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Occurrence: Rare
Aspidella terranovica (field Locality: Lake Baikal
Rock Unit: Kurtun Formation
specimen from Flinders Ranges, Other Names: Probably belongs in
Pteridinium or Ernietta
South Australia) Description: Organism bag-like
Reference: Sokolov, 1972
Occurrence: Very common Type Specimen: Not specified
Locality: Flinders Ranges, South Classification: ?Phylum: Petalonamae
Australia; Avalon Peninsula, Barmia lobatus (from Bekker, 1996,
Newfoundland
pl. 3, no. 4)
Rock Unit: Rawnsley Quartzite,
Wilpena Group; Fermeuse Formation,
Barmia lobatus
St John's Group Occurrence: Rare, two specimens only
Aspidella terranovica
Other Names: Beltanella gilesi, Locality: Askyn River Basin, to the east
(possibly Cyclomedusa radiata, C. of the village of Solontsy, Bashkeria,
davidi, C. gigantea, C. minima, C. southern Urals
delicate, Ediacara flindersi although Rock Unit: Ashynskaya Sequence,
not all researchers would agree with Zeganskaya Series
these synonymies, [Serezhnikova, Description: Oval shaped imprint with
pers. corn., 2006.]), Glaessneria clearly defined concentric zonation,
imperfecta, lrridinitus multiradiatus, best defined in the central disc which
Jampolium wyrshykoowski, Madigania is nearly flattened center; surface
annulata, Medusinites patellaris, is deformed into folds; similar to
Paliella patelliformis, Paramedusiurn
Conornedusites, but does not have
patellaris, Planomedusites patellaris,
a four-fold symmetry; disc definitely
Protodipleurosoma rugulosum, Spriggia
deformed indicating that the surface
annulata, Spriggia wadea, Tateana
was elastic. Diameter 20-28 mm;
inflate, Tirasiana coniforrnis, Tirasiana
concentrallis, Tirasiana disciformis, central zone, 3-6 mm
Reference: Bekker, 1996
?Vendetta larini Beltanella gilesi
Type Specimen: Geological Museum.
Description: Disc-shaped and
preserved in convex relief on the Novosibersk, TSGM 1/11406 (307/89)
bases of beds. Most specimens have Classification: Likely holdfast of a
concentric rings and radial grooves. frond-like organism although originally
Originally described as several species assigned to the Cnidaria
of separate organisms; studies of
thousands of specimens suggest "Beltanella gilesi" (holotype; See
that they simply represent different
preservation styles of the same
structure, perhaps for a few different Aspidella terranovica)
organisms. When compared to the
disc that is attached to the stalk of the Occurrence: Rare
frond-like form Charniodiscus — it is Locality: Ediacara Hills, Flinders
clear that these fossils were all probably Ranges, South Australia; Mackenzie
attachment discs for such fronds. Mountains, Northwest Territories,
Obviously, from the name Cyclomedusa Canada
these forms in the past were Rock Unit: Rawnsley Quartzite,
considered to be body imprints of Wilpena Group; Sheepbed Formation,
jellyfish — but jellyfishes have four-fold Windermere Supergroup
symmetry Other Names: Synonymised with
and mouthparts, features which Aspidella: described originally as an
are not found on these fossils. The anemone
variety of patterns of concentric rings
and grooves probably simply due to Beltanella podolica
different modes of compaction and the Reference: Sprigg, 1947; Gehling et
degree of sedimentary infilling of these al., 2000 and references therein
bulbous holdfasts. Because they were Type Specimen: South Australian
embedded in the substrate, they had a Museum SAM T3-2056
much better chance of being preserved
and this explains their commonness in "Beltanella podolica" (holotype;
the fossil record
same specimen as Bronicella
Reference: Billings, 1872; Gehling et podolica)
al., 2000 and references therein
Type Specimen: Geological Survey of
Canada GSC221 Occurrence: Rare
Classification: Phylum: Petalonamae, Locality: Dniester River, Russia
attachment disc for frond-like organism Ausia fenestrata Rock Unit: Yarysev Formation
Type Specimen: Same specimen as
Ausia fenestrata (holotype) Bronicella podolica
Occurrence: Extremely rare (one hand
specimen with several "individuals")
Beltanella zimimica (TSGM 9/87)
Locality: Farm Plateau, Aus area, Occurrence: Rare Beltanella zimimica
Namibia, southern Africa Locality: Zelem River, Bashkeria, Ural
Rock Unit: Dabis Formation, Nama Mountains
Group Rock Unit: Bakyeievskaya Formation
Description: A sponge-like form, bag- Description: Inflated oval imprint with
like with rows of large pores that occur slightly displaced central disc. The
in well-defined strips. Strips appear surface of the imprint is divided into a
few sectors, which are defined by
radial ribs that originate at the central
to wrap around the body of organism
disc. Along the external border of one
as diagonals while pores themselves
form lines that lead from base to top of of these sectors there are a series of
organism. The pores are large, relative concentric furrows present. Surface of
to those found in poriferans. Organism the imprint has some three dimensions.
was certainly compressable, for in only External border is somewhat deformed
specimen known, which preserves Reference: Bekker, 1992
No 277)
Reference: Pflug, 1972 Fig. 130, Pflug Plate 28) See Ernietta
Type Specimen: Geological Survey of plateauensis
Namibia GSN (F451-H) (Pflug No 282)
Locality: Farm Aar or Plateau, Aus
area, Namibia
Erniobaris epistula" (See
Rock Unit: Dabis Formation, Nama
Fig. 130, Pflug Plate 31) See Ernietta
Group
plateauensis
Reference: Pflug, 1972
Locality: Farm Aar or Plateau, Aus
Type Specimen: Geological Survey
area, Namibia
of Namibia GSN 279 (Pflug No 279),
Rock Unit: Dabis Formation, Nama
Group Erniobaris parietalis Erniodiscus rutilus
but missing from collections in October
Reference: Pflug, 1972 2006 271
Type Specimen: Geological Survey
of Namibia GSN 268 (F449-H) (Pflug "Erniocoris orbitiformis" (See
No 268)
structures as holdfasts that resemble Mackenzie Mountains, Northwest
those of living algae and sponges Territories, Canada
Hiemalora stellaris (PIN 3993/309) Rock Unit: Ust'-Pinega Formation,
Occurrence: Common Valdai Group (Verkovka Formation
Locality: Lyamtsa, Zimnii Bereg, White of Grazhdankin, 2004); Blue Flower
Sea, Russia; Dniester River Basin, Formation, Windermere Supergroup
Podolia, Ukraine; Olenek Uplift, Siberia; Other Names: May be synonymous
Flinders Ranges, South Australia
Rock Unit: Ust'-Pinega Formation, with Pteridinium latum and P nenoxa;
Valdai Group (Lyamtsa, Verkovkaand junior synonym for Onegia nenoxa
Zimnii Gory formations of Grazhdankin, (Grazhdankin, 2004, Ph.D.)
2004); Mogilev Formation; Khatyspyt Description: Large sac-like, bilaterally
Formation; Ediacara Member, Rawnsley symmetric organism regularly
Quartzite, Wilpena Group segmented along its length. Flat
Other Names: Renamed from Pinegia segments curved and divided by fine
stellaris grooves. Arrangement of segments not
Description: Bowl-shaped body with directly opposite. Body appears to have
numerous projections, radiating from been very flexible. Width ranges from 40
outer margin of body. Morphologically to 60 mm and observable length up to
similar form was described by R. 70 mm. Judging by fragments, length
may reach up to 100 mm. Length of
Sprigg as Medusina filamentis, and segments ranges from 4.5 to 6.5 mm,
this name was later synonomized remaining constant on individuals.
with Pseudorhizostomites howchini. Reference: Fedonkin, 1979;
Diameter without "tentacles" ranges Grazhdankin, 2004 (Ph.D.)
from 3 to 40 mm; thickness of tentacles Type Specimen: Geological Institute,
from 0.3 to 2 mm; length of tentacles GIN, Moscow, 4464/147
from 3 to15 mm; max. preserved height, Classification: Although placed by
4 mm many workers in the Petaloname,
Reference: Fedonkin, 1980, 1982 Chistyakov et al., 1984 suggested it
Type Specimen: Paleontological might be a tunicate
Institute (originally Geological Institute,
Moscow, GIN4482/25) lntrites punctatus (TSGM 9/87)
Classification: Family: Hiemaloriidae — Occurrence: ?
