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F rom birth to three months of age is the most sensitive rearing period for the
young calf. With biological, environmental, and nutritional stressors, the
success of this first rearing phase depends on calf managers paying special
attention to detail. Among the most important is establishing a sound calf
nutrition program.
Feeding Colostrum
Feeding calves the correct amount of high-quality colostrum immediately after
birth is the single most important management practice in calf nutrition. Colos-
trum, defined as milk extracted from the mammary gland in the first 24 hours
after birth, contains immunoglobulins that, when absorbed by the calf’s gut,
resist disease challenges. The passing of this protection from dam to calf is called
passive transfer. Research has clearly shown that calves with adequate passive
transfer grow better and have lower mortality and health cost when compared to
calves with failed passive transfer (See Table 3.1).
The immunoglobulins found in colostrum are large proteins. Calves have
openings in the small intestines to accommodate the protein’s absorption—but
21
22
Table 3.1. Passive immunity improves growth and lowers mortality and health cost.
Chapter 3 Passive Transfer
Calf Nutrition
Item Poor Fair Average Good Excellent
Number of calves 51 85 126 195 176
Serum Ig, milligrams per milliliter 0-5 5-10 11-15 16-25 > 25
Four-week gain, pounds per day 0.73 0.81 0.85 0.90 0.92
Feed conversion, pounds feed/pounds gain 2.9 2.6 2.5 2.0 1.9
Scour, days 8.7 6.1 4.7 5.0 4.0
Mortality, percent 33 11 7 3 4
Veterinary Cost, dollars $12.50 $9.85 $7.40 $7.70 $6.20
Source: Data adapted from M.A. Fowler, 1999 PDHGA Proceedings, by P.C. Hoffman, University of Wisconsin.
these openings close shortly after birth. Three types Table 3.2. Nutritional composition of colostrum,
of immunoglobulins (Ig) can be found in colos- transition milk and whole milk.
trum. Immunoglobulin G (IgG) makes up 70 Comparison of colostrum, transition milk (harvested
percent to 80 percent of the immunoglobulins and between 24 and 72 hours) and whole milk from dairy
cows.
helps identify and destroy invading pathogens.
Transition Whole
Immunoglobulin M (IgM) comprises 10 percent to Composition Colostrum Milk Milk
15 percent of immunoglobulins and serves as the Specific gravity 1.05 1.03 1.03
first line of defense against septicemia. Immunoglo- Solids, percent 23.9 14.1 12.9
bulin A (IgA) comprises the remaining 15 percent Protein, percent 14.0 5.1 3.1
of immunoglobulins in colostrum and protects the Casein, percent 4.8 3.8 2.5
IgG, milligrams
mucosal surfaces, such as the intestine, from per milliliters 48.0 15.0 0.6
invasive pathogenic bacteria. Fat, percent 6.7 3.9 3.7
Colostrum also contains vitamins, minerals, Lactose, percent 2.7 4.4 5.0
energy (carbohydrate, fat) and proteins needed for Vitamin A,
calf metabolism, growth and for additional stimula- IU/gallon 46440 17820 5400
Vitamin D,
tion of the calf’s immune system. Hormones
IU/gallon 360 210 140
(insulin) and growth factors (IGF-1) in colostrum Vitamin E,
also aid metabolism. IU/gallon 20 13 4
Milk harvested between 24 and 72 hours is Calcium, percent 0.26 0.15 0.13
termed transition milk, as it composition differs Magnesium,
from colostrum. Milk composition changes again percent 0.04 0.01 0.01
Potassium,
when it is harvested 72 hours after calving—when percent 0.14 0.14 0.15
it is considered whole sellable milk. Table 3.2 Sodium,
shows the composition differences of colostrum, percent 0.07 0.05 0.04
transitional milk and whole milk. Chlorine,
percent 0.12 0.10 0.07
Total minerals,
percent 1.11 0.87 0.74
Three Keys to Source: Adapted from Foley and Otterby (1978), NC205 (1991) and
Colostrum Feeding Quigley (1996) by H. Chester-Jones, University of Minnesota, 2003.
