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the passion for cacti and other succulents

ISSN 2285 – 3987

21June
2017
the passion for cacti and other succulents

summary 21
3 · Editorial 21 | Eduart
4 · Xerophilia 21's Favorite Quote | Xerophilia
5 · The Thelocactus rinconensis complex | Rodrigo H. González G.
55 · Navajoa peeblesiana in habitat | Stefan Nitzschke & Carmen Bechara
65 · Mammillaria bombycina Quehl | Helmut Rogozinski
75 · Notes on Mammillaria aff. candida SB 827 | Elton Roberts
89 · Leuchtenbergia principis Hook. | Juan Miguel Artigas-Azas
99 · Tephrocactus geometricus (A.Cast.) Backeb - part 2. | Robert Bader
107 · Mexican wilderness trough the eyes of an artist. | Leo Rodriguez
123 · Ant-plants of the Solomon Islands Archipelago | Derrick Rowe
131 · Online journals | Xerophilia
132 · Online journals - Huitzilopochtli - Link to the Cactus Explorer website| Xerophilia
133 · Online journals - The Chileans - Link to the Cactus Explorer website | Xerophilia
134 · Abstract în limba română | Xerophilia
136 · Cactus Tour Mexico | Xerophilia
137 · Sedum e Petrosedum | Xerophilia
138 · Interesting offer of cacti seeds from South America | Xerophilia

Founders: Eduart Zimer • Dag Panco • Valentin Posea Nordic representative • Erik Holm
Supporter • Mihai Crisbășanu
Editor: Eduart Zimer Editorial team's e-Mail: xerophilia@xerophilia.ro.
Graphic layout based on Andrea Cattabriga's pattern.
EN edition • Eduart Zimer All rights reserved – no part of this publication may be reproduced in any forms
SP edition & Field researches • Pedro Nájera Quezada or by any means, without written permission of the Editor. All copyrighted
Photo edition • Valentin Posea photographs have been used under the Creative Commons Attribution 4.0
PR & Graphic • Dag Panco International license.

Front cover Back cover

On the edge of the abyss Thelocactus rinconensis ssp.


freudenbergeri,
North Grutas de Garcia, NL.

Photo by Photo by
Leo Rodríguez Manuel “Melo” Salazar

Summary 2 - XEROPHILIA • Volume VI, No. 2 (21), June 2017 | ISSN 2285-3987
summary

the passion for cacti and other succulents

21- june 2017

F
ive years ago, at the end of June 2012, the first

editorial 21
issue of this journal was published online.
A journal is born thanks to its founders! It
was, maybe, the insane act of three passionate
cactophiles who, without knowing whether their
initiative would be supported or not, without
knowing whether they will have the desired continuity or
whether their work will disappear meteorically, without
knowing if they will succeed in overcoming the lack of ex-
perience and reputation or if they will be largely ignored,
without having too much publishing experience these
three enthusiasts decided to share with likeminded peo-
ple free information, every bit of valuable information
they could offer, at no cost.
The coverage of the appearances is good: 21 current
issues and 10 special issues, while two more are being
in the processing stage. Asked whether we, as publish-
ers, have succeeded or not, we are witnessed both by the
number of downloads recorded on each of the issues
and by the number and diversity of the regions of the
world where Xerophilia has been read. now 17! And the Philippines went up from one to 17 as
A journal survives thanks to its readers! Therefore, to well; Malaysia from two to 16. Cuba climbed from one to
our readers, be they many, or maybe only one in their own 15. Venezuela reached 8. Vietnam also climbed from two
country, we feel compelled to thank them equally! From to ... 24! From the first issue until today, the two readers
the well over 100 countries and territories where we have in Namibia and Sri Lanka remain faithful and persistent!
fans, we get an average of more than 2,500 downloads on Unfortunately,we lost the one in Angola...
every issue, with the last six months being topped, in or- As you can see, we have emphasized the small num-
der, by USA, Germany, Mexico, UK and ... Russia - just to list bers of readers in some countries, readers for whom our
the top 5 countries in the ranking, but ... appearance, every three months, is perhaps the only rele-
…but thinking about how hard was for us to gather in- vant information resource in their area. We want to voice
formation before we were connected to the free world, clearly to all our readers, and in particular those listed
without forgetting the others, our great joy is to be able above, that as long as we can publish Xerophilia, we will
to supply this journal to singular readers from Kosovo, not spare our efforts so that they can have a decent read.
Pakistan, New Caledonia, Kyrgyzstan, Uzbekistan, Iraq, A journal exists only thanks to its collaborators! And
Guadeloupe, Barbados, Cape Verde, Armenia, Reunion, if we acknowledge the readers, then, at least but not the
Qatar, Congo, etc… How wonderful is to have a reader last ones, we have to show our gratitude to all the au-
from Congo, an equatorial country! thors, who over the last five years have provided the pa-
In similar conditions, in places it becomes difficult to pers, articles, studies and photographs needed to build
find information you need, we have been adding new this journal. Without them, Xerophilia could not have
summary

readers from year to year, from countries where the pur- existed because they were and are the true artisans of
chase of printed specialized magazines is almost impos- its existence!
sible. Out of respect for these readers, we will make an
enumeration, which is not intended to be a statistic, but
merely illustrates a gratifying phenomenon. As a result,
in 2013, we had two readers in Iran, last year we had 10, Eduart

Editorial
Editorial 33 -- XEROPHILIA
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VI, No.
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(21), June
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2285-3987
summary

Jane Morris Goodall is a British pri-


matologist, ethologist, anthropologist,
and UN Messenger of Peace. Consi-
dered to be the world’s foremost expert
on chimpanzees, Goodall is best known
for her 55-year study of social and fa-
mily interactions of wild chimpanzees in
Gombe Stream National Park, Tanzania.
She is the founder of the Jane Goodall
Institute and the Roots & Shoots pro-
gram, and she has worked extensively
on conservation and animal welfare is-
sues. She has served on the board of
the Nonhuman Rights Project since its
founding in 1996.

Xerophilia 21's
Favorite Quote

Conservation has to become


an absolute priority,
because if we lose nature
that’s the end of us.
We can’t live without it.
summary

Jane Morris Goodall

Editorial
Quote 44 -- XEROPHILIA
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The Thelocactus
summary

rinconensis (Poselg.) Britton & Rose

complex
Rodrigo H. González G. - e-Mail: curadores2012@hotmail.com
Museo del Desierto, Departamento de Investigación, Carlos Abedrop Dávila #3745, Nuevo Centro Metropolitano de Saltil-
lo, C.P. 25022, Saltillo, Coahuila, México

With 42 photos by the author.


With 37 habitat photos offered by Grzegorz Matuszewski, some from the illustration of the book “Thelocac-
tus, systematyka, występowanie i uprawa“ by Grzegorz Matuszewski & Stanislaw Hinz, out of wich:
27 photos of plants in habitat by Grzegorz Matuszewski - web: www.kaktusymeksyku.pl
and 10 photos of plants in habitat by Pavel Pavlíček - web: www.cact.cz
With 7 photos of plants in habitat by Manuel “Melo” Salazar-González
All the photos are labelled under the specific name given to them by their authors.

A
t 40 km west of the city
of Monterrey, close to the
highway to Saltillo, there
is a small town named
Rinconada, Nuevo León.
This is a relatively new
Hacienda de Rinconada - arches of the main town as it is known today
gate. Moorish architecture influence. that the first settlements
were established here in 1583. It is a small
town having today less than 500 inhabi-
tants.
Two relevant events have taken place in
Rinconada N.L. The first one, in 1922, is
Photos on this page by Rodrigo H. González G.

the birth of the famous fighter Blue De-


mond, as he is known world-wide.
The other important event that put
Rinconada on the maps was that in 1855,
when close to the city limits, the German
summary

botanist Poselger found and described a


new species included in the genus Echino-
cactus: Echinocactus rinconensis, Allg. Gar-
tenz 23,18 1855, and named it to honor the
Village of Rinconada, where it was found.

The Thelocactus rinconensis complex 55 -- XEROPHILIA


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Background The question is what about the
Taking into account criteria of taxonomic clas- intermediate populations between
summary

sification, rather than looking for differences be- these two considered subspecies?
tween individuals of the same species, we are The author of this article can almost assure that
seeking for similarities. these intermediate populations are either un-
By studying the wild populations, we will be able known or are not taken completely into account in
to realize the possible distribution patterns of the order to determine a taxonomic place or category,
species over time, and the succession of changes even less if, as discoverer, one has the illusion or
in the morphology of individuals along the differ- motivation to recognise a new taxon that would
ences in altitude, temperature, soil type, pollina- carry the name of its discoverer.
tors and, of course, understand the mechanisms Many times we seek to find the greatest possi-
of seed distribution. ble differences among individuals in a population
This work, through the study of wild populations with mere commercial purposes.
known for the T. rinconensis complex, aims to give The author of this article is convinced of the
arguments to recognize idea that if breeding
or ignore what many lines between individu-
authors have called sub- als of relatively close
species or varieties for populations, whether for
members of the complex important geographical
Thelocactus rinconensis. distances, orographic or
In principle, the author mechanical barriers, are
of this article, starts from not interrupted, should
the appreciative idea not be considered as
that if there is no repro- subspecies, even when
ductive isolation, several apparently their mor-
of the subspecies should pho-physiological cha-
not be considered as racteristics make them
being subspecies, but 1 look very different from
rather they are varieties each other.
or forms of the species. Examples: Thelocac-
Only few of the taxonomic works are based on tus rinconensis ssp. rinconensis and Thelocactus
true field investigations. Much of it is comparative rinconensis ssp. freudenbergeri. Although their
bibliographical work, revisions of herbarium speci- populations grow relatively close, even less than
mens, illustrations elaborated during the scientific 2 km apart, they are plants with many morpho-
expeditions that have been happening since the logical differences. With a high probability it can
eighteenth century or, be ensured that they do
more recently, based on 2 not reproduce with each
the photographic com- other due to the exis-
parison of specimens tence of an important
collected and repro- orographic barrier sepa-
duced in private collec- rating the two popula-
tions in Europe for more tions (Manuel Salazar G.
than 100 years. in Xerophilia-Anul I, nr.
If we visit the wild 3 (3) - Decembrie 2012
populations near the ISSN-3987). It would be
type locality, Rinconada very interesting to have
NL (600 metres above knowledge of reproduc-
sea level) and then we tive testimony between
go to the known popu- both subspecies in labo-
lations of T. rinconen- ratory conditions.
sis fma. nidulans (1450 Astrophytum ornatum
Photos on this page by Rodrigo H. González G.

