Glasnik Hrvatskog botaničkog društva 6(1) 2018.

1. Disk globose or ovoid and purple or dark brown expanding, and it is now found in natural and
Prilozi
in fruit.poznavanju
Stems hirsute flore Hrvatske
or hispid, lower leaves semi-natural
/ Contributions habitats suchofasthe
to the knowledge moist forests,flora
Croatian moist
entire or lobed; plants about 50-150 cm tall meadows, in ditches and along watercourses.
........................................................................ 2 It occurs along river banks and roads and in
Three-lobed Coneflower Rudbeckia trilobadifferent L. (Compositae):
human-made new alien
habitats. species
During in
fieldwork
2. Lower leaves deeply 3-lobed the flora of Bosnia in
or 3-divided, and
the Herzegovina
summer 2017 the species was recorded in
stems branched, hirsute, 50-150 cm tall; a few new localities in the Krivaja valley (Careva
heads many, 10short to 30;scientific minute crown Ćuprija
pappus a communication / kratkoand Kamensko
znanstveno near Olovo as well as
priopćenje
......................................................... R. triloba Maoča and Vozuća near Zavidovići). The species
Semir Maslo (Primary School, Lundåkerskola, Södrahas recently
Storgatan 45,been registered
SE-332 along Sweden;
33 Gislaved, the banks of
2. Leaves neither 3-lobed (corresponding
semmas@edu.gislaved.se) nor 3-divided, author the river
stems / autor Željeznica in the vicinity of the village
za korespondenciju)
simple,
Šemso hispid,
Šarić 30-100 cm
(JP ŠPD-ZDK tall;Zavidovići,
d.o.o heads 1 to Krupac near
3, Izetbegovića
Alije 25,Sarajevo
BA-72220(Fig. 2). It is Bosnia
Zavidovići, yet unknown
and
pappus absent
Herzegovina; .................................. R. hirta how this species was introduced into Bosnia and
semsosumar@gmail.com)
Herzegovina, most likely as a garden escapee.
Maslo, S., Šarić, Š. (2018): Three-lobed Coneflower Rudbeckia triloba L. (Compositae): new alien
species in the flora of Bosnia and Herzegovina. Glas. Hrvat. bot. druš. 6(1): 8-12. 4-7.

Abstract

Three-lobed Coneflower Rudbeckia triloba L. was
discovered in September 2016 during fieldwork
on the banks of the Krivaja River in the vicinity of
village Ribnica near Zavidovići (Central Bosnia).
Three-lobed Coneflower is native in North America
but it has been introduced to some areas beyond
its natural range. This is the first record of this
species for Bosnia and Herzegovina. The paper
presents a short morphological description and
photographs of the species based mainly on the
collected specimens, as well as the distribution
of the taxon. A determination key is given
for Rudbeckia taxa registered in Bosnia and
Herzegovina and neighbouring countries.

8
5
Keywords: alien plants, Coneflower, distribution, morphology, naturalization

Maslo, S., Šarić, Š. (2018): Rudbeckia triloba L. (Compositae): nova strana vrsta u flori Bosne i
Hercegovine. Glas. Hrvat. bot. druš. 6(1): 4-7.

Sažetak
Three-lobed Coneflower Rudbeckia triloba L. was
discovered in September 2016 during fieldwork
on the banks of the Krivaja River in the vicinity of
village Ribnica near Zavidovići (Central Bosnia).
Glasnik
Three-lobedHrvatskog
Coneflowerbotaničkog društva
is native in North America
but it has been introduced to some areas beyond
its natural range. This is the first record of this
species for Bosnia and Herzegovina. The paper
presents a short morphological description and
photographs of the species based mainly on the
collected specimens, as well as the distribution
of the taxon. A determination key 6(1) 2018.
is given
for Rudbeckia taxa registered in Bosnia and
Herzegovina and neighbouring countries.

