Crown completely exposed (to lateral and vertical light within an inverted cone encompassing the crown).
Titre er
Full overhead light (90% or more of the vertical projection of the the crown exposed to vertical ight; lateral ight blocked
‘cone encompassing the crown).
within some or all of an inverted
Grown position 4
‘Some overhead light (10-90% of the vertical projection of the crown exposed to vertical light)
cr 1
Lateral light (less than 10% of the vertical projection of the crown exposed to vertical; crown it laterally)
Grown position 2.5 Grown position 2.0
(high lateral ight) (medium tera ight)
(ow lateral light)
20 45
\ 0
No direct ight (crown not exposed to direct light either vertically or laterally)
Crown position 4
{no overhead light)
Fig. 1: Crown position/illumination score classification sytem
(adapted from: Dawkins, HC (1958) The management of natural tropical high forest with special reference to Uganda.
Institute Peper, Imperial Forestry Institute, University of Oxford, number 34.
Alder D & Synnott TJ (1982) Permanent sample plot techniques for mixed tropical forest. Tropical
Forestry Papers No 25, Oxford Forestry Insitute, University of Oxford.
Clark DA & Ciark DB (1992) Life history diversity of canopy and emergent trees in a neotropical rain forest.
Ecological Monographs 62(3), pp. 315-344.)n