Frost, D. 1985. 110-203. Amphibian Lists of The World For The Hylidae Family.

110 AMPHIBIAN SPECIES OF THE WORLD
Hemisus marmoratun (Peters, 1854). Monatsber. Preuss. Akad. Wiss. Berlin, L854:628.
oRIGINAL NAME: Engystoma marmoratum.
rvnE(s): ZMB.
TypE LocAr-rtv: Cabaceira, Mozambique.
DISTRIBUTIoN: Subsaharan Africa, excluding rainforest, Somalia, and northern and
northeastern part of Rep. South Africa.
coMMENT: Laurent, 1.972, Ann. Mus. R. Afr. Cent., Tervuren, Ser. Octavo, Sci.
Zool., 194:33-38, recognized three subspecies.
Hemisus microscaphus Laurent, L972. Ann. Mus. R. Afr. Cent., Tervuren, Ser. Octavo,
Sci. Zool.,194:38.
rvnE(s): Holotype: MCZ 19396.
TYPE LocALrrY: "lake Zwai, Ethiopia".
DISTRIBUTIoN: Known only from the type locality.
Hemisus olioaceum Laurent, 1963. Copeia, 1963:398.

oRIGINAL NAME: Hemisus oliaaceus.
rvru(s): Holotype: RGMC 40051.
TypE LocALITy: "Mongbwalu, Ituri, northeastern Congo l:Zaire)"
DISTRIBUTIoN: Northeastern Zafte.
Hemisus perreti Laurent, L972. Ann. Mus. R. Afr. Cent., Tervuren, Ser. Octavo, Sci.
Zool., L94:44.
rvnr(s): Holotype: RGMC L97a.
TypE LocALITy: "Lukula, Lower-Congo", Zaire.
DISTRIBUTIoN: Lowet Zaire, north of the estuary.
Hemisus wittei Laurent, 1963. Copeia, L963:397.

rvnE(s): Holotype: RGMC 100840.
TypE LocALITy: "Mabwe, Katanga [:Shaba], southeastern Congo l:Zairel".
DISTRIBUTIoN: Lower Shaba Province, Zaite, and northwestern Zambia.
FAMILY: Hylidae Gray,1825 (1815).

cIrArIoN: Ann. Philos., (2)10:213.
DISTRIBUTIoII:North and South America, the West Indies, and the Australo-Papuan
Region; one species group of Hyla in temperate Eurasia, including extreme northern
Africa and the |apanese Archipelago.
coMMENT: As first formed, the group name was Hylina,by implication a subfamily.
Hylidae is a conserved name in zoology (under provisions of Article 40 of the
International Code of. Zoological Nomenclature, 1985) and for purposes of priority
derives its date from Hylarinia Rafinesque, 1815, Analyse Nat.:78. Duellman, L970,
Monogr. Mus. Nat. Hist. Univ. Kansas, L:L-753, arranged the family into four
subfamilies: Amphignathodontinae, Hemiphractinae, Hylinae, and
Phyllomedusinae. Trueb, L974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 29:L-60,
demonstrated the close relationship of the amphignathodontines and
hemiphractines and suggested that the hemiphractines be included with the
amphignathodontines, an arrangement followed here. On the basis of work done by
Tyler, 197L, Univ. Kansas Publ. Mus. Nat. Hist., L9:3L9-360, and Savage, L973, ln
Vial (ed.), Evol. Biol. Anurans:351-445, Dowling and Duellman, L978, Syst.
Herpetol.,37.L, placed the Australian hylids in the subfamily Pelodryadinae. Lynch,
1971, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 53:L-238, suggested that the
Australian hylids were independently derived from the Myobatrachidae, an
hypothesis, if true, that would make the Hylidae polyphyletic. The taxonomy of this
family was summarized by Duellman, 1977, Das Tierreich, 95:1-225. The Middle
American taxa were treated in detail by Duellman, L970, Monogr. Mus. Nat. Hist.
Univ. Kansas, l:l-752.
coNrRIBUron: William E. Duellman (WED).
REVIEwERS: Werner C. A. Bokermann (WCAB) (Brazil); Ulisses Caramaschi (UC) (Brazil);
Ronald I. Crombie (RIC) (New World); Eduardo Gudynas (EG) (Uruguay and
ANURA: HYLIDAE: HEMIPHRACTINAE 111
adjacent Brazil and Argentina); Marinus S. Hoogmoed (MSH); Enrique La Marca

(ELM) (Northern South America); William F. Pyburn (WFP) (New World); Iay M.
Savage (IMS) (Central America); Norman |. Scott, Ir. (NIS) (Central and South
America); Paulo E. Vanzolini (PEV) (Brazil).
sUBFAMILv: Hemiphractinae Peters, L862.

cITATIoN: Monatsber. Preuss. Akad. Wiss. Berlin, L862:L46.
DrsrRrBUTIoN: Tropical and Andean South America; Panama.
coMMENr: The original spelling of the group name was Hemiphractidae; first use as a
subfamily was by Gadow, 190L, Amphibia and Reptiles:2l0. Duellman , L970,
Monogr. Mus. Nat. Hist. Univ. Kansas, 1:18, included eight genera. Subsequent
workers, i.e., Duellman and Trueb, L976, Occas. Pap. Mus. Nat. Hist. Univ.
Kansas, 58:12, and Maxson, 1977, Syst. Zool., 26:72, placed two of these genera,
Anotheca and Nyctimantis, in the Hylinae. Includes the Amphignathodontinae (of
Boulenger, 1882, Cat. Batr. Sal. Brit. Mus.:449); see comment under Hylidae.
Duellman and Gray, 1983, Herpetologica, 39:333-358, provided a cladogram of the
genera, as did Duellman and Hoogmoed, 1984, Misc. Publ. Mus. Nat. Hist. Univ.
Kansas,75:33.
Amphignathodon Boulenger, L882. Cat. Batr. Sal. Brit. Mus.:450.

TypE spECIrs: Amphignathodon guentheri Boulenger, L882, by monotypy.
DISTRIBUTIoN: As for the single species.
Amphignathodon guentheri Boulenger, 1882. Cat. Batr. Sal. Brit. Mus.:450.
rYPE(s): Holotype: BM L946.9.7 .28.
TypE LocALITy: "Intac", Imbabura Province, Ecuador.
DISTRIBUTIoN: Pacific slopes of Andes in Ecuador and southern Colombia.
coMMENT: This species resembles structurally Gastrotheca angustifrons, Gastrotheca
cornuta, and Gastrotheca dendronastes, from which it differs in the usual
'specific level' characters, plus the unique (for anurans) feature of having true
teeth on the dentary (WED). Probably a member of the Gastrotheca cornuta
group (DCC).
Cryptobatrachus Ruthven, L9L6. Occas. Pap. Mus. ZooL Univ. Michigan,S3:L.
TypE spECIns: Cryptobatrachus boulengeri Ruthven, 19L6.
DISTRIBUTIoN: Northern Andes and Sierra Santa Marta, Colombia.
coMMENT: For an account see Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288:
L55-L62.
Cryptobatrachus boulengeri Ruthven, L916. Occas. Pap. Mus. ZooI. Univ. Michigan,
33:2.
rvnr(s): Holotype: UMMZ 48530.
TypE LocALrrv: San Lorenzo, Departamento Magdalena, Colombia.
DISTRIBUTIoN: Sierra Santa Marta, northern Colombia.
Cryptobatrachus fuhrmanni (Peracca, l9L4). M6m. Soc. NeuchAtel. Sci. Nat., 5:108.
oRIGINAL NAME: Hyla fuhrmanni.
rvnE(s): Unknown.
TypE LocArttv: Gauca (:Heliconia), Departamento Antioquia, Colombia.
DrsrRrBUTroN: Northern parts of the Cordillera Central and Cordillera Oriental of
the Andes in Colombia.
Cryptobatrachus nicefori Cochran and Goin, L970. Bull. U.S. Natl. Mus., 288:156
wrr(s): Holotype: MLS 138.
TypE LocALrtv: La Salina, Departamento Boyacd, Colombia.
DTsTRIBUTIoN: Type locality on eastern slopes of Andes in Colombia.
Flectonotus Miranda-Ribeiro, L920. Rev. Mus. Paulista, Sio Paulo, 12:327.

TypE spECIns: Nototrema pygmaeum Boettger, 1893.
DrsrRrBUrroN: Andes and Cordillera de la Costa of northern Venezuela; Tobago and
Trinidad.
LL2 AMPHIBIAN SPECIES OF THE WORLD
CoMMENT: Genus redefined and reviewed by Duellman and Gray, L983,

Herpetologica, 39:333-358. See comment under Fritziana'
Flectonotus fitzgeraldi (Parker, 1933). Trop. Agric., Trinidad, 11:123.

oRIGINAL NAME: Gastrotheca t'itzgeraldi.
ryPE(s): Holotype: BM 1947 .2-22.41.
TYPE LocAltrv: Mt. Tucutche, Trinidad.
DISTRIBUTIOTv: Trinidad, Tobago, and Peninsula de Paria, Venezuela.
q9MMENT: The taxonomic stalus of mainland and insular populations needs to be
verified (WED).
Flectonotus pygmaeus (Boettger, 1893). Ber. Senckenb. Naturforsch. Ges., 1893:40.

oRIGINAL NAME: N ototrema PYgmaeum.
rypE(s): SMF 2679 designaied lectotype by Mertens, L967 , Senckenb. Biol., 48:43
TypE LoCALrry: Puerto Cabello, Estado Carabobo, Venezuela.
DrsrRrBUTroN: Northern Cordillera Oriental, Colombia; M6rida Andes, and
Cordillera de la Costa, Venezuela.
Fritziana Mello-Leitdo, L920. Rev. Mus. Paulista, Sio Paulo , L2:321,.

rypE spECIrs: Hyla goeldii Boulenger, 1895.
DrsrRrBUTroN: Mou"ntains of southeastern Brazil from Espirito Santo to Sio Paulo;
adjacent coastal islands.
g9MMENT: See Duellman and Gray, L983, Herpetologica,3g:333-358, who redefined
Fritziana as the stem-group of. Flectonotus.
t'issilis (Miranda-Ribeiro, 1920).

Fritziana Rev. Mus. Paulista, Sio Paulo , L2:324.
oRIGINAL NAME: Coelonotus t'issilis.
rypE(s): MZUSP 30A designated lectotype by Bokermann, 1950,PaP.Avulsos Dep
Zool., Sio Paulo , 9:2L8.
TypE LocALtrv: Serra de Maca6, Rio de |aneiro, Brazil.
DrsrRrBUTroN: Mountains of southeastern Brazil (Rio de ]aneiro, Sio Paulo), 500-
1800 m.
SSMMENT: This species was transferred from Flectonotus to Fritziana by Duellman
and Gray, L983, Herpetolo gica, 39:348, on the basis of patristic similarity.
Fritziana goeldii (Boulenger, 1895). Proc. Zool. Soc. London, L894:645.

oRIGINAL NAME: Hyla goeldii.
rvnE(s): Syntypes: BM L947 .2.L2.69 -7 0.
TypE LocArrry: Col6nia Alpina, Teres6polis, Rio de Janeiro, Btazil.
DISTRIBUTIoN: Mountains of southeastern Brazil.
Fritziana ohausi (Wandolleck,ISOT). Abh. Ber. K. Zool. Anthro. Ethno. Mus. Dresden,
lL:L4.
oRIGINAL NAME: Hyla ohausi.
rvrn(s): Holotype: MTKD D2033, destroyed.
TypE LocAlIrv: Petr6polis, Rio de )aneiro, BtaziL.
DrsrRrBUTroN: Mouniains of central Rio de ]aneiro and Sio Paulo, Brazll.
Gastrotheca Fitzinger, L843. Syst. Rept.:30.

TypE SpECIas: Hyla marsupiata Dum6ril and Bibron, L84I, by monotypy.
DISTRTBUTT9N: Panama, northern and western South America southward to northern
Argentina; eastern and southeastern Brazil.
covruritr: Various taxa were named and species grouPs defined by Duellman and
Fritts, '1.972, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 9:L-37; Duellman, L974,
Occas. Pap. Mus. Nat. Hist. Univ. Kansas,22:L-27; Duellman,L974, Occas. Pap.
Mus. Nat. Hist. Univ. Kansas, 27:L-27; and Duellman, L983, Copeia, 1983:868-
874. See Scanlon, Maxson, and Duellman, L980, Evolution, S4:222-229, f.or
discussion of albumin evolution in the genus.
Gastrotheca andaquierzsis Ruiz and HernAndez, 1976. Caldasia, 11:151.

rYPE(s): Holotype: ICN 401.
TypE LocAtirv: Cueva de los Guachoros, Valle del Rio Sauza, ca. 2000 m,
Departamento Huila, Colombia.
DrsrRrBUTroN: Cordillera Oriental of the Andes in Ecuador and southern Colombia.
coMMENT: In the Gastrotheca longipes group.
Gastrotheca angustifrors (Boulenger, 1898). Proc. Zool. Soc. London, L898:124.

oRIGINAL NAME: N ototrema angustifrons.
rypE(s): Syntypes: BM 1947 .2.22.38-39.
TypE LocALITy: "Cachab6" (:Cachabi), Provincia Esmeraldas, Ecuador.
DrsrRrBUTroN: Pacific lowlands of northwestern Ecuador and southwestern
Colombia.
coMMENT: In the Gastrotheca cornuta group.
Gastrotheca argenteoairens (Boettger, 1892). Ber. Senckenb. Naturforsch. Ges., L892:46.

oRIGINAL NAME: Hyla argenteoairens.
rypE(s): SM.F 2676 designated lectotype by Mertens, L967, Senckenb. Biol., 48:41.
TypE LocAurv: Popay6n, Departamento Cauca, Colombia.
DrsrRrBUTIotrt: Southern part of Cordillera Occidental and central and southern
part of Cordillera Central in Colombia.
coMMENT: In the Gastrotheca plumbea group; B. Lutz, L977, Bol. Mus. Nac., Rio de
Janeiro, N.S., Zool., L90:6, recognized a subspecies, Gastrotheca argenteoairens
dunni, from the northern part of the Cordillera Central.
Gastrotheca aureomaculata Cochran and Goin, L970. Bull. U.S. Natl. Mus., 288:L77.
rYPE(s): Holotype: FMNH 6970L.
TypE LocALITy: "in [Departamento] Huila, Colombia, at San Antonio, a small
village 25 kilometers west of San Agustin, at 2,300 meters".
DrsrRrBUrroN: Eastern slopes Cordillera Central in south-central Colombia.
coMMENT: In the Gastrotheca plumbea group. Includes Gastrotheca mertensi Cochran
and Goin, L970, according to Duellman, L983, Herpetologica, 39:105-110.
Gastrotheca bufona Cochran and Goin, L970. Bull. U.S. Natl. Mus., 288:L64.
rvnr(s): Holotype: MLS 344.
TypE LocAl-rry: "Ventanas (about 50 kilometers [by road] northwest of Yarumal),
[Departamento] Antioquia, Colombia".
coMMENT: In the 'Gastrotheca cornuta group. This species was incorrectly associated
with Gasf rotheca weinlandii by Duellman, L974, Occas. Pap. Mus. Nat. Hist.
Univ. Kansas, 27:5, according to Duellman, 1983, Copeia, L983:872.
Gastrotheca caaia Duellman, L974. Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 22:5.
rYPE(s): Holotype: KU 148532.
TypE LOCAIITy: Isla Pequefra, Laguna Cuicocha, 2390 m, Provincia Imbabura,
Ecuador.
DISTRIBUTIoN: Cordillera Occidental in northern Ecuador.
coMMENr: In the Gastrotheca plumbea grouP.
Gastrotheca christiani Laurent, L967. Acta ZooL Lilloana, 22:354.

rvnr(s): Holotype: FML 1359.
TypE LocAt-rry:Monumento Ruta Valle Grande, 50 km northwest of Calilegua,
Provincia Jujuy, Argentina.
DrsrRrBUrroN: Vicinity of type locality on mid-elevational slopes of Andes in
northern Argentina.
coMMENT: In the Gastrotheca marsupiata group. See Cei, 1980, Monit. Zool.ltal.,
N.S., Mon ogr., 2:439 -442.
tL4 AMPHIBIAN SPECIES OF THE WORLD
Gastrotheca chrysosticfa Laurent, L976. Acta Zool. Lilloana, 32:58.

rYPE(s): Holotype: FML 2098.
TypE LocAr-rrv: iralca de San Martin, Serrania de Porongal, Departamento Santa
Victoria, Provincia Salta, Argentina.
DrsrRrBUrroN:Vicinity of type loiality on lower Andean slopes in northern
Argentina.
coMMENT: In the Gastrotheca marsupiata gtoup.
Gastrotheca cornuta (Boulenger, 1898). Proc. ZooI. Soc. London,1898 124'

oRIGINAL NAMEr N ototrema cornutum.
ryPE(s): Holotype: BM 1947 .2.22.49.
TYPE LocALITY: "Cachab6" (:Cachabi), Provincia Esmeraldas, Ecuador'
DrsrRrBUTroN:Pacific lowlands of Ecuador amd Colombia; eastern Panama and the
Caribbean slopes of western Panama.
q6MMENT: In the Castrotheca cornuta group. Includes Gastrotheca ceratophrys
(stejneger , L}LL) according to Duellman, 1983, Copeia, L983:872.
Gastrotheca dendronasfes Duellman, 1983. Copeia, 1983:868.

rypE LocAr.lrv, Rio Calima, 1.5 km (by road) west of Lago Calima, L230 m,
Departamento de Valle, Colombia.
DrsrRrBr]Trow: Pacific slopes of Andes in southern Colombia and northern Ecuador
coMMENT: In the Gastrotheca cornuta grouP.
Gastrotheca ernestoi Miranda-Ribeiro, L920. Rev. Mus. Paulista, Sio Paulo, L2:323.
rYPE(s): Holotype: MZUSP 238.
rypE LocALmv: Macahi 1:14u"u6], Rio de )aneiro, Brazil.
CoMMENT: Probably conspecific with Gastrotheca microdisczs (WED).
Gastrotheca excubitor Duellman and Fritts, 1972. Occas. Pap. Mus. Nat. Hist. Univ.
Kansas, 9:23.
rYPE(s): Holotype: KU L39L94.
TypE LocAlrry: i'northern slope of Abra Acanacu, 31 kilometers (by road) north-
northeast of Paucartambo, Departamento Cuzco, Perti, 3370 m".
DrsrRrBUTroN: Amazonian slopes of Andes in central and southern Peru.
coMMENT: In the Gastrotheca marsupiata gtoup.
Gastrotheca fissipes (Boulenger, 1888). Ann. Mag. Nat. Hist., (6)2:42'

oRIGINAL NAME: N ototrema fissipes.
rypE(s): Holotype: BM L947 .2.22.40.
TypE LocAurv, "Iguarasse" (:IgaraEu), Pernambuco, Btazil.
DrsrRrBUTron: Coaital lowlandJ from Pernambuco to Espirito Santo, Brazil
coMMENT: In the Gastrotheca orsifera grouP.
Gastrotheca galeata Trueb and Duellman, 1,978. Copeia, L978:498.

rYPE(s): Holotype: LSUMZ 32058.
TypE LocAtrry';)'15 km (by road) east of Canchaque, Departamento Piura, Perri.,
L740 m elevation (05" 22' 5,79" 33' W)".
DISTRIBUTIoN: Vicinity of type locality in northern Peru.
coMMENT: In the Gastrotheca marsupiata group.
Gastrotheca gracilis Laurent, L969. Acta ZooL Lilloana, 25:146.

rYPE(s): Holotype: FML 1389.
TypE LocAr-rrv,La Banderita, kilometer 51, Ruta Concepci6n-Andalgal6, Provincia
de Catamarca, Argentina.
DrsrRrBUTroN: Eastern ilopes of Sierra del Aconquija and Taficillo (near Tucum6n),
northwestern Argentina.
coMMENT: In the Gastrotheca marsupiata group. See Cei, 1980, Monit. Zool.ltal.,
N.S., Mon ogr., 2:443-444.
Gastrotheca griswoldi Shreve, L94L. Proc. New England Zool. Club, 18:83.
oRIGINAL NAME: Gastrotheca boliaiana griswoldi.
rYPE(s): Holotype: MCZ 24L02.
TypE LocAttrv: Marainyoc, Departamento Junin, Peru.
DISTRIBUTIoN: Andes of central Peru.
Gastrotheca helenaeDunn, 1944. Caldasia, 2:404.

rvrr(s): Holotype: MLS 268.
TypE LocArrrv: PAramo de Tam6, Departamento Norte de Santander, Colombia
DIsTRIBUTIoN: Cerro Tam6, Colombia and Venezuela.
coMMENT: In the Gastrotheca longipes group.
Gastrotheca humbertoi Lutz, L977. Bo1. Mus. Nac., Rio de |aneiro, N.S., Zool., 290:L
rvrr(s): Formerly in MLS; now lost.
TypE LocAlrrv: El Mirador, 57 km from Mocoa on road from that town to
Sibundoy, Departamento Putumayo, Colombia.
DISTRIBUTIoN: Amazonian slopes of Andes in southern Colombia and Ecuador.
coMMENr: In the Gastrotheca longipes group.
Gastrotheca lojana Parker, 1932. Ann. Mag. Nat. Hist., (L0)9:25.

oRIGINAL NAME: Gastrotheca marsupiata lojana.
rvrr(s): Holotype: BM L947 .2.31. 13.
TypE LocArrrv: Loja, Provincia Loja, Ecuador.
DISTRIBUTIoN: Inter-Andean valleys in southern Ecuador and northern Peru
coMMENT: In the Gastrotheca plumbea group.
Gastrotheca longipes (Boulenger, 1882). Cat. Batr. Sal. Brit. Mus.:418.

oRIGINAL NAME: N ototrema longipes-
rypE(s): Syntypes: BM L947 .2.31.4-5.
TYPE LoCALITY:"Canelos" and "Sarayaclt", Provincia Pastaza, Ecuador.
DISTRIBUTIoN:Upper Amazon Basin in Ecuador and northern Peru.
coMMENr: In the Gastrotheca longipes group.
Gastrotheca marsupiafa (Dum6ril and Bibron, 1841). Erp. G6n., 8:598.

oRIGINAL NAME: Hyla marsupiata.
rvnr(s): Holotype:, MNHNP 4877.
TypE LocALrrv: "Cvzco, [Departamento Cuzco,] au P6rou [:Peru]".
DrsrRrBUTroN: Amazonian drainage systems of Andes from central Peru to
southern Bolivia.
Gastrotheca medemi Cochran and Goin, L970. Bull. U.S. Natl. Mus., 288:L72.
rvrr(s): Holotype: UIli[I[l{Z 123068.
TypE LocALITy: "Cafro GuapAyita, 400 meters, Serrania de La Macarena,
[Departamento] Meta, Colombia".
DISTRIBUTIoN: Serrania de Ia Macarena, Colombia.
coMMENr: In the Gastrotheca oaifera group. Probably conspecific with Gastrotheca
nicet'ori (WED).
Gastrotheca microdiscr.rs (Andersson, 1910). Ark. Zool., 6(9):9.

oRIGINAL NAME: N ototrema microdiscus.
rypE(s): Syntypes: NHRM L480 (2 specimens).
TypE LocALITv: Desvio Ribas, PatanA, Brazil.
DrsrRrBUTIotrl: Coastal regions and mountains of southeastern Brazil from Rio de
]aneiro to Santa Catarina.
g9MMENT: In the Gastrotheca longipes grouP. See comments undet Gastrotheca

ernestoi and Gastrotheca airidis.
Gastrotheca monticola Barbour and Noble,1920. Bull. Mus. Comp. ZooI.,63:426.

rYPE(s)r Holotype: MCZ 5290.
TypE LocAttrv: Huancabamba, Departamento Piura, Peru'
DrsrRrBUTroN: Andean slopes in Huancabamba Depression of southern Ecuador
and northern Peru.
coMMENT: In the Gastrotheca plumbea grouP.
Gastrotheca nicefori Gaige, 1933. Occas. Pap. Mus. Zool. Univ. Michigan,263:1.
rYPE(s): Holotype: UMMZ 73242.
TypE LOCAurv' Pensilvania, Departamento Caldas, Colombia.
DrsrRrBUTroN: Andean slopes of northern Venezuela and northern and central
Colombia; highlands of eastern and central Panama'
coMMENr: In the Castrotheca ouifera group. B. Lutz and Ruiz-C., L977 , Bo1. Mus.
Nac., Rio de Janeiro, N.S., Zool.,289:L2, recognized Gastrotheca nicefori
descampi from the Amazonian slopes of the Cordillera Oriental of the Andes
in souihern Colombia. See comment under Gastrotheca medemi and Gastrotheca
yacambuensis.
Gastrotheca ochoai Duellman and Fritts, L972. Occas. Pap. Mus. Nat. Hist. Univ.
Kansas,9:27.
rYPE(s): Holotype: KU L39202.
TypELocAlrry' 7'southwest base of Cordillera de Vilcanota, west of Rio
Runtumayo, about 3 kilometers north of Chilca, 10 kilometers (by road)
northwesl of ollantaytambo, Departamento Cuzco, Perit, 2760 m".
DTSTRIBUTIoN: Amazonian slopes of Andes in southern Peru.
coMMENT: In the Gastrotheca marsupiata group-
Gastrotheca orophylax Duellman and Pyles, 1980. Occas. Pap. Mus. Nat. Hist. Univ.
Kansas, 84:5.
rypE LocALrry:i'L1 km (by road) east-south-east of Papallacta,2660 m, Provincia
Napo, Ecuador".
DTSTRIBUTIOITI: Upper Amazonian slopes of Andes in Ecuador.
Gastrotheca oaifera (Lichtenstein and Weinland, 1854). Monatsber. Preuss. Akad

Wiss. Berlin, L854:373.
oRIGINAL NAME: N otodelphys oaifera.
rYPE(s): Holotype: ZMB 3073.
TypE LocALITY: Puerto Cabello, Carabobo, Venezuela.
DrSTRrBUTrOr.r: Cordillera de la Costa, northern Venezuela.
coMMENT: In the Gastrotheca oaifera grouP.
Gastrotheca peruana (Boulenger, 1900). Ann. Mag. Nat. Hist., (7)6:18L.

oRIGINAL NAME: N ototrema peruanum.
rypE(s): Syntypes: BM L947 .2.22-42-46.
TypE LocALItv: Caraz, Depattamento Ancash, Peru.
DrsrRrBUTroN: Andes and inter-Andean valleys of northern and central Peru.
Gastrotheca plumbea (Boulenger, 1882). Cat. Sal. Batr. Brit. Mus.:417.

oRIGINAL NAME: N ototrema Plumbeum.
ryPE(s): Holotype: BM L947 .2.31.19.
TypE LocALITY: "Intac", Provincia Imbabura, Ecuador.
DrsrRrBUTroN: Pacific slopes of Andes in northern and central Ecuador.
Gastrotheca psychrophilaDuellman, L974. Occas. Pap. Mus. Nat. Hist. Univ. Kansas,
22:1,5.
TypE LocALIty: "ridge between Loja and Zamora, 2850 m, l3-L4 km E (by road) of
Loja, Provincia Zamora-Chinchipe, Ecuador".
DrsrRrBUTroN: Cordillera Oriental of Andes east of Loja, southern Ecuador.
Gastrotheca riobambae (Fowler, 1913). Proc. Acad. Nat. Sci. Philadelphia, 65:L57.
oRIGINAL NAME: Hyla riobambae.
rYPE(s): Holotype: ANSP l6L6L.
TypE LocAurv: Riobamba, Provincia Chimborazo, Ecuador.
DrsrRrBUTroN: Andes and inter-Andean valleys in northern and central Ecuador
and southern Colombia.
coMMENr: In the Gastrotheca plumbea group. Biochemical and karyological evidence
shows that this taxon is a composite of two or more cryptic species (WED).
Gastrotheca splendens (Schmidt, 1857). Sitzungsber. Akad. Wiss. Wien, Math.
Naturwiss. K1., 34:11.
oRIGINAL NAME: Hyla splendens.
rvrr(s): Holotype: KM 1008/1340.
TypE LocALITy: "Chiriqui-flusse unweit Bocca del toro", Panama; in error according
to Savage and Heyer, 1969, Rev. Biol. Tropical, 16:63, who suggested that the
type may have come from Colombia, Peru, or Bolivia.
?Bolivia, ?Peru, ?Colombia.
DISTRIBUTIow:
coMMENr: Name provisionally applied to Gastrotheca; known only from poorly
preserved type specimen (Duellman, 1970, Monogr. Mus. Nat. Hist. Univ.
Kansas, 1:651).
Gastrotheca testudinea (]im6nez de la Espada, L87L). J. Sci. Math. Phys. Nat., Lisboa,
3:62.
oRIGINAL NAME: N ototrema testudineum.
rYPE(s): Holotype: MNCN 155.
TypE LocArnv: San ]os6 de Moti, Provincia Napo, Ecuador.
DrsrRrBUTIoN: Amazonian slopes of Andes in Colombia, Ecuador, and Peru.
coMMENT: In the Gastrotheca ooifera group.
Gastrotheca airidis A. Lutz and B. Lutz, L939. An. Acad. Brasil. Cienc., 11(1):81.
rYPE(s): Holotype: AL-MN 969.
TypE LocAlrry: Bonito, Serra da Bocaina, border of S5o Paulo and Rio de |aneiro,
Brazil.
DrsrRrBUTroN: Serra do Mar, Rio de ]aneiro and S5o Paulo, southeastern Brazil.
coMMENr: In the Gastrotheca longipes group. Probably conspecific with Gastrotheca
microdiscus (WED).
Gastrotheca walkeri Duellman, L980. Occas. Pap. Mus. Zool. Univ. Michigan, 690:1.
rvnr(s): Holotype: UIll{MZ I17177.
TypE LocAt-rry: "between Estaci6n Biol6gica Rancho Grande and Paso Portachuelo,
Estado Aragua, Venezuela, 1100 m".
DISTRIBUTIoITI: Central part of Cordillera de la Costa, Venezuela.
coMMENT: In the Gastrotheca longipes grouP.
Gastrotheca weinlandii (Steindachner, 1892). Sitzungsber. Akad. Wiss. Wien, Math.

Naturwiss. K1., 101:837.
oRIGINAL NAME: N ototrema weinlandii.
rYPE(s): Holotype: NHMW L6481.
TYPE LOCALITY:,,ECUAdO{,,
DrsrRrBuTror.t: Amazonian slopes of Andes in Colombia, Ecuador, and northern
Peru.
coMMENT: In the Gastrotheca oaifera grouP.
Gastrotheca williamsoni Gaige, 1922. Occas. Pap. Mus. Zool. Univ. Michigan , L07:l
TypE LocAtlrv: San Esteban, Carabobo, Venezuela.
DrsrRrBUTroN: Known only from the type locality on the north-central coastal
lowlands of Venezuela.
coMMENT: In the Gastrotheca longipes grouP.
Gastrotheca yacambuensisYrtztrs,lgT6. Rev. Univ. Centroccid. Lisandro Alvarado,

Tarea Comtn, L978:87.
rvnE(s): Holotype: MBUCV 6015.
Quebrada EI Cedral, Parque Nacional Yacambu, south slope Sierra
TypE LocAr-lrv:
de Portuguesa, Estado Lara, Venezuela, 1700 m'
DrsrRrBurrou,known only from the vicinity of type locality.
q9MMENT: In the Gastrotheca oaifera group. Probably conspecific with Gastrotheca
nicefori (WED).
Hemiphractus W ag\er, L828. Isis von Oken, 21':7 43.

rypE SPECITs: Hemiphractus spixii Wagler, L828 (:Rana scutata Spix, 1,82$, by
subsequent designation of Peters, 1862, Monatsber. Preuss. Akad. Wiss. Berlin
L862:L46.
DrsrRrBUTron: Panama; Pacific slopes of Colombia and northwestern Ecuador; uppe:
Amazon Basin and Amazonian slopes and Andes in Brazil, Ecuador, Peru, ancl
Bolivia.
q9MMENT: For review of the genus see Trueb,1974, Occas. Pap. Mus. Nat. Hist.
Univ. Kansas, 29:l-60.
Hemiphractus bubalus(Jim6nez de la Espada, L87L).J. Sci. Math. Phys. Nat., Lisboa,

3:64.
oRIGINAL NAME: Cerathyla bubalus.
rvrr(s): Holotype: MNCN 176.
TypE LocAltrv: Archidona, Provincia Pastaza, Ecuador.
DrsrRrBUTroN: Upper Amazon Basin and lower Amazonian slopes of Andes in
Ecuador and Colombia.
Hemiphractus fasciatu.s Peters, L862. Monatsber. Preuss. Akad. Wiss. Berlin, 1862:149
rYPE(s): Holotype: ZSM 3510.
7'Pastassa-Tha1, an det Ostseite der Anden in Ecuador" (in error)-
TypE LocAlrry:
DrsrRrBUTroN: Panama, northern Colombia, and Pacific slopes of Colombia and
northwestern Ecuador.
Hemiphractus johnsonl (Noble, L9I7). Bull. Am. Mus. Nat. Hist., 37:798.
oRIGINAL NAME: Cerathyla iohnsoni.
rYPE(s): Holotype: AMNH 1341.
TypE LocAlrry: Santa Rita Creek, 14 mi. north of Mesopotamia, Departamento
Antioquia, Colombia.
Northern Andean slopes of Colombia; upper Amazon Basin and
DrsTRrBUTrOr.r:
Iower Amazonian slopes of Andes from Ecuador to northern Bolivia.
Hemiphractus proboscideus (lim6nez de Ia Espada, l87L). J. Sci. Math. Phys. Nat.,

Lisboa,3:64.
oRIGINAL NAME: Cerathyla proboscidea.
rYPE(s): Holotype: MNCN 173.
TypE LocAt-ttv: Sumaco, Provincia Napo, Ecuador.
DISTRIBUTIoN: Upper Amazon Basin in Ecuador.
coMMENT: See Duellman, L978, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 55:127.
for comment on identification.
ANURA: HYLIDAE: HEMIPHRACTINAE 1.1.9
Hemiphractus scutatus (Spix, L824). Spec. Nov. Testud. Ran. Brasil.:28.

oRIGINAL NAME: Rana scutata.
rYPE(s): Formerly ZSM 37 l0l now lost; see Hoogmoed and Gruber, L983, Spixiana,
Suppl., 9:358-359, for discussion.
TypE LocAltry: Rio Solim6es, Amazonas, Brazll.
DISTRIBUTIoU: Upper Amazon Basin in Ecuador, Peru, and western Brazil.
Stefania Rivero, L968 "L966". Caribb. J. Sci., 6:L42.

TypE spECIrs: Hyla eaansi Boulenger, L904.
DISTRIBUTIoN: Highlands of the Guiana Shield in Venezuela and Guyana.
coMMENr: Duellman and Hoogmoed, L984, Misc. Publ. Mus. Nat. Hist. Univ. Kansas,
75:L-39, revised the genus, provided a cladogram of the species, and defined
two species groups on the basis of overall similarity.
Stefania eaansi (Boulenger, 1904). Proc. Zool. Soc. London, 1904:106.

oRIGINAL NAME: Hyla eaansi.
rvnr(s): Holotype: BM L947 .2.L3.LL.
TypE LocALrty: Groete Creek, Essequibo, Guyana.
DISTRIBUTIoN: Guyana and southeastern Venezuela.
coMMENr: In the Stefania eaansi group. Synonymy includes Stefania scalae
Rivero,
according to Duellman and Hoogmoed, L984, Misc. Publ. Mus. Nat. Hist.
Univ. Kansas,75:9.
Stefania ginesi Rivero, L968 "L966". Caribb. I. Sci., 6:L45.

rvnr(s): Holotype: FMNH 7404L.
TYPE LocALltY: "Chimanta-Tepui, Estado Bolivar, Venezuela, rock outcrops near E.
branch of headwater Rio Tirica, 7300 f.t."
DISTRIBUTIoN: Tepuis (sandstone mountains) in southeastern Venezuela.
coMMENr: In the Stefania ginesi group.
Stefania goizri Rivero, 1968 "L966". Caribb. J. Sci., 6:145.

rYPE(s): Holotype: AMNH 23L93.
TypE LocALIrv: Vegas Falls, Cerro Duida, Territorio Federal Amazonas, Venezuela,
4600 ft.
coMMENr: In the Stefania ginesi group.
Stefania marahuaquensls (Riverc, L96L). BuIl. Mus. Comp. Zool., L26:LL8.

oRIGINAL NAME: Hyla marahuaquensis.
rvrr(s): Holotype: "MCZ 28566.
TypE LocAlIrv: Caflo Caju, Cerro Marahuaca, Territorio Federal Amaz6nas,
Venezuela, about 1200 m.
DISTRIBUTIoN: Cerro Duida and Cerro Marahuaca, southern Venezuela.
coMMENr: In the Stefania eaansi group.
Stefania riae D:uellman and Hoogmoed, 1984. Misc. Publ. Mus. Nat. Hist. Univ.
Kansas,75:23.
rvrr(s): Holotype: KU L74688.
TypE LocALITy: "Cerro Sarisariframa, 1400 m, Estado Bolivar, Venezuela".
coMMENT: In the Stefania eaansi group.
Stefania roraima Duellman and Hoogmoed, 1984. Misc. Publ. Mus. Nat. Hist. Univ
Kansas,75:24.
rvnr(s): Holotype: Univ. Georgetown (Guyana) 10.
TypE LocALITy: "Roraima, Guyana, L4l0 m".
DISTRIBUTIow: Known only from the type locality.
coMMENT: In the Stefania eaansi group.
t20 AMPHIBIAN SPECIES OF THE WORLD
Stefania woodleyi Rivero, L968. Caribb. I. Sci., 6:147.

rvnr(s): Holotype: BM 1967.654 (formerly OUM 3056).
TypE LocAurv: Mt. Kanaima, near the Potaro River, Guyana.
coMMENT: In the Stefania eaansi grouP.
suBFAMILy: Hylinae Gray, 1825 (1815).

cIrArIoN: Ann. Philos., (2)L0:2L3.
North and South America; West Indies; temperate Eurasia, including
DrsrRrBUTroN:
extreme northern Africa and the Japanese Archipelago.
coMMENT: This subfamily probably is an unnatural group containing all those genera
that have not been split off into the other subfamilies, as noted by Duellman,
L970, Monogr. Mus. Nat. Hist. Univ. Kansas, 1:18-20.
AcrisDum6ril and Bibron, 1841. Erp. G6n., 8:506.

rypE spEcrts: Rana gryllus LeConte, L825, by h.at; see Duellman, L970, Monogr. Mus
Nat. Hist. Univ. Kansas, L:645-46, for discussion.
DrsrRrBUrrow: USA east of the Rocky Mountains; southeln Ontario (Pt. Pelee),
Canada; northern Coahuila, Mexico.
Acris crepitans Baird, L854. Proc. Acad. Nat. Sci. Philadelphia, 7:59.
rvrr(s): Not designated.
TypE LocAt-rry: "Northern states generally"; restricted to Albany, Albany County,
New York, USA, by Smith and Taylor, 1950, Univ. Kansas Sci. Bu11., 33:35 (in
error-a locality outside of the known range, according to DueIIman, L970,
Monogr. Mus. Nat. Hist. Univ. Kansas, L:647); restricted to "Potomac River at
Harper's Ferry, West Virginia", USA, by Schmidt, L953, Check List N. Am.
Amph. Rept., Ed. 6:68.
DrsrRrBUTIoN: USA east of Rocky Mountains, except extreme southeastern part;
southern Ontario (Pt. Pelee), Canada; northern Coahuila, Mexico.
Acris gryllus (LeConte, 1825). Ann. Lyc. Nat. Hist. New York, 1:282.
oRIGINAL NAME: Rana gryllus.
rvrr(s): Syntypes: USNM 3564 (7 specimens),5909 (7 specimens).
TypE LocAlIrv: Riceboro, Liberty County, Georgia, USA, according to Schmidt,
L953, Check List N. Am. Amph. Rept., Ed. 6:67.
DTsTRIBUTIow: Southeastern USA.
Allophryne Gaige, 1926. Occas. Pap. Mus. Zool. Univ. Michigan, L76:1.
rypE spECIrs: Allophryne ruthaeni Gaige, L926, by original designation.
DISTRIBUTIoIt: Guyana, Surinam, and northeastern Brazil.
coMMENr: Relationship to other frogs is problematical. Included tentatively in the
Hylidae by Lynch and Freeman, L966, Univ. Kansas Publ. Mus. Nat. Hist., 17:
493-502, and Duellman, L975, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 42:L0.
Dubois, 1983, Bull. Mens. Soc. Linn. Lyon, 52:272, considered Allophryne to be a
member of the Bufonidae, following Laurent, 1979, Bull. Soc. Zool. France, 104:
406,b:ut neither presented any evidence to support this conjecture. Noble, L931,
Biol. Amph.:51"0, considered Allophryne to be a "toothless Centrolenellt". Savage,
L973,lnYial (ed.), Evol. Biol. Anurans:354, considered this species to form a
monotypic family, the Allophrynidae. Unfortunately the name Allophrynidae
does not meet the requirements of Article 13.a (statement of diagnosis or
reference to a diagnosis for new names proposed after 1930) of the 1985
International Code of Zoological Nomenclature, and as such is not a valid
taxonomic name. Although Allophryne is here replaced in the Hylidae, this is
not because we think it belongs there. See Lynch, 1984, Herpetol. Rev., L5:46-
47, f.or a summary of the nomenclatural problem.
ANURA: HYLIDAE: HYLINAE t2L
Allophryne ruthaeni Gaige, L926. Occas. Pap. Mus. Zool. Univ. Michigan, L76:2.
TypE LocALITy: "Tukeit Hill, below Kaiteur Falls, British Guiana [:Guyana]".
DISTRIBUTIow: The Guianan region from Venezuela, through French Guiana,
Guyana, and Surinam to northeastern Brazil.
coMMENT: For discussion see Hoogmoed, L969, ZooI. Meded., Leiden, 44:75-BL
Anotheca Smith, L939. Proc. Biol. Soc. Washington,52:190.

TypE spECIrs: Gastrotheca coronata Stejneger, 1.91L (:Hyla spinosa Steindachner, L864),
by original designation.
Central America and southern Mexico.
DISTRIBUTIoN:
coMMENr: Erroneously placed in the Amphignathodontinae (included in the
Hemiphractinae of this list) by Duellman, L970, Monogr. Mus. Nat. Hist. Univ.
Kansas, 1:18; see Duellman and Trueb,l975, Occas. Pap. Mus. Nat. Hist. Univ.
Kansas, 58:L2, and Maxson, L977, Syst. 2oo1.,26:72.
Anotheca spinosa (Steindachner, 1864). Verh. Zool. Bot. Ges. Wien, L4:239.
oRIGINAL NAME: Hyla spinosa.
rvnr(s): Holotype: NHMW 16101.
TYPE LOCALITY :,,BTaziI,, .
DISTRIBUTIoN:Southeastern Mexico (Atlantic slopes of Oaxaca and Veracruz), Costa

Rica, and western Panama.
coMMENTT For discussion see Duellman, 1970, Monogr. Mus. Nat. Hist. Univ.
Kansas, 1:145-150.
Aparasphenodon Miranda-Ribeiro, 1920. Rev. Mus. Paulista, Sio Paulo, 12:87.

