
icf

7Cb Crus2
6 4
ppf

8Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
Crus2
cbw sf
9a,bCb

4 6
plf PM
9cCb
apmf cbw

cbw 10Cb
3 Cop 7
ECu
Pr MVe SpVe
icp Z chp FVe Cu
sol SolDM 4V SolM
SolIM SolL Mx
sp5 SolVL 10N SolV
2 12N In 5Sol DMSp5 8
Ro
mlf PCRt
dsc/oc IRt
Li
Sp5I
PMn
1 Li 9
AmbC
ts Gi Li
ROb
PrBo
rs
0 C1 RPa GiV 10
CVL
vsc JxO LPGi

IOD
py IOM ml IOPr
-1 asp 11
vert

Interaural -3.84 mm Bregma -12.84 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 141

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 141
5 5
8 2

0 Interaural 10 6cCb

+15 +10 +5 0 -5 Crus1

psf
7 pms 3

7Cb

6 ppf
4

cbw Crus2
8Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 cbw 5
sf
9a,bCb
apmf
4 6
plf 9cCb
PM

3 cbw 10Cb ECu 7

icp Z MVe Cop
SpVe Cu
chp SolDL Mx Pa5
sp5 sol SolI C2
4V SolM
SolVL 10N SolIM SolL 5Sol
SolV
2 12N In 8

dsc/oc
mlf Ro PCRt Sp5I
IRt Li
1 9
AmbSC 12n ts PMn Gi Li
ROb
RVRG
rs
10n GiV CVL
C1
0 10
vsc JxO LPGi

IOM IOD
py ml RPa IOPr
-1 asp 11
vert

Interaural -3.96 mm Bregma -12.96 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4V 4th ventricle 10n vagus n chp choroid plexus GiV gigantocell vent LPGi lat paragig cell PMn paramedian ret nu RVRG rostral ventral rsp SolVL Sol ventrolat
5Sol trig sol trans zn 12N hypoglossal nu Cop copula of pyramis icp inf cerebellar ped ml medial lemniscus pms paramedian sulcus sf 2ary fissure sp5 sp trigeminal tr
6cCb 6c Cb lobule 12n root of 12n Crus1 crus 1 ansiform In intercalated nu mlf med long fasc ppf prepyramidal fiss sol solitary tract Sp5I spinal 5 interpolar
7Cb 7th Cb lobule AmbSC Crus2 crus 2 ansiform IOD IO dorsal nu MVe med vestibular nu psf post superior fiss SolDL Sol dorsolat SpVe spinal vestib nu
8Cb 8th Cb lobule apmf ansoparamed fiss Cu cuneate nu IOM IO medial nu Mx matrix region py pyramidal tract SolI Sol interstitial ts tectospinal tract
9a,bCb 9th Cb lobule, a&b asp ant spinal artery CVL caudoventrolat rt IOPr IO principal nu Pa5 paratrigeminal nu Ro nu of Roller SolIM Sol intermediate vert vertebral artery
9cCb 9th Cb lobule, c C1 C1 adren cells dsc/oc dors sp cer/ol cer IRt intermed ret nu PCRt parvicell ret nu ROb raphe obscurus nu SolL Sol lateral vsc vent spinocer tr
10Cb 10th Cb lobule C2 C2 adren cells ECu ext cuneate nu JxO juxtaolivary nu plf posterolat fissure RPa raphe pallidus nu SolM Sol medial Z nu Z
10N dorsal mo nu 10 cbw cereb white mat Gi gigantocell ret nu Li linear nu PM paramedian lobule rs rubrospinal tract SolV Sol ventral
RBSC5 Book.qxp 16/9/04 16:36 Page 142

4V 4th ventricle 10n vagus n Cop copula of pyramis icp inf cerebellar ped LRt lat reticular nu PMn paramedian ret nu rs rubrospinal tract SolV Sol ventral
5Sol trig sol trans zn 12N hypoglossal nu Crus2 crus 2 ansiform InM intmed of medulla ml medial lemniscus pms paramedian sulcus RVRG rostral ventral rsp SolVL Sol ventrolat
6cCb 6c Cb lobule AmbSL Cu cuneate nu IOD IO dorsal nu mlf med long fasc ppf prepyramidal fiss sf 2ary fissure sp5 sp trigeminal tr
7Cb 7th Cb lobule apmf ansoparamed fiss CVL caudoventrolat rt IODM IO dorsomedial MVe med vestibular nu psf post superior fiss sol solitary tract Sp5I spinal 5 interpolar
8Cb 8th Cb lobule asp ant spinal artery DPGi dors paragigantoc IOM IO medial nu Mx matrix region PSol parasolitary nu SolDL Sol dorsolat SpVe spinal vestib nu
9a,bCb 9th Cb lobule, a&b C1 C1 adren cells dsc/oc dors sp cer/ol cer IOPr IO principal nu Pa5 paratrigeminal nu py pyramidal tract SolI Sol interstitial ts tectospinal tract
9cCb 9th Cb lobule, c C2 C2 adren cells ECu ext cuneate nu IRt intermed ret nu PCRt parvicell ret nu Ro nu of Roller SolIM Sol intermediate vert vertebral artery
10Cb 10th Cb lobule cbw cereb white mat Gi gigantocell ret nu Li linear nu plf posterolat fissure ROb raphe obscurus nu SolL Sol lateral vsc vent spinocer tr
10N dorsal mo nu 10 chp choroid plexus GiV gigantocell vent LPGi lat paragig cell PM paramedian lobule RPa raphe pallidus nu SolM Sol medial Z nu Z

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 142
5 5
8 2

6cCb
0 Interaural 10

+15 +10 +5 0 -5
pms psf
7 3

7Cb
cbw
6 ppf Crus2 4

8Cb
cbw
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
sf

9a,bCb

4 apmf 6
plf 9cCb

pms 10Cb Cop PM

3 cbw ECu 7
icp Z MVe
PSol SpVe Cu
sol SolI chp C2 SolDL Mx Pa5
4V SolM SolL
SolIM
sp5 SolVL SolV 5Sol
2 InM 8
12N 10N
dsc/oc Sp5I
mlf Ro
PCRt
IRt

1 AmbSC Li 9
10n Gi
ts PMn
ROb RVRG
rs
GiV CVL
C1
0 vsc
10
LRt LPGi

IODM IOM IOD

py RPa ml IOPr
-1 asp 11
vert

Interaural -4.08 mm Bregma -13.08 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 143

