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American Antiquity
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ARTICLES
Archaeological theory has traditionally presupposed the existence of "battleship curves" in stylistic
understanding about what governs the width (variant frequency) or length (variant lifespan) of th
these variables, we propose that there is a testable difference between independent decisions, unbi
biased transmission in cultural evolution. We expect independent decision making to be represented b
tribution of variant prevalence in the population. In contrast, unbiased transmission tends to be chara
law or log-normal distribution of prevalence, while biased transmission should deviate significantly f
The difference between these categories may be fundamental to how cultural traits spread and per
analytical predictions for unbiased transmission, we adapt a model of stochastic network growth t
demonstrating the inherent nonlinearity in unbiased transmission, can explain why afew highly popular
to emerge in the course of cultural evolution. For the most part, this model predicts the frequencies
remarkably well over a 400-year span of Linearbandkeramik settlement in the Merzbach valley. Be
quencies of actual motifs are somewhat less than predicted by our unbiased transmission model, w
formist, or pro-novelty, bias in the later phases of the Neolithic Merzbach Valley.
R. Alexander Bentley and Stephen J. Shennan * AHRB Centre for the Evolutionary Analysis of Cultural Behaviour,
Institute of Archaeology, University College London, 31-34 Gordon Square, London WC1-OPY, United Kingdom
459
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460 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
learning, which could not be accounted for by However, there are limits to strictly biology-
genetic transmission or continuity of environment based models of cultural evolution because the ana-
(Boyd and Richerson 1985:46-55). Cultural tradi- logues of the processes that affect gene frequencies
tions, like particular gene distributions, can persist in populations-mutation, selection, and drift-do
in the face of changing environmental conditions not exhaust the set of processes that affect cultural
to which different practices would be better adapted attribute frequencies. It is becoming apparent that
(Boyd and Richerson 1985:55; Edgerton 1971; cultural transmission processes are much more cen-
Guglielmino et al. 1995; Henrich and Gil-White tral to sociocultural evolution than has been
2001; Henrich et al. 2001). assumed before (Boyd and Richerson 1985; Hen
In order for archaeologists to study human rich and Boyd 2001; Henrich and Gil-White 2001
behavior and its history from a Darwinian stand- Henrich 2001). The simplest form, termed unbiase
point, they must address two complications that transmission by Boyd and Richerson (1985), occurs
the existence of cultural traditions poses for evo- when each individual acquires his/her behavior
lutionary change. The first is that differences in simply by copying from another individual within
people's survival and reproductive success can the population. If the process is purely unbiased
occur through natural selection on their cultural transmission, then children are as likely to copy
traditions, not simply on their genes via their genet- other people as to copy their own parents. The result
ically inherited dispositions. The second concerns of unbiased transmission, as we will explore in
cultural transmission, in that the frequencies of cul- detail in this paper, is that each individual is equally
tural attributes may change through time, not only likely to be copied, while each variant is copied in
as a result of natural selection affecting people's proportion to its frequency.
survival, but also as a result of their conscious and A rather different process, which affects the fre-
unconscious decision making. The aim of this paper quency with which information is handed on to
is to examine quantitative patterns that distinguish future generations, or even to next week, and there-
different routes of cultural transmission. In pre- fore affects the frequency of cultural attributes in
dicting quantitatively how a few highly popular the population through time, is prestige-biased
styles may be expected to emerge in the course of transmission (Henrich and Gil-White 2001; cf.