TypE spECIns: Aparasphenodon brunoi Miranda-Ribeiro, 1920.
DISTRIBUTIoN: Coastal region of southeastern Brazil; upper Orinoco Basin, Venezuela
coMMENT: For discussion see Carvalho,1941, Pap. Avulsos Dep. Zool., Sio Paulo, 1:
101-110, and Trueb, 1970, Univ. Kansas Pubi. Mus. Nat. Hist., 78:547-716.
Aparasphenodon brunoi Miranda-Ribeiro, L920. Rev. Mus. Paulista, S5o Paulo, L2:87.
rvnr(s): Holotype: MN 2475.
TypE LocALITy: "S. E. brasileiro"; stated as Rio de ]aneiro , Brazil, by Bokermann,
L966, Lista Anot. Local. Tipo Anf. Brasil.:13.
DISTRIBUTIoN: Coastal lowlands of southeastern Brazil from Espirito Santo to Sio
Paulo.
coMMENT: See Sa4ima and Cardoso, 1980, Iheringia, Zool., 55:3-7.
Aparasphenodon aenezolanus (Mertens, 1950). Senckenb. Bio1., 3L:1.

oRTGTNAL NAME: Corythomantis aenezolana.
rvrr(s): Holotype: SMF 22168.
TypE LocALrrv: San Fernando de Atabapo, Territorio Federal Amazonas, Venezuela
DISTRIBUTIoN: Southwestern Amazonian Venezuela.
coMMENT: For generic assignment see Trueb, 1970, Univ. Kansas Publ. Mus. Nat.
Hist., 18:599. For recent discussion see Paolillo and Cerda, 1983 "198L", Mem.
Soc. Cienc. Nat. La Salle, 4L(Ll5):77-95.
Aplastodiscus A. Ltttz, 1950. ln B. Lutz, Mem. Inst. Oswaldo Cruz, Rio de Janeiro, 46:
612.
TypE spECIns: Aplastodiscus perairidis A. Lutz, 1950.
coMMENT: Discussed by Cei, 1980, Monit. ZooLItal., N.S., Monogr., 2:450-452.
Caramaschi, 1983, Resumos X Congreso Brasil. 2ool.:306-307, presented
summary information on Aplastodiscus as a synonym of Hyla. Resolution of this
must await publication of the complete paper.
r22 AMPHIBIAN SPECIES OF THE WORLD
Aplastodiscus perairidis A. Lut2,1950. In B. Lutz, Mem. Inst. Oswaldo Cruz, Rio de

]aneiro, 46:6L2.
rvnE(s): Holotype: AL-MN 2L32.
TypE LocALlry: Fazenda do Bonito, Serra de Bocaina, Sio ]os6 do Barreiro, Sio
Paulo, Brazil.
DrsrRrBUTroN: Central and southeastern Brazil from Brasilia and Serra da
Mantiqueira and coastal ranges south to Rio Grande do Sul and northeastern
Argentina.
Ar g ent e ohy I a T rueb, L97 0 . Herpeto 1o gica, 26 :255 .
rypE spECIas: Hyla siemersi Mertens, 7937, by original designation.
DrsrRrBUrrow: Drainages and mouth of the Rio ParanA and the Delta La Plata in
Argentina and southern coast of Uruguay; south-central Paraguay; questionably
Amazon Basin in Peru.
coMMENT: For discussion see Trueb, L970, Herpetologica 26:254-267, and Cei, L980,
Monit. Zool.Ital., N.S., Monogr., 2:452-453. See comment under Argenteohyla
altamazonica.
Argenteohyla altamazonica Henle, 1981. Amphibia-Reptilia, 2:134.

rYPE(s): Holotype: ZFMK 29993.
Pucallpa, ca.200 m NN".
TypE LocALITy: "Peru;
coMMENr: Probably a junior synonym of Phrynohyas aenulosa (WED).
Argenteohyla siemersi (Mertens, 1937). Senckenb. Biol., 19:12.

oRIGINAL NAME: Hyla siemersi.
rYPE(s): Holotype: SIll{F 22249.
TypE LocAltrv: Rio La Plata, Provincia Buenos Aires, Argentina.
DrsrRrBUTroN: Drainages and mouth of the Rio Paran6 (Corrientes, Entre Rios, and
Buenos Aires provinces) in Argentina and southern coast of Uruguay; south-
central Paraguay.
Calyptahyla Trueb and Tyler, L974. Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 24:41.
rypE spEcrus: Trachycephalus lichenatus Gosse, L851 (:Hyla uucialis Harlan, L826).
DISTRIBUTIoN: Jamaica, West Indies.
REVIEWERS: Albert Schwartz (AS); Richard Thomas (RT).
Calyptahyla crucialis (Harlan, L826). Am. J. Sci. Arts, L0:64.

oRIGINAL NAME: Hyla crucialis.
rvnr(s): Holotype: ANSP 2L80; see Crombie, 1973, Bull. Zool. Nomencl., 30:4-6.
TYPE LoCALITY: "Jamaica".
Corythomanfis Boulenger, L896. Ann. Mag. Nat. Hist., (5)10:405.
TYPE sPECIns: Corythomantis greeningi Boulenger, 1896.
DTsTRIBUTIoN: Northeastern Brazil.
Corythomantis greenirzgi Boulenger,7896. Ann. Mag. Nat. Hist., (5)10:405.
rvrr(s): Holotype: BM L947 .2.25.97 .
TypE LocAlrry: "Brazil (exact locality unknown)"; catalogue in BM has locality

recorded as "Espirito Santo, BrazTl" (WED).
DrsrRrBUTroN: Xeric and subhumid regions of northeastern Brazil.
69MMENT: For discussion see Carvalho, 194I, Pap. Avulsos Dep. Zool., Sio Paulo,
1:101-110, and Trueb, 1970, Univ. Kansas Publ. Mus. Nat. Hist., L8:546-7L6.
See Sazima and Cardoso, 1980, Iheringia, Zool., 55:3-7.
Hyla Laurenti, 1768. Synops. Rept.:32.

TypE spEcrrs: Hyla airidis Laurenti, 7768 1-11ona arborea Linnaeus, 7758), by
subsequent designation of Stejneger, L907, Bull. U.S. Natl. Mus., 58:75.
DrsrRrBUTroN: Central and southern Europe; eastern Asia; northwestern Africa;
North, Central, and South America; Greater Antiiles in West Indies.
ANURA: HYLIDAE: HYLINAE 123
coMMENr: Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas, l:173-5L8,
reviewed the Mexican and Central American species and discussed species
grouPs. B. Lutz, L973, BraziL. Spec. Hyla, discussed the Brazilian species.
Duellman, L972, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 11:1-3i, reviewed
the Hyla bogotensis group. Duellman, L973, Copeia, L973:5L5-533, reviewed the
Hyla geographica group. Duellman, L974, Occas. Pap. Mus. Nat. Hist.,23:1-40,
reviewed t};.e Hyla paroiceps group. Duellman and Trueb, 1983, }z Rhodin and
Miyata (eds.), Adv. Herpetol. Evol. Biol.:33-51, reviewed the Hyla columbiana
group/ and discussed the phylogenetic relationships of groups of Hyla
characterizedby 30 chromosomes and reduced larval mouthparts (Hyla labialis,
Hyla columbiana, Hyla minuta, Hyla marmorata, Hyla paroiceps, Hyla leucophyllata,
and Hyla microcephala groups). Pyburn and Hall, L984, Herpetologica, 40:366-
372, supplied a key to the species of the Hyla geographica group.
REVIEWERS: Jonathan A. Campbell (]AC) (Mexico and Central America); Gustavo
Casas-Andreu (GCA) (Mexico); Oscar Flores-Villela (OFV) (Mexico).
Hyla acreana Bokermann, L964. Senckenb. Biol., 45:244.

rvnE(s): Holotype: WCAB 1363.
TypE LocALITy: "Tarauac6, Acre, Brazil".
DISTRIBUTIoN: Western Brazil (Acre and Amazonas), northeastern Bolivia, and
southeastern Peru.
coMMENT: In the Hyla marmorata group; see Heyer, L977, Pap. Avulsos ZooI., Sio
Paulo, 3L:L42. See comment under Hyla senicula.
Hyla albicans Bokermann, L967. Neotropica, L3:64.

rvnr(s): Holotype: WCAB L95L9.
TypE LocALrry: Teres6polis, Rio de ]aneiro, Brazil.
Hyla albofrenata A. Lutz, L924. C. R. S6ances Soc. Biol. Paris,90:24L.

rvrr(s): Syntypes: AL-MN 577 (2 specimens, Tijuca), USNM 96478 (Angra dos
Reis), 963L9 (Sumare), 96339-45 (Paineiras).
TypE LocALITy: "Environs of Rio de Janeiro", Btazll.
DISTRIBUTIoN: Mountain slopes of southeastern Brazll (Minas Gerais, Espirito
Santo, Rio de Janeiro, 56o Paulo).
coMMENT: In the Hyla albomarginata group, according to Duellman, L970, Monogr
Mus. Nat. Hist. Univ. Kansas, 7:240.
Hyla alboguttata Bodlenger, 1882. Cat. Batr. Sal. Brit. Mus.:356.

rvnr(s): Syntypes: BM 19 47 .2.L3.47 -48 (Sarayacu), L9 47 .2.3.49 (Canelos).
TYPE LocALITy: "Sarayacr;-" and "Canelos", Provincia Pastaza, Ecuador.
DISTRIBUTIoN: Amazonian Ecuador.
Hyla albolineata A. Lutz and B. Ltttz, 1939. An. Acad. Brasil. Cienc., 7L:69.
rvrr(s): Holotype: AL-MN 1778.
TypE LocAtIty: Teres6polis, Rio de janeiro, Brazil.
coMMENT: In the Hyla albomarginata group, according to Duellman, L970, Monogr
Hyla albomarginata Spix, 1824. Spec. Nov. Testud. Ran. Brasil.:33.

rvnr(s): Holotype: formerly ZSM 237010; destroyed.
TYPELocAlrry: Bahia, Brazil.
DISTRIBUTIoN:Caribbean lowlands of Colombia to Guianas, lower Amazon Basin,
and Atlantic forests of eastern Brazll from Pernambuco to Santa Catarina.
coMMENr: In the Hyla albomarginata group. It is highly likely that the populations
along the Caribbean lowlands are specifically distinct (WED).
fe
Hyla albonigra Nieden, L923. Das Tierreich,46:278'

rvnr(s): Syntyp"t, MNHNP 4817 (2 specimens)'
TypE LocALITY: "Buenos-Ayr6s", Argentina'
DrSrRrBUrroN: ? Bolivia (actording t6 Ni"d"t , L923, Das Tierreich,
46:278)'
,upiu."*ent name fot Hyla zebra Dum6ril and Bibron'
aortr*r, Hyla albonigra is u
L84I, ErP. G6n., 8:575.
Hyla albopunctata spix, L824. Spec.-Nov. Testud. Ran. Brasil.:33.

rypE(s): Formerly in ZSM, n'ow losg KU 100000 designated neotyPe by
Duellman'
!97 L, HerPetolo grca, 27 :402.
TypE LoCArmy: Unkr-o*rr; neotype from BeIo Horizonte, Minas
Gerais' Brazil'
(Provincia
DrsrRrBUTroN: Central and soutireaste rn Brazll; northeastern Argentina
Corrientes); eastern ParaguaY.
coMMENr: In the Hyla albopuictaia grouP. See Cei, L980, Monit' Zool'
Ital'' N'S''
Monogr., 2:462-463.
Hyla albopunctulata Boulenger,lSS?. Cat. Batr. Sal. Brit. Mus.:385.

rvrr(s): Syntypes: including BM 80.12'5'159-L62'
TypE LOCALITY: "Sar ayacu","Provincia Pastaza, Ecuador
(in error ?) and "Ecuador"'
DISTRIBUTIoN: Known only from the type locality'
coMMENr: Questionably in the Hyla bogotensis SrouP'
Hyla albosignata A. Lutz and B. Lut2,1938. An. Acad. Brasil' Cienc', 10:185'
rvnr(s): HolotYPe: AL-MN 722.
TypE LocArrrr, da Serra, Paranapiacaba, Sdo Paulo , Btazil.
ilto
DrsrRrBUTroN: Serra do Mar and Serri da Mantiqueira in southeastern
Brazil.
q9MMENT: In the Hyla albomarginata group, according to Duellman, L970, Monogr'
Mus. Nat. Hist. Univ. Kansas, L:240'
Hyla alborsittata Lichtenstein and Mattens, 1856. Nomencl. Rept. Amph. Mus. zool'
Berolin.:37.
rYPE(s): Not traced (in ZMB if still extant)'
TYPE LocAltrY: Brazil.
DISTRIBUTIoN: Unknown.
Hyla alemani Rivero , L964- Caribb. J. Sci., 4:302'

rYPE(s): HolotyPe: SCN 238.
TypE LocArtrv, Cag..a, Estado Aragua, Venezuela'
coMMENr: Closely allied io Hyla granoia, according to the_original description,-or
a
possible ,yr,6rry*, u..ordi.,{ to Rivero, Lg67 , Mem. Soc. Cienc. Nat. La Salle,
27:7-8.
Hyla altipotensDuellman, Lg68. Univ. Kansas Publ. Mus. Nat. Hist', L7:572'
rYPE(s)r HolotYPe: KU 101001.
(kilometer
rypE LocArtry, "37 kilometers (by road) north of san Gabriel Mixtepec
post L83 on road from Oaxata to Puerto Escondido), Oaxaca, Mexico'
elevation 1860 meters".
DISTRIBUTION: Pacific slopes of Sierra Madre del Sur in Oaxaca, Mexico'
CoMMENT: In the Hyla tieniopus 1rotp. See Duellman, L970, Monogr'
Mus' Nat'
Hist. Univ. Kansas, 1:450-453.
Hyla alaarengai Bokermann, L956. Pap. Avulsos Dep. Zool-, Sio Paulo,12:357'
rYPE(s): HolotyPe: MZUSP 1680.
TypE LOCALTTy: "santa Barbara, pr6ximo a Belo Horizonte, Estado de
Minas Gerais,
BrasiI".
DISTRIBUTIou: Serra do Espinhago in Bahia and Minas Gerais, Brazil'
Hyla alytolylaxDuellman, 1972. Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 11:15
rvnr(s): Holotype: KU 111903.
TypE LocALITy: "Tandapi, Provincia Pichincha, Ecuador, L460 m".
DISTRIBUTIow: Pacific slopes of Andes in southern Colombia and Ecuador.
coMMENT: In the Hyla bogotensis group.
Hyla americana (Dum6ril and Bibron, 1841). Erp. G6n., 8:506.

oRTGTNAL NAME: Litoria americana.
rvnr(s): Holotype: MNHNP 756.
TypE LocALITy: "Nouvelle-Or16dns" (:11ew Orleans), Orleans Parish, Louisiana,
USA.
DISTRIBUTIow: Unknown.
coMMENT: This name has never been associated with a population of anurans
(wED).
Hyla anataliasiasi Bokermann, L973. Rev. Brasil. Biol., 34:593.

rvnr(s): Holotype: WCAB 45272.
rypE LocAury: 80 km north of Paraiso do Norte, Municipio de Brejinho do
Nazar6, GoiAs, Brazil.
DISTRIBUTIoN: Central Brazll (upper Rios Araguaia, Tocantins, and Xingu).
Hyla anceps A-Lutz, L929. C. R. S6ances Soc. Biol. Paris, L0l:943.

rvnr(s): Syntypes: AL-MN L776-77, USNM 9644L-42.
TypE LocALITy: Estrela, Mag6, Rio de Janeiro, Brazll.
DISTRIBUTIoN: Lowlands of Rio de ]aneiro in southeastern Brazil.
coMMENT: For discussion see Cochran, 1955, Bull. U.S. Natl. Mus.,206:188
Hyla andersonii Baird, 1854. Proc. Acad. Nat. Sci. Philadelphia,T:60.

rvns(s): Holotype: USNM 3600.
TypE LocALITy: "Anderson, S[outh]. C[arolina].", USA, in error; designated as
"Aiken County, South Carolina", USA, by Schmidt, 1953, Check List N. Am.
Amph. Rept., Ed. 6:69; see Gosner and Black, L967, Cat. Am. Amph. Rept.,
54.1, who regarded the type locality as unknown.
DISTRIBUTIoN: Atlantic Coastal Plain from central New Jersey to northern Florida,
USA, in scattered populations.
coMMENr: Reviewed by Gosner and Black, L967, Cat. Am. Amph. Rept., 54.L-2.
PRoTECTED srArus: USA EsA-Endangered (11 Nov. L977); Florida population only.
Hyla angustilineata Taylor, 1952. Univ. Kansas Sci. BuIl., 35:850.

rvnr(s): Holotype: USNM 75060.
TypE LocALITy: "La Palma, [Provincia San Jos6,] Costa Rica".
DrsrRrBUrrow: Cordillera of Costa Rica.
coMMENT: In the Hyla pseudopuma group; see Duellman, L970, Monogr. Mus. Nat
Hist. Univ. Kansas, 1:273.
Hyla anisifzl (M6hel:i, L904). Ann. Hist. Nat. Mus. Natl. Hungarici,2:229
oRIGINAL NAME: Hylella anisitzi.
rvrE(s): MNH.
TypE LocAurv: ? Asuncion or Villa Sana, Paraguay.
DTsTRIBUTIoN: Known only from the type locality.
Hyla annectans (Jerdon, 1870). Proc. Asiat. Soc. BengaI,1870:84.

oRIGINAL NAME: P olypedates annectans.
rvnr(s): Holotype: ZMB 3500.
TypE LocAlrrv: Khasi Hills, Assam, India.
DrsrRrBUrIoN: Southern Asia (south of Himalayas) from Assam (India) through
northern Burma, Thailand, and Vietnam to western China.
coMMENr: In the Hyla arborea group; reviewed by Bourret, L942, Batr. Indochine
222-223; Pope, L93L, Bull. Am. Mus. Nat. Hist.,5l:474-477;Liu,1950,

Fieldiana: Zool. Mem., 2:224-23L; and Liu and }{u, 196L, Tailless Amph.
China:130-1323.
REVIEwERS:Leo |. Borkin (LIB); ]. C. Daniel (JCD); Sushil Dutta (SD); Shuqin Hu
(SH); Masafumi Matsui (MM); Ermi Zhao (EZ).
Hyla aperomea Duellman, 1982. Amphibia-Reptilia, 3:153.

TypE LocAr-mv' 8 km NNE of. Balzapata, 1850 m, Departamento Amazonas, Peru
DrsrRrBUTroN: Amazonian slopes of Andes in northern and central Peru.
coMMENT: In the Hyla minima grouP.
Hyla arborea (Linnaeus, L758). Syst. Nat., Ed. 10, l:213.

oRIGINAL NAME: Rana arborea.
rYPE(s): Unknown (see comment).
TypE LocAtITY: "Sub foliis arborum Europae, Americae".
DrsrRIBUTroN: Central and southern Europe east to Caucasus, northwestern coast of
Africa, and Turkey.
coMMENr: Two of Linnaeus' type specimens (NHRM 155) are syntypes of Hyla
punctata (Schneider). Andersson, 1900, Bih. K. Svenska Vetensk. Akad.
Handl., 26:17 , noted a third specimen in the same jar which "seems to me to
be Hyla inframaculata". He noted further "Al1 three specimens thus are from
America, and Linnaeus says also that his type come from this part of the
world". The third specimen was not found by Duellman when he was in the
NHRM in L969. Thus, it seems no European specimen is now listed as a type
specimen (Duellman , 1977, Das Tierreich, 95:31). In the Hyla arborea rtolrP.
Four subspecies (arborea, kretensis, molleri, and schelkownikowi) presently
recognized; other subspecies listed by Duellman, L977, Das Tierreich, 95:31.-
33, have been elevated to specific status.
REVIEwERS: Steven C. Anderson (SCA); Leo ]. Borkin (L]B); Kim M. Howell (KMH);
Shuqin Hu (SH); Masafumi Matsui (MM); Ermi Zhao (EZ).
Hyla arborescandens Taylor, 1939. Univ. Kansas Sci. Bull., 25:388.

rvnr(s): Holotype: UIMNH 25045 (formerly EHT 3135).
TypE LocAlrry: "on a mountain side about 3 km. southwest of Acultzingo,
Veracruz", Mexico.
DrsrRrBUTroN: Sierra Madre Oriental in eastern Mexico (Puebla, Oaxaca, and
Veracruz).
C9MMENT: In the Hyla miotympanum SrouP; see Duellman, L970, Monogr. Mus.
Nat. Hist. Univ. Kansas, 1:380-384.
Hyla arenicolor Cope, 1886. ]. Acad. Nat. Sci. Philadelphia, (2)6:84.

rypE(s): Syntypes: USNM 11410 (five specimens); USNM 11410a designated
lectotype by Gorman, 1960, Herpetologica, 16:218.
TypE LOCALITY: "Northern Sonora", Mexico, or what is now southern Atizona,
USA; restricted to Santa Rita Mountains, Arizona, USA, by Smith and Taylor,
1950, Univ. Kansas Sci. Bu11.,33:354; restricted to Pefia Blanca Springs, 10
miles northwest of Nogales, Santa Ctuz County, Arizona, USA, by Gorman,
L9 60, Herpetolo gica, L6:2L8.
DrsrRrBUTroN: Mountains and plateau areas of USA (southern Utah and western
Colorado southward through eastern Arizona, western New Mexico, and
western Texas) southward in Mexico to MichoacAn, Guerrero, and Oaxaca.
coMMENr: Hyla arenicolor is a replacement name f.or Hyla affinis Baird, L854, Proc.
Acad. Nat. Sci. Phitadelphia,T:6L, which is preoccupied by Hyla affinis Spix,
L824.In the Hyla aersicolor group; for discussion see Duellman, L970, Monogr
Mus. Nat. Hist. Univ. Kansas, L:5L4.
Hyla ariadne Bokermann, L967. Neotropica, 13:61.

rYPE(s): Holotype: WCAB 29492.
TypE LocAlrry: Campo de Fruticultura da Serra da Bocaina, Sio Jos6 do Barreiro,
Sio Paulo, BrazTl.
DISTRIBUTIoN: Serra da Bocaina, Sio Paulo, southeastern Brazil.
Hyla armafa Boulenger, 1902. Ann. Mug. Nat. Hist., (7)L0:394.

rvrr(s): Syntypes: BM 7947 .2. 13.59-60.
TypE LocALrrv: "La Paz, Boiivia, 4000 m."
DISTRIBUTIow: Eastern slopes of Andes in Peru and Bolivia.
Hyla astarfea Bokermann, L967. Rev. Brasil. Biol., 27:157.

TYPE LocAtIrv: Paranapiacaba, Sio Paulo, Btazil.
DISTRIBUTIoN: Serra do Mar, southeastern Brazil.
coMMENT: In the Hyla circumdata group according to the original description.
Hyla auraria Peters, L873. Monatsber. Preuss. Akad. Wiss. Berlin, L873:6L5.
rYPE(s): Formerly in ZSM; now lost.
TYPE LocALITY: "South America".
DISTRIBUTIow: Unknown.
coMMENT: This name has not been associated with a population of anurans
(wED).
Hyla auiaoca Yiosca, L928. Proc. Biol. Soc. Washington, 41:89.

rYPE(s): Holotype: USNM 75017.
TypE LocAt-rry: "outskirts of Mandeville, [St.
Tammany Parish,] Louisiana", LJSA.
DrsrRrBUTroN: Southeastern USA (Mississippi River drainage from southern Illinois
southward; eastern Georgia and adjacent South Carolina); isolated records in
southeastern Oklahoma and central Arkansas.
coMMENT: In the Hyla aersicolor group; for discussion see Smith, L966, Cat. Am.
Rept. Amph., 28.L-2.
Hyla baileyi Cochran, 1953. Herpetologica, 8:LL4.

oRIGINAL NAME: Hyla goughi baileyi.
rYPE(s): Holotype: UM.}I4.Z L06737.
TypE LocArmy: Fazenda PoEo Grande, 8 km north of juquia, Sio Paulo, Brazil.
DISTRIBUTIoN: Atlantic forests of Sio Paulo, Brazil.
Hyla balzani Boulenger, 1898. Ann. Mus. Civ. Stor. Nat. Genova, (2)L9:L32.
rYPE(s): Holotype: MSNG 28872.
TypE LocALITy: "Prov. Yungas, at 1600 metres altitude", Bolivia.
DISTRIBUTIoN: Amazonian slopes of Bolivia and Peru.
Hyla battersbyi Rivero, 195L. Bull. Mus. Comp. Zool., 126:L38.

rvrr(s): Holotype: BM 53.2.4.L65.
TYPE LocAtttv: Caracas, Venezuela.
Hyla benitezi Rivero, L961. Bull. Mus. Comp. Zool., 126:116.
rYPE(s): Holotype: MCZ 28564.
TypE LocAtIry: Cano Wanadi, Cerro Marahuaca, Territorio Amazonas, Venezuela.
DrsrRrBUTrow: Western part of Guyana Highlands in Amazonas, Venezuela.
coMMENr: See Rivero, L97L, Caribb. J. Sci., 11:181-193.
Hyla berthalutzae Bokermann, 1962. Neotropica, 8:84.

TypE LocALrrv "Paranapiacaba, Estado de Sio Paulo, Brasil".
DrsrRrBuTror.r: Coastal lowlands from Espirito Santo to 56o Paulo, and Serra do
Mar, Sio Paulo, Brazil.
CoMMENT: Related to Hyla misera and Hyla fiAna, according to the original
description.
Hyla bifurca Andersson, 1954. Ark. Zool.,37A(2):79'

oRIGINAL NAME: Hyla (Hylella) bifurca.
rvnr(s): Holotype: NHRM L962.
TypE LocArrrv, Rio Pastaza, Provincia Pastaza, Ecuador'
DISTRIBUTIoT.T: Upper Amazon Basin in Colombia, Ecuador, Peru, and
Acre, Btazil
g9MMENT: In the'Hyla leucophyllata group; see Duellman, 1974, Occas. Pap. Mus.
Nat. Hist. Univ. Kansas, 27:L5.
Hylabiobeba Bokermann and sazima, L974. Rev. Brasil. Biol., 33:329.

TypE LOCAurvl Usina, Serra do Cipo, ]aboticatubas, Minas Gerais, Btazil'
DTSTRTBUTTON: serra do EspinhaEo in Minas Gerais, Brazil.
C9MMENT: In the Hyla mirtinsi-Hyla langei grouP, according to the original
description. UC regards this statement as in error and considers this species
to be closel/related to or a iunior synonym of Hyla pardalis'
"ith",
Hyla bipunctata Spix, L824. Spec. Nov. Testud. Ran. Brasil':36'
rvpr(s), Syntypes: ZS}y'^ 2497 /0 (4 specimens) (destroyed); Ho-ogmoed and Gruber,
Lgp3; Spiiiana, Suppl., 9:368, noted that Spix based his description on two
syntypes onlY.
TYPE LoCALITY: "Bahta", BtazTl.
DrsrRrBUTroN: Coastal region of eastern Brazil from Bahia to Rio de Janeiro
Hyla bischofi Boulenger, L887. Ann. Mag' Nat. Hist., (5)20:298'

rypE(s): Syntypes: BM L947 .2.L2.81-82.
rypE'Loc;Lrry; "Mundo Novo, [Taquara,] Rio Grande do Sul", Brazil.
DrsrRrBuTror.r: Southeastern Brazil from Rio de )aneiro to Rio Grande do Sul
Hyla bistincta Cope, L877. Proc. Am. Philos. Soc., L7:87 '
rYPE(s): Holotype: USNM 3226L.
TypE LocALITY: "Vera Crtz most probably", Mexico.
DrsrRrBUTroN:Mountains of Mexico from Durango and Veracruz to Oaxaca.
g9MMENT: In the Hyla bistincta group; for discussion see Duellman, 1970, Monogr
Mus. Nat. Hist. Univ. Kansas, L:457-462.
Hyla boans (Linnaeus , L758). Syst. Nat., Ed. 10, L:213'

oRIGINAL NAME: Rana boans.
rypE(s): Holotype: ZIUIJ 27; designation of RMNH 16603 as neotype by Duellgu.,
L1TL, Heipetoto gica, 27:399, is invalid. See comment under Hyla leucophyllata.
TYPE LoCALITY: "America".
Amazon Basin, upper Orinoco Basin, Guianas, and Pacific lowlands
DrsrRrBUTroN:
of Colombia in south America; eastern Panama; Trinidad.
CoMMENT: In the Hyla boans grouP; for discussion see Duellman, L97L,
Herpetolo gica, 27 :398, and Mertens, L972, Senckenb. Biol., 53:L97 -L98.
Hylabogertae Straughan and Wright, L969. Contrib. Sci. Nat. Hist. Mus. Los Angeles
Co., L69:L.
rYPE(s): Holotype: LACM 44400.
TypE LocAlrry: tributary of Rio Atoyac, below Vivero El Tapanal, L-6 km south of
La Cofradia, Distrito Sola de Vega, Oaxaca, Mexico'
DISTRIBUTIoT: Known only from the type locality'
coMMENr: In the Hyla bisiincta group; for discussion see Duellman , L970, Monogr.
Mus. Nat. Hist. Univ. Kansas, 1:479-482.
ANURA: HYLIDAE: HYLINAE r29
Hyla bogotensis (Peters, 1882). Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1882:
107.
oRIGINAL NAME: Hylonomus bogotensis.
rYPE(s): Holotype: ZMB L0209.
TypE LocAltrv: BogotA, Departamento Cundinamarca, Colombia.
DISTRIBUTIoN: Central and southern part of Cordillera Oriental of Andes and
? southern part of Cordillera Central in Colombia.
coMMENr: In the Hyla bogotensis group; for discussion see Duellman, L972, Occas.
Pap. Mus. Nat. Hist. Univ. Kansas, LL:1-3L, and Ruiz-C. and Lynch, L982,
Caldasia, L3:647 , who considered Hyloscirtus aermiculatus B. Lutz and Ruiz-C.,
1977, to be a synonym.
Hyla bokermanni Goin, L960. Ann. Mag. Nat. Hist. , (L3)2:72L.

rvrr(s): Holotype: WCAB 2881.
TypE LocAtITv: Tarauaca, Acre, Brazil.
DrsrRrBUTroN: Upper Amazon Basin in Colombia, Ecuador, Peru, and western
Brazil.
coMMENr: In the Hyla paraiceps group; see Duellman and Crump, L974, Occas. Pap
Mus. Nat. Hist. Univ. Kansas, 23 14.
Hylabreaifrons Duellman and Crump, 1974. Occas. Pap. Mus. Nat. Hist. Univ.
Kansas, 23:15.
rvnE(s): Holotype: KU 126370.
TypE LocALITy: "Santa Cecilia, Provincia Napo, Ecuador".
DISTRIBUTIoN: Amazon Basin and French Guiana.
coMMENT: In the Hyla paraiceps group.
Hylabromeliacia Schmidt, 1933. Field Mus. Nat. Hist. Publ., Zool.Ser.,20:L9.

rvnr(s): Holotype: FMNH 4718.
TypE LocALITy: "Mountains west of San Pedro Sula", Cort6s, Honduras.
DISTRIBUTIoN: Atlantic slopes of Guatemala and Honduras.
coMMENT:In the Hyla bromeliacia group; for discussion see Duellman, 1970,
Monogr. Mus. Nat. Hist. Univ. Kansas, L:429-434.
Hyla cadaoerina Cope, L866. Proc. Acad. Nat. Sci. Philadelphia, 18:84.
rvnr(s): Syntypes: ANSP L987-88.
TypE LocAtIty: "Tejon Pass", Los Angeles County, California, USA.
DrsrRrBUTroN: Southern California, USA, and northern Baja California, Mexico.
coMMENT: Hyla cadaaerina is a replacement name f.or Hyla nebulosa Hallowell, L854,
Proc. Acad. Nat. Sci. Philadelphia,T:96.In the Hyla eximia group; f.or
discussion of this species see Gaudin, L979, Cat. Am. Amph. Rept., 225.L-2.
Hyla calcarataTroschel, 1848. Amph. In Schomburgk, Reisen Brit. Guiana:650.

rvnr(s): Unknown.
TypE LocALITY: "Britisch-Guiana" 1:6rryana).
DISTRTBUTIoII: Guianas, Amazon Basin, and upper Orinoco Basin in Venezuela and
Brazil.
coMMENr: In the Hyla geographica group; for discussion see Duellman, 1973,
Copeia, L973:522-523.
Hyla callipleura Boulenger, L902. Ann. Mag. Nat. Hist., (7)10:394.

rypE(s): Syntypes: BM L947.2.L3.64-73 (Charuplaya), L947.2.L3.74 (San Ernesto).
TypE LocAr-rry: "Bolivia at Charuplaya (1350 m.) and at San Ernesto, Mapiri district
(500 m.)".
DISTRIBUTIoN: Amazon slopes of Andes in southern Peru and Bolivia.
Hyla carinata Andersson/ 1938. Ark. Zool.,30A(23):22.

rypE(s): Syntypes: NHRM 1874 (3 specimens).
TypE LocAttrv: San ]os6, Tarraco, Bolivia.
Hyla carnifer Duellm an, L969. Herpetolo gica, 25,242'

rYPE(s): HolotYPe: KU 117993.
)'Tur',dapi (formerly known as Corneja Astorga,0 24' S' 78" 51' W)'
TypE LoCArrr",
Provincia Pichincha, Ecuador, L450 m" '
DTsTRTBUTTON: Pacific slopes of Andes in Colombia
and Ecuador.
coMMENr: In the Hyla columbiana grotp'
Hyla caraalhoiPeixoto, 1981. Rev. Brasil' Biol', 41:515'

rvnn(s): HolotYPe: EI 6501.
TypE LOCArrr"''Purque Nacional da Serra dos Org6os, Teres6polis' Rio de |aneiro'
Brazil.
CoMMENT: In the Hyla circumdata group, accolding to the original description'
23L6'
Hyla cembra Caldwell, Lg74. occas. Pap. Mus. Nat. Hist. Univ. Kansas,
rYPE(s): HolotYPe: KU 137035'
TypE LocArrr"' burrrpamento Rio Molino,2L60 m, Oaxaca' Mexico'
coMMENr: In the Hyla bisiincta gtoup according to the original description'
Hyla chaneque D:uellman, L96L. Herpetologica, 17:L'

rYPE(s): HolotyPe: KU 58439.
)'rtr"am above (6.2 kilometers by road south of) Ray6n Mescalapa'
TypE LocArrrr'
Chiapas, Mexico, (elevation 1690 meters)"'
DrsrRrBUTroN: Atlantic slopes in Oaxaca and Chiapas and Pacific
slopes in Chiapas
in southern Mexico.
CoMMENT: In the Hyla taeniopus SrouP; see Duellman, L970, Monogr'
Mus' Nat'
Hist. Univ. Kansas, L:440-445.
Hyla charadricola Duellman , Lg64. Univ. Kansas Publ. Mus' Nat' Hist', L5:478'
rYPE(s): HolotyPe: KU 58414.
)'Rio Totolapa, 14.4 kilometers by road west of Huachinango'
TypE LOCArrrr,
Puebla, Mexico, elevation 2280 meters"'
DrsrRrBUTrON: High mountains of Hidalgo and_Puebla in eastern
Mexico.
CoMMENT: In the"Hyla bistincta glouP; sei Duellman, L970, Monogr'
Mus' Nat
Hyla charazaniYellard,, 1970. Rev. Mus. Argent. Cienc' Nat' Bernardino Rivadavia'
L0:16.
rYPE(s): MACN.
TypE'LOCArtry: Charazafli, Departamento Munecas' Bolivia.
DISTRIBUTIoII: Known only from the type locality'
Hyla chinensls Giinther, 1859 "L858". Cat. Batr. Sal. Coll. Brit. Mus':108'
oRIGINAL NAME: Hyla arborea Yat. chinensis'
rypE(s): Syntypes:"including BM 1947.2.23.93 (* several not traced); Duellman,
L|TT; DjjTierreich, 95I+5, considered BM L947.2.23.93 to be the holotype.
TypE L9CALTTy: "China"; given by Boulen get, L882, Cat. Batr. Sal.
Brit. Mus':382, as
"China" , "Formosa" , and "Chusan?"'
DTSTRTBUTTON: Central and southeastern China
(including Taiwan) to northern
Vietnam.
CoMMENT: In the Hyla arboreagtouP; see Pope, L931, Bull' Am'
Mus' Nat' Hist'' 61:
46G;Liu, L911,"Fi"ldiunu:2ool-. Mem., 2:225; Okada, 1931, Tailless Batr.Japan'
Emp.:54-66; Bourret , Lg42, Batr. Indochine:223 -226; and Liu and Hu,
I96L,
Taiiless Amph. China:132-L34, for discussion'
REVIEWERS: Leo J. Borkin (LlB); Shuqin Hu (SH); Masafumi
Matsui (MM); Ermi
Zhao (EZ).
Hyla chryses Adler, L965. Occas. Pap. Mus. Zool. Univ. Michigan, 642:L.
rYPE(s): Holotype: UI0l4,.I[ltZ L25374.
TypE LocALITy: "between 'Puerto Chico' and 'Asoleadero' (about 45 km airline
west-northwest of Chilpancingo), Guerrero, Mexico, 2540-2600 m."
DISTRIBUTIoN: Sierra Madre del Sur in Guerrero, Mexico.
coMMENr: In the Hyla bistincta group; for discussion see Duellman, L970, Monogr.
Mus. Nat. Hist. Univ. Kansas, I:468-470.
Hyla chrysoscelis Cope, 1880. Bull. U.S. Natl. Mus., L7:29.

oRIGINAL NAME: Hyla femoralis chrysoscelis.
rvnr(s): Holotype: ANSP L3672.
TYPE LocAlrry: Dallas, Texas, USA.
DISTRIBUTIoN: Southeastern North America from eastern North Dakota and
Delaware to southern Texas and northern Florida, USA.
coMMENr: In the Hyla aersicolor group. Hyla chrysoscelis is one of a pair of cryptic
species wit}l. Hyla aersicolor. See Maxson, Pepper, and Maxson, L977, Science,
Washington D.C., L97:L0L2-13, and Johnson, 1966, Texas I. Sci., 18:361.
Hyla cinerea (Schneider,1792). Hist. Amph., l:174.

oRIGINAL NAME: Calamita cinereus.
rvrr(s): Unknown.
TypE LocALITy: "Carolina"; restricted to "Charleston, South Carolina", USA, by
Schmidt, L953, Check List N. Am. Amph. Rept., Ed. 6:69.
DISTRIBUTIoN: Southeastern USA; introduced into northwestern Puerto Rico.
coMMENr: See Duellman and Schwartz, L958, Bull. Florida State Mus., Biol. Sci., 3:
241, f.or discussion. The frequent citation of. Calamita carolinensis Pennant,
L792, as an older name for this species is based on a bibliographic error by
Schneider; Calamita carolinensis has never been proposed (DRF).
Hyla circumdata (Cope, 1867). Proc. Am. Philos. Soc., 11:555.

oRIGINAL NAME: Hypsiboas circumdatus.
rvnr(s): Syntypes: including MCZ 1508, other syntype not traced.
TypE LocArtrv: Rio de Janeiro, Brazil.
DISTRIBUTIoN: Mountains of Espirito Santo, Rio de ]aneiro, Sio Paulo, and Santa
Catarina in southeastern Brazll.
coMMENr: In the Hyla circumdata group.
Hyla claresignata A. Lutz and B. Lutz, L939. An. Acad. Brasil. Cienc., LL:67.
rvnE(s): Holotype: AL-MN L97L.
TypE LocAurv: Teies6polis, Rio de ]aneiro, BrazIL
DISTRIBUTIoN: Serra do Mar in Rio de Janeiro, 56o Paulo, and Paran6, southeastern
Brazil to Misiones Plateau, Argentina.
coMMENT: See Cochran, 1955,8u11. U.S. Natl. Mus., 206:86, and Cei, 1980, Monit.
Zool.Ital., N.S., Monogr., 2:467-469, for discussion.
Hyla clepsydra A. Lutz, L925. C. R. S6ances Soc. Biol., Paris, 93:2LL

rvnr(s): Holotype: AL-MN 976.
TypE LocAtttv: Serra da Bocaina, Sio Paulo, Brazil.
Hyla columbiana Boettger, 1892. Kat. Batr. Samml. Mus. Senckenb. Naturforsch.
Ges.:41.
rypE(s): SMF 2365 designated lectotype by Mertens, L967, Senckenb. Biol., 48:4L
TypE LocAurv: Popayin, Departamento Cauca, Colombia.
DISTRIBUTIoN: Highlands of southern Colombia.
coMMENr: In the Hyla columbiana group; synonymy includes Hyla oariabilis
according to Duellman and Trueb, L983, /n Rhodin and Miyata (eds.), Adv.
Herpetol. Evol. Biol.:33-51.
Hyla colymbaDunn,1931. Occas. Pap. Boston Soc. Nat. Hist., 5:400.

rvnE(s): Holotype: MCZ L0234.
TypE LocAtItv: La Loma, Provincia Bocas del Toro, Panama'
DrsrRrBUTroN: Caribbean slopes of Costa Rica and western Panama and Pacific
slopes of central and eastern Panama.
66MMENT: In the Hyla bogotensis group; see Duellman, 1972, Occas. Pap. Mus. Nat
Hist. Univ. Kansas, ll:21-22.
Hyla crassa (Brocchi, L877). Bull. Soc. Philomath. Paris, (7)1:130.

oRIGINAL NAME: Cauphias crassus.
rYPE(s): Holotype: MNHNP 6331.
TYPE LocAurv: Mexico.
DISTRIBUTION:Cerro San Felipe and Sierra Mixe, Oaxaca, Mexico.
coMMENr: In the Hylabistincla grotp; see Caldwell, 1974, Occas. Pap. Mus. Nat
Hist. Univ. Kansas, 28:L-37.
Hyla crepitans Wied-Neuwied , 1824. Abbild. Naturgesch. Brasil.:pl. 47 , fig. 1.

ivpr(s): Syntypes: including AMNH 785, other syntypes not-traced.
rypE LoCALITY; Tamburil, ]iboya, and Areal da Conquista, Bahia, Btazil-
DrsrRrBUTroN: Central and noitheastern Brazil, Guianas, northern Colombia and
Venezuela, eastern Panama, northern Honduras.
g9MMENT: In the Hyla boans group. This taxon apparently is a composite of severaj
species; see Kluge , L979; Occas. Pap. Mus. Zool. Univ. Michigan, 588:L-24.
Hyla uucifer Wied-Neuwied, 1838. Reise N. Am., l:249.

rYPE(s): Unknown.
TYPE LOCALITY: "Cantonment Leavenworth" (:port Leavenworth), Leavenworth
County, Kansas, USA.
DrsrRrBUTroN: Eastern North America; introduced at Marianao and Canasi, Cuba.
REVIEwERS: A1bert Schwartz (AS); Richard Thomas (RT).
Hyla cyanomma Caldwell, L974. Occas. Pap. Mus. Nat. Hist. Univ. Kansas,28:8
TypE LoCAlIry: 1.2 km north of Cerro Pe16n, 1650 m, Oaxaca, Mexico.
DISTRIBUTIoN: Sierra JuArez, Oaxaca, Mexico.
coMMENr: In the Hyla bistincta group according to the original description.
Hyla cymbalum Bokermann, 1963. Neotropica, 9:27 .

7'Campo Grande, Santo Andr6, Sio Paulo, Brasil"'
TypE LocAltrv,
coMMENr: In the Hyla raddiana (:Hyla pulchella) grotp, according to the original
description. Probably conspecific with Hyla pulchella (UC).
Hyla debilis Taylor, L952. Univ. Kansas Sci. BuIl., 35:880.

