PRINCIPLES OF ANIMAL TAXONOMY

N U M B E R XX O F THE

Columbia Biological Series

EDITED AT C O L U M B I A UNIVERSITY
 GEORGE GAYLORD SIMPSON

The Museum of Comparative Zoology

Harvard University

Principles of Animal Taxonomy

New York and London

Columbia University Press

SBN 231-02427-4
Library of Congress Catalog Card Number: 60-13939
Copyright © 1961 Columbia University Press
First printing 1961
Fourth printing 1969
Printed in the United States of America
Columbia Biological Series

EDITED AT COLUMBIA UNIVERSITY

GENERAL, EDITOR
L. C. Dunn
PROFESSOR OF ZOOLOGY

EDITORIAL BOARD

Theodosius Dobzhansky
DACOSTA PROFESSOR OF ZOOLOCY
Edward S. Hodgson
ASSOCIATE PROFESSOR OF ZOOLOGY

Howard Levene
ASSOCIATE PROFESSOR OF MATHEMATICAL
STATISTICS AND BIOMETRICS

John A. Moore
PROFESSOR OF ZOOLOCY

William C. Steere
PROFESSOR O F BOTANY

/. Herbert Taylor
PROFESSOR O F C E L L BIOLOGY