unbiased cultural transmission, we offer a quanti- indirectly biased transmission of Boyd and Rich-
tative null model, deviations from which can be erson 1985). Prestige-biased transmission takes
meaningfully explained as biased transmission. place when an individual adopts the cultural
attribute of someone who appears to be more suc-
Modes of Transmission
cessful in terms of some locally accepted criterion,
In considering the importance of transmission in if the attribute concerned is not actually the
even
cultural evolution, our focus is to model thereason
evo- for their success. An example is copying the
lution of neutral traits through time, rather mode
than of speech or dress of successful musicians
those strongly subject to natural selection. In orthe
actors. This may have a genuinely positive result
absence of selection forces, neutral traits are sorted
in that it can be an efficient shortcut to becoming
successful oneself. Henrich and Gil-White (2001)
by drift and random differences in transmission
(e.g., Braun 1991; Dunnell 1978; Lipo et al. 1997;
argue that prestige is equivalent to "the size and lav-
Neff 1992; Neiman 1995; O'Brien 1996; O'Brienishness of a given model's clientele." Because this
size and lavishness is the main cue that others use
and Holland 1995). The large-scale consequences
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 461
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462 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
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CULTURAL TRANSMISSION AND NETWORK GROWTH
Ben 463
1
dent decision
O boot and macro quits in any g
more than the
0 file
0.1 - shows that f
._
declined roug
o health agencie
0-
.01
ing campaigns
0.01 0 tic"half-life"
0
[] subpopulation
depend on fac
so on. The dec
0.001 I I I
more educated
0 10 20 30 range from ab
Survival elor's
time degree
(months) t
school educati
Figure 1. Survival times
ferences of
acro c
Internet (after Pastor-Satorr
nential decay
Figl). Only the y-axis is logar
function plots making
as a occur
straight line
In sum, it is f
exponential distribution
exponential d
viruses have a characteristi
highly simplif
months, depending
which is onsubst th
Satorras and tial Vespignanidecay in d 2
nential lifespan
texts. distribut
equation (1) for independ
Case 2: Unbiased Transmission
because a computer (or com
independentlyNeiman's (1995) Model. Neiman (1995)
whether orofferedn
a
An example of
model for use as a this expo
null hypothesis representing the
modem culture is
effects of drift alone, in thein
absence ofthe
other forces
whole, such as natural selection.
smoking may Neiman's treatment
be is an ch
100-
0~
a)
0
o
o
0-
E
E -
-71;~7rI
0 03
-- Some oollege
10
-- Bachelor's degree I I
10 ,
1970 1980 1990 2000
1965 1975 1985 1995
Year Year
Figure 2. The decline of smoking in the U.S. (a) Male Smokers as a percentage of U.S. population in the past 35 years. (b)
Male smokers in the U.S., as classified by education level. The plot has a logarithmic y-axis.
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464 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 465
10 p=0
(a)
a 8
3:
a
_6 6
:t4
2
0
0 100 200 300 400 500
lime step
p = 0.05
10 (b)
8
6 6
0
0 100 200 300 400 500
lime steps
= 0.10 (c)
10
S
us8
a
16
it 4
Figure 3. Assemblage diversity in Neiman's (1995) model, run for 500 time steps. The runs shown represent three dif-
ferent innovation rates: (a) no innovation; (b) pt = .05; and (c) I' = .10. The model shown is for N = 20 individuals.
is a much more difficult task because (1) each indi- geneity F,, which can be solved for), we are left to
vidual can acquire a different variant in each time run it with a few different parameters and observe
step and (2) the frequency of each variant can the results. Figure 3 shows the number of different
change in every time step. Like many highly sim- variants in each time step of a run of the neutral-
plified models of complex dynamics (e.g., Bak trait model using N = 20 individuals and three dif-
1996; Paczuski et al. 1996; Turcotte 1999), the neu- ferent mutation rates, p = 0, p = .05 and p = .10.
tral-trait model is much more complicated than it The model is surprisingly complex, and we note a
would seem from its simple setup. few general characteristics. The first is that it never
Since we are not aware of an analytical solution quite reaches an equilibrium state such that the
for the neutral-trait model in terms of the distribu- homogeneity F, is constant from one time step to
tion of variant frequencies (as opposed to the homo- the next. There appears to be nominal equilibrium,
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466 AMERICAN ANTIQUITY [
1~
0.1 -
'
0.01 -
I
-
._
ca
0.001 - -A
o
?0 A
0.0001 "
9
2
A cl
0.05
0.00001- *
A D1
A 0.10 x
*
0.15
0.000001-
x 0.20 C
0.0000001 I I l
Number of copies
Figure 4. Distribution of the number of variant copies after running Neiman's (1995) model with innov
time steps. N = 20 individuals, beginning with 20 different variants. For the first run, one innovation per tim
introduced (|p = .05). In subsequent trials, the innovation rate was increase by introducing two (pi = .10), t
and finally four (.t = .20) innovations per time step. Each distribution follows an approximately power-law fu
the slope (power-law exponent) increasing with innovation rate. The power-law slope for It = .05 is -1.67
power law fit vs. r2 = .734 for exponential fit); for jI = .10 is -1.75 (r2 = .975 for power law, .787 exponential)
is -1.99 (r2 = .971 for power law, .786 exponential); for ,t = .20 is -2.00 (r2 = .957 for power law, .863 expo
visualization, the dashed line shows a power-law fit to the it = .05 data; the dashed gray line shows an expo
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 467
0.1 -e,.
'"O.,
0.01
0..
.0?