TypE LocALlry' i'Islu Bonita, (American Cinchona Plantation) eastern slope of
V6lcan P6as, [Provincia Heredia,] Costa Rica, 5600 ft'"
DrsrRrBUTroN: Caribbean slopes of Costa Rica and Pacific slopes of southwestern
Panama.
CoMMENT: In the Hyla riaularis gtoup; see Duellman, 1970, Monogr. Mus. Nat.
Hyla decipiens A. Lutz, L925. C. R. S6ances Soc. Biol., Paris, 93:2L2.

rypE(s): Syntypes: AL-MN 90-9L, USNM 96194.
TYPE LocAlIrv: Rio de Janeiro, Rio de ]aneiro, Btazll.
DrsrRrBUTroN: Coastal region of eastern Braztl from Pernambuco to Rio de faneiro.
coMMENr: See B. Lutz, L973, Brazil. Spec. Hyla:208.
Hyla dendroscarta Taylor, 1940. Proc. U.S. Natl. Mus., 89:45.

rYPE(s): Holotype: USNM L03679.
TypE LocALITy: "Cuautlapa, Yeracruz", Mexico.
DISTRIBUTIoIt: Mountains of central Veracruz and northern Oaxaca, Mexico.
coMMENr: In the Hyla bromeliacia group; see Duellman, 1970, Monogr. Mus. Nat.
Hyla dentei Bokermann, L967. Rev. Brasil. Biol., 27:L09.
TypE LocAtrrv: Serra do Navio, Amap6, Brazil.
coMMENr: In the Hyla albopunctata group or Hyla geographica grovp, according to
the original description.
Hyla denticulenta Duellman, L972- Occas. Pap. Mus. Nat. Hist. Univ. Kansas, Ll:22.
TypE LocALITy: "Charta, Departamento Santander, Colombia,2400 m".
DISTRIBUTIoITI: Andes of northern Colombia.
coMMENr: In the Hyla bogotensis group.
Hyla dolloi Werner, 1903. Zool. Anz.,26:253.
ryPE(s): Syntypes: MRHN 648L (2 specimens).
TYPE LocALITY: "Brasil".
DISTRTBUTIow: Unknown.
coMMENT: Name not associated with any known species (WED).
Hyla ebraccata Cope, 1874. Proc. Acad. Nat. Sci. Philadelphia,26:69.
rvnE(s): Holotype: ANSP 2079.
TypE LocALITy: "region of Nicaragua".
DISTRIBUTIoN: Lowlands of southern Mexico and Central America to northwestern
Colombia.
coMMENT: In the Hyla leucophyllata group; see Duellman, L970, Monogr. Mus. Nat.
Hist. Univ. Kansas, L:227.
Hyla echinafa Duellman, L962. Trans. Kansas Acad. Sci., 54:349.

rvrr(s): Holotype: UIMNH 49339.
TypE LocAttry: Campamento Vista Hermosa, 1500 m, Oaxaca, Mexico.
DISTRIBUTIow: Sierra JuArez in northern Oaxaca, Mexico.
coMMENT: In the Hyla miliaria group; see Duellman, L970, Monogr. Mus. Nat. Hist.
Univ. Kansas, L:346-348.
Hyla elegans Wied-Neuwied, L824.Isis von Oken, 6:67L.
rvnr(s): Not traced (?AMNH).
TypE LocALItv: Ponte do Gentio, Rio Alcobaga, Daravelas, Bahia, Btazil, according
to Bokermann, 1966. Lista Anot. Local. Tipo Anf. Brasil.:49.
DISTRIBUTIoN: Atlantic forests from Bahia to Sio Paulo, Brazil.
coMMENr: In the Hyla leucophyllata group. Removed from the synonymy of Hyla
leucophyllata by Caramaschi and Jim, 1982, Cienc. Cult., Supl., 7:848.
Hyla elkejungingerae Hen1e, 1981. Amphibia-Reptilia, 2:L23.
rYPE(s): Holotype: ZFMK 33352.
"Peru: El Boqueron, ca. 1000 m NN".
TypE LocALITy:
Vicinity of the type locality.
DrsrRrBUrroN:
coMMENr: Most likely a synonym of. Osteocephalus taurinus (MSH).
Hyla erythromma Taylor, L937. Proc. Biol. Soc. Washington, 50:48.
rypE(s): Holotype: FMNH 100083 (formerly EHT-HMS 5975).
TypE LocALITy: "in pines along Mexico-Acapulco highway at km. 350, near a
spring known as Agua del Obispo, between Rincon and Cajones", Guerrero,
Mexico.
DISTRIBUTIoN: Mountains of southern Mexico (Sierra Madre del Sur in Guerrero;
Sierra luArez in Oaxaca).
COMMENT: SoIe member of the Hyla erythromma grouPi see Duellman, 1970,
Hyla euphorbiacea Gtinther, 1859 "L858". Cat. Batr. Sal. Coll. Brit. Mus.:109.
rvnr(s): Syntypes: BM L947 .2.24.L5-L6 (Mexico), L947 .2.24.L7 (Cordova),
19 47 .2.24.1 8- 1 9 (Cordilleras).
TypE L96ALITY: "Cordova" (:C6rdoba, Veracruz, Mexico); "Cordilletas"; "Mexico'
DrsrRrBUTroN: Highland.s of southern Mexico (central Vetacruz to Oaxaca) and A1-
Yerapaz, Guatemala.
couurrur, In the Hyla eximia grotp; Hyla bocourfi (Mocquard, 1899), synonymized
with Hyla eupiorbiacea by Duellman , L970, Monogr. Mus. Nat. Hist. Univ.
Kansas, 1:505-510, is probably a distinct species in the highlands of Alta
Yerapaz and Baja Yetapaz, Guatemala (WED and JAC).
Hyla eximiaBaird,1854. Proc. Acad. Nat. Sci. Philadelphia,T:6L'

rypE(s): Syntypes: USNM 3248 (2 specimens).
TypE LocALITY: "Valley of Mexico", Mexico.
DrsrRrBUTroN: Highlands of southwestern USA (Arizona and New Mexico) and
Mexico southward to Guerrero, Morelos, and Ptrebla.
coMMENr: In the Hyla eximia group; includes as a synonym Hyla wrightorum
Taylor, 'tg}g, Univ. Kansas Sci. BuIl., 25:436. For discussion see Duellman,
1970, Monogr. Mus. Nat. Hist. Univ. Kansas, 1:499-505, and Blair, L960,
Southwest. Nat., 5:129-135.
Hyla faber Wied-Neuwied, L821. Reise Brasil., 2:249.

rYPE(s): Unknown (?AMNH).
TypE LocAltrv: Santa In6s ("Fazenda St. Agnes"),Bahia,Brazil.
DrsrRrBUTroN: Eastern to southern Brazil; southeastern Paraguay; Misiones,
Argentina.
coMMENT: In the Hyla boans group; for discussion see B. Lutz, L973, Brazil. Spec.
Hyla:Ll. See also Cei, 1"980, Monit. Zool.Ital., N.S., Monogr., 2:454-459.
Hyla fasciafa Gtinther, 1859 "L858". Cat. Batr. Sal. Coll. Brit. Mus.:100.
rvnr(s): Holotype: BM 58.4.25.22.
TYPE LocALITY: "Andes of Ecuador".
DrsrRrBuTror.i: Guianas and upper Amazon Basin from Ecuador to Bolivia.
coMMENr: In the Hyla geographica groupi see Duellman, L973, Copeia, 1973:523-
526.
Hyla faaosa Cope, 1886. Proc. Am. Philos. Soc., 23:95.

rYPE(s): Holotype: ANSP 11398.
TypE LocAlIrv: Pebas, Departamento Loreto, Peru.
DrsrRrBUTroN: Upper Amizon Basin in Ecuador, Peru, Bolivia, and western Brazi-
French Guiana.
69MMENT: In the Hyla leucophyllata group; see Duellman, L974, Occas. Pap. Mus-
Hyta femoralisBosc,1800. In Daudin, Hist. Nat. Quadrup. Ovip., Livr. 1:10.

rYPE(s): Unknown.
TypE LOCALITy: "La Caroline"; restricted to "Charleston, South Carolina", USA, t'r
Schmidt, 1953, Check List N. Am. Amph. Rept., Ed' 6:71.
DrsrRrBUrroN: Southeastern USA (Attantic Coastal Plain from Virginia to
Louisiana).
coMMENr: In the Hyla oersicolor gtoup.
Hyla fimbrimembra Taylor, L948. Copeia, 1948:233.
rypE(s): Holotype: R. C. Taylor 76L (now FMNH?).
TypE LocAurv: Isla Bonita, Provincia Heredia, Costa Rica.
DrsrRrBUTroN: Caribbean slopes of north-central Costa Rica, from Monteverde to
Volc6n Barba.
coMMENr: In the Hyla miliaria group; see Duellman, L970, Monogr. Mus. Nat. Hist
Univ. Kansas, 1:348.
Hyla fuentei C. Goin and O. Goin, L968. Copeia, 1968:581

rvnr(s): Holotype: CM 442L8.
TypE LocAlIrv: Powakka, Suriname District, Surinam.
Hyla fusca Laurenti, L768. Synops. Rept.:34.

rYPE(s): Unknown.
TYPELocAlIrv: Unknown.
DISTRIBUTIoN:Unknown.
Hyla geographica Spix, 1824. Spec. Nov. Testud. Ran. Brasil.:39.

rypE(s): Formerly ZSM 3510 (lost); see Hoogmoed and Gruber, 1983, Spixiana,
Suppl., 9:370, for discussion.
TypE LocAlIrv: Rio Tef.6, Amazonas, Btazil.
DISTRIBUTIow: Tropical South America east of the Andes; Trinidad.
coMMENr: In the Hyla geographica group; see Duellman, L973, Copeia, L973:526-
530, and Lutz, L973, Brazil. Spec. Hyla:34-37.
Hyla gieslerl Mertens, 1950. Wochenschr. Aquar. Terrarienkd., 44:185.

rvnr(s): Holotype: SMF 4L2L7.
TypE LocAtIrv: Barro Branco, Rio de faneiro [Brazil].
DISTRIBUTIoN: Lowlands of Espirito Santo and Rio de ]aneiro, Brazil.
coMMENT: In the Hyla paruiceps group; see Heyer, 1980, Proc. Biol. Soc
Washington, 93:655, for taxonomic status.
Hyla godmani Gtinther, 1901. Biol. Cent. Am., Rept. Batr.:275.

rvrr(s): Syntypes: BM 1901.12.L9.88-95 (]alapa), and 1901.12.L9.96 (Misantla); nU
L90L.L2.L9.96 designated lectotype by Duellman, 1964, Copeia, L964:455.
TypE LocArtrv: Restricted to Misantla, Veracruz, Mexico,by lectotype designation.
DISTRIBUTIoN: Lowlands of Veracruz, Mexico.
coMMENT: In the Hyla godmani group; see Duellman, L970, Monogr. Mus. Nat.
Hist. Univ. Kansas, 1:356,-359.
Hyla graceae Myers and Duellman, 7982- Am. Mus. Novit., 2752:5.
rvnr(s): Holotype: AMNH 107966.
TypE LocArmv: Cgntinental Divide southeast of Cerro Colorado, 1650 m, Provincia
Chiriqui, Panama.
DISTRIBUTIow: Mountains of eastern Panama.
coMMENr: In the Hyla pseudopuma group.
Hyla grandisonae Goin, L966. Q. J. Ftorida Acad. Sci.,29:39

rypE(s): Holotype: BM 1938.10.3.25.
TYPE LocAtrrv: Mazaruni, Guyana.
Hyla granosa Boulenger, L882. Cat. Batr. Sal. Brit. Mus.:358.

rvnr(s): Syntypes: BM L947.2.L2.93 (Demerara Falls), L947.2.12.94-96 (Santar6m),
L947.2.12.97-98 (Interior of Brazil), L947.2.L2.99 (Canelos); BM L947.2.12.99
designated lectotype by Duellman, 1974, Occas. Pap. Mus. Nat. Hist. Univ.
Kansas,27:8.
TYPE LocALITY: "Demerara Falls", Guyana; "Santarem", Par6, Btazil; "Interior of
Brasil"; "Canelos, [Provincia Pastaza,] Ecuador". Restricted to Canelos by
lectotype designation.
DISTRIBUTIoN: Guianas and Amazon Basin in Brazil, Colombia, Venezuela, Ecuador,
and Peru.
&"
CoMMENT: In the Hyla granosa grouP; see Hoogmoed,1979, Zool. Verh., Leiden,
L72:L-45.
Hyla gratiosa LeConte, 1856. Proc. Acad. Nat. Sci. Philadelphia, 8:146.
rvnrls;, Holotype: ANSP 2089; the status of type(s) is confusing, see Caldwell,
1982, Cat. Am. Amph. Rept., 298.L, for discussion.
TypE LOCALITY: "Lower part of Georgia"; restricted to "Libetty County, Georgia",
USA, by Schmidt, 1953, Check List N. Am. Amph. Rept., Ed.6:7L.
DrsrRrBUTroN: Southeastern USA (North Carolina to eastern Louisiana); isolated
records from southern Kentucky, Tennessee, and northern Alabama.
coMMENr: Reviewed by Caldwell, L982, Cat. Am. Amph. Rept., 298.L-2-
Hyla gryllaf a Duellm an, 197 3. Herpetolo gica, 29 :2L9.

TypE L9CALITY: "Estaci6n Biot6gica Rio Palenque, 55 km north of Quevedo,
Provincia Los Rios [Pichincha], Ecuador,220 m".
DISTRIBUTIoTv: Pacific lowlands of northwestern Ecuador.
coMMENT: In the Hyla miuocephala grouP.
Hyla guentheriBoulenger, 1886. Ann. Mag. Nat. Hist., (5)18:445.

rvrr(s): Holotype: BM 62.5.8.9.
TypE LocALITy: "Rio Grande do Sul", Brazil.
DrsrRrBUTroN: Southeastern Brazil (Rio Grande do Sul and Santa Catarina) and
northeastern Argentina.
coMMENT: Hyla guentheri is a replacement name f.or Hyla leucotaenia Gtinther, 1868,
Proc. Zool. Soc. London, 1868:489. See Lutz, 1973,Btazil. Spec. Hyla:120.
Hyla hallotnellii Thompson , L9L2. Herpetol. Not., 2:2.

rYPE(s): Holotype: CAS 23808.
rypE LocALITv: "Kikaigashima, Loo Choo [:Ryukyu] Islands, lapan".
DISTRIBUTIow: Ryukyu Is., |apan.
coMMENT: In the Hyla arborea group; for discussion see Nakamura and I-Jeno, L963,
Japan. Rept. Amph. Color:35, and Okada, L93L, Tailless Batr. Japan. Emp.:67-
70. See also Matsui and Matsui, L982, Japan. |. Herpetol.,9:79-86-
REVIEWERS: Leo J. Borkin (LlB); Shuqin Hu (SH); Masafumi Matsui (MM); Ermi
Zhao (EZ).
Hyla haraldschultzi Bokermann, L962. Neotropica, 8:88.

rvru(s): Holotype: WCAB 2896.
TypE LocAlrry: "Santa Rita do Weill, Sio Paulo de Olivenga, Estado de Amazonas,
Brasil".
DISTRIBUTIoN: Eastern Peru and Rio Solim6es, Brazil.
Hyla hazelae Taylor, L940. Univ. Kansas Sci. Bull., 26:385.

rurr(s): Holotype: FMNH L0A047 (formerly EHT-HMS L6262).
TypE LocALITy: "Cerro San Felipe, about L0 miles north of Oaxaca, Oaxaca",
Mexico.
DISTRIBUTIow: Mountains of central Oaxaca, Mexico.
coMMENT: In the Hyla hazelae grovp; see Duellman, L970, Monogr. Mus. Nat. Hist.
Univ. Kansas, 1:385.
Hyla heilprini Noble, L923. Am. Mus. Novit., 61:L.

rvru(s): Holotype: AMNH 11401.
TypE LocAl-rry: Lo Bracito [:Los Bracitos], Provincia Pacificador [now Duarte],
Dominican Republic.
DTsTRIBUTIoN: Hispaniola, West Indies.
coMMENr: Questionably in the Hyla albomarginata group; see Trueb and Tyler,
L974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 24:l-60.
REvIEwERS: Albert Schwartz (AS); Richard Thomas (RT)
Hyla helenae Ruthven, L9L9. Occas. Pap. Mus. Zool. Univ. Michigan, 69:t0
rYPE(s): Holotype: U}I{MZ 5268L.
TypE LocAltrv: Valley of the Demerara River, Guyana.
Hyla hobbsi Cochran and Goin, L970. Bull. U.S. Natl. Mus.,288:311.
TypE LOCALITy: "Cafro Goacay6, atributary of the Rio Apoporis, in Amazonas,
Colombia".
DISTRIBUTIoN: Region of Rio Apoporis, Amazonas, Colombia.
coMMENT: In the Hyla punctata group; see Pybutn, 1978, Proc. Biol. Soc.
Washington, 91:123, for taxonomic status.
Hyla hutchinsi Pyburn and Hall, 1984. Herpetologica, 40:366.

unn(s): Holotype: Estaci6n de Biologia Tropical Roberto Franco 3000.
rypE LocAt-rry: "about 2 km SW Umuflapito (0o 34' N, 70" 05' W), Comisaria de
Vaup6s, Colombia".
DrsrRrBUTIoN: Known only from the type locality.
coMMENr: In the Hyla geographica group, according to the original description.
See comment under Hyla.
Hyla hypselops (Cope, 1870). Proc. Am. Philos. Soc., 11:554.

oRIGINAL NAME: Hypsiboas hypselops.
rYPE(s): Unknown.
TypE LocALITy: "Pebas", Departamento Loreto, Peru.
Hyla ibitiguara Cardoso, 1983. Iheringia, Zool., 62:38.

TypE LocALrry: "Fazenda Salto (20 40' 3-46" L6' W;900 m), Alpin6polis, Minas
Gerais, Brasil".
coMMENT: In the Hyla circumdata group, according to the original description.
Hyla imitafor (Barbour and Dunn, l92l). Proc. Biol. Soc. Washington, 34:160.
oRIGINAL NAME: P aludicola imitator.
TYPE LocAurv: Lake Cudajaz (:Lago Codaj6s), Amazonas, Brazil.
Hyla inframaculata Boulenger, L882. Cat. Batr. Sal. Brit. Mus.:354.

rypE(s): Holotype: BM L947 .2.13. 10.
TypE LocALITy: "Santarem", Par6, Btazll.
Hyla intermixta Daudin, 1802. Hist. Nat. Rain. Gren. Crap.:23.

tvnr(s): Formerly in MNHNP; now lost.
TYPE LocALITY: Unknown.
DISTRIBUTIoIS: Unknown.
coMMENT: This name is not assignable to any known species (WED)
Hyla izecksohni lim and Caramaschi, L979. Rev. Brasil. Biol., 39:7L7.
TypE LOCALITv: "Rubiio Jtnior, Botucatu, Sio Paulo, Brasil (aprox. 22" 53' S e 48'
30'w)".
coMMENT: In the Hyla circumdata group according to the original description.

Hyla japonica Giinther, 1859 "L858". Cat. Batr. Sal. Coll. Brit. Mus.:109.
oRIGINAL NAME: Hyla arborea var. iaponica.
ryPE(s): Syntypes: BM 44.2.22.107 (3 specimens).
TYPE LocAltrv: Japan.
DrsrRrBUTroN: Japan, Korea, Mongolia, central (south to Guizhou) and northeasterr.
China, and USSR from Lake Baikal through the Amur and Ussuri river basins
southeast to the coast, Sakhalin I. and Kunashir I.
CoMMENT: In the Hyla arborea BrouP; see Kuramoto, 1980, Copeia, 1980:106.
Chinese population reviewed by Liu and FIu, L96L, Tailless Amph. China:
127-128 (as Hyla arborea). Genetically isolated from European Hyla arborea,
according to Daito, 1968, Zool. Mag., Tokyo, 77:LI7 -127 , and Kawamura and
Nishioka, L977,In Taylor and Guttman (eds.), The Reproductive Biology of
Amphibians:L32.
REVTEwERS: Leo J. Borkin (L]B); Shuqin Hu (SH); Masafumi Matsui (MM); Ermi
Zhao (EZ).
Hyla juanitae Snyder, 1972. J. Herpetol., 6:5.
rypE LocAlmy: 15 km (by road) northeast of San Vicente de Benitez,1070 m,
Guertero, Mexico.
Pacific slopes of Sierra Madre del Sur in Guerrero and Oaxaca,
DrsrRrBUTroN:
Mexico.
coMMENT: In the Hyla melanomma grouP.
Hyla kanaima Goin and Woodley, L965. Zool. ]. Linn. Soc.,48:135.

rYPE(s): Holotype: BM 1965.230.
TypE LocAurv: Mt. Kanaima, Potaro District, Guyana.
DISTRIBUTIoN: Mt. Kanaima and Mt. Roraima, southern Guyana.
coMMENr: In the Hyla geographica grouP.
Hyla labialis Peters, 1863. Monatsber. Preuss. Akad. Wiss. Berlin, 1863:463.
rvru(s): Holotype: ZMB 4913.
TypE LocALrry: Mountains surrounding Bogota, Departamento Cundinamarca,
Colombia.
DrsrRrBUTroN: Andes of central and northeastern Colombia and western
Venezuela.
coMMENr: In the Hyla labialis group; see Cochran and Goin, 7970, Bull. U.S. Natl.
Mus., 288:251. Hyla labialis meridensis Rivero, L96L, Bull. Mus. Comp. Zool.,
126L31, is probably a distinct species (WED).
Hyla lancasteriBarbour, 1928. Proc. New England Zool. Club, 10:31.
rvnr(s): Holotype: MCZ 13062.
TypE LocALIty: Peralta, Provincia Cartago, Costa Rica.
DISTRIBUTIoN: Caribbean slopes of Costa Rica and Panama.
coMMENT: In the Hyla lancasteri group; see Myers and Duellman, L982, Am. Mus.
Novit.,2752:L5.
Hyla lanciformis (Cope, L870). Proc. Am. Philos. Soc., 11:556.
oRIGINAL NAME: Hypsiboas lancif ormis.
TYPE(s): Unknown.
TYPE LoCALITY: "Ecuador" .
DrsTRrBUTror.r: Middle and upper Amazon Basin and north-central Venezuela.

coMMENr: In the Hyla albopunctata group. Population in north-central Venezuela
recognized as Hyla lanciformis guerreroi Rivero, L971, Caribb. J. Sci., 1L:4.
Hyla langei Bokermann, 1965. I. Ohio Herpetol. Soc., 5:49.
TypE LocALtrv: Marumbi, Morretes, Paranl, Brazil.
DISTRIBUTIoIt: Known only from the type locality.
coMMENT: In the Hyla martinsi group according to original description.

Hyla larinopygion Duellman, L973. Herpetolo gica, 29,224-
TypE LOCALITy: "Quebrada Santa Tereza, between Popay6n and Quintana,
Departamento Cauca, Colombia, 2200 m".
DISTRIBUTIow: Cordillera Central, Colombia; Cordillera Oriental in southern
Colombia and northern Ecuador.
coMMENr: In the Hyla larinopygion grouP.
Hylalascinia Rivero, 1969. Caribb. I. Sci.,9:145.

rypE LoCAurv: 1abor, above Delicias (opposite Herran, Colombia), P6ramo de
Tam6, T6chira, Venezuela.
DrsrRrBUrroN: Eastern slopes of northern part of Cordillera Oriental of Andes in
extreme western Venezuela.
coMMENT: In the Hyla bogotensis group.
Hyla leali Bokermann, 1964. Neotropica, 10:3.

TypE LocAt-rry: "Forte Principe [da Beira], Territ6rio Federal de Ronddnia, Brasil".
DISTRIBUTIoN: Upper Amazon Basin in Brazil and Peru.
coMMENT: In the Hyla minima group; for discussion of taxonomic status see Heyer,
1977, Pap. Avulsos Zool., Sio Paulo, 3l:144.
Hyla legleri Taylor, 1958. Univ. Kansas Sci. Bu11., 39:33.

TypE LOCALITy: "15 km WSW San Isidro del General, San ]os6 Province", Costa
Rica.
DrsrRrBUTroN: Pacific slopes of southern Costa Rica and western Panama.
coMMENT: In the Hyla salaadorensis group; see Duellman,1970, Monogr. Mus. Nat
Hyla lemal Rivero, L971. Caribb. J. Sci., 11:183.

rYPE(s): Holotype: UPRM 3L79.
TypE LocAlrrv' Paso del Danto, La Escalera, between El Dorado and Santa Elena de
Uairen, 1300-1400 m, Serrania de Lema, Estado Bolivar, Venezuela.
DrsrRrBUTroN: La Escalera region (Bolivar, Venezuela) and Mount Roraima, Guyana
(MSH).
Hyla leptolineata P. Iiraun and C. Braun, L977. Rev. Brasil. Biol., 37:853.
rYPE(s): Holotype: MRGS 8086.
TypE LOCALITv: "Fortaleza dos Aparados, Municipio de Cambar6 do Sul, Rio
Grande do Sul, Brasil".
DISTRIBUTIoN: Rio Grande do Sul, Brazll.
coMMENr: In the Hyla potytaenia group and most closely related to Hyla guentheri,
according to the original description.
Hyla leucophyllata (Beireis, 1783). Schr. Ges. Naturforsch. Freunde Berlin, 4:L82.
oRIGINAL NAME: Rana leucophyllata.
rypE(s): NHRM 157 (now lost) was regarded as the holotype by Duellman, 1977,
Das Tierreich, 95:69; this was disputed by Bcihme, L98L, Bonn. Zool. Beitr., 32:
283-295, who suggested that the holotype was lost and that NHRM 157 was
one of the original Linnaean syntypes of. Hyla boans.
TYPE LoCALITY: "Surinam".
The Amazon Basin and the Guianas.
DISTRIBUTIoN:
coMMENr: In the Hyla leucophyllata group; for discussion of geographic variation
and synonyms, see Duellman, L974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas,
27:18. See comment under Hyla elegans.
Hyla limai Bokermann, 1962. Neotropica, 8:81.

TypE LocALITy: "Sio Vicente, Estado de Sdo Paulo, Brasil".
coMMENT: Related to Hyla minuta and Hyla tnerneri, according to the original
description.
Hyla lindae Duellman and Altig, 1978. Herpetologica,34:177.

rvrr(s): Holotype: KU 164402.
TvPE LocALrry: "11 km east-southeast (by road) of Papallacta, Provincia Napo,
Ecuador, 2660 m".
Amazonian slopes of Andes
DISTRIBUTIoN: in southern Colombia and northern
Ecuador.
coMMENr: In the Hyla larinopygion group.
Hyla loquax Gaige and Stuart, 1934. Occas. Pap. Mus. Zool. Univ. Michigan,2SL:L
rvrr(s): Holotype: UMIli{Z 75446.
TYPE LocArmv: Ixpuc Aguada, north of La Libertad, Departamento EI Pet6n,
Guatemala.
DISTRIBUTIoN:Atlantic lowlands from southern Veracruz, Mexico, to Costa Rica
coMMENr: In the Hyla godmani group; see Duellman, L970, Monogr. Mus. Nat.
Hyla lorseridgei Rivero, 1961. Bull. Mus. Comp. Zool., L26:108.

TYPELocAltrv: Pico Culebra, Cerro Duida, Territorio Amazonas, Venezuela.
DISTRIBUTIoN:Known only from the type locality.
coMMENT: See La Marca and Smith,1982, Caribb. J. Sci., L8:2L, for discussion of
the use of this name rather than Hyla ginesi.
Hyla luteoocellata Roux, L927. Verh. Naturforsch. Ges. Basel, 38:260.

oRIGINAL NAME: Hyla luteo-ocellata.
rvrr(s): NHMB 3900 designated lectotype by Forcart, L946, Verh. Naturforsch.
Ges. Basel, 56:L24.
TypE LocALITy: "El Mene, Prov. Falcon, Y6n6zu6la".
DISTRIBUTIoN: Northern Venezuela (Lake Maracaibo to Unare Depression).
COMMENT: In the Hyla parrsiceps groupi see Duellman and Crump, 1974, Occas. Pap
Mus. Nat. Hist. Univ. Kansas, 23:l-40.
Hyla lythrodes Savage, L968. Bull. S. California Acad. Sci., 67:L.

rYPE(s): Holotype: LACM 26766.
rYPE LocAurv:2L km SW Amburi at confluence of Rio Lari and Rio Dipnari,
Lim6n, Costa Rica.
DISTRIBUTIoN: Caribbean slopes of southern Costa Rica and western Panama.
coMMENT: In the Hyla uranochroa group; see Myers and Duellman, L982, Am. Mus.
Novit., 2752:2L.
Hyla marginata Boulenger/ L887. Ann. Mug. Nat. Hist. , (5)20:298.

rvnr(s): Holotype: BM L947 .2.12.80.
TypE LocALITy: "Mundo Novo, [Taquara,] Rio Grande do Sul", Brazil.
DISTRIBUTIoN: Southeastern Brazil (northern Rio Grande do Sul and southern Santa
Catarina).
Hyla marianae Dunn, L926. Proc. Boston Soc. Nat. Hist., 38:L29.
rvru(s): Holotype: MCZ LLL22.
TypE LocAlrry: "Spaldings, Clarendon Parish (altitude 2900 feet)",lamaica.
DISTRTBUTToN: Jamaica, West Indies.
coMMENT: In the Hyla wilderi grotp; see Trueb and Tyler, L974, Occas. Pap. Mus
Nat. Hist. Univ. Kansas, 24:L-60.
ANURA: HYLIDAE: HYLINAE t4l
REvIEwERS: Albert Schwartz (AS); Richard Thomas (RT)
Hyla marmorata (Laurenti, L768). Synops. Rept.:29.

oRIGINAL NAME: Bufo marmoratus.
rvnr(s): Unknown.
Amazon Basin in Btazll, Colombia, Ecuador, and Peru; southern
DrsrRIBUTroN:
Venezuela; Guianas.
coMMENT: In the Hyla marmorata group; see Bokermann, L964, Senckenb. Biol., 45
243-254.
Hyla martlnsi Bokermann, L964. Neotropica, L0:67.

rYPE(s): Holotype: WCAB L4759.
TypE LocALITy: "regi6n del Colegio de CaraEa, Serra do CaraEa, Santa Barbara,
Minas Gerais, Brasil".
DISTRIBUTIoN: Mountains of central Minas Gerais, Brazll.
coMMENT: In the Hyla martinsi group.
Hyla mathiassoni Cochran and Goin, L970. Bull. U.S. Natl. Mus., 288:286
rvrr(s): Holotype: FMNH 8L779.
TypE LocAtIrv: "Villavicencio, [Departamento] Meta, Colombia".
DTsTRIBUTIoN: Llanos of central Colombia.
coMMENr: In the Hyla microcephala group.
Hyla melanargyrea Cope, L887. Proc. Am. Philos. Soc., 24:45.

rvnr(s): Syntypes: ANSP LL2L6-L8.
TypE LocAurv: Chapada dos Guimaries, Mato Grosso, Btazil.
DrsrRrBUTroN: Interior basins of Mato Grosso to northeastern Brazil; Surinam
(MSH).
coMMENT: In the Hyla marmorata group; see comment under Hyla senicula.
Hyla melanomma Taylor, L940. Univ. Kansas Sci. Bull., 26:508.

rypE(s): Holotype: FMNH 100074 (formerly EHT-HMS 21578).
TypE LocALtry: "7 miles east of Chilpancingo (Ciudad Bravo), Guerrero, Mexico".
DrsrRrBUTroN: Pacific slopes of Sierra Madre del Sur in Guerrero and Oaxaca;
Atlantic slopes of Chiapas in southern Mexico.
coMMENT: In the Hyla melanomma group; see Duellman, L970, Monogr. Mus. Nat
Hist. Univ. Kansas, L:397-403, who recognized two subspecies.
Hyla melanopleura Bdulenger, LgLz. Ann. Mag. Nat. Hist., (8)10:185.

runr(s): Syntypes: BM L947 .2. 1 3.54-58.
TypE LocALITy: "Huancabamba, [Departamento Pasco,] E. Peru".
DISTRIBUTIoN: Lower Andean slopes in central Peru.
Hyla melanorhabdota (Schneider, L799). Hist. Amph., l:L73.

oRIGINAL NAME: Calamita melanorhabdotus.
unE(s): Unknown.
TYPE LocAurv: Brazil.
DTsTRIBUTIoN: ? Brazil.
coMMENr: Name not assignable to any known species (WED)
Hyla meridionalis Boettger, L874. Abh. Senckenb. Naturforsch. Ges., 9:186.

oRIGINAL NAME: Hyla arborea lrar. meridionalis.
rver(s): SMF 1872 designated lectotype by Mertens, L967, Senckenb. Bio1., 48:4L.
TypE LocAllrv: Orotava Valley Tenerife, Canary Islands [Spain].
DrsrRrBUTroN: Northwestern Africa and southwestern Europe, including Portugal
(Madeira), southern and eastern Spain, southern Ftance, northwestern Italy,
and Balearic and Canary Is.
C9MMENT: In the Hyla arborea SrovP; see Crespo, L973, Arq' Mus' Bocage, (2)3:613-
632.
REVIEwER: Leo ]. Borkin (LIB).
Hyla microcephala Cope, 1885. Ploc Am. Philos. soc., 23:28L.

rvPE(s): Syntypes: USNtVt L3473 (2 specimens); now lost'
TypE LOCAttry: "Chiriqui", Panama (iee comment under Dendrobates
maculatus)'
DISTRIBUTTOI,t: sOutheastern Mexico, central America' northern
and eastern south
America to southeastern Brazil.
CoMMENT: In the Hyla microcephala SrouP; see Duellman, L970,
Monogr' Mus' Nat'
Hist. Univ. Kaisas, L:2lL; B. Litz, {973, Brazil. Species Hyla:2L4; and
Duellman , Lg74, Occas. Pap. Mus. Nat. Hist. Univ' Kansas, 27:9, who
recognized five subspecie{. Uyto cheruei Cope, L894, Proc' Acad. Nat. Sci.
nhilXdelphia,46:L95, was plaied in the syngnyml of Hyla microcephalaby
Savage and Heyer, 1970, Rev. Biol' Tropical, 16:17'
Hyla microderma Pyburn, L977. J. Herpetol', 11:405'

rvnE(s): HolotyPe: UTA A-5012.
rypE LocAlrry:
i'about 2 km S. Yapima (long. 69" 28' W, Iat. 1o 03' N), Comisaria de
Vaup6s, Colombia".
otsrntsUilOr.r: Upper Amazon Basin in southeastern Colombia and Brazil'
coMMENr: In the Hyla geographica grouP.
Hyla microps Peters, L872. Monatsber. Preuss. Akad. Wiss' Berlin, 1872682'
TypELocArrrv, Nova Friburgo, Rio de Janeiro, Brazil'
DTSTRTBUTToN:Atlantic forests of southeastern Brazil.
g9MMENT: In the Hyla paraiceps group; see Duellman and Crump,'l'974, Occas. Pap
Mus. Nat. Hist. Univ. Kansis, 25:L-40, and Heyer, L977, Proc. Biol. Soc.
Washington, 93:365.
Hyla miliaria (Cope,1886). Proc. Am. Philos' Soc', 23:272'

oRIGINAL NAME: Hypsiboas miliarius.
rvnr(s): Holotype: USNM 14L93.
TYPE LocALITY: "Nic atagua" .
DrsrRrBUTroN: Nicaragui through Costa Rica and Panama to north-central
Colombia.
g9MMENT: L970, Monogr_. Mus. Nat. Hist
In the Hyla miliaria group; see Duellman,
Univ. Kansas, i:352-355. Specimens from Honduras previously allocated to
this species are being desciibed by Larry David wilson (wED).
Hyla minima Ahl, 1933. Zool. Anz.,104:28.

rvrE(s): HolotyPe: NHMW L9436.
TypE LocAr-lrv, iaperinha, near Santar6m, Pat6, Btazil'
DISTRIBUTIoT: Central Amazon Basin.
coMMENr: In the Hyla minima grovp; see Duellman, 1982, Amphibia-Reptilia, 3:
159.
Hyla minuscala Riveto, L971. Caribb. ). Sci., 11:1'

tvru(s): HolotyPe: UPRM 3377.
TypE LocAr-mv, Nirgua, Estado Yaracay, Venezuela'
DrsrRrBUrroN: InteJor llanos of Veneiuela eastward through Guianas to Bel6m,
Brazil.
Hyla minufa Peters, L872. Monatsber. Preuss. Akad. wiss. Berlin, 1872:680.
rvnr(s): Syntypes: Z}/.B 7456 (5 specimens).
TvPE LocAtttv: Nova Friburgo, Rio de Janeiro, Brazil'
DrsrRrBuTrorrr: Lowlands east"of Andes from Colombia, Venezuela, and Trinidad
southward to Bolivia, Argentina, and Uruguay'
coMMENT: For discussion see Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288:
277-280, and Cei, 1980, Monit. Zool.Ital., N.S., Monogr., 2:492-495.
Hyla miotympanum Cope, 1863. Proc. Acad. Nat. Sci. Philadelphia, L5:47.
rvnE(s): Syntypes: USNM 6311 and one with no number; now lost.
TypE LocAurv: Jalapa and Mirador, Veracruz, Mexico.
DrsrRrBUTIoN: Highlands from Nuevo Le6n to Oaxaca and central Chiapas in
eastern Mexico.
coMMENT: In the Hyla miotympanum group; see Duellman, L970, Monogr. Mus.
Nat. Hist. Univ. Kansas, L:372.
Hyla mixe Duellman, L965. Herpetologica, 2L:32.

TypE LocALrry: "a stream 4.2 kilometers by road south of Campamento Vista
Hermosa, (near latitude L7" 50' N and longitude 96" 20' W), at an elevation of
1,800 meters on the northern slope of Sierra de ]u6rez, Distrito de Ixtl6n,
Oaxaca, M6xico".
DISTRIBUTIoN:Sierra JuArez and Sierra Madre del Sur in Oaxaca, Mexico.
coMMENT: In the Hyla mixomaculata group; see Duellman,1970, Monogr. Mus. Nat
Hyla mixomaculata Taylor, 1950. Copeia, L950:274.

TypE LocAttry: Coscomatepec, Yetactuz, Mexico.
DISTRIBUTIoN: Central Veracruz in eastern Mexico.
coMMENT: In the Hyla mixomaculata group; see Duellman, L970, Monogr. Mus. Nat
Hist. Univ. Kansas, L:4L7-42I.
Hyla molitor Schmidt, L857. Sitzungsber. Akad. Wiss. Wien, Math. Naturwiss. Kl.,
24:1L.
rvnr(s): Syntypes: NHMW L6494; other syntypes unknown.
TypE LocAl-trv: "Chiriqui-Flusse unweit Bocca del toro", Panama; in error,
according to Duellman, L970, Monogr. Mus. Nat. Hist. Univ. Kansas, 1:650.
DrsrRrBUTroN: ? Bolivia (see comment).
coMMENT: Name not assigned to any known species; see Duellman, L970, Monogr.
Hyla multifasciata Grinther, 1859 "1858". Cat. Batr. Sal. Coll. Brit. Mus.:101.
rvrr(s): Holotype: BM L947 .2.23.6.
TypE LocALITy "Pa4a" , BraziI; stated as Be16m, ParA, Btazll, by Duellman, L977 , Das
Tierreich, 95:78.
DTsTRIBUTIoN: Lowlands of eastern Venezuela, Guianas, and Amazon Basin, Btazil.
coMMENT: In the Hyla albopunctata group.
Hyla musica B. Lutz, L949. Mem. Inst. Oswaldo Cruz, Rio de Janeiro, 46:576.
rvnr(s): Syntypes: MN 3203-19.
TypE LocAurv, Parque Nacional Serra dos Org6os, Teres6polis, Rio de |aneiro,
Brazil.
DISTRIBUTIoN: Serra dos Orgios, southeastern Btazil.
coMMENr: See Cochran, 1955, Bull. U.S. Natl. Mus., 206:170.ln t}".:.e Hyla
albomarginata group, according to Duellman, 1.970, Monogr. Mus. Nat. Hist.
Univ. Kansas, L:240.
Hyla mykter Adler and Dennis, L972. Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 7:8.
rvnE(s): Holotype: KU 137553.
TypE LocALtry "'!.1.4 km (by road) southwest of Puerto deJ Gallo, Guertero,
M6xico (about 35 km iirlir,e northeast of Atoy6c de Alvarez, elevation 1985
meters".
DTsTRIBUTIoN: Vicinity of Cerro Te6tepec, Sierra Madre del Sur, Guerrero, Mexico.
coMMENr: In the Hylabistincta Broup, according to the original description.
Hyla nahdereri B. Lutz and Bokermann, 1963. Copeia, 1963:559.

TypE LocAt.rry, "at Estrada Saraiva, outside and slightly below Sio Bento do Sul, a
small town in northern santa Catarina, Brazll (26" L4', 55" S, 49" 22', 50" W,
800 m alt.)".
DISTRIBUTIoN: Serra do Mar in Santa Catarina, Btazll.
g6MMENT: In the Hyla marmorata grotp, according to the original description.
Hyla nana Boulenger, 1889. Ann. Mus. Civ. Stor. Nat. Genova, (2)7:249.
rypE(s): Syntypes: BM L947.2.12.83-85 and MSNG 2972LA; MSNG 2972LA
designated lectotype by Capocaccia, L957 , Ann. Mus. Civ. Stor. Nat. Genova,
69:2L3.
TypE LOCALITY: "Colonia Resistencia, South Chaco, Argentine Republic".
DrsrRrBUTroN: Northeastern Brazil southward through central Paraguay, northern
Argentina, eastern Bolivia; extreme southern Brazil, Uruguay, and La Plata
Basin in Argentina.
CoMMENT: In the Hyla nana grouP. See Cei, 1980, Monit. Zool.Ital., N.S., Monogr.,
2:495-498. Hyla sanborni Schmidt, 1944, Field Mus. Nat. Hist. Publ., ZooI. Ser.,
29:153 (Holotype: FMNH 9581; Typ" Iocality: Hacienda Alvatez,15 km NE
San Carlos, Uruguay; Distribution: Southern Paraguay, central and eastern
Argentina, and Uruguay) is probably a distinct species (NlS). See Barrio, 1967
Physis, Buenos Aires, 26:521-524, f.or subspecies.
Hyla nanuzae Bokermann and Sazima, L974. Rev. Brasil. Biol., 33:333.
TypE LocAlrry: Kilometer 126, Serra do Cip6, Jaboticatubas, Minas Gerais, Btazll.
Hyla noaaisi Bokermann, 1968. J. Herpetol., L:26.

rvnr(s): Holotype: WCAB 3L743.
TypE LOCALtTy: " 'Fazenda Santo Onofre,' 10 km east of Marac6s, Bahia, Brazil" -
DISTRIBUTIoN: Bahia, Brazrl.

coMMENT: In the Hyla marmorata group; see comment under Hyla senicula.
Hyla nubicola Duellman, 7964. Herpetologica, 19:225.

TypE LocAlrry: "stream 3 kilometers southwest of Huatusco, Vetacruz, Mexico
(elevation L,325 meters)".
DISTRIBUTION: Mountains of central Veracruz in eastern Mexico.
coMMENr: In the Hyla mixomaculata group; see Duellman, 1970, Monogr. Mus. Na:
Hyla ocapia Andersson, 1938. Ark. Zool.,30A(23):L9.
rypE(s): Syntypes: NHRM L873 (7 specimens).
TYPE LocAltrv: Puka Khara, Bolivia.
DISTRIBUTIow: Eastern slopes of Andes in Bolivia.
coMMENT: In the HYla ocaPia grouP.
Hyla olioeiraiBokermann, 1963. Atas Soc. Biol. Rio de Janeiro,T{2):6.

rYPE(s): Holotype: WCAB 1,2306.
TypE LocALITY: "Marac6s, Bahia, Brasil".
DISTRIBUTIoN: Xeric areas of Bahia, Btazll.
coMMENT: Related to Hyla decipiens, according to the original description.
Hyla ornatissima Noble, L923. Zoologica, New York, 3:291.

rYPE(s): Holotype: AMNH L3491.
TypE LocAtltv: Meamu, Mazaruni River, Guyana.
DISTRIBUTIoN: Guianas and Amap6, Brazil.
ANURA: HYLTDAE: HYLINAE t45
C9MMENT: In the Hyla granosa SrouP; see Hoogmoed, L979, Zool. Verh., Leiden,
172:L5
Hyla pachyderma Taylor, L942. Univ. Kansas Sci. Bull., 28:308'

rYPE(s): Holotype: USNM LL5029.
7'Pan de Olla
TypE LocAlrry; [in Puebla], south of TezuitlAn, Veracruz", Mexico
C9MMENT: In the Hylabistincta group; see Due1lman,1970, Monogr. Mus. Nat'
Hist. Univ. Kansas, l:473-475.
Hyta palliata Cope,1863. Proc. Acad. Nat. Sci. Philadelphia, L5:47.

ivnr(s): Formerly Smithsonian Museum (USNM) 6225; now lost.
TYPE LoCALITY: "Paragtay" .

CoMMENT: This name has not been assigned to any known species (WED)'
Hyla palmeri Boulenger, 1908. Ann. Mag. Nat. Hist., (8)2:515'

TypE LocArrrv, lim6nez, Departamento Valle, Colombia.
DrsrRrBUTrow: Central Panama and Pacific slopes of Colombia and Ecuador.
g9MMENT: In the Hyla bogotensis group; see Myers and Duellman, L982, Am. Mus
Novit.,2752:23.
Hyla pantosticta Duellman and Berger, 1982. Herpetologica,33:456.