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 143
5 5
8 2

0 Interaural 10
6cCb
+15 +10 +5 0 -5
7 3
psf

7Cb
6 cbw 4
ppf
Crus2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 cbw 5
8Cb

sf
9a,bCb
4 apmf cbw 6
9cCb
plf
PM
Cop
3 7
10Cb ECu
icp PSol
cu SolIM Cu
SolCe SolDL Pa5
SolL C2 chp SolM Mx E5
sp5 4V
2 dsc/oc SolI 5Sol 8
sol 12N SolV
SolVL
InM 10N Sp5I
Ro
IRt PCRt
mlf
AmbSC
1 9
PMn
Gi Li
LRt ROb
rs ts RVRG
10n
C1 GiV CVL
0 vsc IODM LRtS5 10
LRt

IOM
IOD
py ml IOPr
-1 RPa 11
vert

Interaural -4.20 mm Bregma -13.20 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4V 4th ventricle 12N hypoglossal nu Cu cuneate nu InM intmed of medulla ml medial lemniscus psf post superior fiss sol solitary tract sp5 sp trigeminal tr
5Sol trig sol trans zn AmbSC amb subcomp cu cuneate fasciculus IOD IO dorsal nu mlf med long fasc PSol parasolitary nu SolCe Sol central Sp5I spinal 5 interpolar
6cCb 6c Cb lobule apmf ansoparamed fiss CVL caudoventrolat rt IODM IO dorsomedial Mx matrix region py pyramidal tract SolDL Sol dorsolat ts tectospinal tract
7Cb 7th Cb lobule C1 C1 adren cells dsc/oc dors sp cer/ol cer IOM IO medial nu Pa5 paratrigeminal nu Ro nu of Roller SolI Sol interstitial vert vertebral artery
8Cb 8th Cb lobule C2 C2 adren cells E5 ectotrigeminal nu IOPr IO principal nu PCRt parvicell ret nu ROb raphe obscurus nu SolIM Sol intermediate vsc vent spinocer tr
9a,bCb 9th Cb lobule, a&b cbw cereb white mat ECu ext cuneate nu IRt intermed ret nu plf posterolat fissure RPa raphe pallidus nu SolL Sol lateral
9cCb 9th Cb lobule, c chp choroid plexus Gi gigantocell ret nu Li linear nu PM paramedian lobule rs rubrospinal tract SolM Sol medial
10Cb 10th Cb lobule Cop copula of pyramis GiV gigantocell vent LRt lat reticular nu PMn paramedian ret nu RVRG rostral ventral rsp SolV Sol ventral
10n vagus n Crus2 crus 2 ansiform icp inf cerebellar ped LRtS5 lat retic sub5 ppf prepyramidal fiss sf 2ary fissure SolVL Sol ventrolat
RBSC5 Book.qxp 16/9/04 16:36 Page 144

4V 4th ventricle 12N hypoglossal nu Crus2 crus 2 ansiform Gr gracile nu LRtS5 lat retic sub5 PMn paramedian ret nu rs rubrospinal tract SolM Sol medial
5Sol trig sol trans zn 12n root of 12n Cu cuneate nu InM intmed of medulla ml medial lemniscus pms paramedian sulcus RVRG rostral ventral rsp SolV Sol ventral
7Cb 7th Cb lobule AmbSC amb subcomp cu cuneate fasciculus IOD IO dorsal nu mlf med long fasc ppf prepyramidal fiss sf 2ary fissure SolVL Sol ventrolat
8Cb 8th Cb lobule apmf ansoparamed fiss CVL caudoventrolat rt IOM IO medial nu Mx matrix region PSol parasolitary nu sol solitary tract sp5 sp trigeminal tr
9a,bCb 9th Cb lobule, a&b asp ant spinal artery dsc/oc dors sp cer/ol cer IOPr IO principal nu Pa5 paratrigeminal nu py pyramidal tract SolCe Sol central Sp5I spinal 5 interpolar
9cCb 9th Cb lobule, c C1 C1 adren cells E5 ectotrigeminal nu IOVL IO ventrolat PCRt parvicell ret nu Ro nu of Roller SolDL Sol dorsolat ts tectospinal tract
10Cb 10th Cb lobule cbw cereb white mat ECu ext cuneate nu IRt intermed ret nu plf posterolat fissure ROb raphe obscurus nu SolG Sol gelatinous vert vertebral artery
10N dorsal mo nu 10 Cop copula of pyramis Gi gigantocell ret nu LRt lat reticular nu PM paramedian lobule RPa raphe pallidus nu SolIM Sol intermediate vsc vent spinocer tr

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 144
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
7 pms 3
7Cb
cbw

6 ppf 4
Crus2

cbw
cbw 8Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 apmf 5
sf
9a,bCb

4 cbw 6
PM
9cCb
plf Cop

3 cu 7
10Cb ECu
Gr
SolG Cu
PSol SolI Pa5
SolDL10N 4V SolM sol
dsc/oc sp5 E5
2 SolCe SolV Mx 5Sol 8
SolIM InM SolVL
12N Sp5I
mlf Ro IRt PCRt
1 9
AmbSC 12n LRtS5
Gi
ts PMn
rs ROb RVRG
C1 CVL
0 vsc IOVL 10
LRt
LRt
RPa IOM IOD

-1 py ml IOPr 11
asp
vert

Interaural -4.32 mm Bregma -13.32 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 145

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 145
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
7 pms 7Cb
3

cbw
Crus2

6 ppf 4

8Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
sf
apmf

9a,bCb
cbw
4 cbw 6
9cCb
plf PM
Cop

ECu
3 10Cb 7
cu PSol Cu
Gr Pa5
sol SolG chp SolM SolDL Mx
SolCe 4V 10N
sp5 SolVL SolI SolIM
2 SolV 5Sol 8
12N Sp5I
InM
Ro PCRt
IRt
mlf
1 9
dsc/oc
AmbSC Gi
rs ROb RVRG
ts
C1 CVL

vsc IOVL IRt LRt LRtS5

0 LRt 10

ml IOM IOD
py RPa IOPr
-1 asp 11
vert

Interaural -4.44 mm Bregma -13.44 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