To achieve the desired effects of colostrum, calf
managers should focus on three factors: speed of
feeding, the quantity fed and colostrum quality. calf, with some calves unable to absorb immuno-
globulins 10 hours after birth (See Figure 3.1).
1. Feed colostrum within one hour after birth. 2. Feed colostr um at a rate of 12 per
colostrum cent to 15
percent
Approximately 35 percent of ingested immunoglo- per cent of the calf
percent calf’’s body weight. Large-breed
bulins can be absorbed when calves are fed calves weighing more than 90 pounds at birth
colostrum immediately after birth, but this declines can be fed four quarts at birth. Feed calves
to less than 5 percent absorption of immunoglobu- weighing between 50 and 90 pounds three
lins when calves are fed 20 hours after birth. The quarts at birth. If possible, feed an additional
rate at which gut closure occurs varies from calf to two quarts to all calves 12 hours after birth.
23
Efficiency of absorption of immunoglobulis (%) Table 3.3. Time with dam increases mortality.
50 Maximum absorption Mortality in week one through six months of age Chapter 3
increases the longer calves remain with the cow Calf Nutrition
after birth.
40 Time with Dam Herds Percent
After Birth (hours) Evaluated Mortality
2-6 13 5.2
30 7 to 12 35 9.3
13 to 24 32 10.7
25 to 48 24 20.5
20 Average absorption More than 48 35 14.4
Source: Adapted from NC205 Bulletin, 1991, by H. Chester-Jones,
University of Minnesota, 2003.
10
Colostrum
Supplements and Replacers
Figure 3.2. The refractometer measures the When colostrum quantity is lacking or no
protein level in blood. In week-old colostrum is available, colostrum supplements and
calves this can determine if colostrum
replacers, products derived from dried milk, whey,
provided the desired immune protection.
colostrum or bovine serum, should be considered.
In general, do not consider using these products if daily energy and protein requirements for calves fed 25
adequate natural colostrum is available. At proper milk or milk replacer and starter. Table 3.7 summa- Chapter 3
feeding rates, the colostrum supplements have been rizes the recommended concentration levels for Calf Nutrition
shown to have acceptable IgG absorption. Calf minerals and vitamins in liquid and dry feeds.
managers should use supplements as an extender, The NRC’s base recommendations assume a calf’s
to fortify poor-quality colostrum, or when inad- environment is within its thermal neutral zone
equate amounts of fresh or frozen colostrum exist. (TNZ). Within this zone—which ranges between 59
Colostrum replacers provide more IgG when F to 77 F for calves up to three weeks of age—the
compared to supplements. Injectable immunoglo- amount of body heat produced by the calf equals or
bulin products produced from purified bovine exceeds the heat lost from the body through various
blood may be useful to increase circulating anti- means. When the environmental temperature falls
body levels in calves older than 24 hours that didn’t below the TNZ, the calf’s metabolism and energy use
receive the passive transfer of antibodies. Talk with increases. The lower critical temperature (LCT)
your veterinarian about the use of these products. decreases as the calf ages and as its body mass
increases. Breed, hair coat, wind, precipitation, mud
and solar radiation also affect the TNZ. For example,
Calf Nutrient Requirements for every one degree the temperature drops below 59
After supplying colostrum, calf raisers must work F for a newborn calf, the maintenance energy
to meet the calves’ nutrient requirements in the weeks required increases 1 percent. The NRC also provides
that follow. The National Research Council (NRC) recommendations for feeding calves when tempera-
Nutrient Requirements of Dairy Cattle provides useful tures fall above and below the TNZ.
information on the nutritional needs for both milk-fed Newborn calves can be particularly vulnerable to
and weaned calves. Table 3.6 contains estimates of changes in temperature. A calf is born with a supply
of brown adipose tissue that releases energy as heat, requirements for calves. In these situations, addi-
but in extreme cold conditions the energy reserves tional energy should be provided.
may be used up within hours. When brown fat runs In addition, research has indicated that calves
out, a calf uses the energy designed to maintain its housed outside in cold conditions will consume
body temperature and may fail to gain weight. Table more dry matter from calf starter to maintain body
3.8 shows the effects of cold temperature on energy temperature and growth rates (Figure 3.4).