TypE LocAllrv, b.5 km S Santa Barbara on road to La Bonita, 2650 m, Provincia
Carchi, Ecuador.
DrsrRrBUrroN: Upper Rio Chingual Vatley on eastern slopes of Andes in Colombia
and Ecuador.
coMMENr: In the Hyla larinopygion grouP.
Hyta pardalis Spix, 1824. Spec. Nov. Testud. Ran. Brasil.:34.

rvnr(s): Syntypes: ZSM 2499 (2 specimens), now losU see Hoogmoed and Gruber,
L983,Spixiana, Suppl., 9:366, for discussion.
TypE LocALITY: "Rio de Janeiro", Blazil.
DrsrRrBUTroN: Central and eastern Brazll to Sio Paulo; to Misiones, Argentina.
coMMENr: In the Hyla boans group; see Lutz, L973, Brazil. Spec. Hyla:26-29, and
Cei, 1980, Monit. Zoo!.I1aI.,N.S., Monogr., 2:459-462. See comment under
Hyla biobeba.
Hyla paraiceps Boulenger, 1882. Cat. Batr. Sal. Brit' Mus':393'

rYPE(s): Holotype: BM L947 .2.13.5.
TypE LoCALITY, "Sar ayaclt" , Provincia Pastaza, Ecuador'
DISTRIBUTIOTV:Upper Amazon Basin in Brazil, Colombia, Ecuador, and Peru'
coMMENr: In thi^Hyla paraiceps group; see Duellman and Crump, L974, Occas. Pap
Hyla pauiniensis Heyer, 7977. Pap. Avulsos Zoo1., Sio Paulo, 31:L45.
TYPE LocAlrrv: Boca do Pauini, Amazonas,Brazil'
DrsrRrBUrroN: Known only from the type locality in central Amazon Basin.
coMMENr: In the Hyla paiaiceps group according to the original description'
Hyla pellita Duellman, L968. Univ. Kansas Publ. Mus. Nat. Hist. , 17:568'
TypE LocAury' ''33 kilometers north of San Gabriel Mixtepec, Oaxaca, M6xico,
DrsrRrBUTroN: Pacific slopes of the Sierra Madre del Sur in Oaxaca, Mexico.
coMMENT: In the Hyla mixomaculata group; see Duellman, L970, Monogr. Mus. Nat.
Hist. Univ. Kansas, I:421-423.
Hyla pellucensWerner, 1901. Verh. Zool. Bot. Ges. Wien, 50:600.

TypE LocALITy: "Palmar (100 m., westliche der Anden", Ecuador; Duellman, L977,
Das Tierreich, g5:82, thought this might be Palmar on Rio Chila, Provincia
Manabi, Ecuador.
DISTRIBUTIoN: Pacific lowlands of Colombia and Ecuador.
coMMENr: See Duellman, L971, Herpetologica, 27:212-227, and Duellman/ L974,
Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 27:L-27.
Hyla pentheter Adler, 1965. Occas. Pap. Mus. Zool. Univ. Michigan, 642:5.
rvrr(s): Holotype: UMMZ 125381.
rypE LocALrry: "37 km N of San Gabriel Mixtepec (about 100 km airline SSW of
Oaxaca de Ju6rez), 7700 m", Oaxaca, Mexico.
DrsrRrBUTrow: Pacific slopes of the Sierra Madre del Sur in Oaxaca and Guerrero,
Mexico.
coMMENT: In the Hyla bistincta gtoup. See Duellman, L974, Monogr. Mus. Nat.
Hist. Univ. Kansas, l:462-466.
Hyla phlebodes Stejneger/ L906. Proc. U.S. Natl. Mus., 30:817.

rvrr(s): Holotype: USNM 2997.
TypE LocAt-rry: "San Carlos" (Llanuras de San Carlos), Provincia Alajuela, Costa
Rica.
DrsrRrBUTroN:Caribbean lowlands from southern Nicaragua to Panama and Pacific
Iowlands of eastern Panama and northwestern Colombia.
coMMENT: In the Hyla microcephala group, see Duellman, L970, Monogr. Mus. Nat.
Hist. Univ. Kansas, l:220.
Hyla phyllognatha Melin, 1941. Goteborgs K. Vetensk. Vitterh. Samh. Handl.,

(B)1(a):30.
rvrr(s): Holotype: NHMG 474.
TypE LocAltrv: Roque, Departamento San Martin, Peru.
DrsrRrBUTIoN: Amazonian slopes of Andes in Colombia, Ecuador, and Peru.
coMMENr: In the Hyla bogotensis group; see Duellman, L972, Occas. Pap. Mus. Nat
Hyla picadoi Dunn, L937. Copeia, 1937:L64.

rvrr(s): Holotype: MCZ L6002.
TypE LocAltrv: Volc6n Barba, Provincia Heredia, Costa Rica.
DISTRIBUTIoN: Mountains of Costa Rica and western Panama.
coMMENr: In the Hyla zeteki grotp; see Duellman, L970, Monogr. Mus. Nat. Hist
Univ. Kansas, L:3L9-323.
Hyla piceigularis Ruiz-C. and Lynch, L982. Caldasia, 13:659.
rvrE(s): Holotype: ICN 5180.
TypE LocAr-rry: "Cabeceras del rio Luisito (tributary del rio Oibita), Municipio El
Encino, Departamento Santander, vertiente oriental de la Cordillera,
Colombia: altura 1.750 m.s.n.m., Long. 73o L7' L7' W de Greenwich, Lat. 6o 5'
54',N".
DrsrRrBUrroN: Known only from the type locality in Cordillera Oriental of Andes.
coMMENr: In the Hyla bogotensis group according to the original description.
Hyla picta (Gtinther, 1901). Biol. Cent. Am., Rept. Batr.:286.

oRIGINAL NAME: Hylella picta.
rvnE(s): Holotype: BM 1947 .2.22.62.
TypE LocAltrv: ]alapa, Veracruz, Mexico.
DISTRIBUTIoITI: Atlantic lowlands of Mexico, Guatemala, and Honduras
coMMENT: In the Hyla picta group; see Duellman, L970, Monogr. Mus. Nat. Hist.
Hyla pictipes Cope, L876. J. Acad. Nat. Sci. Philadelphia, (2)8:106.
oRIGINAL NAME: Hyla punctariola pictipes.
rYPE(s): Syntypes: USNM 3063L, 30652.
TypE LocAlIrv: Pico Blanco, Provincia Lim6n, Costa Rica.
DTSTRIBUTION: Mountains of Costa Rica and western Panama.
g9MMENT: Sole member of the Hyla pictipes group; see Duellman, 1970, Monogr.
Mus. Nat. Hist. Univ. Kansas, l:295.
Hyla picturataBoulenger, 1882. Ann. Mag. Nat. Hist., (7)3:276.

TvPE LocALITY: "Paramba, [Province Imbabura,] N. W. Ecuador".
DrsrRrBUTroN: Pacific lowlands of southwestern Colombia and northwestern
Ecuador.
coMMENr: Questionably in the Hyla geographica grouP (WED)'
Hyla pinima Bokermann and sazima, L974"L973". Rev. Brasil. Biol., 33:525.
TypE LocAurv, kilometer 132, Serra do Cip6, ]aboticatubas, Minas Gerais, Btazll.
DISTRIBUTIoN: Minas Gerais and Rio Grande do Sul, Brazll.
g9MMENT: Without known affinities, according to the original description.
Hyla pinorum Taylor, L937. Proc. Biol. Soc. Washington, 50:46.

rypE(s): Holotype: UIMNH 25049 (formerly EHT-HMS 5972).
TypE LocAlrry,
;'in pines on Mexico-Acapulco highway between kilometers 350
and 351, near i spring known as Agua del Obispo, between the towns of
Rincon and Cajones", Guerrero, Mexico.
DrsrRrBUrroN: Pacific slopes of Sierra Madre del Sur in southern Mexico.
coMMENT: In the Hyla melanomma group; see Duellman, L970, Monogr. Mus. Nat.
Hyla platydactyla Boulenget,1905. Ann. Mag. Nat. Hist., (7)L6:183.
rypE(s): Holotype: BM L947 .2.I3.L4.
TypE LocALITY: "Merida, Andes of Venezuela".
DISTRIBUTIoN: M6rida Andes in western Venezuela.
69MMENT: In the Hyla bogotensis group; see Duellman, L972, Occas. Pap. Mus- Nat.
Hist. Univ. Kansas, it't-3t. Differences in calls, morphology, and larval
features in various parts of the M6rida Andes suggest that more than one
species may ,be recognizable and that the nominal taxa Hyla jahni and Hyla
paramica Rivero, 196I, Bull. Mus. Comp. Zool., L26:LI2-L13, may be
recognizable species in the Hyla bogotensis group (WED_); these names were
synoinymized with Hyla platydactyla by Duellman, 1972, Occas. Pap. Mus. Nat
Hist. Unirr. Kansas, 1i:1-31. This synonymy was discussed and supported by
Ruiz and Lynch, 1982, Caldasia, L3:655-657.
Hyla plicafa Brocchi, L877. Bull. Soc. Philomath. Paris, (7)L:L26'

TYPE LoCALITY: "Mexique".
High mountains of Sierra Madre Oriental and Cordillera Volc6nica
DrSrRrBUrroN:
in southern Mexico.
coMMENr: In the Hyla eximia group; see Duellman, L970, Monogr. Mus. Nat. Hist.
Univ. Kansas, l:495-497 .
Hyla polytaenia Cope, L869. Proc. Am. Philos. Soc., lL:L64'
rvnn(s): Syntypes: MCZ 1544 (2 specimens).
TYPF LOCALITY:,,BTASiI,,.
DrsrRrBUTroN: Southeastern Brazil from Minas Gerais to Santa Catarina; eastern
Paraguay and northestern Argentina.
L48 AMPHIBIAN SPECIES OF THE WORLD
coMMENr: In the Hyla polytaenia grortp. See Cei, 1980, Monit. Zool.Ital., N.S.,
Monogr., 2:479-480.
Hyla praestans Dtellman and Trueb, 1983. /n Rhodin and Miyata (eds.), Adv.
Herpetol. Evol. Biol.:43.
TypE LocALITy:"Parque Arqueol6gica San Agustin, 3 km southwest of the village
of San Agustin, Departamento de Huila, Colombia (L'53' N, 76o 16' W),
1750 m".
of Cordillera Central and upper Rio Magdalena
DISTRIBUTIoN: Eastern slopes
Valley, Colombia.
coMMENr: In the Hyla columbiana group.
Hyla prasina Brrmeister, 1856. Erldut. Fauna Brasil.:106.

rvnr(s): Holotype: Zllv{B 4675.
TypE LocALItv: Nova Friburgo, Rio de Janeiro, Brazll.
DISTRIBUTIow: Moderate elevations in southeastern Brazil (Minas Gerais, Rio de
janeiro, and Sio Paulo).
coMMENr: In the Hyla pulchella group, according to Litz, L973, Brazil. Spec.
Hyla:83.
Hyla pseudopseudis Miranda-Ribeiro, 1937. Campo, Rio de ]aneiro, L937:68.

rYPE(s): Holotype: MN 579.
TypE LocArny: Veadeiros, Goi6s, Brazll.
DISTRIBUTIoN: Interior highlands of Brazil.
Hyla pseudopuma Gtinther, 1901. Biol. Cent.Am., Rept. Batr;274.

rypE(s): Syntypes: BM 96.10.8.61-67, 1902.L.28.25-26.
TypE LocAltry: La Palma, Provincia San ]os6, Costa Rica.
coMMENT: In the Hyla pseudopuma group; two subspecies were recognized by
Duellman, L970, Monogr. Mus. Nat. Hist. Univ. Kansas, L:263.
Hyla pugnax Schmidt, L857. Sitzungsber. Akad. Wiss. Wien, Math. Naturwiss. Kl.,
24:LL.
rvnr(s): Holotype: KM 1009.
TypE LocALIty: "Chiriqui-Flusse unweit Bocca del toro", Panama.
DISTRIBUTIoN: Central Panama, Caribbean lowlands and valleys of the Rio Cauca
and Rio Magdalena in Colombia.
coMMENr: In the Hyla boans group; this species was shown to be distinct from
Hyla crepitans by Kluge, L979, Occas. Pap. Mus. Zool. Univ. Michigan, 588:
1-24.
Hyla pulchellaDum{ril and Bibron, 1841. Erp. G6n., 8:588.

rvrr(s): Syntypes: MNHNP 163 and 4836.
TYPE LocALITY: "Mont6vid6o", Uruguay.
DISTRIBUTIow: Southeastern Brazll, Uruguay, Bolivia, southern Paraguay, and
northern Argentina.
coMMENT: In the Hyla pulchella group. Some of the five recognized subspecies mav
be specificallydistinc| see Barrio, 1965, Physis, Buenos Aires, 25:115-128, and
Lutz, L973, Brazll. Spec. Hyla:74-81. See also Cei, 1980, Monit. ZooI.Ital., N.S.,
Monogr., 2:469-476. See comment under Hyla cymbalum.
Hyla pulchrilineata Cope, L869. Proc. Acad. Nat. Sci. Philadelphia, 1L:163.
rvrE(s): Holotype: ANSP L4495.
TypE LocALITy: "East St. Domingo", Hispaniola, West Indies.
DrsrRrBUTroN: Hispaniola, West Indies.
coMMENT: In the Hyla pulchrilineata group; see Trueb and Tyler, 1974, Occas. Pap
Mus. Nat. Hist. Univ. Kansas, 24:L-60.
ANURA: HYLIDAE: HYLINAE t49
REVIEWERS: Albert Schwartz (AS); Richard Thomas (RT).
Hyla punctata (Schneider, 1799). Hist. Amph., L:L70.

oRTGTNAL NAME: Calamita punctatus.
rvnE(s): Syntypes: NHRM 155 (2 specimens).
DrsrRrBUTroN:The Amazon Basin; upper Orinoco Basin; central Brazll; Chaco of
Paraguay, and Argentina; Guianas, northern Colombia and Trinidad.
coMMENT: Hyla punctata rubrolineata Lutz, L95L, Mem. Inst. Oswaldo Cruz, Rio de
]aneiro, 47:307 (reviewed by Cei, 1980, Monit. Zool.Ital., N.S., Monogr., 2:
482-484) may be a recognizable species (WED), as is Hyla hobbsi (see Pyburn,
L978, Proc. Biol. Soc. Washington,9L:L23); both of these were considered to
be synonyms of Hyla punctata by Duellman, 1974, Occas. Pap. Mus. Nat. Hist
Univ. Kansas, 27:l-27. See Hoogmoed and Gruber, L983, Spixiana, Suppl., 9:
365, for discussion of some of the synonyms.
Hyla quadrilineata (Schneider, L799). Hist. Amph., L:L69.

oRTGINAL NAME: Calamita quadrilineatus.
rvnr(s): Unknown.
TYPE LocAlIrv: Unknown.
DISTRIBUTIOT.I: UNKNOWN.
coMMENT: This name has not been assigned to any known population (WED)
Hyla raniceps (Cope, L862). Proc. Acad. Nat. Sci. Philadelphia, L4:353.
oRIGINAL NAME: Hypsiboas raniceps.
rvnr(s): Syntypes: USNM 5403 (3 specimens), 5408, L2L60, L2L72 (2 specimens).
TYPE LoCALITY: "Paraguay".
DISTRTBUTToN: French Guiana, eastern Brazll, Paraguay, northern Argentina, and
eastern Bolivia.
coMMENr: In the Hyla albopunctata group. See Cei, 1980, Monit. ZooI.Ital., N.S.,
Monogr., 2:463-466.
Hyla regilla Barrd and Girard, L852. Proc. Acad. Nat. Sci. Philadelphia, 6:L74.
rvnr(s): Syntypes: USNM 9182 (Puget Sound), USNM 15409 (Sacramento River);
USNM 9182 designated lectotype by ]ameson, Mackey, and Richmond, L966,
Proc. California Acad. Sci., 33:553.
TypE LocAlrrv: Puget Sound, Washington, and Sacramento River, California, USA;
restricted to "Sacramento County, California", USA, by Schmidt, 1953, Cat.
Am. Amph. Rept., Ed. 6:72; restricted to Fort Vancouver, Washington, USA,
by lectotype designation of ]ameson, Mackey, and Richmond, L966, Proc.
California Acad. Sci., 33:553.
DrsrRrBUTroN: Western North America from southern British Columbia (Canada)
and western Montana (USA) to southern Baja California (Mexico).
coMMENT: In the Hyla eximia group; see Jameson, Mackey, and Richmond, L966,
Proc. California Acad. Sci. 33:551-620, and Duellman, 1970, Monogr. Mus.
Nat. Hist. Univ. Kansas, L:484-493i seven subspecies are recognized.
Hyla rhodopepla Gtinther, 1859 "1858". Cat. Batr. Sal. Coll. Brit. Mus.:112.
rypE(s): Holotype : BM L947 .2.23.53.
TYPE LocALITY: "Andes of Ecuador".
DrsrRrBUTroN: Upper Amazon Basin from southern Colombia to Bolivia.
coMMENr: In the Hyla microcephala group; see Duellman, L972, Herpetologica, 28
369-375.
Hyla riaeroi Cochran and Goin, L970. Bull. U.S. Natl. Mus., 288:284.
rYPE(s): Holotype: CM 37433.
TypE LocALITy: "Leticia, Amazonas, Colombia."
DrsrRrBUTroN: Upper Amazon Basin in Colombia, Ecuador, and western Brazil;
probably in adjacent Peru.
coMMENr: In the Hyla minima grottp; see Duellman, 1982, Amphibia-Reptilia, 3:

159.
Hyla riaularis Taylor, 1952. Univ. Kansas Sci' Bull', 35:847 '
rYPE(s): HolotyPe: KU 28L97.
TypE LocAlrry: "American Cinchona Plantation (Isla Bonita), eastern
slope of Volc6n
PoAs, [Provincia Heredia,] Costa Rica, elevation 5,500 ft'"
DTSTRIBUTION, Mo.,t tains of Costa Rica and western Panama.
C9MMENT: In the Hyla rioularis gtottp; see Duellman, L970, Monogr'
Mus' Nat'
Hyla robertmertensi Taylor, L937. Proc. Biol. Soc. Washington,50:43.
"rvnE(s):
Holotype: ruNH 100096 (formerly EHT-HMS 2270).
TypE LocALITY: "near Tapachula, Chiapas", Mexico'
DrsrRrBUTroN: Pacific lo*lands from eistern Oaxaca, Mexico, to El SalvadoU
Cintalapa Valley in Chiapas, Mexico.
coMMENr: In the Hila microciphala group; see Duellman, 1970, Monogr. Mus. Nat'
Hist. Univ. Kansas, L:2L7.
Hyla robertsorum Taylor, L940. Univ. Kansas Sci. Bull., 26:393.

rvrE(s): Holotype: FMNH LOOL}A (formerly EHT-HMS 16264).
TypE LocAtlrv' "El Chico National Park, Hidalgo", Mexico'
DrsrRrBUrroN: Sierra Madre Oriental in eastern Mexico (Puebla and Hidalgo).
ggMMENT: In the Hylabistincta Btoltp; see Duellman, L970, Monogr' Mus. Nat.
Hyla rodriguezi Riveto, L968. Breviora, 307:1.
TypE LOCArrrr, i'uro del Danto, Region de La Escalera, 1400 meters, road from El
Dorado to Santa Elena de Uair6n, Estado Bolivar, Venezuela.
La Escalera region and Gran Sabana, Bolivar, Venezuela'
DISTRIBUTIOw:
coMMENr: This species probibly belongs in Osteocephalus (WED and MSH).
Hyla roeschmanni DeGrys, 1938. Zool. Anz., L24:3L5'

ryPE(s): Formerly in ZM}{; destroyed.
TYPE LocAlIrY: Beni, Bolivia.
coMMENr: This name has not been asiociated with any known population (WED).
Hyla rosenbergiBoulenger, 1898. Proc. Zool. soc. London, L898:123.
ryPE(s): Syntypes: BM L947 .2.L2.7 L-7 5.
TYPE LocALITY: "Cachab6" (:Cachabi), Provincia Esmeraldas' Ecuador'
DrsrRrBUTroN: pacific lowlands from southern Costa Rica to northwestern Ecuador.
g9MMENT: In the Hylaboans group; see Kluge, L979, Occas. Pap. Mus. Zool. Univ.
Michigan, 688:L-24.
Hyla rossalleni Goin, L959. Copeia, 1959:340'
rYPE(s): Holotype: FSM 8501.
TypE LocAurv: Leticia, Comisaria Amazonas, Colombia'
DrsrRrBUTroN: Upper Amazon Basin in Ecuador, Peru, southeastern Colombia, and
northeastern Bolivi a; Btazilian Amazonia.
69MMENT : Hyla rossalleni is a replacement name f.or Hyla alleni Goin, L957 , J.
Washington Acad. Sci.,47:-60. In the Hyla minima StortP; see Duellman, 1982,
Amphibia-RePtilia, 3: 159.
Hyla rubicundula Reinhardt and Ltitken, 1862. Vidensk. Medd. Dansk Naturhist.
Foren., 3:L97.
rypE(s): Syntypes: NHMW t65LL, Zill{UC L440-4L'
TvPE LocAurv, Lagoa Santa, Minas Gerais, Btazll'
DrsrRrBuTror.r: Sotjhern Pari to central Minas Gerais and Sio Paulo, Brazil, and
eastern ParaguaY'
coMMENT: In the Hyla rubicundula group.

Hyla rubracyla Cochran and Goin, L970. Bull. U.S. Natl. Mus., 288:229.
rYPE(s): Holotype: USNM L57820.
TypE LocAt-rry: "Rio Calima, near
C6rdoba, [Departamento] Valle, Colombia".
DISTRIBUTIoN: Pacific lowlands of
Colombia.
coMMENT: In the Hyla albomarginata group; see Duellman, L974, Occas. Pap. Mus
Nat. Hist. Univ. Kansas, 27:13.
Hyla rufioculis Taylor, L952. Univ. Kansas Sci. Bull., 35:827.

TYPE LocALITY: "Isla Bonita (American Cinchona Plantation), eastern slope of
Volcin PoAs, fProvincia Heredia,] Costa Rica".
DISTRIBUTToN: Mountains of Costa Rica.
coMMENT: In the Hyla uranochroa group; see Duellman, L970, Monogr. Mus. Nat.
Hyla rufitela Fouquette, 196L. Fieldiana: Zool.,39:595.

rYPE(s): Holotype: FMNH 13053.
TypE LocArrry: Barro Colorado Island, Canal Zone, Panama.
DISTRIBUTIoN: Caribbean lowlands from Nicaragua to Panama; Golfo Duice region
of Costa Rica.
COMMENT: In the Hyla albomarginata group; see Duellman, L970, Monogr. Mus. Nat
Hyla sabrina Caldwell, 1974. occas. Pap. Mus. Nat. Hist. Univ. Kansas, 23:L2.
rvrE(s): Holotype: KU 137086.
TypE LocAury: 16.6 km south of Vista Hermosa, 2020 m, Oaxaca, Mexico.
DISTRIBUTIoN: Moderate elevations on north slopes of Sierra de JuArez, Oaxaca,
Mexico.
coMMENr: In the Hyla bistincta group according to the original description.
Hyla saloadorensis Mertens, 1952. Senckenb. Biol., 33 169.

rYPE(s): Holotype: SMF 43045.
TYPE LocAurv: Hacienda San ]os6, Sierra MetapAn, Departamento Santa Ana, El
Salvador.
DISTRIBUTIoN: Pacific slopes of EI Salvador and Honduras.
coMMENT: In the Hyla salaadorensis group; see Duellman, L970, Monogr. Mus. Nat.
Hist. Univ. Kansas, L:337-34L.
Hyla sanchiangensis Pope, Lg2g. Am. Mus. Novit., 352:2.

TypE LocALITy: "San Chiang, Chungan Hsien, northwestern Fukien [:Fujian]
Province, China, 3000-3500 feet altitude".
DISTRIBUTIoN: Northwestern Fujian, Guangxi, Guizhou, Anhui, Zhejiang, Hunan,
and Jiangxi, China.
coMMENT: In the Hyla arborea gtoup; closely allied to Hyla chinensis, according to
the original description. See Liu and Hu, L961, Tailless Amph. China:135, and
Ma et al., L982, Nat. Hist., Shanghai, l:37.
REVIEWERS: Leo J. Borkin (LIB); Shuqin Hu (SH); Masafumi Matsui (MM); Ermi
Zhao (EZ).
Hyla sarampiona Ruiz-C. and Lynch, 1982. Caldasia, 13:664.

rYPE(s): Holotype: ICN 7440.
TYPE LocALITy: "Quebrada Sopladero, Parque Nacional Natural de Munchique, 33
km NNE of La Uribe, flanco occidental de Ia Cordillera Occidental, 2.190 m. s.
n. m., Departamento del Cauca, Colombia".
DISTRIBUTIoN: Known only from the type locality on the western slopes of
Cordillera Occidental of Andes.
ir*
152 AMPHIBTAN SPECIES OF THE WORLD
coMMENr: In the Hyla larinopygion group according to original description.
Hyla sarayacuensis Shreve, 1935. Occas. Pap. Boston Soc. Nat. Hist.,8:215.
oRIGINAL NAME: Hyla leucophyllat a saray acuensis.
rYPE(s): Holotype: MCZ L9729.
TypE LocAtlrv: Sarayacu, Provincia Pastaza, Ecuador.
DrsrRrBUTroN: Uppei Amazon Basin in Colombia, Ecuador, Peru, and Brazil.
coMMENT: In the Hyla leucophyllata group; see Duellman, L974, Occas. Pap. Mus
Hyla sarda De Betta, 1853. Mem. Accad. Agric. Commerc. Arti, Verona, 35:284.
rvnr(s): Unknown.
rYPE LocArtrv: Sardinia [Italy].
DTSTRIBUTIoN: Sardinia (Italy), Corsica (France), and nearby islands in the
Tyrrhenian Sea.
coryruENr: In the Hyla arborea group. Elevated from status as a subspecies of Hyla
arboreaby Lanza,1983, Lav. Soc. Ital. Biogeogr., N.S., 8:730.
Hyla sartori Smith, 1951. Herpetologica, 7 :L86.

oRIGINAL NAME: Hyla miuocephala sartori.
rYPE(s): Holotype: UIMNH 20934.
TypE LocAlrry: 1 mi. north of Organos, south of El Treinta, Guerrero, Mexico.
DrsrRrBUTroN:Pacific slopes of southwestern Mexico (]alisco to Oaxaca).
coMMENT: In the Hyla microcephala group; see Duellman, L970, Monogr. Mus. Nat
Hyla saaignyi Audouin, 1827 . Descr. i,gypte, 1:183.

rYPE(s): Unknown.
TYPE LocAr-ITv: Western Syria.
DrsrRrBUTron: Syria and southern Turkey to northern and western Iran, and
southern Armenia and Azerbaijan in USSR; southern Arabian Peninsula.
coMMENr: In the Hyla arborea group. This species was regarded as distinct from
Hyla arboreaby Schneider, L974, Oecologia, Berlin, l4:99-LL0.
REVIEwERS: Steven C. Anderson (SCA); Leo ]. Borkin (LIB).
Hyla sazimai Cardoso and Andrade, 1983 "L982". Rev. Brasil. Biol., 42:589.
rvrr(s): Holotype: MN 4149.
TypE LocAt-rry:"Chapadio da Zagaia, Parque Nacional Serra da Canastra, Sio
Roque de Minas, Minas Gerais, Brasil (46" 50' W 20' 10' S; aPProx. 1350 m
alt.)".
Hyla schubartiBokermann, L963. Rev. Brasil. Biol., 23:249.

TypE LocALITy: "Rond6nia, Territ6rio [now Estado] de Rond6r.ia", Btazil.
DISTRIBUTIoN: Brazilian Amazonia.
Hyla secedens B. Lutz, 1963. Copeia, L963:561.

TYPE LocAtItv: Barro Branco, Rio de Janeiro, Btazll.
DISTRIBUTIoN: Serra dos Orgios, southeastern Brazil
Hyla semiguttata A. Lutz, L925. C. R. S6ances Soc. Biol., Paris, 93:211.

TYPE LocAlrrv: Sio Bento do Sul, Santa Catarina, Brazil.
DrsrRrBUTroN: Highlands of extreme southeastern Brazil (Rio Grande do Sul and
Santa Catarina) and extreme northeastern Argentina (Misiones).
J
coMMENT: See Cei, L980, Monit. Zool.Ital., N.S., Monogr., 2:477.
Hyla senicula Cope,1868. Proc. Acad. Nat. Sci. Philadelphia, 20:111.

rvnr(s): Unknown.
TypE LocALrry: Corcovado, Rio de Janeiro, BrazII; Lutz, L973, Brazll. Spec. Hyla:88,
rendered the type locality as "Laranjeiras, a residential part of Rio de ]aneiro,
below Corcovado Mountain".
DrsrRrBUTrow: Coastal region of southeastern Brazll (Espirito Santo and Rio de
Ianeiro).
coMMENT: In the Hyla marmorata group. Hyla senicula is a replacement name for
Hyla marmorata Burmeister, 1856, Erliiut. Fauna Brasil.:93. See Bokermann,
L964, Senckenb. Biol., 45:252. Lutz, L977 , Brazil. Spec. Hyla:88-93, regarded
Hyla aueana, Hyla melanargyrea, and Hyla noaaisi as subspecies of. Hyla senicula.
Heyer, L977, Pap. Avulsos Zoo1., Sio Paulo, 3l:l4l-L62, re-elevated Hyla
acreana to species status. Caramaschi and |im, 1983, Rev. Brasil. Biol., 43:L98,
disputed the subspecific status of Hyla melanargyrea and Hyla noaaisi, and
elevated them to species status.
Hyla sibleszi Rivero, L97L. Caribb. J. Sci., LL:L82.

rvnr(s): Holotype: UPRM 3L77.
TypE LocArrry: Paso del Danto, La Escalera, between EI Dorado and Santa Elena de
Uair6n, 1300-1400 m., Serrania de Lema, Estado Bolivar, Venezuela.
DrsrRrBUTroN: Moderate elevations in the southwestern Venezuela and adjacent
Guyana.
coMMENT: In the Hyla granosa group; see Hoogmoed, 1978, Zool. Verh., Leiden,
L72:23.
Hyla simpler Boettger, 1901. Ber. Senckenb. Naturforsch. Ges., 1901:53.

oRIGINAL NAME: Hyla chinensis var. simplex.
rYPE(s): Holotype: SI[v4F 2626.
TypE LocArtrv: Phuc-Son, Annam, Vietnam.
DrsrRrBUrroN: Vietnam (Annam and Tonkin) and southern China (Guangxi,
Zhejiang, Guangdong and Hainan I.).
coMMENT: In the Hyla arboreagroup; see Pope, L93L, Bull. Am. Mus. Nat. Hist.,61
477-480, and Liu and Hu, L96L, Tailless Amph. China:128-130.
REVIEwERS: Shuqin Hu (SH); Masafumi Matsui (MM); Ermi Zhao (EZ).
Hyla siopela Duellman,1968. Univ. Kansas Publ. Mus. Nat. Hist., L7:570.
TypE LocALrry: "d small stream on west slope of Cofre de Perote,Yeractvz,
M6xico, elevation 2500-2550 meters".
DrsrRrBUTrow: Sierra Madre Oriental in eastern Mexico (Oaxaca and Veracruz).
coMMENr: In the Hyla bistincta group; see Duellman, 1970, Monogr. Mus. Nat.
Hyla smaragdina Taylor, 1940. Copeia, 1940:18.

rypE(s): Holotype: FMNH 100009 (formerly EHT-HMS L7534).
TypE LocAt-rry: "on a mountain side rising from eastern end of Lake Chapala,6
km. east of Cojumatl6n, Michoac6n", Mexico.
DrsrRrBUTrow:Pacific slopes of western Mexico from Sinaloa to Morelos.
coMMENT: In the Hyla sumichrasti group; see Duellman, L970, Monogr. Mus. Nat.
Hist. Univ. Kansas, L:413-4L6.
Hyla smithri Boulenger, 1901. Zool. Rec., 38:33.

rvrr(s): Syntypes: BM L947 .2.L2.7 6-82.
TypE LocALltv: Cuernavaca, Morelos, Mexico.
DrsrRrBUTrow: Pacific iowlands of Mexico from northern Sinaloa to southern
Oaxaca and in the Balsas Depression from Michoac6n to Puebla.
coMMENT: Hyla smithii is a replacement name f.or Hyla nana Ginther, L901, Biol.
Cent. Am., Rept. Batr.:263. In the Hyla picta grouP; see Duellman, 1970,
Hyla soaresi Caramaschi and ]im, 1983. Rev. Brasil. Biol., 43:195.
rYPE(s): Holotype: J. Iim 5740.
TypE LocALIrv: Picos (approx. 07" 05' S-41'30' W), Piaui, Btazll.
g9MMENT: In the Hyla marmorata group, according to the original description, and
related to Hyla senicula, Hyla melanargyrea, and Hyla nooaisi.
Hyla squirellaBosc,1800. In Daudin, Hist. Nat' Quadrup. Ovip., Livr. 1:9.

rYPE(s): Unknown.
TypE LocALrry: "La Caroline"; restricted to Charleston, South Carolina, USA, by
Harper, L940, Am. Midl. Nat., 23:692-723.
DrsrRrBUTroN: Coastal Plain and Mississippi Valley of southeastern North America;
introduced on Grand Bahama I., Bahama Is.
coMMENr: Reviewed by Martof., L975, Cat. Am. A-Pl. Rept., L68.1-2-
Hyla steinbachiBoulenger, 1905. Ann. Mag. Nat. Hist., (7)16:182.

rypE(s): Syntypes: BM 1947 -2.L3.6L-63.
TypE LocAt-rry: "Province Sara, Departament Santa Cruz de la Sierra, Bolivia".
Hyla subocularis Dunn, L934. Am. Mus. Novit., 747:2.

rYPE(s): Holotype: AMNH 4LlL7.
TypE LocAlrry: Rio Chucunaque at the first creek above the Rio Tuquesa, Provincia
Dari6n, Panama.
DISTRIBUTIoN: Eastern Panama and northern Colombia.
coMMENr: In the Hyla paraiceps grotp; see Duellman and Crump, 1974, Occas. Pap.
Mus. Nat. Hist. Univ. Kansas, 23 1-40.
Hyla sumichrasti (Brocchi, 1879). Bull. Soc. Philomath. Paris, (7)3:20.

oRIGINAL NAME: Exerodonta sumichrasti.
rvnr(s): Formerly in MNHNP; now lost.
TypE LocALIrv: Santa Efigenia, Oaxaca, Mexico.
DrsrRrBUrroN: Pacific slopes of southern Mexico (Chiapas and Oaxaca) and Chiapan
highlands, Mexico.
coMMENr: In the Hyla sumichrasti group, see Duellman, L970, Monogr. Mus. Nat.
Hist. Univ. Kansas, L:409'4L3.
Hyla surinamensis Daudin, 1802. Hist. Nat. Rain. Gren. Ctap.:44.
rvnr(s): None designated; based on Seba, L734, Loc. Nat. Thes. Desc. Icon. Exp.:pl.
70, h9.4.
TYPE LoCALITY: "Surinam."
DISTRIBUTIoN: ? Surinam.
CoMMENT: This name has not been assigned to any known taxon (WED).
Hyla suweonensis Kuramoto, 1980. Copeia, 1980:102.

rvnr(s): Holotype: OMNH 6035.
TypE LocAurv: Office of Rural Development, Suweon, Korea.
DISTRIBUTIoN: South Korea.
coMMENr: In the Hyla arborea groltp according to the original description.
REVIEwERS: Leo J. Borkin (LlB); Shuqin Hu (SH); Masafumi Matsui (MM); Ermi
Zhao (EZ).
Hyla taeniopus Ginther, 1901. Biol. Cent. Am., Rept. Batr.:269.

TypE LocAurv: ]alapa, Veracruz, Mexico.
DISTRIBUTIoN: Atlantic slopes of Sierra Madre Oriental, Mexico.
.d
coMMENT: In the Hyla taeniopus group; see Duellman, L970, Monogr. Mus. Nat.
Hyla thorecfes Adler, 1965. Occas. Pap. Mus. Zool. Univ. Michigan, 642:L0.
rYPE(s): Holotype: UIII4MZ L25390.
rypE LocALITy: "about 37 km N of San Gabriel Mixtepec (about 100 km airline
SSW of Oaxaca de ]u6rez),
Oaxaca, 1700 m", Mexico.
DISTRIBUTIow:Pacific slopes of Sierra Madre del Sur, Oaxaca, Mexico.
coMMENT: In the Hyla hazelae grovp; see Duellman, 1970, Monogr. Mus. Nat. Hist.
Univ. Kansas, 1:388-391.
Hyla thysanota Duellman, L966. Univ. Kansas Publ. Mus. Nat. Hist., L7:259.
rvnE(s): Holotype: USNM 151080.
TypE LocALITy: "Cerro Mali, Darien Province, PanamA, (elevation 1265 meters)".
DISTRIBUTIoN: Type locality in the eastern part of the Serrania de Dari6n, Panama.
coMMENr: In the Hyla miliaria group; see Duellman, 1970, Monogr. Mus. Nat. Hist
Hyla tica Starrett, 1966. BuII. S. California Acad. Sci., 65:23.
rvrr(s): Holotype: UMMZ 122482.
rypE LocALtrv: Stream at 4500 feet on Volc6n Turrialba, Provincia Cartago, Costa
Rica.
DISTRIBUTIow: Mountains of Costa Rica and western Panama.
coMMENT: In the Hyla tica group; see Duellman, L970, Monogr. Mus. Nat. Hist.
Univ. Kansas, l:278-284.
Hyla tintinnabulum Melin, L941. Gdteborgs K. Vetensk. Vitterh. Samh. Handl.,
(B)1(a):2e.
rvnE(s): Syntypes: NHMG 473 (2 specimens).
TypE LocALITy: "Rio Uaup6s (some days' journey north of Ipanor6),fAmazonas,]
Btazil".
Hyla torrenticola Duellman and Altig, L978. Herpetologica, 34:181.
rvrr(s): Holotype: KU L69571.
rypE LocAtItv: "10.3 km west (by road) of EI Pepino, Departamento Putumayo,
Colombia,1,440 m, 01o 11' N, 76o 4l' W".
DISTRIBUTIoN: Eastern slopes of Andes in southern Colombia and northern
Ecuador.
coMMENr: In the Hyla bogotensis group according to the original description.
Hyla triangulum Giinther, L869 "1868". Proc. Zool. Soc. London, 1868:489.
rvrE(s): Holotype: BM 1,947 .2.23.88.
TYPE LocALrrY : "Brazll" .
DISTRIBUTIoN: Upper Amazon Basin in Colombia, Ecuador, Peru, and western

Brazil.
coMMENr: In the Hyla leucophyllata group; see Duellman, L974, Occas. Pap. Mus.
Nat. Hist. Univ. Kansas, 27:L9. Many color morphs of this species have been
recognized taxonomically (WED).
Hyla tritaeniata Bokermann, 1965. Rev. Brasil. Biol., 25:259

rvru(s): Holotype: WCAB 1621L.
TypE LocAltrv: Sio Vicente, CuiabA, Mato Grosso, Brazil
DISTRIBUTIow: Southern Mato Grosso, Btazil.
coMMENT: In the Hyla rubicundula group.
Hyla truncatalzecksohn, 1959. Rev. Brasil. Biol., 19:259.

rvnr(s): Holotype: El L07.
TypE LocAlIrv: Itaguai, Rio de janeiro, Brazll.
DISTRIBUTIoN: Coastal lowlands of Rio de ]aneiro, Brazil
coMMENT: In the Hyla truncata group, according to the original description.

Hyla trux Adler and Dennis, L972. Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 7:1.
TypE LocALrry: "Ll.4 km. (by road) southwest of Puerto del Gallo, Guerero,
M6xico (about 35 km airline northeast of Atoy6c de Alvarez), elevation 1985
meters".
DISTRIBUTIoN: Vicinity of Cerro Te6tepec, Sierra Madre del Sur, Guerrero, Mexico,
L760-2L20 m.
coMMENr: In the Hyla taeniopus group according to the original description.
Hyla tsinlingensis Liu and Hu, L966. ln F{u, Djao, and Liu, Acta Zool. Sinica, l8(1):7a.
rver(s): Holotype: CIB (formerly SIB-AC) 623149.
TypE LocALITy: "Ffou-tseng-tze, Couchih Hsien, Shensi [:Shaanxi], alt. L34L m.",
China.
DISTRIBUTIoN: Gansu, Sichuan, Shaanzi, and Hubei, China.
coMMENT: In the Hyla arborea gtoup and relatedto Hyla annectans, according to the
original description.
REVIEwERS: Leo |. Borkin (LIB); Shuqin Hu (SH); Masafumi Matsui (MM); Ermi
Zhao (EZ).
Hyla tuberculosa Boulenger, L882. Cat. Batr. Sal. Brit. Mus.:355.
rvnu(s): Holotype: BM 1947 .2. 13.34.
TypE LocALITy: "Canelos", Provincia Pastaza, Ecuador.
DISTRIBUTIoN: Upper Amazon Basin in western BrazTl, Colombia, Ecuador, and
Peru.
coMMENr: In the Hyla miliaria group; see Duellman, L974, Occas. Pap. Mus. Nat.
Hist. Univ. Kansas, 27:L-27.
Hyla uranochroa Cope, 1876. J. Acad. Nat. Sci. Philadelphia, (2)8:103.
rvnr(s): Holotype: USNM 2065L.
TypE LocALtry: Near Sipurio, Provincia Lim6n, Costa Rica.
coMMENT: In the Hyla uranochroa group; see Duellman, L970, Monogr. Mus. Nat.
Hyla oalancifer Fischein and Smith, 7956. Herpetologica, 12:78.
rvrE(s): Holotype: UIMNH 35398.
TypE LocAlrry: "VolcAn San Martin, 4500 ft., Yeracruz, Mexico".
DISTRIBUTIoITI: Cloud forests of the Sierra de los Tuxtlas, southern Veracruz,
Mexico.
coMMENT: In the Hyla miliaria group; see Duellman, L970, Monogr. Mus. Nat. Hist.
Univ. Kansas, l:342-346. Specimens previously allocated to this species from
the Sierra de las Minas, Guatemala, are being described by Larry David
Wilson (WED).
Hyla aasta Cope, L87L. Proc. Acad. Nat. Sci. Philadelphia,2S:2L9.
rYPE(s): Holotype: ANSP 2097.
rYPE LocALITY: "Near the city of Santo Domingo [Repriblica Dominicana,] W. I."
DrsrRrBUTroN: Hispaniola, West Indies.
coMMENr: In the Hyla boans group; see Trueb and Tyler, L974, Occas. Pap. Mus.
Nat. Hist. Univ. Kansas, 24:l-60.
REVIEwERS: Albert Schwartz (AS); Richard Thomas (RT).
Hyla aersicolor LeConte, 1825. Ann. Lyc. Nat. Hist. New York, L:28L.
rvnE(s): Unknown.
TypE LocALITy: "Northern States"; restricted to "vicinity of New York, New York".
USA, by Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6:73.
DISTRIBUTIoN: Northeastern USA and extreme southern Canada from eastern
North Dakota and eastern Missouri to southern Maine and Delaware; south-
central USA (eastern Oklahoma, Texas, and southwestern Louisiana).
ANURA: HYLIDAE: HYLINAE L57
coMMENr: In the Hyla uersicolor gro:up. Hyla aersicolor and Hyla chrysoscelis are
sibling species which can only be distinguished by call, karyotypes, or cell
volume. The actual range of each species is poorly understood.
Hyla oigilazs Solano, L97L. Acta Biol. Venezuelica, T:2L2.

rvnr(s): Holotype: MBUCV IV-6163.
TypE LocALrry: Km 20 on road between Coloncito and El Vigia, Estado Zulia,
Venezuela.
Maracaibo Basin, Venezuela.
DISTRIBUTIoN:
coMMENr: This species is a member of Ololygon (ELM).
Hyla airidifusca Laurenti, L768. Synops. Rept.:34.

oRTGINAL NAME: Hyla oiridi-fusca.
rYPE(s): Unknown.
DISTRIBUTION: UNKNOWN.
coMMENr: This name has not been assigned to any known species (WED)
Hyla walkeri Stuart, L954. Proc. Biol. Soc. Washington, 67:L65.

rvnE(s): Holotype: UIll.{MZ 1068L7.
TypE LocArrry: Aserradero San Lorenzo,12 km (airline) slightly east of north of
]a1apa, Jalapa, Guatemala.
DrsrRrBUTrow: Mountains of Chiapas, Mexico, and western and southeastern
Guatemala.
coMMENr: In the Hyla eximia group; see Duellman,1970, Monogr. Mus. Nat. Hist
Hyla wilderi Dunn, L925. Occas. Pap. Boston Soc. Nat. Hist., 5:161.
TypE LocAt-rry: "Moneague, Saint Ann Parish, |amaica; altitude 1200 feet".
coMMENr: In the Hyla wilderi grotp; see Trueb and Tyler, 1974, Occas. Pap. Mus.
Nat. Hist. Univ. Kansas, 24:l-60.
REVIEwERS: Albert Schwartz (AS); Richard Thomas (RT).
Hyla xanthosticta Duellman, L968. Univ. Kansas PubI. Mus. Nat. Hist., 17:562.
TypE LocAlrry: "south Fork of the Rio Las Vueltas on the south slope of VolcAn
Barba, near the northwest base of Cerro Chompipe, Heredia Province, Costa
Rica, elevatirin 2100 meters".
DrsrRrBuTrorv: Known only from the type locality in the central cordillera of Costa
Rica.
coMMENr: In the Hyla rioularis group; see Duellman, 1970, Monogr. Mus. Nat.
Hyla zeteki Gaige, L929. Occas. Pap. Mus. Zool. Univ. Michigan, 207:4.
rvnr(s): Holotype: U}'{MZ 63875.
rypE LocAlrry: Caldera Valley above Boquete, Provincia Chiriqui, Panama.
coMMENT: In the Hyla zetekl group; see Duellman, L970, Monogr. Mus. Nat. Hist
Limnaoedus Mittleman and List, 1953. Copeia, 1953:83.