relationships not presently understood; Locality: Zelem River, Bashkeria, Ural
in the past was placed in the Phylum: Mountains; Winter Coast, White Sea,
Cnidaria; Class: Inordozoa; see H. Russia
Ichnusa cocozzi
pleiomorpha discussion
Ichnusa cocozzi Rock Unit: Badyeievskaya Formation;
Occurrence: ? Ust'Pinega Formation
Locality: Sarrabus area, northeast of Description: Individuals of circular
Cagliari, Sardinia shape, with a wavy surface with a
Rock Unit: San Vito Formation depression in the middle, preserved
Description: Described as resembling on the sole of beds; represent forms
Albumares, with four- or eight-fold that were cylindrical in shape and
symmetry slightly tilted in orientation; diameter
Reference: Debrenne & Naud, 1981 ranges from 1 to 5 mm with smaller
Type Specimen: ? forms predominating; height ranges
Classification: ? Cnidaria from 1 to 2 mm and diameter of central
Inaria karli (holotype) depression is 0.5
Occurrence: Common Reference: Becker, 1992
Locality: Chace Range, Flinders Type Specimen: TSGM/TSNEGR
Ranges, South Australia Leningrad Museum (St Petersburg);
Rock Unit: Rawnsley Quartzite, 9/87
Wilpena Group
Description: Shaped like garlic "Irridinitus multiradiatus"
bulb, with lobate outline and hollow (holotype; See Aspidella terranovica)
center. Bulb's diameter equal in size Occurrence:
to the stalk's height (but stalk may Locality: Dniester River, Podolia,
be incomplete). Radiating internal Ukraine; White Sea Region both from
partitions (?mesenteries) separate outcrop and boreholes; ?Mackenzie
lobes and coalesce in central cavity. Mountains, northwestern Canada;
Number of lobes tends to increase with .
?Yellowhead Platform, southwestern
size, larger specimens having between Canada; Nainital District, Uttar Pradesh,
10 and 18. Most specimens external India lrridinitus multiradiatus
and composite molds on top surface of Rock Unit: Mogilev Formation; Ust'-
beds, while some specimens have basal Pinega Formation, Valdai Group; Twitya
surface cast by infilling sand Formation, Windermere Supergroup;
Reference: Gehling, 1988
Hadrynian Miette Group, Windermere
Type Specimen: South Australian
Supergroup; Krol Formation
Museum SAM P27913
Other Names: Aspidella terranovica
Classification: Some researchers
Inaria karli suggested by Gehling at al., 2000 but
have interpreted Inaria as a primitive
not all researchers agree with this
actinian (anemone) with a tubular
synonymy (Serezhnikova, pers. com,
opening and no tentacles. Others have
2006)
suggested that Inaria could be the
Description: Small rounded casts
remains of a sack-like holdfast of one
of basal part of what may have been
of the frond-like animals that has had
polyp, preserved in positive, but low,
the frond broken away and lost during
hyporelief. Central part of fossils
deposition and subsequent burial, and
commonly caved in. Numerous ridges
thus Phylum: Petalonamae?
radiate from center of fossil out to
Maria n. sp. (species manuscript margins where they intersect a narrow
name, unpublished) marginal ridge. Fossils preserved on
Occurrence: Uncommon top of beds. Diameters reach up to 20
Locality: Lyamtsa, White Sea, Russia mm; diameter of the central depression
Rock Unit: Ust'-Pinega Formation, 2-3 mm; thickness of radial canals, 0.5
Valdai Group (Lyamtsa Formation of mm
Grazhdankin, 2004) Reference: Fedonkin, 1983; Velikanov,
Aseeva & Fedonkin, 1983; Fedonkin,
1985
Description: See Inaria karli for
Type Specimen: Paleontological
description
Institute, Moscow, PIN 3994/524
lvesheadia lobata (field specimen)
Reference: Grazhdankin, 2000, 2004 Occurrence: Common
Type Specimen: Paleontological Locality: Leicestershire, UK.; Avalon
Institute, Moscow, PIN 4716/3 Peninsula, Newfoundland; South
Classification: Phylum: Australia
Petalonamae? Rock Unit: Conception and St. John's
Groups, Newfoundland; Rawnsley
"Inkrylovia lata" (PIN 3992/507; See Inkrylovia lata Quartzite, Wilpena Group lvesheadia lobata
Pteridinium simplex) Other Names: "Ivesia" lobate was a
274
Occurrence: Uncommon, only fifteen preoccupied name and thus lvesheadia
specimens known was proposed to replace it by Boynton
Locality: Suz'ma, White Sea, Russia; & Ford, 1995
structures as holdfasts that resemble Mackenzie Mountains, Northwest
those of living algae and sponges Territories, Canada
Hiemalora stellaris (PIN 3993/309) Rock Unit: Ust'-Pinega Formation,
Valdai Group (Verkovka Formation
of Grazhdankin, 2004); Blue Flower
Occurrence: Common
Formation, Windermere Supergroup
Locality: Lyamtsa, Zimnii Bereg, White
Other Names: May be synonymous
Sea, Russia; Dniester River Basin,
with Pteridinium latum and P nenoxa;
Podolia, Ukraine; Olenek Uplift, Siberia;
junior synonym for Onegia nenoxa
Flinders Ranges, South Australia
(Grazhdankin, 2004, Ph.D.)
Rock Unit: Ust'-Pinega Formation,
Description: Large sac-like, bilaterally
Valdai Group (Lyamtsa, Verkovkaand
symmetric organism regularly
Zimnii Gory formations of Grazhdankin,
segmented along its length. Flat
2004); Mogilev Formation; Khatyspyt
segments curved and divided by fine
Formation; Ediacara Member, Rawnsley
grooves. Arrangement of segments not
Quartzite, Wilpena Group
directly opposite. Body appears to have
Other Names: Renamed from Pinegia
been very flexible. Width ranges from 40
stellaris
to 60 mm and observable length up to
Description: Bowl-shaped body with
70 mm. Judging by fragments, length
numerous projections, radiating from
may reach up to 100 mm. Length of
outer margin of body. Morphologically
segments ranges from 4.5 to 6.5 mm,
similar form was described by R.
remaining constant on individuals.
Reference: Fedonkin, 1979;
Sprigg as Medusina filamentis, and Grazhdankin, 2004 (Ph.D.)
this name was later synonomized Type Specimen: Geological Institute,
with Pseudorhizostomites howchini. GIN, Moscow, 4464/147
Diameter without "tentacles" ranges Classification: Although placed by
from 3 to 40 mm; thickness of tentacles many workers in the Petaloname,
from 0.3 to 2 mm; length of tentacles Chistyakov et al., 1984 suggested it
from 3 to15 mm; max. preserved height, might be a tunicate
4 mm Intrites punctatus (TSGM 9/87)
Reference: Fedonkin, 1980, 1982 Occurrence: ?