Heat stress in calves can occur when ambient
temperatures and relative humidity rises. Calves are
Table 3.8. Required increase in energy needs as less sensitive to heat stress than mature lactating cows
temperature falls. because calves have a greater surface area per
Calculations based on calves less than three weeks of pound of body weight to radiate excess heat. The
age having lower critical temperature (LCT) of 59 F, only nutritional aspect of managing heat stress in
while calves more than three weeks of age have a LCT
calves is to make sure adequate free choice water is
of 50 F.
available at all times. In cases of severe dehydration
Percent increase
Percent increase in metabolizing or when weaning calves in the hot summer
Environmental in metabolizing energy, months, water intake should be controlled to
temperature, energy, more than 3 prevent over consumption and potential water
degrees F birth to 3 weeks weeks of age
toxicity (gorging) problems.
68 0 0
59 13 0
41 40 13
23 68 40 Digestion and
5 94 68 Rumen Development
-13 121 94 Although a calf is born a ruminant, the rumen
Source: Adapted from National Research Council Recommendations
for Dairy Cattle, 2001, and James, 2002, by H. Chester-Jones,
does not function at birth. Of the four stomachs—
University of Minnesota. the rumen, reticulum, omasum, and abomasum—
27
Esophagus
62 Calf starter intake, pounds of DM Chapter 3
through weaning Esophageal Calf Nutrition
25 percent groove
59 Rumen
56 10 percent 5 percent
Omasum Reticulum
53
50
60 percent
47 Abomasum
Outside wooden Outside Inside raised (true stomach)
hutches translucent stalls
hutches Phase 1—Non-ruminant (birth to 21 days of age)
Abomasum is 60 percent of total stomach. The
Source: Adapted from H. Chester-Jones and Ziegler,
University of Minnesota, 1993.
rumen is undeveloped and nonfunctional. Rumen
development is stimulated by dry feed intake.
Figure 3.4. The effect of housing on starter intake.
60 percent
the rumen and reticulum make up 30 percent of Rumen
the stomach, yet remain underdeveloped, sterile Reticulum
and nonfunctional. The abomasum, or true Omasum
stomach, represents 60 percent and omasum the
other 10 percent. This means nutrition for the 30 percent
Abomasum
newborn calf is essentially feeding a single-
stomached, non-ruminant that is unable to digest
Phase 2—(22 to 56-84 days of age)
fiber (See Figure 3.5).
Dry feed intake, especially grain (starter) stimulates
Colostrum fed to the newborn calf bypasses the growth of the rumen microorganisms which produce
rumen and reticulum and flows via the omasum to volatile fatty acids. These acids stimulate growth of
the abomasum along a tube formed by the reflex rumen tissue
closure of a groove, known as the esophageal
groove. The calf’s sucking of milk stimulates this
reflex. Colostrum components, such as milk
proteins, butterfat, vegetable and other animal fats, 85 percent
Rumen
lactose (milk sugar) and glucose, are digested
primarily in the abomasum and small intestine. Reticulum
When milk arrives, the abomasal wall secretes 7 percent Omasum
Abomasum
rennin, an enzyme, which causes the casein and fat
to form a curd, which will digest slowly for 12 to Phase 3—(More than 84 days of age)
18 hours. With just a few enzymes in the aboma- At this stage, the calf can be considered a ruminant.
sum and small intestine breaking down protein and
Source: Raising Dairy Replacements (1991)
fat in the first 48 hours of life, forming the curd
Figure 3.5. Rumen development.
allows the digestive track to efficiently digest and
assimilate the nutrients fed.