masl) west of Saltillo, we will find that there is a and Astrophytum coahuilensis are species being
vast difference between the individuals of both morphologically very similar to each other, how-
populations. The same happens if from the type ever, neither in nature nor in laboratory condi-
locality we travel 45 km south of Saltillo near Ar- tions can be reproduced, are two completely dif-
summary

teaga, Coahuila, (almost at 2,000 masl) to visit the ferent species currently isolated by an important
populations of T. rinconensis fma. phymathothelos geographic distance.
(photo 1), where the plants are less than 10 cm It is the same as in T. rinconensis ssp. hintonii
in diameter and growing flush with the earth, that grows isolated, more than 100 km away from
very much different from the known forms of T. the populations of T. rinconensis, also growing in
rinconensis ssp. rinconensis (photo 2). the heart of the Sierra Madre Oriental mountain

The Thelocactus rinconensis complex 6 - XEROPHILIA • Volume VI, No. 2 (21), June 2017 | ISSN 2285-3987
summary

Morpho-physiological changes in Thelocactus


rinconensis form “paila” in response to envi-
ronmental conditions or soil conditions.

does not cease to be a form or variety of the ssp.


rinconensis. In both cases, there are no important
geographical barriers that prevent reproduction
among these populations, therefore their mor-
pho-physiological characteristics are simply modi-
range, or also with T. rinconensis ssp. multicepha- fications in response to an environmental signal.
lus that grows more than 200 km away from the In contrast Thelocactus rinconensis ssp. hintonii,
populations of T. rinconensis ssp. rinconensis. Both located within the Sierra Madre Oriental moun-
species should be considered as subspecies of the tain range and Thelocactus rinconensis ssp. multi-
complex where the probability of their reproduc- cephalus located more than 200 km away from the
tion with any segment of the complex is almost null. known populations of T. rinconensis ssp. rinconenis,
By studying and knowing the intermediate they should be considered as subspecies, since the
populations* between Thelocactus rinconensis ssp. probability of reproducing with any of the known
rinconensis with the populations of the forms nid- subspecies or forms so far is very little or simply
ulans or “paila”, one can understand and verify a inexistent. In addition, for T. rinconensis ssp. mul-
gradual morphological transformation of a typi- ticephalus there are other important reasons for
Photos on this page by Rodrigo H. González G.

cal form of T. rinconensis ssp. rinconensis to a typi- separating this subspecies from the complex,
cal form of T. rinconensis fma. nidulans , following such as the works published in Bradleya 18: 45-
logical distribution patterns over relatively short 70 January 2000 “A phenetic analysis of the genus
distances, that is to say, there are very probably Thelocactus”, where it is verified that T. rinconensis
summary

mechanisms of reproduction between them. The ssp. multicephalus has a single hypodermic layer
same happens with the populations of T. rinco- and all others in the complex have a multilayered
nensis ssp rinconensis with respect to the so-called hypodermis.
ssp. phymatothelos where altitude and therefore
the climate, undoubtedly plays an important role * Intermediary populations: the studied populations be-
in its morpho-physiological differentiation but it tween the type localities of both subspecies.

The Thelocactus rinconensis complex 7 - XEROPHILIA • Volume VI, No. 2 (21), June 2017 | ISSN 2285-3987
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Photo by Rodrigo H. González G.


summary

Thelocactus rinconensis ssp. rinconensis fma “paila”,


plants with radial spines.

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
complex 88 -- XEROPHILIA
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summary

Thelocactus rinconensis ssp. rinconensis.

Subspecies or forms considered by the (2000) => see page 42.


author for the Thelocactus rinconensis • Thelocactus phymathotelos Poselger ex Ruem-
complex (bold) pler, in Förster, Handb. Cact., ed. 2, 602 (1885).
• Thelocactus rinconensis (Poselger) Britton & - as Thelocactus rinconensis ssp. rinconensis
Rose, Cact. 4: 7 (1923) ssp. rinconensis. Neo- fma. phymatothelos.
type: About 20 km North-east of Saltillo, 22 July • Thelocactus phymatothelos Br. & R. (‘phymato-
1972, E. F.Anderson 3180 (US). thele’), Cact. 4: 8 (1923). (1923).
• Echinocactus lophothele Salm-Dyck, Allg. Gar- • Thelocactus rinconensis var. phymatothelos Glass
tenz. 18: 395 (1850). & Foster, Cact. Succ. J. (US) 49: 246 (1977).
• Echinocactus rinconensis Poselger, Allg.Gartenz. • Thelocactus rinconensis ssp. phymatothele (Po-
23: 18 (1855). selger) Glass, Ident. Guide Threatened Cacti of
• Thelocactus lophothele Br. & R., Bull. Torrey Bot. Mexico 1: TH/RIN (1997).
Club 49: 251 (1922). • Thelocactus rinconensis ssp. phymatothelos (Po-
• Thelocactus lophothele ssp. rinconensis Matusz., selger) Doweld nom. superfl., Sukkulenty 1: 30
Thelocactus 170 (2011). (1999).
• Thelocactus rinconensis ssp. nidulans (Quehl) • Thelocactus rinconensis ssp. freudenbergeri
Glass & Foster, cact..succ.J. (US) 49(6):245 (1977) (Haas) Mosco & Zanovello,CCI 7: 18 (1999).
- as ssp. rinconensis fma. nidulans. • Thelocactus rinconensis var. freudenbergeri Haas,
• Thelocactus lophothele ssp. nidulans Matusz., Kakt. and. Sukk. 43: 96-98 (1992).
Thelocactus 180 (2011). • Thelocactus lophothele ssp. freudenbergeri Ma-
• Thelocactus rinconensis ssp. rinconensis fma. tusz., Thelocactus 184 (2011).
“paila”. • Thelocactus rinconensis ssp. hintonii Luethy,
• Thelocactus rinconensis ssp. palomaensis Kakt. and. Sukk. 48: 39(1997).
Photo by Rodrigo H. González G.

Pavlíček & Zatloukal - Kaktusy (Brno) 40(1): 11 • Thelocactus lophothele ssp. hintonii Matusz.,
summary

(5-13; figs.). 2004. - as ssp. rinconensis fma. pa- Thelocactus 177 (2011).
lomaensis. • Thelocactus rinconensis ssp. multicephalus
• Thelocactus lophothele ssp. palomaensis Matusz., (Halda & Panarotto) Luethy, Kakt. and Sukk. 50:
Thelocactus 188 (2011). 80 (1999).
• Thelocactus rinconensis ssp. icamolensis Halda • Thelocactus multicephalus Halda & Panarotto,
& Kupcak, Acta Mus. Richnov. Sect. natur. 7: 75 Acta Mus. Richnov. Sect. natur. 5: 40 (1998).

The Thelocactus rinconensis complex 9 - XEROPHILIA • Volume VI, No. 2 (21), June 2017 | ISSN 2285-3987
Comparative table with some morpho-physiological characteristics
summary

of each subspecies or form of the Thelocactus rinconensis complex

Central Radial
Name Description Flower Alt.
spine spine
Stem semiglobous-
flattened (5-15 cm 3-4 sometimes 5, the 800-
Thelocactus rinconensis Yellow-white,
height - 8-30 cm lower descending, usually no 1200
ssp. rinconensis slightly pink
diam.) olive green the longest, 4-8 cm long m
color
Stem semiglobous-
Thelocactus rinconensis
flattened (5-10 cm 3-4, straight to slightly White with a 1300-
ssp. rinconensis
height - 5-15 cm curved, 2-5 cm long no pink discer- 1400
fma. phymatothelos
diam.) glaucous gray nible midline m
Ojo Caliente
color
Stem depressed (4
Thelocactus rinconensis 2-3, 1-2 cm long,
cm height - 10cm 1600-
ssp. rinconensis curbed, generally con- White, slightly
diam.) glaucous gray no 2000
fma. phymatothelos centrated at the apex pink
color, purple tuber m
Arteaga and eventually lost
tips
Stem semiglobous- 3-5, 4-8 cm straight No,
Thelocactus rinconensis flattened (8-35cm black to light brown, occa-
750-
ssp. rinconensis height - 8-40 cm slightly curved, lower siona- White or yelow
800 m
El Barreal diam.) olive green descen-ding spine, lly 2-4
color usually the longest
3-4 sometimes 5, 4-5
cm long, light gray-
brown color, the des-
Stem semiglobous- cending lower usually
flattened (5-20 cm the longest, with
Thelocactus rinconensis Occa- 1400-
height - 5-30 cm the age their spines
ssp. rinconensis fma. siona- White 1500
diam.) olive green are disintegrated in
nidulans lly 2-4 m
color irregular filaments and
they are not lost giving
a bird’s nest (nido)
appearance, hence the
“nidulans” name
Stem semiglobous- 3-4 sometimes 5, 4-8
Thelocactus rinconensis flattened (5-20 cm cm long, curved, brown
3-6 1150-
ssp. rinconensis height - 5-30 cm chestnut color, few Pink to
very 1300
fma. palomaensis & diam.) gray olive times the descendant deep-magenta
short m
fma. “paila” green color lower generally the
longest
Stem semiglobous
3-4 sometimes 5, 4-5 No,
to globose flattened
cm long, curved, dark occa- 1550-
Thelocactus rinconensis and usually caespi-
brown, the descendant siona- White-yellow 1700
ssp. multicephalus tose (5-15 cm height
lower generally the lly 2-4 m
- 8-20cm diam.) gray
longest
olive green color
6-8, very thin, 2-7 cm
Stem globose slightly length it could be said
6-7
flattened (8-15 cm to have only one well
Thelocactus rinconensis strait, 850-
height - 10-20cm defined central spine, Magenta
ssp. freudenbergeri 1-6 cm 950 m
diam.) green color the others appear ra-
long
dial, the lower slightly
summary

thicker and decendant


stem semiglobous to
3-4 sometimes 5, 4-8 No,
globose, flattened 1100-
Thelocactus rinconensis cm long, curved, dark occa-
(5-20 cm height) Yelow-white 1300
ssp. hintonii chestnut color, the lo- siona-
(8-25 cm lenght) m
wer usually the longest lly 2-4
green color

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
complex 10
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1/15 Thelocactus rinconensis tural and taxonomic name Thelocactus rincon-
summary

ensis (Poselger) Britton & Rose was replaced


(Poselger) Britton & Rose, Cact. 4: 7 (1923)
by the nomenclatural name Thelocactus lopho-
ssp. rinconensis thele Matusz., arguing the basionym Echino-
La Rinconada cactus lophothele Salm-Dyck, Allg. Gartenz. 18:
395 (1850) had been published five years ear-
lier than Echinocactus rinconensis Poselger, Allg.
Grzegorz Matuszewski - who kindly offer us Gartenz. 23: 18 (1855). Starting from here, in G.
all these pictures labelled under his name as au- Matuszewski’s nomenclature all names are dif-
thor - has a founded opinion in the genus The- ferent.
locactus classification, opinion he has presented This article has not the meaning of discuss-
in his wonderful book “Thelocactus. Systematika, ing this point of view. We just point out the dif-
wystepowanie i uprawa”. With this occasion, the T. ferences between both visions trying to avoid
rinconensis complex has been combined again. confusion and wanting to honor the author of
As a consequence, first of all, the nomencla- the photos we are using here.

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - La Rinconada.

