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• Photosynthesis
• removes CO2 from the atmosphere and
• stores it in plant matter.
• The burning of sugar in the cellular respiration of
almost all organisms releases CO2 back to the
environment.
• Photoautotrophs
• feed us,
• clothe us (think cotton),
• house us (think wood), and
• provide energy for warmth, light, transport, and
manufacturing.
• Chlorophyll
• is an important light-absorbing pigment in
chloroplasts,
• is responsible for the green color of plants, and
• plays a central role in converting solar energy to
chemical energy.
Mesophyll
Leaf
Vein
Mesophyll Cell
CO2
O2
Stoma
Chloroplast
Granum
Thylakoid
Thylakoid space
Stroma
Mesophyll
Leaf
Vein
Mesophyll Cell
CO2
O2
Stoma
Chloroplast
• Thylakoids
• are often concentrated in stacks called grana and
• have an internal compartment called the thylakoid
space, which has functions analogous to the
intermembrane space of a mitochondrion in the
generation of ATP.
Chloroplast
Thylakoid
Thylakoid space
Stroma
Mesophyll Cell
Chloroplast
Chloroplast
Stroma
Granum
Becomes reduced
Becomes oxidized
• In photosynthesis,
• light energy is captured by chlorophyll molecules to
boost the energy of electrons,
• light energy is converted to chemical energy, and
• chemical energy is stored in the chemical bonds of
sugars.
Becomes oxidized
Becomes reduced
Light
NADP +
ADP
+ P
Light
Reactions
(in thylakoids)
Chloroplast
Light
NADP +
ADP
+ P
Light
Reactions
(in thylakoids)
ATP
NADPH
Chloroplast
O2
Light
NADP +
ADP
+ P
Light Calvin
Cycle
Reactions
(in stroma)
(in thylakoids)
ATP
NADPH
Chloroplast
O2 Sugar
Visible light
• Plant pigments
• are built into the thylakoid membrane,
• absorb some wavelengths of light, and
• reflect or transmit other wavelengths.
• We see the color of the wavelengths that are
transmitted. For example, chlorophyll transmits
green wavelengths.
Light
Reflected
light
Chloroplast
Thylakoid
Absorbed
Transmitted
light
light
Light
Reflected
light
Chloroplast
Thylakoid
Absorbed
Transmitted
light
light
Excited state
Heat
Photon
of light
Photon
(fluorescence)
Ground state
Chlorophyll
molecule
Excited state
Heat
Photon
of light
Photon
(fluorescence)
Ground state
Chlorophyll
molecule
Photosystem
Light
Light-harvesting Reaction-center
complexes complex
Primary electron
STROMA acceptor
Thylakoid membrane
THYLAKOID Pigment
SPACE Transfer Pair of molecules
of energy chlorophyll a
molecules
© 2015 Pearson Education, Inc.
7.7 Photosystems capture solar energy
ATP
NADPH
Electron
transport
chain ramp
Photosystem II Photosystem I
© 2015 Pearson Education, Inc.
7.9 VISUALIZING THE CONCEPT: The light
reactions take place within the thylakoid
membranes
• A thylakoid membrane includes numerous copies
of
• the two photosystems and
• the electron transport chain.
Thylakoid
sac
Chloroplast
Light H+ Electron Light
Photosystem transport chain Photosystem NADP+ + H+
NADPH
II I
H+
H+
H+ H+
H+ H+
H2O H+
1
O2 + 2 H+ H+ H+ To
2 H+
H+ Calvin
H+ Cycle
H+ THYLAKOID SPACE H+
H+ H+
H+ H+ H+ H+
H+ STROMA
H+
H+ H+
ADP + P ATP
H+
Light
Photosystem
II
Light H+ Electron
Photosystem transport chain
II
H+
H+
H+
H+
H2O
1 H+
2 O 2 + 2 H+
H+
H+ H+
H+ H+
H2O
1 H+ H+ H+ To
O 2 + 2 H+
2 H+
H+ Calvin
Cycle
H+
H+
H+ H+
H+ H+
H2O
1 H+ H+ H+ To
O 2 + 2 H+
2 H+
H+ Calvin
Cycle
H+
H+ THYLAKOID SPACE H+
H+ H+
H+ H+ H+ H+
H+ STROMA
H+
H+ H+
ADP + P ATP
H+
NADP+
ADP
+ P
Light Calvin Input
Reactions Cycle
ATP 3
Step 1
CO2
NADPH
Carbon fixation
Chloroplast Rubisco
O2 Sugar
6 P
3 P P
3-PGA
RuBP 6 ATP
3 ADP
Step 4 P 6 ADP + P
Regeneration of RuBP CALVIN
3 ATP CYCLE
6 NADPH
6 NADP+
6 P
5 P
G3P
G3P
Step 2
Step 3 Reduction
Release of one
molecule of G3P
1 P
G3P Glucose and other
Output compounds
© 2015 Pearson Education, Inc.