The whey fraction of colostrum—containing water,
minerals, lactose, other proteins and the immunoglo- rumen. Specifically, feeding calf starter at two to
bulins—pass out of the abomasum within a few three days of age stimulates the growth of anaerobic
minutes into the small intestine. Components of the bacteria, which are involved in methane produc-
whey fraction are absorbed into the bloodstream tion, breaking down protein and digesting fiber.
during the first 24 hours after birth. Also, the intake of a highly-digestible calf starter
Within the first day after birth, large concentra- stimulates the development of rumen papillae, or
tions of bacteria (mostly aerobic) start to populate the finger-like projections that absorb nutrients.
the rumen. Feeding dry feeds, such as calf starter, The developing rumen and rumen bacteria need
stimulate the numbers and species of bacteria in the water to be able to ferment dry feeds. Thus offer
28 fresh, clean, free-choice water to all calves. It Pasteurized Waste Milk
Chapter 3 should be remembered that milk bypasses the While saleable whole milk is rarely used, calf
Calf Nutrition rumen and reticulum into the abomasum and does raisers often feed calves excess transition milk and
not help stimulate the growth and development of milk harvested from cows treated with antibiotics.
rumen bacteria. Rumen bacteria need water to This “waste” milk is fed at rates similar to whole
ferment dry calf starter feeds and later forage (See milk—one to two pounds for every 10 to 12
later section on importance of water for calves). pounds of bodyweight.
The Pasteurized Milk Ordinance (PMO) defines
two different methods for pasteurization: batch
Liquid Feeding Options pasteurization at 145 F for 30 minutes (low-
After supplying colostrum, calf raisers must temperature, long-time or LTLT) or high-tempera-
select the type of liquid feed to use in supplying ture, short-time (HTST) pasteurization at 161 F for
nutrients to the calf’s diet. Options include: 15 seconds (usually using a continuous flow
method). Heating results in a reduction in the
Whole Milk concentration of viable bacteria (See Table 3.10).
Whole milk can be the primary liquid feed for The rate of heat inactivation of bacteria increases
calves. If it were dried, whole milk would contain 29 exponentially with time. However pasteurization
percent to 30 percent fat and 25 percent to 26 percent should not be confused with sterilization. Some
protein. This is higher than most milk replacers that heat-tolerant (usually non-pathogenic) bacteria will
typically have 20 percent fat and 18 percent to 22 survive the process. Additionally, if a poor-quality
percent protein. Whole milk will supply ample milk that already has a very high concentration of
protein and energy for calves if one to two pounds is
fed for every 10 to 12 pounds of bodyweight (See
Table 3.9). However, due to its composition, whole
milk may be deficient in trace minerals and vitamins Table 3.10. Time and temperature requirements
(See Table 3.7). This can be remedied by feeding a for pasteurization.
quality calf starter in addition to liquid feeding. Temperature,
Producers rarely use saleable whole milk to feed Fahrenheit Time
to calves as it is not economical. The price of other 145 30 minutes
liquid feed sources is less than using saleable milk. 161 15 seconds
191 1 seconds
If used, ensure the milk is of a uniform tempera-
194 0.5 seconds
ture. Overfeeding or feeding inconsistent amounts 201 0.1 seconds
of whole milk can cause indigestion and scouring. 204 0.05 seconds
Either weigh or measure the milk fed each calf or 212 0.01 seconds
use a container calibrated to the correct volume. Source: Compiled by P.C. Hoffman, University of Wisconsin, 2003.
Table 3.11. Effect of milk replacer feeding rate on calf growth and milk replacer protein levels.
Absorbable Dietary Milk Replacer Crude Protein
Calf Growth Metabolizing Energy Protein Required a Required
(pounds per day) (megacalories per day) (grams per day) (pounds per day) (percent of DM)
0 1.75 28 0.84 8.3
0.50 2.30 82 1.11 18.1
1.00 3.01 136 1.45 22.9
1.50 3.80 189 1.45 22.9
2.00 4.64 243 2.24 26.6
2.50 5.53 297 2.67 27.2
3.00 6.46 350 3.12 27.6
a
Amount of milk replacer DM containing 2.08 Mcals ME/lb DM needed to meet ME requirements, 100 pound calf.
Source: Adapted from Drackley (2000) by H. Chester-Jones, University of Minnesota.
34 calf raisers use commercially-manufactured calf Figure 3.9. Commercial complete calf starter tag.