Photo by Grzegorz Matuszewski


summary

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
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G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - La Rinconada.
2/15 Thelocactus rinconensis
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis
G. Matuszewski: Thelocactus lophothele ssp. lopho- La Rinconada
thele - La Rinconada.

Photos on this page by Grzegorz Matuszewski


summary

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
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3/15 Thelocactus rinconensis
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(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis
Higueras

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - Icamole.

Photo by Grzegorz Matuszewski


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 13
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G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - El Delgado. 4/6 Thelocactus rinconensis
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis
G. Matuszewski: Thelocactus lophothele ssp. lopho-
thele - El Delgado.
El Delgado

Photos on this page by Grzegorz Matuszewski


summary

en or The
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5/15 Thelocactus rinconensis
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(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis
San Antonio de Arista

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - San Antonio De Arista.

Photo by Grzegorz Matuszewski


summary

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
complex 15
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summary

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - San Juan de Arista.
6/15 Thelocactus rinconensis
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis
G. Matuszewski: Thelocactus lophothele ssp. lopho-
thele - San Juan de Arista. San Antonio de Arista

Photos on this page by Grzegorz Matuszewski


summary

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
complex 16
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tion... but it occurs only in cultivation, indicating
7/15 Thelocactus rinconensis this characteristic could be in fact the result of
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923) a horticultural selection. So far, field researches
in Icamole show the same already cited normal
ssp. rinconensis color variation in the flowers of the complex
Icamole Rinconensis, but no dark purple at all.
From six years, the author, organize the ICA
Desert MTB in Icamole, N.L. and vicinities - a well
known mountain bike race. Working on the tracks
Thelocactus rinconensis ssp. icamolensis Halda for the race, and prospecting the hills around the
& Kupcak has been described in 2000*. Since town, the author has tried to identified the so called
then until today, no one could find the dark “icamolensis” at home. In vain! He has find only the
purple (atropurpurea) flowering subspecies in ordinary Thelocactus rinconensis ssp. rinconensis.
its habitat. As mentioned, the variation of the The photos on these four pages are from all
flower colors in the complex Rinconensis is not a around Icamole, where Grzegorz Matuszewski
constant. it can be yellow for the spp. rinconen- has also tried without results to find this subspe-
sis; or pink for the fma. phymatothelos; or white cies in its presumed habitat.
for the fma. nidulans; or magenta for the ssp. As you can see, there are a lot of variations in
freudenbergeri. The magenta color for “icamolen- the shape of the plants and in the color of the
sis” could be a special color in this location, even spines and the flowers... but no magenta flower.
if it is not so far from ssp. freudenbergeri loca- The conclusion: there is not even a form at all.

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - Icamole. Light yellow flowers

Photos on this page by Grzegorz Matuszewski


summary

* “Taproot thin, ca 100 mm long; stem single, much smaller than ssp. rinconensis (up to 200 mm
across and 60mm tall); flowers small, up to 30mm long, with dark purple obovate leaflets.“ - Acta Mus.
Richnov., Sect. Nat. 7(2): 75 (2000).

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 17
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summary

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - Icamole.
Light yellow flowers

8/15 Thelocactus rinconensis


Thelocactus rinconensis ssp. icamolensis (Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis
Icamole
Light pink, pink striped flowers
G. Matuszewski: Thelocactus lophothele ssp. lopho-
thele - Icamole.

Photos on this page by Grzegorz Matuszewski


summary

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
complex 18
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9/15 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis
Icamole

Thelocactus rinconensis ssp. icamolensis

white, light pink striped flowers

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - Icamole.

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - Icamole.

Photos on this page by Grzegorz Matuszewski


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 19
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10/15 Thelocactus rinconensis Thelocactus rinconensis ssp. icamolensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis
Icamole Light yellow flowers

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - Icamole.

Photo by Grzegorz Matuszewski


summary

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
complex 20
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summary

Photo by Rodrigo H. González G.


summary

The populations find in El 11/15 Thelocactus rinconensis


Barreal and El Higo are typical-
ly intermediary forms between (Poselger) Britton & Rose, Cact. 4: 7 (1923)
T. rinconensis ssp rinconensis ssp. rinconensis
and T. rinconensis ssp. rinco-
nensis fma. nidulans. El Barreal

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 21
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summary

Photo by Rodrigo H. González G.


summary

12/15 Thelocactus rinconensis


(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis
El Barreal

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 22
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summary

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - Est Higo.

The populations find in


El Higo and El Barreal are
13/15 Thelocactus rinconensis
typically intermediary forms (Poselger) Britton & Rose, Cact. 4: 7 (1923)
between T. rinconensis ssp.
rinconensis and T. rinconensis
ssp. rinconensis
ssp. rinconensis fma. nidulans. Est El Higo

G. Matuszewski: Thelocactus lophothele ssp. lopho-


thele - Est Higo.

Photos on this page by Grzegorz Matuszewski


summary

en or The
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rinconensis Complex
complex 23
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14/15 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis
Est El Higo

Photo by Grzegorz Matuszewski


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 24
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summary

G. Matuszewski: Thelocactus lophothele ssp. lopho- 15/15 Thelocactus rinconensis


thele - Est Higo.
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis
G. Matuszewski: Thelocactus lophothele ssp. lopho-
thele - Est Higo.
est El Higo

Photos on this page by Grzegorz Matuszewski


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 25
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1/6 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis fma. nidulans
El Chiflon

Photos on this page by Rodrigo H. González G.


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 26
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summary

2/6 Thelocactus rinconensis


(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis fma. nidulans
Puebla

Photos on this page by Rodrigo H. González G.


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 27
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summary

3/6 Thelocactus rinconensis


(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis fma. nidulans
El Chiflon

Photos on this page by Rodrigo H. González G.


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
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summary

4/6 Thelocactus rinconensis Thelocactus rinconensis ssp. rinconensis fma. nidulans


- La Chancaca.
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis fma. nidulans
Thelocactus rinconensis ssp. rinconensis fma. nidulans
La Chancaca & El Chiflon - El Chiflon.

Photos on this page by Rodrigo H. González G.


summary

en or The
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rinconensis Complex
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summary

Matuszewski: Thelocactus lophothele ssp. lophothele


- El Chiflon,.

For the author, the plants of the T. rinconesis 5/6 Thelocactus rinconensis
complex living in El Chiflon, Puebla and La Chan-
caca, even if plants have but only central spines, (Poselger) Britton & Rose, Cact. 4: 7 (1923)
are in fact Thelocactus rinconensis ssp. rinconensis ssp. rinconensis fma. nidulans
fma. nidulans, as its name is referring to that the
oldest thorns forming together what looks like El Chiflon
a bird nest (Latin = nidum).
For Grzegorz Matuszewski, author of these two
and of the following photos, from El Chiflon, this lophothele, in other words it is a comon T. rinco-
population is just a population of T. lophothele ssp. nensis ssp. rinconensis not a fma. “nidulans”.

Matuszewski: Thelocactus lophothele ssp. lophothele


- El Chiflon,.

Photos on this page by Grzegorz Matuszewski


summary

en or The
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rinconensis Complex
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6/6 Thelocactus rinconensis Matuszewski: Thelocactus lophothele ssp. lophothele


- El Chiflon,.
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis fma. nidulans
Matuszewski: Thelocactus lophothele ssp. lophothele
El Chiflon - El Chiflon,.

Photos on this page by Grzegorz Matuszewski


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 31
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summary

Photo by Rodrigo H. González G.


summary

1/1 Thelocactus rinconensis


(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis fma. “paila”
Sierra Paila

en or The
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rinconensis Complex
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1/4 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis fma. palomaensis
La Paloma

Matuszewski: Thelocactus lophothele ssp. palomaen-


sis - La Paloma.

Photos on this page Pavel Pavlíček


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
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summary

Matuszewski: Thelocactus lophothele ssp. palomaen-


2/4 Thelocactus rinconensis
sis - La Luz.
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis fma. palomaensis
Matuszewski: Thelocactus lophothele ssp. palomaen-
sis - La Paloma.
La Luz & La Paloma

Photos on this page Pavel Pavlíček


summary

en or The
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rinconensis Complex
complex 34
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summary

3/4 Thelocactus rinconensis Matuszewski: Thelocactus lophothele ssp. palomaen-


sis - La Luz.
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis fma. palomaensis
Matuszewski: Thelocactus lophothele ssp. palomaen-
La Luz & La Paloma sis - La Paloma.

Photos on this page Pavel Pavlíček


summary

en or The
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rinconensis Complex
complex 35
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summary

Matuszewski: Thelocactus lophothele ssp. palomaen-


4/4 Thelocactus rinconensis
sis - La Luz.
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. rinconensis fma. palomaensis
Matuszewski: Thelocactus lophothele ssp. palomaen-
sis - La Paloma.
La Luz & La Paloma

Photos on this page Pavel Pavlíček


summary

en or The
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rinconensis Complex
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3/7 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis fma. phymatothelos
Ojo Caliente

Photo by Rodrigo H. González G.


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 37
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summary

Photo by Manuel “Melo” Salazar González.


summary

Manuel “Melo” González: Thelocactus rinco-


3/7 Thelocactus rinconensis
nensis ssp. rinconensis fma. phymatothelos - (Poselger) Britton & Rose, Cact. 4: 7 (1923)
Ojo Caliente.
ssp. rinconensis fma. phymatothelos
Ojo Caliente

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 38
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summary

Photo by Manuel “Melo” Salazar González.


summary

3/7 Thelocactus rinconensis Manuel “Melo” González: Thelocactus rinco-


(Poselger) Britton & Rose, Cact. 4: 7 (1923) nensis ssp. rinconensis fma. phymatothelos -
ssp. rinconensis fma. phymatothelos Ojo Caliente.

Ojo Caliente

en or The
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rinconensis Complex
complex 39
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This photo and the one in the next page 4/7 Thelocactus rinconensis
summary

show a very interesting form that Grzegorz Ma-


tuszewski has photographed near Higueras. (Poselger) Britton & Rose, Cact. 4: 7 (1923)
This form is very alike with the Thelocactus ssp. rinconensis
rinconensis ssp. rinconensis fma. phymatothelos
from Ojo Caliente, some four km to the south-
Higueras
west, aff. fma. phymatothelos
Both populations are considered by the au-
thor as intermediate populations between the
Ojo Caliente
typical form of the ssp. rinconensis and the typi-
cal form of the fma. phymatothelos from Artea-
ga. You can see the differences in the photos Almost the same shape, the same body and
displayed in the precedent pages and in the spine color; even the purple tuber tips, charac-
one in the bottom of this page. teristic to the cited form is present.