Type Specimen: Paleontological Locality: Zelem River. Bashkeria, Ural
Institute (originally Geological Institute, Mountains; Winter Coast, White Sea,
Moscow, GIN4482/25) Russia
Classification: Family: Hiemaloriidae — Rock Unit: Badyeievskaya Formation;
relationships not presently understood; Ust'Pinega Formation
in the past was placed in the Phylum: lchnusa cocozzi Description: Individuals of circular
Cnidaria; Class: Inordozoa; see H. shape, with a wavy surface with a
pleiomorpha discussion depression in the middle, preserved
Ichnusa cocozzi on the sole of beds; represent forms
that were cylindrical in shape and
slightly tilted in orientation; diameter
Occurrence: ? ranges from 1 to 5 mm with smaller
Locality: Sarrabus area, northeast of forms predominating; height ranges
Cagliari, Sardinia from 1 to 2 mm and diameter of central
Rock Unit: San Vito Formation depression is 0.5
Description: Described as resembling Reference: Becker, 1992
Albumares, with four- or eight-fold Type Specimen: TSGM/TSNEGR
symmetry Leningrad Museum (St Petersburg);
Reference: Debrenne & Naud, 1981 9/87
Type Specimen: ? "Irridinitus multiradiatus"
Classification: ? Cnidaria
lnaria karli (holotype) (holotype; See Aspidella terranovica)
Occurrence: Common Occurrence:
Locality: Chace Range, Flinders Locality: Dniester River, Podolia,
Ranges, South Australia Ukraine; White Sea Region both from
Rock Unit: Rawnsley Quartzite, outcrop and boreholes; ?Mackenzie
Wilpena Group Mountains, northwestern Canada;
Description: Shaped like garlic
?Yellowhead Platform, southwestern
bulb, with lobate outline and hollow
Canada; Nainital District, Uttar Pradesh,
center. Bulb's diameter equal in size
India
to the stalk's height (but stalk may
Rock Unit: Mogilev Formation; Ust'-
be incomplete). Radiating internal
Pinega Formation, Valdai Group; Twitya
partitions (?mesenteries) separate AIM
Formation, Windermere Supergroup;
lobes and coalesce in central cavity. lrridinitus multiradiatus
Hadrynian Miette Group, Windermere
Number of lobes tends to increase with
Supergroup: Krol Formation
size, larger specimens having between
10 and 18. Most specimens external
and composite molds on top surface of Other Names: Aspidella terranovica
beds, while some specimens have basal suggested by Gehling et al., 2000 but
surface cast by infilling sand not all researchers agree with this
Reference: Gehling, 1988 synonymy (Serezhnikova, pers. com,
Type Specimen: South Australian Inane karli 2006)
Museum SAM P27913 Description: Small rounded casts
Classification: Some researchers of basal part of what may have been
have interpreted Inane as a primitive polyp, preserved in positive, but low,
actinian (anemone) with a tubular hyporelief. Central part of fossils
opening and no tentacles. Others have commonly caved in. Numerous ridges
suggested that Inane could be the radiate from center of fossil out to
remains of a sack-like holdfast of one margins where they intersect a narrow
of the frond-like animals that has had marginal ridge. Fossils preserved on
the frond broken away and lost during top of beds. Diameters reach up to 20
deposition and subsequent burial, and mm; diameter of the central depression
thus Phylum: Petalonamae? 2-3 mm; thickness of radial canals, 0.5
mm
Inaria nA. spA. (species
Reference: Fedonkin, 1983; Velikanov,
manuscript name, unpublished) Aseeva & Fedonkin, 1983; Fedonkin,
Occurrence: Uncommon 1985
Locality: Lyamtsa, White Sea, Russia Type Specimen: Paleontological
Rock Unit: Ust'-Pinega Formation, Institute, Moscow, PIN 3994/524
Valdai Group (Lyamtsa Formation of Ivesheadia lobata (field specimen)
Grazhdankin, 2004) Occurrence: Common
Locality: Leicestershire, UK.; Avalon
Description: See Inane karli for Peninsula, Newfoundland; South
description Australia
Rock Unit: Conception and St. John's
Groups, Newfoundland; Rawnsley
Reference: Grazhdankin, 2000, 2004
Quartzite, Wilpena Group
Type Specimen: Paleontological
Other Names: "Ivesia" lobate was a
Institute, Moscow, PIN 4716/3
preoccupied name and thus lvesheadia
Classification: Phylum: lnkrylovia lata Ivesheadia lobate
was proposed to replace it by Boynton
Petalonamae?
& Ford, 1995
274
"Inkrylovia lata" (PIN 3992/507; See
Description: Irregularly ornamented,
discoidal impressions (5 to 60 cm in
Pteridinium simplex)
diameter). lvesheadia preserved Reference: Bekker, 1996
in both positive and negative relief, Type Specimen: Geological Museum,
varying with size. Smaller "lobate disc" Novosiberisk, TSGM1/11406 (305/89)
morphs of lvesheadia (10-14 cm in Classification: Uncertain, but perhaps
diameter) feature V-shaped ridges, a holdfast
pointing inward, and outlining three
to five dumbbell—shaped, circular Kaisalia levis (holotype)
depressions. Sometimes ridges merge Occurrence: Rare, one specimen
with smooth raised and relatively Locality: Vinozh, Vinnitsa District,
entire margin. Relief of this morph of Ukraine
lvesheadia exceeds that of thickness
of crystal tuff that surrounds it, and
Rock Unit: Mogilev Formation
Other Names: See Kaisalia mensae
is greater than any other fossil forms Description: Discoidal fossil preserved
in the assemblage. Larger "bubble in positive hyporelief with convex middle
disc" forms of lvesheadia (to 60 cm in and central zones, while external zone
diameter) are ornamented by rounded is flattened; weak, concentric wrinkles
to oval craters that are largest near the occur in outer zone; total diameter 150
outside margin. Lobe-shaped hollows Kaisalia levis
mm; diameter of inner zone diameter,
lacking in specimens more than 25 37-40 mm; width of middle zone 30 mm;
width of external zone 25 mm.
cm in diameter, and arrangement Reference: Gureev, 1987
of circular hollows becomes more Type Specimen: Geological Museum,
regular with increasing size. Where Institute of Geological Sciences, Kiev
forms overlap spindle-shaped fossils, No 2127/71
it seems that lvesheadia rested over
spindles at moment of burial. Thus, it is Classification: Phylum: Cnidaria;
likely that "morphs" of lvesheadia were Class: Cyclozoa or perhaps Phylum:
result of collapse of chambered, three- Petalonamae
dimensional body. The "pizza-disc" form
Kaisalia mensae (holotype)
of lvesheadia preserved in positive
Occurrence: Rare (eight specimens)
Locality: Zimnii Bereg, White Sea,
relief; surface of pizza-disc covered Russia
with closely spaced "pustules," each
about a centimeter in diameter. Disc
Rock Unit: Ust'-Pinega Formation,
bordered by low-relief, sharp ridge.
Valdai Group
However, discovery of a stalk extending
Other Names: Perhaps a synonym of
away from the disc, in a few specimens,
Kaisalia levis
suggests that lvesheadia could have
Description: Large discoidal organism
been large holdfast onto which another
with rounded outline. Regular, flat Kaisalia mensae
structure attached. There is no
concentric rings cover entire disc. Rings
evidence to support the idea that these
separated by fine grooves. Divided into
were jellyfish, contradicting this idea of
three concentric zones. Inner part
Boynton & Ford (1995)
makes up about one-fifth of area of disc
Reference: Boynton & Ford, 1995;
and is bounded by one pair of ridges.
Narbonne et a/., 2001
The middle, broader zone makes up
Type Specimen: Holotype in situ
about half of disc. This region has
on Iva Head, Shepshed, Charnwood
seven similar-sized concentric rings all
Forest, Leicestershire SK 477 170;
of similar width. The inner and middle
casts in Leichester University Geology
regions are elevated with respect to
Department collections Accession No.
outer — this is where organism was
115577 and in Leicestershire Museums
thickest. Outer zone much thinner.
Geology Collections Accession No.
Diameter of disc up to 160 mm;
G32/1994, UK
Classification: Phylum: Petalonamae diameter of central part about 60 mm;
diameter of outer zone about 56 mm.