Chapter 3 starters. Figure 3.9 shows a typical tag from
Calf Nutrition commercial calf starter. Choose products that Dairy Calf Starter
contain between 18 percent and 20 percent crude
protein. Use NRC vitamin and mineral recommen- Medicated
For control of cocidiosis in cattle caused by Eimeria
dations as a guide (See Table 3.6). When selecting a
bovis and Eimeria zuernii
calf starter, consider the following:
• Physical for m . Select whole, coarsely-ground,
form Active Drug Ingredient Examples
cracked, crushed, rolled, steam-flaked, or Decoquinate 22.7 mg/lb
Lasalocid 60-90 g/ton
texturized grains. Do not feed high-moisture Monensin sodium 30-45 g/ton
corn in a calf starter as it often heats and gets
moldy. Guaranteed Analysis
Crude Protein, min 18.0%
• Pellet quality
quality.. Avoid pellets that are too hard or (22% for some plans)
too soft; pellets at both extremes will affect Crude Fat, min 2.7%
intake. Crude Fiber, max 4.5%
Acid Detergent Fiber
• Fines. Calves do not like finely-ground mix-
(ADF), max 9.5%
tures. And, fine feeds tend to cake together when Calcium (Ca), min 0.60%
wet, which deters intake. Calcium (Ca), max 1.1%
• Avoid dusty
dusty,, moldy or of f-flavor feeds.
off-flavor Phosphorus (P), min 0.5%
Salt (NaCl), min 0.3%
• Fiber level. The fiber fraction should have a Salt (NaCl), max 0.8%
particle size of at least 1190 microns. Magnesium (Mg), min 0.3%
• Pr otein sour
Protein ces. Proteins such as urea, raw
sources. Selenium (Se), min 0.3 ppm
Vitamin A, min 10,000 IU/lb
soybeans, feather meal should be avoided in calf Vitamin D, min 2,500 IU/lb
starters. Vitamin E, min 75 IU/lb
• Molasses or molasses-based pr oducts. These
products.
Ingredients
products should be included at between
Grain products, plant protein products, molasses
5 percent and 8 percent of the total mixture. products, calcium carbonate, monocalcium phosphate,
Higher levels can adversely affect handling and dicalcium phosphate, magnesium sulfate, potassium
storage, especially when bags become cold in sulfate, salt, animal protein products, sodium bisulfite,
Apsergillus oryzae fermentation product, Bacillus
the winter months. It is important that starter subtilis fermentation product, dextrose, sodium sulfite,
bags are closed after use to prevent feed from potassium sulfite, vitamin E supplement, ferrous
becoming stale. sulfate, zinc sulfate, calcium iodate, cobalt carbonate,
zinc oxide, manganous oxide, copper sulfate,
• Include B vitamins. ethylenediamine duhydroiodide, manganese sulfate,
• Include coccidostats. Work with your sodium selenite, processed grain products, propionic
veterinarian and nutritionist to supply the acid (a preservative), vitamin A acetate, vitamin D3
supplement, vitamin B12 supplement, riboflavin
proper level.
supplement, niacin, calcium pantothenate, thiamine
mononitrate, mendione dimethylpyrimidinol bisulfite
(source of vitamin K activity), folic acid, biotin,
Encourage Starter Intake pyridoxine hydrochloride, magnesium oxide, riboflavin,
choline chloride, ethoxyquin (a preservative)
Begin offering starter at three days of age. Calves
need encouragement! Place a handful of starter in FEEDING DIRECTIONS: Feed the starter at a rate of
the milk pail or on the calf’s muzzle immediately 1 pounds per 100 pounds body to provide 22.7
milligrams of Decoquinate per 100 pounds of body
after it finishes drinking. Place small handful on weight. If starter includes an ionophore such as
any ledge that holds calf buckets. If facilities do not lasalocid, feed 1 milligram per kilogram (2.2 pounds)
allow a feed pail to be used, use alternative systems of body weight per head daily. Introduce calf starter
at 4 days of age and keep fresh to encourage
such as calf starter bottles to encourage early intake.
consumption. Offer clean, fresh water constantly.
