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You’ve Evolved!
Evolution and Origin of Biodiversity
Fossil Record
Fossils are the
preserved remains of
once-living organisms.
Fossils are created when
three (3) events occur:
1.) The organism must
become buried in
sediment; 2.) the
calcium in the bone or
other hard tissue must
mineralize; and 3.) the
surrounding sediment http://dailymail.co.u
must eventually harden Source: http://dailymail.co.uk
to form rock. The process of fossilization probably occurs rarely.
Usually, animals or plant remains will decay or be scavenged
before the process can begin. Only a fraction of the species that
have existed are known from fossils. Nonetheless, the fossils
that have been discovered are sufficient to provide detailed Source: Raven, et al., Biology 9
th
Dating Fossils
By dating the rocks in which fossils occur, we can get an Today, rocks are dated by measuring the degree of decay of certain
accurate idea of how old fossils are. In Darwin’s day, rocks were radioisotopes contained in the rock with a process called absolute
dated by their position with respect to one another with a process dating; the older the rock, the more of its isotopes have decayed.
called relative dating; rocks in deeper strata are generally older. Because radioactive isotopes decay at a constant rate unaltered by
temperature or pressure, the isotopes in a rock act as an internal forms occurred in the Eocene Period, 45-55 million years ago
clock, measuring the time since the rock was formed. This is a (MYA).
more accurate way of dating rocks and provides dates stated in
millions of years, rather than relative dates. When fossils are Modern-day members of the Equidae include horses,
arranged according to their age, from oldest to youngest, they zebras, donkeys, and asses, all of which are large, long-legged,
often provide evidence of successive evolutionary change. At the fast running animals adapted to living on open grasslands. These
largest scale, the fossil record documents the progression of life species, all classified in the genus Equus, are the last living
through time, from the origin of eukaryotic organisms, through the descendants of a long lineage that has produced 34 genera since
evolution of fishes, the rise of land-living organisms, the reign of its origin on the Eocene Period, approximately 55 MYA.
dinosaurs, and on to the origin of humans. Examination of these fossils has provided a particularly well-
documented case of how evolution has proceeded by adaptation
Given the low likelihood of fossil preservation and to changing environments.
recovery, it is not surprising that there are gaps or “missing
links” in the fossil record. Nonetheless, paleontologists
(scientists who study fossils) continue to fill the gaps in fossil
record. Today, fossil record is far more complete, particularly
among vertebrates; fossils have been found linking all the major
groups. For example, a four-legged aquatic mammal that
provides important insights concerning the evolution of whales
and dolphins from land-living hoofed ancestors was discovered.
Similarly, a fossil snake with legs has shed light on evolution of
snakes, which descended from lizards that gradually became
more and more elongated with simultaneous reduction and
eventual disappearance of limbs.
The earliest known members of the horse family are species All these changes may be understood as adaptation to
in the genus Hyracotherium, did not look much like horses at all. changing global climates. In particular, during the late Miocene
Small, with short legs and broad feet, these species occurred in and early Oligocene (20-25 MYA), grasslands became
wooded habitats, where they probably browsed on leaves and widespread in North America where much of horse evolution
herbs and escaped predators by dodging through openings in the occurred. As horses adapted to these habitats, long-distance and
forest vegetation. The evolutionary path from these small high speed locomotion probably became more important to
creatures to the workhorses of today has involved changes in a escape predators and travel great distances.
variety of traits, including: By contrast, the greater flexibility provided by multiple
· Size – the first horses were no bigger than dogs; some toes and shorter limbs, which was advantageous for ducking
was the size of a housecat. The modern equids can through complex vegetation, was not longer beneficial. At the
weigh more than half a ton. same time, horses were eating grasses and other vegetation that
· Toe reduction – the first horses had four (4) toes on contained grit and other hard substances, thus favoring teeth and
its front feet and three (3) on its hind feet encased in skulls better suited for withstanding such materials.
fleshy pads. The feet of modern horses have a single
toe enclosed in a tough bony hoof. Homology
· Tooth size and shape – the teeth of Hyracotherium As vertebrates evolved, the same bones were sometimes put to
were small and relatively simple in shape. Through different uses. For example, the forelimbs of vertebrates are
time, horse teeth have increased greatly in length and homologous structures, that is, structures with different
have developed a complex pattern of ridges on their appearances and functions all derived from the same body part
molars and premolars. Accompanying these changes in a common ancestor.
have been alterations in the shape of the skull that
strengthen the skull to withstand the stresses imposed
by continual chewing.