TypE spECIrs: Hyla ocularis Bosc and Daudin, 1801.
coMMENr: A monotypic genus distinguished by paedomorphic osteological features;
posited by various authors to be related to Hyla, Auis, or Pseudacris; see Franz
and Chantell, 1978, Cat. Am. Amph. Rept., 209.L, for discussion.
Limnaoedus ocularis (Bosc and Daudin, 1801). /n Sonnini and Latreille, Hist. Nat.
Rept., 2:L87.
oRIGINAL NAME: Hyla ocularis.
rvrr(s): Unknown.
TypE LocALITy: "Carolina"; restricted to "vicinity of Charleston", South Carolina,
USA, by Harper, L939, Am. Midl. Nat., 22:L39-L44.
DrsrRrBUTroN: Coastal plain of southeastern Virginia to western and peninsular
Florida, USA.
coMMENr: Reviewed by Franz and Chantell, 1978, Cat. Am. Amph. Rept., 209.1,-2.
Nyctimantls Boulenger, 1882. Cat. Batr. Sal. Brit. Mus.:421.
TypE spECIns Nyctimantis rugiceps Boulenger, L882, by monotypy.
coMMENT: A monotypic genus of uncertain relationships; see Duellman and Trueb,
L976, Occas. Pap. Mus. Nat. Hist. Univ. Kansas,58:1-14.
Nyctimantis rugiceps Boulenger, L882. Cat. Batr. SaI. Brit. Mus.:422.

oRTGINAL NAME: Nyctimantis rugiceps.
rvrr(s): Syntypes: BM 80.12.5.L52 (Ecuador), 80.12.5.L63-L64 (3 specimens)
(Canelos).
TYPE LoCALITY: "Ecuado{' and "Canelos", Pastaza, Ecuador.
coMMENT: For discussion see Duellman and Trueb, L976, Occas. Pap. Mus. Nat.
Ololygon Fitzinger, 1843. Syst. Rept.:31.
rypE spECIns: Hyla strigilata Spix, L824, by monotypy.
DISTRIBUTIoN: Southern Mexico to Argentina and Uruguay.
coMMENT: Fouquette and Delahoussaye, 1977, ]. Herpetol., LL:387-396, removed the
former Hyla rubra group to this genus, discussed species groups (noted in the
individual species accounts) within Ololygon, and discussed intergeneric
relationships, all on the basis of sperm morphology. Osteological characters of
the adults and morphology of larvae support the recognition of Ololygon as a
monophyletic group (WED). A number of species currently placed in Hyla
probably belong in this genus; see comment under Hyla aigilares. See Le6n, 1969,
Univ. Kansas Publ. Mus. Nat. Hist. Univ. Kansas, 18:505-545, for a review of
the Mexican and Central American species (as Hyla).
Ololygon acuminata Cope, 1862. Proc. Acad. Nat. Sci. Philadelphia, 14:354.
oRIGINAL NAME: Hyla acuminata.
rvru(s): Syntypes: USNM 5843, 102700.
TYPE LoCALITY: "Paragtay".
DrsrRrBuTror.r: Southern Mato Grosso and Mato Grosso do Sul (Brazil), Paraguay,
and northern Argentina.
coMMENr: In the Ololygon x-signata group. See Cei, 1980, Monit. ZooLItal., N.S.,
Monogr., 2:484-487.
Ololygon agilis (Cruz and Peixoto, 1983). Rev. Brasil. Biol., 42:72L.
oRIGINAL NAME: Hyla agilis.
rvnr(s): Holotype: El 7 L23.
TypE LocALITy: "Ibiriba (19' L4' 5,39'55' W, alt. aprox. 20 m.), Municipio de
Linhares, Estado do Espirito Santo", Brazil.
DISTRIBUTIoII: Known only from the region of the type locality.
coMMENT: Related to Ololygonberthae, according to the original description, and in
the Ololygon staufferl group by implication.
Ololygon alleni (Cope, L870). Proc. Am. Philos. Soc., LL:162.
oRTGTNAL NAME: Scytopis alleni.
rvnr(s): Syntypes: ANSP 2159 (Pebas),MCZ 1519 (Par6).
TypE LocALITy: "Pebas", Loreto, Peru, and "Parit" (:Bel6m, PatA), Brazil.
DrsrRrBUTroN: At least the Amazon Basin; range undefined at the present time.
coMMENT: In the Ololygon catharinae group.
Ololygon argyreornafa (Miranda-Ribeiro, 1926). Arq. Mus. Nac., Rio de |aneiro, 27:57.
oRIGINAL NAME: Hylodes argyreornatus.
rvrr(s): Syntypes: MN 114 (2 specimens).
TypE LocArtrv: Rio Mutum, Colatina, Espirito Santo, Brazll.
DISTRIBUTIoN: Coastal region from Espirito Santo to Sio Paulo, Brazil.
coMMENT: Not assigned to species group.
Ololygon aurata (Wied-Neuwied, 1821). Reise Brasil., 2:249.

oRIGINAL NAME: Hyla aurata.
rvnr(s): Unknown.
TypE LocArrrv: Santa In6s, Bahia, Brazil.
DISTRIBUTIoI t: Eastern Bahia, Btazll.
Ololygon baumgardneri (Rivero, l96L). Bull. Mus. Comp. ZooI., 126:123.

oRIGTNAL NAME: Hyla baumgardneri.
rypE LocALttv: Casa de ]ulian, between Tabana and Caflo Chana, 609 m., Territorio
Amazonas, Venezuela.
DISTRIBUTIoN: Amazonas, Venezuela.
coMMENT: In the Ololygon staufferi group.
Ololygon berthae (Barrio, L962). Physis, Buenos Aires, 23:L37.

oRIGINAL NAME: Hyla berthae.
rvnr(s): Holotype: LIHUBA 1080.
TypE LocALIrv: PuntaLara, Departamento Buenos Aires, Argentina.
DISTRIBUTIoN: Northeastern Argentina, southern Paraguay, Uruguay, and southern
Brazil.
coMMENr: In the Ololygon staufferi group. See Cei, 1980, Monit. Zool.Ital., N.S.,
Monogr., 2:491-492.
Ololy gon bl airi (F ouquette and Pyburn, 197 2). Herpetolo gica, 28:L7 6.
oRTGTNAL NAME: Hyla blairi.
rvrE(s): Holotype: AMNH 84585.
TypE LocAttty: Near junction of Rio Guaviare and Rio Ariari, Comisaria Vaup6s,
Colombia.
DISTRIBUTIow: Llanos of central Colombia.
coMMENr: In the'Otolygon rubra group.
Ololygon boesemani (Goin, 1966). ZooI. Meded., Leiden, 4l:229.

oRTGTNAL NAME: Hyla boesemani.
rvnE(s): Holotype: RMNH L260L.
TypE LocArtrv: Zanderij, Suriname District, Surinam.
DISTRIBUTIoN: Northern Guyana and Surinam and Amazonas, Venezuela, to
Amazonas and Par6., Brazil.
coMMENr: In the Ololygon x-signata group.
Ololygonboulengeri (Cope, L887). Buil. U.S. Natl. Mus., 32:12.

oRIGINAL NAME: Scytopis boulengeri.
rYPE(s): Holotype: USNM 13974.
TYPE LoCALITY: "Nic aragva" .
DrsrRrBUTroN:Caribbean lowlands of Central America from Nicaragua to Panama;
Pacific lowlands of Costa Rica and eastern Panama.
coMMENT: For discussion (as Hyla boulengeri) see Duellman, L970, Monogr. Mus.
Nat. Hist. Univ. Kansas, l:20A, and Duellman, L972, ZooI. Meded., Leiden, 47:
180-181. In the Ololygon rostrata group.
Ololygon canastrensis (Cardoso and Haddad, 1982). Rev. Brasil. Biol., 42:499.
oRIGINAL NAME: Hyla canastrensis.
TypE LocAltrv: Parque Nacional da Serra da Canastra, 1.300 m, Sio Roque de
20 15'-46o 30' W), Minas Gerais, Btazil.
Minas (approx.
DISTRIBUTIoITI: Known only from the type locality.
coMMENT: Related to Ololygon rubra and Ololygon catharinae groups, according to
Ololygon catharinae (Boulenger, 1888). Ann. Mag. Nat. Hist., (6)l:417.

oRIGINAL NAME: Hyla catharinae.
rvrr(s): Syntypes: BM L947 .2.L2.65-66.
TypE LocALITy: "Serra do Catharina" , Santa Catarina, Brazil.
DISTRIBUTIoIs: Mountains of eastern and southeastern Brazll.
coMMENT: For discussion see (as Hyla catharinae) B. Lutz, L973, Brazll. Spec. Hyla:
183. In the Ololygon catharinae group. Many of the subspecies cited by B. Lutz,
1973, Brazil. Spec. Hyla:L83-189, and by Duellman, 1977, Das Tierreich,95:43-
44, are recognizable species (UC).
Ololygon crospedospila (A. Lutz, 1925). C. R. S6ances Soc. Biol., Paris, 93:2L1.
oRIGINAL NAME: Hyla crospedospila.
rvnr(s): AL-MN 655 designated lectotype by Bokermann, L966, Lista Anot. Local.
Tipo Anf. Brasil.:48.
TypE LocAt-rry: "Estado do Rio e Sao Paulo"; restricted to Campo Belo, Itatiaia, Rio
de ]aneiro, by Bokermann, L966, Lista Anot. Local. Tipo Anf. Brasil.:48.
DrsrRrBUTroN: Coastal mountains of Rio de Janeiro and Sdo Paulo in southeastern
Brazil.
Ololygon cruentomma (Duellman, L972). Copeia, L972:266.

oRIGINAL NAME: Hyla cruentomma.
rvru(s): Holotype: KU L25587.
TypE LocALrry: "Santa Cecilia on the Rio Aguarico, Provincia Napo, Ecuador (0" 02
N-76'58' W), elevation 340 m".
DISTRIBUTIoTv: Upper Amazon Basin in Ecuador and Peru.
coMMENT: In the Ololygon x-signata group.
Lttz, L925). C. R. S6ances Soc. Biol., Paris, 93:2LL.

Ololygon cuspidata (A.
oRIGINAL NAME: Hyla cuspidata.
rYPE(s): Syntypes: AL-MN 299-302.
TypE LoCArmv: Rio de ]aneiro, Brazll.
DTsTRIBUTIoN: Coastal lowlands of Espirito Santo and Rio de |aneiro in
southeastern Brazil.
Ololygon cynocephala (Dum6ril and Bibron, 1841). Erp. G6n., 8:558.

oRIGINAL NAME: Hyla cynocephala.
rvnn(s): Holotype: MNHNP 765.
TYPE LoCALITY: "Guyane" .
DTsTRIBUTIoN: Northern French Guiana.

coMMENr: Recognized as distinct from Ololygon rubraby Lescure, 1976, Bull. Mus.
NatI. Hist. Nat., Paris, (3)377(2ool..)265:492. In the Ololygon rubra group. The
incorrect spelling Hyla cyanocephala has been used by some authors.
Ololygon duartei (B. Lutz, 1951). Hospital, Rio de Janeiro, 39:705.
oRIGINAL NAME: Hyla rubra duartei.
TypE LocAlIrv: Itatiaia, Rio de ]aneiro, Brazil.
DrsrRrBUTIotrt: Mountains of southeastern Brazll; Serra do Espinhago in Minas
Gerais, Brazll.
ANURA: HYLIDAE: HYLINAE r6t
coMMENr: See B. Lutz, 1973, Brazil. Spec. Hyla:161.. Not assigned to species group
Ololygon ehrhardti (Miiller, L924). Zool. Anz., 59:233.

oRTGINAL NAME: Hyla ehrhardti.
rvnr(s): Holotype: formerly ZSM 801192L; destroyed.
TypE LocAtItv: Humboldt, Santa Catarina, Brasil; stated as "Corup6 (:Humboldt),
Santa Catarina" , BrazTl, by Bokermann, L966, Lista Anot. Local. Tipo Anf .
Brasil.:49.
DISTRIBUTIoIt: Known only from the type locality.
coMMENr: Not assigned to species group. Occasionally misspelled as Ololygon
erhardti.
Ololygon elaeochroa (Cope, 1876). J. Acad. Nat. Sci. Philadelphia, (2)8:105.

oRIGINAL NAME: Hyla elaeochroa.
rvnr(s): USNM 30689 designated lectotype by Cochran, L96L, Bull. U.S. Natl. Mus.,
220:53.
TypE LocArny: East foot of mountains near Sipurio, Provincia Lim6n, Costa Rica.
DrsrRrBUTroN: Caribbean lowlands from Nicaragua to western Panama; Pacific
lowlands of Golfo Dulce region in Costa Rica and Panama.
coMMENr: For discussion see Duellman, L970, Monogr. Mus. Nat. Hist. Univ.
Kansas, 1:188-193. In the Ololygon rubra group.
Ololygon epacrorhina (Duellman, 1972). Zool. Meded., Leiden, 47:L82.

oRIGINAL NAME: Hyla epacrorhina.
TypE LocALItv: "Pilcopata, Departamento Cuzco, Perti, 13o 05'5,71" L2'W,
elevation 750 m-"
DrsrRrBUTroN: Amazonian lowlands of southeastern Peru and northeastern Bolivia
coMMENr: In the Ololygon rostrata group.
Ololygon eurydice (Bokermann, 1968). !. Herpetol., L:29.
oRIGINAL NAME: Hyla eurydice.
rYPE(s): Holotype: WCAB 3L795.
TypELocArrrv: Fazenda Santo Onofre, 10 km east of Marac6s, Brazll.
Southern Bahia to Rio de |aneiro, Btazil.
DISTRIBUTIoITi:
Ololygon flatsoguttata (A.Lutz and B. Lutz, 1939). An. Acad. Brasil. Cienc., Ll:75.
oRIGINAL NAME: Hyla flaaoguttata.
rvnn(s): Holotype: AL-MN 2090.
TypE LocArrry: Fazenda do Bonito, Serra da Bocaina, Sio los6' do Barreiro, Sio
Paulo, Brazil.
DrsrRrBUTroN: Coastal range in Brazil from Espirito Santo to Rio Grande do Sul,
Brazil.
coMMENT: In the Ololygon catharinae grouP.
Ololygon funerea (Cope, I874). Proc. Acad. Nat. Sci. Philadelphia,25:L23.

oRIGINAL NAME: Hyla funerea.
rvnr(s): Syntypes: ANSP LL396-97.
TypE LocAlIrv: Moyobamba, Departamento San Martin, Peru.
DISTRIBUTIoN: Upper Amazon Basin in Ecuador and northern Peru.
coMMENr: In the Ololygon rubra group.
Ololygon fuscomarginata (A. Ltttz, L925). C. R. S6ances Soc. Biol., Paris, 93:138.
oRIcINAL NAME: Hyla fuscomarginata.
rvrr(s): Syntypes: AL-MN 845-846, USNM 96964.
TypE LocALITy: "Sio Paulo and Belo Horizonte", Brazil; restricted to Belo
Horizonte, Minas Gerais, BrazIl, by B. Lutz,1973, Brazil. Spec. Hyla:L65.
DISTRIBUTToN: Southern, central, and eastern Brazil, eastern Bolivia, Paraguay and
northwestern Argentina.
$&
69MMENT: For discussion see B. Lutz, 1973, Brazll. Spec. Hyla:L64.In the Ololygon
staufferi grouP.
Ololygon fuscoaaria (A. Lutz, L925). C. R. S6ances Soc. Biol., Paris, 93:212.
oRIGINAL NAME: HYla fuscoaaria.
rypE(s): Syntypes: AL-MN 76, MCZ 15851.
TypE LocAlrrv, Agua Branca (:Agua Limpa), ]uiz de Fora, Minas Gerais, Brazll.
DrsTRrBUTrOr.t: sOuthern and southeastern Brazil, northern Argentina, Paraguay
and eastern Bolivia.
q9MMENT: See B. Lutz,1973, Brazil. Spec. Hyla:127, for discussion. See also Cei,
1980, Monit. Zool.Ital., N.S., Monogr., 2:487-488. In the Ololygon rubra group.
Ololygon garbei (Miranda-Ribeiro, L926). Arq. Mus. Nac., Rio de Janeiro, 27:96.
oRIGINAL NAME: Garbeana garbei.
TypE LocAl-ttv: Eirunep6, Rio JuruA, Amazonas, Btazil.
DrsrRrBUTroN: Middle and upper Amazon Basin in Colombia, Ecuador, northern
Peru, and Brazil.
coMMENr: See Duellman, !972, Zool. Meded., Leiden, 47:L85-186, for discussion
In the Ololygon rostrata grouP.
Ololygon goinorum (Bokermann, t962). Neotropica, 8:86

oRIGINAL NAME: Hyla goinorum.
TypE LocALITy: "Tarauac6, Estado de Acre, Brasil".
DISTRIBUTIoN: Acre, Brazil.
coMMENr: In the Ololygon staufferi grouP.
Ololygon hayii (Barbour, 1909). Proc. New England Zool. Club, 4:51.
oRIGINAL NAME: Hyla hayii.
rYPE(s): Holotype: MCZ 25L3.
TYPE LocAlIrv: Petr6polis, Rio de Janeiro, Btazil-
DrsrRrBUTroN: Serra do Mar from Espirito Santo to Santa Catarina in southeastern
BraziI.
coMMENT: In the Ololygon rubra group.
Ololygon humilis (8. Ltttz, L954). Mem. Inst. Oswaldo Cruz, Rio de ]aneiro, 52:L60
oRIGINAL NAME: Hyla humilis.
TypE LocArtrY: Rio Babi, Nova IguaEu, Rio de Janeiro, Brazll.
DrsrRrBUTroN: Coastal lowlands of Rio de ]aneiro in southeastern Brazil.
coMMENT: In the Ololygon catharinae grouP.
Ololygon kennedyi (Pyburn, 1973). J. Herpetol., T:297.

oRIGINAL NAME: Hyla kennedyi.
rYPE(s): Holotype: UTA A-3697.
TypE LocArrry: About 110 mi. east-southeast of Puerto Gaitan, Departamento Meta,
Colombia.
DISTRIBUTIoI{: Llanos of eastern Colombia.
coMMENT: In the Ololygon rostrata grouP.
Ololygon longilinea (8. Lutz,1968). Pearce-Sellards Ser., Texas Mem. Mus., 10:5.
oRIGINAL NAME: Hyla longilinea.
TypE LocALrry: Morro de Sio Domingos, Pogos de Caldas, Minas Gerais, Brazil.
DISTRIBUTIoN: Southeastern mountains of Minas Gerais, Brazil-
coMMENr: Not assigned to species grouP.
Ololygon machadoi (Bokermann and Sazima, 1973). Rev. Brasil. Biol., 33:52L.
oRIGINAL NAME: Hyla machadoi.
TypE LocArmv: Kilometer 121, Serra do Cip6, Jaboticatubas, Minas Gerais, Brazll.
DISTRIBUTIoN: Serra do Espinhago, Minas Gerais, Brazil.
coMMENT: Related to small species in the Ololygon catharinae group, according to
Ololygon maracaya (Cardoso and Sazima, 1980). Rev. Brasil. Biol., 40:75.
oRIGINAL NAME: Hyla maracaya.
TypE LocALITv: Fazenda Salto, Alpin6polis, Minas Gerais, Brazil.
DISTRIBUTIow: Mountains of southwestern Minas Gerais, Brazil.
coMMENr: Although named as a species of Hyla by Cardoso and Sazima, the
authors stated that the species was a member of Ololygon as recognized by
Fouquette and Delahoussaye/ L977, ]. Herpetol., ll:387-396.
Ololygon mirim (8. Lutz, 1973). Bol. Mus. Nac., Rio de ]aneiro, N.S., Zool.,288:3
oRIGINAL NAME: Hyla mirim.
TypE LocAltrv: Rio Vermelho, Sio Bento do Sul, Santa Catarina, Brazll.
Ololygon nasica (Cope, 1862). Proc. Acad. Nat. Sci. Philadelphia, L4:354.
oRIGTNAL NAME: Hyla nasica.
rvrE(s): Syntypes: USNM 5835, 32371.
TYPE LocALITY: "Paraguay".
DISTRIBUTIoITI: Paraguay, northern Argentina, Uruguay, eastern Bolivia, and
southern Brazil.
coMMENr: In the Ololygon staufferi group. Placement of Ololygon nasica as a
subspecies of Ololygon x-signafa as done by B. Lutz, L973, Brazil. Spec. Hyla
143, seems to be unwarranted (see Fouquette and Delahoussaye, L977,1.
Herpetol., 11:393).
Ololygon nebulosa (Spix, L824). Spec. Nov. Testud. Ran. Brasil.:39.

oRIGINAL NAME: Hyla nebulosa.
rvrr(s): Syntypes: probably ZSM 253Ll0, now losU MN 4055 (holotype of. Hyla
egleri Lutz, L968) designated neotype by Hoogmoed and Gruber, L983,
Spixiana, Suppl., 9:369.
TypE LocALITy: "in sylvis prope flumen Teff6" , Amazonas, Brazil.
DISTRIBUTIow: Southeastern Venezuela, through the Guianas and the lower
Amazon region to Alagoas in northeastern Brazil.
coMMENr: In the Ololygon rostrata group. Until recently this species was known as
Ololygon egleri (Lutz, 1968, Pearce-Sellards Ser., Texas Mem. Mus., 10:8), but
Hoogmoed and Gruber, 1983, Spixiana, Suppl., 9:37L, pointed out that Hyla
nebulosa Spix, L824, is the oldest name for this species.
Ololygon obtriangulata (8. Lutz, L973). Brazll. Spec. Hyla:190.

oRIGINAL NAME: Hyla obtriangulata.
TypE LocAltrv: Brejo da Lapa, Alto Itatiaia, Rio de ]aneiro, Brazil.
DrsrRrBUTroN: Serra da Mantiqueira and Serra do Mar; city of Sio Paulo, Brazil.
coMMENT: Hyla obtriangulata is a replacement name f.or Hyla catharinae simplex B.
Lutz, L968, Pearce-Sellards Ser., Texas Mem. Mus., 12:5. Not assigned to
species group.
Ololygon opalina (B. Lutz, L968). Pearce-Sellards Ser., Texas Mem. Mus., L2:6.
oRIGINAL NAME: Hyla catharinae opalina.
rYPE(s): Syntypes: MN 4037-38.
TypE LocAtlry: Serra dos Orgios, Teres6polis, Rio de ]aneiro , Brazil.
DISTRIBUTIoN: Serra do Mar in Rio de Janeiro, Brazil.
coMMENT: For discussion of species status, see Heyer, 1980, Proc. Biol. Soc
Washington,9S:657. Presumably in t}l.e Ololygon catharinae gtoup.
Ololygon pachychrus (Miranda-Ribeiro, L937). Campo, Rio de ]aneiro, L937:55

oRIGINAL NAME: Hyla pachychrus.
rvnr(s): Syntypes: MN 237 (4 specimens).
TypE LocAltrv: PoEio, Pernambuco, Btazll.
DISTRIBUTIoN: Northeastern Brazil.
Ololygon perpusilla (A. Lutz and B. Lutz, 1939). An. Acad. Brasil. Cienc., LL:78
oRIGINAL NAME: Hyla perpusilla.
TypE LocAtIty: Recreio dos Bandeirantes, Rio de ]aneiro, BrazII.
DISTRIBUTIoN: Southeastern Brazil (Rio de Janeiro and Sio Paulo).
coMMENr: In the Ololygon rubra group.
Ololygon proboscidea (Brongersma, 1933). Zool. Anz., 103:267.

oRIGINAL NAME: Hyla proboscidea.
rvnr(s): Holotype: ZMA 57L0.
TypELocAlrrv: Upper Gran Rio, Surinam.
DISTRTBUTToN:Interior of the Guianas.
coMMENT: In the Ololygon rostratq group; see Duellman, 1972, Zool. Meded.,
Leiden, 47:L86-L87, and Duellman, L972, Occas. Pap. Mus. Nat. Hist. Univ
Kansas, 11:1-31.
Ololygon quinquefasciata (Fowler, 1913). Proc. Acad. Nat. Sci. Philadelphia, 55:160.
oRIGINAL NAME: Hyla quinquefasciata.
rvnn(s): Holotype: ANSP 18115.
TYPE LocALITY: "Mountains above Chimbo"; corrected to Dur6n, Provincia Guayas,
Ecuador, by Duellman, L97L, Herpetologica 27:212-227.
DISTRIBUTIoN: Pacific lowlands of Colombia and Ecuador.
coMMENr: In the Ololygoit rubra group.
Ololygon rizibilis (Bokermann, L964). Rev. Brasil. Biol., 24:430.

oRIGINAL NAME: Hyla rizibilis.
TypE LocALITy: "Campo Grande, Santo Andr6, Sio Paulo, Brasil"
Ololygon rostrata (Peters, 1863). Monatsber. Preuss. Akad. Wiss. Berlin, 1863:466.
oRIGINAL NAME: Hyla rostrata.
rvnr(s): Holotype: ZMB 3175.
TYPE LocAttty: Caracas, Venezuela.
DISTRIBUTIoN: Central Panama eastward on coastal lowlands to French Guiana;
valleys of Rio Cauca and Rio Magdalena in Colombia; llanos of Colombia and
Venezuela.
coMMENr: For discussion see Duellman, L970, Monogr. Mus. Nat. Hist. Univ.
Kansas, 7:204, and Duellman, L972, Zool. Meded., Leiden, 47:L87-L90. In the
Ololy gon rostrat a group.
Ololygon rubra (Laurenti, 1768). Synops. Rept.:S.

oRIGINAL NAME: Hyla rubra.
rYPE(s): None designated; RMNH L52928 considered neotype (pending
designation by Fouquette) by Duellman, L977, Das Tierreich,95:96.
TypE LocALITy: "Ametica"; potential neotype from Paramaribo, Surinam.
DISTRIBUTIoT: Amazon Basin; coastal Brazil (southward to Espirito Santo); Guianas;
coastal northern South America; eastern Panama; Tobago, Trinidad, and St.
Lucia (Lesser Antilles).
coMMENT: There is disagreement whether Laurenti, L768, or Daudin, 1802, is the
proper authority for the name Ololygon rubra; the earliest name is used herein
pending a decision by the International Commission of Zoological
Nomenclature (WED). In the Ololygon rubra group. Ololygon rubra as currently
recognized is a composite of several species (WED).
Ololygon similis (Cochran, L952). J. Washington Acad. Sci., 42:50.

oRIGINAL NAME: Hyla similis.
rver(s): Holotype: USNM 973L7.
TypE LocALIrv: Manguinhos, Rio de |aneiro, Brazil.
DISTRIBUTIoN: Coastal region of Rio de ]aneiro, Brazil.
coMMENr: In the Ololygon rubra group. The placement of Ololygon similis in the
Ololygon rubra group by Fouquette and Delahoussaye, L977, J. Herpetol., 1L:
392, is contrary to the recognition of Ololygon similis as a subspecies of
Ololygon x-signata by B. Lutz, L973, Brazil. Spec. Hyla:L(l.
Ololygon squalirostris (A. Lutz, 1925). C. R. S6ances Soc. Biol., Paris, 93:212.
oRIGINAL NAME: Hyla squalirostris.
rvnr(s): Syntypes: AL-MN 954-955, USNM 96719.
TypE LocAurv: Fazenda do Bonito, Serra da Bocaina, Sio Paulo, Brazil.
DISTRIBUTIoN: Southeastern Brazll, southern Paraguay, Uruguay, and northeastern
Argentina.
coMMENr: In the Ololygon staufferi group.
Ololygon staufferi (Cope, 1865). Proc. Acad. Nat. Sci. Philadelphia, L7:L95.
oRIGINAL NAME: Hyla staufferi.
rvnr(s): Holotype: USNM L53L7.
TypE LocAurv: Orizaba, Veracruz, Mexico.
DISTRIBUTIoN: Mexico (Tamaulipas and Guerrero southward) through Central
America to central Panama.
coMMENT: For di'scussion of two subspecies see Duellman, L970, Monogr. Mus
Nat. Hist. Univ. Kansas, L:L93-203. In the Ololygon staufferi group.
Ololygon strigilata (Spix, 1824). Spec. Nov. Testud. Ran. Brasii.:38.

oRTGTNAL NAME: Hyla strigilata.
rvrr(s): Formerly ZSM 2369/0 (destroyed); see Hoogmoed and Gruber, L983,
Spixiana, Suppl., 9:368-369, for discussion.
TYPELocAltrv: Bahia, BrazTl.
DrsrRrBUTIoN:Northeastern Brazil. "
coMMENr: In the Ololygon catharinae group.
O I oly g a (Duellman, I97 3). Herpetolo gica, 29 :223.

on sugill at
oRTGTNAL NAME: Hyla sugillata.
rvnr(s): Holotype: KU L46444.
TypE LocALITy: "Estaci6n Biol6gica Rio Palenque, 56 km north of Quevedo,
Provincia Los Rios [Pichincha], Ecuador".
DTsTRIBUTIoN: Pacific lowlands of Colombia and northwestern Ecuador.
coMMENT: In the Ololygon rostrata group.
Ololygon trachythorax (Mtiller and Hellmich, 1936). Wiss. Ergebn' Deutsch. Gran
Chaco-Exped., AmPh. RePt., 1:73.
oRIGINAL NAME: Hyla trachYthorax.
rvrr(s): Syntypes: ZSM L56 (2 specimens).
TypE LocALITY: "Apa-Bergland (San Luis)", Paraguay'
coMMENr: The unjustified emended spetling Ololygon trachy-torax has been used.
Although considered a synonym of Ololygon_fuscoaaria by |u!2, L973, Brazil.
Spec. iyla:Ll7, and subsequently by Cei, 1980, Monit. Zool.Ital., N.S.,
Itonogi., 2:487, Fouquette and Detahoussaye, 1977, j. Herpetol., LL:387-396,
not orily considere d,blotygon trachythorax to be a distinct species, but placed it
in a difierent species group (the Ololygon catharinae grovP) from Ololygon
fuscoaaria (in the OlolygonVubra group). Cei did not mention the earlier PaPer
'by
Fouquette and Deiihoussaye so we presume that he did not see it prior to
his monograPh going to Press.
Ololygon trilineata Hoogmoed and Gorzula, 1979. Zool. Meded', Leiden, 54:L93'
rvnE(s): Holotype: RMNH L8357.
TypE LocAurv: L2 km SE El Manteco, Bolivar, Venezuela'
DrsrRrBUTroN: Llanos of eastern Venezuela and savannas of Guyana and Surinam
coMMENT: Not assigned to species SrouP'
ololygon wandae (Pyburn and Fouquette, L97L). J. Herpetol., 5:97.

oRIGINAL NAME: Hyla wandae.
unE(s): Holotype: USNM L92305.
TypE LocAr-r", i2 km north-northeast of Villavicencio, Departamento Meta,
Colombia.
DISTRIBUTIoN: Ecotone of forest and llanos in central Colombia.
coMMENr: In the Ololygon staufferi grouP.
Olotygon x-signata (Spix, L824). Spec. Nov. Testud. Ran. Brasil.:40.

oRIGINAL NAME: HYla x-signata.
rvnr(s): Formerly ZS}.,d 2494/0 (now destroyed).
TYPE LocAltrY: Bahia, Brazil.
DISTRIBUTIoN: Northern Venezuela to Surinam; eastern, southern, and southeastern
Brazil.
coMMENr: In the Ololygon x-signata grouP. See Cei, 1980, Monit. Zool.Ital., N.S.,
Monogr., 2:+88-i{t Hoo[moed and Gruber, L983, Spixiana, Suppl., 9:364,
includid Hyla affinis Spix and Hyla coerulea Spix in synonymy.
Osteocephalzs Steindachner, 7862. Arch. Zool. Anat. Physiol. Wisg' Med',2:77'

rypE spEcrss: Osteocephalus taurinus Steindachner, L862, by original designation.
DrsrRrBUTroN: Guianas, Amazon Basin, upper Orinoco and Magdalena drainages in
Venezuela and Colombia, southeastern Brazil, and northeastern Argentina.
q9MMENT: See Trueb and Duellman,lg7l, Occas. Pap. Mus. Nat. Hist. Univ. Kansas,
l:L-47, for a review of this genus. See comment under Hyla rodriguezi.
Osteocephalus buckleyl (Boulenger, 1882). Cat. Batr. Sal. Brit. Mus.:362'

oRIGINAL NAME: Hyla buckleYi.
rvnr(s): Syntypes:-BM Lg47-.2.L3.36-39 (Sarayacu), L947.2.13.40-4L, L947.2.L3.43-45
iCu"LfoJ), 1947.2.L3.46 (Pallatanga); BM 1947.2.L3.44 designaterl lectotype_by
Trueb and Duellman, Lg7L, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, l':19.
TypE LOCALITY: "SarayactJ" and "CanelOS", PfOvinCia PastaZa, and "Paitanga"
(:Pallatanga), Provincia Chimborazo (in error), Ecuador; type locality
restricted t"o Canelos by Cochran and Goin, L970, Bull. U.S. Natl. Mus.,288:
21.3, and by lectotype designation.
DISTRIBUTION: Amazon Basin from Amap6, Btazil, through the Guianas to
Colombia, Ecuador, Peru, and Bolivia.
ANURA: HYLIDAE: HYLTNAE 167
Osteocephalus langsdorffii (Dumr6ril and Bibron, 1841). Erp. G6n., 8:557.

oRIGINAL NAME: Hyla Langsdorffii.
TYPE LocALITY: "Br6si1".
DrsrRrBUTroN: Atlantic forests from Bahia, Brazil, to northeastern Argentina.
CoMMENT: See Duellman, L974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas,24:20,
for generic allocation.
Osteocephalus leprieuril (Dum6ril and Bibron, 1841). Erp. G6n., 8:553.
oRIGINAL NAME: Hyla Leprieurii.
TypE LocALITy: "Cayenne", French Guiana.
DrsrRrBUTroN: Amazon Basin, in the Guianas and the upper Amazon Basin in
Colombia, Ecuador, Peru, and western Brazil.
Osteocephalus pearsoni (Gaige, 1929). Occas. Pap. Mus. Zool. Univ. Michigan,207:3.
oRIGINAL NAME: Hyla pearsoni.
rvnE(s): Holotype: UM.Ill4.Z 57548.
TypE LocAurv: Upper Rio Beni, below mouth of Rio Mapiri, Departamento El
Beni, Bolivia.
DrsrRrBUTroN: Upper Amazon Basin and lower Andean slopes from central Peru to
northern Bolivia.
Osteocephalus taurinus Steindachner, 1862. Arch. ZooL Anat. Physiol. Wiss. Med.,
2:77.
rYPE(s): Holotype: NHMW L6492.
TypE LocAttrv: Barra do Rio Negro, Man6us, Amazonas, Brazil.
DISTRIBUTIoN: Amazon Basin, upper Orinoco Basin, and the Guianas.
coMMENT: As presently recognized, this taxon consists of two species that differ in
size and advertisement call (WED). See comment under Hyla elkejungingerae.
Osteocephalus oerruciger (Werner, 1901). Verh. Zool. Bot. Ges. Wien, 50:601.
oRIGINAL NAME: Hyla aerrucigera.
rvnr(s): Holotype: ZIli'{B 16589.
TYPE LOCALITY:,,ECUAdOT,, .
DrsrRrBUTroN: Lower Amazonian slopes of Andes from Colombia to central Peru.

CoMMENT: The incorrect epithet aerrucigerushas had widespread use.
Osteopilus Fitzinger, 1843. Syst. Rept.:30.

TypE SPECIts: Trachy,cephalus marmoratus Dum6ril and Bibron, 184L (:Hyla
septentrionalis Dum6ril and Bibron, 1841).
DrsrRrBuTror.r: The Greater Antilles, Bahama Is., and southern Florida (USA).
coMMENT: For a review of the genus see Trueb and Tyler, \974, Occas. Pap. Mus
Nat. Hist. Univ. Kansas, 24:L-50.
REVIEwERS: A1bert Schwartz (AS); Richard Thomas (RT).
Osteopilus brunneus (Gosse, 1851). Naturalist's Sojourn in )amaica:361

oRIGINAL NAME: Hyla brunnea.
rYPE(s): Unknown.
TYPE LocALttv: Savanna-la-Mar, [Westmoreland Parish,] Jamaica.
DISTRIBUTIow: ]amaica, West Indies.
Osteopilus dominicensis (Tschudi, 1838). Classif. Batr.:30.

oRIGINAL NAME: Hypsiboas dominicensis.
rvrr(s): Syntypes: MNHNP 46L4 (4 specimens).
TYPE LocALITY: "Saint-Domingue".
Hispaniola, including Ile de la GonAve, Ile-i-Vache, Ile Grande
DrsrRrBUTroN:
Cayemite, Ile de la Tortue, and Isla Saona (West Indies).
CoMMENT: Peters, L974, Mitt. Zool. Mus. Berlin, 50:299-322, regarded this as a
subspecie s of. Ost eopilus sept entrionalis.
Osteopilus septentrionalis (Dum6ril and Bibron, 1841). Erp. G6n., 8:538.

oRIGINAL NAME: Hyla septentrionalis.
rYPE(s): Holotype: MNHNP 461.2.
TYPE LoCALITY: "Cuba".
DISTRIBUTIoN: West Indies: Cuba, Isla de Pinos (now Isla de ]uventud), Cayman Is..
Bahama Is.; introduced on northwestern Puerto Rico, Virgin Is., and southern
Florida (USA).
coMMENr: Reviewed by Duellman and Crombie, 1970, Cat. Am. Amph. Rept.,
92.1-4 (as Hyla septentrionalis). Hyla septentrionalis is a replacement name for
Trachycephalus marmoratus Dum6ril and Bibron, L841, Erp. G6n., 8:538. See
comment under Ost eopilus dominicensis.
Phrynohyas Fitzinger, L843. Syst. Rept.:30.