Calves should be eating some starter by five to Avoid feeding hay prior to weaning. Wean calves at
seven days of age. By the second week, calves 4-6 weeks of-age when eating a minimum of 2 pounds
should be consuming around 0.25 to 0.50 pounds per day of calf starter.
Table 3.12 provides a guide for starter intake as age rumen epithelium. While it is not necessary to offer
and weight gain increase. Keep the calf starter as forages at less than eight weeks of age, it is impor-
fresh as possible by only feeding the amount of tant to offer hay to calves by 10 to 12 weeks of age.
starter that calves will consume in one day. Clean Fermented forages can be fed to calves, but is
out moldy or spoiled feed and replace with fresh not recommended for calves less than three months
feed daily (Figure 3.10). Clean and sanitize pails old due to the small amount fed and the difficulty
between each calf. Make sure starter buckets are at of keeping fermented forages fresh.
the correct height to facilitate easy access by the calf
and always provide free choice water.
Water
Water is a critical nutrient for calves and should
Feeding Forages be offered, beginning at three to four days of age. In
Although research has shown that calves can order to digest feed, rumen bacteria must have
consume forages prior to weaning, forages are not water. Milk and milk replacer do not qualify as
necessary until a time when calves consume four to water since they by-pass the rumen. Research
five pounds of a high-quality starter, which is clearly indicated that offering free choice, clean,
typically after weaning. The concept that “scratch” fresh water to calves increases body weight gain and
is needed for early rumen development is a myth, starter intake and reduces weaning age. Calves
unless the calf starter is too finely ground. should consume eight to 10 quarts of water per day
After weaning, begin incorporating hay into the at weaning. Tables 3.13 and 3.14 provide water
diet. The forage should be of good quality—alfalfa or consumption guidelines for varying intakes and
alfalfa/grass hay in the range of 125 to 135 relative temperatures.
forage quality and free of mold. Do not feed haylage Continue free choice water during freezing
or forages with protein levels exceeding 22 percent conditions. Typically, calves will drink water within
as these feeds can contain high levels of non–protein one hour of the milk feeding. Remove water from
nitrogen. Young calves are not very efficient at buckets before it freezes.
processing non-protein nitrogen because the rumen
is not fully functioning. Calves fed forages high in
non-protein nitrogen will often scour and look Table 3.13. Water intake and dry matter
unthrifty with symptoms similar to coccidiosis. consumption.
A general recommendation would be to wean Dry Matter Intake Water Intake
(pounds) (gallons)
calves at five to seven weeks of age and offer hay
1.0 0.75
starting at week eight. Monitor starter consumption
2.0 1.5
to ensure that hay is not being overeaten. Long 5.0 2.0 - 4.0
forage is beneficial to promote the growth of the Source: Adapted from M. Cattell, (1999), by H. Chester-Jones,
rumen muscular layer and maintain the health of University of Minnesota.
36
Table 3.14. Estimate water needs by evaluating temperature, milk and starter intake.
Chapter 3 Starter DM Mean Daily High
Calf Nutrition Milk Intake Water Intake1 Intake Water Intake Temperature Water Intake2
(gallons per day) (gallons per day) (pounds per day) (gallons per day) (F) (gallons per day)
0.00 0.99 0.0 0.00 32 0.36
0.07 0.95 0.6 0.07 41 0.40
0.13 0.91 1.1 0.20 50 0.42
0.20 0.87 1.7 0.33 59 0.46
0.26 0.85 2.2 0.46 68 0.62
0.33 0.83 2.8 0.60 77 0.69
0.40 0.77 3.3 0.73 86 0.83
0.46 0.75 3.9 0.93 95 0.99
0.53 0.73 4.4 1.12 — —
0.60 0.70 5.0 1.26 — —
0.66 0.67 5.5 1.46 — —
0.73 0.66 6.1 1.65 — —
0.79 0.65 6.6 1.85 — —
0.86 0.63 — — — —
0.93 0.61 — — — —
1
Calves consuming 2.2 pounds of starter at 70 F.
2
Weaned calves consuming 2.2 lbs of starter per day.
Source: Adapted from J.D. Quigley, Calf Notes, November 2000.