0.001 - 0
2-
0
*.
o 0.0001 -
0 Female names 0
[1 Male names
0.00001 -
. ..... exponential
0.000001 - 0
0.0000001
Frequency (%)
Figure 5. Frequency distribution of first names, from 1990 U.S. Census data. The popularity of a name as a fraction of
the U.S. population. On this log plot, the power-law slope for males names is -1.10 (r2 = .992), and for female names -1.24
(r2 = .976). For comparison, the dashed gray line shows an exponential function fitted to the female name data. For con-
fidentiality reasons the U.S. Census does not provide the data for the most original names, so the distributions are trun-
cated on the left.
a power law for four different innovation rates, p resemble the unbiased copying of variants in the
= .05, p =.10, p =-.15 and p = .20. A higher inno- neutral-trait model. If so, we would expect a power-
vation rate makes the slope steeper, such that high law distribution of name frequencies. Figure 5
frequencies are less likely. shows the frequencies of both female and male first
Most of the plots we show in this paper display names as listed in the 1990 U.S. Census.7 Each dis-
the probability distribution, which is equivalent to tribution-appearing as a straight, downwardly
the fraction of the sample expected to fall within sloping line on the log-log plot-is an approximate
each bin of a histogram. Because most of these dis- power law overall. Interestingly, the slope is steeper
tributions have long tails, we bin the data using log- for female names than for male names, in the same
arithmic bin sizes. In order to plot the probability way the slope is steeper for the higher innovation
distribution using logarithmic bins, the number rate in the model (Figure 4). This suggests that the
within each bin must be normalized by the increas- innovation rate is higher for girls' names, perhaps
ing sizes of the bins. For example, with the top of because girls in American culture are given a wider
the bins at 1, 2, 4, 8, 16... we must then divide the variety of original names than boys, whose names
contents of each bin by 1, 1, 2, 4, 8, . . ,respec- may be more traditional. Further discussion of this
tively. This way, the cumulative version of the interesting topic would lead beyond the focus of
power law in equation (6) still forms a straight line this paper (but see Hahn and Bentley 2003).
with slope -D on a log-log plot. In contrast, an An Analytically Predictive Modelfor Unbiased
exponential probability distribution plots as a Transmission: The Stochastically Growing Net-
straight line when only the y-axis is logarithmic, work. What Neiman (1995) and we have demon-
so it is easily distinguishable from a power law. strated for the neutral-trait model is something
With our observation that the neutral-trait model rather counterintuitive: over time, unbiased trans-
leads to a power-law distribution of variant fre- mission naturally leads to a highly skewed distri-
quencies, we can consider a cultural example in the bution of variant frequencies. If we want to use
way people choose names for their children. Babies unbiased transmission as a null model, we need to
are often named after real people, and if the chance be able to predict explicitly the expected distribu-
a particular name is chosen is proportional to its tions over time. This would be useful because
frequency in the population, the process may biased transmission skews the frequency distribu-
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468 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 469
0 .1 I..""
0.1-
0.01
._
0.001
c,
0
o
0.0001 -
PO
.
0.00001 O DLA
.
* SOC
- --exponential .
0.000001 -
0.0000001
Figure 6. The citations of journal articles on particular topics. The x-axis shows the number of other papers that had
cited each article by the end of 1999. One distribution is for articles (N = 1131) with "self-organized criticality" (SOC)
in the title or keywords from its origin in 1987 (Bak et al. 1987), and the other for articles on "diffusion limited aggre-
gation" (DLA) from its origin in 1981 (Witten and Sander 1981). Despite the two different topics and dates of introduc-
tion, the probability distributions are similar, each with an approximately power-law tail, with slope (exponent) - 2.1 ?
.2 for SOC, and -2.1 ? .2 for DLA. The dashed gray line shows an exponential function for comparison.
variations in more specialized subfields, and each ution of the lifespans of pottery styles, exemplified
new article cites at least most of the papers that by brand names from seventeenth- to early twen-
inspired it. The distribution of the number of times tieth-century New York State, exhibit a "fat tail,"
these papers were cited through 1999 exhibits a i.e., approximately power law or log normal in form
power-law tail (Figure 6), indicating that a scale- for lifespans longer than about 20 years (Figure 7).