G. Matuszewski: Thelocactus lophothele ssp.


lophothele - Higueras.

le z G .
i
sz e w sk
H . G o n zá
summary

rz M a tu
y R o d ri go
o
y G rz e g
P h o to b
P h o to b

en or The
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rinconensis Complex
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5/7 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis
Higueras
aff. fma. phymatothelos
Ojo Caliente

G. Matuszewski: Thelocactus lophothele ssp.


lophothele - Higueras. .

w sk i
zá le z G

a tu sz e
H. Gon
summary

g o rz M
o d ri g o

b y G rz e
b y R
P h o to

P h o to

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rinconensis Complex
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6/7 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis fma. phymatothelos
Arteaga

Photos on this page by Rodrigo H. González G.


summary

en or The
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rinconensis Complex
complex 42
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7/7 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. rinconensis fma. phymatothelos
Arteaga

Photos on this page by Rodrigo H. González G.


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 43
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1/4 Thelocactus rinconensis


(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. feudenbergeri
(Haas) Mosco & Zanovello,CCI 7: 18 (1999).
North Grutas de Garcia

Photos on this page by Rodrigo H. González G.


summary

en or The
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rinconensis Complex
complex 44
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Photos on this page by Rodrigo H. González G.

2/4 Thelocactus rinconensis


summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. feudenbergeri
(Haas) Mosco & Zanovello,CCI 7: 18 (1999).
North Grutas de Garcia

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 45
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Photo by Manuel “Melo” Salazar González.

3/4 Thelocactus rinconensis


summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. feudenbergeri
(Haas) Mosco & Zanovello,CCI 7: 18 (1999).
North Grutas de Garcia

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 46
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4/4 Thelocactus rinconensis
summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. feudenbergeri
(Haas) Mosco & Zanovello,CCI 7: 18 (1999).
North Grutas de Garcia

Photos on this page by Manuel “Melo” Salazar González.


summary

en or The
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Thelocactus rinconensis
rinconensis Complex
complex 47
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summary

1/3 Thelocactus rinconensis Matuszewski: Thelocactus lophothele ssp. hintonii -


Rayones Valey, West Santa Rosa.
(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. hintonii
Luethy, Kakt. and. Sukk. 48: 39 (1997)
Rayones Valley, West Santa Rosa Matuszewski: Thelocactus lophothele ssp. hintonii -
Rayones Valey, West Santa Rosa.

Photos on this page Pavel Pavlíček


summary

en or The
The Thelocactus
Thelocactus rinconensis
rinconensis Complex
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Photo by Manuel “Melo” Salazar González.

2/3 Thelocactus rinconensis


summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. hintonii
Luethy, Kakt. and. Sukk. 48: 39 (1997)
Rayones Valley, West Santa Rosa

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Photo by Manuel “Melo” Salazar González.

3/3 Thelocactus rinconensis


summary

(Poselger) Britton & Rose, Cact. 4: 7 (1923)


ssp. hintonii
Luethy, Kakt. and. Sukk. 48: 39 (1997)
Rayones Valley, West Santa Rosa

en or The
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1/2 Thelocactus rinconensis


(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. multicephalus
Luethy, Kakt. and. Sukk. 48: 39 (1997)
Sandia

Photos on this page by Rodrigo H. González G.


summary

en or The
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2/2 Thelocactus rinconensis


(Poselger) Britton & Rose, Cact. 4: 7 (1923)
ssp. multicephalus
Luethy, Kakt. and. Sukk. 48: 39 (1997)
Sandia

Photos on this page by Rodrigo H. González G.


summary

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Thicker and descending central spine

Photos on this page by Rodrigo H. González G.


reviewed. From the resulting revisions we can
About 15 years ago, through the company that the mention the variation of the color of the flower
summary

author of the text directed, e.g. LANDSCAPE CU- in an area of 10 ha. The flower color varies from
RATORS S.A. of C.V., the rescue and relocation of yellow (predominant in the area) to white, white
native plants, among them several thousand cacti, with pink median line and light pink. This varia-
was carried out. This rescue was carried out to al- tion helps to explain and understand many of the
low starting the construction work of what is cur- confusions made by those who described these
rently a golf course near the type locality. During species
this project more than 3,000 plants of Thelocactus Another interesting observation is what particu-
rinconensis ssp. rinconensis, were rescued. larly defines the T. rinconensis complex: the lower
During this work, the collected specimens were central spine is generally descending and thicker

Rescued plants
summary

than the others. other, a typical characteristic of all the members


Analysing the rescued specimens, it was pos- of the T. rinconenis complex.
sible to verify the diameter ratio of this descend- Studied populations in the field, where the de-
ing spine, which as shown in the photograph, is a scending central spine is partially or totally divi-
fusion of 2 central spines, resulting in the appea- ded are T.rinconensis ssp. rinconensis, T. rinconensis
rance of the lower central spine wider than the fma. nidulans and T. rinconensis fma. “paila”.

The Thelocactus rinconensis complex 53 - XEROPHILIA • Volume VI, No. 2 (21), June 2017 | ISSN 2285-3987
Bibliography
summary

• Bravo-Hollis H & Sánchez-


Mejorada H. 1992. Las cactáceas
de México, Vol. 2, Segunda
Edición. Universidad Nacional
Autónoma de México, México,
D.F.
• Anderson, E. F. 1987: A revision
of the genus Thelocactus B. & R.
(Cactaceae). Bradleya 5: 49-76.
• Britton, N. L. & Rose, J. N. 1923:
The Cactaceae. Vol. 4. Carnegie
Inst. Washington, Washington, D.
C. Doweld, A. B. 1998:
• Bradleya 18:45-70 January 2000
A phenetic analysis of the genus
Thelocactus
• Thelocactus multicephalus Halda
& Panarotto, Acta Mus. Richnov.
sect. Nat. 5: 40 (1998). Type: Feb.
1985, J. J. Halda s. n. (PR).
• Thelocactus rinconensis ssp. mul-
ticephalus (Halda & Panarotto)
Luethy, Kakt. and Sukk. 50: 80
(1999).
• Thelocactus rinconensis (Posel-
ger) Britton &Rose, Cact. 4: 7
(1923). Neotype: About 20 km
• Salm-Dyck, Allg. Gartenz. 18: 395
(1850). Echinocactus lophothele
• Br. & R. (‘phymatothele’), Cact. 4:
8(1923). Thelocactus phymatothe-
los
• Glass & Foster, Cact. Succ. J.
(US) 49: 246 (1977). Thelocactus
rinconensis var. phymatothelos
• Threatened Cacti of Mexico 1: TH/
RIN (1997).
• Thelocactus rinconensis ssp. phy-
matothelos (Poselger) Doweld
nom. superfl., Sukkulenty 1: 30
(1999).
• Thelocactus rinconensis var.
Freudenbergeri Haas, Kakt. and.
Sukk. 43: 96–98 (1992).
• Thelocactus rinconensis ssp. hin-
tonii Luethy, Kakt. and. Sukk. 48:
39(1997).
• John Pilbeam 1996 UK Cactus file
handbook Thelocactus.

Xerophilia’s and author’s


acknowledgements
Photos on this page by Rodrigo H. González G.

• We want to thank Dr. Leccinum J. Garcia-Morales for


his suggestions and for his revision of the text.

Plants with fused spines.


summary

Xerophilia’s and author’s


acknowledgements

• We thank Grzegorz Matuszewski for his over the Author’s acknowledgement


years prompt and constant help with great photos.
• We thank Manuel “Melo” Salazar-González for his • I especially thank to my Professor, Mr. Suguri Kozui, for
past and present collaboration with our journal. the revision of the text and his support in my explorations.

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Navajoa
summary

peeblesiana Croizat

in habitat

Stefan Nitzschke

Carmen Bechara

M
y friend Carmen and I
considered spending this
year’s holiday, in the USA.
However, the planned
centre of attention was
purely touristic, with only
few pure cactus related
stops included in our
travel schedule. We already had Los Angeles
with its stars honouring Walk of Fame and the
Hollywood Sing, the casinos in Las Vegas and
a very recommendable Prince Tribute show,
the Grand Canyon and some of its breath-ta-
summary

king views lightly powdered with snow, as well


as the unfortunately unsuccessful search for
Navajoa maia at the location JB6 which me my
friend Josef Busek kindly suggested me during
the journey.

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Now, it was time for another clean cactus


stop to follow: Navajoa peeblesiana near Hol-
brook Arizona. From good friends I had been
given plenty of details, however, coincidentally
with the indication that in the place mentioned
during the last visits only very few plants were
actually found. Thanks to a good preparation,
summary

by means of Google Earth, the journey was


quickly completed, the ascent to the plateau
intended to be examined was also no witch-
craft, however, now began the particular I had
quite a little anxiety: the actual search.

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After only a few minutes of browsing on


the plateau I could hardly believe my eyes,
the first plant was found and after further
intense search I could find plants of all sizes
in many small micro-populations, consisting
of 4-8 plants. After the agreed time of two
hours of searching for cacti I went all the
summary

way back to Carmen, who had spent the time


at the car. Taking everything into account,
I have found in this rather short period of
time about 80 plants, some of them partly
covered with buds.

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Whether this number of plants seen is


enough to speak of a healthy population, I
leave the cactus experts of the United States
to decide, but I am glad that I cannot confirm
the black-painted statistics supplied by my
informants.
summary

Now, we continued our travel with a short


tourist stop at the Wigwam Motel from the
Disney film Cars, placed was on the way to
another cactus site, which I would like to re-
port in the next issue.

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To be able to present something relevant


related to the genus Navajoa, I consider that
I am still much too new in the subject. For
instance, I could not find in the field great
differences to the also visited Navajoa pee-
blesiana ssp. menzelii and likewise several
connoisseurs came close to the skies by
sending pictures for determination of both
types.
summary

With pleasure I will deal more closely with


the genus in the future and, besides the
Mexican genera, I will now also create a little
space in my collection to be able to observe
plants of the genus having a positive origin.

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Mammillaria
summary

bombycina Quehl

Rog. 179, Rog. 382, Rog. 383 and Rog. 385

Helmut Rogozinski
with photos by Helmut Vogel

M
ammillaria bombycina,
Rog 179, reddish spines,
Sierra El Laurel/Los Ali-
sos, Aguascalientes.
In 1990, we drove from
Jalpa, Aguascalientes,
across Jaltiche de Arriba
towards Alisos, Aguas-
calientes, to find the 80 years long known M.
bombycina, which was rediscovered only a
short time before by Fitz Maurice. From far
summary

away you can see the high-rise rock faces. You


get stopped to the right by steep crags. We Mammillaria bombycina Rog 179 Los Alisos,
see on the way, on a huge rock block, a huge Aguascalientes.
M. bombycina pad with more than a hundred
heads.

Mammillaria
Mammillaria bombycina
bombycina 65
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Mammillaria bombycina Rog 179 Los Alisos,


Aguascalientes.

M. bombycina is fine spined, usually with dark


brown central spines. Probably we were the
first Europeans to see these plants in nature.
Our joy was huge. Afterwards, we inspected
the site, M. bombycina grows almost always -
summary

except for the just described exception - in the


steep rock walls, and taking pictures is possible
only with great security. We continue towards
Ciénegas de los Pinos. Here is a single standing
mountain range.