Embryological Development embryological forms provides convincing evidence that they are
both mollusks
Vestigial Structures
Many organisms
possess vestigial
structures that
have no apparent
function, but that
resemble
structures their Source: Raven, et al., Biology 5th edition figure
21.17
presumed
Source: Raven, et al., Biology 5th edition figure 21.15 ancestors had. Human, for example, possess a complete set of
muscles for wiggling their ears, just as a coyote does. Boa
In many cases, the evolutionary history of an organism can constrictors have hip bones and rudimentary legs. The skeleton
be seen to unfold during its development, with the embryo of a baleen whale, which contains pelvic bones, as other
exhibiting characteristics of the embryos of its ancestors. For mammal skeletons do, even though such bones serve no known
example, early in their development, human embryos function in the whale. It is difficult to understand vestigial
possess gill slits, like fish; at later stage, every human structures such as these as anything other than evolutionary
embryo has a long bony tail, the vestige of which we carry relics, holdovers from the evolutionary past. They argue strongly
to adulthood as the coccyx at the end of our spine. Human for the common ancestry of the members of the groups that share
fetuses even possess a fine fur (called lanugo) during the them, regardless of how different they have subsequently
fifth month of development. These relict developmental become.
forms suggest that our development has evolved with new
instructions layered on top. Evidence from Biochemistry and Genetics
The observation that seemingly different organisms may Blood tests show that human blood is more like the
exhibit similar embryological forms provides indirect, but rabbit’s blood than a frog’s blood or an insect’s blood.
convincing, evidence of a past evolutionary relationship. Slugs Furthermore, human blood is more like an ape blood, and is
and giant ocean squids, for example, do not bear much more like an ape’s blood than rabbit’s blood. Notice that the
superficial resemblance to each other, but the similarity of their closer the groups are in structure, the closer their blood
chemistry. This is one (1) more piece of evidence of relationship As a result, the population will gradually come to
and common ancestral origin. include more and more individuals with advantageous
characteristics. In this way, the population evolves and
Modern genetics also provides evidence of evolution. adapts better to its local circumstances.
All organisms use the same genetic code to synthesize proteins.
A universal genetic code is consistent with the idea that all THREE (3) SAMPLES OF NATURAL SELECTION
organisms evolved from a common ancestor that used that code.
· Selection to Avoid Predators
Theories of Evolution
Selection
· Selection to Match Climatic Conditions uptake of insecticide, whereas alleles at the kdr and dld-r genes
decrease the number of target sites, thus decreasing the binding
Many studies of ability of insecticide. Other alleles enhance the ability of the
selection have focused on insects’ enzymes to identify and detoxify insecticide molecules.
genes encoding enzymes.
Often investigators fined the
enzyme allele frequencies Source: http://new.coolclassrooom.org
vary with latitude, so that one
allele is more common in northern populations, but is
progressively less common at more southern locations. An
example is seen in studies of a fish the mummichog (Fundulus
heteroclitus), which ranges along the eastern coast of North
America. In this fish, geographic variation occurs in allele
frequencies for the gene that produces the enzyme lactate
dehydrogenase, which catalyzes the conversion of pyruvate to
lactate (for release of energy when there is not enough oxygen). th
Raven, et al., Biology 9 edition figure 20.7
Biochemical studies show that enzymes formed by these alleles
function differently at different temperatures, thus explaining
their geographic distributions. The form of the enzyme more
frequent in the north is a better catalyst at low temperatures than
the enzyme from the south. Moreover, studies indicate that at
low temperatures, individuals with the northern allele swim
faster, and presumably survive better, than individuals with the
alternative alleles.
The Relationship and Differences of Natural Selection and suggested to him that lineages change gradually as species
Evolution migrate from one area to another.