free network of papers and their citations has If lifespan is one measure of success, it would
formed (Bentley and Maschner 2000). The reason appear that pottery brands in historical New York
the left-hand side of this distribution is not a power State were prone to the rich-get-richer effect. Pres-
law is the least-cited articles tend to be the newest, tigious styles like Wedgewood, while high-quality
and the production of new articles has fallen behind to be sure, benefited also from their very promi-
what is required to retain the power law all the way nence, and therefore had especially long life spans
to the left. This should become clearer when we (Bentley and Maschner 2001). Hence Figure 7 sug-
discuss the stochastic growth model below. Nev- gests that pottery consumption in the New York
economy was dominated by unbiased transmis-
ertheless, in this network the rich can get richer sim-
ply through unbiased transmission. While Bak et sion. In the next section we adapt a detailed model
al. (1987) continue to be cited by almost all new relevant not just to a static end result as in Figure
papers coming out on self-organized criticality,
7, but also to the dynamics of the formation of these
distributions.
each new report is lucky to be cited once or twice
in the future. Figure 6 shows that this pattern is not The Stochastic Growth Model of Adamic and
Huberman (2000). In this subsection, we describe
unique, as virtually the same distribution has also
a mathematical model of stochastic network
formed in the citations of "diffusion limited aggre-
growth, which we believe to be applicable to unb
gation," another highly influential physics topic of
the last twenty years (Witten and Sander 1981). ased transmission. While the model was design
We can extend the model from this academic to simulate web pages on the Internet, it is gener
enough to apply to people in a society, or hous
citation example to a market economy by explor-
holds in a community. As mentioned above, in orde
ing whether or not the success of brand names fol-
lows a similar stochastic growth process. Bentley
to form a scale-free network-a special case of sto
and Maschner (2001) have shown that the distrib-
chastic network growth-a network of intercon
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470 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
*..,
0.1
.
0.01
:_
,^
PO
,0
I 0.001
0.0001
0.00001
10 100 1000
Lifespan (years
nected Adamic
agents an
must
ple guidelines
g is (Bara
norma
new connections
mally (e.
dist
sites) are give rise
continually t
nectivity
preferentially attac
nodes, andprocess
(2) the in
ne
population of
the agent
probab
However,tions
because th
after
ential
attachment, ca
growing, let
P(ns) _ us
1 (lnn- turn
gt)2
chastic n s t2n 2 26J tgrow
network
connected nodes, A
model the
where gogrowth
is the mean value of g over time, with in
stan-
a node has at
dard deviation time
s. The log-normal function in equa- t
nections it already
tion (8) is not h
as complicated as it may seem at first.
In fact, the log-normal looks like a normal distrib-
n,(t + 1) = ns(t)
ution when the x-axis is made logarithmic.1' An
where g(t) expresse
important aspect of equation (8) is that both the
between agents
position got and width o2t ofare ad
the log-normal peak
can increase with time t, such that
fluctuate it moves outward and
rando
spreads out with
connections in time on equatio
a logarithmic scale.
tiplied by In applying
the this to prehistoric cultural evolution,
curre
This means that
equation (8) predicts that, over time, asa log-normalt
time, thedistribution
resulting
of frequencies will develop among vari- f
copies ofants introduced
a at the
pottery same time into a cultural net-
work dominated
connected node by unbiased transmission.
(e.g., It also
than for
predictsa poorly
that over time their accumulation will
process increase as a whole (the peak moves outward) and
described b
plicative because,
the disparity among them will increase (the ovpeak
the spreads out).
growth Because the distribution
rate g spreads
mu out
develop on
in a logarithmic
the scale, the disparity
distbecomes one
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 471
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472 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
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CULTURAL TRANSMISSION AND NETWORK GROWTH
Bentley 473 an
a
25 - b 25 .
20 - Pess 15
20 -
15 15 - qo
5 5-
II I 0
co
I
(F GSoCI I
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474 AMERICAN ANTIQUITY [Vol.
11,
C)
1
= 0.71
Ce *.
00
r2 .78 100
2 *
tL 5
a
et r20=0.71
10
0 5 10 15 20 0 5 10 15 20
Figure 10. (a) Comparison of the number of houses occupied during eac
motif copies during that phase. (b) comparison with the frequency of the m
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CULTURAL TRANSMISSION AND NETWORK GROWTH
Bentl 475
0.1
0.01
^ 0.001
0.000
0.00001
0.000001 I I
Copies
Figure 12. Frequency distribution of Merzbach pottery motifs. The probability distribution looks somewhat like a power
law (black line), but the data are better fit by a log-normal function (dotted gray line). An exponential function (dashed
gray line) is shown for comparison.