TYPE sPECIns: Hyla zonata Spix, 1824 (:11rna aenulosa Laurenti, 1768), by monotypy
and subsequent designation of the Int. Comm . Zool. Nomencl. , 1958, Opin. 520.
DISTRIBUTIoN: Lowlands of Mexico, Central and South America east of the Andes,
south to northern Argentina.
coMMENT: For a review of the genus see Duellman, 197L, Occas. Pap. Mus. Nat. Hist
Univ. Kansas, 4:L-21.
Phrynohyas coriacea (Peters, L867). Monatsber. Preuss. Akad. Wiss. Berlin, 1867777
oRIGINAL NAME: Hyla coriacea.
rvnr(s): Syntypes: ZMB 5807 (3 specimens).
DISTRIBUTIoN: Surinam and upper Amazon Basin in Ecuador and Peru.
Phrynohyas imitatrix (Miranda-Ribeiro, 1926). Arq. Mus. Nac., Rio de ]aneiro, 27:77
oRIGTNAL NAME: Hyla imitatrix.
rver(s): Syntypes: MN 154 (2 specimens).
TypE LocAtrty: Teres6polis, Rio de ]aneiro, Brazil.
DISTRTBUTToN: States of S5o Paulo and Rio de ]aneiro, Brazil.
Phrynohyas mesophaea (Hensel, 1867). Arch. Naturgesch., 33:154.

oRTGTNAL NAME: Hyla mesophaea.
TypE LocArmy: Porto Alegre, Rio Grande do Sul, Brazil.
Atlantic forests of southeastern Brazil.
DISTRIBUTIoITI:
Phrynohyas resinifictrix (Goeldi, L907). Proc. Zool. Soc. London, 1907:135.

oRIGINAL NAME: Hyla resinifictrix.
rYPE(s): Holotype: BM 1947 .2.23.24.
TYPE LocALITY: "Mission of San Antonio do Prata, at the River Maracani", Brazll.
DISTRIBUTIoN: Surinam, French Guiana and Amazonian Brazil.
coMMENr: Recognized as distinct from Phrynohyas aenulosa by Lescure, L976, Bull
Mus. Natl. Hist. Nat., Paris, (3)377(2ool.)265:503.
Phrynohyas aenulosa (Laurenti, 1768). Synops. Rept.:3l.

oRIGINAL NAME: Rana aenulosa.
rYPE(s): Unknown; based on pl.72in Seba, 1734, Loc. Nat. Thes. Desc. Icon.
Exp., 1.
TYPE LoCALITY: "America".
Lowlands of Mexico, Central America (to 2500 m in Guatemala), to
DISTRIBUTIoN:
Amazon Basin of Btazil, Guianas, and south to ParanA (Brazil), Paraguay,
northern Argentina; Trinidad and Tobago.
coMMENT: See Cei, 1980, Monit. Zool.Ital., N.S., Monogr., 2:444-450, and
Duellman, L970, Monogr. Mus. Nat. Hist. Univ. Kansas, l:L63-L72. See
comment under Argenteohyla altamazonica.
Phyllodytes Wagler, 1830. Nat. Syst. Amph.:202.

rypE spECIrs: Hyla luteola Wied-Neuwied, L824.
DISTRIBUTIou: Eastern Brazll; Trinidad.
coMMENT: The most recent review of the genus was by Bokermann, L966, An. Acad
Brasil. Cienc., 38:335-344.
Phyllodytes acuminatus Bokermann, L966. An. Acad. Brasil. Cienc., 38:342.

rvrr(s): Holotype: WCAB 270L.
TypE LocAtIrv: Mangabeiras, Alagoas, Brazil.
Phyllodytes auratus (Boulenger, L9L7). Ann. Mag. Nat. Hist., (8)20:185.

oRIGINAL NAME: Amphodus auratus.
rvnE(s): Syntypes: BM L9L7 .9.27 .39-4L.
TypE LocAurv: Mt. Tucutche, Trinidad.
coMMENr: See Kenny, L969, Stud. Fauna Curagao and other Caribb. ls., 29:L-78.
Phyllodytes luteolus (Wied-Neuwied, 1824). Abbild. Naturgesch. Brasil.:pl. 4L, hg. 2

oRTGTNAL NAME: Hyla luteola.
rvnr(s): Unknown (?AMNH).
TypE LocAlrrv: R6gencia, Espirito Santo , Brazll (by subsequent designation of
Bokermann,1966, An. Acad. Brasil. Cienc., 38:336).
DrsrRrBUTroN: Coastal region of eastern Brazil from Paraiba to southern Espirito
Santo.
Phyllodytes tuberculosas Bokermann, L966. An. Acad. Brasil. Cienc., 38:339

TypE LocAtItv: Marac6s, Bahia, Brazll.
DISTRIBUTIoN: Sergipe to Bahia, Brazil.
Plectrohyla Brocchi, L877. Bull. Soc. Philomath. Paris, (7)L:92.

rypE spEcrys Plectrohyla guatemalensis Brocchi, L877, by original designation.
Highlands of southeastern Mexico and northern Central America.
DrsrRrBUTroN:
coMMENT: For review of genus and definition of species groups see Duellman,'1.970,
Monogr. Mus. Nat. Hist. Univ. Kansas, l:547-584.
Plectrohyla aaia Stuart, 1952. Proc. Biol. Soc. Washington, 65:6.

wrn(s): Holotype: UIl/.}l{Z L02280.
TypE LocAury: Grhnja Lorena, 10 km airline northwest of Colombia, Departamento
Quetzaltenango [now Quezaltenango], Guatemala.
DrsrRrBUTroN: Pacific slopes of Chiapas, Mexico, and southwestern Guatemala.
coMMENT: In the Plectrohyla guatemalensis group.
Plectrohyla dasypus McCranie and Wilson, 1981. Occas. Pap. Mus. Nat. Hist. Univ
Kansas, 92:1.
TypE LocAllrv: Quebrada Cusuco at EI Cusuco, 6.5 km WSW Buenos Aires,
Departamento Cortes, Honduras.
DISTRIBUTION: SiCTTA OMOA, HONdUTAS.
coMMENT: In the Plectrohyla saSorum grouP.
Plectrohyla glandulosa (Boulenger, 1883). Ann. Mag. Nat. Hist., (5)12:L64-

oRIGINAL NAME: Hyla glandulosa.
rvnr(s): Syntypes: BM 1947 .2.20.40-41'.
TYPE LoCALITY: "Guatemala".
DISTRIBUTIoN: Highlands of Guatemala and El Salvador.
Plectrohyla guatemalensls Brocchi, L877. BuII. Soc. Philomath. Paris, (7)L:92'

rYPE(s): Syntypes: MNHNP 6332 (2 specimens).
TYPE LocAltrv: "Pacicilla" (:Patzicia), Departamento Chimaltenango' Guatemala'
DrsrRrBUrroN: Highlands of Chiapas, Mexico, southeastward to El Salvador and
north-central Honduras.
q9MMENT: In the Plectrohyla guatemalensls group. This taxon consists of at least
three distinct sPecies (WED).
plectrohyla hartwegi Duellman , 1968. Univ. Kansas Pubt. Mus. Nat. Hist. , 17:576.
rYPE(s): Holotype: U}l,4}l4,Z 94428.
)'Burrejonel (9 kitometers west of Chicomuselo), Chiapas, M6xico,
rypE LocArrry,
DrsrRrBUTrou: Pacific slopes of Chiapas and eastern Oaxaca, Mexico; Sierra de las
Minas, Guatemala.
coMMENT: In the Plectrohyla guatemalensls group.
Plectrohylairll Stuart, L942. Occas. Pap. Mus. Zool. Univ. Michigan, 455:4'
rYPE(s): Holotype: Uill{}t{Z 89092.
-Fir,ca
TypE LocAr.rrvr San Francisco, 25 km N of Nebaj, lL75 m, Departamento el
Quiche, Guatemala.
DrSTRrBUrroN: Atlantic slopes of the highlands in western Guatemala and Chiapas.
Mexico.
Plectrohyla lacertosa Bumzahem and Smith, L954. Herpetologica, 10:64.

rYPE(s): Holotype: UIMNH 33693.
TypE LocALITY: "Region del Soconusco", Chiapas, Mexico.
DISTRIBUTIoN: Unknown from any precise locality.
q9MMENT: Known only from the holotype; relationships and status of the species
are unknown (WED).
Plectrohyla matudai Hartweg, L94L. Occas. Pap. Mus. Zool. Univ. Michigan,437:5'
rYPE(s): Holotype: UI[l/.}l,4.Z 88863.
TypE LOCAltry: Cerro Ovando, Distrito Soconusco, Chiapas, Mexico.
DrsrRrBUTroN: Pacific slopes of southeastern Mexico (Oaxaca and Chiapas) and
Guatemala.
Plectrohyla pokomchi Duellman and Campbell, 1984. Copeia, L984:394.

rypE LocAury'
i'Rio Sananj6, 3.5 km E of La Uni6n Barrios, Departamento Baja
Yerapaz, Guatemala, i,585-1.,707 m (15" 11' N, 90o 10' W)".
DrsrRrBUT;oN: Cloud forest above 1400 m in the Sierra de las Minas and the
contiguous Sierra de Xucaneb in eastern Guatemala'
Plectrohyla pycnochila Rabb, L959. Herpetologica, 15:45.

TypE LocALITY: "Coyame, VeractlJzt Mexico" (in error).
DISTRIBUTIoN: Highlands of central Chiapas, Mexico.
coMMENT: In the Plectrohyla guatemalensls group.
Plectrohyla quecchi Stuart, L942. Occas. Pap. Mus. Zool. Univ. Michigan,455:L.
TypE LocAurv: barranca Las Palmas, 2 km north of Finca Los Alpes, Alta Verapaz,
Guatemala, elevation 1015 m.
DISTRIBUTIoN: Atlantic slopes of central Guatemala.
Plectrohyla sagorum Hartweg, L94L. Occas. Pap. Mus. Zool. Univ. Michigan, 437:2.
TypE LocArrry: Cerro Ovando, 1800 m, Distrito Soconusco, Chiapas, Mexico.
DrsrRrBUrroN: Pacific slopes of southeastern Mexico (Chiapas) to EI Salvador.
coMMENr: In the Plectrohyla sagorum grouP.
Plectrohyla tecunumani Duellman and Campbell, 1984. Copeia, L984:391.

TypE LocALtry: "a cave 1 km east of Chemal, Departamento Huehuetenango,
Guatemala, 3,395 m (15' 27' N, 9L" 3I' W)".
DrsrRrBUTroN: Two localities in the Sierra de Cuchumatanes (type locality and 2.6
km N Huehuetenango), Guatemala.
Pseudacris Fitzinger, 1843. Syst. Rept.:31.
TypE spECIrs: Rana nigrita Le Conte, L825, by monotypy.
DrsrRrBUrroN: North America east of the Rocky Mountains from Hudson Bay to the
Gulf of Mexico.
Pseudacris brachyphona (Cope,1889). Bull. U.S. Natl. Mus., 34:34L.
oRIGINAL NAME: Chor ophilus f eriarum br achy ph onus.
rvrr(s): Unknown.
TypE LocALITy: "in west Pennsylvania, near the Kiskiminitas River",
Westmoreland County, Pennsylvania, USA.
DrsrRrBUTroN: Appalachian Mountains from Pennsylvania to Alabama, USA.
coMMENT: Reviewed by Hoffmann, 1980, Cat. Am. Amph. Rept., 234.L-2.
Pseudacris brimleyi Brandt and Walker, L933. Occas. Pap. Mus. ZooI. Univ. Michigan,
262:2.
rvnr(s): Holotype: UI|I/.MZ 7 4316.
TypE LocALITy: "near Washington, [Beaufort County,] North Carolina", USA.
DISTRIBUTIow: Atlantic Coastal Plain from Virginia to Georgia, USA.
coMMENr: Reviewed by Hoffmann, 1983, Cat. Am. Amph. Rept., 3lL.L-2.
Pseudacris clarkii (Baird, 1854). Proc. Acad. Nat. Sci. Philadelphia,T:60.

oRIGINAL NAME: Helocaetes clarkii.
rvnE(s): Syntypes: USNM 33L3,3315, 33L7.
TypE LocAury: Galveston and Indianola, Texas, USA; restricted to "Galveston",
Galveston County, Texas, USA, by Schmidt, L953, Check List N. Am. Amph.
Rept., Ed.6:74.
DrsrRrBUrroN: Central USA from Kansas to the GuIf of Mexico; lower Rio Grande
Valley (Texas, USA, and Tamaulipas, Mexico).
coMMENr: See Duellman, L970, Monogr. Mus. Nat. Hist. Univ. Kansas, L:642-645.
Pseudauis nigrita (Le Conte, L825). Ann. Lyc. Nat. Hist. New York, L:282.
oRIGINAL NAME: Rana nigrita.
rvnr(s): Unknown.
TypE LocAtttv: None given; designated as "Liberty County, Georgia", USA, by
Schmidt, L953, Check List N. Am. Amph. Rept., Ed.6:74.
DrsrRrBUrroN: Atlantic Coastal Plain from North Carolina to Mississippi, USA.
coMMENT: Two subspecies recognized; for discussion see Smith and Smith, 1952,
Am. Midl. Nat., 48:165-180, and Schwartz, L957, Am. Mus. Novit., 1838:1-12.
Pseudacris ornata (Holbrook, 1836). N. Am. Herpetol., Ed. L, L:97.
oRIGINAL NAME: Rana ornata.
rvnr(s): Unknown.
TypELocALrry:"...onlyinSouthCarolina...aboutfourmilesfromCharleston,
between the Cooper and Ashley rivers . . . ", Charleston County, South
Carolina, USA.
DrsrRrBUTroN: Atlantic Coastal P1ain from North Carolina to central Florida and
southeastern Louisiana, USA.
CoMMENT: For discussion see Harper, L937, Am. Midl' Nat', 22:134-149'
Pseudacris streckeri A. H. Wright and A. A. Wright,1923. Handbook Frogs Toads, Ei
l:25.
rypE(s): Syntypes: CU 2485 (5 specimens), now lost according to Smith' L966'
Cat
Am. AmPh. RePt., 27.1.
TypE LOCArtry: Not itut"d; designated (in error) as
"Waco, Mclennan County'
Texas", USA by Schmidt, {953, Check List N. Am. Amph. Rept., Ed.6:76;
revised. to "soinerset, Bexar County, Texas", USA, by Smith, L966, Cat' Am'
Amph. RePt., 27 -L. r . i.
6ir"ontinuous in south-central USA: west-central Illinois;
DrsrRrBUTro*,
southeastern Missouri and adjoining areas; Oklahoma south to the GuIf of
Mexico.
coMMENr: Reviewed by smith, L966, Cat. Am. Amph. Rept., 27.L-2.
Pseuda7is triseriata (wied-Neuwied, 1838). Reise N. Am., L:249.
oRIGINAL NAME: HYla triseriata.
rvrr(s): Lost.
TypE LocAlrry: "Mount Vernon, Ohio River, Indiana", USA; restricted to betweer'
Rush Creek and Big Creek along route from New Harmony to Mt' Vernon'
posey County, Indiana, USA, by Harper L955, Proc. Biol. Soc. Washington, 6!
155.
DrsrRrBUTroN: east of Rocky Mountains from Great Bear Lake and
North America
southeastern Quebec, Canada, southward to northern Florida and Texas, USA
q9MMENT: Four subspecies recognized; for discussion see Smith and Smith, L952'
Am. Midl. Nat., 48:165-180; and schwattz,1957, Am.Mus. Novit., 1838:1-12
Pternohyla Boulenger, L882. Ann. Mag. Nat. Hjst', (5)10:326'
rypE;pECrrs: Ptirnohyla fodiens Boulenger, L882, by monotypy.
DrsrRrBUTroN: South6rn Arizona (USA) southward to Michoacin and Aguascalientes
Mexico.
CoMMENT: See Trueb ,lg7o, Univ. Kansas Publ. Mus. Nat. Hist., L8:546-716,
and
Duellman , Lg7O, Monogr. Mus. Nat. Hist. Univ. Kansas, L:6L8-628. Reviewed b"
Trueb, 1969, Cat. Am. Amph. Rept.,77 'L'
Pternohyla dentata Smith, L957. Herpetologica, 13:1'
tvnr(s): HolotyPe: UIMNH 40551.
TypE LocALrrv:)'8 mi. NE Lagos de Moreno, Jalisco, Mexico"'
DrsrRrBUTroN: Central Mexico (Aguascalientes and adjacent Jalisco).
CoMMENT: Reviewed by Trueb, L969, Cat. Am. Amph. Rept., 77.1-2.
Pternohyla fodiens Boulenger, L882. Ann. Mag. Nat. Hist., (5)10:326.

TYPE LocALITY; "Presidio, [Sinaloa,] W. Mexico"'
DISTRIBUTIoN: Southern Arizona, USA, southward to Michoac6n, Mexico'
coMMENr: Reviewed by Trueb, 1969, Cat. Am. Amph. Rept., 77.3-4.
Ptychohyla Taylor, L944. Univ. Kansas Sci' Bull', 30:41'

irpr ipEcrni: Ptychohyla adipoaentris Taylor, L944 (:Hyla leonhardschultzei Ahl, 1934),
by original designation.
DrsrRr;urroN: Moderlte elevations of southern Mexico (Chiapas, Guerrero, Oaxaca)
southeastward in Central America to northern Nicaragua.
69MMENT: For review of genus see Duellman, 1970, Monogr. Mus. Nat. Hist' Univ'
Kansas 1:518-547.
Ptychohyla euthysanota (Kellogg , L923). Proc. Biol. Soc. Washington, 4L:123'

oRIGINAL NAME: HYla euthYsanota.
rvrE(s): HolotyPe: USNM 73296.
rypE LocArrrvi Los Esemiles, Departamento Chalatenango, El Salvador.
DrsrRrBUTroN: Southeastern Mexico (Chiapas, Oaxaca) and Pacific slopes of
Guatemala and El Salvador.
coMMENT: The two subspecies recognized by Duellman, 1970, Monogr. Mus. Nat
Hist. Univ. Kansas, l:5L8-547, are possibly specifically distinct (WED).
Ptychohyla ignicolor Duellman, L961. Univ. Kansas Publ. Mus. Nat. Hist., 13:352.
rvnr(s): Holotype: U}/.MZ 119603.
TypE LocALtry: "a stream 6 kilometers (by road) south of Vista Ffermosa, Oaxaca,
M6xico (1865 meters)".
DISTRIBUTIoN: Northern slopes of Sierra luArez, Oaxaca, Mexico.
Ptychohyla leonhardschultzei (Ahl, 1.934). Zool. Anz., 106:185.

oRIGINAL NAME: Hyla leonhard-schultzei.
TypE LocAurv: Malinaltepec, Guerrero, Mexico.
DrsrRrBUTroN: Atlantic and Pacific slopes of Oaxaca and Guerrero, Mexico.
Ptychohyla panchoi Duellman and Campbell, L982. Herpetologica, 38:374.

TypE LocArlry: "Aldea Vista Hermosa, Municipio Los Amates, Departamento de
lzabal, Guatemala . . . . Vista Hermosa is situated on the northern escarpment
of the Sierra de las Minas at approximately LSo 17' N, 89" 13' W, at an
elevation of 600 m."
DISTRIBUTIoN: Sierra de las Minas, Guatemala.
Ptychohyla schmidtorum Stuart, L954. Proc. Biol. Soc. Washington, 67:L69.

rYPE(s): Holotype: FMNH 27055.
TypE LocAllry: El Porvenir, 17 km airline west of San Marcos, Departamento San
Marcos, Guatemala.
DISTRIBUTIoN: Atlantic slopes of Chiapas, Mexico, and Pacific slopes of
southeastern Mexico and Guatemala.
coMMENT: The two subspecies recognized by Duellman, 1970, Monogr. Mus. Nat.
Hist. Univ. Kansas, L:5L8-547, are probably distinct species (WED).
Ptychohyla spinipollex (Schmidt, L936). Proc. Biol. Soc. Washington,49:45.

oRIGINAL NAME: Hyla spinipollex.
TypE LocAt.rry: "Mountains behind Ceiba" , Departamento Atlantidad, Honduras.
DrSTRrBUTrorrr: Atlantic slopes of Central American highlands from Guatemala to
northern Nicaragua.
Smilisca Cope, 1865. Proc. Acad. Nat. Sci. Philadelphia, L7:L94.

rypE spECras: Smilisca daulinia Cope, L865 (:Hyla baudinii Dum6ril and Bibron, 1841).
DrsrRrBUTroN:Lowlands of Mexico, extreme southern Texas (USA), Central America
and northwestern South America (exclusive of Amazon Basin).
coMMENT: For review see Duellman and Trueb, t966, Univ. Kansas Publ. Mus. Nat.
Hist., L7:28L-375, and Duellman, L970, Monogr. Mus. Nat. Hist. Univ. Kansas, 1:
585-618. Generic synonymy and key in Duellman,1968, Cat. Am. Amph. Rept.,
58.1-2.
Smiliscabaudinii (Dum6ril and Bibron, 1841). Erp. G6n., 8:564.

oRIGINAL NAME: Hyla Baudinii.
TypE LOCAllryr "Mexique"; restricted to C6rdoba, Veracruz, Mexico, by Smith and
il!l
Taylor, L950, Univ. Kansas Sci. Bull., 33:347; restricted (in error) to "Mexico
City" by Schmidt,1953, Check List N. Am. Amph. Rept., Ed- 6:59.
DrsrRrBUTroN: Extreme southern Texas (USA) and southern Sonora (Mexico) south
in tropical lowlands to Costa Rica.
coMMENr: Reviewed by Duellman, L968, Cat. Am. Amph. Rept., 59.L-2.
sf,[
Smilisca cyanosticta (Smith, 1953). Herpetologica, 8:150.

oRIGINAL NAME: Hyla phaeota cyanosticta.
rYPE(s): Holotype: USNM LLLL47.
TypE LocALITy: "Piedras Negras, Pet6n, Guatemala".
DISTRIBUTIoN: Atlantic slopes of southeastern Mexico
and Guatemala from ]alapa
de Diaz (Mexico) to the Montafias del Mico (Guatemala).
coMMENr: Reviewed by Duellman, 1968, Cat. Am. Amph. Rept., 60.L-2.
Smilisca phaeota (Cope, 1862). Proc. Acad. Nat. Sci. Philadelphia, 14:358.
oRIGTNAL NAME: Hyla phaeota.
rvne(s): Holotype: USNM 4347.
TypE LocALITy: "Turbo, [Intendencia de Choc6, Colombia,] New Granada".
DISTRIBUTIOTT: Caribbean lowlands of Central America from Nicaragua to
northwestern Colombia; Pacific lowlands of southeastern Costa Rica, eastern
Panama, Colombia, and northwestern Ecuador; valleys of the Rio Cauca and
Rio Magdalena, Colombia.
coMMENr: Reviewed by Duellman, L968, Cat. Am. Amph. Rept., 6L.L-2.
Smilisca puma (Cope, 1885). Proc. Am. Philos. Soc., 22:L83.

oRTGTNAL NAME: Hyla puma.
rvnr(s): Holotype: USNM 13735.
TYPE LocALITy: "Nicaragua".
Caribbean lowlands of Costa Rica and adjacent Nicaragua.
DISTRIBUTIoN:
Smilisca srla Duellman and Trueb, 1966. Univ. Kansas Pubt. Mus. Nat. Hist., 17:31.8.
rvnn(s): Holotype: KU 91852.
TYPE LocALtrY: "a small stream at north edge of the village of El Volc6n, Chiriqui,
elevation 1280 meters", Panama.
DISTRIBUTIow: Pacific lowlands of Costa Rica and Panama; Caribbean lowlands of
Panama and Colombia.
Smilisca sordida (Peters, 1863). Monatsber. Preuss. Akad. Wiss. Berlin, L863:460
oRIGINAL NAME: Hyla sordida.
ryPE(s): Syntypes: ZMB 3L4l (2 specimens).
TYPE LocALITy: "Veragua", Panama.
DISTRIBUTToN: Costa Rica and western Panama.
Sphaenorhynchus Tschudi, 1838. Classif. Batr.:7L.

TypE spECIns: Hyla lactea Daudin, 1802, by monotypy.
DISTRIBUTIoII:Amazon and Orinoco basins of South America; Guianas; eastern
Brazil; Trinidad.
Sphaenorhynchus bromelicola Bokermann, L966. Rev. Brasil. Biol., 26L8.

rvnr(s): Holotype: WCAB 3L941..
TYPE LocAtttv: Fazenda Santo Onofre, 10 km east of Marac6s, Bahia, BrazII.
DISTRTBUTToN: Known only from the type locality.
Sphaenorhynchus carneus (Cope, 1868). Proc. Acad. Nat. Sci. Philadelphia, 20:L11.
oRIGINAL NAME: Hylella carnea.
rvnE(s): Lost.
TypE LocALITy: "Napo or upper Maranon", Perlt.
DISTRIBUTIoN: Upper Amazon Basin in southern Colombia, Peru, and Ecuador;
central Amazonia in Brazil.
coMMENr: See Duellman, 1974, Occas. Pap. Mus. Nat. Hist Univ. Kansas, 27:22.
Sphaenorhynchus dorisae (Goin, 1957). Caldasia, 8(36):16.

oRIGINAL NAME: Sphoenohyla dorisae.
rYPE(s): Holotype: FSM 3506.
TypE LocAury: Leticia, Comisaria Amazonas, Colombia.
DISTRIBUTIoN: Upper Amazon Basin in southeastern Colombia, Ecuador, and
Brazilian Amazonia.
Sphaenorhynchus lacteus (Daudin, 1802). Hist. Nat. Rain. Gren. Crap.:20.
oRIGINAL NAME: Hyla lactea.
rYPE(s): Holotype: MNHNP 487L.
TYpE LocALITy: "Brasil".
Amazon and Orinoco basins, Guianas, and Trinidad.
DISTRIBUTIoN:
coMMENr: See Duellman and Lynch, L98L, ]. Herpetol., L5:237-239, for discussion
of nomenclature and synonymy of Sphaenorhynchus eurhostus Rivero, 1969,
Copeia, L969:701. For discussion (as Sphaenorhynchus eurhostus) see Rivero,
1969, Copeia, L969:7 00-7 03.
Sphaenorhynchus orophilus (A. Lutz and B. Lutz,1938). An. Acad. Brasil. Cienc., 10:
t78.
oRIGINAL NAME: Hyla (Sphoenohyla) orophila.
TypE LocArrrv: Quitandinha, Petr6polis, Rio de ]aneiro, Brazil.
DISTRIBUTIoIT: Serra do Mar in Rio de ]aneiro and Sio Paulo, Brazll.
Sphaenorhynchus palustris Bokermann, 1,966. Rev. Brasil. BioI., 26:LG.

rvrr(s): Holotype: WCAB L9235.
TypE LocAttry: Sooretama, Linhares, Espirito Santo, Brazil.
sphaenorhynchus pauloalaini Bokermann, 1973. Rev. Brasil. Biol., z3:592.
TypE LocAr.my: Centro de Pesquisas do Cacau, I1h6us, Bahia, Brazil.
DISTRIBUTToN: Known only from the type locality.
Sphaenorhynchus planicola (A. Lutz and B. Lutz,1938). An. Acad. Brasil. Cienc., 10:182.
oRIGINAL NAME: Hyla (Sphoenohyla) planicola.
rvnE(s): Holotype: AL-MN 2558.
TypE LocALrry: Recreio dos Bandeirantes, Rio de ]aneiro, Brazil.
DISTRIBUTIoN: Coastal lowlands of Rio de ]aneiro, southeastern Brazil.
Sphaenorhynchus prasinus Bokermann, 1973. Rev. Brasil. Biol., 33:589.

TypE LocAlmy: Centro de Pesquisas do Cacau, I1h6us, Bahia, Brazil.
Sphaenorhynchus surdus (Cochran, 1953). Herpetolo gica, 8:1L2.
oRIGINAL NAME: Hyla aurantiaca surda.
rvrE(s): Holotype: UMMZ L06736.
TypE LocAlrry: Curitiba, ParanA, Brazil
Trachycephalus Tschudi, 1838. Classif. Batr.:7 4.
TypE spECIus: Trachycephalus nigromaculafzs Tschudi, 1838, by monotypy.
DISTRIBUTIoN: Eastern Brazil and Pacific lowlands from southern Colombia to
northern Peru.
coMMENr: See Trueb, L970, Univ. Kansas Publ. Mus. Nat. Hist., L8:546-7L6.
c eph alus atl as Bokerman n, 19 66. Neotropic a, 1,2:L20.

T r a chy
TYPE LocALltv: Fazenda Santo Onofre, 10 km east of Marac6s, Bahia, Brazil.
DISTRIBUTToN: Northeastern Brazil.
Trachycephalus jordani (Stejneger and Test, 1891). Proc. U.S. Natl. Mus., l4:L67.
oRIGINAL NAME: Tetraprion iordani.
rvrr(s): Holotype: USNM L2274.
TypE LocAurv: Guayaquil, Provincia Guayas, Ecuador.
DrsrRrBUrrow: Pacific lowlands of South America from southern Colombia to
northern Peru.
Trachycephalus nigromaculatus Tschudi, 1838. Classif. Batr.:7 4.

rvnr(s): Syntypes: MNHNP 4608-9.
TypE LocALITy: "America meridionalis".
DrsrRrBUTroN: Coastal regions of southern Brazil from Espirito Santo to Sio Paulo
interior in Minas Gerais and Goi6s.
Triprion Cope, L866. Proc. Acad. Nat. Sci. Philadelphia, L8:127.

rypE spECIr;s: Pharyngodon petasatus Cope, 1865, by monotypy.
DrsrRrBUTrow: Pacific lowlands of Mexico; Yucatan Peninsula of Mexico and
Guatemala.
CoMMENT: For discussion see Duellman, L970, Monogr. Mus. Nat. Hist. Univ.
Kansas, L:628-640. Triprion is a replacement name f.or Pharyngodon Cope, L865,
Proc. Acad. Nat. Sci. Philadelphia,lT:I93, which is preoccupied.
Triprion petasatus (Cope, 1865). Proc. Acad. Nat. Sci. Philadelphia, L7:L93.
oRIGINAL NAME: Pharyngodon petasatus.
rvrr(s): Holotype: USNM 12287.
TypE LocAtttv: Cenote Tamach6, L7 km north of M6rida, YucatAn, Mexico.
DISTRIBUTIoN: Yucatan Peninsula, Mexico, through El Pet6n, Guatemala, to
northern Honduras.
Triprion spatulatus Gtinther, 1882. Ann. Mug. Nat. Hist., (5)L0:279.

rvnr(s): Syntypes: BM L947 .2.25.79-BL.
TypE LocAt-Itv: Presidio de MazatlAn, Sinaloa, Mexico.
DrsrRrBUTroN: Pacific lowlands of western Ltfexico (Sinaloa to Oaxaca).
CoMMENT: Two subspecies were recognized by Duellman, L970, Monogr. Mus.
Nat. Hist. Univ. Kansas, l:629-637.
suBFAMILy: Pelodryadinae Gtinther, L859 "L858".

cIrArIoN: Cat. Batr. Sal. Coll. Brit. Mus.:119.
DISTRIBUTIoN: The Australo-Papuan region.
coMMENr: As originally formed the group name was Pelodryadidae. On the basis of
work by Tyler, L97L, Univ. Kansas Publ. Mus. Nat. Hist., L9:3L9-360, Savage, 1973
lnYiaI (ed.), Evol. Biol. Anurans:351-445, considered this grouP independently
derived from a "leptodactylid" stock in Australia (convergent with the Hylidae)
and placed it in a distinct family, the Pelodryadidae. This arrangement was,
followed by Fouquette and Delahoussaye, L977 , J. Herpetol., Ll:387 -396, and
Goin, Goin, and Zug, L978,Intr. Herpetol., Ed. 3. The Pelodryadidae was
considered a subfamily of the Hylidae by Duellman, L977, Das Tierreich, 95:1-
225, an arrangement followed by most subsequent authors who recognize this
group. Cogger, Cameron, and Cogger, L983, Zool. Cat' Aust., l(Amph. Rept.):35-
5L, supplied synonymies and literature reviews for the Australian species.
REVTEwERS: Margaret Davies (MD); Michael ]. Tyler (MIT); Richard G. Zweif.el (RGZ)
(New Guinea).
Cyclorana Steindachner, L867. Reise Freg. Novara, Amph.:29.

TypE spECIrs Cyclorana nooaehollandiae Steindachner, L867, by monotypy.
Australia, except extreme southeastern and southwestern parts.
DISTRTBUTIoN:
coMMENr: Australian frogs of the Cyclorana australis group were reviewed by Tyler
and Martin,1975, Trans. R. Soc. S. Aust., 99:93-99, and other species by Tyler
and Martin, 1977, Rec. S. Aust. Mus., L7:26L-276. Tyler, Davies, and King, 1979
ANURA: HYLIDAE: PELODRYADINAE 177
Aust. J. Zool.,27:699-708, reported on karyology. Cyclorana was omitted from

the Myobatrachidae by Heyer and Liem, 1976, Smithson. Contrib. Zool.,233:Ll,
thereby implying transfer to the Hylidae. Transferred formally by Tyler, L978,
Amph. S. Aust.
Cyclorana australis (Gray, L842). ZooL. Misc., 2:56.

oRIGINAL NAME: Alytes australis.
TypE LocAt-rry: "North coast of Australia"; restricted to Port Essington, Northern
Territory, Australia, by Tyler and Martin, L975, Trans. R. Soc. S. Aust., 99:94.
DrsrRrBUTIoN: Kimberley Division (Western Australia) through northern Australia
as far as the Gulf District of Queensland, Australia.
Cyclorana breaipes (Peters, L87L). Monatsber. Preuss. Akad. Wiss. Berlin, 187L:648
oRIGINAL NAME: Chiroleptes breaipes.
rvnr(s): Holotype: ZMB 7063 (not located).
rypE LocAlIrv: Port Bowen [:Port Clinton] Queensland [Australia].
DISTRIBUTIoITi: Northeastern and Gulf region of Queensland, Australia.
coMMENr: Redefined by Tyler and Martin, L977, Rec. S. Aust. Mus., 17:263.
Cyclorana cryptotis Tyler and Martin, L977. Rec. S. Aust. Mus., 17:271.
rYPE(s): Holotype: SAMA R74716.
TypE LocAlIrv: Daly Waters, Northern Territory, Australia.
DISTRIBUTIow: Western Australia and Northern Territory, between latitudes L7" 20'
and30'S.
14o
coMMENr: Redefined by Tyler, Davies, and Martin, L982, Copeia, L982:260.
Cyclorana cultripes Parker, L940. Novit. Zool., 42:22.

rvnE(s): Holotype: BM L947 .2.18.45.
TypE LocAlrrv: Alexandria Station, Northern Territory [Australia].
DISTRIBUTIoN: Lower eastern half of Northern Territory to southwestern
Queensland and possibly northern South Australia, Australia.
coMMENT: See Tyler and Martin, L977, Rec. S. Aust. Mus., 17:26L-276.
Cyclorana longipes Tyler and Martin, 1977. Rec. S. Aust. Mus., 17:271.
rYPE(s)r Holotype: WAM R43258.
TypE LocAltrv: Mitchell Plateau (14' 52' 5,125" 50' E), Kimberley Division,
Western Australia.
DrsrRrBUrroN: Kimberley region of Western Australia and adjacent Northern
Territory to porthern Queensland, Australia.
Cyclorana maculosa Tyler and Martin, L977. Rec. S. Aust. Mus., L7:269.
oRIGINAL NAME: Cyclorana maculosus.
rvrr(s): Holotype: SAMA RL4719.
TypE LocAlIrv: Daly Waters, Northern Territory, Australia.
DrsrRrBUTroN: Northeastern Northern Territory to northwestern Queensland,
Australia.
Cyclorana maini Tyler and Martin, 1977 . Rec. S. Aust. Mus., 17:273.
rvnr(s): Holotype: SAMA Rl5191.
TypE LocAutv: Barrow Creek, Northern Territory, Australia.
DrsrRrBUTroN: Southern half of Northern Territory through central Western
Australia to the western coastal region, Australia.
Cyclorana manya van Beurden and McDonald, 1980. Trans. R. Soc. S. Aust., 104:L93.
rYPE(s): Holotype: QM 134886.
TypE LocAury: Between Coen airport and Deep Creek crossing, 25 km N of Coen
township (13'52' S, 143o 12' E), Queensland, Australia.
DISTRIBUTIoN: Cape York Peninsula, Queensland, Australia.
Cyclorana nooaehollandiae steindachner, 1867. Reise Freg. Novara, Amph.:29.

rYPE(s): Syntypes: NHMW 47L6-L7 (probable)'
TYPE LocALITY; Rockhampton, Queensland [Australia]'
DrsrRrBUrroN: Cape York Peninsula (Queensland) to the New South Wales border
and interiori of Queensland and New South Wales, Australia.
q9MMENT: In the Cyclirana australis complex; see Tyler and Martin, L975, Trans. R'
Soc. Aust.,99:93-99.
cyclorana platycephala (Gtinther, 1873). Ann. Ma8. Nat. Hist., (4)L1:350.
oRIGINAL NAME: Chiroleptes platycephalus.
rvrE(s): Holotype: BM L947 .2.L8.42.
TypE LoCAgrv: Fort Bourke [New South Wales, Australia].
DISTRIBUTIoT: Drier inland regions of Australia, except Victoria'
coMMENT: See comment under Cyclorana slettini'
Cyclorana sleaini Loveridge, 1950. Proc' Biol. Soc. Washington, 63:131'
rvnr(s): Holotype: CAS 82052.
TypE LOCAr.Irv, Noondoo, southeastern Queensland, Australia.
coMMENr: Cyclorana sleaini was includea as a syn_onym of Cyclorana platycephala bv
CoggeriCameron, and. Cogger,1983, Zool. Cat. Aust., l(Amph. Rept.):37,
with no discussion.
Cyclorana oagitus Tyler, Davies, and Martin, 1981. Rec. W. Aust. Mus', 9:148'
rvru(s): HolotyPe: WAM R7L037.
rypE LocAlrrv: ''junction of the Great Northern Highway and the road to Derby,
41 km S of berby,124" 38' E, L7" 44' S, Kimberley Division", Western
Australia.
DTSTRTBUTToN: Kimberley Division, Western Australia, and northwestern Northern
Territory, Australii. The grammatically incorrect epithet aagita has
been used.
Cyclorana aerrucosa Tyler and Martin, 7977. Rec. S. Aust' Mus., L7:257.
ORIGINAL NAME: Cyclorana aerrucosus
rvnr(s): Holotype: QM J18105.
TypE LocAr-lrv, 18 km W of Dalby, Queensland, Australia'
DrsrRrBUTroN: Interior basins of Queensland and New South Wa1es, Australia.
Litoria Tschudi, 1838. Classif. Batr.:36.

rypE spECIns: Litoria freycineti Tschudi, 1838, by monotypy.
DISTRIBUTIoN: New bri.,eu, Moluccan Is., Lesser Sunda Is., Timor, Bismarck
Archipelago, Solomon Is., and Australia, including Tasmania; introduced into
New Caledonia, New Hebrides, Guam, and New Zealand'
g9MMENT: Species groups noted in the species accounts were defined by Tyler and.
Davies, 1978, ^{.ust.-}. Zool., Suppt. Ser., 63:1-47. Papuan species were reviewed
(as Hyla) by Tyler, L968, Zool. Verh., Leiden, 96:L-2A3; Australian state
checklists provided by Tyler, L982, Frogs, Ed' 2'
Litoria adelaidensis (Gray, 1841). Ann. Ma8. Nat' Hist', (2)7:90'
oRIGINAL NAME: Hyla adelaidensis.
rYPE(s): Holotype: BM L947 .2.22.80.
"western Australia"
TYPE LoCALITY: .
Southwestern Western Australia.

DISTRIBUTIoN:
g9MMENT: Discussed by Tyler, Smith, and ]ohnstone, L984, Frogs W. Aust. In the
Lit oria adelaid ensis grouP.
Litoria alboguttata (Gtinther, L867). Ann. Ma8. Nat. Hist., (3)20:54.

oRIGINAL NAME: Chiroleptes alboguttatus.
rypE(s): Syntypes: BM L947.2.26.0-Z (formerly 67.5.6.78-79) (Cape York),
Lg47-.2.18.50-51 (formerly 1936. L2.3.125-126) (Port Denison).
TypE LoCALlry: Port Denison and Cape York, Queensland, Australia.
DTSTRTBUTION: Queensland and New South Wales, Australia.
coMMENT: In the Litoria aurea group. Transferred to Litoria from Cyclorana by Tyler,
1974, Proc. R. Soc. Queensland, 85:27. Maxson, Tyler, and Maxson, 1982, Aust.
l. Zool., 30:643, discussed relationships with Cyclorana.
Litoria albolabris (Wando1leck, 1911). Abh. Ber. K. Zool. Anthro. Ethno. Mus. Dresden,
L3:12.
oRIGINAL NAME: Hyla albolabris.
rvnr(s): NHMW L6976 and SAMAR4947 considered syntypes by Duellman, L977,
Das Tierreich, 95:11.4; Obst, L977, Zool. Abh. Staatl. Mus. Tierkd. Dresden,
34:173,listed MTKD D2220 as holotype (destroyed) and a paratype
in NHMW.
TypE LocALITy: "Eitap6" (:Aitape), West Sepik Province, Papua New Guinea.
coMMENr: In the Litoria albolabris group.
Litoria amboinensis (Horst, 1883). Notes Leyden Mus., 5:239.
oRIGINAL NAME: Hyla amboinensis.
rvnr(s): Syntypes: RMNH 4418 (Amboina, 2 specimens), 44L9 (Misool).
TypE LoCALITY: "Amboina" and "Misool", Moluccas, Indonesia.
DrsrRrBUTroN: New Guinea (southern lowlands, Huon and Vogelkop peninsulas)
and Moluccan Is. (Amboina, Ceram, Misool).
coMMENT: For discussion see Tyler, L968, Zool. Verh., Leiden, 96:27.In the Litoria
peronii group.
Litoria angiana (Boulenger, 1915). Ann. Mag. Nat. Hist., (8)L6:402.
oRIGINAL NAME: Hyla angiana.
rvnE(s): Syntypes: BM L947.2.24.6 and L947.2.30.95-98.
rypE LocALITy: "Angi Lakes (6000 feet) in the Arfak Mountains, . . . Mt. Koebr6
(8000 feet) in the same mountains", Vogelkop Peninsula, Irian Jaya (New
Guinea), Indonesia.
DTsTRIBUTIoN: Mountains of New Guinea.
angiana group.
Litoria arfakiana (Peters and Doria, L878). Ann. Mus. Civ. Stor. Nat. Genova, L3:42L.
oRIGINAL NAME: Hyla (Litoria) arfakiana.
rvnr(s): MSNG 29723A designated lectotype by Capocaccia, 1957 , Ann. Mus. Civ.
Stor. Nat. Genova, 69:213.
TypE LocALITy: "Hatam", Arfak Mountains, Vogelkop Peninsula, Irian laya,
Indonesia (New Guinea).
DISTRIBUTIow: Highlands of New Guinea.
coMMENT: In the "Litoria arfakiana group. See Menzies and Zweif.el, 1974, Am. Mus.
Novit., 2558:1-16.
Litoria aruensis (Horst, 1883). Notes Leyden Mus., 5:242.
oRIGINAL NAME: Hyla aruensis.
rvnE(s): Syntypes: RMNH 4416 (Aru, 2 specimens),44L7 (Misool).
TypE LocALITy: "Aroe-Islands" (:Aru) and "Misoo1", Irian Jaya (New Guinea) and
Moluccas, New Guinea.
DrsrRrBUTroN: New Guinea, Moluccan Is. (Waigeo, Misool), Aru Is., and Louisiade
Archipelago.
aruensis group.
Litoria aurea (Lesson, 1830). Voy. Coquille, Zool.,2:60.
oRIGINAL NAME: Rana aurea.
rvnE(s): MNHNP 4857 tisted as lectotype by Roux-EstEve, L979, BulI. Trim. Soc.
G6ol. Normandie et Amies Mus. F{arve, 66:26.
TypE LocAlrrv: Macquarie River, Bathurst, New South Wales, Australia; lectotype
from Port ]ackson (:Sydney), New South Wales, Australia.
DrsrRrBuTrorrl: Eastern New South Wales, south of the Richmond River, Australia.
coMMENT: In the Litoria aurea group. See Courtice and Grigg, L975, Aust. Zoo- -:
L49-L63, and Maxson, Tyler, and Maxson, L982, Aust. J. Zool.,30:643. CoE:::
Cameron, and Cogger, 1983, Zool. Cat. Aust., l(Amph. Rept.):39, discussej.
types and citation of the description.
Litoria becki (Loveridge, 1945). Proc. Biol. Soc. Washington, 58:53.

oRTGTNAL NAME: Hyla becki.
rypE LocALITy: "Mt. Wilhelm, 5000-8000 feet, Bismarck Range, Madang Divis:::
Chimbu Province, Papua New Guinea.
coMMENT: For discussion see Tyler, L968, Zool. Verh., Leiden, 96:47.In the L::-*;
becki group.
Litoria bicolor (Gray, L842). Zool. Misc., 3:57.

oRIGINAL NAME: Eucnemis bicolor.
TypE LocALIrv: Port Essington, Northern Territory, Australia.
DrsrRrBUTIoN: Coastal Australia from Western Austra'lia to northeastern
Queensland; southern lowlands of New Guinea; Aru Is.
coMMENT: For discussion see Straughan, 1969, Proc. R. Soc. Queensland, 80:{3
Litoria booroolongensis (Moore, 196L). Bull. Am. Mus. Nat. Hist., L2L:292.
oRIGINAL NAME: Hyla booroolongensis.
rvrr(s): Holotype: AM R16006.
TypE LocAurv: Guy Fawkes Creek, Ebor, New South Wales, Australia.
DISTRIBUTIoN: Eastern New South Wales, Australia.
coMMENT: In the Litoria booroolongensis group.
Litoria breaipalmata Tyler, Martin, and Watson, 1972. Proc. Linn. Soc. New South.
Wales, 97:82.
rvnr(s): Holotype: SAMA R11236.
TypE LocAlrrv: Ourimbah Creek, 5 mi. northwest of Gosford, New South \A'al,-
Austraiia.
DISTRIBUTIow: Eastern New South Wales, Australia.
coMMENT: In the Litoria breaipalmata group.
Litoria brongersmai (Loveridge, L945). Proc. Biol. Soc. Washington, 58:56.

oRIGINAL NAME: Hyla brongersmai.
TypE LocALITy: "Parana [:Panara] Valley, central Dutch New Guinea", near
Doorman Top, Snow Mountains, Irian ]aya (New Guinea), Indonesia.
coMMENr: See Tyler, 1968, Zool. Verh., Leiden, 96:54. Questionably in the Lit-*-.r
becki group.
Litoria bulmeri (Tyler, L968). Zool. Verh., Leiden, 96:56.

oRTGTNAL NAME: Hyla bulmeri.
rvnr(s): Holotype: SAMA R5625.
TypE LocALtry: " . . . at an altitude of approximately 7300 ft at GIkm, upper
Aunjung Valley, Schrader Mountains, Eastern Mountains, [Madang Pror-::::
Papua] New Guinea".
DISrRIBUrIotrt: Known only from the type locality.
coMMENr: In the Litoria bulmeri group.
Litoria burrowsi (Scott, L942). Rec. Queen Victoria Mus., 1:5.

oRTGINAL NAME: Hyla burrowsi.
rvrr(s): Holotype: QVM L94L.4L.
rypE LocAurv: Cradle Valley, Tasmania [Australia].
DISTRIBUTIoIS: Southwestern and western Tasmania, Australia
coMMENr: In the Litoria maculata group. See Martin and Littlejohn, L982, Tasman.
Amph.:30, for discussion.
Litoria caerulea (White, L790). J. Voy. New South Wales:248.

oRTGTNAL NAME: Rana caerulea.
rvnr(s): Formerly in Hunterian Mus. (presumed lost); see Tyler and Dobson, L973,
Herpetolo gica, 29 :37 3 -37 5.
TypE LocAtITv: New South Wales, Australia.
DrsrRrBUTroN: Northern and eastern Australia; islands in the Torres Straits; New
Guinea (northern lowlands, southern lowlands, and Vogelkop Peninsula);
New Zealand (introduced).
coMMENr: See Tyler, 1968, Zool. Verh., Leiden, 95:60, and Tyler, Crook, and
Davies, L983, Rec. S. Aust. Mus., 18:424.In the Litoria caerulea grouP. Without
discussion, Cogger, Cameron, and Cogger, 1983, Zool. Cat. Aust., l(Amph.
Rept.):43, recognized Litoria gilleni (:Hyla gilleni Spencer, L896, Rep. Horn
Exp. Cent. Aust.,2:L73; Holotype: NMM D9827; TyP" locality: Alice Springs,
Northern Territory, Australia), usually considered a subspecies or synonym of
Litoria caerulea, as a distinct species.
Litoria capitula (Tyler, 1968). ZooI. Verh., Leiden, 96:64.

oRIGINAL NAME: Hyla capitula.
rvnE(s): Holotype: RMNH 53L7.
TypE LocALITy: "Sam1akki, Tenimber Island", Moluccan Is., Indonesia
DISTRIBUTIoN: Tenimber I., Moluccan Is., Indonesia.
coMMENr: In the Litoria rubella group.
Litoria caaernicola Tyler and Davies,7979. Trans. R. Soc. S. Aust., L03:149.

rvnr(s): Holotype: WAM R43228.
TypE LocAurv: 3 km west of Surveyor's Pool, Mitchell Plateau, Kimberley
Division, Western Australia, Australia.
DrsrRrBUTroN: Gorges bordering the Mitchell Plateau, northern Western Australia.
coMMENT: In the Litoria caerulea grouP. See Tyler, Smith, and ]ohnstone, 1984,
Frogs W. Aust.
Litoria chloris (Boulenger, 1893). Proc. Linn. Soc. New South Wales, (\7:a03
oRTGINAL NAME: Hyla chloris.
TypE LocALItv: Dunoon, Richmond River, New South Wales, Australia.
DISTRIBUTIoN: Eastern Australia (Queensland and New South Wales).
coMMENT: In the Litoria aruensis grouP.
Litoria chloronota (Boulenger, L9L1). Ann. Mag. Nat. Hist., (8)8:55.

oRIGINAL NAME: Hylella chloronota.
rvnr(s): Syntypes: BM L947 .2.3L.20-21.
TypE LocAt-rry: "Arfak Mountains at an altitude of 8000 feet", Vogelkop Peninsula,
Irian Jaya (New Guinea), Indonesia.
coMMENT: See Tyler, 1968, Zool. Verh., Leiden, 96:56.In the Litoria bicolor grouP.
Litoria citropa (Dum6ril and Bibron, 1841). Erp. G6n., 8:600.

oRIGINAL NAME: Hyla citropa.
TypE LocALITy: "Nouvelle-Hollande ... au Port ]ackson [:Sydney]", New South
Wales, Australia.
DrsrRrBUTrow:Mountains and coastal areas of New South Wales and mountains of
Victoria in southeastern Australia.
coMMENT: In the Litoria citropa group. See comment under Litoria jenolanensls. See
also Tyler and Anstis,1975, Rec. S. Aust. Mus., 17:41'. Dendrohyas citropa
Tschudi, 1838, Classif. Batt;7i, is a nomen nudum according to Cogger,

Cameron, and Cogger, L983, Zool. Cat. Aust., l(Amph. Rept.):4l.
Litoria congenita (Peters and Doria , L878). Ann. Mus. Civ. Stor. Nat. Genova, 13:427
oRIGINAL NAME: Hyla (Litoria) congenita.
rypE(s): MSNG 9722A designated lectotype by Capocaccia, 1957, Ann. Mus. Civ'
Stor. Nat. Genova, 69:2L3.
"Yule Island", Central Province, Papua New Guinea.
TypE L96ALITY:
Southern coastal lowlands of New Guinea; Aru Is.
DTSTRIBUTION:
69MMENT: See Ty1er , L958, ZooL. Verh., Leiden, 96:68.In the Litoria rubella SrouP
Litoria contrastens (Ty1er, 1968). Zool. Verh., Leiden, 96:72.

oRIGINAL NAME: Hyla contrastens.
rvrr(s): Holotype: SAMA R5845.
7'Barabttna", near Kundiawa, eastern mountains, Chimbu Province
TypE LocALITy:
Papua New Guinea.
DISTRIBUTIoN: Eastern mountains, Papua New Guinea.
coMMENT: In the Litoria bicolor grouP.
Litoria cooloolensis Liem, L974. Mem. Queensland Mus., 17:169.

