Plant Adaptations • Internal molecular signals in plants include tropisms and photoperiodism. • Tropism: a growth response that results in the curvature of a whole plant toward or away from stimuli due to different rates of cell elongation • Tropisms include: phototropism, gravitropism (geotropism), and thigmotropism Phototropism • Phototropism: plant response to light • Stems and leaves exhibit positive (toward) phototropism • Roots exhibit negative (away) from light Gravitropism (Geotropism) • Gravitropism (geotropism): plant response to gravity • Stems and leaves: exhibit negative gravitropism • Roots: exhibit positive gravitropism Thigmotropism • Thigmotropism: plant response to touch • Ex. Climbing vines; rapid leaf movements • Video link: • https://www.youtube.c om/watch?v=EUUTaiT-b dc Plant Responses to Stress • Drought: plants control water loss by reducing the rate of transpiration by closing stomata • Because soil usually dries from the surface down, drought stimulates deeper root growth to maximize exposure to soil water Plant Responses to Stress • Flooding: overwatered plants may suffocate because the soil lacks the air spaces that provide oxygen for cellular respiration in the roots • Some plants are adapted to wet habitats; they have aerial roots that provide access to oxygen • How do non-aquatic plants cope? Oxygen deprivation triggers apoptosis of some cells in the root, creating air tubes that function as “snorkels”, providing oxygen to the submerged roots. Plant Responses to Stress • Salt Stress: excess NaCl threatens plants for two reasons: 1) roots will lose water; 2) NaCl and certain other ions are toxic to plants in high concentrations • Response: production of compatible solutes that keep the water potential of cells more negative than that of the soil solution. • However: most plants cannot survive salt stress for long • The exceptions are the halophytes: salt-tolerant plants with salt glands that pump salts out of the leaf. Plant Responses to Stress • Heat stress: can denature • Cold Stress: can enzymes decrease the fluidity of • Transpiration is the cell membrane evaporative cooling for • Plants increase the the plant; but could lead proportion of to severe water loss unsaturated fatty acids • Above 40C, plants produce in the membrane to heat-shock proteins that keep the membranes protect other proteins from denaturing. fluid when temperatures drop. Plant Defenses Against Herbivores • Physical defenses include thorns. • Chemical defenses include distasteful or toxic compounds. • Some plants recruit predatory animals to help defend the plant against specific herbivores. Example: parasitoid wasps inject their eggs into caterpillars feeding on the plants. The plants attracted the wasps by releasing compounds from the damaged leaf. Photoperiodism • Photoperiodism: a physiological response to daylength • Ex. Flowering in plants • Photoperiodic plants are classified as long-day plants or short-day plants. This is actually regulated by the hours of darkness. This is called their critical night length. Photoperiodic Plants • Short-day plants: • Long-day plants: flower require a light period when days are 14 hours shorter than 14 hours or longer. to flower • Ex. Spinach, radish, • Ex. Chrysanthemums, lettuce, iris, and many poinsettias, and some cereal crops. soybean varieties. • Generally flower in late • These generally flower spring or early summer in late summer, fall, or winter. Other Plants… • Day-neutral plants: flower when they reach a certain stage of maturity, regardless of day length at the time. • Ex. Tomatoes, rice, dandelions