Natural selection is a process, whereas evolution is the On the Galapagos Islands, Darwin encountered a variety
historical record, or outcome, of change through time. An of different finches on various islands. The 13 species of finches,
example is how giraffes evolved to have longer necks from their although related slightly differed in appearance, particularly in
ancestors. Through the process of natural selection, longer- their beaks. Darwin thought that all these birds had descended
necked giraffes are more successful in survival and pass the from a common ancestor from the South American mainland
long-neck trait on to their offspring. several million years ago. Eating different foods on different
islands, the finches' beaks had changed during their descent.
Natural selection can lead to evolution, but natural
selection is only one (1) of several processes that can result to The diversity of Darwin’s finches:
evolutionary change. The five (5) agents of evolutionary change
are mutation, gene flow, nonrandom mating, and genetic drift. · Ground finches – feed on seeds that they crush in their
powerful beaks
Natural selection can occur without producing
evolutionary change; only if variation is genetically based will · Warbler finches – feed on insects with their smaller and
natural selection lead to evolution. An example is an offspring narrower beaks
who do not resemble its parents because of its experience on
different environments during growth and not because it is · Fruit/bud/cactus finches – feed on fruit/bud/cactus and
different genetically. other insects they attract
Charles Darwin is an English Naturalist who wrote the · Sharp-beaked ground finches – include “vampires” that
book: On the Origin of Species by Means of Natural Selection. sometimes creep up on seabirds and use their sharp beaks
In Darwin’s time, most people believed that the different kinds to pierce seabirds’ skin and drink their blood.
of organisms resulted from direct actions of a creator. Darwin
was a part of a five-year navigational mapping expedition · Wood-pecker finches – pick up a twig, cactus spine, or
around the coast of South America, aboard H.M.S. Beagle. leaf stalk, and trim it into shape with their beaks, and
During the voyage, he studied a wide variety of plants and then poke it into dead branches to pry out grubs.
animals on continents and islands in distant seas. Darwin
repeatedly saw characteristics of similar species varied
somewhat from place to place. These geographical patterns
As basis, for his observations, these assumptions example, a deer that runs swiftly to escape wolves will develop
developed: strong running muscles in its legs. An animal like the giraffe has
1. The Theory of Need to keep reaching up to tall shrubs and trees for food. Each
2. The Theory of Use and Disuse generation of giraffes had a slightly longer neck. After hundreds
3. The Theory of Transmitting Acquired Traits of thousands of years, the short-necked giraffes became the
present-day long necked giraffes. In the case of snakes, the legs
· Theory of Need and short bodies, which were believed to be present in their early
Lamarck believed that organisms change because they ancestors eventually disappeared. It became necessary for
need to. It seemed, according to him, that there is natural snakes to walk through narrow places. They began to stretch
tendency for organisms to make themselves better and more their bodies and to crawl rather than walk.
suited to their environment.
For example, birds developed wings because their · Theory of Acquired Traits
ancestors developed the need to fly. Elephants have long trunks Lamarck explained that an organism acquired a body
for gathering food; lions have powerful claws and jaws for structure (for example, characteristic structure to its offspring.
capturing prey; and deer have long, swift legs for escaping He thought that if an organism acquired a body structure
predators. Therefore, each organism adapts to suit its particular (for example, long limbs, legs, or necks) during its lifetime, it
way of life. could pass on that characteristic structure to its offspring.
Lamarck further believed that changes happened According to him, this is how evolution occurs.
because organisms were able to reshape their bodies in many
new ways. Body, structures, he explained, could be altered- Lamarck and Darwin’s Observations Compared
changed in size or shape depending on the needs of the
organism. Lamarck’s giraffe
For example, birds trying to use their front limbs for 1. Original, short-necked ancestor
flying could eventually change those limbs into wings, the same 2. Keeps stretching neck to reach leaves higher up on tree
way birds with short legs could develop long legs by stretching 3. And stretching
into deeper water. Then, if these animals had no more need for 4. And stretching until neck becomes progressively longer
the structures they had developed through time, these body parts
would decrease in size or eventually disappear.
· Mutation
Mutation is a permanent change in a cell’s DNA. It includes
changes in the nucleotide sequence, alteration of gene
Source: www.tumblr.com position, gene loss or duplication, and insertion of foreign
Darwin’s giraffe sequences. Mutation from one allele to another can
1. Original group exhibits variation in neck length obviously change the proportion of a particular alleles in a
2. Natural selection favors longer necks; better chance to population. Mutation is the ultimate source of genetic
get leaves of trees. Favored characteristics pass on to variation and thus makes evolution possible.
next generation.