Individual rings in outer zone vary in
"Ivesia" lobata (See lvesheadia width from 3 to 6 mm
lobata) Reference: Fedonkin, 1984
Type Specimen: Paleontological
Ivovicia rugulosa (holotype) Institute, Moscow, PIN 4482/284
Classification: Phylum: Cnidaria;
Occurrence: Rare Class Cyclozoa or possibly Phylum:
Locality: Zimnii Bereg, White Sea, Petalonamae, probably holdfasts of
Kharakhtia nessovi
Russia frond-like organism
Rock Unit: Ust'-Pinega Formation,
Valdai Group (Yorga Formation of Kharakhtia nessovi (holotype)
Grazhdankin, 2004)
Occurrence: Rare
Description: Small bilateral animal Locality: Kharakhta River, Russia
of proarticulaten plan with oval shape; Rock Unit: Ust'-Pinega Formation,
body not deep; broad, unsegmented Valdai Group (Verkovka Formation,
head region and long trunk subdivided Grazhdankin, 2004)
into long, narrow segments or isomeres
whose ends are rounded; most
isomeres narrow and flexed posteriorly, Description: Medium to large-sized
but in anterior part of trunk, distal ends bilaterally symmetric metazoan, body
flexed anteriorly instead; axial structure Ivovicia rugulosa rounded in dorsal view, divided into
present in trunk and consists of two two rows of transverse segments
parallel ridges which continue entire (isomeres) positioned in alternating
length of trunk order ("gliding reflection") along its
Reference: lvantsov, in press longitudinal axis. Segments decrease in
Type Specimen: Paleontological size from one end to other, their lateral
Institute. PIN 3993/5504 ends recurved posteriorly. Normally five
Classification: Phylum uncertain, pairs of segments present. Margin of
but in the past has been placed in
the Phylum Proarticulata based on its body covered extensively with coarse
"symmetry of gliding reflection," where radial folds, probably present in living
segments meet along the midline in a organism and not due to postmortem
zigzag fashion distortion. Shows similarity to Yorgia
and Vendia
"Jampolium wyrzhykoowskii"
Ivantsov, Malakhovskaya &
Serezhnikova, 2004
(See Aspidella terranovica
Jurtia paliji; from Bekker, 1996, pl. 3,
no. 2) Type Specimen: Paleontological
Occurrence: One specimen Institute, Moscow, PIN4852/250
Locality: Karanyurt River, Askyn River Jampolium wyrzhykoowskii Classification: Bilaterian but Phylum
Basin, Bashkeria, southern Urals uncertain, but in past has been placed
Rock Unit: Ashynskaya Sequence, in the Phylum Proarticulata based on its Khatyspytia grandis
Zeganskaya Series "symmetry of gliding reflection" where
Description: Disc-shaped imprint, segments meet along the midline in a 275
ovoid, with clearly defined concentric zigzag fashion. Class: Vendiamorpha;
zonation; well-defined peripheral band Family: Uncertain; but see Gehling et
and raised central disc; reminiscent of al. (2005) and Narbonne (2005) for
Khatyspytia grandis (holotype) Kullingia concentrica (tool mark;
Occurrence: Rare
Locality: Khorbosuonka River, Olenek
PIN 3995/1)
Uplift, Siberia
Rock Unit: Khatyspyt Formation Occurrence: Rare
Other Names: Some researchers have Locality: Sweden and possibly
suggested that there are similarities to Khorbusuonka Basin, Olenek Uplift,
Charniodiscus Siberia; Mackenzie Mountains,
Northwest Territories, Canada
Rock Unit: Dividalen Group; Khatyspyt
Description: Colony with large Formation: Sheepbed Formation,
attachment organ; thick base and the
Windermere Supergroup
remainder of the feather-like colony Kullingia concentrica
Other Names: A tether, spin or swing
tapers dorsally to apex; growth from
mark; bio-mechanical trace fossil
axial stem; central axis extends from
base to dorsal (distal) apex; branches Description: A series of concentric
basally thick, tapering distally and flexed rings
towards apex Reference: Foyn & Glaessner 1979;
Reference: Fedonkin, 1985 Jensen, Gehling & Droser, 1998;
Type Specimen: Paleontological Jensen, Gehling, Droser & Grant, 2002
Institute, Moscow, PIN 3995/132 Type Specimen: Swedish Geological
Classification: Phylum: Petalonamae Survey, SGU Type 22, Uppsala
Classification: Once thought to belong
to the Chondrophorina (a free swimming
Kimberella quadrata (holotype and medusoid hydrozoan), but it is now
thought to be a scratch circle, a current
PIN 3993-5542) or wave-induced rotation tool mark of an
Occurrence: Rare in Australia; more anchored, tubular organism, possibly a
common in Russia sabelliditid (Jensen et al., 2002)
Locality: Ediacara Hills, Flinders Lomosovis mmalus (holotype)
Ranges, South Australia; Suz'ma, Occurrence: Uncommon
Karakhta and Solza rivers, Zimnii Bereg,
White Sea, Russia; Nainital Distict, Uttar
Locality: Novodnestrovsky Quarry,
Pradesh, India
Rock Unit: Rawnsley Quartzite, Dniester River, Podolia, Ukraine;
Wilpena Group; Ust'-Pinega Formation, Flinders Ranges,South Australia.
Valdai Group (Verkhovka, Zimnii Gory Rock Unit: Mogilev Formation;
and Yorga formations, Grazhdankin,
2004); ?Krol Formation Ediacara Member, Rawnsley Quartzite,
Other Names: ?Solza, ?Brachina, Wilpena Group
Zolotytsia Description: Large, dendritic, colonial
Description: Elongate, boat-shaped organism. Basal, small discoid
form with complex internal structure; tall attachment. Upper part of stem with
dorsal, flexible yet stiff, unmineralized fine, tubular processes and ends in
shell; frill-like rim around outside of crown of fine, long hair-like structures.
shell — which may have been circular Such processes can also diverge from
tubular structure with protruding "finger- sides of basal stem and often bifurcate
like structures" rather than true frill.
or split in two Surface of organism
Shell sometimes reached up to 15 cm in
flat. Upper part of basal stem has two
length and 5-7 cm in width with height
semicircular processes, both containing
of 3-4 cm. Many specimens, however,
were much smaller. Beyond shell and long, tubular growths or shorter brush-
Kimberella quadrate (holotype) like ends. Near base, stem possesses
tubular ring, may have extended a "foot"
as in modern mollusks that could be two or three processes of differing
extended or contracted. Associated lengths. Processes are commonly thick
with body fossils are radiating, fan- proximally and then narrow. Surface of
shaped sets of paired scratch marks processes and base smooth, although
processes are mainly covered by Lomosovis malus
(Radulichnus), feeding traces of an
animal that fed by extracting nutrients narrow, elongate ridges. Narrow, double
from sea floor sediments — has been grooves, nearly straight, incise stem
reconstructed with paired from side to side. Based on amount of
"scolecodont" -like structures contortion and crumpling of organism,
embedded within a retractable it appears to have been fairly soft
proboscis. More than 800 specimens and flexible. Ends of processes and
known from the White Sea region of offshoots commonly brush-like. Length
Russia, preserving significant of basal stem 60-80 mm; maximum
information about this form, including width 25 to 40 mm; length of processes
not only feeding traces but crawling 60-100 mm; thickness, 6 to 11 mm
traces as well. Originally described as Reference: Fedonkin, 1983; Velikanov,
a siphonophore, then later as pelagic Aseeva & Fedonkin, 1983
medusa, still later as box jellyfish Type Specimen: Paleontological
Reference: Glaessner, 1959; Institute, Moscow, PIN 3994/418
Glaessner & Wade, 1966; Fedonkin & Classification: Phylum: Petalonamae;
Waggoner, 1997 Family: Uncertain
Type Specimen: South Australian Kimberella quadrata White Sea, Russia
Museum SAM P12734
(latex peel; PIN 3993-5542),
"Lorenzinites rarus" (See
Classification: Phylum: Mollusca; Rugoconites enigmaticus)
Class: may be related to living chitons Lossina lissetskii (PIN 3993/5057)
(amphineurans) and monoplacophorans Occurrence: Relatively rare
Locality: Solza River and Zimnii Bereg,
Kuibisia glabra (holotype) White Sea, Russia
Occurrence: Rare Rock Unit: Ust'-Pinega Formation,
Locality: Farm Plateau, Aus Region,
Valdai Group (Verkovkaand Yorga
Namibia
formations of Grazhdankin, 2004)
Rock Unit: Dabis Formation, Nama
Description: Very small, bilateral
Group
metazoan with proarticulate body plan,
Other Names: May be the same as
elongated body, consisting of head
Ernietta plateauensis; also could be the
and trunk, separated by narrow furrow;
external covering of Ausia
Description: Pickford (1995) head. Semicircular in outline, its length
paraphrases Hahn & Pflug: "a solitary 30% of total length of animal; head
'polyp' with approximately conical shape contacts only first pair of isomeres; trunk
about 10 cm long and 3.5 cm broad segmented into short paired isomeres;
at its widest." "Co-axial ribs cover the isomeres with tapering ends; isomeres
surface of the polyp. These ribs are lie oblique to long axis of body; along
all approximately the same shape and axial zone there is no segmentation
size" observed, so individual isomere pairs do
Reference: Hahn & Pflug, 1985 not meet along midline; midline axis with
Type Specimen: Geological Survey of medial, longitudinal depression; surface
Namibia GSN 2 (F545) of head region and midline zone of trunk
Classification: Uncertain, but perhaps is covered by numerous small knobs
Ernietta plateauensis (Runnegar, Kuibisia glabra Reference: Ivantsov, in press Lossina lissetskii
1992) or external surface of Ausia. Type Specimen: Paleontological
Terminology used in description implies Institute. Moscow, PIN 3993/5057
relationship to a polyp-bearing organism Classification: Phylum uncertain,
— which currently is unwarranted but in the past has been placed in the
Phylum Proarticulata based on its
276 "symmetry of gliding reflection" where
segments meet along the midline in a
zigzag fashion; but see discussion in
Gehling et al. (2005) and Narbonne Other Names: Possibly Mawsonites
(2005) concerning dickinsoniomorphs randallensis, and may be a form of
"Madigania annulata" (holotype; Aspidella
See Aspidella terranovica)
Occurrence: ? Description: Distinctive, disc-shaped
organism whose surface is sculptured
Locality: Ediacara Hills, Flinders by many lobes, which unevenly radiate
Ranges, South Australia outward from central disc. Inner lobes
Rock Unit: Rawnsley Quartzite, small and increase in size toward outer
Wilpena Group margin, where clefts between the lobes
Other Names: Synonymized with indent outer, circular edge
Aspidella terranovica and ?Spriggia
annulate, Cyclomedusa davidi
Reference: Sprigg, 1949 Reference: Glaessner & Wade, 1966;
Type Specimen: South Australian Sun, 1986; Droser et al., 2005
Museum SAM T31-2031 Type Specimen: South Australian
Majaella verkhojanica (holotype) Museum SAM P17009/P40943
Occurrence: ? Madigania annulata Classification: Mawsonites may have
been a jellyfish and later a trace fossil,
but it is more likely to be a holdfast
Locality: Maya River, Eastern Siberia
Rock Unit: Yuodoma Formation "Medusina asteroides" (PIN
Description: Skeleton irregularly 3994/263; See Medusinites
discoidal/saucer-shaped, made up asteroides)
of two flat walls often with irregular Medusina asteroides
Occurrence: ?
