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Crown completely exposed (to lateral and vertical light within an inverted cone encompassing the crown). Titre er Full overhead light (90% or more of the vertical projection of the the crown exposed to vertical ight; lateral ight blocked ‘cone encompassing the crown). within some or all of an inverted Grown position 4 ‘Some overhead light (10-90% of the vertical projection of the crown exposed to vertical light) cr 1 Lateral light (less than 10% of the vertical projection of the crown exposed to vertical; crown it laterally) Grown position 2.5 Grown position 2.0 (high lateral ight) (medium tera ight) (ow lateral light) 20 45 \ 0 No direct ight (crown not exposed to direct light either vertically or laterally) Crown position 4 {no overhead light) Fig. 1: Crown position/illumination score classification sytem (adapted from: Dawkins, HC (1958) The management of natural tropical high forest with special reference to Uganda. Institute Peper, Imperial Forestry Institute, University of Oxford, number 34. Alder D & Synnott TJ (1982) Permanent sample plot techniques for mixed tropical forest. Tropical Forestry Papers No 25, Oxford Forestry Insitute, University of Oxford. Clark DA & Ciark DB (1992) Life history diversity of canopy and emergent trees in a neotropical rain forest. Ecological Monographs 62(3), pp. 315-344.) n

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