3. After many generations, the group is still variable, but · Gene Flow
shows a general increase in neck length. Gene flow is the movement of alleles from one (1)
population to another. It can be a powerful agent of change
because members of two different populations may exchange
genetic materials. Sometimes gene flow is obvious as when
animal moves from one place to another. If the characteristics of
the newly arrived animal differ from those of the animals already
Source: www.tumblr.com there, and if the newcomer is adapted well enough to the new
area to survive and mate successfully, the genetic composition
Five (5) Agents of Evolutionary Change of the receiving population may be altered. Other important
There are different types of winter squash such as kinds of gene flow are not obvious such as drifting of gametes
acorn, butternut, Hubbard, pumpkin, and spaghetti squash. or immature stages of plants or marine animals from one place
All are members of the same species, Cucurbita maxima. to another.
Humans were able to produce such a variety of squash.
Both natural selection and artificial selection are
mechanisms of change in the gene pool of a population. The
key difference is that in artificial selection, humans ensure
that individuals with the more desirable traits are allowed to
reproduce. In natural selection, those individuals who are
best suited to their environment survive and reproduce.
Source: evolution.berkeley.edu
· Selection Anthropoids
This agent is already introduced to you in the discussion · Known as the higher primates; include monkeys, apes,
of the evolution theories. Refer to pages 8 and 9 of this handout and humans.
for the discussion of the two (2) types of selection which are · Diurnal, feeding mainly in fruits and leaves.
natural and artificial. · Live in groups of complex interactions.
· Tend to care for their young for prolonged periods,
The Evolution of Humans allowing for a long childhood of learning and brain
In 1897, Charles Darwin published another ground development.
breaking book, “The Descent of Man”. In this book, he · New World monkeys - anthropoids migrated to South
suggested that humans evolved from the same African ape America
ancestors that gave rise to the gorilla and the chimpanzee. Little · Old World monkeys – a group of anthropoids that
fossil evidence existed at that time to support Darwin's case, but remained in Africa.
numerous fossil discoveries made since then strongly support his
hypothesis. Homonoids
The story of human evolution begins around 65 million · Apes have larger brains than monkeys and they lack
years ago, with a group of small, arboreal (species that live in tails. The living apes consists of the gibbon,
trees) mammals called Archonta. These primarily insectivorous orangutan, gorilla, and chimpanzee. Fossils
mammals had large eyes that were most likely nocturnal (active discovered suggest the first possible Hominoid:
at night). Their radiation gave rise to different types of o Ramapithecus – 8 million year-old jaw with
mammals, including bats, tree shrews, and other primates, the teeth found in India. Fossils indicate they are
order of mammals that contains humans. closely related to orangutan.
o Proconsul – has many characteristics of Old
40 million years ago, the Archonta split in to two (2) groups: World monkeys but lacks tail and has apelike
hands, feet, and pelvis.
Prosimians
· Apes evolved knuckle-walking, supporting their
· Looked like a cross between a squirrel and a cat. weight on the back sides of their fingers.
· Common in North America, Europe, Asia, and Africa. · Hominids became bipedal and walking upright.
· Nocturnal, have grasping digits, and binocular vision. Five (5) to 10 million years ago, the world’s climate began
· An example of prosimian that survive today is the tarsier. to get cooler, and the great forests of Africa were largely
replaced with savannas and open woodland. In response to
these changes, the hominids, which were bipedal arose. In
every case where fossils allow determination to be made, the haircut of bone, on the head to anchor its immense
hominids are bipedal, walking upright. Bipedal locomotion jaw muscles.
is the hallmark of hominid evolution. There are two major · Australopithecus afarensis - nicknamed “Lucy”. The
groups of hominids: The Australopithecus and the Homo: skeleton was 40% complete and over 3 million years
old. The shape of the pelvis indicated that Lucy was
Australopithecus a female, and her leg bones proved she walked
· First hominid upright. Her brain was no larger than that of a
· Walked upright and lived in Africa over 3 million chimpanzee, about the size of an orange.