concentric crinkles (Glaessner in
Robison & Teichert, 1979)
Reference: Vologdin & Maslov, 1960 Locality: Ediacara Hills, Flinders
Type Specimen: ? Ranges, South Australia; Zimnii Gory,
White Sea, Russia
Rock Unit: Ediacara Member,
Classification: Family: Suvorovellidae; Rawnsley Quartzite, Wilpenja Group;
with doubtful invertebrate affinities Ust'-Pinega Formation, Valdai Group
Marywadea ovata (SAM P13754; Reference: Sprigg, 1949
Type Specimen: South Australian
1 holotype) Museum SAM T40-2021
Occurrence: Rare (six specimens only)
"Medusina filamentus" (See
Locality: Ediacara Hills, Hinders Majaella verkhojanica
• Ranges, South Australia Medusinites asteroides)
Rock Unit: Rawnsley Quartzite, Locality: Flinders Ranges, South
Medusina filamentus
Wilpena Group Australia
Rock Unit: Ediacara Member,
Rawnsley Quartzite, Wilpena Group
Other Names: Spriggina ovate
Description: Segmented form Reference: Sprigg, 1949
with head distinct from rest of body Type Specimen: South Australian
and composed of regular angular Museum SAM T68
segmental elements that meet along
midline groove. Angular segments "Medusina mawsoni" (holotype;
and head shield resemble those of
arthropod. Has blunt head with sets See Medusinites asteroides)
of ramifying branches extending from Locality: Flinders Ranges, South
center of head to frontal margin. Only Australia
two well-preserved specimens (one Rock Unit: Ediacara Member,
large and one juvenile) and four Rawnsley Quartzite, Wilpena Group
poorly preserved specimens of this
Reference: Sprigg, 1949
form are known. Cephalon wide and
Type Specimen: South Australian
possesses structures that look very
Museum SAM T39
like the digestive caecae in arthropods.
Median lobe flattened and broader than Medusinites applanatus
in Spriggina. No indication that these
Medusina mawsoni
forms had ability to flex side to side.
Although originally thought to be related (holotype?)
to polychaete worms, with further study, Occurrence: Two imprints and
they seem most similar to some kind of counterparts
arthropod Locality: Ryauzyak River, southern
Reference: Glaessner & Wade, 1966 Urals
Type Specimen: South Australian Rock Unit: Ashynskaya Sequence,
Museum SAM P13754 Zeganskaya Series
Classification: Phylum: Arthropoda; Description: Somewhat deformed
Family: Sprigginidae or Phylum: ellipse-shaped imprint with broad
Proarticulata; Class: Cephalozoa; Ma,ywadea ovate peripheral area that terminates with
Family: Sprigginidae (lvantsov, in press) finely divided edges; surface of disc
Mawsonites randellensis (SAM characterized by a number of pustules
of varying sizes, which may be due
24595) to deformation; central zone takes up
Occurrence: Uncommon about one-fourth of total surface area;
Locality: Flinders Ranges, South preserved in negative epirelief; diameter
Australia of imprint varies from 16 to 24 mm.;
Rock Unit: Ediacara Member, central depression varies from 4 to 6
Rawnsley Quartzite, Wilpena Group mm
Other Names: Possibly Mawsonites Reference: Bekker, 1996
spriggi, maybe a form of Aspidella Type Specimen: Geological Museum
Description: Disc-shaped with two Novosibersk, TSGM 1/11406 (312a/87)
distinct concentric zones of equal Classification: Uncertain, but perhaps
width delineated by sharp furrows and holdfasts
surrounding raised center. Central,
middle and outer concentric zones Mawsonites randellensis
marked by fine striae, which radiate
Medusinites asteroides (SAM
outward to edge of organism. Outer
margin scalloped due to raised lobes of P13785/6)
unequal width Occurrence: Common
Reference: Sun, 1986; Jenkins, 1992 Locality: Ediacara Hills, Flinders
Type Specimen: South Australian Ranges, South Australia; Zimnii Coast,
Museum SAM P24594 White Sea, Russia; Dniester River
Classification: Uncertain, see Basin, Podolia, Ukraine; Ryauzyak
Mawsonites spriggi. But, M. Basin, South Urals and other places;
randellensis tends to have only marginal Wernecke Mountains and Mackenzie
lobes, unlike M. spriggi, and has a Mountains, Northwest Canada;
raised central disc, characteristic of Carmarthen, South Wales Medusinites. asteroides
some forms of Aspidella Rock Unit: Rawnsley Quartzite,
Wilpena Group; Ust'-Pinega Formation;
Mawsonites spriggi (holotype) Mogilev Formation; Basa Formation,
Occurrence: Rare Mawsonites spriggi Asha Series; Sheepbed Formation,
Windermere Supergroup
Locality: Ediacara Hills, Flinders Other Names: Medusina asteroides,
Ranges, South Australia M. filamentus, M. mawsoni, Medusina
Rock Unit: Rawnsley Quartzite, mawsoni, Protolyella asteroides
Wilpena Group Description: Small, rounded form with
Type Specimen: South Australian Nadalina yukonensis (holotype)
Museum SAM T39
Classification: Phylum: Cnidaria;
Occurrence: ?
Class: Cyclozoa; Family: Medusinitidae
Locality: Yukon, Canada
Rock Unit: Siltstone Unit 1,
Medusinites paliji (holotype)
Windermere Group
Occurrence: ?
Reference: Narbonne & Hofmann,
Locality: Nemia River, a tributary of the
1987
Dniester River, Ukraine
Rock Unit: Mogilev Formation Type Specimen: Geological Survey of
Other Names: Nemiana simplex Canada GSC 83022
Description: Originally described as Classification: ?
B. simplex but later found to be distinct
form. Discoidal shape with variable size Namacalathus hermanastes
of central knob relative to size of body; Occurrence: Not common
form of central knob irregular; surface Locality: Driedornvlagte Pinacle
of outer zone in large specimens often Reef Complex, southern Namibia and
bears many concentric lines; small elsewhere
forms are similar to members of family Rock Unit: Nama Group
Beltanelloididae. Description: Stem topped by outward
Reference: Gureev, 1987 flaring cup with circular opening at Namacalathus hermanastes
Type Specimen: Geological Museum, top. Cup perforated by six to seven
Institute of Geological Sciences, Kiev openings. Small, with maximum
No. 2127/19 diameter of 2.5 cm
Reference: Grotzinger et al., 2000
Type Specimen: Museum of the
Classification: Phylum: Cnidaria;
Geological Survey of Namibia F314
Class: Cyclozoa; Family: Medusinitidae
Medusinites paliji Classification: Uncertain
Medusinites patellaris (holotype;
Namalia villersiensis (See Ernietta
same specimen as "Planomedusites
patellaris" and "Paramedusium plateauensis; field specimen, 6715)
patellaris; See Aspidella terranovica) Occurrence: Rare
Occurrence: ? Locality: Farms Buchholzbrunn and
Locality: Dniester River, Ukraine Vrede, Namibia Namalia villersiensis
Rock Unit: Mogilev Formation Rock Unit: Dabis Formation, Nama
Other Names: Aspidella terranovica, Group
Planomedusites patellaris, Reference: Germs 1968, 1972;
Paramedusium patellaris Runnegar, 1992
Reference: Sokolov, 1972, 1973; Type Specimen: No number given in
Gehling et al., 2000 Germs, 1968
Type Specimen: Same as
Planomedusinites patellaris, Namapoikia rietoogensis
Paramedusium patellaris Occurrence: Rare
Locality: Driedoornvlagte, southern
Namibia
"Medusinites" sokolovi Rock Unit: Omkyk Member, Zaris
(See Vendella sokolovi)
Formation, Nama Group (ash overlying
Locality: Ladova and Nemia rivers,
this rock unit dated at 548.8 +/-1 million
tributaries of the Dniester River, Ukraine
years)
Rock Unit: Mogilev Formation
Description: Fully mineralized modules
Other Names: Vendella sokolovi
of jointed tubules, about same width
Reference: Gureev, 1987
throughout (about 1 mm in width).