Material andalien
Keywords: methods
plants, Coneflower, distribution, morphology, naturalization

The autumn
Maslo, S., Šarić, floristic surveyRudbeckia
Š. (2018): in 2016 in triloba R. triloba (Fig.
the areaL. (Compositae): nova 1) strana
is short-lived
vrsta uperennial,
flori Bosne withi
of Central Bosnia
Hercegovine. Glas.resulted
Hrvat.inbot. newdruš.records6(1):for4-7. 8. stem branched, 50-150 cm tall, moderately hirsute
the
8-12.
alien flora of Bosnia and Herzegovina, R. triloba. or strigose to subglabrous. Leaves are thin, sharply
Digital photographs and GPS coordinates were toothed to subentire, the basal ones broadly ovate
Sažetak
taken in field. Identification of the specimens was or subcordate and long-petiolate, the cauline
done
U rujnu according
2016. u to Perdue
Bosni (1957), Britton
i Hercegovini je po&prvi Brown mostly narrower
put prirodnog areala.and U radu short-petiolate
se donosi kratak or sessile.
opis
(1970),
u prirodiCronquist
zabilježena (1980),vrsta Urbatsch
Rudbeckia & Cox (2006)
triloba Some of thekarakteristika
L. morfoloških leaves are generally 3-lobed,
vrste, karta distribucije rarely
andobalama
na Weakleyrijeke (2007).The
Krivaje,nomenclature
u blizini Zavidovića. follows kao any iofključ
them za pinnately
determinaciju lobed, the roda
vrsta lobesRudbeckia
generally
the Euro-Med
Vrsta je autohtonachecklist za (EURO+MED
područje 2006).The acute. Headsu many
Sjeverne zabilježenih Bosni (10-30),
i Hercegovini long pedunculate,
i susjednim
specimensaliwere
Amerike, collected
je unešena and stored
u neka područja into in corymbiform arrays, terminating the branches;
the zemljama.
izvan
Herbarium of the National Museum of Bosnia and the disk ovoid, dark purple, 1-1.5 cm wide; rays
Herzegovina
Ključne riječi:(SARA-51805
naturalizacija, and SARA-51806). rudbekija,
rasprostranjenost, 8-12, yellow
straneorvrste orange,biljaka1-2 cm long; receptacular
bracts glabrous, abruptly narrowed to a short but
Results and discussion
Introduction distinct awn tip often shortly surpassing the disc
corollas. Pappus a minute crown; chromosome
Rudbeckia
The genustriloba L. (Sp. Pl.:
Rudbeckia L. 907. 1753), also Species
(Compositae) number n=19 (Cronquist
R. laciniata 1980). Inbeen
has recently its native area
described
known
has 23asspecies
Three-lobed nativeConeflower,
to Northoriginated
Americainand the asR. triloba
invasivegrows tointhemesic to wet woodlands,
country (Vojnikovićthickets, 2015)
eastern part
Mexico of North
(Urbatsch & America,
Cox 2006). andManyis a very rare, and
species pastures, roadsides
is included andpreliminary
in the meadows, check-list generally on of
locally
are naturalized
cultivated asalien in some parts
ornamentals of Europe.
outside wet soils. alien
their invasive The flowering
plant species period(IAS)
stretchesin Bosnia from Juneand
To identify
native this new species
distribution range. inSix theofflora of Bosnia
these to October. The
have Herzegovina plant reproduces
(Maslo 2016). Species by seeds which
R. triloba
and Herzegovina,
been reported asweintroduced offer here the in adjusted
Europe key are dispersed
but has been reported by the wind (Britton &asBrown
in Europe 1970).
introduced
according
only to BrittonR.& hirta
two species, Brown L. (1970), laciniata in The
and R. Cronquist Austria first(Forstner
finding of& this Hüblspecies
1971, Stöhr for Bosnia
