Topic : Survey of superclass AGNATHA in fish Scientific Classification
• Kingdom ------------ Animalia
• Phylum ------------ Chordata • Subphylum --------- Vertebrata • Superclass ---------- Agnatha Introduction • Agnatha is Greek word meaning no jaws . • Consist of both present (cyclostomes) and extinct (conodonts and ostracoderms) species. • According to recent molecular data they are monophyletic . • 2 groups In cyclostomes still survive . Lampreys and hagfish . Modern Agnathans In addition to absence of jaws modern agnathans are characterized by : • Absence of paired fins • Presence of a notochord • 7 or more paired gill pouches Other Characteristics • Fertilization is external. • They are cold blooded or ectothermic . • Cartilaginous skeleton • Heart contain 2 chambers. Metabolism • Ectothermic • Metabolism slow in cold water • Have no distinct stomach but long gut • Sharp teeth but unable to move this limits their possible food types • Hagfish mostly eat dead animals Body Covering • Body covered with skin with neither dermal or epidermal scales • Skin of hagfish has slime glands for defense against enemy fish Appendages • No paired appendages • Extinct species had paired fins • Most do have caudal fin Reproduction • External fertilization • It is believed hagfish have 30 eggs over a lifetime • Most specie are hermaphrodite • Lampreys are able to reproduce once in their lifetime . After external fertilization in lampreys cloacas remain open allowing a fungus to wnter their intestine killing them . Lampreys bury their eggs resulting hatchlings go through 4 years of larval development before becoming adults . 2 classes of Agnathans • Myxini ----------hagfish ----------- 70 specie • Petromyzontida ---------- lampreys ---------- 38 specie Hagfish Introduction and characteristics Introduction • Eel shaped • Slime producing marine fish • Also called slime eels • Only known living organisms that have skull but no vertebral column • Jawless Body features • Paddle like tails • Cartilaginous skulls • Eyes are simple eye spots • No true fins • 6 or 8 barbels and a single nostril • Horny comb shaped teeth Slime • Slime is released from some 100 glands • Works for defense • Slime clog up enemy fish gills so they die Respiration • Respire by taking in water through its pharynx bringing the water through its pharynx bringing the water through internal gill pouches which vary in number from 5 to 16 pairs • Have a well developed dermal capillary network that supplies the skin with oxygen Nervous system • Some morphologists believe that they do not possess a cerebellum while others believe that it is continuous with mid brain • It is now believed that hagfish neuroanatomy is similar to that of lampreys . Common feature of both is absence of myelin in neurons Eye • Lacks a lens • Eye spot can detect light but none can resolve detailed images Circulation and fluid balance • Are known to have one of the lowest blood pressures among vertebrates • Circulatory system also consist of multiple accessory pumps throughout the body which are considered auxiliary hearts • Only known vertebrates with osmoregulation isosmotic to their external environment Reproduction • Females lay from 1 to 30 tough yolky eggs • Do not have larval stage in contrast to lampreys which have a long one • Take as long as 11 months before hatching Commercial use • As food not often eaten but eaten in Korea • Skin in variety of clothing accessories referred to as eel skin . It produces a durable leather suitable for wallets and belts Lampreys Introduction and characteristics Introduction • Lampreys derived from Latin word meaning stone licker • Feed by boring into flesh of other fish to suck their blood Use in research • Used as model organism in biomedical research • Axons of lampreys are particularly large and allow for microinjection of substances for experimental manipulation In human culture • As food • As pests