4V 4th ventricle 12N hypoglossal nu Crus2 crus 2 ansiform InM intmed of medulla ml medial lemniscus ppf prepyramidal fiss sol solitary tract SolVL Sol ventrolat
5Sol trig sol trans zn AmbSC amb subcomp Cu cuneate nu IOD IO dorsal nu mlf med long fasc PSol parasolitary nu SolCe Sol central sp5 sp trigeminal tr
7Cb 7th Cb lobule apmf ansoparamed fiss cu cuneate fasciculus IOM IO medial nu Mx matrix region py pyramidal tract SolDL Sol dorsolat Sp5I spinal 5 interpolar
8Cb 8th Cb lobule asp ant spinal artery CVL caudoventrolat rt IOPr IO principal nu Pa5 paratrigeminal nu Ro nu of Roller SolG Sol gelatinous ts tectospinal tract
9a,bCb 9th Cb lobule, a&b C1 C1 adren cells dsc/oc dors sp cer/ol cer IOVL IO ventrolat PCRt parvicell ret nu ROb raphe obscurus nu SolI Sol interstitial vert vertebral artery
9cCb 9th Cb lobule, c cbw cereb white mat ECu ext cuneate nu IRt intermed ret nu plf posterolat fissure RPa raphe pallidus nu SolIM Sol intermediate vsc vent spinocer tr
10Cb 10th Cb lobule chp choroid plexus Gi gigantocell ret nu LRt lat reticular nu PM paramedian lobule RVRG rostral ventral rsp SolM Sol medial
10N dorsal mo nu 10 Cop copula of pyramis Gr gracile nu LRtS5 lat retic sub5 pms paramedian sulcus sf 2ary fissure SolV Sol ventral
RBSC5 Book.qxp 16/9/04 16:36 Page 146

4V 4th ventricle 12N hypoglossal nu Cu cuneate nu IOBe IO beta subnu ml medial lemniscus ppf prepyramidal fiss sf 2ary fissure SolVL Sol ventrolat
5Sol trig sol trans zn AmbL ambig nu loose cu cuneate fasciculus IOC IO med subnu C mlf med long fasc PSol parasolitary nu SolCe Sol central sp5 sp trigeminal tr
7Cb 7th Cb lobule apmf ansoparamed fiss dsc/oc dors sp cer/ol cer IOD IO dorsal nu Mx matrix region py pyramidal tract SolDL Sol dorsolat Sp5I spinal 5 interpolar
8Cb 8th Cb lobule C1 C1 adren cells ECu ext cuneate nu IOPr IO principal nu Pa5 paratrigeminal nu Ro nu of Roller SolG Sol gelatinous SubP subpostrema area
9a,bCb 9th Cb lobule, a&b cbw cereb white mat Gi gigantocell ret nu IOVL IO ventrolat PCRt parvicell ret nu ROb raphe obscurus nu SolI Sol interstitial ts tectospinal tract
9cCb 9th Cb lobule, c chp choroid plexus Gr gracile nu IRt intermed ret nu plf posterolat fissure RPa raphe pallidus nu SolIM Sol intermediate vsc vent spinocer tr
10Cb 10th Cb lobule Cop copula of pyramis InM intmed of medulla LRt lat reticular nu PM paramedian lobule rs rubrospinal tract SolM Sol medial
10N dorsal mo nu 10 Crus2 crus 2 ansiform IOB IO med subnu B LRtS5 lat retic sub5 pms paramedian sulcus RVRG rostral ventral rsp SolV Sol ventral

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 146
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
7 3
7Cb
pms

6 ppf 4

Crus2
cbw 8Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 5
sf

9a,bCb
apmf
4 6
cbw
9cCb
PM
plf Cop

3 10Cb ECu 7
PSol
cu SubP Gr Cu Pa5
chp SolG
sol SolM SolDL SolI Mx
4V
SolCe 10N 5Sol
2 SolV 8
sp5 SolIM SolVL
12N
InM Sp5I
Ro PCRt
IRt
mlf
1 9
dsc/oc
ROb Gi
AmbL LRtS5
ts RVRG
rs
IOBe LRt
C1
0 IOVL 10
vsc IRt
IOC LRt
ml IOD
py RPa IOB
IOPr
-1 11

Interaural -4.56 mm Bregma -13.56 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 147

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 147
5 5
8 2

0 Interaural 10

+15 +10 +5 0 -5
7 3

7Cb

ppf
6 4

8Cb

Crus2
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

5 sf 5
9a,bCb

apmf cbw cbw

4 9cCb 6
PM
Cop
plf

3 cu ECu 7
10Cb
Cu CuR Pa5
sp5 SolI Gr
AP SubP SolDL
SolC SolM Mx 5Sol
sol SolCe 10N SolV
2 CC SolVL 8
SolIM 12N
Sp5I
PCRt
Ro 12GH
IRt
dsc/oc mlf
1 ROb
9
Gi AmbL
ts
rs RVRG C1/A1 LRtS5
IOK
vsc IOBe 10n
0 LRt IRt 10
RPa IOC
IOB IOD
ml
py IOPr
-1 vert 11

Interaural -4.68 mm Bregma -13.68 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

5Sol trig sol trans zn 12GH 12 geniohyoid Cop copula of pyramis Gr gracile nu LRt lat reticular nu PM paramedian lobule sf 2ary fissure SolV Sol ventral
7Cb 7th Cb lobule 12N hypoglossal nu Crus2 crus 2 ansiform IOB IO med subnu B LRtS5 lat retic sub5 ppf prepyramidal fiss sol solitary tract SolVL Sol ventrolat
8Cb 8th Cb lobule Amb ambiguus nu Cu cuneate nu IOBe IO beta subnu ml medial lemniscus py pyramidal tract SolC Sol commissural sp5 sp trigeminal tr
9a,bCb 9th Cb lobule, a&b AP area postrema cu cuneate fasciculus IOC IO med subnu C mlf med long fasc Ro nu of Roller SolCe Sol central Sp5I spinal 5 interpolar
9cCb 9th Cb lobule, c apmf ansoparamed fiss CuR cun rotundus IOD IO dorsal nu Mx matrix region ROb raphe obscurus nu SolDL Sol dorsolat SubP subpostrema area
10Cb 10th Cb lobule C1/A1 C1 norad/A1 adr dsc/oc dors sp cer/ol cer IOK IO med cap Kooy Pa5 paratrigeminal nu RPa raphe pallidus nu SolI Sol interstitial ts tectospinal tract
10N dorsal mo nu 10 cbw cereb white mat ECu ext cuneate nu IOPr IO principal nu PCRt parvicell ret nu rs rubrospinal tract SolIM Sol intermediate vert vertebral artery
10n vagus n CC central canal Gi gigantocell ret nu IRt intermed ret nu plf posterolat fissure RVRG rostral ventral rsp SolM Sol medial vsc vent spinocer tr
RBSC5 Book.qxp 16/9/04 16:36 Page 148