et al.
L. are considered to be widely naturalized in 2009), Ukraine (Mosyakin & Fedoronchuk Bosnia
(1980), Urbatsch & Cox (2006) and Weakley (2007): and Herzegovina is coming from Central 1999),
Europe (Greuter 2006). In the flora of Bosnia Great in 2016, on the
Britain banks
(Sell of the2006),
& Murrell KrivajaMontenegro
river in the
1.
and Disk elongated only
Herzegovina or cylindric
two species in fruit, yellowish
of the vicinity of the
genus (Stešević village Ribnica
& Jovanović 2008),near HungaryZavidovići (Király (44°
et
Glasnik
or gray.
Rudbeckia, Hrvatskog
Stems
have been botaničkog
essentially
recorded sodruštva
glabrous,far: R. leaves
hirta al.21’ 2009),
10.18” N; 18° 23’ 35.36”
Romania (SirbuE) & (Fig.
Oprea 2). 2010)6(1) 2018.
Individuals
and
and thin,
R. usually
laciniata subglabrous,
(Slavnić 1960, pinnately divided or
Obradović in the population
& Slovenia (Jogan 2013). were numerous, with some
pinnatifid;
Budak 1982, plants about 100-300 cm
Beck-Mannagetta et tall, 1983). plants up to 150 cm high. About twenty flowering
al. usually
colonial; pappus a short crown ..... R. laciniata specimens were recorded at a linear distance of
Material and methods ca 100 m. In Central Bosnia the species is slowly
1. Disk globose or ovoid and purple or dark brown expanding, and it is now found in natural and
in fruit.
The Stems
autumn hirsute
floristic or hispid,
survey in 2016 lower
in the area 41.semi-natural
leaves R. triloba
Disk elongated habitats
(Fig. orissuch
1) as moist
short-lived
cylindric forests,
perennial,
in fruit, moist
yellowishwith
entire orBosnia
of Central lobed;resulted
plants in about
new 50-150
records cm for the meadows,
tall stem branched,
or gray. in ditches
Stems 50-150 and
essentially along
cm tall, watercourses.
moderately
glabrous, hirsute
leaves
........................................................................
alien flora of Bosnia and Herzegovina, R. triloba. It thin,
2 or occurs
strigose along
usually river banks
to subglabrous.
subglabrous, Leaves andare
pinnately roadsthin, and or
sharply
divided in
Digital photographs and GPS coordinates were toothed different tohuman-made
pinnatifid; subentire,
plants about thehabitats.
basal
100-300 onesDuring
cm tall,fieldwork
broadly ovate
usually
2. Lower
taken leaves
in field. deeply 3-lobed
Identification or 3-divided,
of the specimens was or in colonial;
the summer
subcordate pappus 2017
and the species
long-petiolate,
a short crown ...... was the R.recorded
cauline
laciniata in
donestems branched,
according to Perdue hirsute,
(1957), 50-150
Britton & cmBrown a few new
tall; mostly localities
narrower andin short-petiolate
the Krivaja valley or (Careva
sessile.
headsCronquist
(1970), many, 10(1980), to 30; pappus
Urbatsch a& minute
Cox (2006) Ćuprija
crown 1.Some Disk of and Kamensko
the leaves
globose are and
or ovoid near
generally
purpleOlovo3-lobed,
or dark as well as
rarely
brown
and .........................................................
Weakley (2007).The nomenclature R. follows
triloba any Maoča
in of andStems
them
fruit. Vozuća
pinnately near
hirsute Zavidovići).
lobed, the lobes
or hispid, Thegenerally
lower species
leaves
the Euro-Med checklist (EURO+MED 2006).The acute. has recently
entire Heads
or lobed;been
many registered
(10-30),
plants aboutalong
long the banks
pedunculate,
50-150 cm tall of
2. Leaves neither
