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The effects of thinning intensity on the growth and yield of Norway spruce (Picea abies (L.) Karst.)
were investigated in long-term thinning experiments on mineral soil sites in southern Finland. The
measurement period was on average 27 years, and the intensity of the thinnings from below ranged
from heavy thinning (45 per cent of basal area removed) to no thinning. Total stem volume
increment and merchantable volume produced per hectare were the highest on the unthinned plots,
but light and moderate thinning (<30 per cent removed) produced almost as much. Heavy thinning
(>30 per cent removed) reduced the volume increment by about 10 per cent. However, a part of the
total production of unthinned plots was lost through natural mortality. On the thinned plots, natural
mortality was considerably lower compared with the unthinned plots. The average diameter
increment of all the trees, as well as the diameter of the largest trees, clearly increased with
increasing thinning intensity. In contrast, dominant height increment was not affected by thinning.
The stand age at the time of establishment of the experiments had no major effect on the growth
reactions after thinning. Thus, heavy thinning results in earlier thinning yields and a higher
proportion of larger-sized stems at the expense of a somewhat lower total yield.
Introduction
Control of stand density by thinning has been the
major tool in regulating tree growth and improving timber quality. While thinning from below
may increase the merchantable volume of a
stand, usually it does not increase the total
volume increment per unit area (e.g. Assmann,
1954; Carbonnier, 1967; Hasenauer et al., 1997;
Zeide, 2001). Several studies have shown that
volume increment of many tree species does not
decline with decreasing stand density within a
wide range of stand density (e.g. Hamilton, 1981;
Horne et al., 1986). This indicates that thinning
Institute of Chartered Foresters, 2004
Summary
350
F O R E S T RY
and the first results for Norway spruce were published by Ilvessalo (1920). They described the
structure and development of fully stocked,
naturally regenerated pure Norway spruce
stands. A decade later, Cajander (1933) studied
the development of planted, unthinned Norway
spruce stands.
After World War II, selective thinnings, i.e.
removing only the largest trees, were prohibited
and thinnings from below were promoted (Anon.,
1948; Kalela, 1948). At the same time, a research
project was initiated to study the effects of thinnings on the structure and development of
naturally regenerated Norway spruce stands
(Vuokila, 1956; Kallio, 1957). However, the
intensity of the thinnings was low and the
removed trees were heavily suppressed or dying.
In the early 1960s, the thinning intensity in
practical forestry increased mainly due to the
mechanization of harvesting operations. Because
the short- and long-term growth effects of heavy
thinnings were not known, a new research project
was established and growth and yield tables, as
well as thinning guidelines, were prepared for
planted Norway spruce stands (Vuokila and
Vliaho, 1980). During the last decades, growth
models have been developed for forest management planning (e.g. Hynynen et al., 2002).
All the Finnish studies cited above were based
on temporary sample plots (apart from the recent
growth models that were based on remeasured
inventory growth plots) because there was an
urgent demand for growth and yield tables, but
no data available. However, permanent sample
plots provide more reliable results on long-term
growth and stand dynamics. In Finland, the first
thinning experiments were established in the
1920s and 1930s (Ilvessalo, 1932). Some of the
results of these experiments have been published
(Vuokila, 1960, 1977), but the experiments have,
in the main, not fulfilled their expectations due to
natural damage, changes in measurement
methods, subjective classification of many parameters, experimental design, etc. (Vuokila,
1965). In addition, the thinnings applied have
become out-of-date.
In the 1960s, new thinning experiments on
Norway spruce, Scots pine and silver birch (the
so-called HARKAS series) involving more intensive treatments and a better experimental design
were established (Vuokila, 1983). In the 1970s,
T H I N N I N G I N T E N S I T Y A N D G R O W T H O F N O RWAY S P R U C E
Figure 1. Location of the experiments. For explanation of the onset stages, see Materials and methods.
351
352
F O R E S T RY
No.
H100*
6
3
10
4
8
10
8
8
4
5
8
12
3
5
4
4
12
4
8
4
4
34.4
31.6
28.9
31.1
30.2
32.4
32.9
32.7
32.9
30.0
31.0
34.0
36.4
30.2
33.0
33.0
30.0
34.5
34.7
33.0
33.0
1918
1925
1931
1931
1914
1916
1932
1918
1936
1934
1934
1955
1930
1938
1934
1924
1922
1931
1926
1925
1930
1970
1970
1971
1973
1970
1970
1970
1971
1971
1973
1977
1981
1979
1965
1964
1964
1961
1962
1962
1962
1962
1998
1998
1994
1998
1999
1998
1998
1998
1985
1992
2001
1994
1994
1998
1999
1999
1998
1998
1999
1988
1992
3
2
1
2
3
3
2
3
2
2
2
1
3
1
1
2
1
1
1
1
1
4
5
5
5
5
6
6
5
3
4
5
4
5
6
6
6
8
8
8
6
7
Hdom
(m)
No. of
stems ha1
24.3
16.9
13.5
17.0
22.4
23.4
18.1
23.1
16.3
18.0
16.9
12.5
24.1
9.8
12.1
17.4
14.2
14.7
16.0
15.6
14.3
1072
1404
2212
1856
1114
1042
2013
935
2932
1526
2137
2973
1626
3335
1689
1168
2055
3247
2394
2295
3402
Vh001
Vh002
Vh005
Vh009
Vh011
Vh012
Vh013
Vh014
Vh017
Vh040
Vh048
Vh097
Vh098
Ha001
Pu041
Pu042
Nyn1
Nyn2
Nyn3
Nyn4
Nyn5
No. of
plots
Year
of
Establishment
Last
Treatment
No. of
planting
of plots
measured onset measurements
T H I N N I N G I N T E N S I T Y A N D G R O W T H O F N O RWAY S P R U C E
BA
n -1 T
i =1
i * _ BAr i
+ BAt i + 1i /2
T tot
(1)
(2)
353
354
F O R E S T RY
Figure 2. Mean annual diameter increment of all the trees (parts ac) and the 400 by diameter largest trees
ha1 (parts df) on plots of different thinning intensity (U, unthinned; L, light; M, moderate; H, heavy
thinning) and treatment onset stage (early, parts a and d; medium, parts b and e; late onset, parts c and f).