5Sol trig sol trans zn AmbL ambig nu loose Cu cuneate nu IOB IO med subnu B LRtS5 lat retic sub5 pms paramedian sulcus sf 2ary fissure SolVL Sol ventrolat
7Cb 7th Cb lobule AP area postrema cu cuneate fasciculus IOBe IO beta subnu mld med lem decussn ppf prepyramidal fiss sol solitary tract sp5 sp trigeminal tr
8Cb 8th Cb lobule apmf ansoparamed fiss CuR cun rotundus IOC IO med subnu C mlf med long fasc py pyramidal tract SolC Sol commissural Sp5I spinal 5 interpolar
9Cb 9th Cb lobule C1/A1 C1 norad/A1 adr dsc/oc dors sp cer/ol cer IOD IO dorsal nu Mx matrix region Ro nu of Roller SolDL Sol dorsolat SubP subpostrema area
10Cb 10th Cb lobule cbw cereb white mat ECu ext cuneate nu IOK IO med cap Kooy Pa5 paratrigeminal nu ROb raphe obscurus nu SolI Sol interstitial ts tectospinal tract
12GH 12 geniohyoid CC central canal Gi gigantocell ret nu IRt intermed ret nu PCRt parvicell ret nu RPa raphe pallidus nu SolIM Sol intermediate vert vertebral artery
12N hypoglossal nu Cop copula of pyramis Gr gracile nu LRt lat reticular nu plf posterolat fissure rs rubrospinal tract SolM Sol medial vsc vent spinocer tr
a artery Crus2 crus 2 ansiform IOA IO med subnu A LRtPC lat retic parvicell PM paramedian lobule RVRG rostral ventral rsp SolV Sol ventral

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 148
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3

7Cb
pms
ppf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
apmf
8Cb

Crus2
sf
5 5
9a,bCb

cbw
cbw
4 9cCb 6
PM
Cop
plf
a cu ECu
3 7
10Cb Pa5
Gr Cu
sp5 AP CuR
SolIM SolC SubPSolDL
Mx 5Sol
sol 10N SolV SolI
SolVL
2 SolM CC Sp5I 8
12N
dsc/oc PCRt
12GH IRt
Ro
mlf
1 mld 9
AmbL Gi
rs ROb
ts RVRG
IOK C1/A1 LRtS5
vsc
RPa IOBe IOD LRt
0 LRtPC 10
IOC
ml
IOA
py IOB
Interaural -4.80 mm vert
Bregma -13.80 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 149

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 149
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3

7Cb
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 ppf 4
8Cb
apmf

sf
Crus2
5 5
9a,bCb

Cop PM

4 9cCb 6

cbw

ECu
3 cu 10Cb 7
a Gr Cu Pa5
SolM AP
SubP SolDL
CuR
sp5
sol SolC Mx Sp5C
SolV SolI
2 SolIM CC 10N SolVL 8
12N MdD
CeCv
10n IRt Sp5I
Ro
12GH
mlf
1 AmbL mld 9
dsc/oc ts ROb MdV
rs RVRG
IOK C1/A1 LRtS5
IOBe
IOC IOD
0 IOB IOA LRt 10
vsc ml LRtPC
py RPa

Interaural -4.92 mm 12n vert IOD Bregma -13.92 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

7Cb 7th Cb lobule 12n root of 12n Crus2 crus 2 ansiform IOBe IO beta subnu MdV medull ret vent Ro nu of Roller SolI Sol interstitial ts tectospinal tract
8Cb 8th Cb lobule a artery Cu cuneate nu IOC IO med subnu C ml medial lemniscus ROb raphe obscurus nu SolIM Sol intermediate vert vertebral artery
9a,bCb 9th Cb lobule, a&b AmbL ambig nu loose cu cuneate fasciculus IOD IO dorsal nu mld med lem decussn RPa raphe pallidus nu SolM Sol medial
9cCb 9th Cb lobule, c AP area postrema CuR cun rotundus IOK IO med cap Kooy mlf med long fasc rs rubrospinal tract SolV Sol ventral
10Cb 10th Cb lobule C1/A1 C1 norad/A1 adr dsc/oc dors sp cer/ol cer IRt intermed ret nu Mx matrix region RVRG rostral ventral rsp SolVL Sol ventrolat
10N dorsal mo nu 10 cbw cereb white mat ECu ext cuneate nu LRt lat reticular nu Pa5 paratrigeminal nu sf 2ary fissure sp5 sp trigeminal tr
10n vagus n CC central canal Gr gracile nu LRtPC lat retic parvicell PM paramedian lobule sol solitary tract Sp5C spinal 5 caudal
12GH 12 geniohyoid CeCv central cervic nu IOA IO med subnu A LRtS5 lat retic sub5 ppf prepyramidal fiss SolC Sol commissural Sp5I spinal 5 interpolar
12N hypoglossal nu Cop copula of pyramis IOB IO med subnu B MdD medullary ret dors py pyramidal tract SolDL Sol dorsolat SubP subpostrema area
RBSC5 Book.qxp 16/9/04 16:36 Page 150

7Cb 7th Cb lobule 12n root of 12n Cop copula of pyramis Gr gracile nu LRtPC lat retic parvicell PM paramedian lobule sf 2ary fissure SolVL Sol ventrolat
8Cb 8th Cb lobule Amb ambiguus nu Crus2 crus 2 ansiform IOA IO med subnu A LRtS5 lat retic sub5 ppf prepyramidal fiss sol solitary tract sp5 sp trigeminal tr
9a,bCb 9th Cb lobule, a&b AP area postrema Cu cuneate nu IOB IO med subnu B MdD medullary ret dors py pyramidal tract SolC Sol commissural Sp5C spinal 5 caudal
9cCb 9th Cb lobule, c apmf ansoparamed fiss cu cuneate fasciculus IOBe IO beta subnu MdV medull ret vent Ro nu of Roller SolDL Sol dorsolat Sp5I spinal 5 interpolar
10N dorsal mo nu 10 C1/A1 C1 norad/A1 adr CuR cun rotundus IOC IO med subnu C ml medial lemniscus ROb raphe obscurus nu SolI Sol interstitial SubP subpostrema area
10n vagus n cbw cereb white mat CVL caudoventrolat rt IOK IO med cap Kooy mlf med long fasc RPa raphe pallidus nu SolIM Sol intermediate ts tectospinal tract
12GH 12 geniohyoid CC central canal dsc dorsal sp cereb tr IRt intermed ret nu Mx matrix region rs rubrospinal tract SolM Sol medial vert vertebral artery
12N hypoglossal nu CeCv central cervic nu ECu ext cuneate nu LRt lat reticular nu Pa5 paratrigeminal nu RVRG rostral ventral rsp SolV Sol ventral vsc vent spinocer tr