Type Specimen: Geological Museum,
Similar to early Cambrian coralomorphs
Institute of Geological Sciences, Kiev. Medusinites patellaris like Yaworiporia from Siberia. Occur on
No number given in Gureev, 1987.
pinnacle reef
Same specimen as Vendella sokolovi
Reference: Wood, Grotzinger &
Mialsemia semichatovi (holotype) Dickson, 2002
Type Specimen: Geological Survey of
Occurrence: Rare (one specimen) Namibia. GSN F623 Namapoikia rietoogensis
Locality: Zimnii Coast, White Sea, Classification: Relative of corals or
Russia sponges
Nasepia altae (holotype)
Rock Unit: Ust'-Pinega Formation, Occurrence: Rare
Valdai Group (Yorga Formation, Locality: Farms Vrede and Chamis,
Grazhdankin, 2004) Namibia
Other Names: Dzik (2002) has Rock Unit: Kuibis and Schwarzrand
suggested that Mialsemia could subgroups, Nama Group
be congeneric with Rangea and Description: Leaf-like bodies with fine
Bomakellia. Fedonkin (pers. obs., 2006) ribs subparallel to long axis and with
assigns to Rangea clearly marked margins (Glaessner in
Description: Cigar-shaped organism Robison & Teichert, 1979). Like Rangea
with narrow fusiform body and long, flat schneiderhoehni and Pteridinium
appendages on both sides, rounded simplex consists of a bundle of spindle-
at their ends. These double pennate shaped bodies. Nasepia differs from
"appendages" are largest in central both of these taxa in having petaloids
part of the body and decrease in size of smaller size and in configuration
posteriorly and anteriorly. Body is of ribs usally subparallel to long axis
elongate, smooth and not divided Mialsemia semichatovi
of petaloids, not angled. Ribs 0.1 to
into segments. What appears to be 1.0 mm wide; petaloids about 10.5 cm
posterior end tapers to abrupt point. across Nasepia altae
Narrow median ridge on body has two Reference: Germs, 1973
short furrows, one on either side. Two Type Specimen: South African r%.1
short, narrow, ridges radiate away from Museum, SAM K1088.
smooth surface of posterior end of Classification: Uncertain
Nemiana bakeevi (holotype) t',
"thorax." Anterior end of body smoothly Occurrence: ? 6,
rounded. What appears to be dorsal end
of organism crescent-shaped. Closely
spaced tubercles on lateral margin Locality: Zelem River, Bashkeria, Ural
of body at base of each appendage, Mountains
of which there are seven pairs. Rock Unit: Badyeievskaya Formation
Although similar to Bomakellia, there Description: Preserved in hyporelief and
are differences in overall proportions negative epirelief: individuals preserved as
and shape and growth pattern of the convex, oval-shaped individuals with a smooth
"appendages" surface, usually preserved in groups and
Reference: Fedonkin, 1985; Dzik, 2002 rarely as solitary individuals or two or three
Type Specimen: Paleontological individuals together. Lack clearly expressed
Institute. Moscow, PIN3993/401 concentric zonation; diameter ranges from 11
Classification: Originally classified to 22 mm
Reference: Becker, 1992
as belonging to Phylum: Arthropoda. Nadalina yukonensis Nemiana bakeevi
Type Specimen: TSGM/TSNEGR Leningrad
Class Paratrilobita; now serious Museum, St Petersburg; N1/11406 (318/87)
278consideration should be given to Classification: Possibly algae or
possible rangeomorph relationships even a colonial cnidarian (possibly an
(Dzik, 2002) anemone analogue)
Reference: Fedonkin, 1981
Nemiana simplex (holotype) Type Specimen: Geological Institute,
Occurrence: Common in some Moscow. GIN 4482/199
localities Cl assification: Phylum: ?Cni dari a;
Locality: Dniester River, Podolia, Class Cyclozoa or Phylum:
Ukraine; Zimnii Bereg, White Sea,
Petalonamae, attachment disc for frond-
Russia; Khorbusuonka Basin, Olenek
like organism
Uplift, Siberia; Aus region of Namibia;
genus recognized in South Australia Nimbia paula (holotype)
Rock Unit: Mogilev Formation Occurrence: One specimen only
(Yampol Beds), Yaryshev Formation Locality: Pilipy, Danilov Creek a
(Bernasheva Beds) and the Nagoryany tributary of the Dniester River, Ukraine
Formation (Dzhurzhevka Beds); Ust'- Rock Unit: Mogilev Formation
Pinega Formation (Yorga Formation Description: Discoidal organism,
of Grazhdankin, 2004); Khatyspyt
doughnut-shaped with central
Formation; Dabis Formation; Rawnsley
depression; preserved on soles of beds
Quartzite, Wilpena Group
in positive hyporelief; surface smooth:
Other Names: Tirasiana disciformis of
Palij?; Beltanelliformis brunsae; some central zone is knob-like; diameter of
authors consider Beltanelloides as a disc 5 mm; diameter of circular ridge
junior synonym of Nemiana but recent forming the dounut - 1 to 1.5 mm
studies by Leonov (2006 and in press) Reference: Gureev, 1987
show them to be two quite distinct Type Specimen: Geological Museum.
groups with different morphology, Institute of Geological Sciences, Kiev
ecologies and have produced - No number Nimbia paula
quite different taphocenoses - i.e.
preservation styles; Sekwia kaptarenkoe Classification: Phylum: ?Cnidaria;
perhaps Vendella Class Cyclozoa or Phylum:
Description: Impressions of Nemiana Petalonamae, attachment disc for
form positive hyporelief or negative
frond-like organism
epirelief. Casts with smooth-surface,
convex discs, usually arranged in Nemiana simplex Onega stepanovi (PIN 3992/5005)
groups, rarely as isolated individuals. Occurrence: Common
In some better-preserved specimens
and larger ones, fine concentric
grooves preserved, perhaps muscle Locality: Suz'ma (5 km from the mouth
impressions? In some cases, where of the Suz'ma River), Karakhta and
it is possible to separate cast from Solza rivers, Zimnii Bereg, White Sea,
overlying rocks, there appears to be Russia
small, rounded depression on relative Rock Unit: Ust'-Pinega Formation
flat, upper surface, perhaps oral or (Verkhovka, Zimnii Gory and Yorga
mouth opening. Nemiana individuals formations), Valdai Group
often occur in masses, sometimes
covering entire bedding plans for metres Description: Small organism with flat
- apparently a colonial organism. ovate body, except for centrally raised
Nemiana evidently a benthic, sac-
segmented zone that is shifted towards
like organism, round as viewed from
narrower end of body. Central zone
above, preferring muddy sea floors,
with five paired, longitudinally arranged
and positioned partially submerged in
the mud, often in huge concentrations. lobes, each divided by deep, broad
Nemiana concentrations were buried groove. Length of lobes gradually
in place by incoming sand avalanches, decreases toward narrow end of the
with sand filling their inner cavities body. Length 4 to 7.5 mm; width, 2.5 to
3.8 mm; width of lobes about 0.3 mm
Reference: Fedonkin 1977
Reference: Palij, 1976; Fedonkin, 1981 Type Specimen: Geological Institute,
Type Specimen: Kiev State University
Moscow, GIN 4464/57b originally
No 1818
(now part of the collections of the Onega stepanovi (latex peel)
Paleontological Institute, Moscow, PIN
Classification: Possi bly algae or 3992/5049)
even a colonial cnidarian (possibly an Classification: Phylum: Arthropoda;
anemone analogue) Family: ?Vendomiidae or another
variation is Phylum: Proarticulata;
Nimbia dniesteri (PIN 173848) Class: Vendiomorpha; Family
Occurrence: Not common Vendomiidae
Locality: Dniester River near
Novodnestrovskaya Hydroelectric "Onegia nenoxa" (PIN 3992/400;
Power Station, Ukraine PIN 3993-5542); Same specimen as
Rock Unit: Lomosov Beds, Mogilev Pteridinium nenoxa; See Pteridinium
Formation
Description: Small, discoidal organism Nimbia dniesteri
with flat and smooth central part and simplex)
clearly protruding marginal ridge that Occurrence: Common
has a trapezoidal cross-section, a Locality: Suz'ma, White Sea, Russia
structure not present on N. occlusa. Rock Unit: Ust-Pinega Formation,
Shallow, rounded depression in the Valdai Group (Verkovka Formation, Onegia nenoxa
centre of the disc (diameter 2-3 mm). Grazhdankin, 2004)
Diameter of 10 to 30 mm, width of
marginal ridge 3 to 6 mm
Reference: Fedonkin, 1983 Other Names: = Pteridinium simplex.