Treatments marked with the same letter are not significantly different (P 0.05). Stem diameter before the
onset of the treatment was used as the covariate in equation 2.
T H I N N I N G I N T E N S I T Y A N D G R O W T H O F N O RWAY S P R U C E
355
Results
The arithmetic mean annual diameter increment of
all the trees on a plot increased with decreasing
stand density (Figure 2ac). The differences
between the thinning intensities were statistically
significant at all treatment onset stages. Because
the differences of the mean increment may be
caused by different numbers of stems, mean
diameter increment was also calculated for the
400 (by diameter) largest trees ha1. The differences between the treatments in the diameter increment of the largest trees were, however, similar to
those for all the trees (Figure 2df). On the other
hand, thinning intensity had no clear effect on
annual dominant height increment (Figure 3).
On plots with light and moderate thinning
intensity, the annual volume increment per hectare
was about the same as on the unthinned plots at
all treatment onset stages (Figure 4). Compared
with the unthinned plots, a heavy thinning intensity decreased the volume increment per hectare
during the whole measurement period by 8 per
cent at the early onset stage. At the medium onset
stage, a heavy thinning intensity decreased the
volume increment even more (11 per cent), but the
differences between the treatments were not
statistically significant. At the late onset stage, the
volume increments among the individual plots
were very variable and, therefore, the differences
between the thinning intensities were not statistically significant.
In addition to the analysis according to fixed
treatment onset stages, the whole material was
356
F O R E S T RY
T H I N N I N G I N T E N S I T Y A N D G R O W T H O F N O RWAY S P R U C E
357
13
10
7
15
U
L
M
H
17
19
21
23
25
27
29
31
Figure 6. Volume increment on plot i during the whole measurement period in relation to the mean volume
increment on the unthinned plots of the same experiment (ivi/ivu) plotted against the relative average basal
area during the whole measurement period (related to that of the unthinned plots, BAi/BAu, see equation 1).
The thick continuous line is a non-linear curve fitted to the data [ivi/ivu = (BAi/BAu)2/{(0.08 + 0.91BAi/
BAu)2}, R2 = 0.16].
16
358
F O R E S T RY
Table 2: Total yield and its structure on plots of different thinning intensity (U, unthinned; L, light; M,
moderate; H, heavy thinning) and treatment onset stages
Early onset
U
Late onset
374a
172b
29a
575b
377a
140c
28a
545b
365a
182a
20a
567a
408a
161a
20a
589a
396a
145a
17a
558a
374a
150a
18a
542a
560a
109a
10a
679a
556a
99a
9a
664a
552a
101a
9a
662a
541a
84a
7a
632a
339ab
94b
8b
441b
314b
45c
4c
363c
349a
145a
12a
506a
354a
103a
8ab
465ab
335a
50b
5b
390bc
286a
46b
4b
336c
526a
80a
7a
613a
480ab
55ab
5ab
540ab
442b
42b
4ab
488b
419b
25b
3b
447b
25a
72bc
20bc
117c
56b
93c
24c
173d
2a
4a
1a
7a
38ab
44ab
6ab
88ab
44ab
90bc
12b
146b
65b
101c
14b
180b
8a
3a
1a
12a
46ab
28b
3a
77ab
90b
52c
5b
147b
108b
62c
5b
175b
8a
5b
1b
14b
7a
2b
1b
10b
10a
35a
7a
52a
15a
19ab
6a
40a
14a
7b
1b
22a
12a
8b
1b
21a
28a
20a
2a
50a
29a
14ab
1ab
44ab
17a
6bc
1ab
24b
13a
0c
0b
13b
Total volume before the onset of the treatment was used as the covariate in equation 2.
Treatments marked with the same letter (in the same line and within the onset stages) are not significantly
different (P 0.05).
Discussion
Total volume production per unit area was
highest on the unthinned or lightly thinned plots,
irrespective of the treatment onset stage.
However, light and moderate thinning had
almost no effect on volume increment, or total
volume produced during the long measurement
period. Only heavy thinning resulted in a clear
reduction in volume increment. Even though the
basal area of the heaviest treatments was, on
average, 5570 per cent below that on the
Medium onset
T H I N N I N G I N T E N S I T Y A N D G R O W T H O F N O RWAY S P R U C E
359
Figure 7. Cumulative volume of dead trees on plots of different thinning intensity: (a) early; (b) medium;
(c) late onset of the treatment. For explanation of the symbols, see Figure 2.
360
F O R E S T RY
T H I N N I N G I N T E N S I T Y A N D G R O W T H O F N O RWAY S P R U C E
361
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F O R E S T RY
Acknowledgements
We are greatly indebted to many technicians, especially
to Timo Siitonen, who have maintained and measured
the experiments. We also thank Professor Jari Hynynen
for his support and advice, Pentti Niemist and Dr
Anssi Ahtikoski for valuable comments on the manuscript, Dr Helena Henttonen for the annual increment
indices, John Derome for revision of the language and
Sointu Nenola for drawing Figure 1.
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