It should be no surprise that there are more Applying the Stochastic Network Growth Model
copies of older pottery motifs, produced over a to the Merzbach Data
longer period of time, or motifs from times of larger
populations to produce them. What is not neces- We can now model the change in the Merzbach
sarily expected is that these relationships would be motif frequencies by using the model of Adamic
nonlinear. Following our discussion above, our and Huberman (2000). As their model demon-
model of independent decisions implies that each strates, the log-normal distribution such as we see
band type had an equal probability of being copied in Figure 12 may result from the same rich-get-
and that the total number of copies of a variant richer process that produces a power law or, as dis-
should be linearly proportional to its lifetime. cussed above, a combination of log normals if there
Instead, the nonlinear relationship in Figure 11 sug- is not continual increase in the number of agents
gests that older, established styles were copied in in the network. During the LBK era in the
preference to newer styles. Furthermore, Figure 12 Merzbach valley, prehistoric populations rose, sta-
shows a clear log-normal distribution of band-type bilized, and then declined over a period of about
frequencies in the Merzbach valley for its entire 400 years. Hence we would not expect the distrib-
occupational time span. In contrast, the prediction ution in Figure 12 to be a power law because the
for independent decisions is an exponential decay network did not grow continuously throughout the
in the distribution of frequencies (equation [2]). period. Instead, it could be a mixture of log-nor-
Hence we are lead to reject independent decision mal distributions that developed according to the
making as a hypothesis. Instead, the transmission model of Adamic and Huberman (2000).
process appears to be analogous to stochastic net- We can adapt the model of Adamic and Huber-
work growth, in which older, established motifs man (2000) to explain the evolution of pottery motif
were reproduced in proportion with their existing popularity in the Merzbach Valley. In order to do
frequency. This may seem to contradict the result this, we assume that each pottery motif is a node
of Shennan and Wilkinson (2001), who found a pro- of the network, and copies of each motif are con-
novelty bias in the later Merzbach phases. How- nections leading toward that node. The number of
ever, as we discuss below, this result can be used "connections" leading toward each node is equiv-
to explain the difference between the null hypoth- alent to the number of copies of the motif. Sto-
esis of stochastic network growth and the actual chastic network growth implies that new
Merzbach data. connections (motif copies) are preferentially
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476 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
attached to
data.
(copied
Final
nodes (motifs).
go(i) = anh
Mathematical
ber ofApp
occ
can attempt
motif
to mode
that
to equation (8), whi
variables (s
normal we arrive at a model for the distribution of
function motif
that
with frequencies
time. We begun in the same phase:
apply
ested in the detailed
( -\2
r (lnn
section,
P(ns) = 2 exp - anht (10)
comparing
delineate the
nS t2n(2 2c2 t
pottery
in which they were
tion (8) where, to reiterate:
represent th
begun in ns chronolog
is the number of copies of the pottery motif
mize the accumulated
a prioriby time t info
we start nh is the average number
with the of houses in
simthe val-
maintainedley during the time from
t t
which they a and 6 are parameters
were whose values we need
in
occupationto determine. of phas
result of We now look to estimate the parameters a and
incorporat
Thus the a2 from the data. A prediction of equation (10) is
lifetime, d
motifs begun
that the position of the peak in pha
of each log-normaldis-
9, for example. We
tribution in motif frequencies will depend on anht , t
of houses
which isoccupied
just a multiplied by the cumulative num- d
of the study area
ber of houses occupied over a time span t. In Fig-co
km2, so ure 13a we see a can
we good linear relationship
be (r2 = .92) r
porary between nht and ln(ns), where the bar over the
households w
and of all latter
thequantity indicates decorat
the average value of ln(ns).
Since For a log-normal distribution,
larger populati ln(ns) is the posi-
tially tion of the peak
copy a on the x-axis. Hence Figure 13a
particu
growth shows that the log-normal
rate g(i), peaks moved out ana
at a
constant rate, just as predicted
proportional to by equation
the (8). We p
the phase. Using
find the constant a from the slope of the line the
in Fig-
estimate ure 13a, such that ln(ns) = .0466 nht. We also
population
a is a constant.
expect the width of the peak toBecaus
increase with time,
history of
with variance g, we
(2t, so we plot the do
variance in ln(ns)
over vs. t (Figure
time. To 13b). Although the linearity is not as a
find
tion (8), we
good as in Figure will esti
13a, the slope of the best-fit line
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 477
5-
(b)
(a)
4 -
sCa
y = .0466x
R2 = .93
2-
I
.C 1
1-
O0
5 10
O0 50 100 150
Figure 13. (a) Comparison of nht versus the average value of ln(ns) for the Merzbach pottery motifs. The linear relatio
ship between the average value of In(ns) and nht is just as predicted by equation (9). The line has been fitted through t
origin, giving the simple relation In(ns) = .0466nht. (b) Comparison of motif lifetime t versus the variance of In(
motifs begun in the same phase. Equation (9) again predicts a linear relationship (through the origin), for which the b
fit gives variance [ln(n)] = .261t.