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Mammillaria bombycina Rog 383, El Maguey 2044 m, Mammillaria bombycina Rog 383, El Maguey 2044 m,
Aguascalientes. Aguascalientes.

All these old photos on this page are from and 1994 with a Nikon F 100, Fuji Velvia Film, 50 ASA.

Mammillaria bombycina Rog 383, El Maguey 2044 m,


Aguascalientes.

Mammillaria bombycina Rog 383,


coarse spines, El Maguey 2044 m,
Aguascalientes
In March 1994, we drove on the MEX 70 to El
Sauz and from there to El Maguey. The location
is above the village. Here, in the rocky quarries,
we see the yellow-spined M. bombycina. It is an
incredible sight!
The plants are covered by extremely rough
summary

spines. Rarely only, reddish spined plants can


be also seen. Many plants bear carmine pink
colored flowers. The plant mats can become up
to one meter wide. We have so far rarely seen
something so overwhelming.

en or Mammillaria
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Mammillaria bombycina Rog 383, El Maguey 2044 m, Mammillaria bombycina Rog 383, El Maguey 2044 m,
Aguascalientes. Aguascalientes.

All these old photos on this page are from 1994 and made with a Nikon F 100, Fuji Velvia Film, 50 ASA.

Mammillaria higuerensis, El Marguey, 2044 m,


Aguascalientes.

Location: slopes leading to the village, on red-


dish brown boulders, accompanied by trees,
Yucca, Agave, Mammillaria higuerensis Rog 767.
In 2007, we visited the site for a second
time. We had to search for a long time. In the
meantime, the MEX 70 has been upgraded to
an expressway, and the route has been slightly
altered. The village of El Sauz apparently no
longer exists or the road to Maguey was moved
away from it. The signboard used to be locat-
ed about 100 m away from the main road and
could not be seen from the road now in 2007.
summary

At that time I wrote in my travel notes: “I re-


membered the location more beautiful. What
beautiful photos did I take back then. However,
I was also somewhat more mobile then.” In fact,
I did not dare to pass these steep walls in 2007.

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Landscape of Ciénega de los Pinos 2441m, Sierra El


Laurel, Aguascalientes.

Landscape of Ciénega de los Pinos 2441m, Sierra El


Laurel, Aguascalientes.

Mammillaria bombycina, Rog


385, Ciénega de los Pinos 2441m,
Sierra El Laurel, Aguascalientes
In 1994 and 2007 we visited this area again. M.
bombycina is still present, but the large M. bom-
bycina cushion we saw in 1990 has disappeared.
summary

In 2007, the ascent through the seemingly


denser undergrowth was incredibly tedious.
However, when we arrived up on the mountain
plateau we have been copiously compensated
by the view.

en or Mammillaria
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Mammillaria aff. bombycina “bombyperez” Rog 385,


Ciénega de los Pinos, Aguascalientes.

Mammillaria aff. bombycina “bombyperez” Rog 385,


Ciénega de los Pinos, Aguascalientes.

We saw an incredible number of M. bomby-


cina aff., an intermediate form of M. bombycina
and M. perezdelarosae, which Thomas Linzen
had once jokingly designated M. bombyperez.
The size corresponds to that of a M. perezdela-
rosae and some specimens also have somewhat
summary

different spines, but always much brighter.


Some specimens sprouted and many had four
median spines. The flowers have much similari-
ty to M. bombycina, the fruits likewise, the seed
again with M. perezdelarosae.

en or Mammillaria
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Rocks descent with M. densispina. ROG 178 Cienega


de los Pinos.
summary

The descent through the forest into a dry


streambed was then less laborious.
In the descent there are rocks with M. densi-
spina growing alongside M. higuerensis and M.
bombycina aff.

en or Mammillaria
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Mammillaria higuerensis, Ciénega de los Pinos,


2441m, Sierra El Laurel, Aguascalientes.

Mammillaria higuerensis, Colomos, Aguascalientes.


summary

en or Mammillaria
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Mammillaria perezdelarosae, Los Munos, Aguasca-


lientes.

Landscape near El Sauz, Aguascalientes. Mammillaria perezdelarosae, Los Munos, Aguasca-


lientes.
summary

en or Mammillaria
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summary

Mammillaria bombycina Rog 383 El Maguay, 2044 m,


Aguascalientes - a young specimen once H. Rogo-
zinski’s plant, now in Mihai Crisbășanu’s collection,
Bordes, France (photo from May 2017).

en or Mammillaria
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Notes on
summary

Mammillaria
aff. candida SB 827 Guaxcama, SLP

Elton Roberts

With supplementary photos from habitat kindly given to us, in alphabetical order, by:
Grzegorz Matuszewski, 13 photos - web: www.kaktusymeksyku.pl
Stefan Nietzschke, 12 photos - web: www.turbinicarpus.net

M
aybe some year the genus
of this plant will come in
for a landing; it has been Mammmillaria candida, Amparo, Coahuila.
listed as Neomammillaria,
Mammillaria and then in
the subgenus Mammil- 2006) says ‘Seed morphology excludes this spe-
Photo by Grzegorz Matuszewski.

loydia and the latest is the cies from Mammillaria. It may be a stabilized in-
summary

genus Mammilloydia. It is tergeneric hybrid, though Epithelantha seems a


the only plant in the genus Mammilloydia. It is more likely candidate as maternal ancestor than
the seed that makes the difference as the seed Neolloydia.’ That makes me wonder what Mam-
is different from the seed of the genus Mammil- millaria I would have to cross with an Epithelan-
laria. Hunt in the NCL (The New Cactus Lexicon, tha to come up with Mammilloydia candida.

Notes on Mammillaria candida 75


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Mammmillaria candida, spines.

Here is the description from The Mammillaria Mammmillaria candida, El Barreal, Nuevo Leon.
Handbook by R.T. Craig (1963 [1945]):
Body simple and cespitose from the base and
body, globose, later cylindric, sunken at apex,
5 – 7 cm wide. Tubercles arranged in 13 and
21 spirals, pale bluish green, cylindric, nearly
clavate, obtuse at tip, with watery sap, 10 mm
long, 5 mm wide at base. Areoles round with
scant white wool, later becoming naked. Axils
with 4 – 7 white bristles to length of tubercles.
Central spines 8 – 12, 4 – 7 mm long, acicular,
little heavier than radials, straight, smooth, stiff,
white, brown at tip, middle one porrect, spread-
ing. Radial spines over 50, 5 – 9 mm long, slen-
der acicular, straight, smooth, white, horizontal
to somewhat ascending in confusion. Flowers
funnelform, near top, 20 mm long, 15 mm wide.
Outer perianth segments greenish light brown
to nearly white, rose red to brownish mid stripe,
Photos by Grzegorz Matuszewski.

darker ventrally, lanceolate, tip acuminate, mar-


gins ciliate, 10 mm long. Inner perianth seg-
summary

ments muddy rose mid stripe, white margins,


lanceolate, tip acute, margins serrate, 20 mm
long. Filaments and style rose red. Anthers
golden yellow. Stigma lobes 6, purple red. Fruit
red, elongate, seeds black, glossy.

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Photo by Stefan Nitzschke.
Photo by Stefan Nitzschke.
summary

Photo by Grzegorz Matuszewski.


Photo by Stefan Nitzschke.
Mammmillaria candida, La Menchaca, Coahuila. Mammmillaria candida, Galeana, Nuevo Leon.

Mammmillaria candida, San Jose Del Llano,


Tamaulipas.

Photo by Grzegorz Matuszewski.

In his seed list Steven Brack listed the seed as


summary

Mammillaria seed “sp. SB827 Guaxcama, SLP, a


gypsum cliff dweller”. This was years and years
ago and I ordered some seed and I got seed-
lings and in time they turned out to be plants
that match fairly precise the plants of Mammil-
laria candida.

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But when the plants bloomed the flowers did


not quite match any of the Mammillaria candida
that I already had. These plants had flowers
about the same size but they are a wonderful
pink color. After some years I finally made tags
for my plants and gave the plants the name of
Mammillaria candida SB 827. Many years later I
still have that name on them.
When the plants of SB 827 are not in flower
it is impossible to tell the difference in them
and any other Mammillaria candida. In his book
Mammillaria, John Pilbeam does not mention
SB 827 in his list of field collection numbers. I
am not sure if he knew what the plant is or not.
summary

en or Notes on Mammillaria candida 78


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Steven Brack has sold his business known as


Mesa Garden and the new owners are trying

Photo by Stefan Nitzschke.


to continue the business. I see that the above
mentioned seed is still in the catalog under the
old listing but it is also listed as; ‘Mammillaria
Mammmillaria candida, La Escondida, Nuevo Leon. aff. candida SB827 Guaxcama, SLP’. I do not
know how long the seed has been listed both
ways and I do not know if Steven Brack listed
it both ways in the last few years. I see that it
Mammmillaria candida, Amparo, Coahuila. is listed with two catalog numbers, the ‘affinity
candida SB827’ is listed under catalog number
592. The seed listed as sp. SB 827 Guaxcama.
SLP is under catalog number 968.01

Photo by Grzegorz Matuszewski.


summary

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I grew this plant from seed. In one of the pic-


tures you can see a seedling that is about 2.5
cm in diameter and tall. Here the tubercles are
far enough apart that the spine clusters do not
overlap. At this stage of growth there are about
40 radial spines and 4 to 6 central spines that
I can see. I cannot say that the areoles have
scant wool for the areole is obscured with white
wool. When getting larger the plants get a much
denser spination. Werner Reppenhagen in his
book Die Gattung Mammillaria Monographie
(1961/1962) says that the plants have 65 to 119
radial spines that are to 12 mm long. And the
Photos by Grzegorz Matuszewski.

plant has 7 to 16 central spines 1 to 7 mm long.


Pilbeam says that Reppenhagen made about
summary

30 collections of the plants. As can be seen the


spine clusters in the older plant overlap each
other. I have decided not to try and count the
radial spines for I am sure that I cannot do it Mammmillaria candida, La Roca, Tamaulipas.
without getting mixed up.

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Mammmillaria candida, La Menchaca, Coahuila. Mammmillaria candida, La Escondida, Nuevo Leon.

Mammmillaria candida, La Negrita, San Luis Potosi.


It grows first as a solitary
stemmed plant, but it could at any
time start to offset. At this stage
the spines are much denser and
it looks like there are a lot more
than on younger plants. Even
when I enlarge the photo I can-
not come close to counting the
radial spines. I can count some of
the centrals but they also blend
in with some of the radials so it is
impossible to get a correct count.

Photos on this page by Stefan Nitzschke.


summary

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Comparing younger plants with older plants it can be noticed


that the spine clusters on younger plants there are a lot fewer
radial spines than what there is on much older plants. Here a
person could almost count the spines and not come out with a
number that is 30 to 40 or so wrong. The spines are in several la-
yers making clumps here and there on areoles. The radial spines
are chalk white and some are semi translucent. The centrals on

Photos by Grzegorz Matuszewski.


this plant are pinkish at the base and darken to pinkish gray color
at the tips. Some of the central spines remain pinkish from base
to tip and are semi translucent. The radial spines on older plants
are chalk white and so dense that I cannot tell if they are translu-
cent but the centrals remain lighter in color all the way from base
to tip. Mammmillaria candida, Cuevas,
Nuevo Leon.