specimens 3-lobed nor
were collected and3-divided,
stored into stemsthe inthe.....................................................................
river Željeznica
corymbiform arrays, in terminating
the vicinity the of the village
branches; 2
simple, hispid,
Herbarium 30-100 Museum
of the National cm tall; headsof Bosnia 1 toand Krupac
3, the disknear
ovoid, Sarajevo
dark purple,(Fig. 2).1-1.5 It iscm yetwide;unknownrays
pappus absent
Herzegovina
Herzegovina ..................................
(SARA-51805
(SARA-51805 and
andSARA-51806).
SARA-51806). R. hirta 2.8-12,
howLowerthis species
yellow or orange,
leaves was
deeply introduced
1-23-lobed
cm long; into Bosnia
orreceptacular
3-divided, and
Herzegovina,
bracts
stems glabrous,
branched,most likely
abruptly as a garden
narrowed
hirsute, 50-150to escapee.
a short
cm tall;but
Results and discussion distinct
headsawn many, tip 10often
to 30;shortly
pappussurpassing
a minutethe disc
crown
corollas. Pappus a minute crown; chromosome
......................................................... R. triloba
Rudbeckia triloba L. (Sp. Pl.: 907. 1753), also number n=19 (Cronquist 1980). In its native area
known
known as asThree-lobed
Three-lobedConeflower,
Coneflower, originated
originated in the triloba grows
in 2.R.Leaves neither in mesic
3-lobed to wetnorwoodlands,
3-divided,thickets, stems
eastern
the eastern part part
of North America,
of North America,and is anda very very pastures,
is a rare, simple, roadsides
hispid, 30-100 and meadows,
cm tall; heads generally1 to on 3,
locally naturalized
rare, locally alien inalien
naturalized someinparts some of parts
Europe. of wet soils. The
pappus absent flowering period stretches from
................................... June
R. hirta
To identify
Europe. Tothis new species
identify this newinspecies the florainofthe flora to October. The plant reproduces by seeds which
Bosnia
and Herzegovina,
of Bosnia we offer here
and Herzegovina, wetheofferadjusted
here key the are dispersed
R. triloba (Fig.by 1) the wind (Britton
is short-lived & Brown
perennial, with1970).
stem
according
adjusted key to according
Britton & to Brown
Britton (1970),
& Brown Cronquist
(1970), branched, The first50-150 findingcm of tall,
thismoderately
species forhirsute Bosnia or
(1980),
Cronquist Urbatsch
(1980), & Cox (2006) and
Urbatsch & Cox Weakley(2006) and and
(2007): Herzegovina
strigose to subglabrous. is coming fromare
Leaves Central
thin, Bosnia
sharply
Weakley (2007). in 2016, to
toothed onsubentire,
the banks theofbasal
the Krivaja
ones broadly river inovate
the
1. Disk elongated or cylindric in fruit, yellowish vicinity of the village Ribnica near Zavidovići (44°
or gray. Stems essentially glabrous, leaves 21’ 10.18” N; 18° 23’ 35.36” E) (Fig. 2). Individuals
thin, usually subglabrous, pinnately divided or 9 5 in the population were numerous, with some
pinnatifid; plants about 100-300 cm tall, usually plants up to 150 cm high. About twenty flowering
colonial; pappus a short crown ..... R. laciniata specimens were recorded at a linear distance of
ca 100 m. In Central Bosnia the species is slowly
1. Disk globose or ovoid and purple or dark brown expanding, and it is now found in natural and
in fruit. Stems hirsute or hispid, lower leaves semi-natural habitats such as moist forests, moist
Glasnik Hrvatskog botaničkog društva 6(1) 2018.