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 150
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3

7Cb
ppf
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
8Cb

apmf sf
5 Crus2 5

9a,bCb
cbw
4 6
cbw PM
Cop

9cCb
3 ECu 7
cu Pa5
SubP Gr Cu
AP SolIM CuR
sp5 SolDL Mx
sol SolM SolI Sp5C
SolC
2 10N SolV SolVL
8
CC CeCv
12N
MdD Sp5I
10n Ro IRt
12GH
mlf
1 mld 9
ROb MdV
AmbL
dsc rs RVRG IOK C1/A1
12n ts
CVL LRtS5
IOBe LRt
0 RPa IOC LRtPC 10
vsc IOB
ml IOA
12n py
Interaural -5.04 mm vert
Bregma -14.04 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 151

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 151
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4
8Cb

sf
5 5

9a,bCb PM
cbw
Cop
4 6

9cCb
ECu
3 cu 7
Cu
Gr CuR
AP
SolDL Mx
sp5 SolM SubP
sol SolC Sp5C
2 10N SolIM SolVL 8
CC CeCv SolV MdD
12N Sp5I
dsc/oc
IRt
ia
mlf 12GH
1 AmbL 9
RVRG ROb
ts MdV
rs mld CVL
A1/C1 IOK
12n
IOC LRt
IOBe
0 vsc IOB LRtPC 10
ml
RPa IOA
12n
Interaural -5.16 mm vert
py
Bregma -14.16 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

8Cb 8th Cb lobule A1/C1 A1noradr/C1adren Cu cuneate nu ia int arcuate fibers LRt lat reticular nu Mx matrix region sf 2ary fissure SolVL Sol ventrolat
9a,bCb 9th Cb lobule, a&b AmbL ambig nu loose cu cuneate fasciculus IOA IO med subnu A LRtPC lat retic parvicell PM paramedian lobule sol solitary tract sp5 sp trigeminal tr
9cCb 9th Cb lobule, c AP area postrema CuR cun rotundus IOB IO med subnu B MdD medullary ret dors py pyramidal tract SolC Sol commissural Sp5C spinal 5 caudal
10N dorsal mo nu 10 cbw cereb white mat CVL caudoventrolat rt IOBe IO beta subnu MdV medull ret vent ROb raphe obscurus nu SolDL Sol dorsolat Sp5I spinal 5 interpolar
12GH 12 geniohyoid CC central canal dsc/oc dors sp cer/ol cer IOC IO med subnu C ml medial lemniscus RPa raphe pallidus nu SolIM Sol intermediate SubP subpostrema area
12N hypoglossal nu CeCv central cervic nu ECu ext cuneate nu IOK IO med cap Kooy mld med lem decussn rs rubrospinal tract SolM Sol medial ts tectospinal tract
12n root of 12n Cop copula of pyramis Gr gracile nu IRt intermed ret nu mlf med long fasc RVRG rostral ventral rsp SolV Sol ventral vert vertebral artery
vsc vent spinocer tr
RBSC5 Book.qxp 16/9/04 16:36 Page 152

8Cb 8th Cb lobule
9a,bCb 9th Cb lobule, a&b
9cCb 9th Cb lobule, c
10N dorsal mo nu 10
12GH 12 geniohyoid
12N hypoglossal nu
12n root of 12n
A1 A1noradr cells
AP area postrema
cbw cereb white mat
CC central canal
CeCv central cervic nu
Cop copula of pyramis
Ct conterminal nu
Cu cuneate nu
cu cuneate fasciculus
CuR cun rotundus
dsc dorsal sp cereb tr
Gr gracile nu MdD medullary ret dors ROb raphe obscurus nu SolVL Sol ventrolat
ia int arcuate fibers MdV medull ret vent RPa raphe pallidus nu sp5 sp trigeminal tr
IOA IO med subnu A ml medial lemniscus rs rubrospinal tract Sp5C spinal 5 caudal
IOB IO med subnu B mld med lem decussn sf 2ary fissure SubP subpostrema area
IOBe IO beta subnu mlf med long fasc sol solitary tract ts tectospinal tract
IOC IO med subnu C Mx matrix region SolC Sol commissural vert vertebral artery
IRt intermed ret nu PM paramedian lobule SolDL Sol dorsolat vsc vent spinocer tr
LRt lat reticular nu py pyramidal tract SolM Sol medial
LRtPC lat retic parvicell RAmb retroambiguus nu SolV Sol ventral

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 152
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

8Cb
sf
5 5
9a,bCb PM
Cop
cbw

4 6

9cCb
3 7
cu
sp5 Cu
Gr
SubP AP CuR
SolDL Mx
sol SolC SolM
2 SolVL 10N Sp5C 8
SolV CeCv
CC MdD
12N
IRt
ia
12GH
mlf
1 RAmb 9
ROb
mld MdV A1
ts
dsc rs IOBe
LRt
vsc IOB LRtPC
0 ml IOC IOA Ct 10
12n
RPa
12n py Ct
Interaural -5.28 mm vert Bregma -14.28 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 153

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 153
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

8Cb
sf
5 5
9a,bCb
PM
cbw Cop
4 6

9cCb
3 7
cu
Obex
Gr Cu
sp5 gr
CuR Mx
SolC SolDL
sol SolM SolVL
2 SolV 8
Sp5C
CC 10N MdD
12N CeCv
ia IRt
12GH
1 RAmb mlf 9
mld
MdV
ts IOBe A1
dsc rs
vsc ml LRt LRtPC
0 Ct 10
IOC IOB
12n IOA
12n py
Ct
Interaural -5.40 mm vert Bregma -14.40 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