TypE LocAlrrv: Lake Coolamera, Cooloo1a, southeast Queensland, Australia
DISTRIBUTIoN: Southeastern Queensland, Australia.
coMMENrr In the Litoria bicolor grouP.
Litoria coplandi (Tyler, 1968). Rec. S. Aust. Mus., L5:4L6.

ORIGINAL NAME: Hyla coplandi.
rvnr(s): Holotype: WAM RL3722G
TypE LocALtrv: Inverway Station, Northern Territory, Australia.
DrsrRrBUTroN: Northwestern Western Australia and Northern Territory, Australia
coMMENr: In the Litoria coplandi grouP.
Litoria cyclorhynchas (Boulenger, 1882). Cat. Batr. Sal. Brit. Mus.:411.

oRIGINAL NAME: Hyla aurea var. cyclorhynchus.
rypE(s): Syntypes: AMNH 62493, BM 69.7.22.3-7.
TYPE LocALITY: "W. Australia".
DISTRIBUTIow: Southern coastal Western Australia.
q9MMENT: In the Litoria aurea group. Discussed by Tyler, Smith, and Johnstone,
1984, Frogs W. Aust.
Litoria dahlii (Boulenger, L896). Proc. Zool. Soc. London, L895:867 .
oRIGINAL NAME: Chiroleptes dahlii.

rypE(s): syntypes: BM L947.2.L8.49 (formerly 95.LL.L4.25), UZMO K2002.
TypE LocAltrv: Daly River, Northern Territory, Australia.
DrsrRrBUTrow: Northeastern Western Australia, Northern Territory, and
northwestern Queensland, Australia.
g9MMENT: In the Litoria aurea group. For discussion see Tyler, Davies, and King,
1978, Trans. R. Soc. S. Aust., L02:L7-24.
Litoria darlingtoni (Loveridge, L945). Proc. Biol. Soc. Washington, 58:53.

oRIGINAL NAME: Hyla darlingtoni.
TypE L9CALITY: "Mount Wilhe1m, 5000-8000 feet, Bismarck Range, Madang
Division, [Chimbu Province, Papua] New Guinea ".
DISTRIBUTIoN: Eastern mountains of New Guinea.
coMMENr: See Tyler, 1968, Zool. Verh., Leiden, 96:76.In the Litoria peronii grouP
Litoria dentata (Keferstein, 1868). Arch. Naturgesch., 34:284.

oRIGINAL NAME: Hyla dentata.
rvnr(s): Holotype: probably ZFMK (formerly Zool. Mus. Gottingen l23a),
according to Cogger, Cameron, and Cogger, L983, ZooL Cat. Aust., l(Amph.
Rept.):42.
TypE LocAtItv: New South Wales, Australia.
DrsrRrBUTroN: Eastern New South Wales and southeastern Queensland, Australia.
coMMENr: See Moore, L96L, Bull. Am. Mus. Nat. Hist., L2:275.In the Litoria rubella
grouP.
Litoria dorsalis Macleay, 1877. Proc. Linn. Soc. New South Wales, 2:138.
rvnr(s): Unknown.
TypE LocALITy: "Katow", near mouth of Binaturi River, Western Province, Papua
New Guinea; stated as "Mawatta, Binaturi River (as Katow), Papua New
Guinea", by Cogger, Cameron, and Cogger, L983, Zool. Cat. Aust., l(Amph.
Rept.):42.
DISTRIBUTIoN: Southern lowlands, New Guinea.
coMMENT: In the Litoria dorsalis group; see Tyler, 1958, ZooI. Verh., Leiden, 96:80.
See comment under Litoria microbelos.
Litoria dorsiaena (Tyler, L968). Zool. Verh., Leiden, 96:83.

oRIGINAL NAME: Hyla dorsiaena.
rvnE(s): Holotype: SAMA R7901.
TypE LocALrry: "Telefomin, Eastern Mountains, [West Sepik Province, Papua] New
Guinea".
DISTRIBUTIow: Eastern and Arfak mountains, New Guinea.
coMMENT: In the Litoria dorsiaena group.
Litoria eucnemis (Lbnnberg, 1900). Ann. Mag. Nat. Hist., (7)6:579.

oRIGINAL NAME: Hyla eucnemis.
rvnE(s): Syntypes: BM 1947.2.23.87, ZIUU (no number).
TypE LocAurv: Sattelberg, Huon Peninsula, New Guinea.
DrsrRrBUTroN: Southern lowlands and Huon Peninsula, New Guinea; northeastern
coastal Queensland, Australia.
coMMENT: In the Litoria eucnemis group; see Tyler, 1968, Zool. Verh., Leiden, 96:86
With no discussion, Australian populations were recognized as a distinct
species, Litoria Serrata, by Cogger, Cameron, and Cogger, 1983, Zool. Cat.
Aust., l(Amph. Rept.):49. See Tyler, L965, Proc. ZooI. Soc. London, 145:91-
106. See comntent under Litoria genimaculata.
Litoria eaeretti (Boulenger, L897). Ann. Mag. Nat. Hist., (5)19:509.

oRIGINAL NAME: Hyla eaeretti.
rvnr(s): Syntypes: BM 1947 .2.23.60-65 (Savu), 1947.2.23.66-67 (Sumba).
TypE LocAttrv: Savu and Sumba Is., Indonesia.
DISTRIBUTIoN: Timor and adjacent Lesser Sunda Is., Indonesia.
coMMENr: See Tyler , 1968, Zool. Verh., Leiden, 96:90. In the Litoria peronii grouP.
Litoria ewingii (Dum6ril and Bibron, L841). Erp. G6n., 8:597.

oRIGINAL NAME: Hyla Ewingii.
TypE LocALITY: "la terre de Van Di6man" 1:1'utmania), Australia.
Australia and Tasmania; introduced into New Zealand.
DrsrRrBUTroN: Southeastern
coMMENr: See Moore, L96L, Bull. Am. Mus. Nat. Hist.,12:278.In the Litoria ewingii
8rouP.
Litoria fallax (Peters, 1881). Monatsber. Preuss. Akad. Wiss. Berlin, L88L:224.
oRIGINAL NAME: Hylomantis fallax.
rvnr(s): Syntypes: ZMB 5887 (2 specimens), 5925 (2 specimens), 6633 (2
specimens), 9650.
TypE LocArtrv: Port Bowen [:Port Clinton], MacKay, and Rockhampton,
Queensland, Australia.
DISTRIBUTIoN: East coast of Australia and New South Wales.
coMMENr: In the Litoria bicolor group. For discussion (as Litoria glauerti) see
Straughan, 1969, Proc. R. Soc. Queensland, 80:43-54.
Litoria flaaipunctafa Courtice and Grigg, 1975. Aust. Zool., 18:159.

rveE(s): Holotype: AM R40676.
TypE LocArrtv: Booralong Creek Road, 12.8 km W of Guyra, New South Wales,
Australia, 30" 16' S, 151o 33' E.
DISTRIBUTIoN: New England Tableland, New South Wales, Australia.
coMMENT: In the Litoria aurea group. Cogger, Cameron, and Cogger, L983, Zool.
Cat. Aust., l(Amph. Rept.):40, resurrected the name Litoria castanea (:Hyla
castanea Steindachner, 1867 , Reise Freg. Novara, ,A,mph.:67; Types: unknown;
Typ" locality: unknown) and regarded Litoria flaztipunctata as a junior
synonym. We here retain Litoria flaoipunctata f.or this taxon because none of
the collecting localities of the 'Novara' expedition (Gans, L955, Ann. Carnegie
Mus., 33:280) are within the known range of Litoria flaoipunctafa (MlT and
MD).
Litoria freycineti Tschudi, 1838. Classif. Batr.:77.

rvnr(s): Syntypes: MNHNP 246 (2 specimens).
TYPE LocALITY: "Australia".
DISTRIBUTIow: East coast of southern Queensland and New South Wales, Australia
coMMENr: See Moore, 1.96L, BuIl. Am. Mus. Nat. Hist., 12:302.In the Litoria
freycineti group.
Litoria genimaculafa (Horst, 1883). Notes Leyden Mus., 5:240.

oRIGINAL NAME: Hyla genimaculata.
rYPE(s): Holotype: RMNH 4420.
TypE LocAlrrv: "Gebeh" 1:6"tie?) I., west of Waigeo I., Indonesia.
DISTRIBUTIoN: Lowlands of southern New Guinea and adjacent islands.
coMMENT: See Tyler, 1968, Zool.. Verh., Leiden, 96:94.In the Litoria eucnemis
group. Ingram and Covacevich, 198L, Mem. Queensland Mus., 20:29L-396,
without discussion, regarded Australian specimens of. Litoria eucnemis as
representing this species.
Litoria gracilenta (Peters, 1869). Monatsber. Preuss. Akad. Wiss. Berlin, L869:789.
oRIGINAL NAME: Hyla gracilenta.
rvnE(s): Holotype: ZMB 6618.
TypE LocArrry: Port Mackay, Queensland, Australia.
DISTRIBUTIoN: East coast of Australia from
the Cape York Peninsula to New South
Wales; eastern mountains and southern lowlands of New Guinea.
coMMENr: See Moore, 196L, Bull. Am. Mus. Nat. Hist. , L2:265, and Tyler, L968,
Zool. Verh., Leiden, 95:97.In the Litoria aruensis group.
Litoria graminea (Boulenger, 1905). Ann. Mag. Nat. Hist., (7)L6:L83.

oRIGINAL NAME: Hyla graminea.
rvnr(s): Holotype : BM L947 .2.23.3L.
TypE LocAl-rry: "Northern British New Guinea, altitude 900 feet"
DISTRTBUTToN: New Guinea.
coMMENT: See Tyler, L968, Zool. Verh., Leiden, 96:100.
Litoria impura (Peters and Doria, L878). Ann. Mus. Civ. Stor. Nat. Genova, L3:426.
oRIGINAL NAME: Hyla (Litoria) impura.
rvnr(s): MSNG designated lectotype by Capocaccia, L957, Ann. Mus. Civ. Stor.
Nat. Genova,69:2"1.8.
TYPE LoCALITY: "Yule Island in Nova Guinea australi".
DISTRIBUTIoN: Southeastern New Guinea.
coMMENT: In the Litoria thesaurensls group. See Menzies and Ztg, L979,
Micronesica, 15:325-333.
Litoria inermis (Peters, L867). Monatsber. Preuss. Akad. Wiss. Berlin, L867:30.
oRIGINAL NAME: Chiroleptes inermis.
rypE(s): Syntypes: RMNH 1888, AMNH 23582, NHMW 18384 (2 specimens), ZMB
5589; Cogger, Cameron, and Cogger, L983, Zool. Cat. Aust., l(Amph. Rept.):
44, regarded ZMB 5589 as the holotype and regarded the putative syntype
status of other specimens as in error.
TypE LocAurv: Rockhampton, Queensland, Australia.
DISTRIBUTIoIt: Northern and northeastern Australia.
coMMENr: See Straughan, 1969, Proc. R. Soc. Queensland, S0:207.In the Litoria
latopalmata group. Davies, Martin, and Watson, L983, Trans. R. Soc. S. Aust.,
1.07:87, redefined the species.
Litoria infrafrenata (Gtinther, L867). Ann. Mag. Nat. Hist., (3)20:56.

oRIGINAL NAME: Hyla infrafrenata.
rvnE(s): Holotype: BM 1947 .2.24.L7.
TypE LocArrrv: Cape York Peninsula, Queensland, Australia.
DTsTRIBUTIoN: Cape York Peninsula, Queensland, Australia, and New Guinea; New
Ireland, Bismarck Archipelago; introduced into |ava.
coMMENr: See Tyler, L968, ZooL. Verh., Leiden, 96:104.In the Litoria infrafrenata
8rouP.
Litoriairis (Tyler, L962). Rec. S. Aust. Mus., L4:253.
oRIGINAL NAME: Hyla iris.
rvnE(s): Holotype: BM L95L.L206.
TypE LocALITy: "Bamna, near Nondugl (Lat. 5" 49' S; Long. 144" 44' E) at 6,500
feet", Western Highlands Province, Papua New Guinea.
DISTRIBUTIow: Snow and Eastern mountains, New Guinea.
coMMENT: See Tyler,1968, Zool. Verh., Leiden, 96:LlL. In the Litoria nigropunctata
grouP.
Litoria jenolanensis (Copland, L957). Proc. Linn. Soc. New South Wales, 82:97.
oRIGINAL NAME: Ityla jenolanensis.
rvnn(s): Holotype: AM R14412.
TypE LocAury: Bottomless Pit, |enolan Caves, New South Wales, Australia.
DISTRIBUTIoN: New South Wales, Australia.
coMMENT: With no discussion, considered a synonym of Litoria citropa by Cogger,
Cameron, and Cogger, 1983, Zool. Cat. Aust., 1(Amph. Rept.):4l.
Litoria jeruisiensis (Dum6ril and Bibron, 1841). Erp. G6n., 8:580.

oRIGINAL NAME: Hyla jeroisiensis.
TypE LocALITy: "la baie de Jervis" (:Jervis Buy), New South Wales, Australia.
DISTRIBUTIoN: Southeastern New South Wales, Australia.
coMMENT: In the Litoria ewingii group. See White, Whitford, and Watson, 1980,
Aust. Zool.,20:375-390, for discussion of identity of this species.
Litoria jeudii (Werner, 1901). Zool. Anz.,24:99.

oRTGTNAL NAME: Hyla jeudii.
rvnr(s): Holotype: ZMB L6498.
TypE LocALITy: "German New Guinea" (:northern part of Papua New Guinea).
coMMENT: See Tyler, L968, Zool. Verh., Leiden, 96:lL6.In the Litoria jeudii grouP
Litoria kinghorni (Loveridge, 1950). Proc. BioI. Soc. Washington,63:"132.

oRIGINAL NAME: Hyla kinghorni.
rYPE(s): Holotype: AM R13818.
TypE LocArtrv: Ulong, New South Wales, Australia.
DrsrRIBUrIoN: Known only from the type locality.
coMMENT: In the Litoria lesueurii group. With no discussion, considered a synonym
of Litoria lesueurii by Cogger, Cameron, and Cogger, L983, Zool. Cat. Aust.,
1(Amph. Rept.):46.
Litoria latopalmata Gtinther, L857. Ann. Mag. Nat. Hist., (3)20:55.

rypE(s): Syntypes: BM L947 .2.24.27 -28.
TypE LocAtlrv: Port Denison [:Bowen], Queensland, Australia.
DrsrRrBUTrou: Eastern Australia to theinterior of Queensland and northeastern
South Australia.
coMMENT: See Davies, Martin, and Watson, 1,983, Trans. R. Soc. S. Aust., L07:87-
108, for definition of species. In the Litoria latopalmata 9touP.
Litorialesueurii (Dum6ril and Bibron, 1841). Erp. G6n.,8:595.

oRIGINAL NAME: Hyla Lesueurii.
rvnn(s): Syntypes: MNHNP 4845-46.
TypE LocALITy: "port ]ackson" (:Sydney), New South Wales, Australia.
DrsrRrBUTroN: Eastern Australia from northern Queensland to southern Victoria.
coMMENT: In the Litoria lesueurii group. See comments under Litoria kinghorni and
Litoria ainosa.
Litoria leucoaa (Tyler, 1968). Zool. Verh., Leiden, 96:IL9.

oRTGTNAL NAME: Hyla leucoaa.
TypE LocALITy: "Busilmin, Eastern Mountains, [West Sepik Province, Papua] New
Guinea".
DrsrRIBUrIoN: Known only from the type locality.
coMMENr: In the monotypic Litoria leucoaa grouP.
Litoria longicrus (Boulenger, l9l1). Ann. Mag. Nat. Hist., (8)8:55.

oRIGINAL NAME: Hylella longicrus.
TypE LocALITy: "one from Fakfak and the other from Wendessi", Vogelkop
Peninsula, Irian ]aya (New Guinea), Indonesia.
DISTRIBUTIou: Easternmountains and Vogelkop Peninsula, New Guinea.
coMMENr: See Tyler, 1968, Zool. Verh., Leiden, 96:L22.In the Litoria bicolor grouP
Litoria longirostris Tyler and Davies, L977. Copeia, 1977:620.

rypE LocAlIrv: Rocky River, Mcllwraith Ranges, Cape York Peninsula,
DrsrRrBUTroN: Mcllwraith Range, Cape York Peninsula, Queensland, Australia
coMMENr: In the Litoria dorsalis grouP.
Litoria lorica Davies and McDonald, 1979. Trans. R. Soc. S. Aust., I03:L70.
rvrr(s): Holotype: QM J36090.
TypE LocArnv: Alexandra Creek near Thornton Peak (16' 07' S, 145o 20' E),
coMMENr: In the Litoria nannotis grouP.
Litoria louisiadensls (Tyler, L968). Zool. Verh., Leiden, 96:L24'

oRIGINAL NAME: Hyla louisiadensis.
TypE LocAt-rry:"Mount Rossell, Rossell Island, Louisiade Archipelago, [Milne Bay
Province, Papua] New Guinea".
DrsrRrBUTroN: Rossell and Sudest Is., Louisiade Archipelago, Papua New Cuinea.
coMMENT: In the monotypic Litoria louisiadensis group.
Litoria lutea (Boulenger, L887). Proc. Zool. Soc. London, L887:337.

oRTGINAL NAME: Hyla lutea.
rvnr(s): Syntypes: BM L947 .2.23.50-52.
TypE LocAtITv: Faro Island, Solomon Islands.
DISTRIBUTIoN: Northwestern islands of the Solomon Is.
coMMENT: See Tyler, L968, ZooL. Verh., Leiden, 96:L27.In the Litoria thesaurensis
grouP.
Litoria maculata (Spencer, 1901). Proc. R. Soc. New South Wales, (2)L3:L77
oRIGINAL NAME: Hyla maculata.
rYPE(s): Holotype: NMM D8498.
TypE LocAr-lrv: Poowong, Victoria, Australia.
DISTRIBUTIoN: Eastern New South Wales and Victoria, Australia.
coMMENT: In the Litoria maculata grouP.
Litoria meiriana (Tyler, 1969). Rec. S. Aust. Mus., L6:2.

oRIGINAL NAME: Hyla meiriana.
rvnE(s): Holotype: SAMA R9082.
TypE LocAury: 98 miles north of Mainoru, Northern Territory, Australia
DISTRIBUTIoITi: Northern Territory and northern Western Australia.
coMMENr: In the monotypic Litoria meiriana grouP.
Litoria microbelos (Cogger, 1966). Aust. Zool., L3:223.

oRIGINAL NAME: Hyla dorsalis microbelos.
TypE LocAurv: Cairns, Queensland [Australia].
DrsrRrBUTroN: Northern Queensland to coastal northern Western Australia,
Australia.
COMMENT: Considered a synonym of Litoria dorsalis by Cogger, Cameron, and
Cogger, L983, ZooL Cat. Aust., l(Amph. Rept.):42, without discussion. Cogger
and Lindner, L974,In Frith and Calaby (eds.), Aust. CSIRO Div. Wildl. Res.
Tech. Pap.,28:63-L07, also questioned the distinctiveness of this species. See
Tyler and Parker, L972, Trans. R. Soc. S. Aust., 96:16L.
Litoria micromembrana (Tyler, L963). Trans. R. Soc. S. Aust., 86:121.

oRIGINAL NAME: Hyla micromembrana.
TypE LoCALITv: "at an elevation of 7,500 ft. on Mt. Podamp, Wahgi-Sepik Divide,
near Nondugl", Western Highlands Province, Papua New Guinea.
DrsrRrBUTroN: Snow Mountains in eastern Irian Jaya (Indonesia) to Huon
Peninsula (Papua New Guinea), New Guinea.
coMMENT: For discussion see Tyler, t968, Zool. Verh., Leiden, 96:L3L.In the Litoria
becki group.
Litoria modica (Tyler, 1968). Zool. Verh., Leiden, 96:L35-

oRIGINAL NAME: Hyla modica.
TypE LocALITy: "Oruge, Eastern Mountains, [Papua] New Guinea"
DISTRIBUTIoN: Central ranges of Papua New Guinea.
coMMENr: In the Litoria becki group.
Litoria moorei (Copland, L957). Proc. Linn. Soc. New South Wales, 82:83.
oRTGTNAL NAME: Hyla moorei.
rvnr(s): Holotype: WAM R5981.
TypE LocAtttv: Pemberton, Western Australia.
DISTRIBUTIoN: Southwest Western Australia.
coMMENT: In the Litoria aurea group. See Maxson, Tyler, and Maxson, L982, Aust. J
Zool.., 30:543, and Tyler, Smith, and ]ohnstone, 1984, Frogs W. Aust.
Litoria multiplica (Tyler, L964). Am. Mus. Novit., 2L87:2.

oRIGINAL NAME: Hyla multiplica.
TypE LocALITy: "Kassam, elevation 4500 feet, Kratke Mountains, Eastern Highlands
[Province], Australian Trusteeship Territory of New Guinea [now Papua New
Guinea]".
DISTRIBUTIoN: Eastern mountains, New Guinea.
coMMENT: For discussion see Tyler, L968, ZooL Verh., Leiden, 96:139.In the Litoria
aruensis group.
Litoria mystax (van Kampen, 1906). Nova Guinea, 5:L73."

oRTGTNAL NAME: Hyla mystax.
rvnr(s): Holotype: RMNH 4632.
TypE LocALITy: "Moaif", northern lowlands, Irian Jaya (New Guinea), Indonesia.
coMMENT: For discussion see Tyler, L968, Zool. Verh., Leiden, 96:L42.In the Litoria
bicolor group.
Litoria nannotis (Andersson, L9L6). K. Svenska Vetensk. Akad. Handl., ($52:L6.

oRIGINAL NAME: Hyla nannotis.
TypE LocAlrrv: Tully River, Queensland, Australia.
DISTRIBUTIoN: Northeastern Queensland, Australia.
coMMENT: For discussion see Liem, L974, Mem. Queensland Mus., 17:LSl, and
Davies and MacDonald, L979, Trans. R. Soc. S. Aust., L03:l69.In the Litoria
nannotis group.
Litoria napaea (Tyler, L968). Zool. Verh., Leiden, 96:145.

oRIGINAL NAME: Hyla napaea.
TypE LocAt-rry:"at an altitude of 2830 ft at Camp 2, 3-4 miles south-west of
Bernhard Camp, Idenburg River, Snow Mountains, New Guinea", Irian Jaya,
Indonesia.
coMMENT: In the monotypic Litoria napaea group.
Litoria nasuta (Gray, L842). ZooI. Misc., 3:56.

oRTGTNAL NAME: Pelodytes nasutus.
rypE(s): Holotype: BM L947.2.22.8L.
TypE LocArtrv: Port Essington, Northern Territory, Australia.
DISTRIBUTIoN: Coastal Australia from northern Western Australia to southern Neu'
South Wales; southern lowlands and southeast peninsula of New Guinea.
coMMENr: See Moore, L961, Bull. Am. Mus. Nat. Hist. 121:305, and Tyler, 1968,
Zool. Verh., Leiden, 96:L48.In the Litoria t'reycineti group.
Litoria nigrofrenata (Gtinther, L867). Ann. Mag. Nat. Hist., (3)20:56.

oRIGINAL NAME: Hyla nigrofrenata.
rvnr(s): Syntypes: BM L947 .2.23.46-47 .
TypE LocArnv: Cape York Peninsula, Queensland, Australia.

DISTRIBUTIoN: East coast of Cape York Peninsula, northern Australia, and southern
New Guinea.
CoMMENT: For discusssion see Tyler, 1968, Zool. Verh., Leiden, 96:724. In the
Lit oria nigrofr enat a groluP.
Litoria nigropunctata (Meyer, L875). Monatsber. Preuss. Akad. Wiss. Berlin, L874:L39.
oRIGINAL NAME: Hyperolius nigropunctatus.
ryPE(s): Formerly in MTKD; destroyed in Worid War II.
TypE LocAtrv: 'iJobi" (:]apen) I., Geelvink Bay, Irian ]aya (New Guinea),
Indonesia (see comment).
DISTRIBUTIoN: Northern New Guinea and Gebe and Jobi Is.
69MMENT: In the Litoria nigropunctata grotp. See Menzies, L972, Herpetologica, 28:
291-300. Obst, L977, Zool. Abh. Staatl. Mus. Tierkd., Dresden, S4:174,
mentioned "Ansus, lobi" , as the locality for the destroyed holotype.
Litoria nyakalensis Liem, L974. Mem. Queensland Mus., L7:L57.

TypE LoCAury: Henrietta Creek, Palmerston Nat. Pk., 800 m., Queensland,
Australia.
DISTRIBUTIoN: Northern Queensland, Australia.
coMMENT: In the Litoria nannotis group. See Davies and MacDonald, L979, Trans. R.
Soc. S. Aust., 103:L69, for discussion.
Litoria obtusirostris Meyer, L875. Monatsber. Preuss. Akad. Wiss. Berlin, 1874:L39.
ryPE(s): Formerly in MTKD; destroyed in World War II.
TypE LocALIty: "]obi" (:Japen) I., Geelvink Bay, Irian |aya (New Guinea),
coMMENr: For discussion see Tyler, L968, Zool. Verh., Leiden, 96:L55. Obst, L977,
Zool. Abh. Staatl. Mus. Tierkd., Dresden, 34:174, mentioned "Ansus, lob7",
as the locality for the destroyed holotype.
Litoria oenicolen Menzies and Zweif.el, 1974. Am. Mus. Novit., 2558:10.
TypE LocArrryr "beside the Trauna River (a tributary of the Baiyer River) at the
Baiyer River Wildlife Sanctuary (L4l' LL' E,5o 29' S), Western Highlands
District [now Province], Papua New Guinea, elevation 1200 m."
DISTRIBUTIoT: Known only from the type locality.
coMMENT: In the Litoria arfakiana grouP.
Litoria olongburensls'Liem and Ingram, 1977. Victorian Nat., 94:259.

TypE LOCAuty: Coomboo Lake, Fraser Island, Queensland, Australia.
DrsrRrBUTroN: Southeastern Queensland and northeastern New South Wales,
Australia.
coMMENT: In the Litoria dorsalis grouP.
Litoria pallida Davies, Martin, and Watson, 1983. Trans. R. Soc. S. Aust., L07:L0L.
rypE LocAlrry: Gulungul Creek Crossing, Arnhem Highway, Northern Territory,
Australia.
DrsrRrBUTroN: Northern Australia from Broome in Western Australia to Cape York
in Queensland.
Litoria paraewingi Watson, Loftus-Hills, and Littlejohn, L971. Aust. l. Zool., l9:4L4.
rYPE(s): Holotype: NMM D27601.
TypE LocAuryr b'Connors Flat, 8.3 km south-southeast of Yea, Victoria, Australia.
DISTRIBUTIoN: Eastern Victoria, Australia.
coMMENr: In the Litoria ewingii grouP.
Litoria pearsoniana (Copland, l96L). Proc. Linn. Soc. New South Wales, 86:168.
oRIGINAL NAME: Hyla pearsoniana.
rvnr(s): Holotype: AM R18588.
TypE LocArnv: Cedar Creek, E of Mt. Glorious, about 20 mi. NW of Brisbane,
DISTRIBUTIoN: Elevated areas of northeastern New South Wales, Australia.
coMMENr: In the Litoria citropa group. Hyla pearsoniana is a replacement name for
Hyla pearsoni Copland, L960, Proc. Linn. Soc. New South Wales, 85:L54, which
is preoccupied by Hyla pearsoni Gaige, L929. Cogger, Cameron, and Cogger,
1983, Zool. Cat. Aust., l(Amph. Rept.):48, considered Litoria pearsoniana to be a
junior synonym of Litoria phyllochroa but provided no evidence to contradict
evidence of dispecificity provided by Barker and Grigg, L977, Field Guide
Aust. Frogs:54-55 (call differences), or King, L981, Proc. Melbourne Herpetol.
Symp., R. Melbourne Zool. Gardens, Melbourne:L72 (chromosomal
differences).
Litoria peronii (Tschudi, 1838). Classif. Batr.:75.

oRTGTNAL NAME: Dendrohy as peronii.
rvnn(s): Holotype: MNHNP 4809.
TypE LocALITy: "Nov. Holl." (:Australia).
DISTRIBUTIoN: Eastern Australia, south from Mary River (Queensland).
coMMENT: For discussion see Martin, Gartside, Littlejohn, and Loftus-Hills, L979,
Proc. Linn. Soc. New South Wa1es, 103:23. In the Litoria peronii group.
Litoria personata Tyler, Davies, and Watson,'1.978. Trans. R. Soc. S. Aust., 102:151.
rvnE(s): Holotype: SAMA RL6773.
TypE LocArmv: Birndu (12' 32' S, 132" 08' E), southeast of Cannon Hill Station, East
Alligator River Region, Northern Territory, Australia.
DISTRIBUTIoN:Known only from the type locality.
coMMENT: In the Litoria nigrofrenata group.
Litoria phyllochroa (Gtinther, 1863). Proc. ZooI. Soc. London, 1863:251.

oRIGINAL NAME: Hyla phyllochroa.
rvnr(s): Syntypes: BM L947.2.24.3-5, ZMB 4929.
TypE LocAurv: Sydney and Errumanga, New South Wales, Australia.
DISTRIBUTIoN: Eastern part of New South Wales and adjacent Queensland and
Victoria, Australia.
coMMENr: In the Litoria citropa group; see Moore, 196L, Bull. Am. Mus. Nat. Hist.,
L2L:255. See comment under Litoria pearsoniana.
Litoria pratti (Boulenger, L9Ll). Ann. Mag. Nat. Hist., (8)8:56.

oRIGINAL NAME: Hyla pratti.
rypE(s): Syntypes: BM 19 47 .2.23.54-56 (Wendess 1), L9 47 .2.23.57 -58 (Arfak
Mountains).
rypE LocALIry: "Wendessi . . . [and] Arfak Mountains at an altitude of 8000 feet",
Vogelkop Peninsula, Irian ]aya (New Guinea), Indonesia.
DrsrRrBurron: Known only from the type localities.
coMMENT: For discussion see Tyler, L968, Zool. Verh., Leiden, 95:L57.In the Litoria
becki group.
Litoria prora (Menzies, L969). Trans. R. Soc. S. Aust., 93:165.

oRIGINAL NAME: Hyla prora.
rYPE(s): Holotype: UP 1015.
TypE LocALtrv: ".. . near Efogi . . . at an altitude 3,800'; location L47" 38' E;9 9' S
and approximately 37 miles northeast of Port Moresby", Central Province,
Papua New Guinea.
coMMENT: In the monotypic Litoria prora group.
ANURA: HYLIDAE: PELODRYADINAE t9t
Litoria pygmaea (Meyer, L875). Monatsber. Preuss. Akad. Wiss. Berlin, L874:139.
oRIGINAL NAME: Hyperolius pygmaeus.
rvrr(s): Formerly in MTDK; destroyed in World War II.
TypE LocALITy: "]obi" (:]apen) I., Geelvink Bay, Irian Jaya (New Guinea),
Lowlands of New Guinea.
DISTRIBUTIoN:
coMMENT: For discussion see Tyler, 1968, Zool. Verh., 96:159.In the Litoria rubella
group. Obst, L977, Zool. Abh. Staatl. Mus. Tierkd., Dresden,34:L74,
mentioned "Ansus, lobi" as the locality for the destroyed holotype.
Litoria quadrilineata Tyler and Parker, L974. Trans. R. Soc. S. Aust., 98:71.
rvrr(s): Holotype: SAMA R13489.
TypE LocALITy: "]alan Trikora [:Tridora Road], Merauke, Irian Jaya, New Guinea",
Indonesia.
DrsrRIBUrIow: Known only from the type locality.
coMMENT: In the monotypic Litoria quadrilineata gtoup.
Litoria raniformis (Keferstein, L867). Nachr. Ges. Wiss. Gottingen, 18:358.

oRIGINAL NAME: Chirodryas raniformis.
rvnE(s): Holotype: formerly Zool. Mus. Grittingen (now in ZFMK) 93h; presumed
lost.
TypE LocALITy: "New South Wales?" , Australia.
DrsrRrBUTroN: Southeastern Australia (exclusive of coastal New South Wales);
Tasmania.
coMMENr: In the Litoria aurea group. See Courtice and Grigg, 1975, Aust. Zool., 18:
149-L63, and Maxson, Tyler, and Maxson, L982, Aust. J. Zool., 30:643.
Litoria reaelata Ingram, Corben, and Hosmer, L982. Mem. Queensland Mus., 20:635.
rvru(s): Holotype: QM 128233.
TypE LocAtItv: O'Reilly's (28 14' S, 153o 08' E), Lamington Plateau, SE
DrsrRrBUTroN: Southeastern Queensland and northeastern New South Wales,
Australia.
coMMENr: In the Litoria ewingii grouP.
Litoria rheocola Liem, 1974. Mem. Queensland Mus., 17:L52.

rvnr(s): Holotype: QM J2263L.
rypE LoCALITy: Kuranda, 50 km west [actually ca. 33 km by road NW-RGZ] of
Cairns, Queensland, Australia.
coMMENT: In the Litoria nannotis group. See Davies and McDonald, L979, Trans. R.
Soc. S. Aust., L03:L59, for discussion.
Litoria rothii (De Vis, 1884). Proc. Linn. Soc. New South Wales,9:66.
oRTGINAL NAME: Hyla rothii.
rYPE(s): Syntypes: formerly in QM; now lost.
TypE LocAurv: Mackay, Queensland, Australia.
DrsrRrBUTroN: Northern Australia from northeastern Western Australia to
Queensland north of Mary Rive4 southern New Guinea.
coMMENr: In the Litoria peronii grouP.
Litoria rubella (Gray, L842). Zool. Misc., 2:56.

oRIGINAL NAME: Hyla rubella.
rvrr(s): Syntypes: BM 1947 .2.24.7 -9.
TypE LocArrrv: Port Essington, Northern Territory, Australia.
DrsrRrBUTroN: Northern two thirds of Australia and southern New Guinea.
coMMENr: For discussion see Copland, L957 , Proc. Linn. Soc. New South Wales,
82:46.In the Litoria rubella grouP.
1,92 AMPHIBIAN SPECIES OF THE WORLD
Litoria sanguinolenta (van Kampen, 1909). Nova Guinea, 9:33.

oRIGINAL NAME: Hyla sanguinolenta.
rYPE(s):ZMA 5676 designated lectotype by Daan and Hillenius, L966, Beaufortia,
158:121.
TYPE LocALITY: "Sabang, Lorentz River, West New Guinea" (Irian laya,Indonesia).
DISTRIBUTIoN: Southern lowlands in western New Guinea.
coMMENr: For discussion see Tyler, L968, Zool. Verh., Leiden, 96:L64.In the Litoria
infrafrenata group.
Litoria spinifera (Tyler, L968). ZooL. Verh., Leiden, 96:L67.

oRIGINAL NAME: Hyla spinifera.
TypE LocALrry: "at an altitude of approximately 5000 ft at Oruge, Eastern
Mountains, [Papua] New Guinea".
DISTRIBUTIoN: Eastern mountains, New Guinea.
coMMENr: In the Litoria becki grotp.
Litoria splendida Tyler, Davies, and Martin, L977. Trans. R. Soc. S. Aust., 101:133
rvnn(s): Holotype: WAM R56840.
TypE LocArtry: Lake Argyle Tourist Village, Kimberley Division, Western
Australia.
DISTRIBUTToN: Northern Western Australia.
coMMENr: In the Litoria caerulea group.
Litoria subglandulosaTyLer and Anstis, 1983. Trans. R. Soc. S. Aust., L07:L30.

TYrE LocAtrrv: Barwick Creek, Point Lookout, near Ebor, New South Wales,
Australia.
DISTRIBUTIoN: Mountains of northeastern New South Wales and southeastern
coMMENr: In the Litoria citropa group. Litoria subglandulosa is a replacement name
for Litoria glandulosa Tyler and Anstis, L975, Rec. S. Aust. Mus., L7:46, which is
preoccupied by Litoria glandulosa Bell, 1843.
Litoria thesaurensis (Peters, 1877). Monatsber. Preuss. Akad. Wiss. Berlin, L877:421.
oRIGINAL NAME: Hyla thesaurensis.
rypE(s): Syntypes: ZMB 9L2L (2 specimens).
TypE LocArrry: Treasury Island, Solomon Islands.
DISTRIBUTIoN: New Guinea, New Britain, New Ireland, and Solomon Is.
coMMENT: In the Litoria thesaurensis group. See Menzies and Zug,1979,
Micronesi ca, 15:325-333.
Litoria timida Tyler and Parker, 1972. Trans. R. Soc. S. Aust., 96:L57.
rYPE(s): Holotype: SAMA Rl1658.
TypE LocALITy: "Menemsorae, Western District [Province], Papua New Guinea"
DISTRIBUTIow: Eastern and central New Guinea.
coMMENT: In the Litoria dorsalis group.
Litoria tornieri (Nieden, L923). Das Tierreich, 46:228.

oRIGINAL NAME: Hyla tornieri.
rYPE(s): Holotype: BM 1947 .2.22.79.
TYPE LocAtltv: North coast of Australia; corrected to Port Essington, Northern
Territory, Australia, by Boulenger, L882, Cat. Batr. Sal. Brit. Mus.:413.
DISTRIBUTIOTT: Northern and eastern Australia.
coMMENr: In the Litoria latopalmata group. Hyla tornieri Nieden, L923, is a
substitute name for Pelodytes affinis Gray, L842, ZooL. Misc., 3:56. Davies,
Martin, and Watson, L983, Trans. R. Soc. S. Aust., L07:87, reviewed the
species.
{
Litoria tyleri Martin, Watson, Gartside, Littlejohn, and Loftus-Hills, 1979. Proc. Linn
Soc. New South Wales, 103:25.
TypE LocAltrv: Ryan's Swamp, Caves Beach Reserve, 14 km S Huskisson, New
South Wales, Australia.
DrsrRrBUTroN: Coastal southeastern Queensland and northeastern New South
Wales, Australia.
coMMENr: In the Litoria peronii group.
Litoria umbonata Tyler and Davies, 1983. Copeia, 1983:803.

rypE LocALITy: "Baliem River Valley, 1600 m . . . Irian laya, .. . New Guinea",
Indonesia.
DrsrRrBUrIoN: Vicinity of type locality.
coMMENr: In the Litoria wisselensis group.
Litoria aagabunda (Peters and Doria, L878). Ann. Mus. Civ. Stor. Nat. Genova, L3:424
oRTGTNAL NAME: Hyla (Litoria) oagabunda.
rvnr(s): MSNG 29704A designated lectotype by Capocaccia, L957, Ann. Mus. Civ.
Stor. Nat., Genova, 69:2L4.
TypE LocALITy: "Wahai, Seram Island (Moluccas)".
DISTRIBUTIoN: Vogelkop Peninsula, Irian ]aya (New Guinea) and Ceram I.,
Moluccan Is., Indonesia.
coMMENT: For discussion see Tyler, 1968, Zool. Verh., Leiden, 96:L77.In the
monotypi c Litoria aagabunda group.
Litoria aerreauxii (Dum6ri1, 1853). Ann. Sci. Nat., Paris, (3)L9:1,71.

oRIGINAL NAME: Hyla aerreauxii.
rvnr(s): Syntypes: MNHNP 4849 (5 specimens), 4850 (9 specimens).
TYPE LocArnv: Australia.
DrsrRrBUTIow: Eastern Australia.
coMMENr: For discussion see Littlejohn, L965, Evolution, L9:234.In the Litoria
ewingii group.
Litoria ainosa (Lamb, 1911). Ann. Queensland Mus., L0:27.

oRTGTNAL NAME: Hyla uinosa.
TypE LocAurv: Ithaca Creek, Brisbane, Queensland, Australia.
coMMENT: For discussion see Moore, 7961,8u11. Am. Mus. Nat. Hist.,96:2L2.
Without discussion, considered a synonym of Litoria lesueurii by Cogger,
Cameron, and Cogger, 1983, Zool. Cat. Aust., l(Amph. Rept.):46.
Litoria oociaincens Menzies, L972. Herpetologica, 28:296.

rvnn(s): Holotype: UP L957.
TypE LocALITy: "Brown River forestry station, . . . L47" 14' E;10o 48' S, sea level,
about 25 miles NW Port Moresby", Central Province, Papua New Guinea.
DISTRIBUTIoN: Coastal lowlands of southeastern New Guinea.
coMMENT: In the Litoria rubella grouP.
Litoria watjulumensis (Copland, L957). Proc. Linn. Soc. New South Wales, 8?:96.
oRIGINAL NAME: Hyla latopalmata watjulumensis.
rypE(s): Syntypes: WAM Rl1195, LLL98-99, 11633-35, L1896-107, 17932-38.
TypE LocAurv: Watjulum [:Wotjulum] Mission, Western Australia.
DrsrRrBUTroN: Northern Western Australia and Northern Territory, Australia.
coMMENr: In the Litoria nigrofrenata group. The unjustified emendation of the
specific epithet to wotjulumensis was by Tyler, 1968, Rec. S. Aust. Mus., L5:7L3
MIT disputes DRF's interpretation of the International Code of Zoological
Nomenclature.
Litoria wisselensis (Tyler, 1968). Zool. Verh., Leiden, 96:180.

oRIGINAL NAME: Hyla wisselensis.
rvnr(s): Holotype: RMNH 12295.
TypE LocAt.rry: "Enarotali, Lake Paniai, Wissel Lakes, Snow Mountains, [Irian laya,)
New Guinea", Indonesia.
DISTRIBUTIoN: Montane lakes in Snow Mountains, New Guinea.
coMMENT: In the Litoria zaisselensis group; see Tyler and Davies, L983, Copeia, 1983:
803-808.
Litoria wollastoni (Boulenger, L9L4). Trans. Zoo1.. Soc. London, 20:248.
oRIGINAL NAME: Hyla wollastoni.
rvnr(s): Holotype: BM 1947 .2.23.59.
TypE LocALITy: "Camp VI, Utakwa [:Octakwa] R., 2L00 f.t.",Irian ]aya (New
Guinea), Indonesia.
DISTRIBUTIoN: Highlands of New Guinea.
coMMENT: In the Litoria arfakiana group. See Menzies and Zweif.el, 1974, Am. Mus
Novit., 2558:1-16.
Nyctimystes Stejneger, L916. Proc. Biol. Soc. Washington,29:85.
rypE spECIns: Nyctimantis papua Boulenger, L897 , by original designation.
DISTRTBUTToN: New Guinea, eastern islands in Moluccas, and northern Queensland,
Australia.
coMMENT: Genus defined and species listed by Tyler and Davies, L979, Aust. l. Zool.,
27:755-772. See also Zweifel,7983, Am. Mus. Novit., 2759:L-2L.
Nyctimystes aoocalis Zweif.el,1958. Am. Mus. Novit., 1896:6.