Mammmillaria candida, La Mu-


ralla, San Luis Potosi.
Photos by Grzegorz Matuszewski.
summary

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It is also interesting to compare the flowers of younger plants to those of older plants. We have
here pictures of two individual flowers. The flower of the younger plant has 10 petals, the flower on
the older plant has 18 to 20 petals. It is easy to see the muddy pink midstripe up the petals. The style,
stigma and the filaments all look to be the same color of pink. But we also have exceptions; there the
colors are much lighter and the flower looks mostly white. I do have Mammillaria candida that has
white flowers. Before I got SB 827 all the Mammillaria candida plants I had seen had white flowers
with a very narrow pinkish midstripe up the petals. Not all my plants are SB 827 but also Mammillaria
candida with no collection data. Their flowers are different from that of Mammillaria candida SB 827
and also the spine clusters are much denser than what those on SB 827 are. Maybe this is why Steven
Brack has kept the plant with just a collection number on it for all these years and not given it a name.
summary

en or Notes on Mammillaria candida 83


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Photo by Grzegorz Matuszewski.

Photo by Stefan Nitzschke.

Mammmillaria candida, San Jose de Palmas, San Mammmillaria candida, Buenavista, San Luis
Luis Potosi. Potosi.

Mammmillaria candida, North Matehuala, Mammmillaria candida, North Matehuala,


San Luis Potosi. San Luis Potosi.
Photo by Stefan Nitzschke.

Photo by Stefan Nitzschke.

Mammillaria aff. candida SB827 Guaxcama, SLP is the name that is now given for the plant in the
latest seed list from Mesa Garden. If in fact the plant is enough different from Mammillaria candida,
then Steven Brack would be right in listing it as Mammillaria sp. SB 827 Guaxcama SLP for all these
years. If it is different enough someone in time will decide that it should be named even if it is a variety
of Mammillaria candida. The SLP stands for San Luis Potosi one of the Mexican states. Mammillaria
summary

candida comes from a large area and from four different Mexican states. One state that Mammillaria
candida grows in is San Luis Potosi. One area that is mentioned in the books for habitat of M. candida
is Cerritos SLP. Cerritos is only about 12 miles from Guaxcama. Guaxcama is in close vicinity to other
places listed where Mammillaria candida grows; specifically, south of Cerritos and west of Cerritos. So
SB 827 comes from the area where M. candida grows.

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Mammmillaria candida, Rayones, Nuevo Mammmillaria candida, El Barreal, Nuevo


Leon. Leon.

Mammmillaria candida, La Soledad, Nuevo


Leon.

Photos on this page by Grzegorz Matuszewski.


summary

en or Notes on Mammillaria candida 85


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Mammmillaria candida, La Escondida, Nue- Mammmillaria candida, Galeana, Nuevo


vo Leon. Leon.

Mammmillaria candida, San Jose del Sitio,


Nuevo Leon.

Photos on this page by Stefan Nitzschke.


summary

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The elevation of habitat varies from about


1600 feet (488 meters) to as high as 8200 feet
(2500 meters). If the plant you have was from
8000 feet (2438 meters) elevation, it will take
Photo by Grzegorz Matuszewski.

temperatures down to 20 F (-6.6 C) but if the


summary

plant is from only 1600 feet (488 meters) it


should be kept above freezing. As there is no
way of knowing the elevation your plant came
Mammmillaria candida, La Lagunita, Nuevo from then it is wise to keep the plants above
Leon. freezing.

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Photo by Grzegorz Matuszewski.


summary

Give the plants good light for good dense


spine growth, also give a fast draining soil mix.
The plants do not like having soil that stays Mammmillaria candida, La Trinidad, Nuevo
damp too long! Leon.

en or Notes on Mammillaria candida 88


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Leuchtenbergia
summary

principis Hook.

the odd cactus

Juan Miguel Artigas Azas - e-Mail: artigas659@hotmail.com - web: www.juanartigas.org

W
A view from Hidalgo, a habitat of Leuchtenbergia
principis.

hen I first saw the a picture of a flowering plant of what seemed


plant in my rookie days to be what I had saw, but the picture was not
of cactus sightseeing I clear enough so I could be sure about it.
just didn’t know what I then decided to come back to visit the
to think, it would not plants again.
resemble any cactus Luck stroke on me as they were in flower,
I had seen, and I did which was in mid-July. The flowers left no
not considered it to be doubt, it was certainly a cactus. Flowers, steam-
one. The specimen I found was neatly camou- ing from the areole at the tip of the new tu-
flaged among branches under and bush, hard bercles, one or occasionally two in each plant,
to spot. I remember to have taken a good time were big and yellow with very long petals.
examining the plant. Could it be a weird aga- Their silk-like shine under the sun astonished
summary

ve? That was what I decided at the time. It was me with its beauty. Flowers were so promiscu-
less than a year after that first sight that I got ous that quickly exposed the fact that the plant
my copy of the just newly available “The Cac- was not rare in the area, but many were found.
tus Family” book by Edward Anderson. In go- Interestingly enough rarely two clusters near
ing through the pages I stumbled (p. 395) with to each other.

Leuchtenbergia
Leuchtembergia principis
principis 89
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Leuchtenbergia principis, by Walter Hood Fitch, on Plate 4393, On researching the plant, it turns out
in Curtis Botanical Magazine, Volume 74, 1848. it had been described back in 1848 by
Below: One of the first pictures of the species, from The Cac-
William Jackson Hooker (1785-1865), an
taceae, by N. L. Britton & J. N. Rose, volume 3, page 108, “Figu-
re 117a is from a photograph of a plant sent by Dr. Elswood English botanist who was also a skilled
Chaffey from Zacatecas, Mexico, in 1910”. botanical illustrator, as well as member
of the Royal Society. The description of
the plant comes with a beautiful illustra-
tion (Plate 4393) by Walter Hood Fitch of a
flowering individual that leaves no doubt
of its identity.
The plant is based on a thick (five to ten
cm) stem that develops over time, loo-
king as if formed by the persistent bases
of old tubercles. The tubercles are green
and triangular, and as long as 15 centi-
meters, they end in long extensions that
summary

rightly serve the plant as camouflage as


they spread in all directions. They were
likely natural adaptations from old spines.
Public domain.

Public domain.

All this make the plant as high as 30 cm,


which is also the approximate diameter
that they reach.

en or Leuchtenbergia
Leuchtembergia principis
principis 90
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William Hooker got his specimens from the any such name was recorded. Quite a weird
summary

neighborhood of Real del Monte [officially Min- name for a Mexican plant you should think. It
eral del Monte] in the Mexican state of Hidalgo turns out that Leuchtembergia was coined by
by the favor of John Taylor, and recognized it somebody else without telling Hooker (likely
uniqueness by describing a full monotypic John Taylor) or even misleading him, to hon-
genus for the plant. The given generic name or an amateur botanist, the late Eugène de
Leuchtenbergia was told to him to be the name Beauharnais, prince of Eichstätt and Duke of
by which the plant was known in the continent, Leuchtenberg (1781-1824), Napoleon’s step
and he willingly adopted it, although recogniz- son from his first marriage with Joséphine de
ing that he had failed to find the place where Beauharnais.
summary

en or Leuchtenbergia
Leuchtembergia principis
principis 91
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The specific name principis could not had


been more proper for a monotypic genus, as
in Latin it means “leader or the pack”, some-
thing that has proven quite right to our days,
and likely will remain so in view of the exten-
sive exploration that cactus territories without
producing a second form, but you never know.
summary

en or Leuchtenbergia
Leuchtembergia principis
principis 92
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Leuchtembergia principis is actually known as Potosí, Hidalgo and Zacatecas (Anderson, 2001)
the cactus agave, a very suitable common name but we would also have to add at least Nuevo
summary

which made clear to me that my early confusion León and Tamaulipas to that distribution, where
was shared by other people. It is now known I have seen the plant in a number of locations.
that it is very widespread but rare from Coahuila The International Union for the Conservation
to Hidalgo in Chihuahuan desert vegetation. It IUCN, citing a supplied report by Water Fitz-
can be found in Coahuila, Guanajuato, San Luis Maurice, extends the range to Durango.

en or Leuchtenbergia
Leuchtembergia principis
principis 93
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The plant seems to prefer bushy areas in flat


lands, where it is often seen hiding at the base
of short nurturing bushes, with an occasional
plant growing in the open, not very commonly
in my experience tough. It seems susceptible
to the direct sun (as opposed by some reports
that claim it to be a full sun plant). In my experi-
ence exposed plants don’t look too healthy, but
nevertheless, show nice red rims in the triangu-
lar tubercles, not seen in well nurtured plants.
summary

en or Leuchtenbergia
Leuchtembergia principis
principis 94
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summary

L. principis and Lophophora williamsii.

en or Leuchtenbergia
Leuchtembergia principis
principis 95
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After my 2001 experience with wild flowering


plants, it had to pass 12 years before I could
finally see more plants in flower in natural habi-
tat, this in spite of numerous visits to several
populations. Flowers just last for a day, and I
have never been able to predict their blosso-
ming, which regularly occurs between the last
part of June and the first part of July, but it is
apparently absent in dry years.
summary

en or Leuchtenbergia
Leuchtembergia principis
principis 96
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The IUCN lists this plant with registration I have had the opportunity to grow this plant
152572 as of Least Concern, founding the opin- from seeds and it has been a very nice experi-
ion on the extensive field experience of W.A. ence. Seeds are easy to saw in closed clear plas-
Fitz-Maurice. The same report estimates the en- tic containers by spreading some seeds on the
tire population in 500,000 individuals, although surface of a mix of humid peat moss and fine
it is not clear the methodology used for such an coarse sand. Small plants are easy and reason-
estimation. The Mexican government in how- able fast to grow in humid well lighted condi-
ever more conservative, and in the Mexican Of- tions. The plantlets make from the beginning
ficial Norm (NOM-059-SEMARNAT-2001), it is lis- beautiful copies of larger specimens. I am never
summary

ted under category “threatened”. I tend to agree tired to watch this plant and I eagerly wait for
with Fitz, while the plant is rare in any given loca- their yearly blossoming, which always bring to
tion, it is very widespread. For the IUCN, the cur- my memory the incredible beautiful shine of
rent main concern for the conservation of this their yellow silky-like petals, another marvel of
plant in habitat is its illegal collection. the cactus world.

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principis 97
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summary
summary

References
Anderson, Edward F., 2001, The
Cactus Family, Timber Press.

en or Leuchtenbergia
Leuchtembergia principis
principis 98
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Tephrocactus
summary

geometricus (A.Cast. ) Backeb.