8Cb 8th Cb lobule
9a,bCb 9th Cb lobule, a&b
9cCb 9th Cb lobule, c
10N dorsal mo nu 10
12GH 12 geniohyoid
12N hypoglossal nu
12n root of 12n
A1 A1noradr cells
cbw cereb white mat
CC central canal
CeCv central cervic nu
Cop copula of pyramis
Ct conterminal nu
Cu cuneate nu
cu cuneate fasciculus
CuR cun rotundus
dsc dorsal sp cereb tr
Gr gracile nu
gr gracile fasciculus MdD medullary ret dors RAmb retroambiguus nu sp5 sp trigeminal tr
ia int arcuate fibers MdV medull ret vent rs rubrospinal tract Sp5C spinal 5 caudal
IOA IO med subnu A ml medial lemniscus sf 2ary fissure ts tectospinal tract
IOB IO med subnu B mld med lem decussn sol solitary tract vert vertebral artery
IOBe IO beta subnu mlf med long fasc SolC Sol commissural vsc vent spinocer tr
IOC IO med subnu C Mx matrix region SolDL Sol dorsolat
IRt intermed ret nu Obex obex SolM Sol medial
LRt lat reticular nu PM paramedian lobule SolV Sol ventral
LRtPC lat retic parvicell py pyramidal tract SolVL Sol ventrolat
RBSC5 Book.qxp 16/9/04 16:36 Page 154

8Cb 8th Cb lobule
9a,bCb 9th Cb lobule, a&b
9cCb 9th Cb lobule, c
10N dorsal mo nu 10
12GH 12 geniohyoid
12N hypoglossal nu
12n root of 12n
A1 A1noradr cells
cbw cereb white mat dsc dorsal sp cereb tr LRtPC lat retic parvicell rs rubrospinal tract sp5 sp trigeminal tr
CC central canal Gr gracile nu MdD medullary ret dors sf 2ary fissure Sp5C spinal 5 caudal
CeCv central cervic nu gr gracile fasciculus MdV medull ret vent sol solitary tract ts tectospinal tract
Cop copula of pyramis IOA IO med subnu A mld med lem decussn SolC Sol commissural vert vertebral artery
Ct conterminal nu IOB IO med subnu B mlf med long fasc SolDL Sol dorsolat vsc vent spinocer tr
Cu cuneate nu IOC IO med subnu C Mx matrix region SolM Sol medial
cu cuneate fasciculus IRt intermed ret nu py pyramidal tract SolV Sol ventral
CuR cun rotundus LRt lat reticular nu RAmb retroambiguus nu SolVL Sol ventrolat

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 154
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

8Cb
sf

5 5

9a,bCb

Cop
cbw
4 6

9cCb
3 7
sp5 cu
Gr Cu
gr CuR
SolC SolDL Mx
2 SolM SolVL 8
sol 10N Sp5C
CC SolV
CeCv MdD
12N
IRt
ia 12GH
1 RAmb mlf 9
MdV
12n mld A1
dsc rs
ts LRt LRtPC
vsc
0 Ct 10
12n IOC
Ct IOA
12n py IOB
Interaural -5.52 mm vert Bregma -14.52 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 155

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 155
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

sf 8Cb

5 5

9a,bCb

4 cbw 6

3 9cCb 7
cu
Gr
gr Cu
sp5 MnA Mx IB Mx
2 SolC SolDL 8
sol A2 SolM
SolVL 10N SolV Sp5C
CC
12N MdD
CeCv
ia IRt
12GH
1 mld
9
RAmb mlf
12n pyd MdV
A1
rs ts
dsc Ct LRt LRtPC
0 vsc IOM 10
12n 12n py
Ct
Interaural -5.64 mm vert Bregma -14.64 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

8Cb 8th Cb lobule
9a,bCb 9th Cb lobule, a&b
9aCb 9th Cb lobule, a
10N dorsal mo nu 10
12GH 12 geniohyoid
12N hypoglossal nu
12n root of 12n
A1 A1noradr cells
A2 A2 noradr cells
cbw cereb white mat
CC central canal
CeCv central cervic nu
Ct conterminal nu
Cu cuneate nu
cu cuneate fasciculus
dsc dorsal sp cereb tr
Gr gracile nu MdV medull ret vent rs rubrospinal tract sp5 sp trigeminal tr
gr gracile fasciculus mld med lem decussn sf 2ary fissure Sp5C spinal 5 caudal
IB internal basal nu mlf med long fasc sol solitary tract ts tectospinal tract
IOM IO medial nu MnA median acc nu SolC Sol commissural vert vertebral artery
IRt intermed ret nu Mx matrix region SolDL Sol dorsolat vsc vent spinocer tr
LRt lat reticular nu py pyramidal tract SolM Sol medial
LRtPC lat retic parvicell pyd pyramidal decussn SolV Sol ventral
MdD medullary ret dors RAmb retroambiguus nu SolVL Sol ventrolat
RBSC5 Book.qxp 16/9/04 16:36 Page 156

9a,bCb 9th Cb lobule, a&b
9cCb 9th Cb lobule, c
10N dorsal mo nu 10
12GH 12 geniohyoid
12N hypoglossal nu
A1 A1noradr cells
A2 A2 noradr cells
cbw cereb white mat
CC central canal
CeCv central cervic nu
Cu cuneate nu
cu cuneate fasciculus
dsc dorsal sp cereb tr
Gr gracile nu
gr gracile fasciculus MdD medullary ret dors RAmb retroambiguus nu SolVL Sol ventrolat
ia int arcuate fibers MdV medull ret vent rs rubrospinal tract sp5 sp trigeminal tr
IB internal basal nu mlf med long fasc sol solitary tract Sp5C spinal 5 caudal
IOM IO medial nu MnA median acc nu SolC Sol commissural ts tectospinal tract
IRt intermed ret nu Mx matrix region SolDL Sol dorsolat vert vertebral artery
LRt lat reticular nu py pyramidal tract SolM Sol medial vsc vent spinocer tr
LRtPC lat retic parvicell pyd pyramidal decussn SolV Sol ventral

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 156
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

5 5

9a,bCb
cbw
4 6

9cCb
3 7
cu
Gr
sp5 gr MnA Cu
IB Mx
2 sol SolC SolDL 8
SolV A2 SolVL
Sp5C
SolM CeCv
ia 10N MdD
12N
CC 12GH IRt
1 9
RAmb pyd MdV
mlf A1
rs
ts LRt LRtPC
0 dsc IOM
10
vsc
py
Interaural -5.76 mm vert
Bregma -14.76 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 157