years ago Homo
· Had a small brain · African origin
· Weighed about 18 kilograms and were about 1 meter · Have larger brain than Australopithecus and most
tall likely used tools
· Dentition was distinctly hominid, but their brains · Homo habilis – means “handy man”. Found with
were not any larger than those of apes stone tools near its fossil. Its skull suggests a brain
· Australopithecus africanus – a skull of a five-year- volume of 680 cubic centimeters that which is larger
old individual, still with his milk teeth. While the than the Australopithecus. Homo habilis was small
skull had many apelike features such as a projecting in stature, with arms longer than legs and a skeleton
face and a small brain, it had distinctly human much like Australopithecus.
features as well (a rounded jaw unlike pointed jaw of · Homo rudolfensis – the skull which was 1.9 million
apes). The ventral position of the foramen magnum years old, had a brain volume of 750 cubic
(the hole at the base of the skull from which the centimeters and many characteristics if human
spinal cord emerges) suggested that the creature skulls. It was assigned to a different species than
walked upright. Raymond Dart, the anatomy Homo habilis because of its substantial brain
professor who discovered the skull, concluded it was expansion.
a human ancestor. Scientists now estimate the skull · Homo ergaster – have even larger brains that Homo
to be 2.8 million years old. rudolfensis, with skeletons less much like the
· Australopithecus robustus – second kind of Australopithecus and more like a modern human in
Australopithecus, which was unearthed in South both size and proportion.
Africa. It had massive teeth and bones.
· Australopithecus boisei – nicknamed the “Nut- Homo erectus
cracker man”. It had a great bony ridge, a Mohawk · Evolved in Africa and migrated from there to Europe
and Asia.
03 Handout 2 *Property of STI
Page 14 of 15
SH1633
· About 1.5 meters tall, a lot larger than a homo including scrapers, spearheads, and hand axes. They
habilis. lived in huts or caves and took care of their injured
· It had a large brain, about 1000 cubic centimeters and and sick. They buried their dead, often placing food,
walked erect. weapons, and even flowers with the bodies which
· Its skull had prominent brow ridges and a rounded suggests that they believed in life after death. This is
jaw like modern humans. the first evidence of the symbolic thinking
· Lived in tribes of 20 to 50 people, often dwelling in characteristic of modern humans.
caves.
· Successfully hunted animals, butchered those using Homo sapiens
flint and bone tools, and cooked them over fire. · They are the only surviving species of the genus
· Survived for over a million years, longer than any Homo, and are the only surviving hominid. The
species of human. skulls are modern in appearance, with high, short
· Java man – Homo erectus found in Java. Structure of braincases, vertical foreheads with only slight brow
its thigh bone indicated it was an excellent walker ridges, and a cranial capacity of roughly 1550 cc,
and skull cap suggested a very large brain. within the range of modern humans.
o Cro-Magnons – fossils of Homo sapiens that had
· Peking man – Homo erectus found in Peking (now
complex social organization and are thought to
Beijing) China that closely resembled Java man.
have had full language capabilities. They lived by
hunting and made beautiful cave paintings
The evolutionary journey to modern humans entered its
throughout Europe.
final phase when modern humans first appeared in Africa
about 600,000 years ago.
REFERENCES
Capco, C. e. (2010). You and the natural world biology . Quezon City:
Three (3) Species of Modern Humans Phoenix Publishing House, Inc.
· Homo heidelbergensis – has more advanced Evolution theories and biodiversity. (n.d.). Retrieved from
anatomical features than Homo erectus such as a http://www.edu.gov.mb.ca/k12/cur/science/found/gr12_bio/unit3.p
df
bony keel running along the midline of the skull, a In darwin's footsteps. (2014, September 4). Retrieved from
thick ridge over the eye socket, and a large brain. http://www.nytimes.com/2014/08/05/science/in-darwins-
· Homo neanderthalensis – short, stocky, and footsteps.html
powerfully built. Their skulls were massive, with Mason, K. A., Losos, J. B., & Singer, S. R. (2014). Biology 10th edition. New
protruding faces, heavy, bony ridges over the brows, York: McGraw-Hill Education.
and larger brain cases. They made diverse tools,