Material
or and methods
subcordate and long-petiolate, the cauline vicinity of the village Ribnica near Zavidovići (44°
mostly narrower and short-petiolate or sessile. 21’ 10.18” N; 18° 23’ 35.36” E) (Fig. 2). Individuals
SomeThe ofautumn floristic
the leaves aresurvey in 2016
generally in therarely
3-lobed, area in R.
thetriloba (Fig. 1)were
population is short-lived
numerous, perennial,
with some with
of Central
any of them Bosnia resulted
pinnately in new
lobed, records
the lobes for the
generally stem branched,
plants up to 15050-150
cm high. cmAbout
tall, moderately hirsute
twenty flowering
alien flora
acute. Heads of Bosnia and Herzegovina,
many (10-30), R. triloba.
long pedunculate, in or strigose towere
specimens subglabrous.
recordedLeaves
at a linearare thin, sharply
distance of
Digital photographs
corymbiform and GPS coordinates
arrays, terminating the branches; were
the toothed
ca 100 m. to subentire,
In Central the basal
Bosnia theones
speciesbroadly ovate
is slowly
takenovoid,
disk in field.
darkIdentification
purple, 1-1.5 of cm
the wide;
specimens was
rays 8-12, or subcordate
expanding, andandit is long-petiolate,
now found in natural the caulineand
done according
yellow or orange,to1-2 Perdue (1957),
cm long; Britton & Brown
receptacular bracts mostly narrower
semi-natural andsuch
habitats short-petiolate
as moist forests, or sessile.
moist
(1970), Cronquist
glabrous, abruptly (1980),
narrowed Urbatsch
to a short& Cox (2006)
but distinct Some of theinleaves
meadows, ditches areand
generally 3-lobed, rarely
along watercourses.
and tip
awn Weakley (2007).The
often shortly nomenclature
surpassing follows
the disc corollas. any
It of them
occurs pinnately
along river lobed,
banks the andlobesroadsgenerally
and in
the Euro-Med
Pappus a minutechecklist
crown; (EURO+MED
chromosome 2006).The
number acute. Heads
different many (10-30),
human-made habitats.longDuringpedunculate,
fieldwork
specimens
n=19 were collected
(Cronquist 1980). In and stored area
its native into the R. corymbiform
in the summer 2017 arrays,theterminating
species was therecorded
branches; in
Herbarium
triloba grows of in
themesic
National Museum
to wet of Bosnia
woodlands, and
thickets, the
a disk
few newovoid, dark in
localities purple, 1-1.5 valley
the Krivaja cm wide; rays
(Careva
Herzegovina
pastures, (SARA-51805
roadsides and SARA-51806).
and meadows, generally on 8-12, yellow
Ćuprija and or orange, 1-2
Kamensko cm Olovo
near long; receptacular
as well as
wet soils. The flowering period stretches from June bracts glabrous,
Maoča and Vozuća abruptly narrowed toThe
near Zavidovići). a short
speciesbut
Results
to October. and discussion
The plant reproduces by seeds which distinct awn tip
has recently often
been shortly surpassing
registered along the banks the discof
are dispersed by the wind (Britton & Brown 1970). corollas.
the river Pappus
Željeznica a minute crown; of
in the vicinity chromosome
the village
Glasnik
Rudbeckia
The Hrvatskog
first triloba of
finding L.botaničkog
(Sp.
this Pl.: 907.društva
species 1753),
for Bosnia also number near
Krupac n=19Sarajevo
(Cronquist 1980).
(Fig. 2). ItInisitsyet 6(1)
native 2018.
unknownarea
known
and as Three-lobed
Herzegovina Coneflower,
is coming from originated
Central Bosniain the how this grows
R. triloba speciesin was
mesicintroduced
to wet woodlands,
into Bosnia thickets,
and
eastern
in 2016,part of North
on the banks America, and is ariver
of the Krivaja veryinrare,
the pastures, roadsides
Herzegovina, and as
most likely meadows,
a gardengenerally
escapee.on
locally naturalized alien in some parts of Europe. wet soils. The flowering period stretches from June
To identify this new species in the flora of Bosnia to October. The plant reproduces by seeds which
and Herzegovina, we offer here the adjusted key are dispersed by the wind (Britton & Brown 1970).
according to Britton & Brown (1970), Cronquist The first finding of this species for Bosnia
(1980), Urbatsch & Cox (2006) and Weakley (2007): and Herzegovina is coming from Central Bosnia
in 2016, on the banks of the Krivaja river in the
1. Disk elongated or cylindric in fruit, yellowish vicinity of the village Ribnica near Zavidovići (44°
or gray. Stems essentially glabrous, leaves 21’ 10.18” N; 18° 23’ 35.36” E) (Fig. 2). Individuals
thin, usually subglabrous, pinnately divided or in the population were numerous, with some
pinnatifid; plants about 100-300 cm tall, usually plants up to 150 cm high. About twenty flowering
colonial; pappus a short crown ..... R. laciniata specimens were recorded at a linear distance of
ca 100 m. In Central Bosnia the species is slowly
1. Disk globose or ovoid and purple or dark brown expanding, and it is now found in natural and
in fruit. Stems hirsute or hispid, lower leaves semi-natural habitats such as moist forests, moist
entire or lobed; plants about 50-150 cm tall meadows, in ditches and along watercourses.
........................................................................ 2 It occurs along river banks and roads and in
different human-made habitats. During fieldwork
2. Lower leaves deeply 3-lobed or 3-divided, in the summer 2017 the species was recorded in
stems branched, hirsute, 50-150 cm tall; a few new localities in the Krivaja valley (Careva
heads many, 10 to 30; pappus a minute crown Ćuprija and Kamensko near Olovo as well as
......................................................... R. triloba Maoča and Vozuća near Zavidovići). The species
has recently been registered along the banks of
2. Leaves neither 3-lobed nor 3-divided, stems the river Željeznica in the vicinity of the village
simple, hispid, 30-100 cm tall; heads 1 to 3, Krupac near Sarajevo (Fig. 2). It is yet unknown
pappus absent .................................. R. hirta how this species was introduced into Bosnia and
Herzegovina, most likely as a garden escapee.