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 157
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

5 5

9a,bCb
4 6

3 7
9cCb
cu Gr
gr Cu
sp5 MnA Mx Mx
2 IB 8
sol SolC SolM SolVL
SolV Sp5C
A2 MdD
12N CeCv
11N CC IRt
1 9
pyd RAmb
11N mlf MdV A1
rs
dsc ts
0 10
vsc LRt LRtPC
py

Interaural -6.00 mm vert Bregma -15.00 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

9a,bCb 9th Cb lobule, a&b
9cCb 9th Cb lobule, c
11N access nerve nu
12N hypoglossal nu
A1 A1noradr cells
A2 A2 noradr cells
CC central canal
CeCv central cervic nu
Cu cuneate nu
cu cuneate fasciculus MdV medull ret vent SolC Sol commissural
dsc dorsal sp cereb tr mlf med long fasc SolM Sol medial
Gr gracile nu MnA median acc nu SolV Sol ventral
gr gracile fasciculus Mx matrix region SolVL Sol ventrolat
IB internal basal nu py pyramidal tract sp5 sp trigeminal tr
IRt intermed ret nu pyd pyramidal decussn Sp5C spinal 5 caudal
LRt lat reticular nu RAmb retroambiguus nu ts tectospinal tract
LRtPC lat retic parvicell rs rubrospinal tract vert vertebral artery
MdD medullary ret dors sol solitary tract vsc vent spinocer tr
RBSC5 Book.qxp 16/9/04 16:36 Page 158

9a,bCb 9th Cb lobule, a&b cu cuneate fasciculus MdV medull ret vent SolM Sol medial
9cCb 9th Cb lobule, c dsc dorsal sp cereb tr mlf med long fasc sp5 sp trigeminal tr
11N access nerve nu Ge5 gelat caudal sp 5 MnA median acc nu Sp5C spinal 5 caudal
A1 A1noradr cells Gr gracile nu Mx matrix region ts tectospinal tract
A2 A2 noradr cells gr gracile fasciculus pyd pyramidal decussn vert vertebral artery
CC central canal IB internal basal nu RAmb retroambiguus nu vsc vent spinocer tr
CeCv central cervic nu IRt intermed ret nu rs rubrospinal tract
Cu cuneate nu MdD medullary ret dors SolC Sol commissural

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 158
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

5 5

4 6

9a,bCb

3 9cCb 7
MnA
cu Cu
Gr
sp5 gr
2 IB 8
SolC Ge5
SolM Mx
A2 MdD Sp5C

CC CeCv IRt
1 9
RAmb MdV
pyd 11N
mlf A1
rs
0 dsc 10
ts
vsc

Interaural -6.24 mm vert

Bregma -15.24 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 159

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 159
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

5 5

4 6

9a,bCb

3 7

cu Gr
sp5 gr MnA Cu
2 IB 8
SolC Ge5
A2 SolM
MdD Sp5C

CeCv IRt
1 CC 9
RAmb
mlf pyd MdV
11N A1
rs
0 dsc ts 10
vsc

vert
Interaural -6.48 mm Bregma -15.48 mm
7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

9a,bCb 9th Cb lobule, a&b cu cuneate fasciculus MdD medullary ret dors SolC Sol commissural
11N access nerve nu dsc dorsal sp cereb tr MdV medull ret vent SolM Sol medial
A1 A1noradr cells Ge5 gelat caudal sp 5 mlf med long fasc sp5 sp trigeminal tr
A2 A2 noradr cells Gr gracile nu MnA median acc nu Sp5C spinal 5 caudal
CC central canal gr gracile fasciculus pyd pyramidal decussn ts tectospinal tract
CeCv central cervic nu IB internal basal nu RAmb retroambiguus nu vert vertebral artery
Cu cuneate nu IRt intermed ret nu rs rubrospinal tract vsc vent spinocer tr
RBSC5 Book.qxp 16/9/04 16:36 Page 160

11N access nerve nu
A1 A1noradr cells
A2 A2 noradr cells
CC central canal
CeCv central cervic nu
Cu cuneate nu
cu cuneate fasciculus
dcs dors corticosp tr
dsc dorsal sp cereb tr
Ge5 gelat caudal sp 5
Gr gracile nu
gr gracile fasciculus
ia int arcuate fibers
IB internal basal nu
IRt intermed ret nu
MdD medullary ret dors
MdV medull ret vent
mlf med long fasc
MnA median acc nu
pyd pyramidal decussn
RAmb retroambiguus nu
rs rubrospinal tract
SolC Sol commissural
SolM Sol medial
sp5 sp trigeminal tr
Sp5C spinal 5 caudal
ts tectospinal tract
vert vertebral artery
vsc vent spinocer tr

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 160
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

5 5

4 6

3 7

cu
sp5 Gr Cu
gr
2 MnA 8
dcs IB Sp5C
SolC SolM MdD Ge5

A2 IRt
CeCv
ia
1 CC RAmb 9
MdV A1
rs pyd 11N
dsc mlf
vsc ts
0 vert 10

Interaural -6.72 mm Bregma -15.72 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7
RBSC5 Book.qxp 16/9/04 16:36 Page 161

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

+5 0 -5 -10 -15
10 Bregma 0
Figure 161
5 5
9 1

0 Interaural 10

+15 +10 +5 0 -5
8 2

7 3
The Rat Brain in Stereotaxic Coordinates 5th Edition Paxinos & Watson

6 4

5 5

4 6

3 7

cu Gr
sp5 gr Cu
2 MnA 8
dcs IB Sp5C
SolC SolM MdD Ge5
A2
CC CeCv IRt
1 ia 9
MdV
rs
pyd A1
dsc 11N
mlf
vsc
0 ts 10
vert

Interaural -6.96 mm Bregma -15.96 mm

7 6 5 4 3 2 1 0 1 2 3 4 5 6 7

11N access nerve nu
A1 A1noradr cells
A2 A2 noradr cells
CC central canal
CeCv central cervic nu
Cu cuneate nu
cu cuneate fasciculus
dcs dors corticosp tr
dsc dorsal sp cereb tr
Ge5 gelat caudal sp 5
Gr gracile nu
gr gracile fasciculus
ia int arcuate fibers
IB internal basal nu
IRt intermed ret nu
MdD medullary ret dors
MdV medull ret vent
mlf med long fasc
MnA median acc nu
pyd pyramidal decussn
rs rubrospinal tract
SolC Sol commissural
SolM Sol medial
sp5 sp trigeminal tr
Sp5C spinal 5 caudal
ts tectospinal tract
vert vertebral artery
vsc vent spinocer tr
RBSC5 Book.qxp 16/9/04 16:36 Page 162

1-10 spinal cord layers IML intermediolateral cell column *Fig 162a, 162b, 162c, 162d, 162e are In G. Paxinos(Ed), The Nervous System,
CeCv central cervical nucleus IMM intermediomedial cell column reproduced from Molander and Grant Second Edition, Academic Press,
cu cuneate fasciculus LatC lateral cervical nucleus (1995) with permission of the authors. SanDiego.
dl dorsolateral fasciculus LSp lateral spinal nucleus Users of these figures should cite
gr gracile fasciculus dcs dorsal corticospinal tract Molander, C. and Grant, G., 1995,
Spinal cord cytoarchitecture.