TypE LocAl-rry: "east side of Goodenough Island, D'Entrecasteaux Group, [Milne
Bay Province,] Territory of Papua [:Papua New Guinea], New Guinea".
DISTRIBUTIoN: Known only from the type loca1ity.
Nyctimystes cheesmani Tyler, L964. Zool. Abh. Staatl. Mus. Tierkd., Dresden, 27:268.
rvnr(s): Holotype: BM L947 .2.24.3L.
TypE LocAt-rry: "Mondo, Papua (5000 ft.)", Central Province, Papua New Guinea.
DISTRIBUTIoN: Eastern New Guinea and D'Entecasteaux Is.
coMMENT: For discussion see Zweif.el,1958, Am. Mus. Novit., 7896:2L. Nyctimystes
cheesmani is a replacement name for Hyla montana Parker, L936, Ann. Mag.
Nat. Hist., (10)17:80.
Nyctimystes dayi (Gtinther, L897). Novit. Zool., 4:406-
oRIGINAL NAME: Hyla dayi.
rvnr(s): Unknown.
TypE LocAlIrv: Bartle Frere Mountains, Queensland, Australia.
coMMENT: For discussion see Liem, 1974, Mem. Queensland Mus., L7:L56.
Regarded as a species inquirenda by Cogger, Cameron, and Cogger, L983, ZooL
Cat. Aust., 1(Amph. Rept.):260. See comments under Nyctimystes hosmeri,
Nyctimystes tympanocryptis, and Nycfimystes aestigia.
Nyctimystes daymani Zweifel,1958. Am. Mus. Novit., 1896:8.
TypE LocAt-rry: "North slope of Mt. Dayman, Territory of Papua, [Milne Bay
Province, Papua] New Guinea".
DISTRIBUTIow: Mountains of eastern New Guinea.
Nyctimystes disrupta Tyler, L963. Rec. Aust. Mus., 26:LL8.
rvrE(s): Holotype: AM R15923.
TypE LocALITy: "at an elevation of 6,000 feet in the Kaironk Valley, Schrader
Mountains, Australian Trusteeships Territory of New Guinea" (Madang
Province, Papua New Guinea).
DrsrRrBUTroN: Highlands of Huon Peninsula and central Papua New Guinea.
coMMENT: See Zweif.el, L983, Am. Mus. Novit., 2759:8-L2.
Nyctimystes fluoiatilis Zweif.el, 1958. Am. Mus. Novit., 1896:11'

TypE LocA1-rry,
i'B"rnhard Camp, Idenburg River, Netherlands New Guinea" (Irian
Jaya, Indonesia).
Nyctimystes foricula Tyler, L963. Rec. Aust. Mus., 26:120'

rvrr(s): Holotype: AM Rl5904.
TypE LocAgry
i'b"tween 5,000 feet and 6,000 feet in the Kaironk Valley, Schrader
Mountains, Australian Trusteeships Territory of New Guinea" (Madang
Province, Papua New Guinea).
DISTRIBUTIoN: Mountains of eastern New Guinea.
Nyctimystes granti (Boulenger, L9L4). Trans. Zool. soc. London,20:249.

oRIGINAL NAME: Nyctimantis granti.
rYPE(s): Holotype: BM L947 .2.24.47 .
TypE LocAlrry, "Camp VIa, Utakwa [:Octakwa] R., 3000 ft.", Irian ]aya (New
Guinea), Indonesia.
DISTRIBUTIoN: Mountains of New Guinea.
CoMMENT: For discussion see Zweifel, 1958, Am. Mus. Novit., 1896:13.
Nyctimystes gularis Parker, L936. Ann. Ma8. Nat. Hist., (L0)17:78'

ryPE(s): Holotype: BM L947 .2.24.29.
TypE L9CALITY, "Mondo, Papua (5000 ft.)", Central Province, Papua New Guinea.
DISTRIBUTIoN: Mountains of southeastern New Guinea.
coMMENT: For discussion see Zweif.el, L958, Am. Mus. Novit., L896:L4, and Tyler,
L9 62, Copeia, L9 62:435 - 435.
Nyctimystes hosmeri Tyler, L964. Trans. R. Soc. S. Aust., 88:111.

TypE LocArrrv: 1ully Falls, Cape York Peninsula, Queensland, Australia.
DrsrRrBUTroN: Atheiton Tableland and adjacent mountains, Queensland, Australia.
g9MMENT: Ingram and Covacevich, 198L, Mem. Queensland Mus., 20:298, implied
that this form is synonymous with Nycf imystes dayi.
Nyctimystes humeralis (Boulenger, L912). Zool. ]ahrb., l(suppl. 15):216.

oRIGINAL NAME: Hyla humeralis.
rypE(s): Holotype:,,8M L947 .2.23.48.
rYPE LocAtIrv, i'Mudew [:Madiu], St' ]oseph [Angabunga] River' British New
Guinea, between 2000 and 3000 feet", Central Province, Papua New Guinea.
COMMENT: For discussion see Zwetfel,1958, Am. Mus. Novit., L896:16.
Nyctimystes kubori Zweifel,1958. Am. Mus. Novit., 1895:18'

5000 and 8000 feet,
TypE LOCAury, ''K,rbor Mountains near Kt P . . . between
north-eastern New Guinea" (Chimbu Province, Papua New Guinea).
coMMENT: For discussion see Tyler, 1963, Rec. Aust' Mus', 26:L23'
Nyctimystes montana (Peters and Doria, L878\. Ann. Mus. Civ. Stor. Nat. Genova, L3
423.
oRIGINAL NAME: Hyla (Litoria) montana.
rYPE(s): Holotype: MSNG 29720.
TypE LoCALIrr,
i'Hutam", Arfak Mountains, Vogelkop Peninsula, Irian Jaya (New
Guinea), Indonesia.
coMMENT: For discussion see Tyler, L964, Zool. Abh. Staatl. Mus. Tierkd., Dresden,
27:265.
Nyctimystes narinosa Zweifel,1958. Am. Mus. Novit., 1896:26.

TypE LocAt-rry: "Mt. Hagen in the Wahgi Valley region of north-east New Guinea
at an elevation of over 8000 feet", Western Highlands Province, Papua New
Guinea.
DISTRIBUTIoN: Mountains of central New Guinea.
Nyctimystes obsoleta (Lonnberg, 1900). Ann. Mag. Nat. Hist., (7)6:580.
oRIGINAL NAME: Hyla obsoleta.
rvnr(s): ZIUU (no number).
TypE LocALITy: "Simbang", Huon Peninsula, Morobe Province, Papua New
Guinea.
coMMENT: For discussion see Tyler, L965, Proc. Zool. Soc. London, L45:92.
Nyctimystes papua (Boulenger, L897). Ann. Mag. Nat. Hist., (6)L9:L2.

oRTGTNAL NAME: Nyctimantis papua.
rvnn(s): BM 96.10.31.50 designated lectotype by Zweif.el, L983, Am. Mus. Novit.,
2759:4.
TypE LocALITy: "Mount Victoria, Owen Stanley Range, [Papua] New Guinea".
DrsrRIBUTroN: Type locality on the western slope of Mt. Victoria, Papua New
Guinea.
coMMENT: For discussion see Zweif el, 1983, Am. Mus. Novit., 2759:4-8.
Nyctimystes perimetri Zweif.el,1958. Am. Mus. Novit., L896:3L.
rvrr(s): Holotype: AMNH 60081.
TypE LocALrry:"at an elevation of 250-350 meters... on the west slope of Mt.
Riu, Sudest (Tagula) Island, Louisiade Archipelago, Territory of Papua New
Guinea" (Milne Bay Province, Papua New Guinea).
DISTRIBUTIoN: Sudest and Rossel Is., Louisiade Archipelago, New Guinea.
Nyctimystes persimilis Zweifel, 1958. Am. Mus. Novit., L896:34.

rvrr(s): Holotype: AMNH 56838.
TypE LocAt-rry: "north slope of Mt. Dayman, Territory of Papua, New Guinea, at
an elevation of 1570 meters" (Milne Bay Province, Papua New Guinea).
Nyctimystespulchra (Wandolleck, 1911). Abh. Ber. K. Zool. Anthro. Ethno. Mus.

Dresden, 13(6):L2.
oRIGINAL NAME: Hyla pulchra.
rvnu(s): Syntypes: MTKD D22L9 (4 specimens), destroyed.
TypE LocALITy: "Sacksackhiitte", Torricelli Mountains, West Sepik Province, Papua
New Guinea.
DrsrRrBUTroN: Mountains of New Guinea.
coMMENT: For discussion see Tyler, L964, Zool. Abh. Staatl. Mus. Tierkd., Dresden
27:267, and Zweifel, 1980, BuIl. Am. Mus. Nat. Hist., 165:400.
Nyctimystes rueppelli (Boettger, 1895). Zool. Anz., l8:L37.

oRIGINAL NAME: Hyla rueppelli.
rvrr(s): SMF 2614 designated lectotype by Mertens, 1967, Senckenb. Biol., 48:42.
TypE LocALITy: "Kau, N-Halmaheira, Moluccas", Indonesia.
DISTRIBUTIoN: Halmahera and Morotai Is., Moluccan Is., Indonesia.
coMMENT: For discussion see Zweifel, 1958, Am. Mus. Novit., 7896:34.
Nyctimystes semipalmat a P arker, 1936. Ann. Mag. Nat. Hist., ( 10)17:83.

TypE LocALITv: "Kokoda (1200 ft.)", Northern Province, Papua New Guinea.
DrsrRrBUTroN: Papua New Guinea.
ANURA: HYLIDAE: PHYLLOMEDUSINAE L97
CoMMENT: See Zweifel, 1958, Am. Mus. Novit., 1896:38, and Zweifel, 1980, Bull.
Am. Mus. Nat. Hist., L65:400.
Nyctimystes trachydermis Zweifel, L983. Am. Mus. Novit., 2759:L2.

TypE LocAt-rry:"at an elevation of about 1250 m. at Gapaia Creek, trail between
Garaina and Saureli, Morobe Province, Papua New Guinea".
DTsTRIBUTIoN: Mountains of eastern New Guinea.
Nyctimystes tyleri Zweif.el,, 1983. Am. Mus. Novit., 2759:L6.

rypE(s): Holotype: AMNH 82878 (erroneously given as 82867 in the original
publication-RGZ).
TypE LocAt,rry: "elevation of about 1280 m. at Gapaia Creek, between Garaina and
Saureli, Morobe Province, Papua New Guinea".
DISTRIBUTIoN: Type locality in the highlands of eastern New Guinea.
Nyctimystes tympanocryptis (Andersson,1916). K. Svenska Vetensk. Akad. Handl.,

(4)52:19.
oRTGINAL NAME: Hyla tympanocryptis.
TypE LocAurv: Malanda, Queensland, Australia.
DTsTRIBUTIoN: Cape York Peninsula, Queensland, Australia.
coMMENr: See Tyler, 1968, Ark. Zool., (2)20:501. Ingram and Covacevich, L98L,
Mem. Queensland Mus., 20:298, implied that this form is synonymous with
Nyctimystes dayi.
Nyctimystes oestigia Tyler, L964. Trans. R. Soc. S. Aust., 88:113.

rvnr(s): Holotype: NHMW L7187.
TypE LocArrry: Mount Bartle Frere, Cape York Peninsula, Queensland, Australia.
DISTRIBUTIoN: Northeastern Queensland, Australia.
coMMENT: Ingram and Covacevich, L981, Mem. Queensland Mus., 20:298, implied
that this form is synonymous with Nycf imystes dayi.
Nyctimystes zweifeli Tyler, L967. Trans. R. Soc. S. Aust., 9L:191.

TypE LocAt-rry:"Telefomen [:Telefomin], Western Highlands, New Guinea" (West
Sepik Province, Papua New Guinea).
DISTRIBUTIoN: Known only from the vicinity of the type locality.
suBFAMILy: Phyllomedtsinae Giinther, L859 "L858".

cIrArIoN: Cat. Batr. SaI. Coll. Brit. Mus.:120.
DTsTRIBUTIoN: Mexico to Argentina.
ff
coMMENT: As originally formed, the group name was Phyllomedusidae. Frogs referred
to this subfamily possess distinctive morphological, biochemical, behavioral, and
reproductive features (Duellman,L968, Univ. Kansas Publ. Mus. Nat. Hist., 18:3-
ll!l
10). Bagnara and Ferris, L973,1. Exp. Zool.,190:367-372, suggested that similar
melanosomes in some Phyllomedusa and some Litoria might be indicative of a close
relationship between the Phyllomedusinae and the Pelodryadinae. The
immunological evidence of Maxson, L975, Experientia, S2:L149-1150, does not
refute sucli a relationship, but suggests that the divergence has not been recent.
Osteological and myological evidence of Tyler and Davies, L978, Herpetologica,
34:2L9-224, does not refute such a hypothesized relationship (DCC).
REVIEwER: David C. Cannatella (DCC).
Agalychnis Cope, 1864. Proc. Acad. Nat. Sci. Philadelphia, L6:181..

rypE spECIus: Hyla callidryas Cope, 1862 (see comment).
DrsrRrBUTrow: Southern Mexico, Central America, Pacific lowlands of Colombia and
northwestern Ecuador, and Amazonian Ecuador.
CoMMENT: For discussion see Duellman, L970, Monogr. Mus. Nat. Hist. Univ.
Kansas, 1:87-130. The designation of Agalychnis moreletii as the type species of
Agalychnis by Taylor, L952, Univ. Kansas Sci. Bull., 35:80L, and the comments br'
Duellman,lgTO, Monogr. Mus. Nat. Hist. Univ. Kansas, L:87-88, are in error;
see Cope, 1864, Proc. Acad. Nat. Sci. Philadelphia, 15:181.
Agalychnis annae (Duellman, L963). Rev. Biol. Tropical, LL:1.

oRIGINAL NAME: Phyllomedusa annae.
TypE LocALITY: "TaPanti, Cartago Province, Costa Rica, L200 meters".
DISTRIBUTIoII: Caribbean slopes and Meseta Central of Costa Rica.
Agalychnis calcarifer Boulenger, L902. Ann. Mag. Nat. Hist., (7)9:52.

rypE(s): Holotype: BM 19 47 .2.24.22.
TypE LOCALITv: "Rio Durango, 350 feet", Provincia Esmeraldas, Ecuador.
DrsrRrBUTroN: Caribbean slopes of Costa Rica and Panama and Pacific lowlands of
Colombia and northwestern Ecuador.
CoMMENT: See Myers and Duellman, L982, Am. Mus. Novit., 2752:25-25.
Agalychnis callidryas (Cope, L862). Proc. Acad. Nat. Sci. Philadelphia, 14:359.
oRIGINAL NAME: Hyla callidryas.
rYPE(s): Holotype: ANSP 209L.
TypE LOCALITy: "Darien" (Provincia), Panama; corrected to C6rdoba, Yeracruz,
Mexico, by Smith and Taylor, '1.950, Univ. Kansas Sci. BulI., 33:347; correction
considered unjustified by Duellman, L970, Monogr. Mus. Nat. Hist. Univ.
Kansas, L:102.
DrsrRrBUTroN: Atlantic lowlands of Veracruz and Oaxaca, Mexico, southeastward on
the Caribbean lowlands to central Panama; Pacific lowlands of southern Costa
Rica and eastern Panama.
CoMMENT: Savage and Heyer, 1967, Beitr. Neotrop. Fauna,50:11L-L3L, analyzed
intraspecific variation.
Agalychnis craspedopus (Funkhouser, L957). Occas. Pap. Nat. Hist. Mus. Stanford
Univ., 5:23.
oRIGINAL NAME: Phyllomedusa craspedopus.
rYPE(s): Holotype: CAS-SU 10310.
TypE LocALlry: "Chicherota, Rio Bobonaza, Napo-Pastaza [now Pastaza] Province,
eastern Ecuador (Lat.2" 22' 5., Long.76" 38'W.), at an altitude of about 250
meters".
DrsrRrBUTroN: Known only from the type series and RMNH material from
Montalvo (MSH), in Amazonian Ecuador.
Agalychnis litodryas (Duellman and Trueb, L967). Copeia, L967:L25.

oRIGINAL NAME: Phyllomedusa litodry as.
unr(s): Holotype KU 96L49.
TypE LocALtry: "L km west-southwest of the junction of the Rio Mono and the Ric
Tuira, Dari6n Province, Panam6, elevation 130 m".
DISTRIBUTIoN: Eastern Panama to northwestern Ecuador.
Agalychnis moreletii (Dum6ril, 1853). Ann. Sci. Nat., Paris, (3)19:169.

oRIGINAL NAME: Hyla moreletii.
rvnE(s): Syntypes: MNHNP 428 (parchment label 767) (2 specimens).
TypE LOCAtIryi "Verapaz", Guatemala; restricted to "CobAn, [Departamento] Alta
Yerapaz", Guatemala,by Smith and Taylot, L950, Univ. Kansas Sci. 8u11., 33:
3L7.
DrsrRrBUTroN: Atlantic and Pacific slopes from Veracruz and Guerrero, Mexico, to
Guatemala and El Salvador.
ANURA: HYLIDAE: PHYLLOMEDUSINAE t99
Agalychnis saltator Taylor, 1955. Univ. Kansas Sci. Bull., 37:527.

rvnn(s): Holotype: KU 35615.
TypE LocALtry "4 km NNE of TilarAn l:Finca San Bosco], [Provincia] Guanacaste,
Costa Rica".
DrsrRrBUTIoN: Caribbean lowlands of Nicaragua and Costa Rica.
Agalychnis spurrelli Boulenger, L9L3. Proc. Zool. Soc. London, 1913:1024.

rvnE(s): Syntypes: BM 1,947 .2.24.24-25.
TypE LocAt-rry: "Pefra Lisa, Condoto, altitude 300 feet", Provincia Choc6,
Colombia.
Southeastern Costa Rica to the Pacific lowlands of Colombia
DrsrRrBUTroN:
Pachymedusa Duellman,1968. Univ. Kansas Publ. Mus. Nat. Hist., L8:5.

rypE spECIns: Phyllomedusa dacnicolor Cope, L864, by original designation.
DTsTRIBUTIoN: As for the single species.
coMMENT: See Pyburn, L970, Copeia, L970:209-2L8, f.or arguments against
recognition at generic level. Pachymedusa is the sister-taxon of other
phyllomedusines (DCC).
Pachymedusa dacnicolor (Cope, L864). Proc. Acad. Nat. Sci. Philadelphia, 1,5:181.
oRTGINAL NAME: Phyllomedusa dacnicolor.
rvnr(s): Formerly in USNM; now lost.
TYPE LoCALITY: "near Colima", Mexico.
DISTRIBUTIoN: Pacific lowlands of Mexico from southern Sonora to the Isthmus of
Tehuantepec.
coMMENT: See Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas, 1:81, for
review.
Phyllomedusa Wagler, 1830. Nat. Syst. Amph.:201.

TypE spECIns: Rana bicolor Boddaert, L772.
DISTRIBUTIoN: Costa Rica, Panama, Pacific slopes of Colombia, and South America
including Trinidad, southward to northern Argentina.
east of the Andes,
coMMENr: Although species groups have not been defined for all species,
Cannatella, L982, Copeia, 1982:501-513, has discussed and defined the
Phyllomedusa perinesos group, and Cannatella, 1980, Occas. Pap. Mus. Nat. Hist
Univ. Kansas, 87:L-40, has discussed and defined the Phyllomedusa buckleyi
grouP.
Phyllomedusa aspera (Peters, 1872). Monatsber. Preuss. Akad. Wiss. Berlin, L872:772.
oRTGINAL NAME: Ilylomantis aspera.
rvnE(s): Syntypes: ZMB7507 (4 specimens).
TypE LocAury: Bahia,Brazil; probably Caravelas, Bahia, Brazil, according to
Bokermann, 1965, Lista Anot. Local. Tipo Anf. Brasil.:58.
Phyllomedusa ayeaye (8. Lutz, L966). Copeia, 1965:236.

oRTGTNAL NAME: Pithecopus ayeaye.
TypE LocALITy: "Morro do Ferro, a hill inside an ancient volcanic crater, at Pogos
de Caldas, Minas Gerais", Brazil'
DISTRIBUTIoN: Mountains of Minas Gerais, Brazil.
coMMENr: In the Phyllomedusa hypocondrialis grouP.
Phyllomedusa bahiana A. Lutz, 1925. C. R. S6ances Soc. Biol., Paris, 93:139.

TypE LocALITy: "Bahia",Brazil; probably Salvador, Bahia, Btazil,, according to
Bokermann, L956, Lista Anot. Local. Tipo Anf. Brasil.:82.
DrsrRrBUTroN: Central and eastern Brazil in the states of Bahia and Minas Gerais
coMMENr: In the Phyllomedusa burmeisteri gtoup.
Phyllomedusa baltea Duellman and Toft, 1979. Herpetologica, 35:64.

TypE LocALrrY: "Laguna, on the west slope of Serrania de Sira, 1280 m,
Departamento Hu6nuco, Peri".
DISTRIBUTIoN:Serrania de Sira, Peru.
coMMENr: In the Phyllomedusa perinesos grouP.
Phyllomedusa bicolor (Boddaert, L772). Epistola . . . de Rana bicolore:L9.

oRIGINAL NAME: Rana bicolor.
rvrr(s): Unknown (originally in collection of ]ohann Albert Schlosser).
TypE LOCArrry: Restricted to "Surinam" by Funkhouset, L957, Occas. Pap. Nat.
Hist. Mus. Stanford Univ., 5:38.
DrsrRrBUTrow: Amazon Basin in Brazil, Colombia, and Peru; the Guianan region of
Venezuela and the Guianas.
coMMENr: Reviewed by Duellman, 1974, Herpetologica, 30:L05-1L2.
Phyllomedusa boliaiana Boulenger, 1902.Ann. Mag. Nat. Hist. , (7)10:395-

rvnr(s): Syntypes: BM .2.22.32-33.
1947
TypE LocALITy: "Chulumani, Bolivia, 2000 metres".
DrsrRrBUTroN: Amazonian slopes of Andes and lowlands in Bolivia, northern
Argentina, and western Mato Grosso and R6ndonia, Brazil.
coMMENT: See Cannatella, 1983, Proc. Biol. Soc. Washington,96:59-66.
Phyllomedusa buckleyi Boulenger, 1882. Cat. Batr. SaI. Brit. Mus.:425.

rvru(s): Holotype: BM L947 .2.22.35.
TypE LocALITy: "Sar ay aclt, [Provincia P astaza,f Ecuador".
DrsrRrBUTroN: Upper Amazon Basin and lower Amazonian slopes of the Andes in
Ecuador.
coMMENr: In the Phyllomedusa buckleyi group.
Phyllomedusa burmeisteri Boulenger, 1882. Cat. Batr. Sal. Brit. Mus.:428.

rvnr(s): BM L947.2.22.22 designated lectotype by Funkhouser, L957, Occas. Pap.
Nat. Hist. Mus. Stanford Univ.,5:56.
TypE LocALITY: "Rio |aneiro", "Btazil", and "Oran Salta, Buenos Ayres"; restricted
to "the city of Rio de ]aneiro, specifically the Tijuca", Brazil, by Funkhouser,
L957, Occas. Pap. Nat. Hist. Mus. Stanford Univ., 5:56.
DISTRIBUTIoN: Central and southeastern Brazil.
coMMENr: In the Phyllomedusa burmeisteri group.
Phyllomedusa centralis Bokermann, 1965. Rev. Brasil. Biol., 25:258.

rvrr(s): Holotype: WCAB 15380.
TypE LocALITy: "Chapada dos Guimaries, Mato Grosso, Brasil".
coMMENr: In the Phyllomedusa hypocondrialis grouP.
Phyllome dusa co chr anae Bokermann, 19 66. Herpetolo gica, 22:293.

TypE LocAt-rry: "Campo de Fruticultura da Bocaina, Serra da Bocaina, Sio ]ose do
Barreiro, state of Sio Paulo , Brazil, collected at 1,600 m."
DISTRIBUTIoIS: Coastal mountains, Sio Paulo, Brazil.
coMMENT: In the Phyllomedusa guttata group; see Cannatella, 1980, Occas. Pap.
Phyllomedusa coelestrs (Cope, 1874). Proc. Acad. Nat. Sci. Philadelphia,26:12L.
oRIGINAL NAME: Pithecopus coelestis.
rvnr(s): Holotype: ANSP 11384; incorrectly reported as lost by Duellman, L974,
Herpetologica, 30: 105-LL2 (DCC).
TypE LocAt-rry: "Moyabamba [:Moyobamba], [Departamento San Martin,] Peru".
ANURA: HYLIDAE: PHYLLOMEDUSINAE 201
Phyllomedusa distincta B. Lutz, 1950. Mem. Inst. Oswaldo Cruz, Rio de )aneiro, 48:608
oRTGTNAL NAME: Phyllome dusa (P ithe copus) burmeist eri distinct a.
rYPE(s): Holotype: AL-MN 775.
TypE LocALITY: "Rio Vermelho", Santa Catarina, Brazil.
DISTRIBUTIoN: Southeastern Brazil (Sio Paulo and Santa Catarina).
coMMENT: In the Phyllomedusa burmeisteri group.
Phyllome dusa duellm ani Cannatella, L982. Copeia, 1982:502.

TypE LoCALITv: "8 km NNE (by road) Balzapata, Depto. Amazonas, Perti, L,850 m,
05" 47', 5,77" 48', W".
DrsrRrBUTroN: Known only from the type locality on Amazonian slopes of Andes
coMMENT: In the Phyllomedusa perinesos grouP.
P hyllome dusa e cu at orian a Cannatella, L982. Copeia, 1982:507 .
rYPE(s): Holotype: USNM 2L5750.

TypE LocAlrry: "Agua Rica, a one-house posada on the trail between Lim6n and
Gualeceo, slightty south and west of Lim6n, Provincia de Morona-Santiago,
Ecuador, L,890 m, 03o 02' 5,78" 27' W".
DrsrRrBUTroN: Known only from the type locality on the Amazonian slopes of the
Andes.
coMMENT: In the Phyllomedusa perinesos grouP.
Phyllomedusa exilis Cru2,1980. Rev. Brasil. Biol., a0(a):683.

rYPE(s): Holotype: EI 5584.
TypE LocALITy: "SantaTeteza, Espirito Santo, Brasil".
coMMENr: In the Phyllomedusa guttata group, according to the original description.
Phyllomedusa fimbriafa (Miranda-Ribeiro, 1923). Bo1. Mus. Nac., Rio de Janeiro, 1:4.
oRIGINAL NAME: Phrynomedusa fimbriata.
rYPE(s): Holotype: MZUSP 3L6.
TypE LocALITy: "Alto da Serra", Patanapiacaba, Sio Paulo, Btazil.
DrsrRrBUTroN: Serra do Mar from Rio de ]aneiro to Santa Catarina, southeastern
Brazil.
coMMENr: In the Phyllomedusa fimbriata group. See Cannatella, 1980, Occas. Pap.
Phyllomedusa guttata A. Lutz, L925 "L924". C. R. S6ances Soc. Biol., Paris, 90:241.
ivnE(s): Syntypes: AL-MN 297, USNM 96244-46 (Tijuca); AL-MN 449, USNM
963L7-L8 (Sumar6); USNM 96338 (Paineiras)'
TypE LOCALITv: "dans les montagnes, prds de Rio"; implicitly restricted to "the
Carioca mountain at Tijuca, Rio de faneiro", Brazil,by Lutz andLttz, 1939,
An. Acad. Brasil. Cienc., LL:219-263.
DrsrRrBUTroN: Coastal mountains from Rio de Janeiro to Sio Paulo, Brazil.
coMMENTt In the Phyllomedusa guttata group; see Cannatella, 1,980, Occas. Pap.
Phytlomedusa hypocondrialis (Daudin, 1802). Hist. Nat. Rain. Gren. Crap:Z9.

oRIGINAL NAME: Hyla hypocondrialis.
rYPE(s): Unknown.
South America east of the Andes from Bolivia, Colombia, and the
DrsrRrBUTroN:
Guianas southward to Argentina, Paraguay, and southeastern Brazil.
coMMENr: See Cei, 1980, Monit. Zool.Ital., N.S., Monogr., 2:437-439. In the
Phyllomedusa hypocondrialis group. The unjustified emendation hypochondrialis
has been widely used.
,lbt,
Phyllomedusa iheringii Boulenger, 1885. Ann. Mag. Nat. Hist., (5)16:88.

rvrr(s): Syntypes: BM 1947.2.22.26-3L, ZFMK 28560.
TypE LocAt-rry:"S. Lorenzo, on the southern border of the Lagoa dos Patos", Rio
Grande do SuI, Brazil.
DISTRIBUTIoN: Extreme southern Brazil (Rio Grande do Sul), Uruguay, and
Argentina (Misiones).
CoMMENT: Reviewed by Cei, 1980, Monit. Zool.Ital., N.S., Monogr.,2:435-436.In
the Phyllomedusa burmeisterl group.
Phyllomedusa jandaia Bokermann and Sazima, L978. Rev. Brasil. Biol., 38:927 .
TypE LocAt-rry: "Km L26 da estrada de Vespasiano a ConceiEio do Mato Dentro,
1.500 m, |aboticatubas, Minas Gerais, Brasil".
DrsrRrBUrroN: Type locality in the Serra do Espinhago, Minas Gerais, Brazil.
coMMENr: In the Phyllomedusa guttata group, according to the original description
Phyllomedusa lemur Boulenger, 1882. Cat. Batr. Sal. Brit. Mus.:425.

rvnr(s): Holotype: BM 1947.2.22.37.
TYPE LoCALITY:"Costa Rica".
DISTRIBUTIoN:Moderate elevations in Costa Rica and Panama.
coMMENr: In the Phyllomedusa buckleyl group.
Phyllomedusa marginata lzecksohn and Cruz, L976. Rev. Brasil. Biol., 36:257.
rvnr(s): Holotype: El 5L77.
TypE LocALITy: "SantaTereza, Estado do Espirito Santo, Brasil".
coMMENr: In the Phyllomedusa fimbriata group, according to the original
description; see Cannatella,1980, Occas. Pap. Mus. Nat. Hist. Univ. Kansas,
87:L-40.
Phyllomedusa medinae Funkhouser, 1962. Copeia, 1962:588.

rvnr(s): Holotype: EBRG (no number).
TypE LocAt.rry: "the biological station 'F{enry Pittie{", Estado Aragua, Venezuela
DISTRIBUTIoI{: Known only from the type locality.
Phyllomedusa palliata Peters, L872. Monatsber. Preuss. Akad. Wiss. Berlin L872:773
rvrr(s): Holotype: ZMB 7181.
TYPE LocALITY: "LJcayali", Peru.
DISTRIBUTIoN: Upper Amazon Basin in Ecuador and Peru.
coMMENT: Reviewed by Duellman, L974, Herpetologica, 30:105-112.
p erinesos Duellm an, L97 3. Herpetolo gica, 29 :226.

Phyllom e dusa
rvnr(s): Holotype: KU 1,46562.
TypE LocAt.rry: "Rio Salado, about 1 km upstream from the Rio Coca, Provincia
Napo, Ecuador, 1410 m".
DISTRIBUTIoN: Amazonian slopes of Andes in Napo Province, Ecuador.
coMMENT: In the Phyllomedusa perinesos group. See Cannatella, L982, Copeia, L98L
501-513.
Phyllomedusa psilopygion Cannatella, 1980. Occas. Pap. Mus. Nat. Hist. Univ. Kansas.
87:32.
TypE LocALITy: "8 km W Danubio, Rio AnchicayA, Departamento de Valle,
Colombia, 300 m, (03 37' N, 76o 47' W)".
DISTRIBUTIoN: Pacific lowlands of southern Colombia.
ANURA: HYPEROLIIDAE 203
Phyllomedusa rohdei Mertens, L926. Senckenb. Biol., 8:140.

TypE LocALITy: "Rio de ]aneiro, Brasilien".
DISTRIBUTIoN: Lowlands of southeastern Brazil (Rio de ]aneiro and Sio Paulo).
coMMENr: In the Phyllomedusa hypocondrialis group.
Phyllomedusa sauaagii Boulenger, L882. Cat. Batr. SaI. Brit. Mus.:429.

rvru(s): Syntypes: BM 1947 .2.25.83-84.
TypE LocALITy: "Oran Salta" and "Buenos Ayres", Argentina.
DISTRIBUTIoU: The Chacoan region of eastern Bolivia, northern Paraguay, Mato
Grosso do Sul (Brazil), and northern Argentina.
coMMENr: See Cei, L980, Monit. Zool.Ital., N.S., Monogr., 2:426-434.
Phyllomedusa tarsius (Cope, 1868). Proc. Acad. Nat. Sci. Philadelphia,20:113.

oRIGINAL NAME: Pithecopus tarsius.
rvnE(s): Formerly Smithsonian Museum (USNM) 6652; now lost.
TypE LocALITy: "Rio Napo, or Upper Amazon, below the mouth of the fotme{',
Departamento Loreto, Peru.
DISTRIBUTIoN: Amazon Basin in southern Colombia, Ecuador, and Peru.
coMMENr: Closely related to Phyllomedusa trinitatis and Phyllomedusa nenustt,
according to Duellman, 1974, Herpetologica, 30:L05-L12.
Phyllomedusa tomopterna (Cope, 1868). Proc. Acad. Nat. Sci. Philadelphia,20:LL2.

oRIGINAL NAME: Pithecopus tomopternus.
rvnr(s): Syntypes: Smithsonian Museum (USNM) 6651(2 specimens), now lost.
TypE LocALITy: "Rio Napo, or Upper Amazon, below the mouth of the forme{',
Departamento Loreto, Peru.
DISTRIBUTIoII: Upper Amazon Basin in Ecuador and Peru; Guianan region from
southeastern Venezuela to French Guiana; Brazilian Amazonia.
coMMENT: Reviewed by Duellman, L974, Herpetologica, 30:L05-114.
Phyllomedusa trinitatls Mertens, 1926. Senckenb. Biol., 8:145.

TypE LocALITy: "Port of Spain", Trinidad.
DISTRIBUTIoIt: Trinidad and the coastal mountains in northern Venezuela.
Phyllomedusa aaillanfi Boulenger, 1882. Cat. Batr. Sal. Brit. Mus.:427.
rvnr(s): Holotype: BM L947 .2.22.34.
TypE LocALITy: "Santarem, [Par6,] Brazil".
DISTRIBUTIoN: Guihnas and Amazon Basin from northeastern Brazil to Colombia,
Ecuador, Peru, and northern Bolivia.
coMMENT: See Duellman, 1974, Herpetologica, 30:105-LL2.
Phyllomedusa aenusta Duellman and Trueb, 1967. Copeia, L967:L28.

TypE LocALrry: "1km west-southwest of the junction of the Rio Mono and the Rio
Tuira, Dari6n Province, PanamA, elevation 130 m".
F.{MILv:Hyperoliidae Laurent, 1943.

cIrArIoN: 8u11. Mus. R. Hist. Nat. Belg., L9(28):L6.
DISTRIBUTIoN: Africa and nearby islands south of the Sahara; Madagascar; Seychelles Is.
in the Indian Ocean.
coMMENT: As first formed, the group name was Hyperoliinae. Laurent, 1951, Rev. ZooI.
Bot. Afr., 45:119, was the first to consider this group a distinct family, including the
Arthroleptinae and Astylosterninae in the Hyperoliidae, an arrangement considered

Frost, D. 1985. 110-203. Amphibian Lists of The World For The Hylidae Family.

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Frost, D. 1985. 110-203. Amphibian Lists of The World For The Hylidae Family.

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110 AMPHIBIAN SPECIES OF THE WORLD

Hemisus olioaceum Laurent, 1963. Copeia, 1963:398.

Hemisus wittei Laurent, 1963. Copeia, L963:397.

FAMILY: Hylidae Gray,1825 (1815).

adjacent Brazil and Argentina); Marinus S. Hoogmoed (MSH); Enrique La Marca

sUBFAMILv: Hemiphractinae Peters, L862.

Amphignathodon Boulenger, L882. Cat. Batr. Sal. Brit. Mus.:450.

Flectonotus Miranda-Ribeiro, L920. Rev. Mus. Paulista, Sio Paulo, 12:327.

CoMMENT: Genus redefined and reviewed by Duellman and Gray, L983,

Flectonotus fitzgeraldi (Parker, 1933). Trop. Agric., Trinidad, 11:123.

Flectonotus pygmaeus (Boettger, 1893). Ber. Senckenb. Naturforsch. Ges., 1893:40.

Fritziana Mello-Leitdo, L920. Rev. Mus. Paulista, Sio Paulo , L2:321,.

t'issilis (Miranda-Ribeiro, 1920).

Fritziana goeldii (Boulenger, 1895). Proc. Zool. Soc. London, L894:645.

Gastrotheca Fitzinger, L843. Syst. Rept.:30.

Gastrotheca andaquierzsis Ruiz and HernAndez, 1976. Caldasia, 11:151.

Gastrotheca angustifrors (Boulenger, 1898). Proc. Zool. Soc. London, L898:124.

Gastrotheca argenteoairens (Boettger, 1892). Ber. Senckenb. Naturforsch. Ges., L892:46.

Gastrotheca christiani Laurent, L967. Acta ZooL Lilloana, 22:354.

Gastrotheca chrysosticfa Laurent, L976. Acta Zool. Lilloana, 32:58.

Gastrotheca cornuta (Boulenger, 1898). Proc. ZooI. Soc. London,1898 124'

Gastrotheca dendronasfes Duellman, 1983. Copeia, 1983:868.

Gastrotheca fissipes (Boulenger, 1888). Ann. Mag. Nat. Hist., (6)2:42'

Gastrotheca galeata Trueb and Duellman, 1,978. Copeia, L978:498.

Gastrotheca gracilis Laurent, L969. Acta ZooL Lilloana, 25:146.

Gastrotheca helenaeDunn, 1944. Caldasia, 2:404.

Gastrotheca lojana Parker, 1932. Ann. Mag. Nat. Hist., (L0)9:25.

Gastrotheca longipes (Boulenger, 1882). Cat. Batr. Sal. Brit. Mus.:418.

Gastrotheca marsupiafa (Dum6ril and Bibron, 1841). Erp. G6n., 8:598.

Gastrotheca microdiscr.rs (Andersson, 1910). Ark. Zool., 6(9):9.

g9MMENT: In the Gastrotheca longipes grouP. See comments undet Gastrotheca

Gastrotheca monticola Barbour and Noble,1920. Bull. Mus. Comp. ZooI.,63:426.

Gastrotheca oaifera (Lichtenstein and Weinland, 1854). Monatsber. Preuss. Akad

Gastrotheca peruana (Boulenger, 1900). Ann. Mag. Nat. Hist., (7)6:18L.

Gastrotheca plumbea (Boulenger, 1882). Cat. Sal. Batr. Brit. Mus.:417.

coMMENT: In the Gastrotheca plumbea grouP.

Gastrotheca weinlandii (Steindachner, 1892). Sitzungsber. Akad. Wiss. Wien, Math.

coMMENT: In the Gastrotheca oaifera grouP.

Gastrotheca yacambuensisYrtztrs,lgT6. Rev. Univ. Centroccid. Lisandro Alvarado,

Hemiphractus W ag\er, L828. Isis von Oken, 21':7 43.

Hemiphractus bubalus(Jim6nez de la Espada, L87L).J. Sci. Math. Phys. Nat., Lisboa,

Hemiphractus proboscideus (lim6nez de Ia Espada, l87L). J. Sci. Math. Phys. Nat.,

Hemiphractus scutatus (Spix, L824). Spec. Nov. Testud. Ran. Brasil.:28.

Stefania Rivero, L968 "L966". Caribb. J. Sci., 6:L42.

Stefania eaansi (Boulenger, 1904). Proc. Zool. Soc. London, 1904:106.

Stefania ginesi Rivero, L968 "L966". Caribb. I. Sci., 6:L45.

Stefania goizri Rivero, 1968 "L966". Caribb. J. Sci., 6:145.

Stefania marahuaquensls (Riverc, L96L). BuIl. Mus. Comp. Zool., L26:LL8.

Stefania woodleyi Rivero, L968. Caribb. I. Sci., 6:147.

suBFAMILy: Hylinae Gray, 1825 (1815).

AcrisDum6ril and Bibron, 1841. Erp. G6n., 8:506.

Anotheca Smith, L939. Proc. Biol. Soc. Washington,52:190.

DISTRIBUTIoN:Southeastern Mexico (Atlantic slopes of Oaxaca and Veracruz), Costa

Aparasphenodon Miranda-Ribeiro, 1920. Rev. Mus. Paulista, Sio Paulo, 12:87.

Aparasphenodon aenezolanus (Mertens, 1950). Senckenb. Bio1., 3L:1.

Aplastodiscus perairidis A. Lut2,1950. In B. Lutz, Mem. Inst. Oswaldo Cruz, Rio de

Argenteohyla altamazonica Henle, 1981. Amphibia-Reptilia, 2:134.

Argenteohyla siemersi (Mertens, 1937). Senckenb. Biol., 19:12.

Calyptahyla crucialis (Harlan, L826). Am. J. Sci. Arts, L0:64.

TypE LocAlrry: "Brazil (exact locality unknown)"; catalogue in BM has locality

Hyla Laurenti, 1768. Synops. Rept.:32.

Hyla acreana Bokermann, L964. Senckenb. Biol., 45:244.

Hyla albicans Bokermann, L967. Neotropica, L3:64.