Part 2: the habitat pictorial

Robert Bader

A
s we have announced in Xe-
rophilia issue No.20, we con-
tinue to present the strange
Tephrocactus geometricus, in
a habitat pictorial taken on a
segment of Ruta National 60,
traversing Paso de San Fran-
cisco, in Catamarca province,
summary

Argentina, not so far from the Chilean border.


Without having a proper text, this pictorial
intends only to emphasis the variability of the
shape of this species in a very small areal which
was inspected.

Thephrocactus
Tephrocactusgeometricus
geometricus - part 2 99
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summary

RB 3268
Tephrocactus geometricus fma. inermis
1820 m
27°36’54.4 S - 67°42’40 W

en or Thephrocactus
Tephrocactusgeometricus
geometricus - part 2 100
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summary

RB 3269
Tephrocactus geometricus
1820 m
27°36’54.4 S - 67°42’40 W

en or Thephrocactus
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geometricus - part 2 101
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RB 3271
Tephrocactus geometricus
1885 m
27°38’00.1 S - 67°43’24 W
summary

en or Thephrocactus
Tephrocactusgeometricus
geometricus - part 2 102
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RB 3274
summary

Tephrocactus geometricus
2000 m
27°39’43.4 S - 67°45’41 W

en or Thephrocactus
Tephrocactusgeometricus
geometricus - part 2 103
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RB 3275
Tephrocactus geometricus fma. minimus
2920 m
27°44’38.1 S - 67°59’53 W

T. geometricus is not rare and it is easy to find


along Ruta National 60 to Paso San Francisco.
As you already could see, in deeper regions
from about 1800 m, the species forms large
groups, partly with strong thorns, but also al-
most naked.
At higher altitudes at 2900m you will only
find small plants. These sprouted little or not at
all. We call this form fma. minimus.
The accompanying plants that are found
in the same habitat are Echinopsis leucantha,
summary

Pterocactus spec., Tunilla soehrensii, Puna bon-


nieae, Cumulopuntia boliviana and Lobivia fama-
timensis var. bonnieae (if one is lucky enough
to see it). These species mostly grow in gritty
gravel but they are not on growing on rock.

en or Thephrocactus
Tephrocactusgeometricus
geometricus - part 2 104
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summary

RB 3275
Tephrocactus geometricus fma. minimus
2920 m
27°44’38.1 S - 67°59’53 W

en or Thephrocactus
Tephrocactusgeometricus
geometricus - part 2 105
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RB 3275
Tephrocactus geometricus fma. minimus
summary

2920 m
27°44’38.1 S - 67°59’53 W

en or Thephrocactus
Tephrocactusgeometricus
geometricus - part 2 106
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Mexican wilderness
summary

through the eyes


of an artist

Leo Rodriguez

Some of these pictures have been displayed on National Geographic site.

S
ometimes, wandering
in the wild means more
than anything else, al-
lowing us to see and to
enjoy all what Mother Na-
ture can offer us, and she
gives us a lot of wonder-
summary

ful gifts, from landscapes


and living beings, to colors, sounds and Echinocactus platyacanthus, after the rain, waiting
perfumes. in the light mist a full sun bath - Sierra del Doctor,
On the following pages we will pre- Queretaro.
sent you only images…

Mexican wilderness through the


Mexican
eyes of
wilderness
an artist 107
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summary

Another E. platyacanthus, a young plant after the


rain - Sierra del Doctor, Queretaro.

en or Mexican wilderness through the


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eyes of
wilderness
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As if they want to have a better view on the magni-


ficent Sierra del Doctor, some plants grow on the
edge of high cliffs and rocks.

en or Mexican wilderness through the


Mexican
eyes of
wilderness
an artist 109
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summary

Both photos on this page are images of Rapicactus


subteraneus‘ kingdom in Nuevo Leon.

en or Mexican wilderness through the


Mexican
eyes of
wilderness
an artist 110
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summary

Aztekium hintonii, growing on gypsum-rock cliffs,


in the dry season, as you can see the testimony of
the dried maniples of Selaginella. Look at the giant
one, on the edge of the cliff! It must have watched
a century long secret things unfolding...

en or Mexican wilderness through the


Mexican
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an artist 111
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Above, Mammillaria perezdelarosae ssp. andersonia-


na, growing on mineral soil, in Zacatecas; Below, so
alike and so different, its sister, M. perezdelarosae
ssp. perezdelarosae, living on a similar kind of soil,
in Aguascalientes. Look at these plants so different
from what we can see in culture!
summary

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Big and old Aztekium hintonii and its more younger


descendants, on the same gypsum-rock cliff, in
Nuevo Leon.

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Above and below, Mammillaria perezdelarosae ssp.


perezdelarosae, plants living protected by some
dried moss, in Aguascalientes. Again, look at these
plants so different from what we can see in culture!
summary

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Mammillaria perezdelarosae ssp. perezdelarosae,


plants living protected by some dried moss, in
Aguascalientes.

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Yucca in Zacatecas.

Go and search for Rapicactus subteraneus! You can


find him, here in... Nuevo Leon.
summary

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After the rain, this Hehtia is waiting for sunshine


between the rocks..

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Both pictures are a testimony of the power that life


can have in the most arid places: look at these spe-
cies growing together on a sterile gypsum-rock cliff.

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The unique Aztekium hintonii is that kind of species


you can see thousands of times with the same
curiosity and satisfaction.

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The evening in Zacatecas...

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I am big! I am beautiful! I am Aztekium hintonii!

en or Mexican wilderness through the


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an artist 121
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=> Not big, but delicate. And yes, beautiful! ... Mam-
millaria prezdelarosae spp. perezdelarosae.

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Ant-plants
summary

of the
Solomon Islands
Archipelago
Derrick Rowe

Photos by Brendan Cleaver

M
yrmecodia tuberosa is
an extremely widespread
and variable ochlospe-
cies that currently is con-
sidered to extend from
central Vietnam south
through Indochina, the
summary

Malayan peninsula, New


Guinea and its islands, the Philippines, Indo-
nesia, Northeast Australia and as far as the
Solomon Island archipelago in the south west Ant plants provide habitats for ant colonies at most
Pacific Ocean. heights in forest canopy.

Ant-plants ofof
Ant-plants the Solomon
the SolomonIslands
Islans 123
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Myrmecodia tuberosa “salomonensis”. M. tuberosa “salomonensis”

M. tuberosa “salomonensis” – old and young plants


on the same tree.
summary

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M. tuberosa “salomonensis” – two plants with longer stems. M. tuberosa “salomonensis” - growing at soil level.

M. tuberosa “salomonensis” – short and un-branched stem.

Very simply an ochlospe-


cies is a species very difficult
to fit neatly into our human
classification system. Conse-
quently, although this ‘spe-
cies’ has some morphologi-
cal similarities throughout
its vast range, it is so variable
that Huxley & Jebb in their
1993 revision of Myrmeco-
dia added botanically unof-
ficial ‘trinomials’ in an effort
to document this huge di-
summary

versity. (Huxley, C. R; Jebb,


M. H. P. 1993. The tuberous
epiphytes of the Rubiaceae
5. A revision of Myrmecodia.
Blumea 37, pp271- 334.)

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M. tuberosa “salomonensis” - growing at soil level. M. tuberosa “salomonensis”.

M. tuberosa “salomonensis” - cut through a


windfall plant.
summary

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M. tuberosa “salomonensis”.

Thus they named the variant (their word) found in the Solomon Is-
lands as Myrmecodia tuberosa ‘salomonensis’ and yes that is the correct
Latinised spelling of the trinomial. However, single quotation marks are
summary

correctly intended for horticultural cultivars, so their usage by Huxley


& Jebb is potentially confusing. I prefer to use double quotation marks
which are botanically legal. Indeed, cultivators may add whatever they
like to a scientific plant name as long as double quotation marks are
used.

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Hydnophytum forbesii – a thorny example. Illustra-


tion from Curtis’s Botanical Magazine.

Squamellaria kajewskii
I visited Bougainville Island in 2011, primarily to
find and photograph among others the amaz-
ing species Hydnophytum kajewskii which thanks
to the recent fascinating study by Dr Guillaume
Chomicki et al., is now Squamellaria kajewskii.
summary

Public domain.

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Around that time, I was sent photographs
summary

taken on Santa Isabel Island by Brendon Clea-


ver a helicopter pilot who had worked there
prior to my expedition. So, not only are these
photographs world firsts on the WWW. They
also show what may prove to be genetic variances
between Solomon island populations of M. tube-
rosa “salomonensis”. Certainly, Santa Isabel pop-
ulations show distinct dissimilarities to those

I photographed in the northern Solomon Is-


lands. Yet they are not absolute differences be-
cause although Bougainville Island plants were
predominantly very spiny, a few specimens
were completely spineless. Comparisons may
be made with my Bougainville Island images in
Xerophilia Special Issue No. 3 - January 2014,,
see pages 42- 45.
Santa Isabel Island sits in the middle and
eastern side of the long narrow archipelago, so
some distance from Bougainville Island in the
north. From Arawa on Bougainville Island near
where I took my photos to central Santa Isabel
Island is around 300km (186 miles.) – see map.
Hydnophytum kajewskii was made world
famous if only in ant-plant circles by Doctors
summary

Camilla Huxley and Matthew Jebb’s in the


following online report.
Recently this species and others have been Squamellaria kajewskii - on Santa Isabel Island, Cen-
placed in the genus Squamellaria in a fasci- tral Solomon Islands. These are world first photos
nating study by Guillaume Chomicki et al. on the web.

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Another windfall specimen. It is very probably a Another habitat.


Hydnophytum species.