Type Specimen: Paleontological "P nenoxa." Junior synonyms include
Institute, Moscow, PIN 3994/384-A Archangelia valdaica, lnkrylovia late,
Classification: Phylum: ?Cnidaria; Suzmites (according to Grazhdankin,
Class Cyclozoa or Phylum: 2004 Ph.D. thesis)
(holotype)
Occurrence: Relatively rare Classification: Phylum: Porifera.
They may be the oldest known Porifera
Locality: Ediacara Hills, Flinders
Ranges, South Australia; Zimnii Bereg, Palaeoplatoda segmentata (GIN
White Sea, Russia; Dniester River
Basin, Podolia, Ukraine 4464/103)
Rock Unit: Rawnsley Quartzite, Occurrence: Rare
Wilpena Group; Ust'-Pinega Formation, Locality: Suz'ma, (5 km upstream from
Valdai Group (Verkovka and Zimnii the river mouth), White Sea, Russia
Gory formations of Grazhdankin, 2004); Rock Unit: Ust'-Pinega Formation,
Mogilev Formation Valdai Group (Verkovka Formation,
Description: Irregularly rounded-oval Grazhdankin 2004)
in shape and sculptured by strong Other Names: Could be a Dickinsonia?
concentric grooves. Rounded shape Description: Organism has flattened,
interrupted by triangular notch on one leaf-shaped body with undulating
side and a suture-line (the axis) that Ovatoscutum concentricum margin, indicating elasticity during
arises at top of notch, then continues life of the organism. Body narrow in
to opposite margin. Individual grooves middle. What appears to be bottom
broadest near axis and narrow on either or ventral side of body covered with
side of notch fine, peripheral, slightly curved ridges.
Reference: Glaessner & Wade, 1966; Segmented into fine, transverse ridges
Wade, 1971; Fedonkin, 1985, 2002 present on one side ( bottom or ventral
Type Specimen: South Australian surface?) but only weakly on other.
Museum SAM P13770 Length, more than 70 mm; width, 30
Classification: Thought to belong mm; length of individual segments, 0.6
to the Phylum: Cnidaria; Family: mm; width of median ridge, 2 mm
Chondroplidae originally but more Reference: Keller & Fedonkin, 1977
recently suggested that it may be Type Specimen: Geological Institute
related to the dickinsoniomorphs GIN 4464/101
Classification: Phylum Proarticulata;
Class: Dipleurozoa; Family:
Palaeopascichnus delicatus Dickinsoniidae (Fedonkin, 1979);
Phylum: Petalonamae (Ivantsov, pers.
(field specimen) corn., 2006, but classification not clear);
Occurrence: Uncommon in South see discussion for Dickinsonia costata
Australia; common in White Sea section "Paliella patelliformis" (Ukraine
Locality: Suz'ma, Zimnii Bereg, White specimen; See Aspidella terranovica)
Sea, Russia; Dniester Basin, Podolia; Occurence: Common in Podolia but
South Australia; Newfoundland; Wales. rarer in the White Sea sequence
Namibia Locality: Zimnii Bereg, White Sea,
Rock Unit: Ust'-Pinega Formation, Russia; Dniester River Basin, Podolia,
Valdai Group (Lyamtsa, Verkhovka, Ukraine, Ryauzyak River, South Urals Palaeoplatoda segmentata
Zimnii Gory and Yorga formations; and Central Urals
Kanilov Formation of Grazhdankin, Rock Unit: Ust'-Pinega Formation,
2004), Mogilev Formation; Ediacara Palaeopascichnus delicatus Valdai Group (Verkovka Formation of
Member, Rawnsley Quartzite, Wilpena Grazhdankin, 2004); Lomozov Beds,
Group; Fermeuse Formation; ?Name Mogilev Formation; Basa Formation,
Group Asha Series; Cherny Kamen Formation,
Sylvista Series
Description: First described as a Other Names: Aspidella terranovica
meander trace, Form A, by Glaessner. suggested by Gehling et al., 2000, but
Consists of arcuate to looped ribs and not all researchers agree with synonymy
grooves. However, many specimens (Serezhnikova, pers. com.)
show bifurcation of the sets of arcuate Description: Flat, circular body
loops and appear to represent growth composed of central tubercle with
series of serial sets of sausage-shaped flattened tip and broad, smooth outer Paliella patelliformis
tubes on bedding plane zone separated from distinct inner
Reference: Palij, 1976; Gehling et al., zone. These are separated by narrow
2000; Seilacher et al., 2003; Droser et annular groove. Radial lines present in
al., 2005 only largest specimens (diameters from
5 to 45 mm with fine, paired grooves
spaced at about 0.5 mm) in outer, but
Type Specimen: Ukranian Academy
not inner zone. Paliella is similar to
of Sciences, Institute of Geological
Cyclomedusa but differs in not having
Sciences, Kiev, IGN1907/7
concentric groove over entire structure
Classification: Probably some form
or in central part. Small specimens
of collapsed serial beads or tubes
of Paliella similar to Medusinites
growing on the sea floor. Regarded
asteroides but differ in not having
as xenophyophores, giant protozoans
distinct radial grooves that cut outer
that reach up to 25 cm today, by Dolf
zone. Specimens occur in groups or in
Seilacher. By others regarded as a
pairs, where they are pressed close to
trace fossil, but there now seems to be
each other, often rimmed by common
a case for considering it some sort of
narrow zone — suggesting that this
body fossil which needs further study
form may have reproduced by simple
and clarification. Droser et al. (2005)
division. Another type of branching
suggested that other taxa described
reproduction is suggested by some
originally as trace fossil might also
specimens that have long, stem-like
chord which extends from central part
be much the same sort of organism: Palaeophragmodictya reticulate of larger form to smaller form at end of
Yelovichnus, Intrites, some Neonereites, chord, presumably the offspring
280Orbisiana, and possibly Harlaniella Reference: Fedonkin, 1980
Type Specimen: Geological Institute,
Palaeophragmodictya reticulata Moscow, GIN 4482/51
"Papillionata eyrie" (holotype; See Parvancorina minchami (sang
Occurrence: ?
Locality: Zimnii Bereg, White Occurrence: Rare
Sea, Russia (Verkovka Formation, Locality: Novodnestrovskaya
Grazhdankin, 2004) Hydroelectric Station, Dniester River,
Rock Unit: Ust'-Pinega Formation, Ukraine
Valdai Group Rock Unit: Lomosov Beds, Mogilev
Other Names: Pinegia is a pre- Formation
occupied generic name of a Permian Other Names: Similarities to Valdaina
insect described by A.V. Martynov in plumose, ?Pteridinium latum and other
1928, a junior synonym of Hiemalora Pteridinium "species", lnkrylovia late Podolimirus mirus
stellaris, reassigned by Fedonkin in Description: Large, bilaterally
1982 symmetric organism with proarticulate
Reference: Fedonkin, 1980, 1982 body plan, composed of paired,
Type Specimen: Geological Institute, posteriorly flexed segments (isomeres),
Moscow, GIN 4482/25 wide proximally and narrowing distally.