Figure 1. Three-lobed
Three-lobedConeflower
ConeflowerRudbeckia
Rudbeckiatriloba L. L.
triloba in in
thethe vicinity
vicinity of village
of village Kamensko
Kamensko nearnear Olovo
Olovo and
and on the banks of Krivaja River near Zavidovići (Photos: Š. Šarić).
on the banks of Krivaja River near Zavidovići (Photos by Šemso Šarić).

According to Richardson et al. (2000), the10 5

observation period is too short to understand and
declare a state of naturalized species. Therefore it
can be considered an alien casual, probably not yet
as naturalized, waiting for further field investigations
to achieve the proper status attribution.
 According to Richardson et al. (2000), the
observation period is too short to understand and
declare a state of naturalized species. Therefore it
Glasnik
Figure 1. Hrvatskog botaničkog
Three-lobed Coneflower društvatriloba L.can
Rudbeckia be considered
in the an alien
vicinity of village Kamensko near 6(1)
casual, probably 2018.
not
Olovo yet
and
on the banks of Krivaja River near Zavidovići (Photosasbynaturalized,
Šemso Šarić).waiting for further field investigations
to achieve the proper status attribution.

Referencesto Richardson et al. (2000), the Acknowledgements

According
observation period is too short to understand and
• Beck-Mannagetta,
declare G., species.
a state of naturalized Maly, K., Bjelčić,
Therefore it We• Perdue,
Wewould
would R.like
like E.
to (1957):
thank
to thank toSynopsis
Aldin
to Aldin ofBoškailo
Rudbeckia
Boškailo for thefor
canŽ.be(1983): Flora
considered anBosnae et Hercegovinae
alien casual, probably not yet 4, the subgenus
mapping
mapping Rudbeckia.
of distribution Rhodora
of species
of distribution as59:
of species well293-299.
as Jessica
well as
as Sympetalae 4: 1-188,
naturalized, waiting Zemaljski
for further field muzej Bosne Jessica
investigations • Richardson,
Andersson D.forM.,
for improving
Andersson Pyšek,
the
improving English
theP.,English
ofRejmánek,
this paper.
of this
i Hercegovine
to achieve u Sarajevu,
the proper Sarajevo.
status attribution. M., Barbour, M. G., Panetta, F. D., West C.
paper.
• Britton, N. L., Brown, A. (1970): An illustrated J. (2000): Naturalization and invasion of alien
Flora of the northern United States, Canada and
Acknowledgements plants: concepts and definitions. Diversity &
the British Possessions. New York. Distributions 6: 93-107.
• Cronquist,
We would like to A. thank
(1980): Asteraceae,
to Aldin BoškailoVolume
for the • Sell, P., Murrell, G. (2006): Flora of Great
I, Vascular
mapping flora ofofthe
of distribution Southeastern
species United
as well as Jessica Britain and Ireland: Volume 4. Cambridge Figure 2. The
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Andersson University
improving ofthe
North Carolina
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of this paper. University Press, Cambridge. Bosnia and Herz
Chapel Hill, N.C. • Sirbu, C., Oprea, A. (2010): Contribution to and Maoča, 2. R
• Euro+Med (2006): Euro+Med PlantBase - the the Knowledge of the Alien Flora of Romania:
information resource for Euro-Mediterranean Rudbeckia triloba L. and Senecio inaequidens 6
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• Forstner,W., Hübl, E. (1971): Ruderal-, Figure • Slavnić,2. TheŽ. distribution
(1960): Onofimmigration,
Rudbeckia trilobaexpansionL. in
Segetal- und Adventivflora von Wien. Verlag Bosnia and and
naturalization
Herzegovina: of1.some
Careva adventive plants in
Ćuprija, Kamensko
Notring, Wien. andBosnia
Maoča,and Herzegovina.
2. Ribnica, 3. Vozuća Godišnjak Biološkog
and 4. Krupac.
• Greuter, W. (2006): Compositae (pro parte instituta Univerziteta u Sarajevu 13(1-2): 117-
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