0.1
0.01
>-4
0.001
I
0
2
sv
0.0001
, 7 ' 0 "P
- - % --- ". X
0.00001
Copies
Figure 14. The modelled distribution of Merzbach pottery motif frequency, after equation (12). As the phases get older,
the peak of the log-normal distribution moves toward the right and also spreads out. Note that since the spread is along
a logarithmic axis, motifs from phases 6 or 7 are likely to be tens or hundreds of times more frequent than those from
phases 11-13.
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478 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
0.1
8 or 12). Since the data are limited, we had to com-
bine phases 9-13 to have enough data for a his-
* A
0.01
A togram. We want to keep our model relevant to the
*x archaeological data that we were forced to com-
bine like this, so for comparability we generated
^ 0.001 +~~* ~the model distribution by binning the modeled
a * X probabilities in the same way,13 and also combin-
0.0001
ing them to get a prediction for phases 6, 7 and
9-13. Figure 16 compares the model and the data
Phase 6 and shows that the model and data are close for
0.00001
Phase 7 phase 7 and phases 9-13.
A
Phases 9-11 With phase 6, however, the model does not fit
0.000001
well, as the actual distribution has a higher pro-
1 100 10oo 1000o portion of low-frequency motifs. Even so, the data
Copies from phase 6 confirm the general expectation of our
model that the log-normal peak moves outward
Figure 15. Frequenc
motifs for three time y distribtion
spans of Merach pttey and spreads out as the phases get older. For this rea-
by initial phase.
son, the only motif with over a thousand recovered
copies comes from phase 6. The reason for the mis-
his process is that the most-pop- match between model and data is that many band
important about tf
ular motifs from older phases are likely to have types originating in phase 6 went extinct well before
thousands of time -s more copies than the popular phase 15 (Figure 17), violating our model assump-
motifs from younlger phases. tion that all motifs lasted until the end of the occu-
Now we can c ompare the model and the data. pation. We note that the pro-novelty bias observed
Because there are not enough data from each phase by Shennan and Wilkinson (2001) for the late occu-
to show a distribi ution, Figure 15 shows the fre- pation phases helps to explain this, although sev-
quency distributicns for motifs begun in phase 6, eral of the band-types from phase 6 were abandoned
phase 7, and phases 9-13 (no motifs from phases already in phase 7. Whatever the reason for this, if
0.1
0.01
0.001
a
;.
?
0.0001
0.00001
0.000001
Copies
Figure 16. The modelled distributions versus the actual data from the Merzbach pottery. Model 1 assumes all band types
survived until the end of the occupation at phase 15. The data on several phases had to be combined in order to have
enough data for a histogram. For comparability, we calculated the modelled distributions by combining the same phases
and binning data the same way as for the actual data (see note 7).
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 479
Huberman (2000) and the later-phase Merzbach copied preferentially, such that the probability that
a cultural trait is chosen simply depends on how
pottery data, considering that the model uses only
two variables, the phase begun and the average prevalent it is already. By being more available
0.1
0.01 --'
0.001 -
0.00001
Model phase 6
Mll
Model phase 7
Model phases 9-13
0.000001
Copies
Figure 18. Model 2 vs. actual data from the Merzbach band types. Model 2 is the same as Model 1 in Figures 15 and 17
except that the actual lifespans of the band types were used rather than assuming all survived to phase 15.
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480 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
IM/
;,IvY/t/ _
I>// /
..."
K211=
1,1/
1'
- -
Figure 19. Modeling a case of unbiased transmission as stochastic network growth. In our example from Neolithic
Germany, we model LBK households as the nodes of the network, and copies of pottery motifs, which we take to repre-
sent the different household identities, as connections leading toward each node (which can be one's own household, as
represented by arrows curving back again). LBK motifs after Frirdich (1994), LBK longhouse after Liuning (1984).
and/or visible, more trusted and/or sought-after, a sion, as exemplified by trial simulations of
popular cultural trait may gain an advantage Neiman's (1995) adaptation of the neutral-trait
through unbiased transmission simply by virtue of model, generates an approximately power-law dis-
its being popular. In addition, the most popular tribution of variant frequencies. In order to make
traits are usually the ones that got into the growth quantitative predictions, this effect can be modeled
process early. A modern analogy is the success of as a process of stochastic network growth after
a brand of cola begun in the early 1900s vs. a new Adamic and Huberman (2000), which also gener-
brand begun in the 1980s-they both taste the same, ates a power law or log-normal distribution of vari-
but there is a distinct advantage to having gotten ant prevalence. In the stochastic network model,
started first. those agents with more of a particular attribute are
In this paper, we have explored models of the more likely to acquire even more of it in the future,
evolution of variant frequencies evolving under at a rate proportional to what they already have.
