MEDICAL

EIGHTEENTH
EDITION

MICROBIOLOGY
Commissioning Editor: Jeremy Bowes/Timothy Horne
Development Editor: Carole McMurray
Project Manager: Andrew Riley
Designer: Stewart Larking
Illustration Manager: Jennifer Rose
Illustrator: Antbits
MEDICAL
EIGHTEENTH
EDITION

MICROBIOLOGY
A GUIDE TO MICROBIAL INFECTIONS:
PATHOGENESIS, IMMUNITY, LABORATORY DIAGNOSIS AND CONTROL

EDITED BY

David Greenwood BSc PhD DSc FRCPath

Emeritus Professor of Antimicrobial Science, University of Nottingham Medical School,
Nottingham, UK

Mike Barer BSc MBBS MSc PhD FRCPath

Professor of Clinical Microbiology, Department of Infection, Immunity and Inflammation,
University of Leicester Medical School, Leicester, UK

Richard Slack MA MB BChir FRCPath FRSPH DRCOG

Formerly Senior Lecturer, Division of Microbiology, University of Nottingham Medical School,
Nottingham, UK; Honorary Consultant in Communicable Disease Control and Regional
Microbiologist, Health Protection Agency, East Midlands, UK

Will Irving MA, MB BChir, MRCP, PhD, FRCPath

Professor and Honorary Consultant in Virology, University of Nottingham and Nottingham
University Hospitals NHS Trust, Department of Microbiology, Queen’s Medical Centre,
Nottingham, UK

EDINBURGH  LONDON  NEW YORK  OXFORD  PHILADELPHIA  ST LOUIS  SYDNEY  TORONTO  2012