Much was made of the scaphoid (boat) shape of artii (sic.) Fosberg, Lloydia 3, fig5 (1940.) These are
the tuber of this species in the above report and Brass 2548 and Kajewski 2389, one collected at Be-
in Odoardo Beccari’s type illustration in Malesia rande, the other on the Berande River, (Guadalcanal
2, yet their tuber forms often vary. Certainly, on Island) the field notes indicate a plant with branches
Bougainville Island a number of specimens that I pendulous from a tumid (swollen) stock inhabited by
photographed had quite rotund tubers as did one great numbers of small brown ants; the branches are
particularly massive thus venerable specimen. more than a meter long. Kajewski describes the fruit
See page 48 onward, in Xerophilia Special Issue as cream-colored, thickest at the base, tapering to a
No. 3 - January 2014. blunt point 8 mm. long 3 mm. in diameter.”
Here we have another of Brendon’s photo- From, Merr. & L. M. Perry. (Elmer Drew Merrill &
graphs of a windfall specimen surviving on Santa Lily May Perry) in Journal of the Arnold Arbore-
Isabel Island. It is very probably a Hydnophytum tum, vol 26, p18, (1945.)
species but it differs from the specimens of Hyd- Type Collection, Faro Island, Solomon Islands.
nophytum longistylum that I photographed on Bou- Its leaves and the presence of inhabitant ants are
gainville Island which have much rougher tubers. features shared with H. moseleyanum but Australian
See page 41 onward of Xerophilia Special Issue and New Guinea specimens, those that I am most
No. 3 - January 2014. familiar with, have red fruits and mostly a smooth
Although nothing has yet been officially pu- epidermis, except for some Cape York Peninsula
blished, there are hints on herbarium sheets that populations in the far north that have suggestions
Jebb & Huxley intend to sink a number of older of root-like ‘spines’. Yet as noted above it seems
names which includes H. longistylum into H. mose- it is to be lumped into H. moseleyanum. However,
leyanum. This will provide forms with quite mu- its muricate tuber and sometimes-tuberous roots
ricate (thorny) tubers to the particularly smooth- differ from Australian and New Guinea mainland
surfaced, almost shiny tubers seen in Australian H. examples of H. moseleyanum subsp. moseleyanum.
moseleyanum specimens. Thus this taxon and others surely show why
Thus the specimen above may be a form of keeping an accurate provenance of plants brought
H. moseleyanum; however, I suspect further field into cultivation is important.
work in the Solomon Islands may produce more
species. H. moseleyanum Becc. in Malesia
Furthermore, it now seems doubtful that Jebb Raccolta 2, p150, (1885). (See also)
& Huxley’s revision of Hydnophytum will ever be Type, Manus Island, Admiralty Islands, on the
published. Modern research using genetics has equator north of New Guinea Island.
probably made their work obsolete as has been Solomon Islands collection.
shown by Dr Guillaume Chomicki’s above work on Synonyms. When finally published some of
Squamellaria. these may have infra generic status:
H. brassii, S. Moore
H. longistylum Becc. (Odoardo Beccari) H. camporum S. Moore
in Malesia Raccolta 2, p152, (1885.) H. forbesii Hook f. (A very rough surfaced form.)
“Epiphytic on beach trees, common, stems numerous H. longistylum Becc.
on a large tuberous base, irregular in form and vary- H. mindanaense Elmer
ing greatly in size, with an uneven muricate (covered H. oblongum (Benth) Becc.
with short, sharp points) surface pierced by nume- H. papuanum Becc. (See Forster, P. I. in Illustra-
rous entrance-holes of the small brown ants which ted Handbook of Succulent Plants: Dicotyledons,
inhabit it. Stems 1 m or more long, often galled (?) p435, 2002.)
summary

the nodes swollen; leaves very thick and fleshy, the H. robustum Rech.
veins obscure (visible when dry); flowers white; fruit H. stewartii Fosberg,
yellow, (?) about 9 mm. long, 4 mm. diameter, with H. moseleyanum Becc. var. agatifolium (Val.) com.
two large white seeds enclosed in mucilaginous pulp. nov., Jebb & C. R. Huxley in press. (See # 2 & 3 of
From Guadalcanal Island, there are two collections, the following herbarium sheets.) Synonym H.
which in all details agree with Hydnophytum Stew- agatifolium Val.

en or Ant-plants of the Solomon Islands 130 - XEROPHILIA • Volume VI, No. 2 (21), June 2017 | ISSN 2285-3987
Online

Xerophilia
summary

magazines

Acc Aztekium Journal (Romanian) -


The Romanian Acc Aztekium journal.
Latest issue: No 46, March 2017.

Sansevieria Online (German) - The


free online journal about the genus
Sansevieria. Latest issue: No 5 (1),
2017.

Succulentopi@ (French) - The quar-


terly free online journal published by
the French site Le Cactus Francophone
finally made its comeback after more
than one year of absence. Latest
issue: No 16, May 2017.

Sukkulenten (German) - Monthly


free online journal of the FGaS -
Fachgesellschaft andere Sukkulenten
(formerly Avonia-News). Latest issue:
Vol. 10, No 6, June 2017.
summary

The Cactus Explorer (English) - The


first free online C&S journal. Latest
issue: No 18, April 2017.

Magazines
Online magazines 131
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on
n l i ne) p ag inati
(O ous
5 4 - 9733 h continu
20 it
i nt ) ISSN wsletter w
(Pr ne
5 4 - 9725 mmillaria rt, GB
20 a o
ISSN asional M ilborne P
c M
an oc id Hunt,
v
© Da
Huitzilopochtli
(who is a national Aztec deity of war, sun, human sacrifice
and the patron of the city of Tenochtitlan)

is an occasional Mammillaria
newsletter published by David
Hunt since March 2009. This
journal started to be published
few years after finalizing the
immense amount of work put into
The New Cactus Lexicon. There
are only 11 editions published so
far, in sequential page numbering,
but further issues are planned.
Graham Charles has introduced a
link on his The Cactus Explorers
website allowing free access to
digitized versions.
Last issue March 2017.
summary

Magazines
Online magazines 132
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The Chileans
is a journal dedicated to South American cacti rers such as Ritter, Horst, or Buining. The group
published by a group founded in 1965, founded was in fact very active and weekly meetings
by John Donald, David Whiteley and Harry Mid- were held, where talks were given by members,
dleditch. The aim was to exchange informa- followed by discussions. The weekly meetings
tion, share photographs and allow to exchange were held until 2003. With the mid-1970’s The
plants. The journal started to be published in Chileans appeared once or twice a year, and
1966, in a time when more information was with 1985 (excepting for two editions in 2006)
summary

becoming available and access to remote habi- only once a year. Graham Charles was involved
tats was much easier than in previous decades. in the production of the journals since 1994. He
Very popular, the journal was appearing several has introduced two links on his The Cactus Ex-
times a year and included exquisite information plorers website allowing free access to digitized
on new species just discovered by explo- versions of this bibliographical marvel!

Magazines
Online magazines 133
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ABSTRACT - scurtă prezentare a articolelor
summary

Complexul Thelocactus rinconensis pagina 5


Rodrigo H. González G. ~ cu foto de: Grzegorz Matuszewski, Pavel Pavlíček &
Manuel “Melo” Salazar González

Complexul T. rinconensis este unul dintre cele mai fascinante din cadrul genului Thelocactus. Forma
plantelor, dimensiunile și dispunerea spinilor, variațiile de culoare ale epidermei, florile, caracteristi-
cile de sol pe care cresc plantele în habitat, toate aceste elemente au fascinat colecționarii, de când
a apărut specia și până azi. Textul prezintă opinia biologului Rodrigo H. Gonzalez G. asupra modului
cum ar trebui să fie clasați membrii complexului. Mai mult de 80 de fotografii completează textul.

Navajoa peeblesiana Croizat., in habitat pagina 55


Stefan Nitzschke & Carmen Bechara

La al treilea articol în paginile Xerophiliei, prietenul nostru din Germania, ne împărtășește atât mo-
mentul când a găsit Navajoa peeblesiana, cât și câteva fotografii ale acestor plante care cresc într-un
habitat, cu spărturi de pietre și stânci rotunjite, mărturii ale ultimelor glaciațiuni.

Mammillaria bombycina Quehl. pagina 65


Helmut Rogozinski

Încă o foarte interesantă prezentare, care vine în completarea articolului din numărul trecut.

Note despre Mammillaria aff. candida SB 827 pagina 75


Elton Roberts ~ cu foto din habitat de: Grzegorz Matuszewski & Stefan Nitzschke

Notele lui Elton Roberts au devenit un serial neîntrerupt despre cultura unor specii, ajutând redacția
să-și îndeplinească unul din scopuri: să ofere cititorilor ei, materiale și puncte de vedere diferite privind
îngrijirea cactușilor și suculentelor. Materialul este însoțit de o frumoasă galerie de poze din habitat.

Leuchtenbergia principis Hook. pagina 89


Juan Miguel Artigas Azas

După articolul său despre Mammillaria bombycina, în acest număr autorul ne prezintă, așa cum
spune el însuși, o specie ciudată: cactusul agavă.

Thephrocactus geometricus (A.Cast.) Backeb. - partea 2 - Habitatul pagina 99


Robert Bader

Așa cum anunțam, în numărul trecut, la articolul lui Elton Roberts despre cultura T. geometricus,
continuăm în acest număr cu prezentarea speciei, în habitat: patru locații ale lui Robert Bader.

Natura mexicana prin ochii unui artist pagina 107


summary

Leo Rodriguez

Mai cunoscut cititorilor români prin seria de gravuri pe care le-a creat special pentru Xerophilia și
care v-au fost prezentate, cu trei ani în urmă, ocazie cu care, datorită succesului au fost repede epui-
zate, Leo Rodriguez ne prezintă, în numărul de față, o serie de fotografii făcute în excursiile sale.

Romanian abstract
Online magazines 134 -- XEROPHILIA
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March
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Plante-furnicar din Arhipelagul Insulelelor Solomon pagina 123
summary

Derrick Rowe

Un alt grupaj de fotografii și de texte, aparținând unui autor care are expertiză și autoritate în do-
meniu: Myrmecodia, Squamellaria, Hydnophytum...

Huitzilopochtli »»» un link spre site-ul The Cactus Explorer pagina 132
The Chilean »»» un link spre site-ul The Cactus Explorer pagina 133

Cele două linkuri de mai sus vă vor îndrepta spre o pagină a site-ului susmenționat permițându-vă
să accesați o serie de 11 broșuri editate de David Hunt despre genul Mammillaria (primul) și o serie de
73 de linkuri către o publicație consacrată cactușilor din Chile (al doilea).
summary

Romanian abstract
Online magazines 135
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CACUS & SUCCULENT FIELD TOUR MEXICO
Biologo. Rodrigo H. Gonzalez G.
Rio Yukon 419 Col. Del Valle CP 66220
San Pedro Garza Garcia N.L.
+52 81 83353764
+52 81 8115996184
rodrigo@curadoresdepaisaje.com
SEDUM
E PETROSEDUM
che nascono spontaneamente
in Italia

Author: Massimo Afferni, 2016.

Format: 21 x 29.7 cm, 160 pages, more


than 100 color photos, soft cover, text
in Italian.

The Sedum and the Petrosedum belong


to the few genera of succulent plants in
Italy. Sometimes inconspicuous, howe-
ver, manage to impress when they are
in full bloom. In this book Massimo Af-
ferni, passionate about these genera,
makes known they to us, by analyzing
all its forms and varieties supported by
high-quality images.

Published by the Cactus & Co.

To purchase the book contact e-Mail:


associazionecactusco.info@gmail.com
Cacti seeds from South America
Greatest selection from Volker Schädlich
The Chaco in Paraguay volker@gymnos.de
Bolivia
Argentina
Brazil www.gymnos.de

Acanthocalycium, Borzicactus, Cereus, Cleistocactus,


Echinopsis, Frailea, Gymnocalycium, Harrisia, Lobivia,
Opuntia, Oreocereus, Parodia, Soehrensia, Trichocereus,
Weingartia etc.
summary
summary

Advertising 139 - XEROPHILIA • Volume VI, No. 2 (21), June 2017 | ISSN 2285-3987
Xerophilia – Volume VI, No. 2 (21), June 2017

www.xerophilia.ro
ISSN 2285-3987

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