independent decisions and under unbiased trans- Conformist-biased transmission implies that there
mission. Independent decision making is expected is an extra preference for the most popular trait, in
to exhibit an exponential distribution of variant addition to this frequency dependence. Conformist-
prevalence in the population. Unbiased transmis- biased transmission should produce a highly
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 481
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482 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 483
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484 AMERICAN ANTIQUITY [Vol. 68, No. 3, 2003
Neff, H. Notes
1992 Ceramics and Evolution. Archaeological Method and
Theory 4:141-193. 1. After having thought of this example, we discovered
Neiman, F. D. that Henrich (2001) had also used Michael Jordan to make
1995 Stylistic Variation in Evolutionary Perspective: Infer- the same point. Our choices of the same person for an illus-
ences from Decorative Density and Interassemblage Dis- trative example demonstrates prestige-biased transmission in
tance in Illinois Woodland and Ceramic Assemblages. itself!
American Antiquity 60:7-36. 2. The Taylor series is a method of approximating a func-
Newman, M. E. J.
tion by the sum of its derivatives. The function e-x has the
2001 The Structure of Scientific Collaboration Networks.
Taylor series expansion
Proceedings of the National Academy of Sciences USA
98:404-409. 2 3
x x2 x3
O'Brien, M. J. e- =1 ---+ +...
1! 2! 3!
1996 The Foundations of Evolutionary Archaeology. In
If pp.
Evolutionary Archaeology, edited by M. J. O'Brien, x is small, then the terms on the right are ne
17-23. University of Utah Press, Salt Lake City. have e-x - 1 - x.
O'Brien, M. J., and T. Holland 3. The distribution has leveled off in recent years, and the
1995 The Nature and Premise of a Selection-based Archae-
distribution may be better described as S-shaped (Henrich
ology. In Evolutionary Archaeology: Methodological
2001). Nonetheless, most of the decline has been exponential.
Issues, edited by P. A. Teltser, pp. 175-200. University of
4. Neiman uses a term for the effective population, Ne
Arizona Press, Tucson.
Paczuski, M., S. Maslov, and P. Bak which accounts for the fact that not all individuals in the pop-
1996 Avalanche Dynamics in Evolution, Growth, andulation are potential carriers of a certain behaviour. For sim-
Depinning Models. Physical Review E 53:414-443. plicity, we refer only to a population N without losing the
Pastor-Satorras, R., and A. Vespignani character of his model.
2001 Epidemic Spreading in Scale-free Networks. Physi- 5. In this case, the chance of not being copied faithfully
cal Review Letters 86:3200-3203.
by a particular person is [1 - (1 - I)/N], and the chance of
Plotkin, H. C.
being ignored by everyone is [1 - (1 - p)/NIN. Therefore, the
1994 The Nature of Knowledge: Concerning Adaptations,
probability Ps that anyone's own personal variant (not some-
Instinct and the Evolution of Intelligence. Allen Lane, Lon-
don.
one else's identical version) survives to the next time step is
the same as the chance that it is not totally ignored:
Pocklington, R., and M. L. Best
1997 Cultural Evolution and Units of Selection in Repli-
cating Text. Journal of Theoretical Biology 188:79-87.
Shennan, S. J., and J. R. Wilkinson
2001 Ceramic Style Change and Neutral Evolution: A Case
If N is large, then (I - u)/N is small and we can make the Tay-
Study from Neolithic Europe. American Antiquity
lor series approximation (see note 2) that 1 - (1 - p)/N - e("-
66:577-594.
l)/N and equation (3) becomes:
Simon, H. A.
1955 On a Class of Skew Distribution Functions. Biomet- p _1 N
rica 42:425-440.
p,1- e N =1 -e-1.
Stehli, P.
1989 Merzbachtal-Umwelt und Geschichte Einer Band-
keramischen Siedlungskammer. Germania 67:51-76.Thus Ps is approximately constant. Note that u < 1, so ps gets
1994 Chronologie der Bandkeramik im Mcrzbachtal.smaller
In as p gets larger. The probability P(t) of being copied
Die Bandkeramik im Merzbachtal auf der Aldenhovenerfor t consecutive time steps is:
Platte, edited by J. Liining and P. Stehli, pp. 79-192.