Figure 162a*

1
gr
gr 2 1
cu dl cu 2
3
3
4 4

dcs
dcs 5 LatC LSp
5
CeCv
10 IMM CeCv 7
7 LSp
LatC 10 IMM
9 8
9 9 8 9
9
9

C1
C3

1
dl gr
cu 2
3 gr 1
4 dl 2
cu 3
4
dcs 5 LSp LSp
LatC
5
dcs
CeCv
7 6
10 IMM
10 IMM 7
9 8 9
9
8
9 9
9

C2 C4
RBSC5 Book.qxp 16/9/04 16:36 Page 163

Figure 162b*
dl 1
2 dl 1
3 gr
gr
4 2
cu LSp cu 3
5 4 LSp

dcs 6
dcs 5

7 6
10 IMM
9 IMM
10 7
8 9 9 9
9 9 9 8
9 9
9

C5 C7

dl gr dl
1
cu gr 1
2
3 2
4 cu 4 LSp
LSp 3
5 5
dcs dcs
6 6

IMM IMM 7 9
7 9
10
8 10 8
9 9 9
9 9

C6 C8

1-10 spinal cord layers IML intermediolateral cell column *Fig 162a, 162b, 162c, 162d, 162e are In G. Paxinos(Ed), The Nervous System,
CeCv central cervical nucleus IMM intermediomedial cell column reproduced from Molander and Grant Second Edition, Academic Press,
cu cuneate fasciculus LatC lateral cervical nucleus (1995) with permission of the authors. SanDiego.
dl dorsolateral fasciculus LSp lateral spinal nucleus Users of these figures should cite
gr gracile fasciculus dcs dorsal corticospinal tract Molander, C. and Grant, G., 1995,
Spinal cord cytoarchitecture.
RBSC5 Book.qxp 16/9/04 16:36 Page 164

1-10 spinal cord layers IML intermediolateral cell column *Fig 162a, 162b, 162c, 162d, 162e are In G. Paxinos(Ed), The Nervous System,
CeCv central cervical nucleus IMM intermediomedial cell column reproduced from Molander and Grant Second Edition, Academic Press,
cu cuneate fasciculus LatC lateral cervical nucleus (1995) with permission of the authors. SanDiego.
D dorsal nucleus (Clarke) LSp lateral spinal nucleus Users of these figures should cite
dl dorsolateral fasciculus dcs dorsal corticospinal tract Molander, C. and Grant, G., 1995,
gr gracile fasciculus Spinal cord cytoarchitecture.

Figure 162c*

gr
dl gr 1
2 dl
3 1
cu cu 2
4
3 4
5 LSp LSp
dcs IML 5
6
9 dcs IML
IMM 7 7
10 D
8 IMM
9 9 10 8
9
9

T1 T5

dl gr
1
2 dl 1
cu gr 2
3 4 4
3
LSp
5 LSp
5
dcs dcs
IML D IML
D 7 7
IMM
IMM
10 8 10 8
9 9
9 9

T3 T10
RBSC5 Book.qxp 16/9/04 16:36 Page 165

Figure 162d*

dl 1
2 dl 1
dl 1 3
2 gr 4 gr 2
gr 3 4 3
LSp
LSp 5 4
5 dcs dcs
dcs D 6 5 LSp
IML
IML 10 6
10 7
IMM 10
IMM 7 9 9
9 IMM 7

8 8
9 8
9 9
9 9
9

L1 L3 L5

dl 1 dl 1
2
gr 4 2
3 gr gr 1
LSp 3 dl
5 4 dcs 2 3
dcs 4
LSp
D 5 5
IML dcs LSp
10 6 6
IMM 7
10 10
9 7
IMM 7 IMM 9
8 9 9
9 8 9
8 9 9
9 9
9 9

L2 L4 L6

1-10 spinal cord layers
CeCv central cervical nucleus
D dorsal nucleus (Clarke)
dl dorsolateral fasciculus
gr gracile fasciculus
IML intermediolateral cell column
IMM intermediomedial cell column
LatC lateral cervical nucleus
LSp lateral spinal nucleus
dcs dorsal corticospinal tract
*Fig 162a, 162b, 162c, 162d, 162e are In G. Paxinos(Ed), The Nervous System,
reproduced from Molander and Grant Second Edition, Academic Press,
(1995) with permission of the authors. SanDiego.
Users of these figures should cite
Molander, C. and Grant, G., 1995,
Spinal cord cytoarchitecture.
RBSC5 Book.qxp 16/9/04 16:36 Page 166

1-10 spinal cord layers
CeCv central cervical nucleus
dl dorsolateral fasciculus
gr gracile fasciculus
IML intermediolateral cell column
IMM intermediomedial cell column
LatC lateral cervical nucleus
LSp lateral spinal nucleus
dcs dorsal corticospinal tract
*Fig 162a, 162b, 162c, 162d, 162e are In G. Paxinos(Ed), The Nervous System,
reproduced from Molander and Grant Second Edition, Academic Press,
(1995) with permission of the authors. SanDiego.
Users of these figures should cite
Molander, C. and Grant, G., 1995,
Spinal cord cytoarchitecture.

Figure 162e*

gr 1 gr 1
dcs 2 dl dcs 2
dl 3 3
4 4
5 LSp
5
LSp
6 10
7
10
IMM
IMM 7
8
9
8 9
9

S1 S3

gr gr 1
1 dsc 2
2 3
dcs 3
4 4
5 LSp
5
LSp
10 10
7
7 IMM
IMM
8
9
8
9

S2 S4