Planomedusites grandis Segments diverge from median axis at
(Geological Museum GM 2127-70) about 80-85 degrees and then curve
Occurrence: Not common, 10 abruptly, running parallel to long axis
specimens only of organism; length of organism @ 170
Locality: Dniester River, Podolia, mm, width at least 100 mm; width of first
Ukraine proximal segment, 22 mm; nine or fewer
Rock Unit: Mogilev Formation pairs of isomeres present
Description: Discoidal form, preserved Reference: Fedonkin, 1983; Velikanov,
both as hemisphere and as flattened Aseeva & Fedonkin, 1983
fossil on sole of beds in positive Type Specimen: Paleontological
hyporelief; flattened forms show weakly Institute. Moscow, PIN 3994/417
preserved peripheral ridges; central Classification: Phylum uncertain,
knob normally conical; diameter of but in the past has been placed in
Planomedusites grandis
whole form ranges from 22 to 125 mm; the Phylum Proarticulata based on
diameter of central knob ranges from 5 its "symmetry of gliding reflection"
to 8 mm where segments meet along the
Reference: Sokolov, 1972, 1973 midline in a zigzag fashion. Class:
Type Specimen: Geological Museum, Vendiomorpha; Family: Vendomiidae
Institute of Geological Sciences, Kiev but in past classification was placed
— no number available in the Petalonamae
"Protodipleurosoma rugulosa"
Description: Organism with small clearly divided into two broad zones, 1 '''' W1 . '*
ovoid body of which more than one-third separated by depression. Outer zone . , , )
occupied by a broad, semicircular has sharp relief and regular radial I Veprina undosa
"head" region with two symmetrical ribs, separated by relatively narrow
depressions, which could be the furrows. Inner zone, similar in width
remains of eyes? Median ridge to outer zone, has less regular ribbing
originates in this area; paired, elongate and extends around an elongated
segments diverge from this median depression in center of this form.
ridge and become smaller towards Elongated central depression imparts
narrower end of body. Length, 4 mm; bilateral character to body. Diameter
width, 3 mm; width of median ridge, 0.2 approximately 60 mm; width of outer
mm; number of segments, six ribbed zone 10-15 mm; width of the
Reference: Keller & Fedonkin, 1977; inner ribbed zone 6-11 mm; width of
lvantsov, in press central depression, 2-4 mm; width of the
Type Specimen: Geological Institute ribs in outer zone 1.5-3 mm; width of
GIN 4464/57; Paleontological Institute, the fine ridges, 0.3-0.5 mm and length
Moscow, PIN3992/57 up to 15 mm. As noted in Treatise of
Invertebrate Paleontology, Veprina has
Classification: Uncertain some similarities to Peytoia from the
Ventogyrus chistyakovi (PIN mid-Cambrian Burgess Shale Fauna of
Canada
Vendomia menneri Reference: Fedonkin, 1980
4564/1009) Type Specimen: Geological Institute,
Occurrence: Moderate abundance at Moscow, GIN 4482/29
type locality Classification: Phylum: Cnidaria;
Locality: Yarnema, Onega River, White Class: Cyclozoa; Family: Uncertain
Sea, Russia
Rock Unit: Ust'-Pinega Formation, Vladimissa missarzhevskii
Valdai Group (Verkovka Formation of (holotype)
Grazhdankin, 2004). Some researchers Occurrence: Rare
have suggested it belongs to the Locality: Zimnii Bereg, White Sea,
uppermost part of the Vendian complex Russia (5 km north of Zimnegorsky
(Erga or Padun Formation) Lighthouse)
Other Names: Cf Arborea sp.
(Christyakov et al., 1984); not all
Rock Unit: Ust'-Pinega Formation,
researchers agree
Valdai Group (Yorga Formation of
Grazhdankin, 2004)
Description: Internal sand casts,
usually with egg-shape, bipolar form;
Description: Flat, leaf-shaped
three-fold, radially situated modules,
organism broad in middle with acutely
each with a large basal chamber
pointed anterior and posterior ends.
and two meridional rows of smaller
Margin finely scalloped, and two rows
chambers decreasing in size towards
of small, longitudinal tubercles present.
apical end. Three longitudinal and
Body seems to have considerable
numerous transverse soft septa
thickness (several mm). Body surface
subdivide internal space into chambers;
smooth. Along mid-body are two rows
each small chamber with some
of similar sized, rounded tubercles,
triangular minor septa; on each side of
closely spaced. Each row oriented
each major septa are complex, identical Ventogyrus chistyakovi Vladimissa missarzhevskii
longitudinally and contains four
systems of branching channels coursing
tubercles. No mouth discerned. Length
along the axial rod of this structure
about 45 mm, width, 32 mm; width
of tubercles, 1.5 mm with maximum
with alternating lateral branches, which spacing of 2 mm
bifurcate up to five times before they Reference: Fedonkin, 1985
reach outer surface; internal space of Type Specimen: Paleontological
each module subdivided into two rows Institute, Moscow, PIN 3993/204
of chambers by longitudinal septum Classification: Thought to be related
and transverse septa originating from to worms in the Platyhelminthes,
the latter; modules have a single similar in form to living rhabdocoelan
large, non-paired chamber at wide end; turbellarians
internal sides of longitudinal septa join
edges of axial pyramid; largest septa Wadea genA. nov.
of the first order extends from axial (See Rugoconites tenuirugosus)
pyramid to outer side of module, defines
chambers of first order alternating
relative to each other by about half of
Wigwamiella enigmatica
(holotype)
their length; transverse septa of second Occurrence: Rare
order are about four times shorter than Locality: Mount Scott Range, South
first order septa, and so only partially Australia
subdivide space of first order chamber; Rock Unit: Ediacara Member,
septae of the third and fourth orders Rawnsley Quartzite, Wilpena Group
are even shorter and divide space Other Names: Possibly
of corresponding chambers of the Pseudorhyzostomites, an escape
second and next smaller chambers; Wigwamiella enigmatica
structure
each face of axial pyramid and both Description: Circular to oval in shape,
surfaces of longitudinal septum bear 291
with defined outer rim and furrows
imprints of regularly branching channels
of distribution system, which consists
of the major branches; each branch
radiating from a center. Furrows structure of the isomeres but axial lobe
irregularly branch and thin towards rim not visible
where series of smaller, finer furrows Reference: Ivantsov, 1999
tend to parallel main ones. Central Type Specimen: Paleontological
part often strongly depressed (negative Institute, Moscow, PIN 3993/5024
hyporelief) Classification: Bilaterian but Phylum
Reference: Runnegar, 1991 Uncertain: in the past has been placed
Type Specimen: South Australian in the Phylum Proarticulata based on its
Museum SAM P27978 "symmetry of gliding reflection" where
segments meet along the midline in a
zigzag fashion. Class: Cephalozoa;
Classification: Phylum: Cnidaria or
Family: Yorgiidae
Unknown — could be pull away structure
when attachment bulb dislodged from Zolotytsia biserialis (holotype)
the sea bottom Occurrence: Rare
Locality: Zimnii Bereg, White Sea.
Windermeria aitkeni (holotype) Russia; Dniester River Basin, Podolia,
Occurrence: Rare, single specimen Ukraine; Nainital district, Uttar Pradesh,
Locality: Sekwi Brook North, India
Rock Unit: Ust'-Pinega Formation,
Mackenzie Mountains, Northwestern Valdai Group (Verkovka Formation
Canada of Grazhdankin, 2004); Bernashevka
Rock Unit: Upper Blueflower Beds, Mogilev Formation; Krol
Formation, Windermere Group Formation
Description: Small (16.4 mm x 7.9 Description: Two distinct rows of
mm), bipolar, parallel-sided, segmented oval shaped bodies on either side of
ovoid with semicircular terminations. mid-line groove. Each row divided by
Eight nearly equal-sized segments deep groove, narrower than median
arranged subtransverse to medial furrow groove. Bodies wide at one end and
in opposite arrangement (i.e. symmetry narrower at other. Oval bodies not
of "gliding reflection" — that is, segments symmetric, possibly due to postmortem
on one side of the midline do not match deformation
Windermeria aitkeni Reference: Fedonkin, 1981
up, they are offset)
Reference: Narbonne, 1994 Type Specimen: Paleontological
Type Specimen: Geological Survey of Institute, Moscow, PIN 3993/343
Classification: Phylum: Petalonamae; Zolotytsia biserialis
Canada, GSC 102374
Classification: Primitive bilaterian; Family: Uncertain
Family Dickinsonidae
www.vend.paleo.ru/pub/Chistyakov_et
al_1984.pdf
Yelovichnus gracilis (holotype)
Occurrence: ?
Locality: Zimnii Bereg, White Sea,
Russia