Habelt, Bonn. P(t) = p.
Turcotte, D. L.
6. Neiman (1995:14) arrives at this formula through the
1997 Fractals and Chaos in Geology and Geophysics. Cam-
approximation that N is very large and p is small. If we
bridge University Press, Cambridge.
assume p is not quite so small, it would follow from Neiman
1999 Self-organized Criticality. Reports on Progress in
Physics 62:1377-1429. (1995: equation[2]) that ne= (2Np + 1)/(1 - 21). From this we
United States Census Bureau predict and average of 3.2 variants for p = 0.05 and 6 variants
for p = 0.10, which are in good agreement with Figures 3b
1995 "Documentation and Methodology for Frequently
and 3c.
Occurring Names in the U.S. - 1990." <http://www.cen-
sus.gov/ftp/pub/genealogy/www/freqnames.html> 7. The frequencies of names among 6.3 million persons,
Wilkinson, J. R. about one fortieth of the U.S. population, were collected in
1997 Ceramics as Viral Quasi-Species. Unpublished M.A. the 1990 Census from sample areas around the country. These
dissertation, Institute of Archaeology, University College
data, with 3519 different surnames, 4275 female and 1219
London.
Witten, T. A., and L. M. Sander male first names, are on a publicly accessible file (U.S.
1981 Diffusion LimitedAggregation, a Kinetic Critical Phe-
Census Bureau 1995).
nomenon. Physical Review Letters 47:1400-1403. 8. Kimura and Crow (1964) predicted the equilibrium
neutral allele (variant) frequency distribution ?(x) in a large
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Bentley and Shennan] CULTURAL TRANSMISSION AND NETWORK GROWTH 485
population as: that the position of the peak occurs at ln(ns) = got, and the wid
of the peak (2t is set on a logarithmic scale.
(x) = x-l( - x) - 1
12. To produce a histogram to be displayed with a loga-
where x is the frequency of a given allele and 0rithmic
is a measure
x-axis, the data must be converted to log form befor
of the variability in a population equal to 4NA, (as in the
they are text,
binned. The bin values can then be converted back
N is the effective population size and u is the neutral mutation
regular form to display on the plots. To produce a compar
rate). ble distribution from the model, we bin the model distributio
9. Some readers may want a more explicit definition of in the same way. The modeled probability of being betwee
our network metaphor, having perhaps asked, "does the vari- xI and x2 (x1 < x2) is equal to the area under the probabili
ant copy also have a corresponding link?" For our particular curve between xi and x2. This is the same the cumulative d
model, the answer is no, not an incoming link. By our defini- tribution at x, subtracted from its value at x2. The cumulativ
tion, a copy has only an outgoing link that points to the vari- log-normal function (Turcotte 1997:36) is:
ant, which for metaphorical purposes could be considered the 1 Inx- y
"blueprint" of the variant. An additional copy adds another F(x) = -erfc
2 2o
incoming link to the variant, but not to existing copies. So
only the variant nodes are included when we tally how many where erfc is the complimentary er
incoming links the different nodes have, since only the vari- to equation (16), we have
ant nodes, and not their copies, have any incoming links. This 1 (ln x-0.0466n~
definition, which we hold to consistently in our analysis, is P(n,2 > x)= erfc -( 46
0.2534t
practically necessary. Among archaeological remains, we
could never hope to identify links to individual variant copies for the probability that the num
(like we can with journal references, for example). If we x. The probability that ns is withi
could, it would add enormously to our sample (later copies ference between the cumulative
potentially having many outgoing links, just like journal arti-
cles obligated to have large bibliographies), but our model P(x P(x <n erfc
< n, < X<2)= = 1- 0er (ln x2
2 0.2534t
could easily accommodate this change in the nature of the
data (see Bentley and Maschner 2000).
Another option for our study in particular (discussed later 1 erfc _(lnxl - 0.0466nht)
--erfc -
in this paper) is to model LBK houses as the nodes, and 2 0.2534t
copies of the motifs that may represent household identities
as the network links. While this function would b
10. The "Web of Science" (WOS) database (www.webof- it is simple to do on any spr
science.com) lists journal articles published between 1987 error function, such as Excel
and the end of 1999 with certain keywords. Articles before 13. To combine phases w
1987 were found using the Social Science Citations Index. from each phase given by e
Both databases are maintained by the Institute of Scientific distribution of motif frequ
Information. have:
11. Note the similarity between equation (8) and the nor-
_1 _ _ (n ns -anht)2
P(n= S) = exp -
mal (bell-curve) distribution,
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