Figure 7.10-1
H2O CO2
Light
NADP+
ADP
+ P
Light Calvin
Reactions Cycle
ATP
NADPH
Chloroplast
O2 Sugar
Rubisco
6 P
3 P P
3-PGA
RuBP 6 ATP
3 ADP
Step 4 6 ADP + P
Regeneration CALVIN
3 ATP CYCLE
of RuBP 6 NADPH
6 NADP+
6 P
5 P
G3P G3P
Step 2
Step 3 Reduction
Release of one
molecule of G3P
1 P
G3P Glucose and other
Output compounds
© 2015 Pearson Education, Inc.
Figure 7.10-3-1
Input
3
Step 1
CO2
Carbon fixation
Rubisco
6 P
3 P P
3-PGA
RuBP
CALVIN
CYCLE
Rubisco
6 P
3 P P
3-PGA
RuBP 6 ATP
6 ADP + P
CALVIN
CYCLE
6 NADPH
6 NADP+
6 P
G3P
Step 2
Reduction
Rubisco
6 P
3 P P
3-PGA
RuBP 6 ATP
6 ADP + P
CALVIN
CYCLE
6 NADPH
6 NADP+
6 P
5 P
G3P G3P
Step 2
Step 3 Reduction
Release of one
molecule of G3P
1 P
G3P Glucose and other
Output compounds
© 2015 Pearson Education, Inc.
Figure 7.10-3-4
Input
3
Step 1
CO2
Carbon fixation
Rubisco
6 P
3 P P
3-PGA
RuBP 6 ATP
3 ADP
Step 4 6 ADP + P
Regeneration CALVIN
3 ATP CYCLE
of RuBP 6 NADPH
6 NADP+
6 P
5 P
G3P G3P
Step 2
Step 3 Reduction
Release of one
molecule of G3P
1 P
G3P Glucose and other
Output compounds
© 2015 Pearson Education, Inc.
7.11 EVOLUTION CONNECTION: Other
methods of carbon fixation have evolved in
hot, dry climates
• Most plants use CO2 directly from the air, and
carbon fixation occurs when the enzyme rubisco
adds CO2 to RuBP.
• Such plants are called C3 plants because the first
product of carbon fixation is a three-carbon
compound, 3-PGA.
Bundle-
sheath CO2 CO2
cell
Calvin Calvin
Cycle Cycle
Sugar Sugar
C4 plant Day CAM plant
Sugarcane Pineapple
Bundle-
sheath CO2 CO2
cell
Calvin Calvin
Cycle Cycle
Sugar Sugar
C4 plant Day CAM plant
Sugarcane
Pineapple
H2O CO2
Chloroplast
Light
NADP+
ADP
+ P
Light
Reactions
RuBP
Photosystem II Calvin
Cycle 3-PGA
Electron (in stroma)
transport chain
Thylakoids
Photosystem I ATP Stroma
NADPH G3P
Cellular
respiration
Cellulose
Starch
O2 Sugars Other organic
compounds
Southern
tip of
South
America
Antarctica
September 2012
Light
energy
6 CO2 6 H2O C6H12O6 6 O2
Chloroplast CO2
H 2O
Light
NADP+ Stroma
Thylakoids ADP
+ P
Light Calvin
Reactions Cycle
ATP
NADPH
O2 Sugar
Photosynthesis
converts
includes both
(a)
(b) (c)
to
in which in which
chemical
energy
light-excited
CO2 is fixed
H2O is split electrons of
to RuBP
chlorophyll
and and then
(e)
(f) to produce
producing
Mitochondrion Chloroplast
Intermembrane H+ c.
space
Membrane
Matrix d.
a.
b. e.